172 lines
19 KiB
XML
172 lines
19 KiB
XML
<document id="A2C35A4BCE41E64E64CF81C7016C9A09" ID-CLB-Dataset="21156" ID-DOI="10.11646/zootaxa.4661.2.3" ID-GBIF-Dataset="6c12aca1-7350-4265-9ded-6d415152f181" ID-ISSN="1175-5326" ID-Zenodo-Dep="3998528" ID-ZooBank="95722A8A-E02C-47E5-AA9B-777B6CA226E6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1598306200512" checkinUser="felipe" docAuthor="Fukuda, Yuki, Ogawa, Ryo & Hori, Michio" docDate="2019" docId="1D1C8783DF778E0C698632668BB5FEB2" docLanguage="en" docName="zootaxa.4661.2.3.pdf.imf" docOrigin="Zootaxa 4661 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calochroa hamiltoniana" docType="treatment" docVersion="6" lastPageNumber="291" masterDocId="E125FFFBDF648E18691134438958FFF1" masterDocTitle="The reclassification of Sophiodela and other tiger beetles (Coleoptera, Cicindelidae) based on the structure of the everted internal sac of the male genitalia" masterLastPageNumber="308" masterPageNumber="271" pageNumber="290" updateTime="1698845354460" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3130DC25CA51CA3DBDDB37E6BF6B9489">The reclassification of Sophiodela and other tiger beetles (Coleoptera, Cicindelidae) based on the structure of the everted internal sac of the male genitalia</mods:title>
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<mods:namePart id="FBBDDA61A4739CB64D0C0073AAA87AC7">Fukuda, Yuki</mods:namePart>
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<mods:affiliation id="2FFA7F75E9778239077B5C42221CFFAD">Kurayoshi, 111 - 8, Kamogawa-town, Tottori, 682 - 0923 Japan E-mail: yuki 03 go 3 @ gmail. com</mods:affiliation>
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<mods:nameIdentifier id="216E796E0EBEE5FDAA376F568DEC00E7" type="email">yuki03go3@gmail.com</mods:nameIdentifier>
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<mods:namePart id="841EC52528F12DA689197BBB638127DA">Ogawa, Ryo</mods:namePart>
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<mods:affiliation id="F1CEC99559C2B60615A43763A2647764">Matsuyama, 6 - 5 - 8, Hakusuidai, Ehime, 791 - 0113 Japan E-mail: ailmasbl 0854 @ yahoo. co. jp</mods:affiliation>
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<mods:namePart id="9D3CC846D658D4500276F41266F68E97">Hori, Michio</mods:namePart>
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<mods:affiliation id="3D7E0D4C83A3ABE2B27831498B8D8166">Wakayama, 6 - 1 - 2 Takajyomachi, Wakayama, 640 - 8135 Japan E-mail: hori @ terra. zool. kyoto-u. ac. jp</mods:affiliation>
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<mods:date id="74B5EC607789588B3BAAE71FBB1C8F3F">2019</mods:date>
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<mods:number id="B69C3506DCCD855CD6C2DD84FF92E01D">4661</mods:number>
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<treatment id="1D1C8783DF778E0C698632668BB5FEB2" ID-DOI="http://doi.org/10.5281/zenodo.4337415" ID-GBIF-Taxon="167293593" ID-Zenodo-Dep="4337415" LSID="urn:lsid:plazi:treatment:1D1C8783DF778E0C698632668BB5FEB2" httpUri="http://treatment.plazi.org/id/1D1C8783DF778E0C698632668BB5FEB2" lastPageId="20" lastPageNumber="291" pageId="19" pageNumber="290">
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<subSubSection id="DDAF651EDF778E0B698632668BC9F9CE" box="[151,657,1573,1599]" pageId="19" pageNumber="290" type="nomenclature">
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<paragraph id="950A3695DF778E0B698632668BC9F9CE" blockId="19.[151,657,1573,1635]" box="[151,657,1573,1599]" pageId="19" pageNumber="290">
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<heading id="CE4281F9DF778E0B698632668BC9F9CE" box="[151,657,1573,1599]" level="2" pageId="19" pageNumber="290">
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<taxonomicName id="52B54D16DF778E0B698632668BC9F9CE" ID-CoL="PXDS" authority="(Thomson, 1857)" baseAuthorityName="Thomson" baseAuthorityYear="1857" box="[151,657,1573,1599]" class="Insecta" family="Carabidae" genus="Calochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="290" phylum="Arthropoda" rank="species" species="hamiltoniana">
