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<document id="27765B10353171384E6BBE7CFC4FDD64" ID-DOI="10.11646/zootaxa.4038.1.1" ID-GBIF-Dataset="066e84f1-206b-40cb-a615-2126696b813d" ID-ISSN="1175-5326" ID-Zenodo-Dep="289876" ID-ZooBank="1B62B78C-AA59-4417-A4FC-1CC9CED745E0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461316934850" checkinUser="plazi" docAuthor="Janák, Jiří &amp; Bordoni, Arnaldo" docDate="2015" docId="039D87F55579FF84FF4AF8AAFCE6BCCA" docLanguage="en" docName="zt04038p094.pdf" docOrigin="Zootaxa 4038 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Thyreocephalus guineensis Bernhauer 1912, comb. nov." docType="treatment" docVersion="7" lastPageNumber="70" masterDocId="FFA4FF8D553BFFC1FFDDFFA6FFFFBE25" masterDocTitle="Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="94" masterPageNumber="1" pageNumber="67" updateTime="1698618126966" updateUser="plazi">
<mods:mods id="1EF4FA1BB801F5FBE53E9A31B2F0E162" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="65C9318B59D17B6EE338145E299D1887">
<mods:title id="BA60F1AB28EC6608583DEEC207C904CD">Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
</mods:titleInfo>
<mods:name id="F0346DA810B5666EA4261DE9C4654AE9" type="personal">
<mods:role id="56C091792D731842AA636386AB19C7CC">
<mods:roleTerm id="76E047616F16B1B275F2B05014997F70">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EC62726A4196AD01841AE6E2FA706F63">Janák, Jiří</mods:namePart>
</mods:name>
<mods:name id="6CA00D734EF0898CAB55875CFFDD877F" type="personal">
<mods:role id="7E5670CC33FD3DDCFB4B41D1E29E0BC5">
<mods:roleTerm id="02215A5F2E90B8DB01D76BB307B96A4E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6E3DF6E960AADC41986BDAD881B120B5">Bordoni, Arnaldo</mods:namePart>
</mods:name>
<mods:typeOfResource id="8B90D4D241AAAEBFBC56B6BF7EA5A48C">text</mods:typeOfResource>
<mods:relatedItem id="95025B853BBE208DFBBD6728864C62A4" type="host">
<mods:titleInfo id="D5AF1C78410EA155B045E53F50E0D144">
<mods:title id="458B15395954F2AB97CB5AD74949419C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2F61C427C5E9167B5A9B2978FE2A9505">
<mods:date id="9E63C3B95F6FF535360D2949DEB53AE8">2015</mods:date>
<mods:detail id="2CA97A763D9F5B8D51762E0CA141E450" type="volume">
<mods:number id="CF4F4FFB8EF83C193ACAA6B99B212A86">4038</mods:number>
</mods:detail>
<mods:detail id="681DC1B8C334B530A7747A72223BFE45" type="issue">
<mods:number id="37DDA440D0D937C4DD281BDECA431554">1</mods:number>
</mods:detail>
<mods:extent id="C7CFD8556B4B502015C22A17F0B6B49C" unit="page">
<mods:start id="83E4BD067424FDCEC875DCC693EC250B">1</mods:start>
<mods:end id="0AA6B291CFC13BFB01D8E71DE2D8D9D9">94</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2AFC13DA0A51582CDB9D73BE4C57CEF9">journal article</mods:classification>
<mods:identifier id="A33026AAE7CBF917B3F059C21621EECF" type="DOI">10.11646/zootaxa.4038.1.1</mods:identifier>
<mods:identifier id="F7E4A50B0E9243E7FBDDB173ACC05E7D" type="GBIF-Dataset">066e84f1-206b-40cb-a615-2126696b813d</mods:identifier>
<mods:identifier id="24E1AC6779BA6257532F7D6CEF166858" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7CE2114D4534C8522FA5332CB15F5C4E" type="Zenodo-Dep">289876</mods:identifier>
<mods:identifier id="C8D05101B345FC0D7C7E8C5694BE2661" type="ZooBank">1B62B78C-AA59-4417-A4FC-1CC9CED745E0</mods:identifier>
</mods:mods>
<treatment id="039D87F55579FF84FF4AF8AAFCE6BCCA" ID-DOI="http://doi.org/10.5281/zenodo.6101687" ID-GBIF-Taxon="127693925" ID-Zenodo-Dep="6101687" LSID="urn:lsid:plazi:treatment:039D87F55579FF84FF4AF8AAFCE6BCCA" httpUri="http://treatment.plazi.org/id/039D87F55579FF84FF4AF8AAFCE6BCCA" lastPageId="69" lastPageNumber="70" pageId="66" pageNumber="67">
<subSubSection id="C32E65685579FF83FF4AF8AAFE45B962" pageId="66" pageNumber="67" type="nomenclature">
