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<document ID-DOI="http://dx.doi.org/10.3897/dez.69.84080" ID-Pensoft-Pub="1860-1324-2-151" ID-Pensoft-UUID="B7D8CC48BD32502C819369D75ADA50C8" ID-ZooBank="3B245B5371564A3F96672F2CD756779A" ModsDocID="1860-1324-69-2-151" checkinTime="1658871268840" checkinUser="pensoft" docAuthor="Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia &amp; Prous, Marko" docDate="2022" docId="4D49FDB220E1535699067E44863869D5" docLanguage="en" docName="DeutEntomolZeit 69(2): 151-218" docOrigin="Deutsche Entomologische Zeitschrift 69 (2)" docPubDate="2022-07-26" docSource="http://dx.doi.org/10.3897/dez.69.84080" docTitle="Pristiphora coactula" docType="treatment" docVersion="1" id="B7D8CC48BD32502C819369D75ADA50C8" lastPageNumber="151" masterDocId="B7D8CC48BD32502C819369D75ADA50C8" masterDocTitle="Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta)" masterLastPageNumber="218" masterPageNumber="151" pageNumber="151" updateTime="1658871268840" updateUser="pensoft">
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<mods:title>Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta)</mods:title>
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<mods:namePart>Liston, Andrew</mods:namePart>
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<mods:affiliation>Senckenberg Deutsches Entomologisches Institut, Eberswalder Str. 90, 15374 Muencheberg, Germany</mods:affiliation>
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<mods:namePart>Mutanen, Marko</mods:namePart>
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<mods:affiliation>Ecology and Genetics Research Unit, PO Box 3000, 90014 University of Oulu, Oulu, Finland</mods:affiliation>
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<mods:namePart>Heidemaa, Mikk</mods:namePart>
<mods:affiliation>Estonian Naturalists' Society, Struve 2, Tartu 51003, Estonia</mods:affiliation>
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<mods:namePart>Blank, Stephan M.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kiljunen, Niina</mods:namePart>
<mods:affiliation>Ecology and Genetics Research Unit, PO Box 3000, 90014 University of Oulu, Oulu, Finland</mods:affiliation>
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<mods:namePart>Taeger, Andreas</mods:namePart>
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<mods:namePart>Viitasaari, Matti</mods:namePart>
<mods:affiliation>Alkutie 41 E, 00660 Helsinki, Finland</mods:affiliation>
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<mods:namePart>Vikberg, Veli</mods:namePart>
<mods:affiliation>Liinalammintie 11 as. 6, 14200 Turenki, Finland</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wutke, Saskia</mods:namePart>
<mods:affiliation>Department of Environmental and Biological Sciences, University of Eastern Finland, 80101 Joensuu, Finland</mods:affiliation>
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<mods:namePart>Prous, Marko</mods:namePart>
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<mods:affiliation>Senckenberg Deutsches Entomologisches Institut, Eberswalder Str. 90, 15374 Muencheberg, Germany &amp; Ecology and Genetics Research Unit, PO Box 3000, 90014 University of Oulu, Oulu, Finland &amp; Department of Zoology, Institute of Ecology and Earth Sciences, University of Tartu, Vanemuise 46, 51014 Tartu, Estonia</mods:affiliation>
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<mods:date>2022</mods:date>
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<treatment LSID="urn:lsid:plazi:treatment:4D49FDB220E1535699067E44863869D5" httpUri="http://treatment.plazi.org/id/4D49FDB220E1535699067E44863869D5" lastPageNumber="151" pageId="0" pageNumber="151">
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<taxonomicName LSID="4D49FDB2-20E1-5356-9906-7E44863869D5" authority="(Ruthe, 1859)" baseAuthorityName="Ruthe" baseAuthorityYear="1859" class="Insecta" family="Tenthredinidae" genus="Pristiphora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristiphora coactula" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="coactula">Pristiphora coactula (Ruthe, 1859)</taxonomicName>
</paragraph>
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<paragraph pageId="0" pageNumber="151">
<taxonomicName authorityName="Ruthe" authorityYear="1859" class="Insecta" family="Tenthredinidae" genus="Nematus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nematus coactulus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="coactulus">Nematus coactulus</taxonomicName>
Ruthe, 1859: 307-308. ♀. Holotype. Type locality: Iceland.
</paragraph>
<paragraph pageId="0" pageNumber="151">
<taxonomicName authority="Konow, 1890" authorityName="Konow" authorityYear="1890" class="Insecta" family="Tenthredinidae" genus="Lygaeonematus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lygaeonematus (Lygaeotus) trochantericus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="trochantericus" subGenus="Lygaeotus">Lygaeonematus (Lygaeotus) trochantericus</taxonomicName>
Lindqvist, 1952: 101-102. ♀. Holotype (http://id.luomus.fi/GL.7708) and paratypes (♀, ♂). Type locality: Finland, Utsjoki, Outakoski. syn. nov.