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<emphasis id="A7C1EA87DF778E0B6986326688EFF9CE" box="[151,439,1573,1599]" italics="true" pageId="19" pageNumber="290">Calochroa hamiltoniana</emphasis>
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<emphasis id="A7C1EA87DF778E0B68AE32668BC9F9CE" bold="true" box="[447,657,1573,1599]" pageId="19" pageNumber="290">
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(
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<bibRefCitation id="F1244B64DF778E0B68D632668BD0F9CE" author="Thomson, J." box="[455,648,1573,1599]" pageId="19" pageNumber="290" pagination="321 - 325" refId="ref20642" refString="Thomson, J. (1857) Voyage entomologique a Londres. Archives Entomologiques, 1, 321 - 325." type="journal article" year="1857">Thomson, 1857</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="DDAF651EDF778E0B6986320A8869F992" box="[151,305,1608,1635]" pageId="19" pageNumber="290" type="description">
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<paragraph id="950A3695DF778E0B6986320A8869F992" blockId="19.[151,657,1573,1635]" box="[151,305,1608,1635]" pageId="19" pageNumber="290">
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(
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<figureCitation id="0D8E2A10DF778E0B698E320A89A3F992" box="[159,251,1608,1635]" captionStart="FIGURE 5" captionStartId="9.[151,250,1803,1828]" captionTargetBox="[159,1428,178,1767]" captionTargetId="figure@9.[151,1435,168,1778]" captionTargetPageId="9" captionText="FIGURE 5. Habitus of Calochroa and Cicindela (Pancallia). A, Calochroa schillhammeri Wiesner; B, Calochroa fabriciana (Horn); C, Calochroa hamiltoniana (Thomson); D, Cicindela (Pancallia) angulicollis Horn; E–F, C. (P.) cyanea Fabricius stat. nov. (E, female; F, male); G, C. (P.) princeps Vigors; H, C. (P.) shivah Parry." figureDoi="http://doi.org/10.5281/zenodo.3998538" httpUri="https://zenodo.org/record/3998538/files/figure.png" pageId="19" pageNumber="290">Figs 5C</figureCitation>
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,
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<figureCitation id="0D8E2A10DF778E0B6817320A8870F992" box="[262,296,1609,1635]" captionStart="FIGURE 9" captionStartId="22.[151,250,1945,1970]" captionTargetBox="[151,1435,181,1920]" captionTargetId="figure@22.[151,1435,181,1920]" captionTargetPageId="22" captionText="FIGURE 9. Everted internal sac of Calochroa. A, Calochroa assamensis (Parry); B, C. hamiltoniana (Thomson); C, C. bicolor bicolor (Fabricius); D, C. fabriciana (Horn); E, C. flavomaculata (Hope). Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3998546" httpUri="https://zenodo.org/record/3998546/files/figure.png" pageId="19" pageNumber="290">9B</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="DDAF651EDF778E0B698632CE8C11F8B5" pageId="19" pageNumber="290" type="reference_group">
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<paragraph id="950A3695DF778E0B698632CE8B38F957" blockId="19.[151,1436,1677,2041]" box="[151,608,1677,1702]" pageId="19" pageNumber="290">
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<taxonomicName id="52B54D16DF778E0B698632CE8B04F957" ID-CoL="V86P" authority="Thomson, 1857: 323" authorityName="Thomson" authorityPageNumber="323" authorityYear="1857" box="[151,604,1677,1702]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="290" phylum="Arthropoda" rank="species" species="hamiltoniana">
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<emphasis id="A7C1EA87DF778E0B698632CE88DFF954" box="[151,391,1677,1701]" italics="true" pageId="19" pageNumber="290">Cicindela hamiltoniana</emphasis>
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<bibRefCitation id="F1244B64DF778E0B689C32CE8B04F957" author="Thomson, J." box="[397,604,1677,1702]" pageId="19" pageNumber="290" pagination="321 - 325" refId="ref20642" refString="Thomson, J. (1857) Voyage entomologique a Londres. Archives Entomologiques, 1, 321 - 325." type="journal article" year="1857">Thomson, 1857:323</bibRefCitation>
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</taxonomicName>
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||
.
