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<document id="DBA79A1CCB230A5CAA614F821145F88B" ID-DOI="10.1080/00222930500089271" ID-ISSN="1464-5262" ID-Zenodo-Dep="4581592" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614874687196" checkinUser="carolina" docAuthor="Askew, R. R. &amp; Nieves-Aldrey, J. L." docDate="2005" docId="3C4A87B1FFB0935DFDE8FF62FBE6D8C0" docLanguage="en" docName="JNATHIST.39.25.2331-2338.pdf" docOrigin="Journal of Natural History 39 (25)" docSource="http://dx.doi.org/10.1080/00222930500089271" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Ricasia Askew &amp; Nieves-Aldrey 2005, gen. nov." docType="treatment" docVersion="3" lastPageNumber="2332" masterDocId="C073FFC9FFB3935EFFF5FFAEFFB5DE4E" masterDocTitle="A new genus and species of Pteromalidae (Hymenoptera, Chalcidoidea) from Spain, parasitic in cynipid galls on Fumaria" masterLastPageNumber="2338" masterPageNumber="2331" pageNumber="2332" updateTime="1698880109161" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="B51EC6B68B61A71B7EC74EC0FCD0B888">A new genus and species of Pteromalidae (Hymenoptera, Chalcidoidea) from Spain, parasitic in cynipid galls on Fumaria</mods:title>
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<mods:name id="D2CA6857661295869CFE4BF67167CA84" type="personal">
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<mods:namePart id="952D17F5D38C18C58A062F0A54B54714">Askew, R. R.</mods:namePart>
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<mods:namePart id="27AB45BBAF9F5E7D1F5B7CF8BA9B79F6">Nieves-Aldrey, J. L.</mods:namePart>
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<mods:typeOfResource id="DC3D184B46AD6F2C2FFF633305E98549">text</mods:typeOfResource>
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<mods:title id="D0008A3520BC4EFDAF70740009C9BEB8">Journal of Natural History</mods:title>
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<mods:part id="58C06EE9A9F141C49718E5DD2214D398">
<mods:date id="B3BBF18CD38A2D159DDBEA91E631219C">2005</mods:date>
<mods:detail id="81FF0E957945145DE2BD18C517EC3283" type="pubDate">
<mods:number id="53ACFF58C3C0D3AE737659F43F06C211">2005-06-29</mods:number>
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<mods:detail id="41CE2DBBBE7227303D3005E9AD21254D" type="volume">
<mods:number id="11E3C95425B1790402E9B8687669D615">39</mods:number>
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<mods:detail id="7DD96209CD8E8E5E96EF432FB38BB6F1" type="issue">
<mods:number id="0A90AA6383A0B4FB181580A0B6996F57">25</mods:number>
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<mods:start id="A67C77AD84334D6697C5A474B525B1FB">2331</mods:start>
<mods:end id="282C65D4F72CEF1A0AF5E90411FDDF5D">2338</mods:end>
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<mods:classification id="14F1D9078882E873B329913BCC0978B5">journal article</mods:classification>
<mods:identifier id="73FC2EA613C375B58A2CFB19F43BC350" type="DOI">10.1080/00222930500089271</mods:identifier>
<mods:identifier id="6A1C73A2AD604B0FA371EC07522091BF" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="D81A5E5D4BC5B95D920FE84BE03FE1B6" type="Zenodo-Dep">4581592</mods:identifier>
</mods:mods>
<treatment id="3C4A87B1FFB0935DFDE8FF62FBE6D8C0" LSID="urn:lsid:plazi:treatment:3C4A87B1FFB0935DFDE8FF62FBE6D8C0" httpUri="http://treatment.plazi.org/id/3C4A87B1FFB0935DFDE8FF62FBE6D8C0" lastPageNumber="2332" pageId="3" pageNumber="2332">
<subSubSection id="FCF9652CFFB0935DFDE8FF62FD40DEAA" box="[541,757,204,228]" pageId="3" pageNumber="2332" type="nomenclature">
<paragraph id="B45C36A7FFB0935DFDE8FF62FD40DEAA" blockId="3.[526,770,137,228]" box="[541,757,204,228]" pageId="3" pageNumber="2332">
<heading id="EF1481CBFFB0935DFDE8FF62FD40DEAA" box="[541,757,204,228]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="2332" reason="2">
