treatments-xml/data/F5/42/AA/F542AA7FFFD8FFE400C784413F2BFC32.xml
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<document id="66C00AA294D23D319DABC8D4AE248DD3" ID-CLB-Dataset="27612" ID-DOI="10.11646/zootaxa.4551.1.5" ID-GBIF-Dataset="7f5b691a-88a8-471f-a777-d19bb03ba162" ID-ISSN="1175-5326" ID-Zenodo-Dep="2622617" ID-ZooBank="A5D621EC-1FD3-4565-BA9F-B7B2BF837620" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554269426536" checkinUser="plazi" docAuthor="Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H. &amp; Mohammad, Salwa K." docDate="2019" docId="F542AA7FFFD8FFE400C784413F2BFC32" docLanguage="en" docName="zootaxa.4551.1.5.pdf" docOrigin="Zootaxa 4551 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atylotus shagrensis Abu El-Hassan &amp; Badrawy &amp; Fadl &amp; Mohammad 2019, sp. nov." docType="treatment" docVersion="9" lastPageNumber="85" masterDocId="097BD207FFDDFFE2005082023A5DFFBB" masterDocTitle="Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species" masterLastPageNumber="86" masterPageNumber="79" pageNumber="84" updateTime="1698716044556" updateUser="ExternalLinkService">
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<mods:title id="2F06DC32907F1980AE20FB16595382E8">Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species</mods:title>
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<mods:namePart id="E584A423719CB69E999E356BE0FBFAE5">Abu El-Hassan, Gawhara M. M.</mods:namePart>
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<mods:namePart id="2DF6B3FB03B497D0D1A647A4F1F3E666">Badrawy, Haitham B. M.</mods:namePart>
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<mods:namePart id="EE7241D81D89F6D3A6D8680E6383B0DD">Fadl, Hassan H.</mods:namePart>
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<mods:namePart id="632C015C7C422CCC59792C9334527EC1">Mohammad, Salwa K.</mods:namePart>
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<mods:date id="C490AF938A488B60BB6B4A4037825F72">2019</mods:date>
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<treatment id="F542AA7FFFD8FFE400C784413F2BFC32" ID-DOI="http://doi.org/10.5281/zenodo.5923771" ID-GBIF-Taxon="154532843" ID-Zenodo-Dep="5923771" LSID="urn:lsid:plazi:treatment:F542AA7FFFD8FFE400C784413F2BFC32" httpUri="http://treatment.plazi.org/id/F542AA7FFFD8FFE400C784413F2BFC32" lastPageId="6" lastPageNumber="85" pageId="5" pageNumber="84">
<subSubSection id="35F148E2FFD8FFE700C784413B56F9C5" pageId="5" pageNumber="84" type="nomenclature">
<paragraph id="7D541B69FFD8FFE700C784413BBAF9E6" blockId="5.[151,487,1603,1662]" box="[151,487,1603,1629]" pageId="5" pageNumber="84">
<heading id="261CAC05FFD8FFE700C784413BBAF9E6" bold="true" box="[151,487,1603,1629]" fontSize="11" level="1" pageId="5" pageNumber="84" reason="1">
<emphasis id="4F9FC77BFFD8FFE700C784413BBAF9E6" bold="true" box="[151,487,1603,1629]" pageId="5" pageNumber="84">
<taxonomicName id="BAEB60EAFFD8FFE700C784413BDFF9E6" ID-CoL="9HN3D" authority="Abu El-Hassan &amp; Badrawy &amp; Fadl &amp; Mohammad, 2019" authorityName="Abu El-Hassan &amp; Badrawy &amp; Fadl &amp; Mohammad" authorityYear="2019" box="[151,386,1603,1629]" class="Insecta" family="Tabanidae" genus="Atylotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="species" species="shagrensis" status="sp. nov.">
<emphasis id="4F9FC77BFFD8FFE700C784413BDFF9E6" bold="true" box="[151,386,1603,1629]" italics="true" pageId="5" pageNumber="84">Atylotus shagrensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54AC7A00FFD8FFE701D984413BBAF9E6" box="[393,487,1603,1629]" pageId="5" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="7D541B69FFD8FFE700C784673B56F9C5" blockId="5.[151,487,1603,1662]" box="[151,267,1637,1662]" pageId="5" pageNumber="84">(Figs 2, 7)</paragraph>
</subSubSection>
<subSubSection id="35F148E2FFD8FFE700C784AF3E91F97D" box="[151,1228,1709,1735]" pageId="5" pageNumber="84" type="materials_examined">
<paragraph id="7D541B69FFD8FFE700C784AF3E91F97D" blockId="5.[151,1436,1709,2025]" box="[151,1228,1709,1735]" pageId="5" pageNumber="84">
<emphasis id="4F9FC77BFFD8FFE700C784AF3BE4F97D" bold="true" box="[151,441,1709,1734]" pageId="5" pageNumber="84">
Type material.