<paragraph id="8B8B36E35579FF83FF4AF8AAFCA8B903" blockId="66.[151,855,1804,1863]" box="[151,855,1804,1830]" pageId="66" pageNumber="67">
<heading id="D0C3818F5579FF83FF4AF8AAFCA8B903" bold="true" box="[151,855,1804,1830]" fontSize="11" level="1" pageId="66" pageNumber="67" reason="1">
<emphasis id="B940EAF15579FF83FF4AF8AAFCA8B903" bold="true" box="[151,855,1804,1830]" pageId="66" pageNumber="67">
<taxonomicName id="4C344D605579FF83FF4AF8AAFD36B903" ID-CoL="56RW7" authority="Bernhauer, 1912" authorityName="Bernhauer" authorityYear="1912" box="[151,713,1804,1830]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="guineensis" status="comb. nov.">
<emphasis id="B940EAF15579FF83FF4AF8AAFE1EB903" bold="true" box="[151,481,1804,1830]" italics="true" pageId="66" pageNumber="67">Thyreocephalus guineensis</emphasis>
(Bernhauer, 1912)
</taxonomicName>
<taxonomicNameLabel id="A273578A5579FF83FD0DF8AAFCA8B903" box="[720,855,1804,1830]" pageId="66" pageNumber="67">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8B36E35579FF83FF4AF888FE45B962" blockId="66.[151,855,1804,1863]" box="[151,442,1838,1863]" pageId="66" pageNumber="67">(Figs. 166, 167, 172–177)</paragraph>
</subSubSection>
<subSubSection id="C32E65685579FF83FF4AF8D1FCD1B9A8" box="[151,814,1911,1933]" pageId="66" pageNumber="67" type="reference_group">
<paragraph id="8B8B36E35579FF83FF4AF8D1FCD1B9A8" blockId="66.[151,814,1911,1933]" box="[151,814,1911,1933]" pageId="66" pageNumber="67">
<treatmentCitationGroup id="AB2411CD5579FF83FF4AF8D1FCD1B9A8" box="[151,814,1911,1933]" pageId="66" pageNumber="67">
<taxonomicName id="4C344D605579FF83FF4AF8D1FDB3B9A8" ID-CoL="3CBNB" authority="Bernhauer, 1912a: 183" authorityName="Bernhauer" authorityPageNumber="183" authorityYear="1912" box="[151,588,1911,1933]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="guineensis">
<emphasis id="B940EAF15579FF83FF4AF8D1FEA2B9A8" box="[151,349,1911,1933]" italics="true" pageId="66" pageNumber="67">Eulissus guineensis</emphasis>
<treatmentCitation id="0A9510F25579FF83FEBEF8D1FDB3B9A8" author="Bernhauer" box="[355,588,1911,1933]" page="183" pageId="66" pageNumber="67" year="1912">
<bibRefCitation id="EFA54B125579FF83FEBEF8D1FDE5B9A8" author="Bernhauer" box="[355,538,1911,1933]" pageId="66" pageNumber="67" refString="Bernhauer, M. (1912 a) Beitrag zur Staphylinidenfauna von Afrika. Entomologische Mitteilungen, 1, 177 - 183, 203 - 209." type="journal article" year="1912" yearSuffix="a">Bernhauer, 1912a</bibRefCitation>
: 183
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0A9510F25579FF83FD8BF8DEFCD5B9A8" author="Herman" box="[598,810,1911,1933]" page="3627" pageId="66" pageNumber="67" year="2001">
<bibRefCitation id="EFA54B125579FF83FD8BF8DEFD15B9A8" author="Herman" box="[598,746,1911,1933]" pageId="66" pageNumber="67" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3). Bulletin of the American Museum of Natural History, 265, 3021 - 3840." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 3627
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32E65685579FF82FF4AF81FFD24BFD1" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67" type="materials_examined">
<paragraph id="8B8B36E35579FF83FF4AF81FFD4FB9F7" blockId="66.[151,1436,1977,2038]" box="[151,688,1977,2002]" pageId="66" pageNumber="67">
<emphasis id="B940EAF15579FF83FF4AF81FFECAB9F7" bold="true" box="[151,309,1977,2002]" pageId="66" pageNumber="67">
<typeStatus id="548F88415579FF83FF4AF81FFF2DB9F7" box="[151,210,1977,2002]" pageId="66" pageNumber="67">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32376735579FF83FEE1F81FFE72B9F7" box="[316,397,1977,2002]" name="Guinea-Bissau" pageId="66" pageNumber="67">Guinea</collectingCountry>
(without additional data).