<taxonomicName id="73E34D24FFB0935DFDE8FF62FDCADEAA" authority="Askew &amp; Nieves-Aldrey, 2005" authorityName="Askew &amp; Nieves-Aldrey" authorityYear="2005" box="[541,639,204,228]" class="Insecta" family="Pteromalidae" genus="Ricasia" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="2332" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="8697EAB5FFB0935DFDE8FF62FDCADEAA" bold="true" box="[541,639,204,228]" italics="true" pageId="3" pageNumber="2332">Ricasia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9DA457CEFFB0935DFD7DFF62FD40DEAA" box="[648,757,204,228]" pageId="3" pageNumber="2332" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FCF9652CFFB0935DFF88FEB2FBE6D8C0" pageId="3" pageNumber="2332" type="description">
<paragraph id="B45C36A7FFB0935DFF88FEB2FF40DF7A" blockId="3.[125,245,284,308]" box="[125,245,284,308]" pageId="3" pageNumber="2332">
<emphasis id="8697EAB5FFB0935DFF88FEB2FF40DF7A" box="[125,245,284,308]" italics="true" pageId="3" pageNumber="2332">Description</emphasis>
</paragraph>
<paragraph id="B45C36A7FFB0935DFF88FEE3FED1DC06" blockId="3.[125,1173,333,910]" pageId="3" pageNumber="2332">
Head and thorax with very fine, engraved reticulate sculpture and numerous quite conspicuous hair-pits. Head with occiput not margined; foramen magnum relatively high (
<figureCitation id="2CD82A22FFB0935DFF70FE20FF4BDFE8" box="[133,254,398,422]" captionStart="Figure 1" captionStartId="4.[138,198,1416,1435]" captionTargetBox="[183,1140,133,1380]" captionTargetId="figure-33@4.[183,1140,133,1380]" captionTargetPageId="4" captionText="Figure 1. Riυasia fumariae sp. n., adult female. (A) Antenna; (B) detail of antennal clava; (C) head in anterior view; (D) head in posterior view; (E) mesosoma in dorsal view; (F) mesosoma in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4581594" httpUri="https://zenodo.org/record/4581594/files/figure.png" pageId="3" pageNumber="2332">Figure 1D</figureCitation>
); head in dorsal view with frons flat, not produced between eyes and scrobal depression not visible; antenna inserted distinctly below middle of face; clypeus marked off by a groove (
<figureCitation id="2CD82A22FFB0935DFEE5FE61FE3DDFA9" box="[272,392,463,487]" captionStart="Figure 1" captionStartId="4.[138,198,1416,1435]" captionTargetBox="[183,1140,133,1380]" captionTargetId="figure-33@4.[183,1140,133,1380]" captionTargetPageId="4" captionText="Figure 1. Riυasia fumariae sp. n., adult female. (A) Antenna; (B) detail of antennal clava; (C) head in anterior view; (D) head in posterior view; (E) mesosoma in dorsal view; (F) mesosoma in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4581594" httpUri="https://zenodo.org/record/4581594/files/figure.png" pageId="3" pageNumber="2332">Figure 1C</figureCitation>
), its anterior margin straight; anterior tentorial pits distinct; labrum in about same plane as clypeus. Mandibles (
<figureCitation id="2CD82A22FFB0935DFD60FE41FCBEDC49" box="[661,779,495,519]" captionStart="Figure 1" captionStartId="4.[138,198,1416,1435]" captionTargetBox="[183,1140,133,1380]" captionTargetId="figure-33@4.[183,1140,133,1380]" captionTargetPageId="4" captionText="Figure 1. Riυasia fumariae sp. n., adult female. (A) Antenna; (B) detail of antennal clava; (C) head in anterior view; (D) head in posterior view; (E) mesosoma in dorsal view; (F) mesosoma in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4581594" httpUri="https://zenodo.org/record/4581594/files/figure.png" pageId="3" pageNumber="2332">Figure 1C</figureCitation>
) bidentate. Antenna (
<figureCitation id="2CD82A22FFB0935DFBE3FE41FB3EDC49" box="[1046,1163,495,519]" captionStart="Figure 1" captionStartId="4.[138,198,1416,1435]" captionTargetBox="[183,1140,133,1380]" captionTargetId="figure-33@4.[183,1140,133,1380]" captionTargetPageId="4" captionText="Figure 1. Riυasia fumariae sp. n., adult female. (A) Antenna; (B) detail of antennal clava; (C) head in anterior view; (D) head in posterior view; (E) mesosoma in dorsal view; (F) mesosoma in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4581594" httpUri="https://zenodo.org/record/4581594/files/figure.png" pageId="3" pageNumber="2332">Figure 1A</figureCitation>
) similar in both sexes, 12-segmented, formula 11163 (verified by both light microscope and SEM examination).