<materialsCitation id="CD831134FFD8FFE7011C84AF3BE4F97D" ID-GBIF-Occurrence="2234217300" box="[332,441,1709,1734]" collectingDate="1933-04-24" collectionCode="PPDD" country="Egypt" location="Wadi Shagra" municipality="Mersa Alam" pageId="5" pageNumber="84" specimenCount="1" typeStatus="holotype">
<typeStatus id="A250A5CBFFD8FFE7011C84AF3BE4F97D" box="[332,441,1709,1734]" pageId="5" pageNumber="84" type="holotype">Holotype</typeStatus>
</materialsCitation>
</emphasis>
female,
<collectingCountry id="05FC5BF9FFD8FFE7024A84AD383CF97C" box="[538,609,1711,1735]" name="Egypt" pageId="5" pageNumber="84">Egypt</collectingCountry>
:
<location id="78344DB2FFD8FFE7023B84AF38A3F97D" LSID="urn:lsid:plazi:treatment:F542AA7FFFD8FFE400C784413F2BFC32:78344DB2FFD8FFE7023B84AF38A3F97D" box="[619,766,1709,1734]" country="Egypt" municipality="Mersa Alam" name="Wadi Shagra" pageId="5" pageNumber="84">Wadi Shagra</location>
, near
<collectingMunicipality id="9D308113FFD8FFE7036F84AD3993F97D" box="[831,974,1709,1734]" pageId="5" pageNumber="84">Mersa Alam</collectingMunicipality>
,
<collectingDate id="1911C441FFD8FFE7038984AC3E3EF97D" box="[985,1123,1710,1734]" pageId="5" pageNumber="84" value="1933-04-24">
<date id="09553DA9FFD8FFE7038984AC3E3EF97D" box="[985,1123,1710,1734]" pageId="5" pageNumber="84" value="1933-04-24">24. IV. 1933</date>
</collectingDate>
(PPDD).
</paragraph>
</subSubSection>
<subSubSection id="35F148E2FFD8FFE7009784D038B1F88F" pageId="5" pageNumber="84" type="diagnosis">
<paragraph id="7D541B69FFD8FFE7009784D03EFBF8AA" blockId="5.[151,1436,1709,2025]" pageId="5" pageNumber="84">
<emphasis id="4F9FC77BFFD8FFE7009784D03B1CF950" bold="true" box="[199,321,1746,1771]" pageId="5" pageNumber="84">Diagnosis.</emphasis>
Eyes with short yellow pubescence, frontal index 2.6, calli completely absent, R
<subScript id="E16F192CFFD8FFE7054B84E23F7EF955" attach="left" box="[1307,1315,1760,1774]" fontSize="6" pageId="5" pageNumber="84">4</subScript>
with long appendix, abdominal tergites 13 dark reddish brown with black spots, tergites 47 blackish.
</paragraph>
<paragraph id="7D541B69FFD8FFE70097851E38B1F88F" blockId="5.[151,1436,1709,2025]" box="[199,748,1820,1845]" pageId="5" pageNumber="84">
<emphasis id="4F9FC77BFFD8FFE70097851E3B7BF88E" bold="true" box="[199,294,1820,1845]" pageId="5" pageNumber="84">Length:</emphasis>
(female) body
<quantity id="BA13B68CFFD8FFE70182851E3865F88F" box="[466,568,1820,1845]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="5" pageNumber="84" unit="mm" value="12.5">12.5 mm</quantity>
, wing
<quantity id="BA13B68CFFD8FFE702D2851E38B5F88F" box="[642,744,1820,1845]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="5" pageNumber="84" unit="mm" value="11.6">11.6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="35F148E2FFD8FFE4009785423E4DFDFE" lastPageId="6" lastPageNumber="85" pageId="5" pageNumber="84" type="description">
<paragraph id="7D541B69FFD8FFE7009785423B04F8E2" blockId="5.[151,1436,1709,2025]" box="[199,345,1856,1881]" pageId="5" pageNumber="84">
<emphasis id="4F9FC77BFFD8FFE7009785423B04F8E2" bold="true" box="[199,345,1856,1881]" pageId="5" pageNumber="84">Description.</emphasis>
</paragraph>
<paragraph id="7D541B69FFD8FFE7009785663F7CF852" blockId="5.[151,1436,1709,2025]" pageId="5" pageNumber="84">
<emphasis id="4F9FC77BFFD8FFE7009785663B5AF8C6" bold="true" box="[199,263,1892,1917]" pageId="5" pageNumber="84">Head</emphasis>
: dark reddish-brown. Eyes with short yellow pubescence, without bands; frons parallel-sided, blackish with golden-yellow hairs, frontal index 2.6, calli completely absent, with whitish ocellar tubercle; face and genae polished brown tomentose; palpi yellowish with mixed white and a few black hairs; antennae pale reddish-yellow becoming darker towards apex, basal two segments with golden-yellow hairs, basal flagellomere broad.