</paragraph>
<paragraph id="8B8B36E35579FF82FF1AF87BFF03BEF0" blockId="66.[151,1436,1977,2038]" lastBlockId="67.[151,1436,151,501]" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67">
<emphasis id="B940EAF15579FF83FF1AF87BFE0CB9D3" bold="true" box="[199,499,2013,2038]" pageId="66" pageNumber="67">
<typeStatus id="548F88415579FF83FF1AF87BFEFDB9D3" box="[199,258,2013,2038]" pageId="66" pageNumber="67">Type</typeStatus>
material.
<typeStatus id="548F88415579FF83FE4EF87BFE0CB9D3" box="[403,499,2013,2038]" pageId="66" pageNumber="67" type="syntype">Syntype</typeStatus>
</emphasis>
(sex indet.):
<collectingCountry id="F32376735579FF83FD77F87BFCE5B9D3" box="[682,794,2013,2038]" name="Guinea-Bissau" pageId="66" pageNumber="67">
<emphasis id="B940EAF15579FF83FD77F87BFCE5B9D3" bold="true" box="[682,794,2013,2038]" pageId="66" pageNumber="67">GUINEA</emphasis>
</collectingCountry>
: “
<collectingCountry id="F32376735579FF83FCE2F87BFC68B9D3" box="[831,919,2013,2038]" name="Guinea-Bissau" pageId="66" pageNumber="67">Guinea</collectingCountry>
”, “
<typeStatus id="548F88415579FF83FC1EF878FBCEB9D3" box="[963,1073,2014,2038]" pageId="66" pageNumber="67" type="syntype">Syntypus</typeStatus>
”, “
<taxonomicName id="4C344D605579FF83FB82F878FAD0B9D3" authority="Brnh." authorityName="Brnh." box="[1119,1327,2013,2038]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="guineensis" status="sp. nov.">
<emphasis id="B940EAF15579FF83FB82F878FB2AB9D0" box="[1119,1237,2014,2037]" italics="true" pageId="66" pageNumber="67">guineensis</emphasis>
, Brnh.
</taxonomicName>
<taxonomicNameLabel id="A273578A5579FF83FA9CF878FA75B9D3" box="[1345,1418,2014,2038]" pageId="66" pageNumber="67" rank="species">n. sp.</taxonomicNameLabel>
”, “
<taxonomicName id="4C344D605578FF82FF7EFF31FD8EBE95" authority="Bernhauer" authorityName="Bernhauer" box="[163,625,151,176]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="guineensis">
<emphasis id="B940EAF15578FF82FF7EFF31FE26BE95" box="[163,473,151,176]" italics="true" pageId="67" pageNumber="68">Thyreocephalus guineensis</emphasis>
(Bernhauer)
</taxonomicName>
, J. Janák det. 2006” (
<collectionCode id="ED25AE265578FF82FC5FFF3EFC49BE95" LSID="urn:lsid:biocol.org:col:33211" box="[898,950,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="67" pageNumber="68">DEI</collectionCode>
). The specimen is devoid of the genital segment.
</paragraph>
<paragraph id="8B8B36E35578FF82FF1AFF79FD24BFD1" blockId="67.[151,1436,151,501]" pageId="67" pageNumber="68">
<emphasis id="B940EAF15578FF82FF1AFF79FCECBEDD" bold="true" box="[199,787,223,248]" pageId="67" pageNumber="68">
Additional material examined.
<collectingCountry id="F32376735578FF82FDB7FF79FCECBEDD" box="[618,787,223,248]" name="Cameroon" pageId="67" pageNumber="68">CAMEROON</collectingCountry>
</emphasis>
: 1 spec.: Joko (
<collectionCode id="ED25AE265578FF82FBDEFF46FBA6BEDD" box="[1027,1113,224,248]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="67" pageNumber="68">MFNB</collectionCode>
).
