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<document id="6A1AFA080AB686A0D629DD57A2C3AE77" ID-DOI="10.25226/bboc.v142i2.2022.a5" ID-ISSN="2513-9894" ID-Zenodo-Dep="12535806" ID-ZooBank="F94D7878-8828-48D4-A402-080589626BE5" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1719341608131" checkinUser="felipe" docAuthor="Halley, Matthew R." docDate="2022" docId="F273857CFFC9D10D59B2A5A0FE526637" docLanguage="en" docName="BullBritOrnitholClub.142.2.209-223.pdf" docOrigin="Bulletin of the British Ornithologists’ Club 142 (2)" docSource="http://dx.doi.org/10.25226/bboc.v142i2.2022.a5" docStyle="DocumentStyle:EFA7F293C51FD4806C5290561C16F223.2:BullBritOrnitholClub.2017-.journal_article" docStyleId="EFA7F293C51FD4806C5290561C16F223" docStyleName="BullBritOrnitholClub.2017-.journal_article" docStyleVersion="2" docTitle="Sphenopsis frontalis Oleaginous" docType="treatment" docVersion="1" lastPageNumber="222" masterDocId="0E4AFD04FFC6D102585AA521FFCD6356" masterDocTitle="Taxonomic status of the Western Hemispingus Sphenopsis ochracea (Thraupidae) and a review of species limits in the genus Sphenopsis P. L. Sclater, 1861" masterLastPageNumber="223" masterPageNumber="209" pageNumber="222" updateTime="1734544480592" updateUser="carolina" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="AF3E0DE8C5DD8AB1F8119B1246B29871">Taxonomic status of the Western Hemispingus Sphenopsis ochracea (Thraupidae) and a review of species limits in the genus Sphenopsis P. L. Sclater, 1861</mods:title>
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<mods:namePart id="E70DF2A63FF08C337AFA50B449CE0D64">Halley, Matthew R.</mods:namePart>
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<mods:title id="ECE4F62B27F831EA793F191CB550322E">Bulletin of the British Ornithologists’ Club</mods:title>
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<mods:date id="93D9B5612B51F3645A3601936AFD2F77">2022</mods:date>
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<mods:number id="DE597685E89D04B0A867966F68B1477B">2022-06-03</mods:number>
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<mods:classification id="995C88F7BE2C8E48B6CF7AFA42255468">journal article</mods:classification>
<mods:identifier id="C2718CA1C7D939BC728EF1FFB8BD5C78" type="DOI">10.25226/bboc.v142i2.2022.a5</mods:identifier>
<mods:identifier id="64ED898A17AA15FBE7C97A8EE0FF4074" type="ISSN">2513-9894</mods:identifier>
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</mods:mods>
<treatment id="F273857CFFC9D10D59B2A5A0FE526637" LSID="urn:lsid:plazi:treatment:F273857CFFC9D10D59B2A5A0FE526637" httpUri="http://treatment.plazi.org/id/F273857CFFC9D10D59B2A5A0FE526637" lastPageNumber="222" pageId="15" pageNumber="222">
<subSubSection id="32C067E1FFC9D10D59B2A5A0FCCB63FB" pageId="15" pageNumber="222" type="nomenclature">
<paragraph id="7A65346AFFC9D10D59B2A5A0FCCB63FB" blockId="15.[488,820,129,198]" pageId="15" pageNumber="222">
<taxonomicName id="BDDA4FE9FFC9D10D59B2A5A0FD4C63FB" ID-CoL="9GBHG" authority="(von Tschudi) Oleaginous" authorityName="Oleaginous" baseAuthorityName="von Tschudi" baseAuthorityYear="1844" class="Aves" family="Thraupidae" genus="Sphenopsis" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="frontalis">
<emphasis id="48AEE878FFC9D10D59B2A5A0FD6663C2" bold="true" box="[488,683,129,148]" italics="true" pageId="15" pageNumber="222">Sphenopsis frontalis</emphasis>
(von Tschudi) Oleaginous
</taxonomicName>
<taxonomicName id="BDDA4FE9FFC9D10D5ADDA5BBFCCB63FB" authorityName="Cabanis" authorityYear="1851" box="[647,774,154,173]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="genus">Hemispingus</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="32C067E1FFC9D10D5A37A592FD626390" box="[621,687,179,198]" pageId="15" pageNumber="222" type="description">
<paragraph id="7A65346AFFC9D10D5A37A592FD626390" blockId="15.[488,820,129,198]" box="[621,687,179,198]" pageId="15" pageNumber="222">
(
