treatments-xml/data/D3/4C/F6/D34CF6924CFA656FA325799EB741CB18.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.19.4098" ID-PMC="PMC3597002" ID-Pensoft-Pub="1314-2003-19-31" ID-Pensoft-UUID="FFBDFFE13B62FFA81373FFBB967CFFAF" ID-PubMed="23717189" ID-Zenodo-Dep="576153" ModsDocID="1314-2003-19-31" checkinTime="1451251957823" checkinUser="pensoft" docAuthor="Snow, Neil" docDate="2012" docId="D34CF6924CFA656FA325799EB741CB18" docLanguage="en" docName="PhytoKeys 19: 31-49" docOrigin="PhytoKeys 19" docPubDate="2012-12-28" docSource="http://dx.doi.org/10.3897/phytokeys.19.4098" docTitle="Rhodamnia waigeoensis N. Snow 2012, sp. nov." docType="treatment" docVersion="6" id="FFBDFFE13B62FFA81373FFBB967CFFAF" lastPageNumber="42" masterDocId="FFBDFFE13B62FFA81373FFBB967CFFAF" masterDocTitle="Five new species of Rhodamnia (Myrtaceae, Myrteae) from New Guinea" masterLastPageNumber="49" masterPageNumber="31" pageNumber="39" updateTime="1668140689891" updateUser="ExternalLinkService">
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<mods:title>Five new species of Rhodamnia (Myrtaceae, Myrteae) from New Guinea</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Snow, Neil</mods:namePart>
<mods:affiliation>Herbarium Pacificum, Bishop Museum, 1525 Bernice St., Honolulu, HI 96821 USA &amp; 527 S. Oakes St, Helena, MT 59601 USA</mods:affiliation>
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<mods:title>PhytoKeys</mods:title>
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<mods:date>2012</mods:date>
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<mods:number>2012-12-28</mods:number>
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<mods:number>19</mods:number>
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<mods:start>31</mods:start>
<mods:end>49</mods:end>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.19.4098</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.19.4098</mods:identifier>
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<treatment ID-GBIF-Taxon="152024469" LSID="urn:lsid:plazi:treatment:D34CF6924CFA656FA325799EB741CB18" httpUri="http://treatment.plazi.org/id/D34CF6924CFA656FA325799EB741CB18" lastPageId="11" lastPageNumber="42" pageId="8" pageNumber="39">
<subSubSection pageId="8" pageNumber="39" type="nomenclature">
<paragraph pageId="8" pageNumber="39">
<taxonomicName LSID="D34CF692-4CFA-656F-A325-799EB741CB18" authority="N. Snow" authorityName="N. Snow" authorityYear="2012" class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhodamnia waigeoensis" order="Myrtales" pageId="8" pageNumber="39" phylum="Tracheophyta" rank="species" species="waigeoensis" status="sp. nov.">Rhodamnia waigeoensis N. Snow</taxonomicName>
<taxonomicNameLabel pageId="8" pageNumber="39">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Island of New Guinea, showing localities of new species. Triangle (▲) = Rhodamnia asekiensis; Square (■) = Rhodamnia daymanensis; Diamond (◆) = Rhodamnia makumak (at bottom of line); Closed circle (⚫) = Rhodamnia toratot (at top of line); Inverted triangle (▼) = Rhodamnia waigeoensis (upper left, at bottom of line); Open circles (⚪) = Rhodamnia sharpeana (left end of line [upper] and bottom of line [lower, on truncated eastern half of Tagula Island])." httpUri="https://binary.pensoft.net/fig/10217" pageId="8" pageNumber="39">Figures 1</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Rhodamnia waigeoensis N. Snow. Photo of the the holotype at A (P. van Royen 5556)" httpUri="https://binary.pensoft.net/fig/10222" pageId="8" pageNumber="39">6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="39" type="latin">
<paragraph pageId="8" pageNumber="39">
<emphasis italics="true" pageId="8" pageNumber="39">Closely resembling but differing from</emphasis>
Rhodamnia novoguineensis 
<emphasis italics="true" pageId="8" pageNumber="39">by its thicker and more rigid pedicels, thickly coriaceous leaves, basal acrodromous venation, densely yellowish abaxial laminar indumentum, and solitary flowers</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="39" type="type">
<paragraph pageId="8" pageNumber="39">Type.</paragraph>
<paragraph pageId="8" pageNumber="39">Indonesia. Waigeo Island, Go Isthmus, path from Poean Bay to Fofak Bay, 17 Feb 1955, P. van Royen 5556(holotype: A! [bar code no. 00307477]; isotypes: CANB!, K!, L n.v.).</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="41" pageId="8" pageNumber="39" type="description">
<paragraph pageId="8" pageNumber="39">Description.</paragraph>
<paragraph lastPageId="10" lastPageNumber="41" pageId="8" pageNumber="39">
Trees 5-7 m; girth to 15 cm. Branchlets terete to slightly compressed, the epidermis later becoming fissured; indumentum densely sericeous, 
<pageBreakToken pageId="9" pageNumber="40" start="start">mostly</pageBreakToken>
yellowish or somewhat ferrugineous but becoming more whitish with age. Leaves opposite, evenly distributed along branchlets, discolorous, glossy above and below, the nacreous sheen below imparted by the dense, tightly appressed greenish-white indumentum. Colleters absent. Petioles 5-6.5 mm long, somewhat flat 
<pageBreakToken pageId="10" pageNumber="41" start="start">tened</pageBreakToken>
above, densely sericeous (or somewhat tomentose with age), the indumentum yellowish but aging whitish. Leaf blades (3.5-)6.0-10.0 cm long, (1.8-)2.5-3.7 cm wide, narrowly ovate to ovate, surface flat or slightly wavy; base cuneate, apex acuminate and occasionally somewhat falcate, tip acute; venation perfect basal acrodromous; secondary veins numerous but thin, ca. 0.8-2.0 mm apart; marginal nerve prominent, mostly 0.7-0.9 mm from mid-leaf margins; margins flat; adaxial surface sparsely sericeous, midvein flush throughout, oil glands invisible; abaxial surface densely sericeous with greenish-whitish indumentum but this mostly not obscuring venation, midvein raised throughout, oil glands invisible. Inflorescence (limited material) a 3-flowered cyme, terminal, solitary (one per leaf subtending leaf); peduncle ca. 5 mm, stiff, terete in transsection, densely yellowish-orangish sericeous; pedicel to ca. 3 mm long, indumentum as per peduncle. Bracteoles 2, narrowly triangular and stiffly erect, ca. 2 mm long, ca. 0.5 mm wide, sericeous, sometimes persisting into fruit. Hypanthium cupulate, densely sericeous, oil glands absent, texture smooth. Calyx lobes 4, 2.3-3.5 mm long, broadly ovate, sericeous above, densely sericeous below, more or less reflexed in fruit. Petals 4, 5.5-7.0 mm long, width uncertain (material scanty), apparently obovate to broadly obovate, white (based on specimen label), sparsely sericeous above, densely sericeous below. Staminal disk 2.5-3.5 mm wide, densely short-hairy. Ovary apex densely short-hairy. Stamen number uncertain but almost certainly greater than 20, filaments and anthers red (from specimen label); anthers subcylindrical (material scanty), ca. 0.5 mm. Stigma not seen. Locule 1, placentation parietal, placentas 2, ovules numerous. Fruit (reportedly immature) globose-subglobse, up to 8 mm long and 9.5 mm wide, light green when immature, densely sericeous but indumentum thinning with maturity. Seeds irregularly angular, up to 4 mm long (small sample), up to 9 per fruit, crowded; seed coat hard. Embryos not seen.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/10222" pageId="10" pageNumber="41" start="Figure 6" startId="F6">
<paragraph pageId="10" pageNumber="41">
<emphasis bold="true" pageId="10" pageNumber="41">Figure 6.</emphasis>
<taxonomicName authorityName="N. Snow" authorityYear="2012" class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="10" pageNumber="41" phylum="Tracheophyta" rank="species" species="waigeoensis">
<emphasis italics="true" pageId="10" pageNumber="41">Rhodamnia waigeoensis</emphasis>
</taxonomicName>
N. Snow. Photo of the the holotype at A (
<emphasis italics="true" pageId="10" pageNumber="41">P. van Royen 5556</emphasis>
)
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="10" pageNumber="41" type="phenology">
<paragraph pageId="10" pageNumber="41">Phenology.</paragraph>
<paragraph pageId="10" pageNumber="41">Flowering February; fruiting in January and February.</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="41" type="distribution">
<paragraph pageId="10" pageNumber="41">Distribution.</paragraph>
<paragraph pageId="10" pageNumber="41">
Waigeo Island, Indonesia; from ca. 10-150 m elevation in xerophytic, 
<taxonomicName genus="Myrtaceae" lsidName="" pageId="10" pageNumber="41" rank="genus">Myrtaceae</taxonomicName>
-dominated vegetation at lower elevations behind and upslope of the village of Waifoi, and from transitional forests dominated by 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Decaspermum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="10" pageNumber="41" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="10" pageNumber="41">Decaspermum</emphasis>
</taxonomicName>
J.R. Forst. &amp; G. Forst. (
<taxonomicName genus="Myrtaceae" lsidName="" pageId="10" pageNumber="41" rank="genus">Myrtaceae</taxonomicName>
) or 
<taxonomicName authorityName="Blume" authorityYear="1856" class="Insecta" family="Crambidae" genus="Vatica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="rassak">
<emphasis italics="true" pageId="10" pageNumber="41">Vatica rassak</emphasis>
</taxonomicName>
Blume(= 
<taxonomicName class="Insecta" family="Crambidae" genus="Vatica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="papuana">
<emphasis italics="true" pageId="10" pageNumber="41">Vatica papuana</emphasis>
</taxonomicName>
Schum. &amp; Hollr. [synonym]) (
<taxonomicName genus="Dipterocarpaceae" lsidName="" pageId="10" pageNumber="41" rank="genus">Dipterocarpaceae</taxonomicName>
) at the higher elevation (ca. 150 m).
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="41" type="conservation status">
<paragraph pageId="10" pageNumber="41">Conservation status.</paragraph>
<paragraph pageId="10" pageNumber="41">
Data Deficient given the lack of recent information or collections. 
<taxonomicName authorityName="N. Snow" authorityYear="2012" class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="10" pageNumber="41" phylum="Tracheophyta" rank="species" species="waigeoensis">
<emphasis italics="true" pageId="10" pageNumber="41">Rhodamnia waigeoensis</emphasis>
</taxonomicName>
is presently known only from two collections. The specimen on the type label indicates that the species was common locally at the time of its collection nearly sixty years ago. A vegetation type similar to that of the type gathering occurs on the island of Rauki, where the species also may occur. While the reported ethnobotanical use of 
<taxonomicName authorityName="N. Snow" authorityYear="2012" class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="10" pageNumber="41" phylum="Tracheophyta" rank="species" species="waigeoensis">
<emphasis italics="true" pageId="10" pageNumber="41">Rhodamnia waigeoensis</emphasis>
</taxonomicName>
for cigarette making may lend the species some protection, it also may have encouraged overexploitation.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="41" type="vernacular name">
<paragraph pageId="10" pageNumber="41">Vernacular name.</paragraph>
<paragraph pageId="10" pageNumber="41">
<emphasis italics="true" pageId="10" pageNumber="41">Kikir</emphasis>
(in the Malayan language).
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="41" type="ethnobotany">
<paragraph pageId="10" pageNumber="41">Ethnobotany.</paragraph>
<paragraph pageId="10" pageNumber="41">The herbarium label indicates that the leaves are used for making cigarettes.</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="42" type="comments">
<paragraph pageId="11" pageNumber="42">
<pageBreakToken pageId="11" pageNumber="42" start="start">Comments</pageBreakToken>
.
</paragraph>
<paragraph pageId="11" pageNumber="42">
<taxonomicName authorityName="N. Snow" authorityYear="2012" class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="waigeoensis">
<emphasis italics="true" pageId="11" pageNumber="42">Rhodamnia waigeoensis</emphasis>
</taxonomicName>
belongs in the 
<normalizedToken originalValue="“pearly”">&quot;pearly&quot;</normalizedToken>
group of species given its nacreous indumentum (
<bibRefCitation author="Snow, N" journalOrPublisher="Systematic Botany Monographs" pageId="12" pageNumber="43" pagination="1 - 69" refId="B14" refString="Snow, N, 2007. Systematics of the Australian species of Rhodamnia (Myrtaceae). Systematic Botany Monographs 82: 1 - 69" title="Systematics of the Australian species of Rhodamnia (Myrtaceae)." volume="82" year="2007">Snow 2007</bibRefCitation>
). 
<bibRefCitation author="Scott, AJ" journalOrPublisher="Kew Bulletin" pageId="12" pageNumber="43" pagination="429 - 459" publicationUrl="10.2307/4110145" refId="B11" refString="Scott, AJ, 1979. A revision of Rhodamnia (Myrtaceae). Kew Bulletin 33: 429 - 459, 10.2307/4110145" title="A revision of Rhodamnia (Myrtaceae)." url="10.2307/4110145" volume="33" year="1979">Scott (1979)</bibRefCitation>
included the type gathering of 
<taxonomicName authorityName="N. Snow" authorityYear="2012" class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="waigeoensis">
<emphasis italics="true" pageId="11" pageNumber="42">Rhodamnia waigeoensis</emphasis>
</taxonomicName>
in 
<taxonomicName authorityName="A. J. Scott" authorityYear="1979" class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="novoguineensis">
<emphasis italics="true" pageId="11" pageNumber="42">Rhodamnia novoguineensis</emphasis>
</taxonomicName>
A.J. Scott and the paratype gathering in 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="pachyloba">
<emphasis italics="true" pageId="11" pageNumber="42">Rhodamnia pachyloba</emphasis>
</taxonomicName>
A.J. Scott.
</paragraph>
<paragraph pageId="11" pageNumber="42">
<taxonomicName authorityName="N. Snow" authorityYear="2012" class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="waigeoensis">
<emphasis italics="true" pageId="11" pageNumber="42">Rhodamnia waigeoensis</emphasis>
</taxonomicName>
differs from 
<taxonomicName authorityName="A. J. Scott" authorityYear="1979" class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="novoguineensis">
<emphasis italics="true" pageId="11" pageNumber="42">Rhodamnia novoguineensis</emphasis>
</taxonomicName>
by its thickly coriaceous leaves (vs. thinly coriaceous in 
<taxonomicName authorityName="A. J. Scott" authorityYear="1979" class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="novoguineensis">
<emphasis italics="true" pageId="11" pageNumber="42">Rhodamnia novoguineensis</emphasis>
</taxonomicName>
), consistently basal acrodromous leaf venation (vs. even or uneven suprabasal acrodromous in 
<taxonomicName authorityName="A. J. Scott" authorityYear="1979" class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="novoguineensis">
<emphasis italics="true" pageId="11" pageNumber="42">Rhodamnia novoguineensis</emphasis>
</taxonomicName>
), dense abaxial laminar indumentum with yellowish trichomes (vs. relatively sparse and whitish in 
<taxonomicName authorityName="A. J. Scott" authorityYear="1979" class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="novoguineensis">
<emphasis italics="true" pageId="11" pageNumber="42">Rhodamnia novoguineensis</emphasis>
</taxonomicName>
), solitary flowers (vs. triads or few-flowered racemes in 
<taxonomicName authorityName="A. J. Scott" authorityYear="1979" class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="novoguineensis">
<emphasis italics="true" pageId="11" pageNumber="42">Rhodamnia novoguineensis</emphasis>
</taxonomicName>
), thicker (0.5-0.7 mm wide) and rigid pedicels (vs. ca. 0.3 mm thick and flaccid in 
<taxonomicName authorityName="A. J. Scott" authorityYear="1979" class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="novoguineensis">
<emphasis italics="true" pageId="11" pageNumber="42">Rhodamnia novoguineensis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph pageId="11" pageNumber="42">
Waigeo Island is part of the Raja Ampat Islands of Indonesian New Guinea. The region harbors unusual vegetation assemblages (van 
<bibRefCitation author="Royen, P van" journalOrPublisher="Nova Guinea, Botany" pageId="12" pageNumber="43" pagination="24 - 62" refId="B10" refString="Royen, P van, 1960. Sertulum Papuanum 3. The vegetation of some parts of Waigeo Island. Nova Guinea, Botany 5: 24 - 62" title="Sertulum Papuanum 3. The vegetation of some parts of Waigeo Island." volume="5" year="1960">Royen 1960</bibRefCitation>
), has high rates of endemism (
<bibRefCitation author="Supriatna, J (editor)" journalOrPublisher="Phytotaxa" pageId="12" pageNumber="43" refId="B21" refString="Supriatna, J (editor), 1999. The Irian Jaya Biodiversity Conservation Priority-Setting Workshop. Final Report. Conservation International, Washington, DC." title="The Irian Jaya Biodiversity Conservation Priority-Setting Workshop. Final Report. Conservation International, Washington, DC." year="1999">Supriatna 1999</bibRefCitation>
), and was the subject of relatively recent rapid-assessment surveys (
<bibRefCitation author="Takeuchi, W" journalOrPublisher="Sida" pageId="12" pageNumber="43" pagination="1099 - 1144" refId="B24" refString="Takeuchi, W, 2003a. A community-level floristic reconnaissance of the Raja Ampat Islands in New Guinea. Sida 20: 1099 - 1144" title="A community-level floristic reconnaissance of the Raja Ampat Islands in New Guinea." volume="20" year="2003 a">Takeuchi 2003a</bibRefCitation>
). Van Royen (1960: 54-56) summarized the vegetation on portions of Waigeo Island using six broad categories. One of these, xerophytic vegetation, is described as having three variants, one being dominated by 
<taxonomicName genus="Myrtaceae" lsidName="" pageId="11" pageNumber="42" rank="genus">Myrtaceae</taxonomicName>
.
</paragraph>
<paragraph pageId="11" pageNumber="42">
The label of the type specimen refers directly to the xerophytic vegetation located behind the small village of Waifoi on the east bank of Majalibit Bay. Takeuchi (2003a,b) reported that the Waigeo ultrabasic vegetation resembles the pioneer communities on the ultrabasics at the Kamilai Wildlife Management Area (KWMA) in the Bowutu Mountains (Morobe Province, Papua New Gueina). Communities at KWMA can be topographically unstable due to landslides, but in general appearance and composition are similar to those on Waigeo. However, 
<bibRefCitation author="Takeuchi, W" journalOrPublisher="Sida" pageId="12" pageNumber="43" pagination="1491 - 1559" refId="B25" refString="Takeuchi, W, 2003b. Botanical summary of a lowland ultrabasic flora in Papua New Guinea. Sida 20: 1491 - 1559" title="Botanical summary of a lowland ultrabasic flora in Papua New Guinea." volume="20" year="2003 b">Takeuchi (2003b)</bibRefCitation>
believes the vegetation on the Waigeo ultrabasics is primarily caused by fire succession.
</paragraph>
<paragraph pageId="11" pageNumber="42">
A xerophytic vegetation similar to that occuring on the hills upslope of Waifoi, the village near the type collection, was encountered elsewhere by van Royen (1960: 39, 41) in the Kambelay Hills and the Go Isthmus of Waigeo Island. This general type of xerophytic vegetation is said to recur on Rauki Island, which lies northwest of 
<normalizedToken originalValue="Kabaré">Kabare</normalizedToken>
Bay, where it occurs at the higher elevations (probably less than ca. 40 m, but reported by van Royen [p. 45] as 25 m) along the southern end of the island (van 
<bibRefCitation author="Royen, P van" journalOrPublisher="Nova Guinea, Botany" pageId="12" pageNumber="43" pagination="24 - 62" refId="B10" refString="Royen, P van, 1960. Sertulum Papuanum 3. The vegetation of some parts of Waigeo Island. Nova Guinea, Botany 5: 24 - 62" title="Sertulum Papuanum 3. The vegetation of some parts of Waigeo Island." volume="5" year="1960">Royen 1960</bibRefCitation>
: 44) at ca. 
<geoCoordinate degrees="0" direction="south" minutes="52" orientation="latitude" precision="925" value="-0.8666667">0°52'S</geoCoordinate>
, 
<geoCoordinate degrees="130" direction="east" minutes="56" orientation="longitude" precision="925" value="130.93333">130°56'E</geoCoordinate>
(coordinates based on Google Earth 
<normalizedToken originalValue="™">(TM)</normalizedToken>
[accessed 2 June 2009]). (Rauki Island has been known previously as Rawak, Rawah or Lawak [van 
<bibRefCitation author="Royen, P van" journalOrPublisher="Nova Guinea, Botany" pageId="12" pageNumber="43" pagination="24 - 62" refId="B10" refString="Royen, P van, 1960. Sertulum Papuanum 3. The vegetation of some parts of Waigeo Island. Nova Guinea, Botany 5: 24 - 62" title="Sertulum Papuanum 3. The vegetation of some parts of Waigeo Island." volume="5" year="1960">Royen 1960</bibRefCitation>
: 43]). The substrates underlying the xerophytic vegetation of Rauki include ultrabasic outcrops among the more prevalent limestone (van 
<bibRefCitation author="Royen, P van" journalOrPublisher="Nova Guinea, Botany" pageId="12" pageNumber="43" pagination="24 - 62" refId="B10" refString="Royen, P van, 1960. Sertulum Papuanum 3. The vegetation of some parts of Waigeo Island. Nova Guinea, Botany 5: 24 - 62" title="Sertulum Papuanum 3. The vegetation of some parts of Waigeo Island." volume="5" year="1960">Royen 1960</bibRefCitation>
: 45).
</paragraph>
<paragraph pageId="11" pageNumber="42">
Van Royen (1960: 32) described the soils underlying the xerophytic vegetation on Waigeo as &quot;sandy brown clays with much limestone&quot;. The relatively open vegetation on the slopes was indicated as being spare of trees but conspicuous in its presence of shrubby 
<taxonomicName genus="Myrtaceae" lsidName="" pageId="11" pageNumber="42" rank="genus">Myrtaceae</taxonomicName>
. Noted specifically for 
<taxonomicName genus="Myrtaceae" lsidName="" pageId="11" pageNumber="42" rank="genus">Myrtaceae</taxonomicName>
(van 
<bibRefCitation author="Royen, P van" journalOrPublisher="Nova Guinea, Botany" pageId="12" pageNumber="43" pagination="24 - 62" refId="B10" refString="Royen, P van, 1960. Sertulum Papuanum 3. The vegetation of some parts of Waigeo Island. Nova Guinea, Botany 5: 24 - 62" title="Sertulum Papuanum 3. The vegetation of some parts of Waigeo Island." volume="5" year="1960">Royen 1960</bibRefCitation>
: 32, 55, 59) were 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Baeckia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="frutescens">
<emphasis italics="true" pageId="11" pageNumber="42">Baeckia frutescens</emphasis>
</taxonomicName>
L., 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Myrtella" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="beccarii">
<emphasis italics="true" pageId="11" pageNumber="42">Myrtella beccarii</emphasis>
</taxonomicName>
F. Muell. and 
<taxonomicName authorityName="Baill" authorityYear="1876" baseAuthorityName="Bl." class="Magnoliopsida" family="Myrtaceae" genus="Decaspermum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="rubrum">
<emphasis italics="true" pageId="11" pageNumber="42">Decaspermum rubrum</emphasis>
</taxonomicName>
(Blume) Baill. (as 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Decaspermum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="fruticosum">
<emphasis italics="true" pageId="11" pageNumber="42">Decaspermum fruticosum</emphasis>
</taxonomicName>
J. R. Forst. &amp; G. Forst.var. 
<emphasis italics="true" pageId="11" pageNumber="42">rubrum</emphasis>
, a nomenclatural change that was apparently never validly published [
<bibRefCitation author="Scott, AJ" journalOrPublisher="Kew Bulletin" pageId="12" pageNumber="43" pagination="149 - 165" publicationUrl="10.2307/4108492" refId="B12" refString="Scott, AJ, 1985. Decaspermum (Myrtaceae) in New Guinea. Kew Bulletin 40: 149 - 165, 10.2307/4108492" title="Decaspermum (Myrtaceae) in New Guinea." url="10.2307/4108492" volume="40" year="1985">Scott 1985</bibRefCitation>
; 
<bibRefCitation author="Govaerts, R" journalOrPublisher="Proceedings of the National Academy of Sciences (USA)" pageId="12" pageNumber="43" refId="B5" refString="Govaerts, R, Sobral, M, Ashton, P, Barrie, F, Holst, BK, Landrum, LL, Matsumoto, K, Mazine, FF, Nic, Lughadha E, Proenca, C, Soares-Silva, LH, Wilson, PG, Lucas, E, 2008. World Checklist of Myrtaceae. Royal Botanic Gardens, Kew." title="World Checklist of Myrtaceae. Royal Botanic Gardens, Kew." year="2008">Govaerts et al. 2008</bibRefCitation>
]), and &quot; 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="trinervia">
<emphasis italics="true" pageId="11" pageNumber="42">Rhodamnia trinervia</emphasis>
</taxonomicName>
Reinw. ex Blume 
<emphasis italics="true" pageId="11" pageNumber="42">&quot;</emphasis>
. However, the collection number (5556) that van Royen (1960: 59) cited for 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="trinervia">
<emphasis italics="true" pageId="11" pageNumber="42">Rhodamnia trinervia</emphasis>
</taxonomicName>
represents the holotype of 
<taxonomicName authorityName="N. Snow" authorityYear="2012" class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="waigeoensis">
<emphasis italics="true" pageId="11" pageNumber="42">Rhodamnia waigeoensis</emphasis>
</taxonomicName>
, and 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="trinervia">
<emphasis italics="true" pageId="11" pageNumber="42">Rhodamnia trinervia</emphasis>
</taxonomicName>
is now considered to be a synonym of 
<taxonomicName authorityName="Miq" authorityYear="1855" baseAuthorityName="Benth." class="Magnoliopsida" family="Myrtaceae" genus="Rhodamnia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="11" pageNumber="42" phylum="Tracheophyta" rank="species" species="rubescens">
<emphasis italics="true" pageId="11" pageNumber="42">Rhodamnia rubescens</emphasis>
</taxonomicName>
(Benth.) Miq. (e.g., 
<bibRefCitation author="TPL, (The Plant List: A working list of all plant species)" journalOrPublisher="Sida" pageId="12" pageNumber="43" refId="B27" refString="TPL, (The Plant List: A working list of all plant species), 2012. http://www.theplantlist.org/; accessed 5 October 2012." title="http: // www. theplantlist. org /; accessed 5 October 2012." year="2012">TPL 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="42" type="specimen examined">
<paragraph pageId="11" pageNumber="42">Specimen examined.</paragraph>
<paragraph pageId="11" pageNumber="42">West Papua (Papua Barat; as Radjah Ampat on label), Waigeo Island, Waifoi on E bank of Majalibit [= Mayalibit] Bay, 18 Jan 1955, P. van Royen 5227 (L).</paragraph>
</subSubSection>
</treatment>
</document>