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<document id="F0365D59E2BF07C2CABEC3BD8154D7AF" ID-DOI="10.11646/zootaxa.3846.2.3" ID-GBIF-Dataset="f453051f-b811-4fec-9a94-ae0134550831" ID-ISSN="1175-5326" ID-Zenodo-Dep="250125" ID-ZooBank="B31B3F36-052A-477D-882B-8E555F74832A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460465051053" checkinUser="plazi" docAuthor="Nieves-Aldrey, J. L. &amp; Butterill, P. T." docDate="2014" docId="B776D1305E1C496CFF43FA90FD0FFBC7" docLanguage="en" docName="zt03846p234.pdf" docOrigin="Zootaxa 3846 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lithonecrus Nieves-Aldrey &amp; Butterill" docType="treatment" docVersion="8" lastPageNumber="224" masterDocId="4B4FA9485E1D496FFFD4FFC6FFE9FFF0" masterDocTitle="First evidence of cynipids from the Oceanian Region: the description of Lithonecrus papuanus a new genus and species of cynipid inquiline from Papua New Guinea (Hymenoptera: Cynipidae, Synergini)" masterLastPageNumber="234" masterPageNumber="221" pageNumber="222" updateTime="1698304805645" updateUser="plazi">
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<mods:title id="9F9C62AA0814266B4FDF96357209A297">First evidence of cynipids from the Oceanian Region: the description of Lithonecrus papuanus a new genus and species of cynipid inquiline from Papua New Guinea (Hymenoptera: Cynipidae, Synergini)</mods:title>
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<mods:namePart id="744902137C39B6E283794E8355615D07">Nieves-Aldrey, J. L.</mods:namePart>
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<mods:namePart id="8A6D86CF4C35C374346920AE1C47ECE6">Butterill, P. T.</mods:namePart>
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<mods:date id="3A5AB9633238502612C9CA363DCE282D">2014</mods:date>
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<mods:number id="E8C3CC697C37F7E15C4EC37BD255728F">3846</mods:number>
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<treatment id="B776D1305E1C496CFF43FA90FD0FFBC7" ID-DOI="http://doi.org/10.5281/zenodo.6143081" ID-GBIF-Taxon="119513922" ID-Zenodo-Dep="6143081" LSID="urn:lsid:plazi:treatment:B776D1305E1C496CFF43FA90FD0FFBC7" httpUri="http://treatment.plazi.org/id/B776D1305E1C496CFF43FA90FD0FFBC7" lastPageId="3" lastPageNumber="224" pageId="1" pageNumber="222">
<subSubSection id="77C533AD5E1C496EFF43FA90FEE3FA62" pageId="1" pageNumber="222" type="nomenclature">
<paragraph id="3F6060265E1C496EFF43FA90FD3AFA80" blockId="1.[151,723,1366,1426]" box="[151,723,1366,1392]" pageId="1" pageNumber="222">
<heading id="6428D74A5E1C496EFF43FA90FD3AFA80" bold="true" box="[151,723,1366,1392]" fontSize="11" level="1" pageId="1" pageNumber="222" reason="1">
<emphasis id="0DABBC345E1C496EFF43FA90FD3AFA80" bold="true" box="[151,723,1366,1392]" pageId="1" pageNumber="222">
<taxonomicName id="F8DF1BA55E1C496EFF43FA90FD9DFA80" authority="Nieves-Aldrey &amp; Butterill" authorityName="Nieves-Aldrey &amp; Butterill" box="[151,628,1366,1392]" class="Hexapoda" family="Cynipidae" genus="Lithonecrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="0DABBC345E1C496EFF43FA90FEC0FA80" bold="true" box="[151,297,1366,1392]" italics="true" pageId="1" pageNumber="222">Lithonecrus</emphasis>
Nieves-Aldrey &amp; Butterill
</taxonomicName>
, gen. n.