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</paragraph>
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<paragraph id="950A3695DF778E0B698632EF8D7AF935" blockId="19.[151,1436,1677,2041]" box="[151,1058,1707,1732]" pageId="19" pageNumber="290">
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<taxonomicName id="52B54D16DF778E0B698632EF8B6FF935" ID-CoL="V98L" authority="Chaudoir, 1865: 61" authorityName="Chaudoir" authorityPageNumber="61" authorityYear="1865" box="[151,567,1707,1732]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="290" phylum="Arthropoda" rank="species" species="flavovittata">
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<emphasis id="A7C1EA87DF778E0B698632EF882BF935" box="[151,371,1708,1732]" italics="true" pageId="19" pageNumber="290">Cicindela flavovittata</emphasis>
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<bibRefCitation id="F1244B64DF778E0B686B32E88B6FF935" author="Chaudoir, M. de" box="[378,567,1707,1732]" pageId="19" pageNumber="290" refId="ref17676" refString="Chaudoir, M. de (1865) Catalogue de la collection de Cicindeletes de M. le Baron de Chaudoir. Nys, Bruxelles, 64 pp. https: // doi. org / 10.5962 / bhl. title. 134840" type="book" year="1865">Chaudoir, 1865:61</bibRefCitation>
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</taxonomicName>
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[synonymy by
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||
<bibRefCitation id="F1244B64DF778E0B6BC532E88D40F935" author="Acciavatti, R. E. & Pearson, D. L." box="[724,1048,1707,1732]" pageId="19" pageNumber="290" pagination="77 - 353" refId="ref17273" refString="Acciavatti, R. E. & Pearson, D. L. (1989) The tiger beetle Genus Cicindela (Coleoptera, Insecta) from the indian subcontinent. Annals of Carnegie Museum, 58, 77 - 353." type="journal article" year="1989">Acciavatti & Pearson 1989:127</bibRefCitation>
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||
].
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||
</paragraph>
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||
<paragraph id="950A3695DF778E0B698632888B04F912" blockId="19.[151,1436,1677,2041]" box="[151,604,1738,1763]" pageId="19" pageNumber="290">
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||
<taxonomicName id="52B54D16DF778E0B698632888B00F912" ID-CoL="PXDS" authority=": Rivalier 1961: 127" authorityName="Rivalier" authorityPageNumber="127" authorityYear="1961" box="[151,600,1738,1763]" class="Insecta" family="Carabidae" genus="Calochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="290" phylum="Arthropoda" rank="species" species="hamiltoniana">
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<emphasis id="A7C1EA87DF778E0B6986328888C8F912" box="[151,400,1739,1763]" italics="true" pageId="19" pageNumber="290">Calochroa hamiltoniana</emphasis>
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||
:
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||
<bibRefCitation id="F1244B64DF778E0B688C32898B00F912" author="Rivalier, E." box="[413,600,1738,1763]" pageId="19" pageNumber="290" pagination="121 - 149" refId="ref20230" refString="Rivalier, E. (1961) Demembrement du genre Cicindela Linne, IV. Faune indomalaise. Revue francaise d'Entomologie, 28, 121 - 149." type="journal article" year="1961">Rivalier 1961:127</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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||
<paragraph id="950A3695DF778E0B698632AA8A04F8F0" blockId="19.[151,1436,1677,2041]" box="[151,860,1769,1794]" pageId="19" pageNumber="290">