</paragraph>
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<subSubSection pageId="0" pageNumber="151" type="notes">
<paragraph pageId="0" pageNumber="151">Notes.</paragraph>
<paragraph pageId="0" pageNumber="151">
The nuclear sequence data obtained for this study revealed three main clusters within the 
<taxonomicName authorityName="Hartig" authorityYear="1837" baseAuthorityName="Hartig" baseAuthorityYear="1837" class="Insecta" family="Tenthredinidae" genus="Pristiphora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristiphora carinata" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="151">Pristiphora carinata</emphasis>
</taxonomicName>
group: 
<taxonomicName genus="P." lsidName="P. carinata" pageId="0" pageNumber="151" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="151">P. carinata</emphasis>
</taxonomicName>
, 
<taxonomicName genus="P." lsidName="P. coactula" pageId="0" pageNumber="151" rank="species" species="coactula">
<emphasis italics="true" pageId="0" pageNumber="151">P. coactula</emphasis>
</taxonomicName>
, and 
<emphasis italics="true" pageId="0" pageNumber="151">
<taxonomicName genus="P." lsidName="P. borea" pageId="0" pageNumber="151" rank="species" species="borea">P. borea</taxonomicName>
+ 
<taxonomicName genus="P." lsidName="P. groenblomi" pageId="0" pageNumber="151" rank="species" species="groenblomi">P. groenblomi</taxonomicName>
+ 
<taxonomicName genus="P." lsidName="P. albilabris" pageId="0" pageNumber="151" rank="species" species="albilabris">P. albilabris</taxonomicName>
</emphasis>
(Fig. 
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Maximum likelihood tree of Pristiphora carinata group based on nuclear genes (NaK and POL 2). Numbers at branches show SH-aLRT support (%) / ultrafast bootstrap support (%) values. Values of only well supported branches (&gt; 90 for both) and of P. coactula clade with moderate support are shown. Letters &quot; f &quot; stand for &quot; female &quot;, &quot; m &quot; for &quot; male &quot;, and &quot; l &quot; for larva. Numbers at the end of the tip labels refer to sequence length and the number of heterozygous positions." figureDoi="10.3897/dez.69.84080.figure30" httpUri="https://binary.pensoft.net/fig/719541" pageId="0" pageNumber="151">30</figureCitation>
). No nuclear sequence data are yet available for 
<taxonomicName genus="P." lsidName="P. breadalbanensis" pageId="0" pageNumber="151" rank="species" species="breadalbanensis">
<emphasis italics="true" pageId="0" pageNumber="151">P. breadalbanensis</emphasis>
</taxonomicName>
(Cameron, 1882b) and 
<taxonomicName genus="P." lsidName="P. lativentris" pageId="0" pageNumber="151" rank="species" species="lativentris">
<emphasis italics="true" pageId="0" pageNumber="151">P. lativentris</emphasis>
</taxonomicName>
(Thomson, 1871). A nearly perfect match morphologically to the 
<taxonomicName genus="L." lsidName="L. trochantericus" pageId="0" pageNumber="151" rank="species" species="trochantericus">
<emphasis italics="true" pageId="0" pageNumber="151">L. trochantericus</emphasis>
</taxonomicName>
holotype is ZMUO.035514, which falls within the 
<taxonomicName genus="P." lsidName="P. coactula" pageId="0" pageNumber="151" rank="species" species="coactula">
<emphasis italics="true" pageId="0" pageNumber="151">P. coactula</emphasis>
</taxonomicName>
cluster based on nuclear DNA (Fig. 
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Maximum likelihood tree of Pristiphora carinata group based on nuclear genes (NaK and POL 2). Numbers at branches show SH-aLRT support (%) / ultrafast bootstrap support (%) values. Values of only well supported branches (&gt; 90 for both) and of P. coactula clade with moderate support are shown. Letters &quot; f &quot; stand for &quot; female &quot;, &quot; m &quot; for &quot; male &quot;, and &quot; l &quot; for larva. Numbers at the end of the tip labels refer to sequence length and the number of heterozygous positions." figureDoi="10.3897/dez.69.84080.figure30" httpUri="https://binary.pensoft.net/fig/719541" pageId="0" pageNumber="151">30</figureCitation>
). There are two main clusters based on COI sequences, one of which contains only 
<taxonomicName genus="P." lsidName="P. borea" pageId="0" pageNumber="151" rank="species" species="borea">
<emphasis italics="true" pageId="0" pageNumber="151">P. borea</emphasis>
</taxonomicName>
(Konow, 1904) and 
<taxonomicName genus="P." lsidName="P. groenblomi" pageId="0" pageNumber="151" rank="species" species="groenblomi">
<emphasis italics="true" pageId="0" pageNumber="151">P. groenblomi</emphasis>
</taxonomicName>
(Lindqvist, 1952) and the other one all species (Fig. 
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Maximum likelihood tree of Pristiphora carinata group based on mitochondrial COI gene. Numbers at branches show SH-aLRT support (%) / ultrafast bootstrap support (%) values. Values of only well supported branches (&gt; 90 for both) are shown. Letters &quot; f &quot; stand for &quot; female &quot;, &quot; m &quot; for &quot; male &quot;, and &quot; l &quot; for larva. Numbers at the end of the tip labels refer to sequence length and the number of ambiguous positions." figureDoi="10.3897/dez.69.84080.figure31" httpUri="https://binary.pensoft.net/fig/719542" pageId="0" pageNumber="151">31</figureCitation>
). Within the COI cluster containing all species (Fig. 