</emphasis>
</heading>
</paragraph>
<paragraph id="3F6060265E1C496EFF43FABFFEE3FA62" blockId="1.[151,723,1366,1426]" box="[151,266,1401,1426]" pageId="1" pageNumber="222">
(
<figureCitation id="A7E47CA35E1C496EFF4BFABFFEE8FA62" box="[159,257,1401,1426]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1877,1899]" captionStartId-1="5.[151,250,1950,1972]" captionStartId-2="6.[151,250,1787,1809]" captionStartId-3="7.[151,250,1940,1962]" captionStartId-4="9.[151,250,1927,1949]" captionTargetBox-0="[151,1436,195,1851]" captionTargetBox-1="[196,1400,195,1932]" captionTargetBox-2="[151,1436,193,1774]" captionTargetBox-3="[174,1417,447,1920]" captionTargetBox-4="[151,1436,375,1902]" captionTargetId-0="figure@2.[151,1436,195,1857]" captionTargetId-1="figure@5.[196,1401,195,1932]" captionTargetId-2="figure@6.[151,1436,193,1775]" captionTargetId-3="figure@7.[174,1418,447,1921]" captionTargetId-4="figure@9.[151,1436,375,1902]" captionTargetPageId-0="2" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="9" captionText-0="FIGURE 1. Lithonecrus papuanus sp. nov. (A) female, head anterior view; (B) detail of setae on face (C) male, head anterior view; (D) detail of setae on face; (E) magnification of a single seta (F) female, head posterior view." captionText-1="FIGURE 2. Lithonecrus papuanus sp. nov. (A) female antenna anterior view (B) female antenna posterior view; (C) detail of a plate of basiconic sensillae on F 10; (D) detail of last female antennal flagellomere showing four types of antennal sensillae (see text); (E) male antenna; (F G) details of the first male antennomeres." captionText-2="FIGURE 3. Lithonecrus papuanus sp. nov., female (A) head dorsal view; (B) mesosoma dorsal view; (C) mesosoma lateral view; (D) detail of mesopleuron, the arrow point the ventral margin of mesopleural triangle, angulated in posterior third; (E) pronotum frontal view (F) propodeum, the arrow point the crest limiting posteriorly the deep groove present dorsally on the lateral propodeal areas." captionText-3="FIGURE 4. Lithonecrus papuanus sp. nov., female. (A) metatarsal claw; (B) metasoma lateral view; (C) metasoma ventral view; (D) detail of hypopygium lateral view; (E) micropuncture sculpture on posterior part of last metasomal terga; (F) detail of the sculpture." captionText-4="FIGURE 5. Lithonecrus papuanus sp. nov., (A) female habitus; (B) male habitus; (C) female forewing." httpUri-0="https://zenodo.org/record/250126/files/figure.png" httpUri-1="https://zenodo.org/record/250127/files/figure.png" httpUri-2="https://zenodo.org/record/250128/files/figure.png" httpUri-3="https://zenodo.org/record/250129/files/figure.png" httpUri-4="https://zenodo.org/record/250130/files/figure.png" pageId="1" pageNumber="222">Figs 15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="77C533AD5E1C496EFF43FA07FA88FA2A" box="[151,1377,1473,1498]" pageId="1" pageNumber="222" type="materials_examined">
<paragraph id="3F6060265E1C496EFF43FA07FA88FA2A" blockId="1.[151,1437,1473,2002]" box="[151,1377,1473,1498]" pageId="1" pageNumber="222">
<emphasis id="0DABBC345E1C496EFF43FA07FEC2FA2A" bold="true" box="[151,299,1473,1498]" pageId="1" pageNumber="222">
<typeStatus id="E064DE845E1C496EFF43FA07FF3BFA2A" box="[151,210,1473,1498]" pageId="1" pageNumber="222">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="F8DF1BA55E1C496EFEEDFA07FC8BFA2A" authority="Nieves-Aldrey &amp; Butterill" authorityName="Nieves-Aldrey &amp; Butterill" box="[313,866,1473,1498]" class="Hexapoda" family="Cynipidae" genus="Lithonecrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="222" phylum="Arthropoda" rank="species" species="papuanus" status="sp. nov.">
<emphasis id="0DABBC345E1C496EFEEDFA07FDDBFA29" box="[313,562,1473,1498]" italics="true" pageId="1" pageNumber="222">Lithonecrus papuanus</emphasis>
Nieves-Aldrey &amp; Butterill
</taxonomicName>
,
<taxonomicNameLabel id="1698014F5E1C496EFCB9FA04FC43FA29" box="[877,938,1474,1497]" pageId="1" pageNumber="222" rank="species">sp. n.</taxonomicNameLabel>
, by present designation and monotype.