<figureCitation id="E2E128EFFFC9D10D5A29A592FD656390" box="[627,680,179,198]" captionStart="Figure 5" captionStartId="10.[144,204,873,892]" captionTargetBox="[144,1164,141,858]" captionTargetId="figure-258@10.[141,704,126,858]" captionTargetPageId="10" captionText="Figure 5 (left). Lateral view of Oleaginous Hemispingus Sphenopsis frontalis (Chapman, 1923). From left to right, (1) ANSP 141983, adult female S. f. frontalis prepared by Kjell von Sneidern on 20 March 1939 at ‘La Costa, Huila, Colombia’. (2) ANSP 185826, adult male S. f. frontalis, prepared by Tristan J. Davis on 22 July 1992 at ‘Panguri; ca. 12 km NE San Francisco del Vergel’, Zamora Chinchipe, Ecuador. (3) ANSP 83780, adult male S. f. frontalis, collected on 15 September 1922 at ‘Baeza, Ecuador’. (4) ANSP 67191, adult male S. f.hanieli, collected on 3 March 1914 at ‘Galeparo; Curro Del Avito, Venezuela’ (Matthew R. Halley) Figure 6 (right). Lateral view of Piura Hemispingus Sphenopsis piurae (Chapman, 1923). From left to right, (1) ANSP 116357, adult female prepared by M. A. Carriker, Jr., on 21 June 1933 at ‘Palambla, D. Piura’, Peru. (2) ANSP 116352, adult male, prepared by M. A. Carriker, Jr., on 22 June 1933 at ‘Palambla, D. Piura’, Peru. (3) ANSP 116358, adult male, prepared by M. A. Carriker, Jr., on 24 August 1933 at ‘Chira, D. Cajamarca’, Peru (Matthew R. Halley)" figureDoi="http://doi.org/10.5281/zenodo.12535815" httpUri="https://zenodo.org/record/12535815/files/figure.png" pageId="15" pageNumber="222">Fig. 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="32C067E1FFC9D10D58CAA5C3FC02620F" pageId="15" pageNumber="222" type="reference_group">
<paragraph id="7A65346AFFC9D10D58CAA5C3FDC763A0" blockId="15.[144,1165,226,990]" box="[144,522,226,246]" pageId="15" pageNumber="222">
<taxonomicName id="BDDA4FE9FFC9D10D58CAA5C3FDC863A0" authority="von Tschudi, 1844" authorityName="von Tschudi" authorityYear="1844" box="[144,517,226,246]" class="Aves" family="Vireonidae" genus="Hylophilus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="frontalis">
<emphasis id="48AEE878FFC9D10D58CAA5C3FE9E63A3" bold="true" box="[144,339,226,245]" italics="true" pageId="15" pageNumber="222">Hylophilus frontalis</emphasis>
von Tschudi, 1844
</taxonomicName>
.
</paragraph>
<paragraph id="7A65346AFFC9D10D589AA5DDFCF96271" blockId="15.[144,1165,226,990]" pageId="15" pageNumber="222">
<typeStatus id="A5618AC8FFC9D10D589AA5DDFEED6259" box="[192,288,252,271]" pageId="15" pageNumber="222" type="lectotype">Lectotype</typeStatus>
: MHNN 92.8817, in
<collectingMunicipality id="9A01AE10FFC9D10D59B9A5DDFD306258" box="[483,765,251,271]" pageId="15" pageNumber="222">Muséum d’Histoire Naturelle</collectingMunicipality>
de Neuchâtel, Switzerland (
<emphasis id="48AEE878FFC9D10D5C57A5DAFBE66258" box="[1037,1067,251,270]" italics="true" pageId="15" pageNumber="222">fide</emphasis>
Hellmayr 1936: 423): eastern slope of
<location id="7F0562B1FFC9D10D599BA435FDCD6271" LSID="urn:lsid:plazi:treatment:F273857CFFC9D10D59B2A5A0FE526637:7F0562B1FFC9D10D599BA435FDCD6271" box="[449,512,276,295]" country="Switzerland" municipality="Museum d'Histoire Naturelle" name="Andes" pageId="15" pageNumber="222" stateProvince="Junin">Andes</location>
,
<collectingRegion id="B81EFA88FFC9D10D5A50A435FDF26271" box="[522,575,276,295]" country="Peru" name="Junin" pageId="15" pageNumber="222">Junín</collectingRegion>
,
<collectingCountry id="02CD74FAFFC9D10D5A12A434FDB9627E" box="[584,628,277,296]" name="Peru" pageId="15" pageNumber="222">Peru</collectingCountry>
(Desfayes 1994: 90).
</paragraph>
<paragraph id="7A65346AFFC9D10D58CAA40CFC646260" blockId="15.[144,1165,226,990]" box="[144,937,298,321]" pageId="15" pageNumber="222">
<emphasis id="48AEE878FFC9D10D58CAA40CFF276216" bold="true" box="[144,234,301,320]" italics="true" pageId="15" pageNumber="222">Sphenops</emphasis>
[
<emphasis id="48AEE878FFC9D10D58ADA40FFEC36216" box="[247,270,302,320]" italics="true" pageId="15" pageNumber="222">sic</emphasis>
]
<emphasis id="48AEE878FFC9D10D5941A40CFEA26216" bold="true" box="[283,367,301,320]" italics="true" pageId="15" pageNumber="222">ignobilis</emphasis>
Sclater 1861: 160 (see Coues 1879: 278 for publication date).
<superScript id="8DAF9922FFC9D10D5BF9A40BFC646260" attach="none" box="[931,937,298,310]" fontSize="5" pageId="15" pageNumber="222">3</superScript>
</paragraph>
<paragraph id="7A65346AFFC9D10D58CAA464FC02620F" blockId="15.[144,1165,226,990]" box="[144,975,325,345]" pageId="15" pageNumber="222">
<taxonomicName id="BDDA4FE9FFC9D10D58CAA464FDF8620F" authority="P. L. Sclater, 1862 a: 379" authorityName="P. L. Sclater" authorityPageNumber="379" authorityYear="1862" box="[144,565,325,345]" class="Aves" family="Thraupidae" genus="Sphenopsis" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="ignobilis">
<emphasis id="48AEE878FFC9D10D58CAA464FE99620E" bold="true" box="[144,340,325,344]" italics="true" pageId="15" pageNumber="222">Sphenopsis ignobilis</emphasis>
P. L. Sclater, 1862a: 379
</taxonomicName>
(see Duncan 1937: 72 for publication date).