<collectingCountry id="F32376735578FF82FBA6FF79FED7BF38" name="Central African Republic" pageId="67" pageNumber="68">
<emphasis id="B940EAF15578FF82FBA6FF79FED7BF38" bold="true" pageId="67" pageNumber="68">CENTRAL AFRICAN REPUBLIC</emphasis>
</collectingCountry>
: 4 spec.: Uamgebiet, Bosum,
<date id="FF8A10235578FF82FDA5FEA2FCF7BF38" box="[632,776,260,285]" pageId="67" pageNumber="68" value="1914-04-01" valueMax="1914-04-10" valueMin="1914-04-01">1-10.iv.1914</date>
, S. Tessmann (
<collectionCode id="ED25AE265578FF82FC72FEA2FBFCBF39" box="[943,1027,260,284]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="67" pageNumber="68">MFNB</collectionCode>
,
<collectionCode id="ED25AE265578FF82FBD3FEA2FBB1BF39" box="[1038,1102,260,284]" pageId="67" pageNumber="68">JJRC</collectionCode>
); 3 spec.: Uamgebiet, Bosum,
<date id="FF8A10235578FF82FF4AFE81FEC9BF65" box="[151,310,295,320]" pageId="67" pageNumber="68" value="1914-04-21" valueMax="1914-04-30" valueMin="1914-04-21">21–30.iv.1914</date>
S. Tessmann (
<collectionCode id="ED25AE265578FF82FE3CFE8EFDCABF65" box="[481,565,296,320]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="67" pageNumber="68">MFNB</collectionCode>
,
<collectionCode id="ED25AE265578FF82FD9FFE8EFD7ABF65" box="[578,645,296,320]" pageId="67" pageNumber="68">ABFI</collectionCode>
,
<collectionCode id="ED25AE265578FF82FD4FFE8EFD2DBF65" box="[658,722,296,320]" pageId="67" pageNumber="68">JJRC</collectionCode>
).
<collectingCountry id="F32376735578FF82FD3AFE81FB17BF65" box="[743,1256,295,320]" name="Democratic Republic of the Congo" pageId="67" pageNumber="68">
<emphasis id="B940EAF15578FF82FD3AFE81FB17BF65" bold="true" box="[743,1256,295,320]" pageId="67" pageNumber="68">DEMOCRATIC REPUBLIC OF CONGO</emphasis>
</collectingCountry>
: 1 spec.:
<collectingCountry id="F32376735578FF82FA87FE81FA67BF65" box="[1370,1432,295,320]" name="Democratic Republic of the Congo" pageId="67" pageNumber="68">Zaire</collectingCountry>
, Buje,
<date id="FF8A10235578FF82FF3DFEEAFE6CBF40" box="[224,403,332,357]" pageId="67" pageNumber="68" value="1975-03-27" valueMax="1975-04-04" valueMin="1975-03-27">27.iii–4.iv.1975</date>
, A. Simonetta (
<collectionCode id="ED25AE265578FF82FD8EFEEAFD66BF41" box="[595,665,332,356]" pageId="67" pageNumber="68">ABFI</collectionCode>
);
<specimenCount id="9D32FD6A5578FF82FD6EFEEAFD1DBF41" box="[691,738,332,356]" pageId="67" pageNumber="68" type="generic">3 ♂</specimenCount>
: Route Kamituga à Bukavu,
<quantity id="4CCC9B065578FF82FB95FEEAFB67BF41" box="[1096,1176,332,356]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="67" pageNumber="68" unit="m" value="1700.0">1700m</quantity>
,
<date id="FF8A10235578FF82FB77FEEAFAEBBF40" box="[1194,1300,332,357]" pageId="67" pageNumber="68" value="1949-04-05">5.iv.1949</date>
, N. Leleup (
<collectionCode id="ED25AE265578FF82FF43FED6FF0ABFAD" box="[158,245,368,392]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="67" pageNumber="68">MRAC</collectionCode>
,
<collectionCode id="ED25AE265578FF82FEDFFED6FEBDBFAD" box="[258,322,368,392]" pageId="67" pageNumber="68">JJRC</collectionCode>
).
<collectingCountry id="F32376735578FF82FE8BFEC9FE41BFAD" box="[342,446,367,392]" name="Ghana" pageId="67" pageNumber="68">
<emphasis id="B940EAF15578FF82FE8BFEC9FE41BFAD" bold="true" box="[342,446,367,392]" pageId="67" pageNumber="68">GHANA</emphasis>
</collectingCountry>
: 1 ♀: Manso, Amenfi,
<date id="FF8A10235578FF82FD16FEC9FCB9BFAD" box="[715,838,367,392]" pageId="67" pageNumber="68" value="1968-04" valueMax="1968-06-31" valueMin="1968-04-00">iv–vi.1968</date>
, ex coll. Breuning (
<collectionCode id="ED25AE265578FF82FBF5FED6FB7EBFAD" box="[1064,1153,368,392]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="67" pageNumber="68">MRAC</collectionCode>
).
<collectingCountry id="F32376735578FF82FB48FEC9FAA7BFAD" box="[1173,1368,367,392]" name="Ivory Coast" pageId="67" pageNumber="68">
<emphasis id="B940EAF15578FF82FB48FEC9FAA7BFAD" bold="true" box="[1173,1368,367,392]" pageId="67" pageNumber="68">IVORY COAST</emphasis>
</collectingCountry>
: 1 ♀:
<collectingCountry id="F32376735578FF82FF4AFE32FED2BF88" box="[151,301,404,429]" name="Ivory Coast" pageId="67" pageNumber="68">Côte d'Ivoire</collectingCountry>
, Zepréghé, Daloa,
<date id="FF8A10235578FF82FDD8FE32FDA1BF88" box="[517,606,404,429]" pageId="67" pageNumber="68" value="1962-03">iii.1962</date>
, J. Decelle (
<collectionCode id="ED25AE265578FF82FD37FE32FCBCBF89" box="[746,835,404,428]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="67" pageNumber="68">MRAC</collectionCode>
).
<collectingCountry id="F32376735578FF82FC8BFE32FC4DBF88" box="[854,946,404,429]" name="Niger" pageId="67" pageNumber="68">
<emphasis id="B940EAF15578FF82FC8BFE32FC4DBF88" bold="true" box="[854,946,404,429]" pageId="67" pageNumber="68">NIGER</emphasis>
</collectingCountry>
: 3 spec.: Oberssanga Kag., Beri b. Garnot,
<date id="FF8A10235578FF82FF4AFE11FEBABFF5" box="[151,325,439,464]" pageId="67" pageNumber="68" value="1913-02-26" valueMax="1913-03-02" valueMin="1913-02-26">26.ii–2.iii.1913</date>
, S. Tessmann (
<collectionCode id="ED25AE265578FF82FE2DFE1EFDBCBFF5" box="[496,579,440,464]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="67" pageNumber="68">MFNB</collectionCode>
,
<collectionCode id="ED25AE265578FF82FD8DFE1EFD6ABFF5" box="[592,661,440,464]" pageId="67" pageNumber="68">ABFI</collectionCode>
).
<collectingCountry id="F32376735578FF82FD74FE11FCDBBFF5" box="[681,804,439,464]" name="Nigeria" pageId="67" pageNumber="68">
<emphasis id="B940EAF15578FF82FD74FE11FCDBBFF5" bold="true" box="[681,804,439,464]" pageId="67" pageNumber="68">NIGERIA</emphasis>
</collectingCountry>
:
<specimenCount id="9D32FD6A5578FF82FCE9FE1EFCA0BFEA" box="[820,863,440,464]" pageId="67" pageNumber="68" type="generic">3 ♂</specimenCount>
: Gashaka, Gumti NP,
<quantity id="4CCC9B065578FF82FB83FE11FB5EBFF5" box="[1118,1185,439,464]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="67" pageNumber="68" unit="m" value="560.0">560m</quantity>
, Kwano,
<geoCoordinate id="EE0050245578FF82FAD0FE1EFA8BBFF5" box="[1293,1396,440,464]" direction="north" orientation="latitude" pageId="67" pageNumber="68" precision="925" value="7.3333335">07°20´N</geoCoordinate>
