297 lines
36 KiB
XML
297 lines
36 KiB
XML
<document id="4AD126254ABE84621A2044324178AD7B" ID-DOI="10.5281/zenodo.183668" ID-GBIF-Dataset="27497a71-464b-4099-a848-921524e9beda" ID-ISSN="11755326" ID-Zenodo-Dep="183668" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459926501376" checkinUser="plazi" docAuthor="Brusa, Francisco, Damborenea, Cristina & Noreña, Carolina" docDate="2008" docId="B67787C93A27FFF9FF7DF85F6F8DF9E5" docLanguage="en" docName="zt01861p016.pdf" docOrigin="Zootaxa 1861" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Gieysztoria atalaya Brusa, Damborenea & Noreña, 2008, n. sp." docType="treatment" docVersion="10" lastPageNumber="6" masterDocId="4A4EFFB13A25FFFCFFEAFFCA6E3CF82F" masterDocTitle="“ Dalyellioida ” (Platyhelminthes, Rhabdocoela) from the Río de la Plata estuary in Argentina, with the description of two new species of Gieysztoria" masterLastPageNumber="16" masterPageNumber="1" pageNumber="3" updateTime="1698229560359" updateUser="plazi">
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<mods:mods id="DF7630487546F7427812CDA83FAE6BDA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F5C683768687707F30920FE100B2AB01">
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<mods:title id="DF747C521AA51E2F6AB54746EFCE4D94">“ Dalyellioida ” (Platyhelminthes, Rhabdocoela) from the Río de la Plata estuary in Argentina, with the description of two new species of Gieysztoria</mods:title>
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<mods:name id="B9FD950317A6D8F5CEE036BC676D41BA" type="personal">
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<mods:roleTerm id="6BF8A84BAE2AE18892DA4D61D716CA31">Author</mods:roleTerm>
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<mods:namePart id="5BDCD4498A35E99ECE37669DD7E241B0">Brusa, Francisco</mods:namePart>
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<mods:name id="AD3BA5ABB756BA38E67512B8DB0F75AF" type="personal">
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<mods:roleTerm id="47BCB3AE088F29077512A467113C5545">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1F40149472F05FB1836EB780D3504C29">Damborenea, Cristina</mods:namePart>
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</mods:name>
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<mods:name id="799DA63364BE4F1DC25EF74AA0526E76" type="personal">
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<mods:roleTerm id="EE43B24C61294A5B6FBC9CED10CF8E84">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8564DA7B3F9948897E738EF0A69C602A">Noreña, Carolina</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="61BB1FE46EBE9447FD4D1F0D9DABCD5F">text</mods:typeOfResource>
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<mods:title id="06475952ECCEDA8D9B83A0113DAE0D3A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="2D49BFD06A5673EBA25F7FCA86DE23EB">
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<mods:date id="314F86F2AB3FB145411BEBBAB0BCCE6D">2008</mods:date>
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<mods:detail id="E3F533D2FA7BCCE7630B2A889B98AA70" type="volume">
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<mods:number id="5185DBC8C4F3F90AD5FD00A3FC5AC5FB">1861</mods:number>
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<mods:start id="30DB497136545E09216941B055370D6C">1</mods:start>
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<mods:classification id="1EAD5639591C6BD284564E6C1BE9AFEC">journal article</mods:classification>
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<mods:identifier id="07C0A3E220E28DAF57D1F04D2A1F05F5" type="DOI">10.5281/zenodo.183668</mods:identifier>
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<mods:identifier id="154637DD88DE24376685406984A87DA6" type="ISSN">11755326</mods:identifier>
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<treatment id="B67787C93A27FFF9FF7DF85F6F8DF9E5" ID-DOI="http://doi.org/10.5281/zenodo.5618718" ID-GBIF-Taxon="119374344" ID-Zenodo-Dep="5618718" LSID="urn:lsid:plazi:treatment:B67787C93A27FFF9FF7DF85F6F8DF9E5" httpUri="http://treatment.plazi.org/id/B67787C93A27FFF9FF7DF85F6F8DF9E5" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
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<subSubSection id="76C465543A27FFFEFF7DF85F6F69FFF8" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="3E6136DF3A27FFFEFF7DF85F6FFAFF80" blockId="2.[151,454,1941,2007]" box="[151,454,1941,1967]" pageId="2" pageNumber="3">
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<heading id="652981B33A27FFFEFF7DF85F6FFAFF80" bold="true" box="[151,454,1941,1967]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<emphasis id="0CAAEACD3A27FFFEFF7DF85F6FFAFF80" bold="true" box="[151,454,1941,1967]" pageId="2" pageNumber="3">
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||
<taxonomicName id="F9DE4D5C3A27FFFEFF7DF85F6F41FF80" ID-CoL="3FZT9" box="[151,381,1941,1967]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="2" pageNumber="3" phylum="Platyhelminthes" rank="species" species="atalaya" status="sp. nov.">
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<emphasis id="0CAAEACD3A27FFFEFF7DF85F6F41FF80" bold="true" box="[151,381,1941,1967]" italics="true" pageId="2" pageNumber="3">Gieysztoria atalaya</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="179957B63A27FFFEFE6EF85F6FFAFF80" box="[388,454,1941,1967]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="3E6136DF3A27FFFEFF7DF8776F69FFF8" blockId="2.[151,454,1941,2007]" box="[151,341,1981,2007]" pageId="2" pageNumber="3">