</paragraph>
</subSubSection>
<subSubSection id="77C533AD5E1C496EFF13FA23FD42F9D1" pageId="1" pageNumber="222" type="etymology">
<paragraph id="3F6060265E1C496EFF13FA23FD42F9D1" blockId="1.[151,1437,1473,2002]" pageId="1" pageNumber="222">
<emphasis id="0DABBC345E1C496EFF13FA23FEAFFA0E" bold="true" box="[199,326,1509,1534]" pageId="1" pageNumber="222">Etymology</emphasis>
. A combination of two words: “
<emphasis id="0DABBC345E1C496EFD6CFA23FCEAFA0E" box="[696,771,1509,1534]" italics="true" pageId="1" pageNumber="222">Litho”</emphasis>
from the host tree species
<taxonomicName id="F8DF1BA55E1C496EFBE2FA23FB54FA0E" box="[1078,1213,1509,1534]" class="Magnoliopsida" family="Fagaceae" genus="Lithocarpus" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="222" phylum="Tracheophyta" rank="genus">
<emphasis id="0DABBC345E1C496EFBE2FA23FB54FA0E" box="[1078,1213,1509,1534]" italics="true" pageId="1" pageNumber="222">Lithocarpus</emphasis>
</taxonomicName>
and “
<emphasis id="0DABBC345E1C496EFAD7FA20FAB4FA0D" box="[1283,1373,1510,1533]" italics="true" pageId="1" pageNumber="222">necrus”</emphasis>
from the end word of the related genus
<taxonomicName id="F8DF1BA55E1C496EFDC7F9CFFD4DF9D2" box="[531,676,1545,1570]" class="Insecta" family="Cynipidae" genus="Saphonecrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="0DABBC345E1C496EFDC7F9CFFD4DF9D2" box="[531,676,1545,1570]" italics="true" pageId="1" pageNumber="222">Saphonecrus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="77C533AD5E1C496CFF43F9EBFD0FFBC7" lastPageId="3" lastPageNumber="224" pageId="1" pageNumber="222" type="description">
<paragraph id="3F6060265E1C496EFF43F9EBFEC8F9B6" blockId="1.[151,1437,1473,2002]" box="[151,289,1581,1606]" pageId="1" pageNumber="222">
<emphasis id="0DABBC345E1C496EFF43F9EBFEC8F9B6" bold="true" box="[151,289,1581,1606]" pageId="1" pageNumber="222">Description</emphasis>
</paragraph>
<paragraph id="3F6060265E1C496EFF13F997FB85F822" blockId="1.[151,1437,1473,2002]" pageId="1" pageNumber="222">
<emphasis id="0DABBC345E1C496EFF13F997FEEDF99A" box="[199,260,1617,1642]" italics="true" pageId="1" pageNumber="222">Head</emphasis>
. Rounded in females (
<figureCitation id="A7E47CA35E1C496EFDD2F997FDACF99A" box="[518,581,1617,1642]" captionStart="FIGURE 1" captionStartId="2.[151,250,1877,1899]" captionTargetBox="[151,1436,195,1851]" captionTargetId="figure@2.[151,1436,195,1857]" captionTargetPageId="2" captionText="FIGURE 1. Lithonecrus papuanus sp. nov. (A) female, head anterior view; (B) detail of setae on face (C) male, head anterior view; (D) detail of setae on face; (E) magnification of a single seta (F) female, head posterior view." httpUri="https://zenodo.org/record/250126/files/figure.png" pageId="1" pageNumber="222">Fig. 1</figureCitation>
A), oval in males (
<figureCitation id="A7E47CA35E1C496EFCCAF997FCB6F99A" box="[798,863,1617,1642]" captionStart="FIGURE 1" captionStartId="2.[151,250,1877,1899]" captionTargetBox="[151,1436,195,1851]" captionTargetId="figure@2.[151,1436,195,1857]" captionTargetPageId="2" captionText="FIGURE 1. Lithonecrus papuanus sp. nov. (A) female, head anterior view; (B) detail of setae on face (C) male, head anterior view; (D) detail of setae on face; (E) magnification of a single seta (F) female, head posterior view." httpUri="https://zenodo.org/record/250126/files/figure.png" pageId="1" pageNumber="222">Fig. 1</figureCitation>
C). Slightly pubescent in females, more heavily in males; scattered long setae on the face medially, the lateral areas, and upper part of frons; in males the setation is dense and whitish, the setae being differentiated, longer and broader basally (
<figureCitation id="A7E47CA35E1C496EFBC9F95FFB8DF942" box="[1053,1124,1689,1714]" captionStart="FIGURE 1" captionStartId="2.[151,250,1877,1899]" captionTargetBox="[151,1436,195,1851]" captionTargetId="figure@2.[151,1436,195,1857]" captionTargetPageId="2" captionText="FIGURE 1. Lithonecrus papuanus sp. nov. (A) female, head anterior view; (B) detail of setae on face (C) male, head anterior view; (D) detail of setae on face; (E) magnification of a single seta (F) female, head posterior view." httpUri="https://zenodo.org/record/250126/files/figure.png" pageId="1" pageNumber="222">Figs 1</figureCitation>