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Maximum likelihood tree of Pristiphora carinata group based on mitochondrial COI gene. Numbers at branches show SH-aLRT support (%) / ultrafast bootstrap support (%) values. Values of only well supported branches (&gt; 90 for both) are shown. Letters &quot; f &quot; stand for &quot; female &quot;, &quot; m &quot; for &quot; male &quot;, and &quot; l &quot; for larva. Numbers at the end of the tip labels refer to sequence length and the number of ambiguous positions." figureDoi="10.3897/dez.69.84080.figure31" httpUri="https://binary.pensoft.net/fig/719542" pageId="0" pageNumber="151">31</figureCitation>
), 
<taxonomicName genus="P." lsidName="P. borea" pageId="0" pageNumber="151" rank="species" species="borea">
<emphasis italics="true" pageId="0" pageNumber="151">P. borea</emphasis>
</taxonomicName>
, 
<taxonomicName genus="P." lsidName="P. groenblomi" pageId="0" pageNumber="151" rank="species" species="groenblomi">
<emphasis italics="true" pageId="0" pageNumber="151">P. groenblomi</emphasis>
</taxonomicName>
, and 
<taxonomicName genus="P." lsidName="P. albilabris" pageId="0" pageNumber="151" rank="species" species="albilabris">
<emphasis italics="true" pageId="0" pageNumber="151">P. albilabris</emphasis>
</taxonomicName>
(Boheman, 1852) (
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" lsidName="Betula" order="Fagales" pageId="0" pageNumber="151" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="151">Betula</emphasis>
</taxonomicName>
feeders) tend to separate from 
<taxonomicName genus="P." lsidName="P. coactula" pageId="0" pageNumber="151" rank="species" species="coactula">
<emphasis italics="true" pageId="0" pageNumber="151">P. coactula</emphasis>
</taxonomicName>
(
<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" lsidName="Salix" order="Malpighiales" pageId="0" pageNumber="151" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="151">Salix</emphasis>
</taxonomicName>
) and 
<taxonomicName genus="P." lsidName="P. carinata" pageId="0" pageNumber="151" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="151">P. carinata</emphasis>
</taxonomicName>
(Hartig, 1837) (
<taxonomicName class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" lsidName="Vaccinium" order="Ericales" pageId="0" pageNumber="151" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="151">Vaccinium</emphasis>
</taxonomicName>
). Based on the specimens having nuclear data, the species (mainly females) of the 
<taxonomicName authorityName="Hartig" authorityYear="1837" baseAuthorityName="Hartig" baseAuthorityYear="1837" class="Insecta" family="Tenthredinidae" genus="Pristiphora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristiphora carinata" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="151">Pristiphora carinata</emphasis>
</taxonomicName>
group may be separated by the following key, although it might not always work for all specimens, particularly 
<taxonomicName genus="P." lsidName="P. coactula" pageId="0" pageNumber="151" rank="species" species="coactula">
<emphasis italics="true" pageId="0" pageNumber="151">P. coactula</emphasis>
</taxonomicName>
and 
<taxonomicName genus="P." lsidName="P. borea" pageId="0" pageNumber="151" rank="species" species="borea">
<emphasis italics="true" pageId="0" pageNumber="151">P. borea</emphasis>
</taxonomicName>
. Excluded from the key are the (sub)arctic species 
<taxonomicName genus="P." lsidName="P. breadalbanensis" pageId="0" pageNumber="151" rank="species" species="breadalbanensis">
<emphasis italics="true" pageId="0" pageNumber="151">P. breadalbanensis</emphasis>
</taxonomicName>
and 
<emphasis italics="true" pageId="0" pageNumber="151">
<taxonomicName genus="P." lsidName="P. lativentris" pageId="0" pageNumber="151" rank="species" species="lativentris">P. lativentris</taxonomicName>
. 
<taxonomicName class="Insecta" family="Tenthredinidae" genus="Pristiphora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristiphora" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="genus">Pristiphora</taxonomicName>
<taxonomicName authorityName="Thomson" authorityYear="1871" baseAuthorityName="C.G.Thomson" baseAuthorityYear="1871" class="Insecta" family="Tenthredinidae" genus="Pristiphora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristiphora lativentris" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="lativentris">Pristiphora lativentris</taxonomicName>
</emphasis>
may have somewhat different serrulae from the other species (almost papilliform, see fig. 215 in 
<bibRefCitation DOI="https://doi.org/10.3897/jhr.59.12656.figure3" author="Prous, M" journalOrPublisher="Journal of Hymenoptera Research" pageId="0" pageNumber="151" pagination="1 - 190" refId="B150" refString="Prous, M, Kramp, K, Vikberg, V, Liston, A, 2017. North-Western Palaearctic species of Pristiphora (Hymenoptera, Tenthredinidae). Journal of Hymenoptera Research 59: 1 - 190, DOI: https://doi.org/10.3897/jhr.59.12656.figure3" title="North-Western Palaearctic species of Pristiphora (Hymenoptera, Tenthredinidae)." url="https://doi.org/10.3897/jhr.59.12656.figure3" volume="59" year="2017">Prous et al. 2017</bibRefCitation>
). The identity of 
<taxonomicName authorityName="Cameron" authorityYear="1882" baseAuthorityName="Cameron" baseAuthorityYear="1882" class="Insecta" family="Tenthredinidae" genus="Pristiphora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristiphora breadalbanensis" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="breadalbanensis">
<emphasis italics="true" pageId="0" pageNumber="151">Pristiphora breadalbanensis</emphasis>
</taxonomicName>
(most similar to 
<taxonomicName genus="P." lsidName="P. borea" pageId="0" pageNumber="151" rank="species" species="borea">
<emphasis italics="true" pageId="0" pageNumber="151">P. borea</emphasis>
</taxonomicName>
and 
<taxonomicName genus="P." lsidName="P. coactula" pageId="0" pageNumber="151" rank="species" species="coactula">
<emphasis italics="true" pageId="0" pageNumber="151">P. coactula</emphasis>
</taxonomicName>
) needs further research to confirm if characters (e.g. structure of median mesoscutal lobes) mentioned by 
<bibRefCitation author="Benson, RB" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="151" refId="B11" refString="Benson, RB, 1958. Hymenoptera, Symphyta. Handbooks for the Identification of British Insects 6(2c): 139-258." title="Hymenoptera, Symphyta. Handbooks for the Identification of British Insects 6 (2 c): 139 - 258." year="1958">Benson (1958)</bibRefCitation>
to separate this species are reliable.