</paragraph>
<paragraph id="7D541B69FFDBFFE4009782953B6DFEA4" blockId="6.[151,1436,151,905]" pageId="6" pageNumber="85">
<emphasis id="4F9FC77BFFDBFFE4009782953B7DFF0B" bold="true" box="[199,288,151,176]" pageId="6" pageNumber="85">Thorax</emphasis>
: blackish-brown with long and dense whitish and golden-yellow hairs; mesonotum including scutellum with dense long yellowish hairs, notopleural lobes reddish-brown with yellow hairs; legs reddish-yellow, fore tibia only blackish; wings hyaline with brown veins and a long appendix to R
<subScript id="E16F192CFFDBFFE4043282EC3E37FF47" attach="left" box="[1122,1130,238,252]" fontSize="6" pageId="6" pageNumber="85">4</subScript>
; halteres pale brown with whitish knob.
</paragraph>
<paragraph id="7D541B69FFDBFFE40097832F3EEAFE0E" blockId="6.[151,1436,151,905]" pageId="6" pageNumber="85">
<emphasis id="4F9FC77BFFDBFFE40097832F3B61FEFD" bold="true" box="[199,316,301,326]" pageId="6" pageNumber="85">Abdomen</emphasis>
: with the first three tergites dark reddish-brown, the 1
<superScript id="8A9EB621FFDBFFE403EA83283999FE83" attach="left" box="[954,964,298,312]" fontSize="6" pageId="6" pageNumber="85">st</superScript>
with black spots around the base of the scutellum, 2
<superScript id="8A9EB621FFDBFFE4017083523B6DFEE5" attach="left" box="[288,304,336,350]" fontSize="6" pageId="6" pageNumber="85">nd</superScript>
and 3
<superScript id="8A9EB621FFDBFFE4012483523BDCFEE5" attach="left" box="[372,385,336,350]" fontSize="6" pageId="6" pageNumber="85">rd</superScript>
with large black spots on the middle of each segment and a small black triangular spot on the sides of the posterior margin of each segment, tergites 47 blackish, all tergites covered with long golden-yellow and whitish hairs, venter reddish-brown and covered with long golden-yellow and white hairs.
</paragraph>
<paragraph id="7D541B69FFDBFFE4009783C23B2AFE62" blockId="6.[151,1436,151,905]" box="[199,375,448,473]" pageId="6" pageNumber="85">Male unknown.</paragraph>
<paragraph id="7D541B69FFDBFFE4009783E638BAFE46" blockId="6.[151,1436,151,905]" box="[199,743,484,509]" pageId="6" pageNumber="85">
<emphasis id="4F9FC77BFFDBFFE4009783E63BF5FE46" bold="true" box="[199,424,484,509]" pageId="6" pageNumber="85">Local distribution:</emphasis>
W. Shagra (Eastern Desert).
</paragraph>
<paragraph id="7D541B69FFDBFFE40097800A380BFD9A" blockId="6.[151,1436,151,905]" box="[199,598,520,545]" pageId="6" pageNumber="85">
<emphasis id="4F9FC77BFFDBFFE40097800A3859FD9A" bold="true" box="[199,516,520,545]" pageId="6" pageNumber="85">Geographical distribution.</emphasis>
<collectingCountry id="05FC5BF9FFDBFFE4025B800B380FFD9A" box="[523,594,521,545]" name="Egypt" pageId="6" pageNumber="85">Egypt</collectingCountry>
.