</paragraph>
</subSubSection>
<subSubSection id="32C067E1FFC9D10D589AA47EFBBB623E" box="[192,1142,348,370]" pageId="15" pageNumber="222" type="materials_examined">
<paragraph id="7A65346AFFC9D10D589AA47EFBBB623E" blockId="15.[144,1165,226,990]" box="[192,1142,348,370]" pageId="15" pageNumber="222">
<typeStatus id="A5618AC8FFC9D10D589AA47EFED16224" box="[192,284,351,370]" pageId="15" pageNumber="222" type="holotype">Holotype</typeStatus>
: NHMUK 1885.6.12.876: ‘in Brasilia’ =
<collectingRegion id="B81EFA88FFC9D10D5AD3A47EFD1D6224" box="[649,720,351,370]" country="Venezuela" name="Merida" pageId="15" pageNumber="222">Mérida</collectingRegion>
,
<collectingCountry id="02CD74FAFFC9D10D5A83A47EFCF76224" box="[729,826,351,370]" name="Venezuela" pageId="15" pageNumber="222">Venezuela</collectingCountry>
(see
<collectorName id="D72F51BCFFC9D10D5B30A47EFC7E6224" box="[874,947,351,370]" pageId="15" pageNumber="222">Paynter</collectorName>
&amp;
<collectorName id="D72F51BCFFC9D10D5B95A47EFBC56224" box="[975,1032,351,370]" pageId="15" pageNumber="222">Storer</collectorName>
1970: 265).
<superScript id="8DAF9922FFC9D10D5C2AA47DFBBB623E" attach="none" box="[1136,1142,348,360]" fontSize="5" pageId="15" pageNumber="222">4</superScript>
</paragraph>
</subSubSection>
<subSubSection id="32C067E1FFC9D10D58CAA456FC8362DC" box="[144,846,375,395]" pageId="15" pageNumber="222" type="reference_group">
<paragraph id="7A65346AFFC9D10D58CAA456FC8362DC" blockId="15.[144,1165,226,990]" box="[144,846,375,395]" pageId="15" pageNumber="222">
<taxonomicName id="BDDA4FE9FFC9D10D58CAA456FDA562DD" authority="P. L. Sclater, 1862 b: 110" authorityName="P. L. Sclater" authorityPageNumber="110" authorityYear="1862" box="[144,616,375,395]" class="Aves" family="Passerellidae" genus="Chlorospingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="oleagineus">
<emphasis id="48AEE878FFC9D10D58CAA456FE4962DC" bold="true" box="[144,388,375,394]" italics="true" pageId="15" pageNumber="222">Chlorospingus oleagineus</emphasis>
P. L. Sclater, 1862b: 110
</taxonomicName>
, Taczanowski 1884: 516.
</paragraph>
</subSubSection>
<subSubSection id="32C067E1FFC9D10D589AA4B1FEEE62EA" pageId="15" pageNumber="222" type="materials_examined">
<paragraph id="7A65346AFFC9D10D589AA4B1FEEE62EA" blockId="15.[144,1165,226,990]" pageId="15" pageNumber="222">
<typeStatus id="A5618AC8FFC9D10D589AA4B1FED162F5" box="[192,284,400,419]" pageId="15" pageNumber="222" type="holotype">Holotype</typeStatus>
: NHMUK 1885.6.12.877: ‘In Nov. Granada int.’ =
<collectingRegion id="B81EFA88FFC9D10D5AADA4B1FCF662F5" box="[759,827,400,419]" country="Colombia" name="Bogota" pageId="15" pageNumber="222">Bogotá</collectingRegion>
,
<collectingCountry id="02CD74FAFFC9D10D5B1FA4B1FC6C62F5" box="[837,929,400,419]" name="Colombia" pageId="15" pageNumber="222">Colombia</collectingCountry>
(see
<collectorName id="D72F51BCFFC9D10D5B89A4B0FBD462F2" box="[979,1049,401,420]" pageId="15" pageNumber="222">Warren</collectorName>
&amp;
<collectorName id="D72F51BCFFC9D10D5C62A4B1FB4062F5" box="[1080,1165,400,419]" pageId="15" pageNumber="222">Harrison</collectorName>
1971: 402).
</paragraph>
</subSubSection>
<subSubSection id="32C067E1FFC9D10D58CAA4E3FD5A61CD" pageId="15" pageNumber="222" type="reference_group">
<paragraph id="7A65346AFFC9D10D58CAA4E3FED162B8" blockId="15.[144,1165,226,990]" pageId="15" pageNumber="222">
<taxonomicName id="BDDA4FE9FFC9D10D58CAA4E3FDAA6283" authority="Sclater &amp; Salvin 1871: 784" authorityName="Sclater &amp; Salvin" authorityPageNumber="784" authorityYear="1871" box="[144,615,450,469]" class="Aves" family="Passerellidae" genus="Chlorospingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="ignobilis">
<emphasis id="48AEE878FFC9D10D58CAA4E3FEBE6283" bold="true" box="[144,371,450,469]" italics="true" pageId="15" pageNumber="222">Chlorospingus ignobilis</emphasis>
Sclater &amp; Salvin 1871: 784
</taxonomicName>
(see Dickinson 2005 for publication date), Sclater &amp; Salvin 1879: 504.
</paragraph>
<paragraph id="7A65346AFFC9D10D58CAA4D2FC6C6151" blockId="15.[144,1165,226,990]" box="[144,929,499,519]" pageId="15" pageNumber="222">
<taxonomicName id="BDDA4FE9FFC9D10D58CAA4D2FD9D6150" authority="Taczanowski 1884: 517" authorityName="Taczanowski" authorityPageNumber="517" authorityYear="1884" box="[144,592,499,519]" class="Aves" family="Passerellidae" genus="Chlorospingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="frontalis">
<emphasis id="48AEE878FFC9D10D58CAA4D2FEBF6150" bold="true" box="[144,370,499,518]" italics="true" pageId="15" pageNumber="222">Chlorospingus frontalis</emphasis>
Taczanowski 1884: 517
</taxonomicName>
, von Berlepsch &amp; Hellmayr 1905: 8.