</paragraph>
<caption doi="10.3897/dez.69.84080.figure30" httpUri="https://binary.pensoft.net/fig/719541" pageId="0" pageNumber="151" start="Figure 30" startId="F30">
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">Figure 30.</emphasis>
Maximum likelihood tree of 
<taxonomicName authorityName="Hartig" authorityYear="1837" baseAuthorityName="Hartig" baseAuthorityYear="1837" class="Insecta" family="Tenthredinidae" genus="Pristiphora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristiphora carinata" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="151">Pristiphora carinata</emphasis>
</taxonomicName>
group based on nuclear genes (NaK and POL2). Numbers at branches show SH-aLRT support (%) / ultrafast bootstrap support (%) values. Values of only well supported branches (&gt;90 for both) and of 
<taxonomicName genus="P." lsidName="P. coactula" pageId="0" pageNumber="151" rank="species" species="coactula">
<emphasis italics="true" pageId="0" pageNumber="151">P. coactula</emphasis>
</taxonomicName>
clade with moderate support are shown. Letters &quot;f &quot; stand for 
<normalizedToken originalValue="“female”">&quot;female&quot;</normalizedToken>
, 
<normalizedToken originalValue="“m”">&quot;m&quot;</normalizedToken>
for 
<normalizedToken originalValue="“male”">&quot;male&quot;</normalizedToken>
, and 
<normalizedToken originalValue="“l”">&quot;l&quot;</normalizedToken>
for larva. Numbers at the end of the tip labels refer to sequence length and the number of heterozygous positions.
</paragraph>
</caption>
<caption doi="10.3897/dez.69.84080.figure31" httpUri="https://binary.pensoft.net/fig/719542" pageId="0" pageNumber="151" start="Figure 31" startId="F31">
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">Figure 31.</emphasis>
Maximum likelihood tree of 
<taxonomicName authorityName="Hartig" authorityYear="1837" baseAuthorityName="Hartig" baseAuthorityYear="1837" class="Insecta" family="Tenthredinidae" genus="Pristiphora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristiphora carinata" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="151">Pristiphora carinata</emphasis>
</taxonomicName>
group based on mitochondrial COI gene. Numbers at branches show SH-aLRT support (%) / ultrafast bootstrap support (%) values. Values of only well supported branches (&gt;90 for both) are shown. Letters &quot;f &quot; stand for 
<normalizedToken originalValue="“female”">&quot;female&quot;</normalizedToken>
, 
<normalizedToken originalValue="“m”">&quot;m&quot;</normalizedToken>
for 
<normalizedToken originalValue="“male”">&quot;male&quot;</normalizedToken>
, and 
<normalizedToken originalValue="“l”">&quot;l&quot;</normalizedToken>
for larva. Numbers at the end of the tip labels refer to sequence length and the number of ambiguous positions.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="151">
<table inLine="true" pageId="0" pageNumber="151">
<tr pageId="0" pageNumber="151">
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="151">a</emphasis>
Pterostigma distinctly darker than costa 
<emphasis bold="true" pageId="0" pageNumber="151">b</emphasis>
Legs largely orange or reddish 
<emphasis bold="true" pageId="0" pageNumber="151">c</emphasis>
In female, valvifer 2 and terga 9-10 black or slightly pale 
<emphasis bold="true" pageId="0" pageNumber="151">d</emphasis>
In male, sternum 9 blac
</td>
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">
<taxonomicName genus="P." lsidName="P. albilabris" pageId="0" pageNumber="151" rank="species" species="albilabris">
<emphasis italics="true" pageId="0" pageNumber="151">P. albilabris</emphasis>
</taxonomicName>
(Boheman, 1852) ♂♀ and 
<taxonomicName genus="P." lsidName="P. groenblomi" pageId="0" pageNumber="151" rank="species" species="groenblomi">
<emphasis italics="true" pageId="0" pageNumber="151">P. groenblomi</emphasis>
</taxonomicName>
(Lindqvist, 1952) ♂♀ in part
</td>
</tr>
<tr pageId="0" pageNumber="151">
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="151">aa</emphasis>
Pterostigma similarly pale as costa or somewhat darker than costa 
<emphasis bold="true" pageId="0" pageNumber="151">bb</emphasis>
Legs largely black to pale 
<emphasis bold="true" pageId="0" pageNumber="151">cc</emphasis>
In female, valvifer 2 and terga 9-10 extensively pale 
<emphasis bold="true" pageId="0" pageNumber="151">dd</emphasis>
In male, sternum 9 black to pale
</td>
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">2</td>
</tr>
<tr pageId="0" pageNumber="151">
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">2(1)</td>
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="151">a</emphasis>
Pterostigma somewhat darker than costa 
<emphasis bold="true" pageId="0" pageNumber="151">b</emphasis>
Legs largely orange or reddish 
<emphasis bold="true" pageId="0" pageNumber="151">c</emphasis>