</paragraph>
<paragraph id="B45C36A7FFB0935DFF6DFDFFFBCBDCA5" blockId="3.[125,1173,333,910]" pageId="3" pageNumber="2332">
Pronotum long (
<figureCitation id="2CD82A22FFB0935DFE97FDFFFE62DC27" box="[354,471,593,617]" captionStart="Figure 1" captionStartId="4.[138,198,1416,1435]" captionTargetBox="[183,1140,133,1380]" captionTargetId="figure-33@4.[183,1140,133,1380]" captionTargetPageId="4" captionText="Figure 1. Riυasia fumariae sp. n., adult female. (A) Antenna; (B) detail of antennal clava; (C) head in anterior view; (D) head in posterior view; (E) mesosoma in dorsal view; (F) mesosoma in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4581594" httpUri="https://zenodo.org/record/4581594/files/figure.png" pageId="3" pageNumber="2332">Figure 1E</figureCitation>
), about as long as mesoscutum, without an offset collar. Mesoscutum with complete, deep notauli (
<figureCitation id="2CD82A22FFB0935DFD87FDDFFD5CDCC7" box="[626,745,625,649]" captionStart="Figure 1" captionStartId="4.[138,198,1416,1435]" captionTargetBox="[183,1140,133,1380]" captionTargetId="figure-33@4.[183,1140,133,1380]" captionTargetPageId="4" captionText="Figure 1. Riυasia fumariae sp. n., adult female. (A) Antenna; (B) detail of antennal clava; (C) head in anterior view; (D) head in posterior view; (E) mesosoma in dorsal view; (F) mesosoma in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4581594" httpUri="https://zenodo.org/record/4581594/files/figure.png" pageId="3" pageNumber="2332">Figure 1E</figureCitation>
). Scutellum with frenal area marked off by a shallow groove. Metapleuron not reaching hind wing cavity (
<figureCitation id="2CD82A22FFB0935DFBFAFD3CFB30DCE4" box="[1039,1157,658,682]" captionStart="Figure 1" captionStartId="4.[138,198,1416,1435]" captionTargetBox="[183,1140,133,1380]" captionTargetId="figure-33@4.[183,1140,133,1380]" captionTargetPageId="4" captionText="Figure 1. Riυasia fumariae sp. n., adult female. (A) Antenna; (B) detail of antennal clava; (C) head in anterior view; (D) head in posterior view; (E) mesosoma in dorsal view; (F) mesosoma in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4581594" httpUri="https://zenodo.org/record/4581594/files/figure.png" pageId="3" pageNumber="2332">Figure 1F</figureCitation>
). Propodeum (
<figureCitation id="2CD82A22FFB0935DFEE8FD1CFE24DC84" box="[285,401,690,714]" captionStart="Figure 1" captionStartId="4.[138,198,1416,1435]" captionTargetBox="[183,1140,133,1380]" captionTargetId="figure-33@4.[183,1140,133,1380]" captionTargetPageId="4" captionText="Figure 1. Riυasia fumariae sp. n., adult female. (A) Antenna; (B) detail of antennal clava; (C) head in anterior view; (D) head in posterior view; (E) mesosoma in dorsal view; (F) mesosoma in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4581594" httpUri="https://zenodo.org/record/4581594/files/figure.png" pageId="3" pageNumber="2332">Figure 1E</figureCitation>
) medially shorter than half the length of scutellum; spiracles distant from anterior border. Metacoxa dorsally pilose; metatibia with two apical spurs.