;
<geoCoordinate id="EE0050245578FF82FAA5FE1EFF30BFD0" direction="east" orientation="longitude" pageId="67" pageNumber="68" precision="925" value="11.583333">11° 35´E</geoCoordinate>
, 25.4.–
<date id="FF8A10235578FF82FEFDFE7AFE7ABFD0" box="[288,389,476,501]" pageId="67" pageNumber="68" value="2011-05-03">3.5.2011</date>
, Kremitovský (
<collectionCode id="ED25AE265578FF82FDECFE7AFD8FBFD1" box="[561,624,476,500]" pageId="67" pageNumber="68">JJRC</collectionCode>
,
<collectionCode id="ED25AE265578FF82FDA1FE7AFD30BFD1" box="[636,719,476,500]" pageId="67" pageNumber="68">VKBC</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="DF4B666B5578FF82FF4AF80BFF1FB9C7" httpUri="https://zenodo.org/record/289915/files/figure.png" pageId="67" pageNumber="68" targetBox="[155,1431,566,1938]" targetPageId="67">
<paragraph id="8B8B36E35578FF82FF4AF80BFF1FB9C7" blockId="67.[151,1434,1965,2018]" pageId="67" pageNumber="68">
<emphasis id="B940EAF15578FF82FF4AF80BFED8B9E6" bold="true" box="[151,295,1965,1987]" pageId="67" pageNumber="68">FIGURE 172</emphasis>
. Distribution of species in
<taxonomicName id="4C344D605578FF82FD9DF80BFCACB9E6" box="[576,851,1965,1987]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="guineensis">
<emphasis id="B940EAF15578FF82FD9DF80BFCACB9E6" box="[576,851,1965,1987]" italics="true" pageId="67" pageNumber="68">Thyreocephalus guineensis</emphasis>
</taxonomicName>
group (● =
<taxonomicName id="4C344D605578FF82FC0EF808FB22B9E6" authority="(Bernhauer)" authorityName="Bernhauer" baseAuthorityName="Bernhauer" box="[979,1245,1965,1987]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="guineensis">
<emphasis id="B940EAF15578FF82FC0EF808FBA7B9E6" box="[979,1112,1966,1987]" italics="true" pageId="67" pageNumber="68">T. guineensis</emphasis>
(Bernhauer)
</taxonomicName>
, ▲ =
<taxonomicName id="4C344D605578FF82FAC2F808FF2AB9C7" authority="Janak" authorityName="Janak" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="paraferox">
<emphasis id="B940EAF15578FF82FAC2F808FA65B9E6" box="[1311,1434,1965,1987]" italics="true" pageId="67" pageNumber="68">T. paraferox</emphasis>
Janák
</taxonomicName>
).
</paragraph>
</caption>
<caption id="DF4B666B557FFF85FF4AF80AFE67B9C4" httpUri="https://zenodo.org/record/289916/files/figure.png" pageId="68" pageNumber="69" targetBox="[187,1391,207,1926]" targetPageId="68">
<paragraph id="8B8B36E3557FFF85FF4AF80AFE67B9C4" blockId="68.[151,1436,1964,2017]" pageId="68" pageNumber="69">
<emphasis id="B940EAF1557FFF85FF4AF80AFE99B9E7" bold="true" box="[151,358,1964,1986]" pageId="68" pageNumber="69">FIGURES 173–177</emphasis>
.
<taxonomicName id="4C344D60557FFF85FEAFF80AFCF8B9E7" authority="(Bernhauer)" authorityName="Bernhauer" baseAuthorityName="Bernhauer" box="[370,775,1964,1987]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="guineensis">
<emphasis id="B940EAF1557FFF85FEAFF80AFD7BB9E6" box="[370,644,1964,1987]" italics="true" pageId="68" pageNumber="69">Thyreocephalus guineensis</emphasis>
(Bernhauer)
</taxonomicName>
; 173, head and pronotum; 174, labrum; 175, male tergite 10; 176, sternite 9; 177, aedeagus.