Metafemur completely pale 
<emphasis bold="true" pageId="0" pageNumber="151">d</emphasis>
In female, terga 2-8 and sterna black 
<emphasis bold="true" pageId="0" pageNumber="151">e</emphasis>
In male, sternum 9 (always?) black
</td>
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">
<taxonomicName genus="P." lsidName="P. groenblomi" pageId="0" pageNumber="151" rank="species" species="groenblomi">
<emphasis italics="true" pageId="0" pageNumber="151">P. groenblomi</emphasis>
</taxonomicName>
♂♀ in part
</td>
</tr>
<tr pageId="0" pageNumber="151">
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="151">aa</emphasis>
Pterostigma similarly pale as costa 
<emphasis bold="true" pageId="0" pageNumber="151">bb</emphasis>
Legs largely black to yellowish 
<emphasis bold="true" pageId="0" pageNumber="151">cc</emphasis>
Metafemur black to pale 
<emphasis bold="true" pageId="0" pageNumber="151">dd</emphasis>
In female, terga 2-8 and sterna black or partly pale (starting from tergum 2 and sternum 2) 
<emphasis bold="true" pageId="0" pageNumber="151">ee</emphasis>
In male, sternum 9 black to pale ... ♀ (males of the following species not separated)
</td>
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">3</td>
</tr>
<tr pageId="0" pageNumber="151">
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">3(2)</td>
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="151">a</emphasis>
Valvula 3 in dorsal view gradually narrowing, without invagination and with sharp tip (see figs 98-99 in 
<bibRefCitation DOI="https://doi.org/10.3897/jhr.59.12656.figure3" author="Prous, M" journalOrPublisher="Journal of Hymenoptera Research" pageId="0" pageNumber="151" pagination="1 - 190" refId="B150" refString="Prous, M, Kramp, K, Vikberg, V, Liston, A, 2017. North-Western Palaearctic species of Pristiphora (Hymenoptera, Tenthredinidae). Journal of Hymenoptera Research 59: 1 - 190, DOI: https://doi.org/10.3897/jhr.59.12656.figure3" title="North-Western Palaearctic species of Pristiphora (Hymenoptera, Tenthredinidae)." url="https://doi.org/10.3897/jhr.59.12656.figure3" volume="59" year="2017">Prous et al. 2017</bibRefCitation>
) 
<emphasis bold="true" pageId="0" pageNumber="151">b</emphasis>
Usually only terga 8-10 or 9-10 extensively pale, but sometimes more (starting from tergum 5)
</td>
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">
<taxonomicName genus="P." lsidName="P. carinata" pageId="0" pageNumber="151" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="151">P. carinata</emphasis>
</taxonomicName>
(Hartig, 1837)
</td>
</tr>
<tr pageId="0" pageNumber="151">
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="151">aa</emphasis>
Valvula 3 in dorsal view more or less truncate, with or without indistinct invagination and with broader tip 
<emphasis bold="true" pageId="0" pageNumber="151">bb</emphasis>
Usually terga 8-10 or more (starting from tergum 2) at least partly pale
</td>
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">4</td>
</tr>
<tr pageId="0" pageNumber="151">
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">4(3)</td>
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="151">a</emphasis>
Valvula 3 short, truncate and usually with indistinct invagination (Fig. 
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Tip of abdomen of Pristiphora carinata group females in dorsal view. A. P. borea (ZMUO. 035517); B. P. borea (ZMUO. 033457); C. P. coactula (ZMUO. 046522); D. P. coactula (ZMUO. 035246)." figureDoi="10.3897/dez.69.84080.figure33" httpUri="https://binary.pensoft.net/fig/719544" pageId="0" pageNumber="151">33C, D</figureCitation>
) 
<emphasis bold="true" pageId="0" pageNumber="151">b</emphasis>
Abdomen 
<emphasis italics="true" pageId="0" pageNumber="151">usually</emphasis>
becoming gradually paler from base to apex, dorsally 
<emphasis italics="true" pageId="0" pageNumber="151">usually</emphasis>
starting from tergum 7, laterally and ventrally from tergum 2 and sternum 2 
<emphasis bold="true" pageId="0" pageNumber="151">c</emphasis>
Metafemur 
<emphasis italics="true" pageId="0" pageNumber="151">usually</emphasis>
completely pale 
<emphasis bold="true" pageId="0" pageNumber="151">d</emphasis>
Clypeus 
<emphasis italics="true" pageId="0" pageNumber="151">usually</emphasis>
mostly pale
</td>
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">
<taxonomicName genus="P." lsidName="P. coactula" pageId="0" pageNumber="151" rank="species" species="coactula">
<emphasis italics="true" pageId="0" pageNumber="151">P. coactula</emphasis>
</taxonomicName>
(Ruthe, 1859)
</td>
</tr>
<tr pageId="0" pageNumber="151">
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="151">aa</emphasis>
Valvula 3 
<emphasis italics="true" pageId="0" pageNumber="151">usually</emphasis>
longer, slightly narrowed at apex and without invagination (Fig. 