C, 1D, 1E). Gena slightly expanded behind compound eyes. Clypeus indistinct, ventral margin sinuate, not projecting over mandibles. Anterior tentorial pits visible; epistomal sulcus and clypeo-pleurostomal lines indistinct. Face with wide, strong and blunt irradiating striae from clypeus, almost reaching ventral margin of eye and ventral margin of toruli; The facial striae extend dorsally branched in several rows on the lateral area of the frons, reaching the lateral ocelli, but absent medially (
<figureCitation id="A7E47CA35E1C496EFE89F88BFE76F896" box="[349,415,1869,1894]" captionStart="FIGURE 1" captionStartId="2.[151,250,1877,1899]" captionTargetBox="[151,1436,195,1851]" captionTargetId="figure@2.[151,1436,195,1857]" captionTargetPageId="2" captionText="FIGURE 1. Lithonecrus papuanus sp. nov. (A) female, head anterior view; (B) detail of setae on face (C) male, head anterior view; (D) detail of setae on face; (E) magnification of a single seta (F) female, head posterior view." httpUri="https://zenodo.org/record/250126/files/figure.png" pageId="1" pageNumber="222">Fig. 1</figureCitation>
A). Vertex abruptly limited at the occiput by an occipital carina situated just behind the lateral ocelli (
<figureCitation id="A7E47CA35E1C496EFEE5F8B7FE9DF87A" box="[305,372,1905,1930]" captionStart="FIGURE 3" captionStartId="6.[151,250,1787,1809]" captionTargetBox="[151,1436,193,1774]" captionTargetId="figure@6.[151,1436,193,1775]" captionTargetPageId="6" captionText="FIGURE 3. Lithonecrus papuanus sp. nov., female (A) head dorsal view; (B) mesosoma dorsal view; (C) mesosoma lateral view; (D) detail of mesopleuron, the arrow point the ventral margin of mesopleural triangle, angulated in posterior third; (E) pronotum frontal view (F) propodeum, the arrow point the crest limiting posteriorly the deep groove present dorsally on the lateral propodeal areas." httpUri="https://zenodo.org/record/250128/files/figure.png" pageId="1" pageNumber="222">Fig. 3</figureCitation>
A); the occipital carina is well defined and complete, reaching ventral margin of face (
<figureCitation id="A7E47CA35E1C496EFAEEF8B7FA90F87A" box="[1338,1401,1905,1930]" captionStart="FIGURE 1" captionStartId="2.[151,250,1877,1899]" captionTargetBox="[151,1436,195,1851]" captionTargetId="figure@2.[151,1436,195,1857]" captionTargetPageId="2" captionText="FIGURE 1. Lithonecrus papuanus sp. nov. (A) female, head anterior view; (B) detail of setae on face (C) male, head anterior view; (D) detail of setae on face; (E) magnification of a single seta (F) female, head posterior view." httpUri="https://zenodo.org/record/250126/files/figure.png" pageId="1" pageNumber="222">Fig. 1</figureCitation>
F). Occiput coriaceous dorsally. Gula relatively long; distance between occipital foramen and oral foramen clearly longer than the height of the occipital foramen. Hypostomal sulci well visible (
<figureCitation id="A7E47CA35E1C496EFBDEF87FFBA3F822" box="[1034,1098,1977,2002]" captionStart="FIGURE 1" captionStartId="2.[151,250,1877,1899]" captionTargetBox="[151,1436,195,1851]" captionTargetId="figure@2.[151,1436,195,1857]" captionTargetPageId="2" captionText="FIGURE 1. Lithonecrus papuanus sp. nov. (A) female, head anterior view; (B) detail of setae on face (C) male, head anterior view; (D) detail of setae on face; (E) magnification of a single seta (F) female, head posterior view." httpUri="https://zenodo.org/record/250126/files/figure.png" pageId="1" pageNumber="222">Fig. 1</figureCitation>
F).
</paragraph>
<caption id="6BA030AE5E1F496DFF43F893FB6DF87A" httpUri="https://zenodo.org/record/250126/files/figure.png" pageId="2" pageNumber="223" targetBox="[151,1436,195,1851]" targetPageId="2">
<paragraph id="3F6060265E1F496DFF43F893FB6DF87A" blockId="2.[151,1436,1877,1930]" pageId="2" pageNumber="223">
<emphasis id="0DABBC345E1F496DFF43F893FEE4F89A" bold="true" box="[151,269,1877,1899]" pageId="2" pageNumber="223">FIGURE 1</emphasis>
.
<taxonomicName id="F8DF1BA55E1F496DFECEF893FE14F89B" box="[282,509,1877,1899]" class="Hexapoda" family="Cynipidae" genus="Lithonecrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="223" phylum="Arthropoda" rank="species" species="papuanus" status="sp. nov.">
<emphasis id="0DABBC345E1F496DFECEF893FE14F89B" box="[282,509,1877,1899]" italics="true" pageId="2" pageNumber="223">Lithonecrus papuanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1698014F5E1F496DFDD7F890FDBAF89B" box="[515,595,1878,1899]" pageId="2" pageNumber="223" rank="species">sp. nov.</taxonomicNameLabel>
(A) female, head anterior view; (B) detail of setae on face (C) male, head anterior view; (D) detail of setae on face; (E) magnification of a single seta (F) female, head posterior view.
</paragraph>
</caption>
<paragraph id="3F6060265E1F496CFF13F870FBDBFF08" blockId="2.[151,1436,1974,2035]" lastBlockId="3.[151,1437,151,1079]" lastPageId="3" lastPageNumber="224" pageId="2" pageNumber="223">
Female antenna 13-segmented (
<figureCitation id="A7E47CA35E1F496DFDE3F870FD6CF83F" box="[567,645,1974,1999]" captionStart="FIGURE 2" captionStartId="5.[151,250,1950,1972]" captionTargetBox="[196,1400,195,1932]" captionTargetId="figure@5.[196,1401,195,1932]" captionTargetPageId="5" captionText="FIGURE 2. Lithonecrus papuanus sp. nov. (A) female antenna anterior view (B) female antenna posterior view; (C) detail of a plate of basiconic sensillae on F 10; (D) detail of last female antennal flagellomere showing four types of antennal sensillae (see text); (E) male antenna; (F G) details of the first male antennomeres." httpUri="https://zenodo.org/record/250127/files/figure.png" pageId="2" pageNumber="223">Figs. 2</figureCitation>
A, 2B); flagellum broadened towards apex; with relatively long, erect setae and placodeal sensilla visible on flagellar segments F6F11. Last three flagellar segments presenting wide areas with basiconic setae (
<figureCitation id="A7E47CA35E1E496CFE18FF51FDFCFF40" box="[460,533,151,176]" captionStart="FIGURE 2" captionStartId="5.[151,250,1950,1972]" captionTargetBox="[196,1400,195,1932]" captionTargetId="figure@5.[196,1401,195,1932]" captionTargetPageId="5" captionText="FIGURE 2. Lithonecrus papuanus sp. nov. (A) female antenna anterior view (B) female antenna posterior view; (C) detail of a plate of basiconic sensillae on F 10; (D) detail of last female antennal flagellomere showing four types of antennal sensillae (see text); (E) male antenna; (F G) details of the first male antennomeres." httpUri="https://zenodo.org/record/250127/files/figure.png" pageId="3" pageNumber="224">Figs 2</figureCitation>
B, 2C). Pedicel 1.8 as long as wide, broader but shorter (0.7) than F1; F1 1.7 as long as F2 (
<figureCitation id="A7E47CA35E1E496CFEF2FF7AFE84FF24" box="[294,365,188,213]" captionStart="FIGURE 2" captionStartId="5.[151,250,1950,1972]" captionTargetBox="[196,1400,195,1932]" captionTargetId="figure@5.[196,1401,195,1932]" captionTargetPageId="5" captionText="FIGURE 2. Lithonecrus papuanus sp. nov. (A) female antenna anterior view (B) female antenna posterior view; (C) detail of a plate of basiconic sensillae on F 10; (D) detail of last female antennal flagellomere showing four types of antennal sensillae (see text); (E) male antenna; (F G) details of the first male antennomeres." httpUri="https://zenodo.org/record/250127/files/figure.png" pageId="3" pageNumber="224">Fig. 2</figureCitation>
A). Male antenna 15-segmented, not broadened towards apex (
<figureCitation id="A7E47CA35E1E496CFB9EFF7AFB78FF24" box="[1098,1169,188,213]" captionStart="FIGURE 2" captionStartId="5.[151,250,1950,1972]" captionTargetBox="[196,1400,195,1932]" captionTargetId="figure@5.[196,1401,195,1932]" captionTargetPageId="5" captionText="FIGURE 2. Lithonecrus papuanus sp. nov. (A) female antenna anterior view (B) female antenna posterior view; (C) detail of a plate of basiconic sensillae on F 10; (D) detail of last female antennal flagellomere showing four types of antennal sensillae (see text); (E) male antenna; (F G) details of the first male antennomeres." httpUri="https://zenodo.org/record/250127/files/figure.png" pageId="3" pageNumber="224">Fig. 2</figureCitation>
E). F1 dorsally curved, excavated in the middle and expanded apically (
<figureCitation id="A7E47CA35E1E496CFD66FF19FD11FF08" box="[690,760,223,248]" captionStart="FIGURE 2" captionStartId="5.[151,250,1950,1972]" captionTargetBox="[196,1400,195,1932]" captionTargetId="figure@5.[196,1401,195,1932]" captionTargetPageId="5" captionText="FIGURE 2. Lithonecrus papuanus sp. nov. (A) female antenna anterior view (B) female antenna posterior view; (C) detail of a plate of basiconic sensillae on F 10; (D) detail of last female antennal flagellomere showing four types of antennal sensillae (see text); (E) male antenna; (F G) details of the first male antennomeres." httpUri="https://zenodo.org/record/250127/files/figure.png" pageId="3" pageNumber="224">Figs 2</figureCitation>
F, 2G). F1 more than
<date id="4B6146E65E1E496CFC3EFF26FBEFFF08" box="[1002,1030,224,248]" pageId="3" pageNumber="224">2x</date>
F2.