</paragraph>
<paragraph id="7D541B69FFDBFFE40097802E3E4DFDFE" blockId="6.[151,1436,151,905]" box="[199,1040,556,581]" pageId="6" pageNumber="85">
<emphasis id="4F9FC77BFFDBFFE40097802E3B9BFDFE" bold="true" box="[199,454,556,581]" pageId="6" pageNumber="85">Specimens examined.</emphasis>
Wadi Shagra,
<date id="09553DA9FFDBFFE4023A802F38A9FDFE" box="[618,756,557,581]" pageId="6" pageNumber="85" value="1933-04-24">24. IV. 1933</date>
(
<specimenCount id="6BEDD0E0FFDBFFE40356802F3973FDFF" box="[774,814,557,581]" pageId="6" pageNumber="85" type="female">1 ♀</specimenCount>
,
<typeStatus id="A250A5CBFFDBFFE4036C802E39FDFDFE" box="[828,928,556,581]" pageId="6" pageNumber="85" type="holotype">holotype</typeStatus>
) (PPDD).
</paragraph>
</subSubSection>
<subSubSection id="35F148E2FFDBFFE4009780523906FCFA" pageId="6" pageNumber="85" type="etymology">
<paragraph id="7D541B69FFDBFFE400978052392AFDD2" blockId="6.[151,1436,151,905]" box="[199,887,592,617]" pageId="6" pageNumber="85">
<emphasis id="4F9FC77BFFDBFFE4009780523B0CFDD2" bold="true" box="[199,337,592,617]" pageId="6" pageNumber="85">Etymology:</emphasis>
This new species is named after its
<typeStatus id="A250A5CBFFDBFFE402B68053394BFDD2" box="[742,790,593,617]" pageId="6" pageNumber="85">type</typeStatus>
locality.
</paragraph>
<paragraph id="7D541B69FFDBFFE4009780763906FCFA" blockId="6.[151,1436,151,905]" pageId="6" pageNumber="85">
<emphasis id="4F9FC77BFFDBFFE4009780763B68FD36" bold="true" box="[199,309,628,653]" pageId="6" pageNumber="85">Affinities</emphasis>
:
<taxonomicName id="BAEB60EAFFDBFFE4011780763879FD36" authority="Abu El-Hassan &amp; Badrawy &amp; Fadl &amp; Mohammad, 2019" authorityName="Abu El-Hassan &amp; Badrawy &amp; Fadl &amp; Mohammad" authorityYear="2019" box="[327,548,628,653]" class="Insecta" family="Tabanidae" genus="Atylotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="85" phylum="Arthropoda" rank="species" species="shagrensis" status="sp. nov.">
<emphasis id="4F9FC77BFFDBFFE4011780763879FD36" box="[327,548,628,653]" italics="true" pageId="6" pageNumber="85">Atylotus shagrensis</emphasis>
</taxonomicName>
<emphasis id="4F9FC77BFFDBFFE4027E807638D6FD36" bold="true" box="[558,651,628,653]" pageId="6" pageNumber="85">
<taxonomicNameLabel id="54AC7A00FFDBFFE4027E807638D6FD36" box="[558,651,628,653]" pageId="6" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is closely related to the Egyptian species
<taxonomicName id="BAEB60EAFFDBFFE4042A80773A85FD0A" authority="(Wiedemann, 1828)" baseAuthorityName="Wiedemann" baseAuthorityYear="1828" class="Insecta" family="Tabanidae" genus="Atylotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="85" phylum="Arthropoda" rank="species" species="agricola">
<emphasis id="4F9FC77BFFDBFFE4042A80773EA6FD36" box="[1146,1275,628,653]" italics="true" pageId="6" pageNumber="85">A. agricola</emphasis>
(
<bibRefCitation id="197A6698FFDBFFE4055D80763A8DFD0A" author="Wiedemann, C." pageId="6" pageNumber="85" refId="ref6440" refString="Wiedemann, C. (1828) Ausssereuropaische zweiflugelige Insecten. Als Fortsetzung des Meigenschen Werks. Erster Theil, Schulz, Hamm, xxxii + 608 pp." type="book" year="1828">Wiedemann, 1828</bibRefCitation>
)
</taxonomicName>
and the European species
<taxonomicName id="BAEB60EAFFDBFFE4024D809B39CBFD0A" authority="(Villeneuve), 1920" authorityName="" authorityYear="1920" baseAuthorityName="Villeneuve" box="[541,918,664,689]" class="Insecta" family="Tabanidae" genus="Atylotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="85" phylum="Arthropoda" rank="species" species="loewianus">
<emphasis id="4F9FC77BFFDBFFE4024D809B38EEFD0A" box="[541,691,664,689]" italics="true" pageId="6" pageNumber="85">A. loewianus</emphasis>
(
<bibRefCitation id="197A6698FFDBFFE40297809A39CBFD0A" author="Villeneuve, J." box="[711,918,664,689]" pageId="6" pageNumber="85" pagination="60 - 65" refId="ref6220" refString="Villeneuve, J. (1920) Sur Ochrops (Atylotus) fulvus Meigen. Annales de la Societe Entomologique de Belgique, 1920, 60 - 65." type="journal article" year="1920">Villeneuve), 1920</bibRefCitation>
</taxonomicName>
because of the parallel-sided frons and the abdominal pattern. The new species can be easily separated from
<taxonomicName id="BAEB60EAFFDBFFE403F480BF3E7BFD6E" authorityName="Wiedemann" authorityYear="1828" baseAuthorityName="Wiedemann" baseAuthorityYear="1828" box="[932,1062,700,725]" class="Insecta" family="Tabanidae" genus="Atylotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="85" phylum="Arthropoda" rank="species" species="agricola">
<emphasis id="4F9FC77BFFDBFFE403F480BF3E7BFD6E" box="[932,1062,700,725]" italics="true" pageId="6" pageNumber="85">A. agricola</emphasis>
</taxonomicName>
by the absence of frontal calli, blackish colour of frons, polished brown colour of subcallus and reddish-brown colour of the abdomen.