</paragraph>
<paragraph id="7A65346AFFC9D10D58CAA72DFCD66149" blockId="15.[144,1165,226,990]" box="[144,795,524,544]" pageId="15" pageNumber="222">
<taxonomicName id="BDDA4FE9FFC9D10D58CAA72DFCDA6149" authority="von Berlepsch &amp; Hellmayr 1905: 9" authorityName="von Berlepsch &amp; Hellmayr" authorityPageNumber="9" authorityYear="1905" box="[144,791,524,544]" class="Aves" family="Passerellidae" genus="Chlorospingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="subSpecies" species="frontalis" subSpecies="ignobilis">
<emphasis id="48AEE878FFC9D10D58CAA72DFE016149" bold="true" box="[144,460,524,543]" italics="true" pageId="15" pageNumber="222">Chlorospingus frontalis ignobilis</emphasis>
von Berlepsch &amp; Hellmayr 1905: 9
</taxonomicName>
.
</paragraph>
<paragraph id="7A65346AFFC9D10D58CAA704FC72616E" blockId="15.[144,1165,226,990]" box="[144,959,549,569]" pageId="15" pageNumber="222">
<taxonomicName id="BDDA4FE9FFC9D10D58CAA704FD95616E" authority="von Berlepsch 1912: 1094" authorityName="von Berlepsch 1912:" authorityPageNumber="1094" authorityYear="1912" box="[144,600,549,569]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="frontalis">
<emphasis id="48AEE878FFC9D10D58CAA704FEA9616E" bold="true" box="[144,356,549,568]" italics="true" pageId="15" pageNumber="222">Hemispingus frontalis</emphasis>
von Berlepsch 1912: 1094
</taxonomicName>
, Isler &amp; Isler 1987: 77, Hilty 2011: 167.
</paragraph>
<paragraph id="7A65346AFFC9D10D58CAA71FFD0A6107" blockId="15.[144,1165,226,990]" box="[144,711,574,593]" pageId="15" pageNumber="222">
<taxonomicName id="BDDA4FE9FFC9D10D58CAA71FFD0E6107" authority="von Berlepsch 1912: 1094" authorityName="von Berlepsch" authorityPageNumber="1094" authorityYear="1912" box="[144,707,574,593]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="subSpecies" species="frontalis" subSpecies="oleagineus">
<emphasis id="48AEE878FFC9D10D58CAA71FFE026107" bold="true" box="[144,463,574,593]" italics="true" pageId="15" pageNumber="222">Hemispingus frontalis oleagineus</emphasis>
von Berlepsch 1912: 1094
</taxonomicName>
.
</paragraph>
<paragraph id="7A65346AFFC9D10D58CAA776FED161D5" blockId="15.[144,1165,226,990]" pageId="15" pageNumber="222">
<taxonomicName id="BDDA4FE9FFC9D10D58CAA776FD0E613C" authority="von Berlepsch 1912: 1094" authorityName="von Berlepsch" authorityPageNumber="1094" authorityYear="1912" box="[144,707,598,618]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="subSpecies" species="frontalis" subSpecies="ignobilis">
<emphasis id="48AEE878FFC9D10D58CAA776FE0E613F" bold="true" box="[144,451,598,618]" italics="true" pageId="15" pageNumber="222">Hemispingus frontalis ignobilis</emphasis>
von Berlepsch 1912: 1094
</taxonomicName>
, Chapman 1925: 13, Hellmayr 1936: 424, Hilty 2011: 167.
</paragraph>
<paragraph id="7A65346AFFC9D10D58CAA7A9FD5A61CD" blockId="15.[144,1165,226,990]" box="[144,663,648,668]" pageId="15" pageNumber="222">
<taxonomicName id="BDDA4FE9FFC9D10D58CAA7A9FD5E61CD" authority="Hellmayr &amp; von Seilern, 1914: 87" authorityName="Hellmayr &amp; von Seilern" authorityPageNumber="87" authorityYear="1914" box="[144,659,648,668]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="hanieli">
<emphasis id="48AEE878FFC9D10D58CAA7A9FE9E61CD" bold="true" box="[144,339,648,667]" italics="true" pageId="15" pageNumber="222">Hemispingus hanieli</emphasis>
Hellmayr &amp; von Seilern, 1914: 87
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="32C067E1FFC9D10D589AA780FB40619B" pageId="15" pageNumber="222" type="materials_examined">
<paragraph id="7A65346AFFC9D10D589AA780FB40619B" blockId="15.[144,1165,226,990]" pageId="15" pageNumber="222">
<typeStatus id="A5618AC8FFC9D10D589AA780FED161E2" box="[192,284,673,692]" pageId="15" pageNumber="222" type="holotype">Holotype</typeStatus>
: ZSM 13925, in Zoologische Staatssammlung, Munich (
<emphasis id="48AEE878FFC9D10D5B6CA780FC9961E2" box="[822,852,673,692]" italics="true" pageId="15" pageNumber="222">fide</emphasis>
<collectorName id="D72F51BCFFC9D10D5B06A780FC7B61E2" box="[860,950,673,692]" pageId="15" pageNumber="222">Hellmayr</collectorName>
1936): Galipán,
<location id="7F0562B1FFC9D10D5C0CA780FED56199" LSID="urn:lsid:plazi:treatment:F273857CFFC9D10D59B2A5A0FE526637:7F0562B1FFC9D10D5C0CA780FED56199" country="Venezuela" name="Cerro del Avila" pageId="15" pageNumber="222">Cerro del Ávila</location>
, Distrito Federal,
<collectingCountry id="02CD74FAFFC9D10D59E5A79BFDED619B" box="[447,544,698,717]" name="Venezuela" pageId="15" pageNumber="222">Venezuela</collectingCountry>
(see
<collectorName id="D72F51BCFFC9D10D5A15A79BFD64619B" box="[591,681,698,717]" pageId="15" pageNumber="222">Hellmayr</collectorName>
&amp;
<collectorName id="D72F51BCFFC9D10D5A9EA79BFCE2619B" box="[708,815,698,717]" pageId="15" pageNumber="222">von Seilern</collectorName>
1914: 87,
<collectorName id="D72F51BCFFC9D10D5BD3A79BFC1F619B" box="[905,978,698,717]" pageId="15" pageNumber="222">Paynter</collectorName>
&amp;
<collectorName id="D72F51BCFFC9D10D5BB6A79BFBE8619B" box="[1004,1061,698,717]" pageId="15" pageNumber="222">Storer</collectorName>
1970: 266).
