The reclassification of Sophiodela and other tiger beetles (Coleoptera, Cicindelidae) based on the structure of the everted internal sac of the male genitalia Author Fukuda, Yuki Kurayoshi, 111 - 8, Kamogawa-town, Tottori, 682 - 0923 Japan E-mail: yuki 03 go 3 @ gmail. com yuki03go3@gmail.com Author Ogawa, Ryo Matsuyama, 6 - 5 - 8, Hakusuidai, Ehime, 791 - 0113 Japan E-mail: ailmasbl 0854 @ yahoo. co. jp Author Hori, Michio Wakayama, 6 - 1 - 2 Takajyomachi, Wakayama, 640 - 8135 Japan E-mail: hori @ terra. zool. kyoto-u. ac. jp text Zootaxa 2019 2019-08-28 4661 2 271 308 journal article 21156 10.11646/zootaxa.4661.2.3 6c12aca1-7350-4265-9ded-6d415152f181 1175-5326 3998528 95722A8A-E02C-47E5-AA9B-777B6CA226E6 Calochroa hamiltoniana ( Thomson, 1857 ) ( Figs 5C , 9B ) Cicindela hamiltoniana Thomson, 1857:323 . Cicindela flavovittata Chaudoir, 1865:61 [synonymy by Acciavatti & Pearson 1989:127 ]. Calochroa hamiltoniana : Rivalier 1961:127 . Cicindela ( Calochroa ) hamiltoniana : Acciavatti & Pearson 1989:122 . Calochroa hamiltoniana : Wiesner 1992:108 . Specimens examined. (N = 1) [ South India ] 1♂ ( MHJ ), Anamalai Hills, Topslip, V. 1977 , C.V. Nidek leg. Description . Internal sac everted dorso-apically in right lateral view. DA largest in bladders; VA as large as VB, B, BLR and VLR; DLL smallest in bladders. VA shortly and roundly everted apically, vested with short spines, which are weakly chitinous. VLR roundly everted apically and weakly chitinous. DA well swollen dorsally. DLL roundly swollen right laterally. VB roundly swollen left latero-ventrally, vested numerously with short spines, which are forming a band. BLR roundly swollen left laterally, vested numerously with short spines, which are weakly chitinous. B roundly swollen right laterally. Flagellum very elongate, S-shaped. Medial tooth short and cylindrical, extending left laterally, sinuous at apex. Remarks. This species shares the characteristics of the position of each bladder with Calochroa assamensis ( Parry, 1844 ) , but is distinguished by the following characteristics: VA everted apically; DLL not swollen; VB with short spines; medial tooth dextral extending, sinuous.