</paragraph>
</caption>
<subSubSection id="C32E6568557EFF84FF1AFF31FBE9BC18" pageId="69" pageNumber="70" type="description">
<paragraph id="8B8B36E3557EFF84FF1AFF31FDFEBFAD" blockId="69.[151,1437,151,752]" pageId="69" pageNumber="70">
<emphasis id="B940EAF1557EFF84FF1AFF31FE8BBE95" bold="true" box="[199,372,151,176]" pageId="69" pageNumber="70">Redescription.</emphasis>
Body length
<quantity id="4CCC9B06557EFF84FDD6FF3EFD78BE8A" box="[523,647,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="69" pageNumber="70" unit="mm" value="18.0" valueMax="19.0" valueMin="17.0">17–19 mm</quantity>
; length from anterior margin of head to posterior margin of elytra:
<quantity id="4CCC9B06557EFF84FAAFFF3EFF14BEF1" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="69" pageNumber="70" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">10– 11 mm</quantity>
. Dark reddish brown; elytra with a large yellow, lateral, anterior patch (Fig. 166); abdominal segment 7 completely or only on posterior third light red, segment 8 and genital segment light red. Head and pronotum and related punctation as in Figs. 167, 173; labrum as in
<figureCitation id="130F2A66557EFF84FD38FEA2FCB9BF39" box="[741,838,260,285]" captionStart="FIGURES 173 – 177" captionStartId="68.[151,264,1964,1986]" captionTargetBox="[187,1391,207,1926]" captionTargetId="figure@68.[177,1397,193,1940]" captionTargetPageId="68" captionText="FIGURES 173 – 177. Thyreocephalus guineensis (Bernhauer); 173, head and pronotum; 174, labrum; 175, male tergite 10; 176, sternite 9; 177, aedeagus." httpUri="https://zenodo.org/record/289916/files/figure.png" pageId="69" pageNumber="70">Fig. 174</figureCitation>
. Pronotum shorter and narrower than head, markedly dilated anteriorly, with sinuate sides. Elytra as long and wide as pronotum, with very scarcely marked humeral angles. Surface with punctation arranged in four series, two near suture, one median and one lateral. Abdomen with fine, deep and dense punctation.
</paragraph>
<paragraph id="8B8B36E3557EFF84FF1AFE32FD55BFD0" blockId="69.[151,1437,151,752]" pageId="69" pageNumber="70">
<collectingCountry id="F3237673557EFF84FF1AFE32FEFBBF88" box="[199,260,404,429]" name="Malta" pageId="69" pageNumber="70">Male</collectingCountry>
. Tergite 10 and sternite 9 of male genital segment as in
<figureCitation id="130F2A66557EFF84FCA0FE32FBDDBF88" box="[893,1058,404,429]" captionStart="FIGURES 173 – 177" captionStartId="68.[151,264,1964,1986]" captionTargetBox="[187,1391,207,1926]" captionTargetId="figure@68.[177,1397,193,1940]" captionTargetPageId="68" captionText="FIGURES 173 – 177. Thyreocephalus guineensis (Bernhauer); 173, head and pronotum; 174, labrum; 175, male tergite 10; 176, sternite 9; 177, aedeagus." httpUri="https://zenodo.org/record/289916/files/figure.png" pageId="69" pageNumber="70">Figs. 175, 176</figureCitation>
. Aedeagus (
<figureCitation id="130F2A66557EFF84FB70FE32FAF0BF89" box="[1197,1295,404,429]" captionStart="FIGURES 173 – 177" captionStartId="68.[151,264,1964,1986]" captionTargetBox="[187,1391,207,1926]" captionTargetId="figure@68.[177,1397,193,1940]" captionTargetPageId="68" captionText="FIGURES 173 – 177. Thyreocephalus guineensis (Bernhauer); 173, head and pronotum; 174, labrum; 175, male tergite 10; 176, sternite 9; 177, aedeagus." httpUri="https://zenodo.org/record/289916/files/figure.png" pageId="69" pageNumber="70">Fig. 177</figureCitation>
), very large,
<quantity id="4CCC9B06557EFF84FF4AFE1EFF11BFEA" box="[151,238,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="69" pageNumber="70" unit="mm" value="3.4">3.4 mm</quantity>
long, oblong and proportionally narrow, with broad median lobe and long parameres; inner sac tape-like, very long, folded on itself, covered with scales.
</paragraph>
<paragraph id="8B8B36E3557EFF84FF1AFE59FBE9BC18" blockId="69.[151,1437,151,752]" pageId="69" pageNumber="70">
<emphasis id="B940EAF1557EFF84FF1AFE59FE32BC3D" bold="true" box="[199,461,511,536]" pageId="69" pageNumber="70">Differential diagnosis.</emphasis>
<taxonomicName id="4C344D60557EFF84FE09FE59FCFCBC3D" box="[468,771,511,536]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="guineensis">
<emphasis id="B940EAF1557EFF84FE09FE59FCFCBC3D" box="[468,771,511,536]" italics="true" pageId="69" pageNumber="70">Thyreocephalus guineensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C344D60557EFF84FC4AFDA7FBDFBC3D" box="[919,1056,512,536]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="paraferox">
<emphasis id="B940EAF1557EFF84FC4AFDA7FBDFBC3D" box="[919,1056,512,536]" italics="true" pageId="69" pageNumber="70">T. paraferox</emphasis>
</taxonomicName>
in elytra with large yellow lateral patch and in different shape of the aedeagus with apically rounded median lobe.