</paragraph>
<paragraph id="B45C36A7FFB0935DFF6DFD5DFCC2DD62" blockId="3.[125,1173,333,910]" pageId="3" pageNumber="2332">
Fore wing (
<figureCitation id="2CD82A22FFB0935DFEEAFD5DFE23DD45" box="[287,406,755,779]" captionStart="Figure 2" captionStartId="5.[125,185,1408,1427]" captionTargetBox="[170,1127,133,1372]" captionTargetId="figure-69@5.[170,1127,133,1372]" captionTargetPageId="5" captionText="Figure 2. Riυasia fumariae sp. n. (A) Adult metasoma in ventral view; (B) mesosoma in posterodorsal view; (C) fore wing; (D) fully grown larva in ventral view; (E) fully grown larva, head in anterior view; (F) fully grown larva, oral region." figureDoi="http://doi.org/10.5281/zenodo.4581596" httpUri="https://zenodo.org/record/4581596/files/figure.png" pageId="3" pageNumber="2332">Figure 2C</figureCitation>
) with postmarginal vein much longer than marginal vein; stigmal vein only a little shorter than marginal vein; stigma not enlarged.
</paragraph>
<paragraph id="B45C36A7FFB0935DFF6DFC9BFEB9DDC0" blockId="3.[125,1173,333,910]" pageId="3" pageNumber="2332">
Petiole of gaster short and transverse, smooth; gaster with posterior margins of tergites almost straight; hypopygium of female short, its tip at about one-third of gaster length (
<figureCitation id="2CD82A22FFB0935DFF70FCD8FF4ADDC0" box="[133,255,886,910]" captionStart="Figure 2" captionStartId="5.[125,185,1408,1427]" captionTargetBox="[170,1127,133,1372]" captionTargetId="figure-69@5.[170,1127,133,1372]" captionTargetPageId="5" captionText="Figure 2. Riυasia fumariae sp. n. (A) Adult metasoma in ventral view; (B) mesosoma in posterodorsal view; (C) fore wing; (D) fully grown larva in ventral view; (E) fully grown larva, head in anterior view; (F) fully grown larva, oral region." figureDoi="http://doi.org/10.5281/zenodo.4581596" httpUri="https://zenodo.org/record/4581596/files/figure.png" pageId="3" pageNumber="2332">Figure 2A</figureCitation>
).
</paragraph>
<paragraph id="B45C36A7FFB0935DFF88FC52FEB4DA5A" blockId="3.[125,257,1020,1044]" box="[125,257,1020,1044]" pageId="3" pageNumber="2332">
<emphasis id="8697EAB5FFB0935DFF88FC52FEB4DA5A" box="[125,257,1020,1044]" italics="true" pageId="3" pageNumber="2332">
<typeStatus id="6B588805FFB0935DFF88FC52FF06DA5D" box="[125,179,1020,1043]" pageId="3" pageNumber="2332">Type</typeStatus>
species
</emphasis>
</paragraph>
<paragraph id="B45C36A7FFB0935DFF88FB82FE5EDA2B" blockId="3.[125,1172,1068,1125]" pageId="3" pageNumber="2332">
<emphasis id="8697EAB5FFB0935DFF88FB82FE82DA0A" box="[125,311,1068,1092]" italics="true" pageId="3" pageNumber="2332">
Riυasia
<taxonomicName id="73E34D24FFB0935DFF22FB82FE82DA0A" authority="Askew &amp; Nieves-Aldrey, 2005" authorityName="Askew &amp; Nieves-Aldrey" authorityYear="2005" box="[215,311,1068,1092]" class="Insecta" family="Pteromalidae" genus="Ricasia" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="2332" phylum="Arthropoda" rank="species" species="fumariae" status="sp. nov.">fumariae</taxonomicName>
</emphasis>
<taxonomicNameLabel id="9DA457CEFFB0935DFECBFB82FEC9DA0A" box="[318,380,1068,1092]" pageId="3" pageNumber="2332" rank="species">sp. n.</taxonomicNameLabel>
The genus is named after the locality where host galls were collected, the species after the host plant.
</paragraph>
<paragraph id="B45C36A7FFB0935DFF88FB4EFF50DAB6" blockId="3.[125,229,1248,1272]" box="[125,229,1248,1272]" pageId="3" pageNumber="2332">
<emphasis id="8697EAB5FFB0935DFF88FB4EFF50DAB6" box="[125,229,1248,1272]" italics="true" pageId="3" pageNumber="2332">Comment</emphasis>
</paragraph>
<paragraph id="B45C36A7FFB0935DFF88FABFFBE6D8C0" blockId="3.[125,1173,1297,1678]" pageId="3" pageNumber="2332">
The correct placement of
<emphasis id="8697EAB5FFB0935DFE32FABFFDAFDB67" box="[455,538,1297,1321]" italics="true" pageId="3" pageNumber="2332">Riυasia</emphasis>
within
<taxonomicName id="73E34D24FFB0935DFD74FABFFCAFDB67" authorityName="Dalman" authorityYear="1820" box="[641,794,1297,1321]" class="Insecta" family="Pteromalidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="2332" phylum="Arthropoda" rank="family">Pteromalidae</taxonomicName>
proved difficult to determine. Applying
<bibRefCitation id="D0724B56FFB0935DFF1AFA9FFE0CDB07" author="Graham MWR de V." box="[239,441,1329,1353]" pageId="3" pageNumber="2332" pagination="1 - 908" refId="ref2772" refString="Graham MWR de V. 1969. The Pteromalidae of north-western Europe (Hymenoptera: Chalcidoidea). Bulletin of the British Museum (Natural History) Entomology (Supplement) 16: 1 - 908." type="journal article" year="1969">Graham’s (1969)</bibRefCitation>
keys it runs to
<taxonomicName id="73E34D24FFB0935DFD9BFA9FFC98DB07" authorityName="Walker" authorityYear="1833" box="[622,813,1329,1353]" class="Insecta" family="Pteromalidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="2332" phylum="Arthropoda" rank="subFamily" subFamily="Miscogasterinae">Miscogasterinae</taxonomicName>
, but in the key to tribes of this subfamily, progress beyond the second couplet is baulked by the combination of a relatively long postmarginal vein and bidentate mandibles, on the one hand (as in Micradelini), and metallic coloration and small but developed fore wing speculum on the other. In Micradelini there is no metallic coloration, notably dense pilosity on the eyes and dorsal surface of the thorax, clavate antennae with two or three anelli and transverse funicle segments. Relationship to Micradelini seems distant and
<emphasis id="8697EAB5FFB0935DFCE3FA5AFCDCD842" box="[790,873,1524,1548]" italics="true" pageId="3" pageNumber="2332">Riυasia</emphasis>
has more the appearance of Ormocerini (upgraded to
<taxonomicName id="73E34D24FFB0935DFE1CF9BAFD34D862" authorityName="Walker" authorityYear="1833" box="[489,641,1556,1580]" class="Insecta" family="Pteromalidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="2332" phylum="Arthropoda" rank="subFamily" subFamily="Ormocerinae">Ormocerinae</taxonomicName>
by
<bibRefCitation id="D0724B56FFB0935DFD49F9BAFCD6D862" author="Boucek Z" box="[700,867,1556,1580]" pageId="3" pageNumber="2332" refId="ref2725" refString="Boucek Z. 1988. Australasian Chalcidoidea (Hymenoptera): a biosystematic revision of fourteen families, with a reclassification of species Wallingford: CAB International. 832 p." type="book" year="1988">Bouček 1988</bibRefCitation>
). However, none of the European genera included in this taxon by
<bibRefCitation id="D0724B56FFB0935DFD99F99BFC90D803" author="Graham MWR de V." box="[620,805,1589,1613]" pageId="3" pageNumber="2332" pagination="1 - 908" refId="ref2772" refString="Graham MWR de V. 1969. The Pteromalidae of north-western Europe (Hymenoptera: Chalcidoidea). Bulletin of the British Museum (Natural History) Entomology (Supplement) 16: 1 - 908." type="journal article" year="1969">Graham (1969)</bibRefCitation>
, nor any of the genera placed in
<taxonomicName id="73E34D24FFB0935DFF88F9FBFEA0D823" authorityName="Walker" authorityYear="1833" box="[125,277,1621,1645]" class="Insecta" family="Pteromalidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="2332" phylum="Arthropoda" rank="subFamily" subFamily="Ormocerinae">Ormocerinae</taxonomicName>
by
<bibRefCitation id="D0724B56FFB0935DFEB4F9FBFE44D823" author="Boucek Z" box="[321,497,1621,1645]" pageId="3" pageNumber="2332" refId="ref2725" refString="Boucek Z. 1988. Australasian Chalcidoidea (Hymenoptera): a biosystematic revision of fourteen families, with a reclassification of species Wallingford: CAB International. 832 p." type="book" year="1988">Bouček (1988)</bibRefCitation>
, have an antenna with just a single anellus followed by six funicle segments and a three-segmented clava, nor a large, campaniform pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>
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