</paragraph>
</subSubSection>
<subSubSection id="32C067E1FFC9D10D58CAA7F2FBCD604E" pageId="15" pageNumber="222" type="reference_group">
<paragraph id="7A65346AFFC9D10D58CAA7F2FE5061A9" blockId="15.[144,1165,226,990]" pageId="15" pageNumber="222">
<taxonomicName id="BDDA4FE9FFC9D10D58CAA7F2FDB061B0" authority="Chapman 1921: 122" authorityName="Chapman" authorityPageNumber="122" authorityYear="1921" box="[144,637,722,742]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="subSpecies" species="frontalis" subSpecies="frontalis">
<emphasis id="48AEE878FFC9D10D58CAA7F2FE7161B3" bold="true" box="[144,444,722,742]" italics="true" pageId="15" pageNumber="222">Hemispingus frontalis frontalis</emphasis>
Chapman 1921: 122
</taxonomicName>
, Chapman 1925: 13, Zimmer 1947: 17, Isler &amp; Isler 1987: 77, Hellmayr 1936: 423.
</paragraph>
<paragraph id="7A65346AFFC9D10D58CAA625FBCD604E" blockId="15.[144,1165,226,990]" box="[144,1024,772,792]" pageId="15" pageNumber="222">
<taxonomicName id="BDDA4FE9FFC9D10D58CAA625FDA2604E" authority="Chapman, 1925: 13" authorityName="Chapman" authorityPageNumber="13" authorityYear="1925" box="[144,623,772,792]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="subSpecies" species="frontalis" subSpecies="iteratus">
<emphasis id="48AEE878FFC9D10D58CAA625FE786041" bold="true" box="[144,437,772,791]" italics="true" pageId="15" pageNumber="222">Hemispingus frontalis iteratus</emphasis>
Chapman, 1925: 13
</taxonomicName>
, Hellmayr 1936: 425, Isler &amp; Isler 1987: 77.
</paragraph>
</subSubSection>
<subSubSection id="32C067E1FFC9D10D589AA63CFD74601F" pageId="15" pageNumber="222" type="materials_examined">
<paragraph id="7A65346AFFC9D10D589AA63CFD74601F" blockId="15.[144,1165,226,990]" pageId="15" pageNumber="222">
<typeStatus id="A5618AC8FFC9D10D589AA63CFED16066" box="[192,284,797,816]" pageId="15" pageNumber="222" type="holotype">Holotype</typeStatus>
: AMNH 188022, in
<location id="7F0562B1FFC9D10D59A9A63CFC4D6066" LSID="urn:lsid:plazi:treatment:F273857CFFC9D10D59B2A5A0FE526637:7F0562B1FFC9D10D59A9A63CFC4D6066" box="[499,896,797,817]" country="Venezuela" county="Mt. Turumiquire" municipality="Carapas" name="American Museum of Natural History" pageId="15" pageNumber="222" stateProvince="Sucre">American Museum of Natural History</location>
,
<location id="7F0562B1FFC9D10D5BCAA63FFC366066" LSID="urn:lsid:plazi:treatment:F273857CFFC9D10D59B2A5A0FE526637:7F0562B1FFC9D10D5BCAA63FFC366066" box="[912,1019,797,817]" country="Venezuela" county="Mt. Turumiquire" municipality="Carapas" name="New York" pageId="15" pageNumber="222" stateProvince="Sucre">New York</location>
:
<collectingMunicipality id="9A01AE10FFC9D10D5C51A63CFB976066" box="[1035,1114,797,816]" pageId="15" pageNumber="222">Carapas</collectingMunicipality>
,
<collectingCounty id="93044CE6FFC9D10D5C31A63FFEF6601F" pageId="15" pageNumber="222">Mt. Turumiquire</collectingCounty>
,
<collectingRegion id="B81EFA88FFC9D10D5919A617FEB4601F" box="[323,377,822,841]" country="Venezuela" name="Sucre" pageId="15" pageNumber="222">Sucre</collectingRegion>
,
<collectingCountry id="02CD74FAFFC9D10D59D9A617FE29601F" box="[387,484,822,841]" name="Venezuela" pageId="15" pageNumber="222">Venezuela</collectingCountry>
(see LeCroy 2012: 68).