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Tip of abdomen of Pristiphora carinata group females in dorsal view. A. P. borea (ZMUO. 035517); B. P. borea (ZMUO. 033457); C. P. coactula (ZMUO. 046522); D. P. coactula (ZMUO. 035246)." figureDoi="10.3897/dez.69.84080.figure33" httpUri="https://binary.pensoft.net/fig/719544" pageId="0" pageNumber="151">33A</figureCitation>
), but sometimes not distinguishable from 
<taxonomicName genus="P." lsidName="P. coactula" pageId="0" pageNumber="151" rank="species" species="coactula">
<emphasis italics="true" pageId="0" pageNumber="151">P. coactula</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Tip of abdomen of Pristiphora carinata group females in dorsal view. A. P. borea (ZMUO. 035517); B. P. borea (ZMUO. 033457); C. P. coactula (ZMUO. 046522); D. P. coactula (ZMUO. 035246)." figureDoi="10.3897/dez.69.84080.figure33" httpUri="https://binary.pensoft.net/fig/719544" pageId="0" pageNumber="151">33B</figureCitation>
) 
<emphasis bold="true" pageId="0" pageNumber="151">bb</emphasis>
Abdomen 
<emphasis italics="true" pageId="0" pageNumber="151">usually</emphasis>
slightly or extensively pale only at apex, dorsally 
<emphasis italics="true" pageId="0" pageNumber="151">usually</emphasis>
terga 8-10, laterally 
<emphasis italics="true" pageId="0" pageNumber="151">usually</emphasis>
terga 7-10, ventrally 
<emphasis italics="true" pageId="0" pageNumber="151">usually</emphasis>
sternum 7 
<emphasis bold="true" pageId="0" pageNumber="151">cc</emphasis>
Metafemur black to completely pale 
<emphasis bold="true" pageId="0" pageNumber="151">dd</emphasis>
Clypeus mostly black to mostly pale
</td>
<td colspan="1" pageId="0" pageNumber="151" rowspan="1">
<taxonomicName genus="P." lsidName="P. borea" pageId="0" pageNumber="151" rank="species" species="borea">
<emphasis italics="true" pageId="0" pageNumber="151">P. borea</emphasis>
</taxonomicName>
(Konow, 1904)
</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="151">
Examples of lancets of 
<taxonomicName genus="P." lsidName="P. borea" pageId="0" pageNumber="151" rank="species" species="borea">
<emphasis italics="true" pageId="0" pageNumber="151">P. borea</emphasis>
</taxonomicName>
, 
<taxonomicName genus="P." lsidName="P. carinata" pageId="0" pageNumber="151" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="151">P. carinata</emphasis>
</taxonomicName>
, and 
<taxonomicName genus="P." lsidName="P. coactula" pageId="0" pageNumber="151" rank="species" species="coactula">
<emphasis italics="true" pageId="0" pageNumber="151">P. coactula</emphasis>
</taxonomicName>
are shown in Fig. 