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<figureCitation id="A6E52A5A3A27FFFEFF7DF8776F39FFF8" box="[151,261,1981,2007]" captionStart="FIGURE 1" captionStartId="4.[151,255,1915,1939]" captionTargetBox="[223,1405,281,1849]" captionTargetId="figure@4.[151,1436,260,1905]" captionTargetPageId="4" captionText="FIGURE 1. Gieysztoria atalaya n. sp. A, photograph of the stylet. B, dorsal general view in vivo. C, reconstruction of the stylet (a, b, and c indicate the three types of spines). D, sagittal reconstruction. Scales A and C: 50 m. Scales B and D: 100 m. For abbreviations see Figure 8." httpUri="https://zenodo.org/record/183669/files/figure.png" pageId="2" pageNumber="3">Figures 1</figureCitation>
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,
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<figureCitation id="A6E52A5A3A27FFFEFEF9F8776F1DFFF8" box="[275,289,1981,2007]" captionStart="FIGURE 7" captionStartId="13.[151,255,1842,1866]" captionTargetBox="[184,1411,197,1812]" captionTargetId="figure@13.[171,1415,194,1818]" captionTargetPageId="13" captionText="FIGURE 7. Microphotographs of sagittal sections of Gieysztoria hymanae. A, general view. B, detail of genital region. Microphotograps of Gieysztoria atalaya sp. n. Sagittal sections. C, general view. D, detail of digestive system. E, detail of genital region. F: horizontal section of stylet region. Scale A – F: 50 m. For abbreviations see Figure 8." httpUri="https://zenodo.org/record/183675/files/figure.png" pageId="2" pageNumber="3">7</figureCitation>
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C–F
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</paragraph>
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</subSubSection>
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<subSubSection id="76C465543A26FFFFFF7DFF5D6AEAF955" pageId="3" pageNumber="4" type="materials_examined">
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<paragraph id="3E6136DF3A26FFFFFF7DFF5D6BADF89D" blockId="3.[151,1439,151,2037]" box="[151,1425,151,178]" pageId="3" pageNumber="4">
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<emphasis id="0CAAEACD3A26FFFFFF7DFF5D6F37F89E" bold="true" box="[151,267,151,177]" pageId="3" pageNumber="4">Material.</emphasis>
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<typeStatus id="E165887D3A26FFFFFEF9FF526FB9F89D" box="[275,389,152,178]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
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: one specimen
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<emphasis id="0CAAEACD3A26FFFFFDDCFF526CBEF89E" box="[566,642,152,177]" italics="true" pageId="3" pageNumber="4">in toto</emphasis>
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mounted in polyvinyl–lactophenol
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<collectionCode id="58CFAE1A3A26FFFFFBCEFF526A5DF89D" box="[1060,1121,152,178]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="3" pageNumber="4">MLP</collectionCode>
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5420,
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<taxonomicName id="F9DE4D5C3A26FFFFFB5BFF526B30F89D" box="[1201,1292,152,178]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="14" phylum="Platyhelminthes" rank="species" species="atalaya" status="sp. nov.">Atalaya</taxonomicName>
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<date id="4A60101F3A26FFFFFAFEFF526BADF89D" box="[1300,1425,152,178]" pageId="3" pageNumber="4" value="1921-03-02">21–02–03.</date>
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</paragraph>
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<paragraph id="3E6136DF3A26FFFFFF2FFF0A6DC9F92D" blockId="3.[151,1439,151,2037]" pageId="3" pageNumber="4">
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<typeStatus id="E165887D3A26FFFFFF2FFF0A6F07F8F5" box="[197,315,192,218]" pageId="3" pageNumber="4" type="paratype">Paratypes</typeStatus>
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: Four specimens, sagittally cut
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<collectionCode id="58CFAE1A3A26FFFFFD59FF0A6CCCF8F5" box="[691,752,192,218]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="3" pageNumber="4">MLP</collectionCode>
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5340, 5341,
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<taxonomicName id="F9DE4D5C3A26FFFFFC65FF0A6DD6F8F5" box="[911,1002,192,218]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="14" phylum="Platyhelminthes" rank="species" species="atalaya" status="sp. nov.">Atalaya</taxonomicName>
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<date id="4A60101F3A26FFFFFC19FF0A6A57F8F5" box="[1011,1131,192,218]" pageId="3" pageNumber="4" value="2001-03-05">05–03–01</date>
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,
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<date id="4A60101F3A26FFFFFB92FF0A6AD7F8F5" box="[1144,1259,192,218]" pageId="3" pageNumber="4" value="2002-02-02">020202.</date>
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One specimen
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<emphasis id="0CAAEACD3A26FFFFFF7DFF226EDFF92E" box="[151,227,232,257]" italics="true" pageId="3" pageNumber="4">in toto</emphasis>
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mounted in polyvinyllactophenol,
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<collectionCode id="58CFAE1A3A26FFFFFD62FF226CF8F92D" box="[648,708,232,258]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="3" pageNumber="4">MLP</collectionCode>
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5421,
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<taxonomicName id="F9DE4D5C3A26FFFFFCFFFF226D4CF92D" box="[789,880,232,258]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="14" phylum="Platyhelminthes" rank="species" species="atalaya" status="sp. nov.">Atalaya</taxonomicName>
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<date id="4A60101F3A26FFFFFC92FF226DC9F92D" box="[888,1013,232,258]" pageId="3" pageNumber="4" value="1910-03-04">10–04–03.</date>