</paragraph>
</subSubSection>
<subSubSection id="C32E6568557EFF84FF1AFDE1FD70BCA1" pageId="69" pageNumber="70" type="distribution">
<paragraph id="8B8B36E3557EFF84FF1AFDE1FD70BCA1" blockId="69.[151,1437,151,752]" pageId="69" pageNumber="70">
<emphasis id="B940EAF1557EFF84FF1AFDE1FE9FBC45" bold="true" box="[199,352,583,608]" pageId="69" pageNumber="70">Distribution.</emphasis>
The species is distributed in
<collectingCountry id="F3237673557EFF84FD18FDEEFCC2BC45" box="[709,829,584,608]" name="Cameroon" pageId="69" pageNumber="70">Cameroon</collectingCountry>
,
<collectingCountry id="F3237673557EFF84FC90FDE1FB8ABC45" box="[845,1141,583,608]" name="Central African Republic" pageId="69" pageNumber="70">Central African Republic</collectingCountry>
,
<collectingCountry id="F3237673557EFF84FB58FDE1FF1BBCA1" name="Democratic Republic of the Congo" pageId="69" pageNumber="70">Democratic Republic of Congo</collectingCountry>
,
<collectingCountry id="F3237673557EFF84FF32FDCAFEC4BCA0" box="[239,315,620,645]" name="Ghana" pageId="69" pageNumber="70">Ghana</collectingCountry>
,
<collectingCountry id="F3237673557EFF84FE9BFDCAFE79BCA0" box="[326,390,620,645]" name="Niger" pageId="69" pageNumber="70">Niger</collectingCountry>
and
<collectingCountry id="F3237673557EFF84FE60FDCAFDEEBCA0" box="[445,529,620,645]" name="Nigeria" pageId="69" pageNumber="70">Nigeria</collectingCountry>
(
<figureCitation id="130F2A66557EFF84FDFDFDCAFD7DBCA1" box="[544,642,620,645]" captionStart="FIGURE 172" captionStartId="67.[151,250,1965,1987]" captionTargetBox="[155,1431,566,1938]" captionTargetId="figure@67.[151,1436,553,1944]" captionTargetPageId="67" captionText="FIGURE 172. Distribution of species in Thyreocephalus guineensis group (● = T. guineensis (Bernhauer), ▲ = T. paraferox Janák)." httpUri="https://zenodo.org/record/289915/files/figure.png" pageId="69" pageNumber="70">Fig. 172</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E6568557EFF84FF1AFD29FCE6BCCA" pageId="69" pageNumber="70" type="discussion">
<paragraph id="8B8B36E3557EFF84FF1AFD29FCE6BCCA" blockId="69.[151,1437,151,752]" pageId="69" pageNumber="70">
<emphasis id="B940EAF1557EFF84FF1AFD29FEC4BC8D" bold="true" box="[199,315,655,680]" pageId="69" pageNumber="70">Remarks.</emphasis>
<collectingCountry id="F3237673557EFF84FE99FD36FE9CBC8D" box="[324,355,656,680]" name="American Samoa" pageId="69" pageNumber="70">As</collectingCountry>
the superior line of pronotal hypomeron is turning downwards well before middle, joining or almost joining inferior line next to front margin of procoxae and continuing onto anterior margin of pronotum, the species was transferred from
<taxonomicName id="4C344D60557EFF84FE02FD71FDC3BCD5" box="[479,572,727,752]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAF1557EFF84FE02FD71FDC3BCD5" box="[479,572,727,752]" italics="true" pageId="69" pageNumber="70">Eulissus</emphasis>
</taxonomicName>
to
<taxonomicName id="4C344D60557EFF84FDBDFD71FCEDBCD5" box="[608,786,727,752]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAF1557EFF84FDBDFD71FCEDBCD5" box="[608,786,727,752]" italics="true" pageId="69" pageNumber="70">Thyreocephalus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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