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<taxonomicName id="52B54D16DF778E0B698632AA8A00F8F0" ID-CoL="V86P" authority=": Acciavatti & Pearson 1989: 122" authorityName="Acciavatti & Pearson" authorityPageNumber="122" authorityYear="1989" box="[151,856,1769,1794]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="290" phylum="Arthropoda" rank="species" species="hamiltoniana" subGenus="Calochroa">
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<emphasis id="A7C1EA87DF778E0B698632AA89A2F8F0" box="[151,250,1769,1793]" italics="true" pageId="19" pageNumber="290">Cicindela</emphasis>
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||
(
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<emphasis id="A7C1EA87DF778E0B681932AA882BF8F0" box="[264,371,1769,1793]" italics="true" pageId="19" pageNumber="290">Calochroa</emphasis>
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||
)
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<emphasis id="A7C1EA87DF778E0B689232AA8B52F8F0" box="[387,522,1769,1793]" italics="true" pageId="19" pageNumber="290">hamiltoniana</emphasis>
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||
:
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||
<bibRefCitation id="F1244B64DF778E0B6B0732AA8A00F8F0" author="Acciavatti, R. E. & Pearson, D. L." box="[534,856,1769,1794]" pageId="19" pageNumber="290" pagination="77 - 353" refId="ref17273" refString="Acciavatti, R. E. & Pearson, D. L. (1989) The tiger beetle Genus Cicindela (Coleoptera, Insecta) from the indian subcontinent. Annals of Carnegie Museum, 58, 77 - 353." type="journal article" year="1989">Acciavatti & Pearson 1989:122</bibRefCitation>
|
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</taxonomicName>
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.
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||
</paragraph>
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<paragraph id="950A3695DF778E0B6986334B8B05F8D1" blockId="19.[151,1436,1677,2041]" box="[151,605,1799,1824]" pageId="19" pageNumber="290">
|
||
<taxonomicName id="52B54D16DF778E0B6986334B8B02F8D1" ID-CoL="PXDS" authority=": Wiesner 1992: 108" authorityName="Wiesner" authorityPageNumber="108" authorityYear="1992" box="[151,602,1799,1824]" class="Insecta" family="Carabidae" genus="Calochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="290" phylum="Arthropoda" rank="species" species="hamiltoniana">
|
||
<emphasis id="A7C1EA87DF778E0B6986334B88C8F8D1" box="[151,400,1800,1824]" italics="true" pageId="19" pageNumber="290">Calochroa hamiltoniana</emphasis>
|
||
:
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||
<bibRefCitation id="F1244B64DF778E0B688C33448B02F8D1" author="Wiesner, J." box="[413,602,1799,1824]" pageId="19" pageNumber="290" refId="ref20965" refString="Wiesner, J. (1992) Checklist of the Tiger Beetles of the World. Verlag Erna Bauer, Keltern, 364 pp." type="book" year="1992">Wiesner 1992:108</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="950A3695DF778E0B6986336A8C11F8B5" blockId="19.[151,1436,1677,2041]" box="[151,1353,1833,1861]" pageId="19" pageNumber="290">
|
||
<emphasis id="A7C1EA87DF778E0B6986336A88CEF8B5" bold="true" box="[151,406,1833,1860]" pageId="19" pageNumber="290">
|
||
<collectingRegion id="5771F877DF778E0B6986336A884DF8B5" box="[151,277,1833,1860]" pageId="19" pageNumber="290">Specimens</collectingRegion>
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||
examined.