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</paragraph>
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<paragraph id="3E6136DF3A26FFFFFF2FFEC56C44F97D" blockId="3.[151,1439,151,2037]" pageId="3" pageNumber="4">
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<emphasis id="0CAAEACD3A26FFFFFF2FFEC56FB6F906" bold="true" box="[197,394,271,297]" pageId="3" pageNumber="4">Other material:</emphasis>
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Fifteen specimens, sagittally cut
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<collectionCode id="58CFAE1A3A26FFFFFCE5FEDA6D77F905" box="[783,843,272,298]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="3" pageNumber="4">MLP</collectionCode>
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5342, 5343, 5344, 5723, 5724,
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<taxonomicName id="F9DE4D5C3A26FFFFFB57FEDA6B24F905" box="[1213,1304,272,298]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="14" phylum="Platyhelminthes" rank="species" species="atalaya" status="sp. nov.">Atalaya</taxonomicName>
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<date id="4A60101F3A26FFFFFAF5FEDA6BABF905" box="[1311,1431,272,298]" pageId="3" pageNumber="4" value="2001-03-05">05–03–01</date>
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,
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<date id="4A60101F3A26FFFFFF7DFEF26F39F97D" box="[151,261,312,338]" pageId="3" pageNumber="4" value="2002-02-02">020202</date>
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,
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<date id="4A60101F3A26FFFFFEFBFEF26F43F97D" box="[273,383,312,338]" pageId="3" pageNumber="4" value="2002-04-04">040402</date>
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,
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<date id="4A60101F3A26FFFFFE61FEF26FC5F97D" box="[395,505,312,338]" pageId="3" pageNumber="4" value="1921-04-02" valueMax="1921-04-03" valueMin="1921-04-02">210203</date>
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,
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<date id="4A60101F3A26FFFFFDEFFEF26C44F97D" box="[517,632,312,338]" pageId="3" pageNumber="4" value="1910-03-04">100403.</date>
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</paragraph>
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<paragraph id="3E6136DF3A26FFFFFF2FFE956AEAF955" blockId="3.[151,1439,151,2037]" box="[197,1238,351,378]" pageId="3" pageNumber="4">
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<emphasis id="0CAAEACD3A26FFFFFF2FFE956F57F956" bold="true" box="[197,363,351,377]" pageId="3" pageNumber="4">
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<typeStatus id="E165887D3A26FFFFFF2FFE956F3EF956" box="[197,258,351,377]" pageId="3" pageNumber="4">Type</typeStatus>
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locality.
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</emphasis>
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<taxonomicName id="F9DE4D5C3A26FFFFFE98FEAA6FF1F955" box="[370,461,352,378]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="14" phylum="Platyhelminthes" rank="species" species="atalaya" status="sp. nov.">Atalaya</taxonomicName>
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(
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<geoCoordinate id="5BEA50183A26FFFFFE37FEAA6C40F955" box="[477,636,352,378]" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="1" value="-35.01489">35º00’53.6”S</geoCoordinate>
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–
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<geoCoordinate id="5BEA50183A26FFFFFD67FEAA6D15F955" box="[653,809,352,378]" direction="west" orientation="longitude" pageId="3" pageNumber="4" precision="1" value="-57.53425">57º32’3.3”W</geoCoordinate>
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), Buenos Aires province,
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<collectingCountry id="46C9764F3A26FFFFFBB3FEAA6AEDF955" box="[1113,1233,352,378]" name="Argentina" pageId="3" pageNumber="4">Argentina</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="76C465543A26FFFFFF2FFE4D6D61F98D" box="[197,861,391,418]" pageId="3" pageNumber="4" type="etymology">
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<paragraph id="3E6136DF3A26FFFFFF2FFE4D6D61F98D" blockId="3.[151,1439,151,2037]" box="[197,861,391,418]" pageId="3" pageNumber="4">
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<emphasis id="0CAAEACD3A26FFFFFF2FFE4D6F6EF98E" bold="true" box="[197,338,391,417]" pageId="3" pageNumber="4">Etymology.</emphasis>
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The species name refers to the
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<typeStatus id="E165887D3A26FFFFFD2FFE426CC4F98D" box="[709,760,392,418]" pageId="3" pageNumber="4">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="76C465543A26FFFFFF2FFE656DF9FE39" pageId="3" pageNumber="4" type="description">
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<paragraph id="3E6136DF3A26FFFFFF2FFE656A41FA35" blockId="3.[151,1439,151,2037]" pageId="3" pageNumber="4">
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<emphasis id="0CAAEACD3A26FFFFFF2FFE656F61F9E6" bold="true" box="[197,349,431,457]" pageId="3" pageNumber="4">Description.</emphasis>
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Live adult specimens 430–575 µm long and 95–115 µm wide. Anterior end rounded. Five or six caudal papillae at abruptly tapering posterior end. Rhabdites arranged in groups of 3–4 along body. Rigid cilia at anterior and posterior ends of body longer than those on remaining epidermis.