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Lancets of Pristiphora carinata group. A. P. coactula (ZMUO. 031490); B. P. borea (ZMUO. 033284); C. P. carinata (ZMUO. 031554)." figureDoi="10.3897/dez.69.84080.figure32" httpUri="https://binary.pensoft.net/fig/719543" pageId="0" pageNumber="151">32A-C</figureCitation>
, but more specimens need to be examined to check if there are any consistent differences between the species. Morphological differences between the males of 
<taxonomicName genus="P." lsidName="P. borea" pageId="0" pageNumber="151" rank="species" species="borea">
<emphasis italics="true" pageId="0" pageNumber="151">P. borea</emphasis>
</taxonomicName>
, 
<taxonomicName genus="P." lsidName="P. carinata" pageId="0" pageNumber="151" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="151">P. carinata</emphasis>
</taxonomicName>
, and 
<taxonomicName genus="P." lsidName="P. coactula" pageId="0" pageNumber="151" rank="species" species="coactula">
<emphasis italics="true" pageId="0" pageNumber="151">P. coactula</emphasis>
</taxonomicName>
are not clear. Externally, it seems that 
<taxonomicName genus="P." lsidName="P. coactula" pageId="0" pageNumber="151" rank="species" species="coactula">
<emphasis italics="true" pageId="0" pageNumber="151">P. coactula</emphasis>
</taxonomicName>
tends to be paler (clypeus, pronotal angles, tegula, metafemur, and sternum 9 completely or mostly pale) than 
<taxonomicName genus="P." lsidName="P. borea" pageId="0" pageNumber="151" rank="species" species="borea">
<emphasis italics="true" pageId="0" pageNumber="151">P. borea</emphasis>
</taxonomicName>
and 
<taxonomicName genus="P." lsidName="P. carinata" pageId="0" pageNumber="151" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="151">P. carinata</emphasis>
</taxonomicName>
. The dorso-apical margin of the paravalva of 
<taxonomicName genus="P." lsidName="P. borea" pageId="0" pageNumber="151" rank="species" species="borea">
<emphasis italics="true" pageId="0" pageNumber="151">P. borea</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Penis valves of Pristiphora carinata group. A. P. carinata (ZMUO. 031419); B. P. borea (DEI-GISHym 80148); C. P. groenblomi (DEI-GISHym 80210); D. P. coactula (ZMUO. 039225); E. P. coactula (DEI-GISHym 84186); F. P. albilabris (ZMUO. 032465)." figureDoi="10.3897/dez.69.84080.figure34" httpUri="https://binary.pensoft.net/fig/719545" pageId="0" pageNumber="151">34B</figureCitation>
) may be more strongly inclined basally compared to 
<taxonomicName genus="P." lsidName="P. carinata" pageId="0" pageNumber="151" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="151">P. carinata</emphasis>
</taxonomicName>
and 
<taxonomicName genus="P." lsidName="P. coactula" pageId="0" pageNumber="151" rank="species" species="coactula">
<emphasis italics="true" pageId="0" pageNumber="151">P. coactula</emphasis>
</taxonomicName>
, but differences between the penis valves of the latter two species are not clear (Fig. 
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Penis valves of Pristiphora carinata group. A. P. carinata (ZMUO. 031419); B. P. borea (DEI-GISHym 80148); C. P. groenblomi (DEI-GISHym 80210); D. P. coactula (ZMUO. 039225); E. P. coactula (DEI-GISHym 84186); F. P. albilabris (ZMUO. 032465)." figureDoi="10.3897/dez.69.84080.figure34" httpUri="https://binary.pensoft.net/fig/719545" pageId="0" pageNumber="151">34A, D, E</figureCitation>
). The most distinctive penis valve in the 
<taxonomicName genus="P." lsidName="P. carinata" pageId="0" pageNumber="151" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="151">P. carinata</emphasis>
</taxonomicName>
group seems to belong to 
<taxonomicName genus="P." lsidName="P. albilabris" pageId="0" pageNumber="151" rank="species" species="albilabris">
<emphasis italics="true" pageId="0" pageNumber="151">P. albilabris</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Penis valves of Pristiphora carinata group. A. P. carinata (ZMUO. 031419); B. P. borea (DEI-GISHym 80148); C. P. groenblomi (DEI-GISHym 80210); D. P. coactula (ZMUO. 039225); E. P. coactula (DEI-GISHym 84186); F. P. albilabris (ZMUO. 032465)." figureDoi="10.3897/dez.69.84080.figure34" httpUri="https://binary.pensoft.net/fig/719545" pageId="0" pageNumber="151">34F</figureCitation>
), which has the most distinctly inclined dorso-apical margin of paravalva. Overall shape of penis valve of 
<taxonomicName genus="P." lsidName="P. groenblomi" pageId="0" pageNumber="151" rank="species" species="groenblomi">
<emphasis italics="true" pageId="0" pageNumber="151">P. groenblomi</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Penis valves of Pristiphora carinata group. A. P. carinata (ZMUO. 031419); B. P. borea (DEI-GISHym 80148); C. P. groenblomi (DEI-GISHym 80210); D. P. coactula (ZMUO. 039225); E. P. coactula (DEI-GISHym 84186); F. P. albilabris (ZMUO. 032465)." figureDoi="10.3897/dez.69.84080.figure34" httpUri="https://binary.pensoft.net/fig/719545" pageId="0" pageNumber="151">34C</figureCitation>
) is most similar to 
<taxonomicName genus="P." lsidName="P. borea" pageId="0" pageNumber="151" rank="species" species="borea">
<emphasis italics="true" pageId="0" pageNumber="151">P. borea</emphasis>
</taxonomicName>
, but it may be larger.
</paragraph>
<caption doi="10.3897/dez.69.84080.figure32" httpUri="https://binary.pensoft.net/fig/719543" pageId="0" pageNumber="151" start="Figure 32" startId="F32">
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">Figure 32.</emphasis>
Lancets of 
<taxonomicName authorityName="Hartig" authorityYear="1837" baseAuthorityName="Hartig" baseAuthorityYear="1837" class="Insecta" family="Tenthredinidae" genus="Pristiphora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristiphora carinata" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="151">Pristiphora carinata</emphasis>
</taxonomicName>
group. 