|
||
</emphasis>
|
||
(N = 1)
|
||
<materialsCitation id="25DD3CC8DF778E0B68E6336A8C11F8B5" ID-GBIF-Occurrence="2848419311" box="[503,1353,1833,1861]" collectingDate="1977-05" collectorName="C. V. Nidek" country="India" location="South" pageId="19" pageNumber="290" specimenCount="1" specimenCount-male="1" stateProvince="South">
|
||
<emphasis id="A7C1EA87DF778E0B68E6336A8BCFF8B5" bold="true" box="[503,663,1833,1860]" pageId="19" pageNumber="290">
|
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[
|
||
<collectingRegion id="5771F877DF778E0B68EF336A8B1EF8B5" box="[510,582,1833,1860]" pageId="19" pageNumber="290">South</collectingRegion>
|
||
<collectingCountry id="EDA27605DF778E0B6B5C336A8BD7F8B5" box="[589,655,1833,1860]" name="India" pageId="19" pageNumber="290">India</collectingCountry>
|
||
]
|
||
</emphasis>
|
||
<specimenCount id="83B3FD1CDF778E0B6B8C33698B98F8B5" box="[669,704,1834,1860]" pageId="19" pageNumber="290" type="male">1♂</specimenCount>
|
||
(
|
||
<collectionCode id="F3A4AE50DF778E0B6BDE33698A54F8B5" box="[719,780,1834,1860]" pageId="19" pageNumber="290">MHJ</collectionCode>
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||
), Anamalai Hills, Topslip,
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||
<date id="E10B1055DF778E0B6D2633688DC9F8B5" box="[1079,1169,1834,1861]" pageId="19" pageNumber="290" value="1977-05">
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||
<collectingDate id="F14FE9BDDF778E0B6D2633688DC9F8B5" box="[1079,1169,1834,1861]" pageId="19" pageNumber="290" value="1977-05">V. 1977</collectingDate>
|
||
</date>
|
||
,
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||
<collectorName id="38405343DF778E0B6D8D33698C40F8B5" box="[1180,1304,1834,1860]" pageId="19" pageNumber="290">C.V. Nidek</collectorName>
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||
leg.
|
||
</materialsCitation>
|
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DDAF651EDF778E0C69D6330C8BB5FEB2" lastPageId="20" lastPageNumber="291" pageId="19" pageNumber="290" type="description">
|
||
<paragraph id="950A3695DF778E0B69D6330C8BD6F87D" blockId="19.[151,1436,1677,2041]" pageId="19" pageNumber="290">
|
||
<emphasis id="A7C1EA87DF778E0B69D6330C8813F899" box="[199,331,1871,1896]" italics="true" pageId="19" pageNumber="290">Description</emphasis>
|
||
. Internal sac everted dorso-apically in right lateral view. DA largest in bladders; VA as large as VB, B, BLR and VLR; DLL smallest in bladders.
|
||
</paragraph>
|
||
<paragraph id="950A3695DF778E0C69AD33D58C2BFF26" blockId="19.[151,1436,1677,2041]" lastBlockId="20.[151,1437,153,323]" lastPageId="20" lastPageNumber="291" pageId="19" pageNumber="290">VA shortly and roundly everted apically, vested with short spines, which are weakly chitinous. VLR roundly everted apically and weakly chitinous. DA well swollen dorsally. DLL roundly swollen right laterally. VB roundly swollen left latero-ventrally, vested numerously with short spines, which are forming a band. BLR roundly swollen left laterally, vested numerously with short spines, which are weakly chitinous. B roundly swollen right laterally. Flagellum very elongate, S-shaped. Medial tooth short and cylindrical, extending left laterally, sinuous at apex.</paragraph>
|
||
<paragraph id="950A3695DF708E0C69D634A28BB5FEB2" blockId="20.[151,1437,153,323]" pageId="20" pageNumber="291">
|
||
<emphasis id="A7C1EA87DF708E0C69D634A28869FF0A" box="[199,305,225,251]" italics="true" pageId="20" pageNumber="291">Remarks.</emphasis>
|
||
This species shares the characteristics of the position of each bladder with
|
||
<taxonomicName id="52B54D16DF708E0C6D8F34A28875FEEE" authority="(Parry, 1844)" baseAuthorityName="Parry" baseAuthorityYear="1844" class="Insecta" family="Carabidae" genus="Calochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="291" phylum="Arthropoda" rank="species" species="assamensis">
|
||
<emphasis id="A7C1EA87DF708E0C6D8F34A28CC4FF0A" box="[1182,1436,225,251]" italics="true" pageId="20" pageNumber="291">Calochroa assamensis</emphasis>
|
||
(
|
||
<bibRefCitation id="F1244B64DF708E0C698E3546887DFEEE" author="Parry, F. J. S." box="[159,293,261,287]" pageId="20" pageNumber="291" pagination="454" refId="ref20030" refString="Parry, F. J. S. (1844) A decade of new Coleoptera from Assam. The Annals and Magazine of Natural History, 14, 454." type="journal article" year="1844">Parry, 1844</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, but is distinguished by the following characteristics: VA everted apically; DLL not swollen; VB with short spines; medial tooth dextral extending, sinuous.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |