treatments-xml/data/F5/2B/87/F52B87F65E166161FF74DAF1FC951DB8.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.5584838" ID-GBIF-Dataset="b9f36fb1-1c9d-4af8-96ca-d57973b94862" ID-GBIF-Taxon="192711759" ID-ISSN="1997-3500" ID-Zenodo-Dep="5584838" approvalRequired="1" checkinTime="1619623649741" checkinUser="donat" docAuthor="Seifert, Bernhard" docDate="2021" docId="F52B87F65E166161FF74DAF1FC951DB8" docLanguage="en" docName="myrmecNews.31_133-179_printable.pdf" docOrigin="Myrmecological News 31" docSource="http://zoobank.org/0e55c0d7-531a-48d7-a078-148b96bd461d" docTitle="Formica santschii WHEELER 1913" docType="treatment" docVersion="3" lastPageNumber="165" masterDocId="0912FF8E5E366141FFC9DA41FF91193E" masterDocTitle="A taxonomic revision of the Palaearctic members of the Formica rufa group (Hymenoptera: Formicidae) - the famous mound-building red wood ants" masterLastPageNumber="179" masterPageNumber="133" pageId="32" pageNumber="165" updateTime="1643410080396" updateUser="ExternalLinkService">
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<mods:title>A taxonomic revision of the Palaearctic members of the Formica rufa group (Hymenoptera: Formicidae) - the famous mound-building red wood ants</mods:title>
</mods:titleInfo>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Seifert, Bernhard</mods:namePart>
</mods:name>
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<mods:title>Myrmecological News</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-04-28</mods:number>
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<mods:detail type="volume">
<mods:number>31</mods:number>
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<mods:start>133</mods:start>
<mods:end>179</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.25849/myrmecol.news_031:133</mods:identifier>
<mods:identifier type="GBIF-Dataset">b9f36fb1-1c9d-4af8-96ca-d57973b94862</mods:identifier>
<mods:identifier type="ISSN">1997-3500</mods:identifier>
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<taxonomicName authority="WHEELER, 1913" authorityName="WHEELER" authorityYear="1913" box="[189,626,176,199]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="165" phylum="Arthropoda" rank="species" species="santschii">
<emphasis bold="true" box="[189,626,176,199]" italics="true" pageId="32" pageNumber="165">
Formica santschii
<bibRefCitation author="WHEELER" box="[429,626,177,198]" firstAuthor="WHEELER" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="32" pageNumber="165" pagination="379 - 565" part="53" refId="ref47626" refString="WHEELER, W. M. 1913: A revision of the ants of the genus Formica (LINNE) MAYR. - Bulletin of the Museum of Comparative Zoology 53: 379 - 565." title="A revision of the ants of the genus Formica (LINNE) MAYR" type="journal article" year="1913">WHEELER, 1913</bibRefCitation>
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</paragraph>
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<subSubSection pageId="32" pageNumber="165" type="reference_group">
<paragraph blockId="32.[151,772,176,2023]" pageId="32" pageNumber="165">
<treatmentCitation box="[151,636,209,230]" pageId="32" pageNumber="165">
<taxonomicName authority="WHEELER, 1913" authorityName="WHEELER" authorityYear="1913" box="[151,636,209,230]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="165" phylum="Arthropoda" rank="variety" species="rufa" variety="santschii">
<emphasis box="[151,300,209,230]" italics="true" pageId="32" pageNumber="165">Formica rufa</emphasis>
var.
<emphasis box="[355,454,209,230]" italics="true" pageId="32" pageNumber="165">santschii</emphasis>
<bibRefCitation author="WHEELER" box="[461,636,209,230]" firstAuthor="WHEELER" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="32" pageNumber="165" pagination="379 - 565" part="53" refId="ref47626" refString="WHEELER, W. M. 1913: A revision of the ants of the genus Formica (LINNE) MAYR. - Bulletin of the Museum of Comparative Zoology 53: 379 - 565." title="A revision of the ants of the genus Formica (LINNE) MAYR" type="journal article" year="1913">WHEELER, 1913</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[
<typeStatus box="[648,699,209,230]" pageId="32" pageNumber="165">type</typeStatus>
investigation]
</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="165" type="discussion">
<paragraph blockId="32.[151,772,176,2023]" pageId="32" pageNumber="165">
<taxonomicName authorityName="WHEELER" authorityYear="1913" box="[189,392,273,294]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="165" phylum="Arthropoda" rank="species" species="santschii">
<emphasis box="[189,392,273,294]" italics="true" pageId="32" pageNumber="165">Formica santschii</emphasis>
</taxonomicName>
is
<bibRefCitation author="WHEELER" box="[426,614,273,295]" firstAuthor="WHEELER" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="32" pageNumber="165" pagination="379 - 565" part="53" refId="ref47626" refString="WHEELER, W. M. 1913: A revision of the ants of the genus Formica (LINNE) MAYR. - Bulletin of the Museum of Comparative Zoology 53: 379 - 565." title="A revision of the ants of the genus Formica (LINNE) MAYR" type="journal article" year="1913">WHEELER (1913)</bibRefCitation>
s replacement name for
<treatmentCitation box="[259,647,305,326]" pageId="32" pageNumber="165">
<taxonomicName authority="SANTSCHI, 1911" authorityName="Santschi" authorityYear="1911" box="[259,647,305,326]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="165" phylum="Arthropoda" rank="variety" species="rufa" variety="alpina">
<emphasis box="[259,335,305,326]" italics="true" pageId="32" pageNumber="165">F. rufa</emphasis>
var.
<emphasis box="[392,463,305,326]" italics="true" pageId="32" pageNumber="165">alpina</emphasis>
<bibRefCitation author="SANTSCHI" box="[471,647,305,326]" firstAuthor="SANTSCHI" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="32" pageNumber="165" pagination="349 - 350" part="1911" refId="ref46386" refString="SANTSCHI, F. 1911: Une nouvelle variete de Formica rufa L. (Hym. Formicidae). - Bulletin de la Societe Entomologique de France 1911: 349 - 350." title="Une nouvelle variete de Formica rufa L. (Hym. Formicidae)" type="journal article" year="1911">SANTSCHI, 1911</bibRefCitation>
</taxonomicName>
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, which is a junior primary homonym of
<treatmentCitation pageId="32" pageNumber="165">
<taxonomicName authority="WHEELER, 1909" authorityName="WHEELER" authorityYear="1909" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="165" phylum="Arthropoda" rank="variety" species="adamsi" variety="alpina">
<emphasis box="[464,647,337,358]" italics="true" pageId="32" pageNumber="165">Formica adamsi</emphasis>
var.
<emphasis box="[700,771,337,358]" italics="true" pageId="32" pageNumber="165">alpina</emphasis>
<bibRefCitation box="[151,332,369,390]" pageId="32" pageNumber="165">WHEELER, 1909</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<bibRefCitation author="SANTSCHI" box="[344,529,369,391]" firstAuthor="SANTSCHI" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="32" pageNumber="165" pagination="349 - 350" part="1911" refId="ref46386" refString="SANTSCHI, F. 1911: Une nouvelle variete de Formica rufa L. (Hym. Formicidae). - Bulletin de la Societe Entomologique de France 1911: 349 - 350." title="Une nouvelle variete de Formica rufa L. (Hym. Formicidae)" type="journal article" year="1911">SANTSCHI (1911)</bibRefCitation>
described his
<taxonomicName authorityName="Santschi" authorityYear="1911" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="165" phylum="Arthropoda" rank="variety" species="rufa" variety="alpina">
<emphasis box="[694,771,369,390]" italics="true" pageId="32" pageNumber="165">F. rufa</emphasis>
var.
<emphasis box="[199,270,401,422]" italics="true" pageId="32" pageNumber="165">alpina</emphasis>
</taxonomicName>
from the Alps north of Sondrio. Investigated were
<specimenCount box="[214,457,433,455]" pageId="32" pageNumber="165" type="worker" typeStatus="syntype">two syntype workers</specimenCount>
on separate pins stored in NHM Basel and labelled in F. Santschi´s handwriting. The larger specimen with CW
<quantity box="[433,526,496,518]" metricMagnitude="0" metricUnit="m" metricValue="1.754" pageId="32" pageNumber="165" unit="mm" value="1754.0">1754 µm</quantity>
is depicted in AntWeb (
<bibRefCitation accessDate="2021-01-15" author="ANTWEB" box="[160,319,529,551]" firstAuthor="ANTWEB" pageId="32" pageNumber="165" publicationUrl="https://www.antweb.org&gt;" refId="ref44610" refString="ANTWEB 2021: AntWeb Version 8.54. 4. - &lt;https: // www. antweb. org&gt;, retrieved on 15 January 2021." title="AntWeb Version 8.54. 4." type="url" year="2021">
<collectionCode box="[160,257,529,550]" pageId="32" pageNumber="165">ANTWEB</collectionCode>
2021
</bibRefCitation>
) under CASENT0912249 and is labelled “Valteline. Galli-Valerio”, “
<taxonomicName authority="Sant." authorityName="Santschi" authorityYear="1911" box="[436,599,561,583]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="165" phylum="Arthropoda" rank="species" species="alpina">Formica alpina</taxonomicName>
€ Sant.”, “type”. The smaller, damaged specimen with CW =
<quantity box="[642,736,592,614]" metricMagnitude="0" metricUnit="m" metricValue="1.685" pageId="32" pageNumber="165" unit="mm" value="1685.0">1685 µm</quantity>
on the other pin is labelled “Valteline. Galli-Valerio”, “
<taxonomicName authority="Sant." authorityName="Santschi" authorityYear="1911" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="165" phylum="Arthropoda" rank="species" species="alpina">Formica alpina</taxonomicName>
€ Sant.” By morphology and geography, the
<taxonomicName authorityName="WHEELER" authorityYear="1913" box="[151,279,689,710]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="165" phylum="Arthropoda" rank="species" species="santschii">
<emphasis box="[151,279,689,710]" italics="true" pageId="32" pageNumber="165">F. santschii</emphasis>
</taxonomicName>
types could either belong to
<taxonomicName authorityName="ZETTERSTEDT" authorityYear="1838" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="165" phylum="Arthropoda" rank="species" species="lugubris">
<emphasis italics="true" pageId="32" pageNumber="165">Formica lugubris</emphasis>
</taxonomicName>
or
<taxonomicName authorityName="SEIFERT" authorityYear="1996" box="[236,484,721,742]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="165" phylum="Arthropoda" rank="species" species="paralugubris">
<emphasis box="[236,484,721,742]" italics="true" pageId="32" pageNumber="165">Formica paralugubris</emphasis>
</taxonomicName>
. The specimens were run as wild-cards in an LDA collecting
<specimenCount box="[557,696,753,775]" pageId="32" pageNumber="165" type="worker">369 workers</specimenCount>
of the
<taxonomicName authorityName="ZETTERSTEDT" authorityYear="1838" box="[151,273,785,806]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="165" phylum="Arthropoda" rank="species" species="lugubris">
<emphasis box="[151,273,785,806]" italics="true" pageId="32" pageNumber="165">F. lugubris</emphasis>
</taxonomicName>
morphs A1 and A3 from the Alps in class 1, and
<specimenCount box="[198,332,817,839]" pageId="32" pageNumber="165" type="worker">298 workers</specimenCount>
of
<taxonomicName authorityName="SEIFERT" authorityYear="1996" box="[364,537,817,838]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="165" phylum="Arthropoda" rank="species" species="paralugubris">
<emphasis box="[364,537,817,838]" italics="true" pageId="32" pageNumber="165">F. paralugubris</emphasis>
</taxonomicName>
in class 2. The eleven characters available in the
<taxonomicName authorityName="WHEELER" authorityYear="1913" box="[462,593,849,870]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="165" phylum="Arthropoda" rank="species" species="santschii">
<emphasis box="[462,593,849,870]" italics="true" pageId="32" pageNumber="165">F. santschii</emphasis>
</taxonomicName>
types were
<collectionCode box="[733,766,849,870]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="32" pageNumber="165">CS</collectionCode>
, CL / CW
<subScript attach="left" box="[251,289,890,904]" fontSize="6" pageId="32" pageNumber="165">1750</subScript>
, SL /
<collectionCode box="[362,392,881,902]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="32" pageNumber="165">CS</collectionCode>
<subScript attach="left" box="[392,430,890,904]" fontSize="6" pageId="32" pageNumber="165">1750</subScript>
, SL / Smax
<subScript attach="left" box="[564,602,890,904]" fontSize="6" pageId="32" pageNumber="165">1750</subScript>
, PeW /
<collectionCode box="[696,726,881,902]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="32" pageNumber="165">CS</collectionCode>
<subScript attach="left" box="[726,764,890,904]" fontSize="6" pageId="32" pageNumber="165">1750</subScript>
, nSc
<subScript attach="left" box="[191,229,922,936]" fontSize="6" pageId="32" pageNumber="165">1750</subScript>
, nCH
<subScript attach="left" box="[294,332,922,936]" fontSize="6" pageId="32" pageNumber="165">1750</subScript>
, OccHL
<subScript attach="left" box="[424,462,922,936]" fontSize="6" pageId="32" pageNumber="165">1750</subScript>
, mPnHL
<subScript attach="right" box="[565,603,922,936]" fontSize="6" pageId="32" pageNumber="165">1750</subScript>
, nMet
<subScript attach="left" box="[676,714,922,936]" fontSize="6" pageId="32" pageNumber="165">1750</subScript>
, and MetHL
<subScript attach="left" box="[230,268,954,968]" fontSize="6" pageId="32" pageNumber="165">1750</subScript>
. With these data, the larger
<typeStatus box="[597,685,945,966]" pageId="32" pageNumber="165">syntype</typeStatus>
worker was allocated to class 1 with p = 0.9674 but the smaller one with p = 0.9551 to class 2. It appears questionable if the
<specimenCount box="[193,328,1041,1063]" pageId="32" pageNumber="165" type="worker">two workers</specimenCount>
mounted on separate pins belong to the same nest sample. In order to save a stable nomenclature, a
<typeStatus box="[168,267,1105,1127]" pageId="32" pageNumber="165">lectotype</typeStatus>
of
<taxonomicName authorityName="WHEELER" authorityYear="1913" box="[297,418,1105,1126]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="165" phylum="Arthropoda" rank="species" species="santschii">
<emphasis box="[297,418,1105,1126]" italics="true" pageId="32" pageNumber="165">F. santschii</emphasis>
</taxonomicName>
is fixed by present decision in the undamaged larger specimen labelled “CASENT0912249”.
</paragraph>
</subSubSection>
</treatment>
</document>