</paragraph>
<paragraph id="3F6060265E1E496CFF13FEC3FB22FDCC" blockId="3.[151,1437,151,1079]" pageId="3" pageNumber="224">
<emphasis id="0DABBC345E1E496CFF13FEC3FED6FEEC" box="[199,319,261,284]" italics="true" pageId="3" pageNumber="224">Mesosoma</emphasis>
. Pronotum relatively broad medially, measuring nearly 1/5 of the length of the outer lateral margin. Pronotal plate (
<figureCitation id="A7E47CA35E1E496CFE95FEE1FE6BFEB0" box="[321,386,295,320]" captionStart="FIGURE 3" captionStartId="6.[151,250,1787,1809]" captionTargetBox="[151,1436,193,1774]" captionTargetId="figure@6.[151,1436,193,1775]" captionTargetPageId="6" captionText="FIGURE 3. Lithonecrus papuanus sp. nov., female (A) head dorsal view; (B) mesosoma dorsal view; (C) mesosoma lateral view; (D) detail of mesopleuron, the arrow point the ventral margin of mesopleural triangle, angulated in posterior third; (E) pronotum frontal view (F) propodeum, the arrow point the crest limiting posteriorly the deep groove present dorsally on the lateral propodeal areas." httpUri="https://zenodo.org/record/250128/files/figure.png" pageId="3" pageNumber="224">Fig. 3</figureCitation>
E) relatively well developed, with lateral margins present posteriorly and only faint anteriorly; posterior part of pronotal plate with antero-lateral margins quite well extended to the sides of pronotum. Lateral margins of pronotum rounded, without lateral pronotal carinae (
<figureCitation id="A7E47CA35E1E496CFCAAFEA9FC2DFE78" box="[894,964,367,392]" captionStart="FIGURE 3" captionStartId="6.[151,250,1787,1809]" captionTargetBox="[151,1436,193,1774]" captionTargetId="figure@6.[151,1436,193,1775]" captionTargetPageId="6" captionText="FIGURE 3. Lithonecrus papuanus sp. nov., female (A) head dorsal view; (B) mesosoma dorsal view; (C) mesosoma lateral view; (D) detail of mesopleuron, the arrow point the ventral margin of mesopleural triangle, angulated in posterior third; (E) pronotum frontal view (F) propodeum, the arrow point the crest limiting posteriorly the deep groove present dorsally on the lateral propodeal areas." httpUri="https://zenodo.org/record/250128/files/figure.png" pageId="3" pageNumber="224">Fig. 3</figureCitation>
C). In lateral view, pronotum very short. Mesoscutum with notauli broad, percurrent (
<figureCitation id="A7E47CA35E1E496CFD43FE52FD32FE5C" box="[663,731,404,429]" captionStart="FIGURE 3" captionStartId="6.[151,250,1787,1809]" captionTargetBox="[151,1436,193,1774]" captionTargetId="figure@6.[151,1436,193,1775]" captionTargetPageId="6" captionText="FIGURE 3. Lithonecrus papuanus sp. nov., female (A) head dorsal view; (B) mesosoma dorsal view; (C) mesosoma lateral view; (D) detail of mesopleuron, the arrow point the ventral margin of mesopleural triangle, angulated in posterior third; (E) pronotum frontal view (F) propodeum, the arrow point the crest limiting posteriorly the deep groove present dorsally on the lateral propodeal areas." httpUri="https://zenodo.org/record/250128/files/figure.png" pageId="3" pageNumber="224">Fig. 3</figureCitation>
B). Median mesoscutal impression invisible. Anteroadmedian signae visible. Parapsidal signa deep, reaching posterior margin of pronotum. Transscutal fissure narrow. Scutellar foveae large, obliquely separated, with smooth sculpture but crossed by some strong carinae (
<figureCitation id="A7E47CA35E1E496CFAF9FE1AFA92FE04" box="[1325,1403,476,501]" captionStart="FIGURE 2" captionStartId="5.[151,250,1950,1972]" captionTargetBox="[196,1400,195,1932]" captionTargetId="figure@5.[196,1401,195,1932]" captionTargetPageId="5" captionText="FIGURE 2. Lithonecrus papuanus sp. nov. (A) female antenna anterior view (B) female antenna posterior view; (C) detail of a plate of basiconic sensillae on F 10; (D) detail of last female antennal flagellomere showing four types of antennal sensillae (see text); (E) male antenna; (F G) details of the first male antennomeres." httpUri="https://zenodo.org/record/250127/files/figure.png" pageId="3" pageNumber="224">Fig. 2</figureCitation>
B). Mesopleural triangle subrectangular, ventral margin not straight and angulated in the posterior third (
<figureCitation id="A7E47CA35E1E496CFAD6FE39FAA0FDE8" box="[1282,1353,511,536]" captionStart="FIGURE 3" captionStartId="6.[151,250,1787,1809]" captionTargetBox="[151,1436,193,1774]" captionTargetId="figure@6.[151,1436,193,1775]" captionTargetPageId="6" captionText="FIGURE 3. Lithonecrus papuanus sp. nov., female (A) head dorsal view; (B) mesosoma dorsal view; (C) mesosoma lateral view; (D) detail of mesopleuron, the arrow point the ventral margin of mesopleural triangle, angulated in posterior third; (E) pronotum frontal view (F) propodeum, the arrow point the crest limiting posteriorly the deep groove present dorsally on the lateral propodeal areas." httpUri="https://zenodo.org/record/250128/files/figure.png" pageId="3" pageNumber="224">Figs 3</figureCitation>
C, 3D). Mesopleuron with longitudinal striae and coriaceous sculpture between the striae (
<figureCitation id="A7E47CA35E1E496CFBE5FDE2FB91FDCC" box="[1073,1144,548,573]" captionStart="FIGURE 3" captionStartId="6.[151,250,1787,1809]" captionTargetBox="[151,1436,193,1774]" captionTargetId="figure@6.[151,1436,193,1775]" captionTargetPageId="6" captionText="FIGURE 3. Lithonecrus papuanus sp. nov., female (A) head dorsal view; (B) mesosoma dorsal view; (C) mesosoma lateral view; (D) detail of mesopleuron, the arrow point the ventral margin of mesopleural triangle, angulated in posterior third; (E) pronotum frontal view (F) propodeum, the arrow point the crest limiting posteriorly the deep groove present dorsally on the lateral propodeal areas." httpUri="https://zenodo.org/record/250128/files/figure.png" pageId="3" pageNumber="224">Figs 3</figureCitation>
C, 3D).
</paragraph>
<paragraph id="3F6060265E1E496CFF13FD81FBEEFD00" blockId="3.[151,1437,151,1079]" pageId="3" pageNumber="224">
<emphasis id="0DABBC345E1E496CFF13FD81FDDBFD90" box="[199,562,583,608]" italics="true" pageId="3" pageNumber="224">Metapectal-propodeal complex.</emphasis>
Metapleural sulcus meeting posterior margin of mesopectus at about the height of posterior subalar pit. Lateral propodeal carinae distinct, broad, slightly convergent posteriorly (
<figureCitation id="A7E47CA35E1E496CFAEDFDAAFA95FD74" box="[1337,1404,620,645]" captionStart="FIGURE 3" captionStartId="6.[151,250,1787,1809]" captionTargetBox="[151,1436,193,1774]" captionTargetId="figure@6.[151,1436,193,1775]" captionTargetPageId="6" captionText="FIGURE 3. Lithonecrus papuanus sp. nov., female (A) head dorsal view; (B) mesosoma dorsal view; (C) mesosoma lateral view; (D) detail of mesopleuron, the arrow point the ventral margin of mesopleural triangle, angulated in posterior third; (E) pronotum frontal view (F) propodeum, the arrow point the crest limiting posteriorly the deep groove present dorsally on the lateral propodeal areas." httpUri="https://zenodo.org/record/250128/files/figure.png" pageId="3" pageNumber="224">Fig. 3</figureCitation>
F). Median propodeal areas smooth and pubescent. Dorsally, on the lateral propodeal areas, a deep groove is present, extending from the propodeal spiracles to the dorsal part of the lateral propodeal carinae, the groove being limited posteriorly by a crest (
<figureCitation id="A7E47CA35E1E496CFE46FD11FE3DFD00" box="[402,468,727,752]" captionStart="FIGURE 3" captionStartId="6.[151,250,1787,1809]" captionTargetBox="[151,1436,193,1774]" captionTargetId="figure@6.[151,1436,193,1775]" captionTargetPageId="6" captionText="FIGURE 3. Lithonecrus papuanus sp. nov., female (A) head dorsal view; (B) mesosoma dorsal view; (C) mesosoma lateral view; (D) detail of mesopleuron, the arrow point the ventral margin of mesopleural triangle, angulated in posterior third; (E) pronotum frontal view (F) propodeum, the arrow point the crest limiting posteriorly the deep groove present dorsally on the lateral propodeal areas." httpUri="https://zenodo.org/record/250128/files/figure.png" pageId="3" pageNumber="224">Fig. 3</figureCitation>
F). Nucha dorsally strongly sulcate longitudinally.
</paragraph>
<paragraph id="3F6060265E1E496CFF13FD3BFBE9FCE4" blockId="3.[151,1437,151,1079]" box="[199,1024,764,789]" pageId="3" pageNumber="224">
<emphasis id="0DABBC345E1E496CFF13FD3BFEEAFCE4" box="[199,259,765,788]" italics="true" pageId="3" pageNumber="224">Legs.</emphasis>
Metatarsal claw simple, without basal acute lobe or tooth (
<figureCitation id="A7E47CA35E1E496CFC4DFD3AFC35FCE4" box="[921,988,764,789]" captionStart="FIGURE 4" captionStartId="7.[151,250,1940,1962]" captionTargetBox="[174,1417,447,1920]" captionTargetId="figure@7.[174,1418,447,1921]" captionTargetPageId="7" captionText="FIGURE 4. Lithonecrus papuanus sp. nov., female. (A) metatarsal claw; (B) metasoma lateral view; (C) metasoma ventral view; (D) detail of hypopygium lateral view; (E) micropuncture sculpture on posterior part of last metasomal terga; (F) detail of the sculpture." httpUri="https://zenodo.org/record/250129/files/figure.png" pageId="3" pageNumber="224">Fig. 4</figureCitation>
A).