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</paragraph>
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<paragraph id="3E6136DF3A26FFFFFF2FFDE26C74FA46" blockId="3.[151,1439,151,2037]" pageId="3" pageNumber="4">
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Pharynx
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<emphasis id="0CAAEACD3A26FFFFFEC5FDE26F42FA6E" box="[303,382,552,577]" italics="true" pageId="3" pageNumber="4">in vivo</emphasis>
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130 µm long and 100 µm wide, with four dorsal papillae at anterior margin of pharynx in contrast to other
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<taxonomicName id="F9DE4D5C3A26FFFFFEB6FD856FD8FA47" box="[348,484,591,616]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="4" phylum="Platyhelminthes" rank="genus">
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<emphasis id="0CAAEACD3A26FFFFFEB6FD856FD8FA47" box="[348,484,591,616]" italics="true" pageId="3" pageNumber="4">Gieysztoria</emphasis>
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</taxonomicName>
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species.
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</paragraph>
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<paragraph id="3E6136DF3A26FFFFFF2FFDBC6C8BFAF1" blockId="3.[151,1439,151,2037]" pageId="3" pageNumber="4">Broad nerve tracts from brain to anterior region observed in sagittal sections. Conspicuous nucleus evident at peripheral anterior and posterior regions of brain. Black eyes formed by very numerous pigment spheres arranged in a kidneyshaped structure.</paragraph>
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<paragraph id="3E6136DF3A26FFFFFF2FFD266FCEFB54" blockId="3.[151,1439,151,2037]" pageId="3" pageNumber="4">Male reproductive system formed by two noncompact testes situated in posterior body region. Spermatozoids observed at periphery of testes. Vasa deferentia projecting from rostral portion of testes toward anterior region and apically or subapically entering seminal vesicle; the latter relatively small and continuous with similarsized prostate vesicle.</paragraph>
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<paragraph id="3E6136DF3A26FFFFFF2FFC436F50FCF3" blockId="3.[151,1439,151,2037]" pageId="3" pageNumber="4">
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Sclerotic stylet approximately 62 µm long and 30 µm wide, with welldefined fibrouslike proximal belt showing a clear window on one side (figure 1A, C). Belt open at side opposite window. Three
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<typeStatus id="E165887D3A26FFFFFB1AFC7A6B12FBE5" box="[1264,1326,944,970]" pageId="3" pageNumber="4">types</typeStatus>
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of spines originating from belt. A) One spine (spine “a” in figure 1C, length 23 µm) issuing from under window – broad at the base and tapering towards blunt distal end. Spine hollow, situated between the other two groups of spines. B) On one side of stylet, a group of approximately ten thick spines (spines “b” in figure 1C), originating from belt margin, all of similar shape and length. Very thin spines scattered among them. C) On other side of spine “a”, a third group of spines (spines “c” in figure 1C) arranged in several rows forming a “brush”. Approximately 10 spines in proximal row, with broad base and rapidly tapering so as to present triangular shape (figure 1C).