<taxonomicName id="BAEB60EAFFDBFFE4051080E23B53FCA6" authority="Abu El-Hassan &amp; Badrawy &amp; Fadl &amp; Mohammad, 2019" authorityName="Abu El-Hassan &amp; Badrawy &amp; Fadl &amp; Mohammad" authorityYear="2019" class="Insecta" family="Tabanidae" genus="Atylotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="85" phylum="Arthropoda" rank="species" species="shagrensis" status="sp. nov.">
<emphasis id="4F9FC77BFFDBFFE4051080E23B53FCA6" italics="true" pageId="6" pageNumber="85">Atylotus shagrensis</emphasis>
</taxonomicName>
<emphasis id="4F9FC77BFFDBFFE4014A81063B24FCA6" bold="true" box="[282,377,772,797]" pageId="6" pageNumber="85">
<taxonomicNameLabel id="54AC7A00FFDBFFE4014A81063B24FCA6" box="[282,377,772,797]" pageId="6" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from
<taxonomicName id="BAEB60EAFFDBFFE402D981073943FCA6" authorityName="Villeneuve" authorityYear="1920" baseAuthorityName="Villeneuve" baseAuthorityYear="1920" box="[649,798,772,797]" class="Insecta" family="Tabanidae" genus="Atylotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="85" phylum="Arthropoda" rank="species" species="loewianus">
<emphasis id="4F9FC77BFFDBFFE402D981073943FCA6" box="[649,798,772,797]" italics="true" pageId="6" pageNumber="85">A. loewianus</emphasis>
</taxonomicName>
by the hairy eyes, absence of frontal calli, colour of subcallus and frons, and reddish-brown colour of the abdomen.
</paragraph>
</subSubSection>
<subSubSection id="35F148E2FFDBFFE40097814E3F2BFC32" pageId="6" pageNumber="85" type="discussion">
<paragraph id="7D541B69FFDBFFE40097814E3F2BFC32" blockId="6.[151,1436,151,905]" pageId="6" pageNumber="85">
<emphasis id="4F9FC77BFFDBFFE40097814E3B66FCDE" bold="true" box="[199,315,844,869]" pageId="6" pageNumber="85">Remarks.</emphasis>
Four female specimens of
<taxonomicName id="BAEB60EAFFDBFFE402C7814E3E5DFCDE" authority="Villeneuve" authorityName="Villeneuve" box="[663,1024,844,869]" class="Insecta" family="Tabanidae" genus="Atylotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="85" phylum="Arthropoda" rank="species" species="loewianus">
<emphasis id="4F9FC77BFFDBFFE402C7814E3929FCDE" box="[663,884,844,869]" italics="true" pageId="6" pageNumber="85">Atylotus loewianus</emphasis>
Villeneuve
</taxonomicName>
, which were collected in
<collectingCountry id="05FC5BF9FFDBFFE4051A814E3FC1FCDE" box="[1354,1436,844,869]" name="Croatia" pageId="6" pageNumber="85">Croatia</collectingCountry>
(
<date id="09553DA9FFDBFFE400CF81723B7AFC32" box="[159,295,880,905]" pageId="6" pageNumber="85" value="1996-08-08">8.VIII.1996</date>
), were sent to us for examination by Prof. Dr. Stjepan Krčmar from his personal collection (SKC).
</paragraph>
</subSubSection>
</treatment>
</document>