</paragraph>
<paragraph id="3F6060265E1E496CFF13FCE7FBF5FC57" blockId="3.[151,1437,151,1079]" pageId="3" pageNumber="224">
<emphasis id="0DABBC345E1E496CFF13FCE7FEDAFCC8" box="[199,307,801,824]" italics="true" pageId="3" pageNumber="224">Forewing</emphasis>
. Radial cell open along anterior margin; R1 and Rs stopping close to the anterior margin of wing (
<figureCitation id="A7E47CA35E1E496CFF4BFC80FF0CFCAF" box="[159,229,838,863]" captionStart="FIGURE 5" captionStartId="9.[151,250,1927,1949]" captionTargetBox="[151,1436,375,1902]" captionTargetId="figure@9.[151,1436,375,1902]" captionTargetPageId="9" captionText="FIGURE 5. Lithonecrus papuanus sp. nov., (A) female habitus; (B) male habitus; (C) female forewing." httpUri="https://zenodo.org/record/250130/files/figure.png" pageId="3" pageNumber="224">Fig. 5</figureCitation>
C); R1 forming roughly a 90o angle with anterior margin of wing and R1+Sc; Medial and cubital veins virtually invisible; areolet not present. When visible, the M+Cu1 vein is situated rather higher than apical part of cubital vein (Cu1a). Apical margin of wing with a fringe of long setae (
<figureCitation id="A7E47CA35E1E496CFC62FC48FC10FC57" box="[950,1017,910,935]" captionStart="FIGURE 5" captionStartId="9.[151,250,1927,1949]" captionTargetBox="[151,1436,375,1902]" captionTargetId="figure@9.[151,1436,375,1902]" captionTargetPageId="9" captionText="FIGURE 5. Lithonecrus papuanus sp. nov., (A) female habitus; (B) male habitus; (C) female forewing." httpUri="https://zenodo.org/record/250130/files/figure.png" pageId="3" pageNumber="224">Fig. 5</figureCitation>
C).
</paragraph>
<paragraph id="3F6060265E1E496CFF13FC72FD0FFBC7" blockId="3.[151,1437,151,1079]" pageId="3" pageNumber="224">
<emphasis id="0DABBC345E1E496CFF13FC72FED5FC3B" box="[199,316,948,971]" italics="true" pageId="3" pageNumber="224">Metasoma</emphasis>
(
<figureCitation id="A7E47CA35E1E496CFE9BFC74FE7EFC3B" box="[335,407,946,971]" captionStart="FIGURE 4" captionStartId="7.[151,250,1940,1962]" captionTargetBox="[174,1417,447,1920]" captionTargetId="figure@7.[174,1418,447,1921]" captionTargetPageId="7" captionText="FIGURE 4. Lithonecrus papuanus sp. nov., female. (A) metatarsal claw; (B) metasoma lateral view; (C) metasoma ventral view; (D) detail of hypopygium lateral view; (E) micropuncture sculpture on posterior part of last metasomal terga; (F) detail of the sculpture." httpUri="https://zenodo.org/record/250129/files/figure.png" pageId="3" pageNumber="224">Fig. 4</figureCitation>
B). T1 only half ring shaped dorsally; strongly sulcate. T2+3 covering almost the entire metasoma; smooth and shining, except for a narrow posterior band with micropunctures present (
<figureCitation id="A7E47CA35E1E496CFAD6FC10FAA4FC1F" box="[1282,1357,982,1007]" captionStart="FIGURE 4" captionStartId="7.[151,250,1940,1962]" captionTargetBox="[174,1417,447,1920]" captionTargetId="figure@7.[174,1418,447,1921]" captionTargetPageId="7" captionText="FIGURE 4. Lithonecrus papuanus sp. nov., female. (A) metatarsal claw; (B) metasoma lateral view; (C) metasoma ventral view; (D) detail of hypopygium lateral view; (E) micropuncture sculpture on posterior part of last metasomal terga; (F) detail of the sculpture." httpUri="https://zenodo.org/record/250129/files/figure.png" pageId="3" pageNumber="224">Figs 4</figureCitation>
E, 4F). Projecting part of hypopygial spine short, 1.5 times as long as high in profile (
<figureCitation id="A7E47CA35E1E496CFBFAFC3CFB90FBE3" box="[1070,1145,1018,1043]" captionStart="FIGURE 4" captionStartId="7.[151,250,1940,1962]" captionTargetBox="[174,1417,447,1920]" captionTargetId="figure@7.[174,1418,447,1921]" captionTargetPageId="7" captionText="FIGURE 4. Lithonecrus papuanus sp. nov., female. (A) metatarsal claw; (B) metasoma lateral view; (C) metasoma ventral view; (D) detail of hypopygium lateral view; (E) micropuncture sculpture on posterior part of last metasomal terga; (F) detail of the sculpture." httpUri="https://zenodo.org/record/250129/files/figure.png" pageId="3" pageNumber="224">Figs 4</figureCitation>
C, 4D); hypopygial spine ventrally with two widely spaced rows of long setae.
</paragraph>
</subSubSection>
</treatment>
</document>