</paragraph>
</subSubSection>
<subSubSection id="32C067E1FFC9D10D58CAA66EFC8F602D" pageId="15" pageNumber="222" type="reference_group">
<paragraph id="7A65346AFFC9D10D58CAA66EFB476034" blockId="15.[144,1165,226,990]" box="[144,1162,846,866]" pageId="15" pageNumber="222">
<taxonomicName id="BDDA4FE9FFC9D10D58CAA66EFDAC6034" authority="Chapman 1925: 13" authorityName="Chapman" authorityPageNumber="13" authorityYear="1925" box="[144,609,846,866]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="subSpecies" species="frontalis" subSpecies="hanieli">
<emphasis id="48AEE878FFC9D10D58CAA66EFE616037" bold="true" box="[144,428,846,866]" italics="true" pageId="15" pageNumber="222">Hemispingus frontalis hanieli</emphasis>
Chapman 1925: 13
</taxonomicName>
, Hellmayr 1936: 425, Isler &amp; Isler 1987: 77, Hilty 2011: 167.
</paragraph>
<paragraph id="7A65346AFFC9D10D58CAA646FC8F602D" blockId="15.[144,1165,226,990]" box="[144,834,871,891]" pageId="15" pageNumber="222">
<taxonomicName id="BDDA4FE9FFC9D10D58CAA646FE56602C" baseAuthorityName="Tschudi" baseAuthorityYear="1844" box="[144,411,871,891]" class="Aves" family="Tyrannidae" genus="Hemisphingus" isUncertain="true" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="frontalis">
<emphasis id="48AEE878FFC9D10D58CAA646FEDA602C" bold="true" box="[144,279,871,890]" italics="true" pageId="15" pageNumber="222">Hemisphingus</emphasis>
[
<emphasis id="48AEE878FFC9D10D597EA649FEF6602C" box="[292,315,872,890]" italics="true" pageId="15" pageNumber="222">sic</emphasis>
]
<emphasis id="48AEE878FFC9D10D5912A646FE56602C" bold="true" box="[328,411,871,890]" italics="true" pageId="15" pageNumber="222">frontalis</emphasis>
</taxonomicName>
<emphasis id="48AEE878FFC9D10D59FAA646FDD4602C" bold="true" box="[416,537,871,890]" italics="true" pageId="15" pageNumber="222">flavidorsalis</emphasis>
Phelps &amp; Phelps, Jr., 1953: 140.
</paragraph>
</subSubSection>
<subSubSection id="32C067E1FFC9D10D589AA6A1FEB56093" pageId="15" pageNumber="222" type="materials_examined">
<paragraph id="7A65346AFFC9D10D589AA6A1FEB56093" blockId="15.[144,1165,226,990]" pageId="15" pageNumber="222">
<typeStatus id="A5618AC8FFC9D10D589AA6A1FED160C5" box="[192,284,896,915]" pageId="15" pageNumber="222" type="holotype">Holotype</typeStatus>
: Colección Ornitológica Phelps, Caracas (COP 55625, ‘on deposit’ at AMNH according to
<collectorName id="D72F51BCFFC9D10D589AA6B8FECD60FA" box="[192,256,921,940]" pageId="15" pageNumber="222">Phelps</collectorName>
&amp;
<collectorName id="D72F51BCFFC9D10D5940A6B8FE9760FA" box="[282,346,921,940]" pageId="15" pageNumber="222">Phelps</collectorName>
1953): Cerro Jurustaca,
<location id="7F0562B1FFC9D10D5A63A6BBFD1860FB" LSID="urn:lsid:plazi:treatment:F273857CFFC9D10D59B2A5A0FE526637:7F0562B1FFC9D10D5A63A6BBFD1860FB" box="[569,725,921,941]" country="Venezuela" municipality="Sierra de Perija" name="upper rio Negro" pageId="15" pageNumber="222" stateProvince="Zulia">upper río Negro</location>
,
<collectingMunicipality id="9A01AE10FFC9D10D5A84A6B8FCA360FA" box="[734,878,921,940]" pageId="15" pageNumber="222">
Sierra
<collectorName id="D72F51BCFFC9D10D5B40A6B8FCA360FA" box="[794,878,921,940]" pageId="15" pageNumber="222">de Perijá</collectorName>
</collectingMunicipality>
,
<collectingRegion id="B81EFA88FFC9D10D5B2CA6B8FC6760FA" box="[886,938,921,940]" country="Venezuela" name="Zulia" pageId="15" pageNumber="222">Zulia</collectingRegion>
,
<collectingCountry id="02CD74FAFFC9D10D5BE9A6B8FBD960FA" box="[947,1044,921,940]" name="Venezuela" pageId="15" pageNumber="222">Venezuela</collectingCountry>
(see
<collectorName id="D72F51BCFFC9D10D5C19A6BBFB4160FB" box="[1091,1164,922,941]" pageId="15" pageNumber="222">Paynter</collectorName>
&amp;
<collectorName id="D72F51BCFFC9D10D588CA693FEC26093" box="[214,271,946,965]" pageId="15" pageNumber="222">Storer</collectorName>
1970: 265).
</paragraph>
</subSubSection>
<subSubSection id="32C067E1FFC9D10D58CAA6EAFC62673F" pageId="15" pageNumber="222" type="reference_group">
<paragraph id="7A65346AFFC9D10D58CAA6EAFC8B6088" blockId="15.[144,1165,226,990]" box="[144,838,971,990]" pageId="15" pageNumber="222">
<taxonomicName id="BDDA4FE9FFC9D10D58CAA6EAFD676088" authority="Isler &amp; Isler 1987: 77" authorityName="Isler &amp; Isler" authorityPageNumber="77" authorityYear="1987" box="[144,682,971,990]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="subSpecies" species="frontalis" subSpecies="flavidorsalis">
<emphasis id="48AEE878FFC9D10D58CAA6EAFE2E6088" bold="true" box="[144,483,971,990]" italics="true" pageId="15" pageNumber="222">Hemispingus frontalis flavidorsalis</emphasis>
Isler &amp; Isler 1987: 77
</taxonomicName>
, Hilty 2011: 167.
</paragraph>
<paragraph id="7A65346AFFC9D10D589AA6D2FC62673F" blockId="15.[144,1165,1011,1378]" pageId="15" pageNumber="222">
SPECIMENS EXAMINED:
<emphasis id="48AEE878FFC9D10D5990A6D2FD896750" box="[458,580,1011,1030]" italics="true" pageId="15" pageNumber="222">S. f. frontalis</emphasis>
(
<specimenCount id="6CDCFFE3FFC9D10D5A0FA6D2FD796750" box="[597,692,1011,1030]" count="6" pageId="15" pageNumber="222" type="male">six males</specimenCount>
,
<specimenCount id="6CDCFFE3FFC9D10D5A98A6D2FCF46750" box="[706,825,1011,1030]" count="5" pageId="15" pageNumber="222" type="female">five females</specimenCount>
, one unsexed):
<collectingCountry id="02CD74FAFFC9D10D5B80A6D2FBF46750" box="[986,1081,1011,1030]" name="Colombia" pageId="15" pageNumber="222">Colombia</collectingCountry>
:
<collectingRegion id="B81EFA88FFC9D10D5C1CA6D2FB446750" box="[1094,1161,1011,1030]" country="Colombia" name="Caldas" pageId="15" pageNumber="222">Caldas</collectingRegion>
: Salento (female: ANSP 154474);
<collectingRegion id="B81EFA88FFC9D10D59E4A12DFE366749" box="[446,507,1036,1055]" country="Colombia" name="Cauca" pageId="15" pageNumber="222">Cauca</collectingRegion>
:
<collectorName id="D72F51BCFFC9D10D5A59A12DFD916749" box="[515,604,1036,1055]" pageId="15" pageNumber="222">El Tambo</collectorName>
(male: ANSP 141981); San Antonio (female: ANSP 141982);
<collectingRegion id="B81EFA88FFC9D10D58CAA105FF056761" box="[144,200,1060,1079]" country="Colombia" name="Huila" pageId="15" pageNumber="222">Huila</collectingRegion>
:
<collectorName id="D72F51BCFFC9D10D588BA104FEF06761" box="[209,317,1060,1080]" pageId="15" pageNumber="222">La Candela</collectorName>
(male: ANSP 155593, 155594; unsexed: 155591);
<collectorName id="D72F51BCFFC9D10D5B5DA104FC97676E" box="[775,858,1061,1080]" pageId="15" pageNumber="222">La Costa</collectorName>
(female: ANSP 141983);
<collectingRegion id="B81EFA88FFC9D10D5C1EA105FB446761" box="[1092,1161,1060,1079]" country="Colombia" name="Tolima" pageId="15" pageNumber="222">Tolima</collectingRegion>
: Toche (female: ANSP 154472).
<collectingCountry id="02CD74FAFFC9D10D59E8A11CFDC96706" box="[434,516,1085,1104]" name="Ecuador" pageId="15" pageNumber="222">Ecuador</collectingCountry>
:
<collectingRegion id="B81EFA88FFC9D10D5A56A11FFD8E6707" box="[524,579,1086,1105]" country="Ecuador" name="Napo" pageId="15" pageNumber="222">Napo</collectingRegion>
: Baeza (male: ANSP 83779, 83780).
<collectingCountry id="02CD74FAFFC9D10D5BCBA11FFC726707" box="[913,959,1086,1105]" name="Peru" pageId="15" pageNumber="222">Peru</collectingCountry>
:
<collectingRegion id="B81EFA88FFC9D10D5B92A11CFC366706" box="[968,1019,1085,1104]" country="Peru" name="Junin" pageId="15" pageNumber="222">Junín</collectingRegion>
(female: ANSP 91539).
<emphasis id="48AEE878FFC9D10D588DA177FEF4673F" box="[215,313,1110,1129]" italics="true" pageId="15" pageNumber="222">
S. f.
<taxonomicName id="BDDA4FE9FFC9D10D58A5A177FEF4673F" authorityName="Hellmayr &amp; von Seilern" authorityYear="1914" box="[255,313,1110,1129]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="hanieli">hanieli</taxonomicName>
</emphasis>
(
<specimenCount id="6CDCFFE3FFC9D10D591CA177FE6D673F" box="[326,416,1110,1129]" count="1" pageId="15" pageNumber="222" type="male">one male</specimenCount>
):
<collectingCountry id="02CD74FAFFC9D10D59F5A177FDDE673F" box="[431,531,1110,1129]" name="Venezuela" pageId="15" pageNumber="222">Venezuela</collectingCountry>
: ‘
<collectingMunicipality id="9A01AE10FFC9D10D5A45A177FDB7673F" box="[543,634,1110,1129]" pageId="15" pageNumber="222">Galeparo</collectingMunicipality>
,
<location id="7F0562B1FFC9D10D5ADEA177FCD3673F" LSID="urn:lsid:plazi:treatment:F273857CFFC9D10D59B2A5A0FE526637:7F0562B1FFC9D10D5ADEA177FCD3673F" box="[644,798,1110,1129]" country="Colombia" municipality="Galeparo" name="Curro del Avito'" pageId="15" pageNumber="222" stateProvince="Caldas">Curro del Avito’</location>
(ANSP 67191).