<emphasis bold="true" pageId="0" pageNumber="151">A.</emphasis>
<taxonomicName genus="P." lsidName="P. coactula" pageId="0" pageNumber="151" rank="species" species="coactula">
<emphasis italics="true" pageId="0" pageNumber="151">P. coactula</emphasis>
</taxonomicName>
(ZMUO.031490); 
<emphasis bold="true" pageId="0" pageNumber="151">B.</emphasis>
<taxonomicName genus="P." lsidName="P. borea" pageId="0" pageNumber="151" rank="species" species="borea">
<emphasis italics="true" pageId="0" pageNumber="151">P. borea</emphasis>
</taxonomicName>
(ZMUO.033284); 
<emphasis bold="true" pageId="0" pageNumber="151">C.</emphasis>
<taxonomicName genus="P." lsidName="P. carinata" pageId="0" pageNumber="151" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="151">P. carinata</emphasis>
</taxonomicName>
(ZMUO.031554).
</paragraph>
</caption>
<caption doi="10.3897/dez.69.84080.figure33" httpUri="https://binary.pensoft.net/fig/719544" pageId="0" pageNumber="151" start="Figure 33" startId="F33">
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">Figure 33.</emphasis>
Tip of abdomen of 
<taxonomicName authorityName="Hartig" authorityYear="1837" baseAuthorityName="Hartig" baseAuthorityYear="1837" class="Insecta" family="Tenthredinidae" genus="Pristiphora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristiphora carinata" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="151">Pristiphora carinata</emphasis>
</taxonomicName>
group females in dorsal view. 
<emphasis bold="true" pageId="0" pageNumber="151">A.</emphasis>
<taxonomicName genus="P." lsidName="P. borea" pageId="0" pageNumber="151" rank="species" species="borea">
<emphasis italics="true" pageId="0" pageNumber="151">P. borea</emphasis>
</taxonomicName>
(ZMUO.035517); 
<emphasis bold="true" pageId="0" pageNumber="151">B.</emphasis>
<taxonomicName genus="P." lsidName="P. borea" pageId="0" pageNumber="151" rank="species" species="borea">
<emphasis italics="true" pageId="0" pageNumber="151">P. borea</emphasis>
</taxonomicName>
(ZMUO.033457); 
<emphasis bold="true" pageId="0" pageNumber="151">C.</emphasis>
<taxonomicName genus="P." lsidName="P. coactula" pageId="0" pageNumber="151" rank="species" species="coactula">
<emphasis italics="true" pageId="0" pageNumber="151">P. coactula</emphasis>
</taxonomicName>
(ZMUO.046522); 
<emphasis bold="true" pageId="0" pageNumber="151">D.</emphasis>
<taxonomicName genus="P." lsidName="P. coactula" pageId="0" pageNumber="151" rank="species" species="coactula">
<emphasis italics="true" pageId="0" pageNumber="151">P. coactula</emphasis>
</taxonomicName>
(ZMUO.035246).
</paragraph>
</caption>
<caption doi="10.3897/dez.69.84080.figure34" httpUri="https://binary.pensoft.net/fig/719545" pageId="0" pageNumber="151" start="Figure 34" startId="F34">
<paragraph pageId="0" pageNumber="151">
<emphasis bold="true" pageId="0" pageNumber="151">Figure 34.</emphasis>
Penis valves of 
<taxonomicName authorityName="Hartig" authorityYear="1837" baseAuthorityName="Hartig" baseAuthorityYear="1837" class="Insecta" family="Tenthredinidae" genus="Pristiphora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristiphora carinata" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="151">Pristiphora carinata</emphasis>
</taxonomicName>
group. 
<emphasis bold="true" pageId="0" pageNumber="151">A.</emphasis>
<taxonomicName genus="P." lsidName="P. carinata" pageId="0" pageNumber="151" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="151">P. carinata</emphasis>
</taxonomicName>
(ZMUO.031419); 
<emphasis bold="true" pageId="0" pageNumber="151">B.</emphasis>
<taxonomicName genus="P." lsidName="P. borea" pageId="0" pageNumber="151" rank="species" species="borea">
<emphasis italics="true" pageId="0" pageNumber="151">P. borea</emphasis>
</taxonomicName>
(DEI-GISHym80148); 
<emphasis bold="true" pageId="0" pageNumber="151">C.</emphasis>
<taxonomicName genus="P." lsidName="P. groenblomi" pageId="0" pageNumber="151" rank="species" species="groenblomi">
<emphasis italics="true" pageId="0" pageNumber="151">P. groenblomi</emphasis>
</taxonomicName>
(DEI-GISHym80210); 
<emphasis bold="true" pageId="0" pageNumber="151">D.</emphasis>
<taxonomicName genus="P." lsidName="P. coactula" pageId="0" pageNumber="151" rank="species" species="coactula">
<emphasis italics="true" pageId="0" pageNumber="151">P. coactula</emphasis>
</taxonomicName>
(ZMUO.039225); 
<emphasis bold="true" pageId="0" pageNumber="151">E.</emphasis>
<taxonomicName genus="P." lsidName="P. coactula" pageId="0" pageNumber="151" rank="species" species="coactula">
<emphasis italics="true" pageId="0" pageNumber="151">P. coactula</emphasis>
</taxonomicName>
(DEI-GISHym84186); 
<emphasis bold="true" pageId="0" pageNumber="151">F.</emphasis>
<taxonomicName genus="P." lsidName="P. albilabris" pageId="0" pageNumber="151" rank="species" species="albilabris">
<emphasis italics="true" pageId="0" pageNumber="151">P. albilabris</emphasis>
</taxonomicName>
(ZMUO.032465).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>