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</paragraph>
|
||
<paragraph id="3E6136DF3A26FFFFFF2FFB236DF9FE39" blockId="3.[151,1439,151,2037]" pageId="3" pageNumber="4">Female reproductive system comprising an ovary located on right side of body and dorsal to intestine. Oocytes arranged in two or three series at proximal part of ovary, and uniserially at distal part (figure 1D). Ovary continued into a short oviduct; oviduct widening to form a seminal receptacle connected to uterus by a long common duct. Uterus with thick walls, curved to open into the genital atrium. Egg in uterus large (greater diameter 152 µm, lesser diameter 93 µm), oval, with operculum at one of its ends. Vitellaria smooth (figure 1B); anteriorly reaching base of pharynx and ventral at this level; becoming dorsal at posterior end, there merging to enter the common duct. Bursa copulatrix with very thick muscular walls, opening into common duct. Gonopore surrounded by a strong sphyncter and cement glands.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="76C465543A26FFF9FF2FF9E86F8DF9E5" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="discussion">
|
||
<paragraph id="3E6136DF3A26FFFFFF2FF9E86DD9FF52" blockId="3.[151,1439,151,2037]" pageId="3" pageNumber="4">
|
||
<emphasis id="0CAAEACD3A26FFFFFF2FF9E86F6DFE13" bold="true" box="[197,337,1570,1596]" pageId="3" pageNumber="4">Discussion.</emphasis>
|
||
Based mainly on the configuration and shape of the male stylet,
|
||
<taxonomicName id="F9DE4D5C3A26FFFFFB80F9E96AD9FE13" box="[1130,1253,1571,1596]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="4" phylum="Platyhelminthes" rank="species" species="atalaya">
|
||
<emphasis id="0CAAEACD3A26FFFFFB80F9E96AD9FE13" box="[1130,1253,1571,1596]" italics="true" pageId="3" pageNumber="4">G. atalaya</emphasis>
|
||
</taxonomicName>
|
||
could be compared with three species:
|
||
<taxonomicName id="F9DE4D5C3A26FFFFFE22F9816AB7FE4A" authority="(Beklemischev 1926) Luther 1955" authorityName="(Beklemischev 1926) Luther" authorityYear="1955" box="[456,1163,1611,1637]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="4" phylum="Platyhelminthes" rank="species" species="pavimentata">
|
||
<emphasis id="0CAAEACD3A26FFFFFE22F9816CD5FE4B" box="[456,745,1611,1636]" italics="true" pageId="3" pageNumber="4">Gieysztoria pavimentata</emphasis>
|
||
(
|
||
<bibRefCitation id="5A4F4B2E3A26FFFFFD10F9816DD4FE4A" author="Beklemischev" box="[762,1000,1611,1637]" pageId="3" pageNumber="16" refString="Beklemischev, V. N. (1926) Sur la morphologie de l´organe copulateur du genre Dalyellia. 1. D. virgulifera (Plotn.) et D. pavimentata n. sp. [in Russian]. Bulleti ∏ de l ' I ∏ Ƽtitut deƼ reCherCheƼ Ki 0 l 0 gifueƼ de Perm. 4, 237 - 248." type="journal article" year="1926">Beklemischev 1926</bibRefCitation>
|
||
)
|
||
<bibRefCitation id="5A4F4B2E3A26FFFFFC13F9816AB7FE4A" author="Luther" box="[1017,1163,1611,1637]" pageId="3" pageNumber="16" refString="Luther, A. (1955) Die Dalyelliiden (Turbellaria Neorhabdocoela) Eine Monographie. Acta Zoologica Fennica, 87, 1 - 337." type="journal article" year="1955">Luther 1955</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F9DE4D5C3A26FFFFFB71F9816F94FEA2" authority="(Plotnikov 1906) Luther 1955" authorityName="(Plotnikov 1906) Luther" authorityYear="1955" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="4" phylum="Platyhelminthes" rank="species" species="virgulifera">
|
||
<emphasis id="0CAAEACD3A26FFFFFB71F9816B06FE4B" box="[1179,1338,1611,1636]" italics="true" pageId="3" pageNumber="4">G. virgulifera</emphasis>
|
||
(
|
||
<bibRefCitation id="5A4F4B2E3A26FFFFFAA0F9816F3BFEA2" author="Plotnikov" pageId="3" pageNumber="16" refString="Plotnikov, W. (1906) Die rhabdocolen Turbellarien der Umgebung des Goktscha-Sees. Zoologische Jahrbucher, Systematik. Abteilung fur Systematik, Okologie und Geographie der Tiere, 23, 395 - 400, t. 22." type="journal article" year="1906">Plotnikov 1906</bibRefCitation>
|
||
)
|
||
<bibRefCitation id="5A4F4B2E3A26FFFFFEFDF9B96F94FEA2" author="Luther" box="[279,424,1651,1677]" pageId="3" pageNumber="16" refString="Luther, A. (1955) Die Dalyelliiden (Turbellaria Neorhabdocoela) Eine Monographie. Acta Zoologica Fennica, 87, 1 - 337." type="journal article" year="1955">Luther 1955</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F9DE4D5C3A26FFFFFE08F9B96DF4FEA2" authority="(Gieysztor 1931) Luther 1955" authorityName="(Gieysztor 1931) Luther" authorityYear="1955" box="[482,968,1651,1677]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="4" phylum="Platyhelminthes" rank="species" species="beltrani">
|
||
<emphasis id="0CAAEACD3A26FFFFFE08F9B96C5CFEA3" box="[482,608,1651,1676]" italics="true" pageId="3" pageNumber="4">G. beltrani</emphasis>
|
||
(
|
||
<bibRefCitation id="5A4F4B2E3A26FFFFFD9AF9B96D1BFEA2" author="Gieysztor" box="[624,807,1651,1677]" pageId="3" pageNumber="16" refString="Gieysztor, M. (1931) Contribution a la connaissance des Turbellaries Rhabdocoeles (Turbellaria Rhabdocoela) d'Espagne. Bulletin International de L'Academie Polonaise des Sciences et des Lettres. Classe des Sciences Mathematiques et Naturelles. Serie B. Sciences Naturelles, 2, 125 - 153, t. 13 - 14." type="journal article" year="1931">Gieysztor 1931</bibRefCitation>
|
||
)
|
||
<bibRefCitation id="5A4F4B2E3A26FFFFFCDDF9B96DF4FEA2" author="Luther" box="[823,968,1651,1677]" pageId="3" pageNumber="16" refString="Luther, A. (1955) Die Dalyelliiden (Turbellaria Neorhabdocoela) Eine Monographie. Acta Zoologica Fennica, 87, 1 - 337." type="journal article" year="1955">Luther 1955</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="F9DE4D5C3A26FFFFFC32F9B96AC4FEA3" box="[984,1272,1651,1676]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="4" phylum="Platyhelminthes" rank="species" species="pavimentata">
|
||
<emphasis id="0CAAEACD3A26FFFFFC32F9B96AC4FEA3" box="[984,1272,1651,1676]" italics="true" pageId="3" pageNumber="4">Gieysztoria pavimentata</emphasis>
|
||
</taxonomicName>
|
||
is the species with the most similar stylet to that of the new species.