</paragraph>
</subSubSection>
<subSubSection id="32C067E1FFC9D10D589AA14EFE526637" pageId="15" pageNumber="222" type="discussion">
<paragraph id="7A65346AFFC9D10D589AA14EFE526637" blockId="15.[144,1165,1011,1378]" pageId="15" pageNumber="222">
COMMENTS: In addition to rectifying the identity of ZMUC 104925 (see introduction), which was a source of confusion for García-Moreno
<emphasis id="48AEE878FFC9D10D5A5DA1A9FDE167CD" box="[519,556,1160,1179]" italics="true" pageId="15" pageNumber="222">et al</emphasis>
. (2001), my study confirms that
<emphasis id="48AEE878FFC9D10D5B39A1A9FC1967CD" box="[867,980,1160,1179]" italics="true" pageId="15" pageNumber="222">
S. f.
<taxonomicName id="BDDA4FE9FFC9D10D5BD6A1A9FC1967CD" authorityName="von Berlepsch 1912:" authorityYear="1094" box="[908,980,1160,1179]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="frontalis">frontalis</taxonomicName>
</emphasis>
and
<emphasis id="48AEE878FFC9D10D5C5FA1A9FBA567CD" box="[1029,1128,1160,1179]" italics="true" pageId="15" pageNumber="222">
S. f.
<taxonomicName id="BDDA4FE9FFC9D10D5C74A1A9FBA567CD" authorityName="Hellmayr &amp; von Seilern" authorityYear="1914" box="[1070,1128,1160,1179]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="hanieli">hanieli</taxonomicName>
</emphasis>
are divergent and diagnosable, relative to other
<taxonomicName id="BDDA4FE9FFC9D10D5A63A181FD2967E5" box="[569,740,1184,1203]" pageId="15" pageNumber="222">
<emphasis id="48AEE878FFC9D10D5A63A181FD5467E5" box="[569,665,1184,1203]" italics="true" pageId="15" pageNumber="222">Sphenopsis</emphasis>
species
</taxonomicName>
, and from each other. The adult plumage of
<emphasis id="48AEE878FFC9D10D58CAA19BFF39679A" box="[144,244,1209,1229]" italics="true" pageId="15" pageNumber="222">
S. f.
<taxonomicName id="BDDA4FE9FFC9D10D58E0A198FF39679A" authorityName="Hellmayr &amp; von Seilern" authorityYear="1914" box="[186,244,1209,1228]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="hanieli">hanieli</taxonomicName>
</emphasis>
differs from
<emphasis id="48AEE878FFC9D10D5929A19BFE28679A" box="[371,485,1209,1229]" italics="true" pageId="15" pageNumber="222">
S. f.
<taxonomicName id="BDDA4FE9FFC9D10D59C7A198FE28679A" authorityName="von Berlepsch 1912:" authorityYear="1094" box="[413,485,1209,1228]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="frontalis">frontalis</taxonomicName>
</emphasis>
in colour, especially on the ventral surface, and has ‘a sharply defined superciliary stripe’ as emphasised in the original description (‘Ein scharf abgesetzter Superciliarstreif’, Hellmayr &amp; von Seilern 1914: 87). Given the surprising degree of homoplasy in plumage colour uncovered by phylogenomic analysis (Price-Waldman 2019), re-evaluating the taxonomic status of
<emphasis id="48AEE878FFC9D10D5B9CA025FBEA6641" box="[966,1063,1284,1303]" italics="true" pageId="15" pageNumber="222">
S. f.
<taxonomicName id="BDDA4FE9FFC9D10D5BB7A025FBEA6641" authorityName="Hellmayr &amp; von Seilern" authorityYear="1914" box="[1005,1063,1284,1303]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="hanieli">hanieli</taxonomicName>
</emphasis>
, and other
<taxonomicName id="BDDA4FE9FFC9D10D58CAA03CFF236679" authorityName="von Berlepsch 1912:" authorityYear="1094" box="[144,238,1308,1328]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="frontalis">
<emphasis id="48AEE878FFC9D10D58CAA03CFF236679" box="[144,238,1308,1328]" italics="true" pageId="15" pageNumber="222">S. frontalis</emphasis>
</taxonomicName>
subspecies not included in this study, should be prioritised by future researchers. Here, following tradition, I have treated
<emphasis id="48AEE878FFC9D10D5926A017FE2C661E" box="[380,481,1333,1353]" italics="true" pageId="15" pageNumber="222">
S. f.
<taxonomicName id="BDDA4FE9FFC9D10D59FDA014FE2C661E" authorityName="Hellmayr &amp; von Seilern" authorityYear="1914" box="[423,481,1333,1352]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="hanieli">hanieli</taxonomicName>
</emphasis>
as a subspecies of
<emphasis id="48AEE878FFC9D10D5AC1A017FCC3661E" box="[667,782,1333,1353]" italics="true" pageId="15" pageNumber="222">
S. f.
<taxonomicName id="BDDA4FE9FFC9D10D5A9CA014FCC3661E" authorityName="von Berlepsch 1912:" authorityYear="1094" box="[710,782,1333,1352]" class="Aves" family="Thraupidae" genus="Hemispingus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="222" phylum="Chordata" rank="species" species="frontalis">frontalis</taxonomicName>
</emphasis>
, although additional data may support classifying it at species rank.
</paragraph>
</subSubSection>
</treatment>
</document>
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