|
||
<taxonomicName id="F9DE4D5C3A26FFFFFCDAF9516DDFFE9B" box="[816,995,1691,1716]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="4" phylum="Platyhelminthes" rank="species" species="pavimentata">
|
||
<emphasis id="0CAAEACD3A26FFFFFCDAF9516DDFFE9B" box="[816,995,1691,1716]" italics="true" pageId="3" pageNumber="4">G. pavimentata</emphasis>
|
||
</taxonomicName>
|
||
was considered by
|
||
<bibRefCitation id="5A4F4B2E3A26FFFFFB3EF9516B46FE9A" author="Luther" box="[1236,1402,1691,1717]" pageId="3" pageNumber="16" refString="Luther, A. (1955) Die Dalyelliiden (Turbellaria Neorhabdocoela) Eine Monographie. Acta Zoologica Fennica, 87, 1 - 337." type="journal article" year="1955">Luther (1955)</bibRefCitation>
|
||
as belonging to the Inaequales group, subgroup Fenestrate. Both species possess a robust spine, but in
|
||
<taxonomicName id="F9DE4D5C3A26FFFFFACEF9096BA0FEF3" box="[1316,1436,1731,1756]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="4" phylum="Platyhelminthes" rank="species" species="atalaya">
|
||
<emphasis id="0CAAEACD3A26FFFFFACEF9096BA0FEF3" box="[1316,1436,1731,1756]" italics="true" pageId="3" pageNumber="4">G. atalaya</emphasis>
|
||
</taxonomicName>
|
||
this spine occupies a central position on the girdle (spine “a” figure 1C) whereas in
|
||
<taxonomicName id="F9DE4D5C3A26FFFFFB46F9216B5DFF2B" box="[1196,1377,1771,1796]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="4" phylum="Platyhelminthes" rank="species" species="pavimentata">
|
||
<emphasis id="0CAAEACD3A26FFFFFB46F9216B5DFF2B" box="[1196,1377,1771,1796]" italics="true" pageId="3" pageNumber="4">G. pavimentata</emphasis>
|
||
</taxonomicName>
|
||
it is located on the right side of the stylet (spine “d” of
|
||
<bibRefCitation id="5A4F4B2E3A26FFFFFD0BF8D96D4FFF02" author="Luther" box="[737,883,1811,1837]" pageId="3" pageNumber="16" refString="Luther, A. (1955) Die Dalyelliiden (Turbellaria Neorhabdocoela) Eine Monographie. Acta Zoologica Fennica, 87, 1 - 337." type="journal article" year="1955">Luther 1955</bibRefCitation>
|
||
, figure 41 G, H and 42B). In addition to the different location, the central spine of
|
||
<taxonomicName id="F9DE4D5C3A26FFFFFDAFF8F16C83FF7B" box="[581,703,1851,1876]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="4" phylum="Platyhelminthes" rank="species" species="atalaya">
|
||
<emphasis id="0CAAEACD3A26FFFFFDAFF8F16C83FF7B" box="[581,703,1851,1876]" italics="true" pageId="3" pageNumber="4">G. atalaya</emphasis>
|
||
</taxonomicName>
|
||
is clearly larger and differs in shape from the rest of spines, which is not the case of spine “d” (after
|
||
<bibRefCitation id="5A4F4B2E3A26FFFFFD84F8A96CC3FF52" author="Luther" box="[622,767,1891,1917]" pageId="3" pageNumber="16" refString="Luther, A. (1955) Die Dalyelliiden (Turbellaria Neorhabdocoela) Eine Monographie. Acta Zoologica Fennica, 87, 1 - 337." type="journal article" year="1955">Luther 1955</bibRefCitation>
|
||
) in
|
||
<taxonomicName id="F9DE4D5C3A26FFFFFCC7F8A96DE2FF53" box="[813,990,1891,1916]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="4" phylum="Platyhelminthes" rank="species" species="pavimentata">
|
||
<emphasis id="0CAAEACD3A26FFFFFCC7F8A96DE2FF53" box="[813,990,1891,1916]" italics="true" pageId="3" pageNumber="4">G. pavimentata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3E6136DF3A26FFF8FF2FF8416C5AF8F5" blockId="3.[151,1439,151,2037]" lastBlockId="4.[151,1436,152,218]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
|
||
Furthermore, the constellation of the smaller spines of
|
||
<taxonomicName id="F9DE4D5C3A26FFFFFCA3F8416DC6FF8B" box="[841,1018,1931,1956]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="4" phylum="Platyhelminthes" rank="species" species="pavimentata">
|
||
<emphasis id="0CAAEACD3A26FFFFFCA3F8416DC6FF8B" box="[841,1018,1931,1956]" italics="true" pageId="3" pageNumber="4">G. pavimentata</emphasis>
|
||
</taxonomicName>
|
||
is different from that of
|
||
<taxonomicName id="F9DE4D5C3A26FFFFFAF6F8416BA9FF8B" box="[1308,1429,1931,1956]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="4" phylum="Platyhelminthes" rank="species" species="atalaya">
|
||
<emphasis id="0CAAEACD3A26FFFFFAF6F8416BA9FF8B" box="[1308,1429,1931,1956]" italics="true" pageId="3" pageNumber="4">G. atalaya</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="F9DE4D5C3A26FFFFFF7DF8796F6BFFE3" box="[151,343,1971,1996]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="4" phylum="Platyhelminthes" rank="species" species="pavimentanta">
|
||
<emphasis id="0CAAEACD3A26FFFFFF7DF8796F6BFFE3" box="[151,343,1971,1996]" italics="true" pageId="3" pageNumber="4">G. pavimentanta</emphasis>
|
||
</taxonomicName>
|
||
bears 10 spines on the girdle edge, followed by 2 or 3 irregular rows of spines on the wall of the copulatory organ. In contrast, the spines of
|
||
<taxonomicName id="F9DE4D5C3A26FFFFFD22F8116D7DFFDB" box="[712,833,2011,2036]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="3" pageNumber="4" phylum="Platyhelminthes" rank="species" species="atalaya">
|
||
<emphasis id="0CAAEACD3A26FFFFFD22F8116D7DFFDB" box="[712,833,2011,2036]" italics="true" pageId="3" pageNumber="4">G. atalaya</emphasis>
|
||
</taxonomicName>
|
||
are arranged in two groups, the first group (spines “b”) formed by about 10 spines and the second group (spines “c”) formed by three or four rows of thin spines with similar shape, but different length.
|
||
</paragraph>
|
||
<caption id="6AA166573A21FFF8FF7DF8B16C0DFFF7" httpUri="https://zenodo.org/record/183669/files/figure.png" pageId="4" pageNumber="5" targetBox="[223,1405,281,1849]" targetPageId="4">
|
||
<paragraph id="3E6136DF3A21FFF8FF7DF8B16C0DFFF7" blockId="4.[151,1436,1915,2008]" pageId="4" pageNumber="5">
|
||
<emphasis id="0CAAEACD3A21FFF8FF7DF8B16F26FFBC" bold="true" box="[151,282,1915,1939]" pageId="4" pageNumber="5">FIGURE 1.</emphasis>
|
||
<taxonomicName id="F9DE4D5C3A21FFF8FEC8F8B66FC9FFBC" box="[290,501,1916,1939]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="species" species="atalaya" status="sp. nov.">
|
||
<emphasis id="0CAAEACD3A21FFF8FEC8F8B66FC9FFBC" box="[290,501,1916,1939]" italics="true" pageId="4" pageNumber="5">Gieysztoria atalaya</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0CAAEACD3A21FFF8FE16F8B16C06FFBC" bold="true" box="[508,570,1915,1939]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="179957B63A21FFF8FE16F8B16C06FFBC" box="[508,570,1915,1939]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A, photograph of the stylet. B, dorsal general view
|
||
<emphasis id="0CAAEACD3A21FFF8FB84F8B66A8AFFBC" box="[1134,1206,1916,1939]" italics="true" pageId="4" pageNumber="5">in vivo</emphasis>
|
||
. C, reconstruction of the stylet (a, b, and c indicate the three types of spines). D, sagittal reconstruction. Scales A and C: 50 m. Scales B and D: 100 m. For abbreviations see Figure 8.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3E6136DF3A20FFF9FF2FFF526F8DF9E5" blockId="5.[151,1436,152,458]" pageId="5" pageNumber="6">
|
||
The stylets of
|
||
<taxonomicName id="F9DE4D5C3A20FFF9FE86FF526C6AF89E" box="[364,598,152,177]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="beltrani">
|
||
<emphasis id="0CAAEACD3A20FFF9FE86FF526C6AF89E" box="[364,598,152,177]" italics="true" pageId="5" pageNumber="6">Gieysztoria beltrani</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F9DE4D5C3A20FFF9FD65FF526D17F89E" box="[655,811,152,177]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="virgulifera">
|
||
<emphasis id="0CAAEACD3A20FFF9FD65FF526D17F89E" box="[655,811,152,177]" italics="true" pageId="5" pageNumber="6">G. virgulifera</emphasis>
|
||
</taxonomicName>
|
||
are also similar to that of
|
||
<taxonomicName id="F9DE4D5C3A20FFF9FBB6FF526AEFF89E" box="[1116,1235,152,177]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="atalaya">
|
||
<emphasis id="0CAAEACD3A20FFF9FBB6FF526AEFF89E" box="[1116,1235,152,177]" italics="true" pageId="5" pageNumber="6">G. atalaya</emphasis>
|
||
</taxonomicName>
|
||
; both have a belt with a middle window and a large spine issuing from under it. In
|
||
<taxonomicName id="F9DE4D5C3A20FFF9FC79FF0A6A2CF8F6" box="[915,1040,192,217]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="beltrani">
|
||
<emphasis id="0CAAEACD3A20FFF9FC79FF0A6A2CF8F6" box="[915,1040,192,217]" italics="true" pageId="5" pageNumber="6">G. beltrani</emphasis>
|
||
</taxonomicName>
|
||
the large spine under the window (
|
||
<bibRefCitation id="5A4F4B2E3A20FFF9FF75FF226F64F92D" author="Gieysztor" box="[159,344,232,258]" pageId="5" pageNumber="16" refString="Gieysztor, M. (1931) Contribution a la connaissance des Turbellaries Rhabdocoeles (Turbellaria Rhabdocoela) d'Espagne. Bulletin International de L'Academie Polonaise des Sciences et des Lettres. Classe des Sciences Mathematiques et Naturelles. Serie B. Sciences Naturelles, 2, 125 - 153, t. 13 - 14." type="journal article" year="1931">Gieysztor 1931</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="5A4F4B2E3A20FFF9FE8EFF226FC9F92D" author="Luther" box="[356,501,232,258]" pageId="5" pageNumber="16" refString="Luther, A. (1955) Die Dalyelliiden (Turbellaria Neorhabdocoela) Eine Monographie. Acta Zoologica Fennica, 87, 1 - 337." type="journal article" year="1955">Luther 1955</bibRefCitation>
|
||
) bears small spines, contrasting with the smooth large spine of
|
||
<taxonomicName id="F9DE4D5C3A20FFF9FB37FF226B69F92E" box="[1245,1365,232,257]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="atalaya">
|
||
<emphasis id="0CAAEACD3A20FFF9FB37FF226B69F92E" box="[1245,1365,232,257]" italics="true" pageId="5" pageNumber="6">G. atalaya</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="F9DE4D5C3A20FFF9FA69FF226EC8F906" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="beltrani">
|
||
<emphasis id="0CAAEACD3A20FFF9FA69FF226EC8F906" italics="true" pageId="5" pageNumber="6">G. beltrani</emphasis>
|
||
</taxonomicName>
|
||
, the rest of the spines are grouped at the sides of the large spine, with six spines on one side and a variable number on the other.
|
||
<emphasis id="0CAAEACD3A20FFF9FE12FEF26CADF97E" box="[504,657,312,337]" italics="true" pageId="5" pageNumber="6">G. v i rg u l i f e r a</emphasis>
|
||
has two groups of spines at both sides of the smooth large central spine, similarly to the condition in
|
||
<taxonomicName id="F9DE4D5C3A20FFF9FDDDFEAA6C8DF956" box="[567,689,352,377]" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="atalaya">
|
||
<emphasis id="0CAAEACD3A20FFF9FDDDFEAA6C8DF956" box="[567,689,352,377]" italics="true" pageId="5" pageNumber="6">G. atalaya</emphasis>
|
||
</taxonomicName>
|
||
, but the groups differ in number and shape of the spines. In
|
||
<taxonomicName id="F9DE4D5C3A20FFF9FA69FEAA6F28F98E" class="Rhabditophora" family="Dalyelliidae" genus="Gieysztoria" kingdom="Animalia" order="Rhabdocoela" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="virgulifera">
|
||
<emphasis id="0CAAEACD3A20FFF9FA69FEAA6F28F98E" italics="true" pageId="5" pageNumber="6">G. virgulifera</emphasis>
|
||
</taxonomicName>
|
||
the first group is formed by 4–7 spines on one side and only two robust, scimitarshaped spines, on the other (
|
||
<bibRefCitation id="5A4F4B2E3A20FFF9FEE7FE7A6F9FF9E5" author="Luther" box="[269,419,432,458]" pageId="5" pageNumber="16" refString="Luther, A. (1955) Die Dalyelliiden (Turbellaria Neorhabdocoela) Eine Monographie. Acta Zoologica Fennica, 87, 1 - 337." type="journal article" year="1955">Luther 1955</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |