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<document id="3F197DE810EB424920749A48B04705F2" ID-CLB-Dataset="26928" ID-DOI="10.11646/zootaxa.4596.1.1" ID-GBIF-Dataset="cb288575-93a8-4e59-86fd-ee6bae784b6a" ID-ISSN="1175-5326" ID-Zenodo-Dep="2659091" ID-ZooBank="B88EA161-04D3-4D50-B3FD-E98A60DA3740" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1556872200693" checkinUser="plazi" docAuthor="A, Contreras-Félix Gerardo &amp; Oscar, Francke B." docDate="2019" docId="022787FEFF8C1969FF16398CFB7EFEFE" docLanguage="en" docName="zootaxa.4596.1.1.pdf" docOrigin="Zootaxa 4596 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Vaejovis maculosus , Sissom 1989" docType="treatment" docVersion="14" lastPageNumber="43" masterDocId="FE1EFF86FFA41943FF813C15FFA7FF9B" masterDocTitle="Taxonomic revision of the “ mexicanus ” group of the genus Vaejovis C. L. Koch, 1836 (Scorpiones: Vaejovidae)" masterLastPageNumber="100" masterPageNumber="1" pageNumber="41" updateTime="1701996836513" updateUser="plazi">
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<mods:title id="57A44D38475E27DA1B5A7D78B84C2760">Taxonomic revision of the “ mexicanus ” group of the genus Vaejovis C. L. Koch, 1836 (Scorpiones: Vaejovidae)</mods:title>
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<mods:namePart id="0346C099A05A4A7F43B79888A95A5177">A, Contreras-Félix Gerardo</mods:namePart>
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<taxonomicName id="4D8E4D6BFF8C196BFF16398CFD8EFA28" ID-CoL="7F8PJ" authority="Sissom, 1989" authorityName=", Sissom" authorityYear="1989" box="[151,553,1433,1459]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="maculosus">
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<emphasis id="B8FAEAFAFF8C196BFF16398CFE25FA28" bold="true" box="[151,386,1433,1459]" italics="true" pageId="40" pageNumber="41">Vaejovis maculosus</emphasis>
<bibRefCitation id="EE1F4B19FF8C196BFE08398CFD8EFA28" author="Sissom, W. D." box="[393,553,1433,1459]" pageId="40" pageNumber="41" pagination="131 - 157" refId="ref65485" refString="Sissom, W. D. (1989) Systematic studies on Vaejovis granulatus Pocock and Vaejovis pusillus Pocock, with descriptions of six new related species (Scorpiones, Vaejovidae). Revue Arachnologique, 8 (9), 131 - 157." type="journal article" year="1989">Sissom, 1989</bibRefCitation>
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<figureCitation id="12B52A6DFF8C196BFF1639AEFEA5FA4F" box="[151,258,1467,1492]" captionStart="FIGURE 4" captionStartId="12.[151,250,1698,1720]" captionTargetBox="[162,1430,957,1668]" captionTargetId="figure@12.[150,1436,949,1676]" captionTargetPageId="12" captionText="FIGURE 4. Known distribution of all the species of Vaejovis, included in the “mexicanus” group." httpUri="https://zenodo.org/record/2659101/files/figure.png" pageId="40" pageNumber="41">
Figures
<date id="FE301028FF8C196BFF7339A9FEA5FA4F" box="[242,258,1468,1492]" pageId="40" pageNumber="41" value="2013-01-04">4</date>
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,
<figureCitation id="12B52A6DFF8C196BFE8F39AEFE93FA4F" box="[270,308,1467,1492]" captionStart="FIGURE 13" captionStartId="22.[151,250,989,1011]" captionTargetBox="[151,1435,193,968]" captionTargetId="figure@22.[151,1435,193,968]" captionTargetPageId="22" captionText="FIGURE 13. Comparison in shapes on carapace in species within “mexicanus” group: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659119/files/figure.png" pageId="40" pageNumber="41">13i</figureCitation>
,
<figureCitation id="12B52A6DFF8C196BFEC139AEFECBFA4F" box="[320,364,1467,1492]" captionStart="FIGURE 14" captionStartId="23.[151,250,1558,1580]" captionTargetBox="[340,1236,345,1537]" captionTargetId="figure@23.[340,1247,337,1537]" captionTargetPageId="23" captionText="FIGURE 14. Comparison of the different shapes in the caudal gland on species within the “mexicanus” group (each caudal gland, if present, is highlighted in red): a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov." httpUri="https://zenodo.org/record/2659121/files/figure.png" pageId="40" pageNumber="41">14h</figureCitation>
,
<figureCitation id="12B52A6DFF8C196BFEF939AEFE3AFA4F" box="[376,413,1467,1492]" captionStart="FIGURE 15" captionStartId="24.[151,250,1398,1420]" captionTargetBox="[340,1247,195,1374]" captionTargetId="figure@24.[340,1247,195,1374]" captionTargetPageId="24" captionText="FIGURE 15. Comparison in shapes and trichobothrial patterns in the pedipalp femur, in species within “mexicanus” group: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659123/files/figure.png" pageId="40" pageNumber="41">
<date id="FE301028FF8C196BFEF939AEFE3AFA4F" box="[376,413,1467,1492]" pageId="40" pageNumber="41" value="2015-01">15i</date>
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,
<figureCitation id="12B52A6DFF8C196BFE2839AEFE68FA4F" box="[425,463,1467,1492]" captionStart="FIGURE 16" captionStartId="25.[151,250,1480,1502]" captionTargetBox="[340,1247,258,1458]" captionTargetId="figure@25.[340,1247,258,1458]" captionTargetPageId="25" captionText="FIGURE 16. Comparison in shapes and trichobothrial patterns in the pedipalp patella, in species within “mexicanus” group: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659125/files/figure.png" pageId="40" pageNumber="41">
<date id="FE301028FF8C196BFE2839AEFE68FA4F" box="[425,463,1467,1492]" pageId="40" pageNumber="41" value="2016-01">16i</date>
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<figureCitation id="12B52A6DFF8C196BFE5A39AEFDA6FA4F" box="[475,513,1467,1492]" captionStart="FIGURE 17" captionStartId="26.[151,250,1585,1607]" captionTargetBox="[335,1252,335,1567]" captionTargetId="figure@26.[335,1252,335,1567]" captionTargetPageId="26" captionText="FIGURE 17. Comparison in shapes and trichobothrial patterns in the pedipalp chela of males in species within “mexicanus” group: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659127/files/figure.png" pageId="40" pageNumber="41">
<date id="FE301028FF8C196BFE5A39AEFDA6FA4F" box="[475,513,1467,1492]" pageId="40" pageNumber="41" value="2017-01">17i</date>
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<figureCitation id="12B52A6DFF8C196BFD8C39AEFD95FA4F" box="[525,562,1467,1492]" captionStart="FIGURE 18" captionStartId="27.[151,250,1747,1769]" captionTargetBox="[340,1247,589,1726]" captionTargetId="figure@27.[340,1247,589,1726]" captionTargetPageId="27" captionText="FIGURE 18. Comparison in the different shapes and trichobothrial patterns in the pedipalp chela of females, that selected species within “mexicanus” group present: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659129/files/figure.png" pageId="40" pageNumber="41">
<date id="FE301028FF8C196BFD8C39AEFD95FA4F" box="[525,562,1467,1492]" pageId="40" pageNumber="41" value="2018-01">18i</date>
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,
<figureCitation id="12B52A6DFF8C196BFDBE39AEFDC3FA4F" box="[575,612,1467,1492]" captionStart="FIGURE 19" captionStartId="29.[151,250,1805,1827]" captionTargetBox="[319,1255,877,1784]" captionTargetId="figure@29.[319,1267,877,1784]" captionTargetPageId="29" captionText="FIGURE 19. Comparison on the shapes in the hemispermatophore in species within “mexicanus” group: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659131/files/figure.png" pageId="40" pageNumber="41">
<date id="FE301028FF8C196BFDBE39AEFDC3FA4F" box="[575,612,1467,1492]" pageId="40" pageNumber="41" value="1919-01">19i</date>
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,
<figureCitation id="12B52A6DFF8C196BFDF139A9FD2BFA4F" box="[624,652,1468,1492]" captionStart="FIGURE 27" captionStartId="41.[151,250,1870,1892]" captionTargetBox="[341,1241,661,1849]" captionTargetId="figure@41.[340,1247,661,1849]" captionTargetPageId="41" captionText="FIGURE 27. Habitus of Vaejovis maculosus (♂): a—dorsal view; b—ventral view. Scale bar = 1 cm." httpUri="https://zenodo.org/record/2659147/files/figure.png" pageId="40" pageNumber="41">27</figureCitation>
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<emphasis id="B8FAEAFAFF8C196BFF163A10FEFAF981" box="[151,349,1540,1562]" italics="true" pageId="40" pageNumber="41">Vaejovis maculosus</emphasis>
<bibRefCitation id="EE1F4B19FF8C196BFEE53A11FE57F981" author="Sissom, W. D." box="[356,496,1540,1562]" pageId="40" pageNumber="41" pagination="131 - 157" refId="ref65485" refString="Sissom, W. D. (1989) Systematic studies on Vaejovis granulatus Pocock and Vaejovis pusillus Pocock, with descriptions of six new related species (Scorpiones, Vaejovidae). Revue Arachnologique, 8 (9), 131 - 157." type="journal article" year="1989">Sissom, 1989</bibRefCitation>
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–144, Fig: 29–36, 74–75;
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, 144, 153, map 71;
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<bibRefCitation id="EE1F4B19FF8C196BFF6C3A37FEDDF9A3" author="Sissom, W. D." box="[237,378,1570,1592]" pageId="40" pageNumber="41" pagination="503 - 553" refId="ref65622" refString="Sissom, W. D. (2000) Family Vaejovidae. In: Fet, V., Sissom, W. D., Lowe, G. &amp; Braunwalder, M., Catalog of the scorpions of the world (1758 - 1997). New York Entomological Society, New York, pp. 503 - 553." type="book chapter" year="2000">Sissom, 2000</bibRefCitation>
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<bibRefCitation id="EE1F4B19FF8C196BFE363A37FD35F9A3" author="Soleglad, M. &amp; Fet, V." box="[439,658,1570,1592]" pageId="40" pageNumber="41" pagination="1 - 115" refId="ref66110" refString="Soleglad, M. E &amp; Fet, V. (2008) Contributions to Scorpion Systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae). Euscorpius, 71, 1 - 115." type="journal article" year="2008">Soleglad &amp; Fet, 2008</bibRefCitation>
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, 77, 100.
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<emphasis id="B8FAEAFAFF8C196BFF163A71FEE1F9E6" bold="true" box="[151,326,1636,1661]" pageId="40" pageNumber="41">Type material.</emphasis>
<materialsCitation id="3AE63CB5FF8C196BFECF3A71FBEEF939" ID-GBIF-Occurrence="2237797910" collectingDate="1984-05-28" collectionCode="AMNH" collectorName="NE Zacatepec. &amp; Cols. &amp; D. Sissom &amp; C. S. Colwell. &amp; Examined." country="Mexico" location="Totalco" pageId="40" pageNumber="41" specimenCount="1" stateProvince="Puebla" typeStatus="holotype">
<typeStatus id="5535884AFF8C196BFECF3A71FE11F9E6" box="[334,438,1636,1661]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="F2997678FF8C196BFD843A70FDDFF9E6" box="[517,632,1637,1661]" name="Mexico" pageId="40" pageNumber="41">MEXICO</collectingCountry>
:
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, from
<quantity id="4D769B0DFF8C196BFC9F3A71FCDEF9E6" box="[798,889,1636,1661]" metricMagnitude="4" metricUnit="m" metricValue="1.207008" metricValueMax="1.609344" metricValueMin="0.804672" pageId="40" pageNumber="41" unit="mi" value="7.5" valueMax="10.0" valueMin="5.0">5–10 mi</quantity>
toward
<location id="8F516033FF8C196BFC583A71FB97F9E6" LSID="urn:lsid:plazi:treatment:022787FEFF8C1969FF16398CFB7EFEFE:8F516033FF8C196BFC583A71FB97F9E6" box="[985,1072,1636,1661]" country="Mexico" name="Totalco" pageId="40" pageNumber="41" stateProvince="Puebla">Totalco</location>
, along MEX highway 140, hills
<collectorName id="277B533EFF8C196BFF163A9FFE9BF93A" box="[151,316,1674,1697]" pageId="40" pageNumber="41">NE Zacatepec.</collectorName>
<date id="FE301028FF8C196BFEC23A9CFE1CF93A" box="[323,443,1673,1697]" pageId="40" pageNumber="41" value="1984-05-28">
<collectingDate id="EE74E9C0FF8C196BFEC23A9CFE1CF93A" box="[323,443,1673,1697]" pageId="40" pageNumber="41" value="1984-05-28">V-28-1984</collectingDate>
</date>
<collectorName id="277B533EFF8C196BFE433A9CFDA7F939" box="[450,512,1673,1698]" pageId="40" pageNumber="41">Cols.</collectorName>
:
<collectorName id="277B533EFF8C196BFD8A3A9FFDD9F939" box="[523,638,1673,1698]" pageId="40" pageNumber="41">D. Sissom</collectorName>
and
<collectorName id="277B533EFF8C196BFD343A9CFCF4F939" box="[693,851,1673,1698]" pageId="40" pageNumber="41">C. S. Colwell.</collectorName>
(
<collectionCode id="EC9FAE2DFF8C196BFCE03A9CFC1AF93A" box="[865,957,1673,1697]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="40" pageNumber="41">AMNH</collectionCode>
).
<collectorName id="277B533EFF8C196BFC513A9CFBEEF939" box="[976,1097,1673,1698]" pageId="40" pageNumber="41">Examined.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8A3136E8FF8C196BFF463ABBFCD2F971" blockId="40.[151,1436,1636,2022]" pageId="40" pageNumber="41">
<materialsCitation id="3AE63CB5FF8C196BFF463ABBFCFBF95E" ID-GBIF-Occurrence="2237797898" box="[199,860,1708,1734]" collectingDate="1984-05-28" collectionCode="AMNH" collectorName="NE Zacatepec. &amp; Cols. &amp; D. Sissom &amp; C. S. Colwell. &amp; Examined." country="Mexico" location="Totalco" pageId="40" pageNumber="41" specimenCount="1" specimenCount-female="1" stateProvince="Puebla" typeStatus="paratype">
<typeStatus id="5535884AFF8C196BFF463ABBFE9EF95D" box="[199,313,1710,1734]" pageId="40" pageNumber="41" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9C88FD61FF8C196BFEC33AB8FECBF95E" box="[322,364,1709,1733]" pageId="40" pageNumber="41" type="female">1 ♀</specimenCount>
, same data as holotype (
<collectionCode id="EC9FAE2DFF8C196BFDFC3AB8FD7EF95E" box="[637,729,1709,1733]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="40" pageNumber="41">AMNH</collectionCode>
), examined
</materialsCitation>
;
<materialsCitation id="3AE63CB5FF8C196BFCE73AB8FA3FF95E" ID-GBIF-Occurrence="2237797823" box="[870,1432,1708,1733]" collectingDate="1984-05-28" collectionCode="CAS" collectorName="NE Zacatepec. &amp; Cols. &amp; D. Sissom &amp; C. S. Colwell. &amp; Examined." country="Mexico" location="Totalco" pageId="40" pageNumber="41" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Puebla" typeStatus="paratype">
<specimenCount id="9C88FD61FF8C196BFCE73AB8FC35F95E" box="[870,914,1709,1733]" pageId="40" pageNumber="41" type="male">1 ♂</specimenCount>
,
<specimenCount id="9C88FD61FF8C196BFC1F3AB8FC60F95E" box="[926,967,1709,1733]" pageId="40" pageNumber="41" type="female">1 ♀</specimenCount>
, same data as holotype (
<collectionCode id="EC9FAE2DFF8C196BFB583AB8FAB3F95E" box="[1241,1300,1709,1733]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="40" pageNumber="41" type="Museum">CAS</collectionCode>
), examined
</materialsCitation>
;
<materialsCitation id="3AE63CB5FF8C196BFF163AC4FCD5F971" ID-GBIF-Occurrence="2237797939" box="[151,882,1745,1770]" collectingDate="1984-05-28" collectionCode="WDS" collectorName="NE Zacatepec. &amp; Cols. &amp; D. Sissom &amp; C. S. Colwell. &amp; Examined." country="Mexico" location="Totalco" pageId="40" pageNumber="41" specimenCount="18" specimenCount-female="4" specimenCount-male="14" stateProvince="Puebla" typeStatus="paratype">
<specimenCount id="9C88FD61FF8C196BFF163AC4FF76F972" box="[151,209,1745,1769]" pageId="40" pageNumber="41" type="male">14 ♂</specimenCount>
,
<specimenCount id="9C88FD61FF8C196BFF5C3AC4FEA0F972" box="[221,263,1745,1769]" pageId="40" pageNumber="41" type="female">4 ♀</specimenCount>
and 1 J#, same data as holotype (
<collectionCode id="EC9FAE2DFF8C196BFDFE3AC4FD65F972" box="[639,706,1745,1769]" pageId="40" pageNumber="41">WDS</collectionCode>
), not examined
</materialsCitation>
.
</paragraph>
<paragraph id="8A3136E8FF8C196BFF463AE1FE1FF805" blockId="40.[151,1436,1636,2022]" pageId="40" pageNumber="41">
<emphasis id="B8FAEAFAFF8C196BFF463AE1FDB1F896" bold="true" box="[199,534,1780,1805]" pageId="40" pageNumber="41">Other specimens examined.</emphasis>
<taxonomicName id="4D8E4D6BFF8C196BFD9E3AE3FD5BF896" authorityName=", Sissom" authorityYear="1989" box="[543,764,1781,1805]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="maculosus">
<emphasis id="B8FAEAFAFF8C196BFD9E3AE3FD5BF896" box="[543,764,1781,1805]" italics="true" pageId="40" pageNumber="41">Vaejovis maculosus</emphasis>
</taxonomicName>
:
<materialsCitation id="3AE63CB5FF8C196BFC8F3AE0FEF8F8CE" ID-GBIF-Occurrence="2237797909" collectingDate="2006-05-07" collectionCode="CNAN-SC" collectorName="El Limon &amp; Cols. &amp; O. Francke &amp; A. Ballesteros." country="Mexico" elevation="2348" latitude="19.48289" location="El Limon" longLatPrecision="1" longitude="-97.36631" municipality="Totalco" pageId="40" pageNumber="41" specimenCode="CNAN-SC3638" specimenCount="2" specimenCount-female="2" stateProvince="Veracruz de Ignacio de la Llave">
<collectingCountry id="F2997678FF8C196BFC8F3AE0FC26F896" box="[782,897,1781,1805]" name="Mexico" pageId="40" pageNumber="41">MEXICO</collectingCountry>
:
<collectingRegion id="484AF80AFF8C196BFC0F3AE3FC52F896" box="[910,1013,1782,1805]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="40" pageNumber="41">Veracruz</collectingRegion>
,
<collectingMunicipality id="6A55AC92FF8C196BFB823AE1FBFDF896" box="[1027,1114,1780,1805]" pageId="40" pageNumber="41">Totalco</collectingMunicipality>
,
<quantity id="4D769B0DFF8C196BFBE63AE0FB01F896" box="[1127,1190,1780,1805]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="40" pageNumber="41" unit="km" value="2.0">2 km</quantity>
. SW from
<location id="8F516033FF8C196BFAA63AE1FA3FF896" LSID="urn:lsid:plazi:treatment:022787FEFF8C1969FF16398CFB7EFEFE:8F516033FF8C196BFAA63AE1FA3FF896" box="[1319,1432,1780,1805]" country="Mexico" latitude="19.48289" longLatPrecision="1" longitude="-97.36631" municipality="Totalco" name="El Limon" pageId="40" pageNumber="41" stateProvince="Veracruz de Ignacio de la Llave">
<collectorName id="277B533EFF8C196BFAA63AE1FA3FF896" box="[1319,1432,1780,1805]" pageId="40" pageNumber="41">El Limón</collectorName>
</location>
, highway 140 (
<geoCoordinate id="EFBA502FFF8C196BFEBC3B0CFE76F8AA" box="[317,465,1817,1841]" direction="north" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="19.48289">N 19.48289°</geoCoordinate>
,
<geoCoordinate id="EFBA502FFF8C196BFE5F3B0FFDDFF8A9" box="[478,632,1817,1842]" direction="west" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="-97.36631">W 97.36631°</geoCoordinate>
, elev.
<quantity id="4D769B0DFF8C196BFD423B0CFC87F8AA" box="[707,800,1817,1841]" metricMagnitude="3" metricUnit="m" metricValue="2.348" pageId="40" pageNumber="41" unit="m" value="2348.0">
<elevation id="01A3D1DBFF8C196BFD423B0CFC87F8AA" box="[707,800,1817,1841]" metricMagnitude="3" metricUnit="m" metricValue="2.348" pageId="40" pageNumber="41" unit="m" value="2348.0">2348 m</elevation>
</quantity>
.).
<date id="FE301028FF8C196BFCBA3B0CFC0EF8A9" box="[827,937,1817,1842]" pageId="40" pageNumber="41" value="2006-05-07">
<collectingDate id="EE74E9C0FF8C196BFCBA3B0CFC0EF8A9" box="[827,937,1817,1842]" pageId="40" pageNumber="41" value="2006-05-07">V-7-2006</collectingDate>
</date>
.
<collectorName id="277B533EFF8C196BFC373B0CFC52F8A9" box="[950,1013,1817,1842]" pageId="40" pageNumber="41">Cols.</collectorName>
:
<collectorName id="277B533EFF8C196BFB833B0CFB26F8A9" box="[1026,1153,1817,1842]" pageId="40" pageNumber="41">O. Francke</collectorName>
and
<collectorName id="277B533EFF8C196BFB3C3B0CFAC1F8A9" box="[1213,1382,1817,1842]" pageId="40" pageNumber="41">A. Ballesteros.</collectorName>
<specimenCount id="9C88FD61FF8C196BFAEE3B0CFA3BF8AA" box="[1391,1436,1817,1841]" pageId="40" pageNumber="41" type="female">2 ♀</specimenCount>
(
<specimenCode id="DA289E93FF8C196BFF1F3B29FEFFF8CE" box="[158,344,1852,1877]" collectionCode="CNAN-SC" pageId="40" pageNumber="41">CNAN-SC3638</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3AE63CB5FF8C196BFEEB3B28FDA0F8CE" ID-GBIF-Occurrence="2237797826" box="[362,519,1853,1877]" collectionCode="AMNH" country="Mexico" pageId="40" pageNumber="41" specimenCount="2" specimenCount-female="2" stateProvince="Veracruz de Ignacio de la Llave">
<specimenCount id="9C88FD61FF8C196BFEEB3B28FE33F8CE" box="[362,404,1853,1877]" pageId="40" pageNumber="41" type="female">2 ♀</specimenCount>
(
<collectionCode id="EC9FAE2DFF8C196BFE233B28FE59F8CE" box="[418,510,1853,1877]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="40" pageNumber="41">AMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3AE63CB5FF8C196BFD903B28FE13F805" ID-GBIF-Occurrence="2237797943" collectingDate="2013-06-18" collectionCode="CNAN-SC" collectorName="Cerro de la Cruz &amp; Cols. &amp; O. Francke &amp; J. Mendoza &amp; R. Monjaraz &amp; J. Bokma &amp; G. Contreras." country="Mexico" elevation="2685" latitude="19.47055" location="Cerro de la Cruz" longLatPrecision="1" longitude="-97.26994" municipality="San Martin Tenextepec" pageId="40" pageNumber="41" specimenCode="CNAN-SC3632" specimenCount="14" specimenCount-female="9" specimenCount-male="5" stateProvince="Veracruz de Ignacio de la Llave">
<collectingCountry id="F2997678FF8C196BFD903B28FD23F8CE" box="[529,644,1853,1877]" name="Mexico" pageId="40" pageNumber="41">MEXICO</collectingCountry>
:
<collectingRegion id="484AF80AFF8C196BFD0F3B2BFD53F8CE" box="[654,756,1854,1877]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="40" pageNumber="41">Veracruz</collectingRegion>
,
<collectingMunicipality id="6A55AC92FF8C196BFD7E3B28FBA2F8CE" box="[767,1029,1852,1877]" pageId="40" pageNumber="41">San Martin Tenextepec</collectingMunicipality>
,
<location id="8F516033FF8C196BFB913B28FB6EF8CE" LSID="urn:lsid:plazi:treatment:022787FEFF8C1969FF16398CFB7EFEFE:8F516033FF8C196BFB913B28FB6EF8CE" box="[1040,1225,1852,1877]" country="Mexico" latitude="19.47055" longLatPrecision="1" longitude="-97.26994" municipality="San Martin Tenextepec" name="Cerro de la Cruz" pageId="40" pageNumber="41" stateProvince="Veracruz de Ignacio de la Llave">
<collectorName id="277B533EFF8C196BFB913B28FB6EF8CE" box="[1040,1225,1852,1877]" pageId="40" pageNumber="41">Cerro de la Cruz</collectorName>
</location>
(
<geoCoordinate id="EFBA502FFF8C196BFB593B28FAD0F8CE" box="[1240,1399,1852,1877]" direction="north" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="19.47055">N 19.470551°</geoCoordinate>
,
<geoCoordinate id="EFBA502FFF8C196BFA033B2BFEACF8E1" direction="west" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="-97.26994">W 97.26994°</geoCoordinate>
, elev.
<quantity id="4D769B0DFF8C196BFED03B74FE0BF8E2" box="[337,428,1889,1914]" metricMagnitude="3" metricUnit="m" metricValue="2.685" pageId="40" pageNumber="41" unit="m" value="2685.0">
<elevation id="01A3D1DBFF8C196BFED03B74FE0BF8E2" box="[337,428,1889,1914]" metricMagnitude="3" metricUnit="m" metricValue="2.685" pageId="40" pageNumber="41" unit="m" value="2685.0">2685 m</elevation>
</quantity>
.).
<date id="FE301028FF8C196BFE443B74FDEBF8E2" box="[453,588,1889,1913]" pageId="40" pageNumber="41" value="2013-06-18">
<collectingDate id="EE74E9C0FF8C196BFE443B74FDEBF8E2" box="[453,588,1889,1913]" pageId="40" pageNumber="41" value="2013-06-18">VI-18-2013</collectingDate>
</date>
.
<collectorName id="277B533EFF8C196BFDD63B74FD32F8E1" box="[599,661,1889,1914]" pageId="40" pageNumber="41">Cols.</collectorName>
:
<collectorName id="277B533EFF8C196BFD213B74FCB8F8E1" box="[672,799,1889,1914]" pageId="40" pageNumber="41">O. Francke</collectorName>
,
<collectorName id="277B533EFF8C196BFCAA3B77FC08F8E1" box="[811,943,1889,1914]" pageId="40" pageNumber="41">J. Mendoza</collectorName>
,
<collectorName id="277B533EFF8C196BFC3B3B77FBEDF8E1" box="[954,1098,1889,1914]" pageId="40" pageNumber="41">R. Monjaraz</collectorName>
,
<collectorName id="277B533EFF8C196BFBD53B77FB19F8E1" box="[1108,1214,1889,1914]" pageId="40" pageNumber="41">J. Bokma</collectorName>
and
<collectorName id="277B533EFF8C196BFB743B74FA2FF8E2" box="[1269,1416,1889,1913]" pageId="40" pageNumber="41">G. Contreras.</collectorName>
<specimenCount id="9C88FD61FF8C196BFA0F3B74FF09F806" pageId="40" pageNumber="41" type="male">5 ♂</specimenCount>
,
<specimenCount id="9C88FD61FF8C196BFF3B3B90FF43F806" box="[186,228,1925,1949]" pageId="40" pageNumber="41" type="female">9 ♀</specimenCount>
(
<specimenCode id="DA289E93FF8C196BFF723B90FE0AF805" box="[243,429,1925,1950]" collectionCode="CNAN-SC" pageId="40" pageNumber="41">CNAN-SC3632</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C2946563FF8C1969FF463BBCFB5DFF2B" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="8A3136E8FF8C196AFF463BBCFBDBFDFB" blockId="40.[151,1436,1636,2022]" lastBlockId="41.[151,1437,151,608]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<emphasis id="B8FAEAFAFF8C196BFF463BBCFEE6F859" bold="true" box="[199,321,1961,1986]" pageId="40" pageNumber="41">Diagnosis.</emphasis>
Adults
<quantity id="4D769B0DFF8C196BFE1C3BBCFD96F85A" box="[413,561,1961,1986]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.3" metricValueMin="1.5" pageId="40" pageNumber="41" unit="mm" value="19.0" valueMax="23.0" valueMin="15.0">15 to 23 mm</quantity>
long (Fig: 27-a, b), conspicuously mottled throughout. Carapace (Fig:
<date id="FE301028FF8C196BFADF3BBCFA29F859" box="[1374,1422,1961,1986]" bridgedPair="-" pageId="40" pageNumber="41" value="2013-01">13-i</date>
), shorter than metasomal segment V (Carapace L/MS-V 0.91±0.01 [0.91±0.08]), longer than femur (Carapace L/ Femur L 1.28±0.07 [1.29±0.05]); anterior margin of carapace straight. Mesosomal tergite VII with a pair of well developed, granular carinae present from middle of segment and not reaching posterior margin. Pectinal teeth 16–17 [14–15]; sternite V on males usually with a conspicuous shiny whitish patch, half oval-shaped at posterior margin; sternite VII with a pair of vestigial median carinae and 12 to 14 setae (mode: 12) [12–16 (mode 15)]. Metasomal segment I sub-quadrangular in dorsal view (MSI L/
<geoCoordinate id="EFBA502FFF8D196AFCEB3D3DFC19FEDB" box="[874,958,296,320]" direction="west" orientation="longitude" pageId="41" pageNumber="42" precision="555" value="-0.74">W 0.74</geoCoordinate>
±0.04 [0.71±0.05]); intercarinal spaces on segments I–IV smooth, may present some scattered granules; segment V with noticeably darker coloration than rest of metasoma. Vesicle slender (Fig:
<date id="FE301028FF8D196AFDA03D7AFDE8FE13" box="[545,591,367,392]" bridgedPair="-" pageId="41" pageNumber="42" value="2014-01">14-i</date>
) (L/
<geoCoordinate id="EFBA502FFF8D196AFD013D65FD74FE13" box="[640,723,367,392]" direction="west" orientation="longitude" pageId="41" pageNumber="42" precision="555" value="-1.96">W 1.96</geoCoordinate>
±0.3 [1.85±0.09]) and thin (W/D 1.25±0.19 [1.3±0.13]), ventral surface smooth; with two median longitudinal lighter bands, slightly differentiated in color; dorsal surface smooth and flat, with a slender, long and diffuse whitish vesicular gland. Pedipalp femur (Fig:
<date id="FE301028FF8D196AFBE93DA2FB32FE4B" box="[1128,1173,439,464]" bridgedPair="-" pageId="41" pageNumber="42" value="2015-01">15-i</date>
) and patella (Fig:
<date id="FE301028FF8D196AFAE73DA2FA34FE4B" box="[1382,1427,439,464]" bridgedPair="-" pageId="41" pageNumber="42" value="2016-01">16-i</date>
) always less than three times longer than wide (Femur L/
<geoCoordinate id="EFBA502FFF8D196AFCD63DC9FC16FE6E" box="[855,945,476,501]" direction="west" orientation="longitude" pageId="41" pageNumber="42" precision="555" value="-2.79">W 2.79</geoCoordinate>
±0.06 [2.92±0.08]; patella L/
<geoCoordinate id="EFBA502FFF8D196AFA8E3DC9FACDFE6E" box="[1295,1386,476,501]" direction="west" orientation="longitude" pageId="41" pageNumber="42" precision="555" value="-2.76">W 2.76</geoCoordinate>
±0.2 [2.68±0.1]); patella dorsointernal carina granulose, composed by a single, continuous row of granules. Pedipalp chela (Fig: 17,
<date id="FE301028FF8D196AFEC13E31FEC9FDA6" box="[320,366,548,573]" bridgedPair="-" pageId="41" pageNumber="42" value="2018-01">18-i</date>
) on male greatly inflated; less than twice as long as wide (CM L/
<geoCoordinate id="EFBA502FFF8D196AFBCA3E31FB3AFDA6" box="[1099,1181,548,573]" direction="west" orientation="longitude" pageId="41" pageNumber="42" precision="555" value="-1.75">W 1.75</geoCoordinate>
±0.1 [1.86±0.6]), keelless, with the surface smooth. Pedipalp chela dentate margin of fingers on males straight.
</paragraph>
<caption id="DEF16660FF8D196AFF163B5BFB3AF8FF" box="[151,1181,1870,1892]" httpUri="https://zenodo.org/record/2659147/files/figure.png" pageId="41" pageNumber="42" startId="41.[151,250,1870,1892]" targetBox="[341,1241,661,1849]" targetPageId="41">
<paragraph id="8A3136E8FF8D196AFF163B5BFB3AF8FF" blockId="41.[151,1181,1870,1892]" box="[151,1181,1870,1892]" pageId="41" pageNumber="42">
<emphasis id="B8FAEAFAFF8D196AFF163B5BFEB8F8FF" bold="true" box="[151,287,1870,1892]" pageId="41" pageNumber="42">FIGURE 27.</emphasis>
Habitus of
<taxonomicName id="4D8E4D6BFF8D196AFE173B5AFDFBF8FF" authorityName=", Sissom" authorityYear="1989" box="[406,604,1870,1892]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="maculosus">
<emphasis id="B8FAEAFAFF8D196AFE173B5AFDFBF8FF" box="[406,604,1870,1892]" italics="true" pageId="41" pageNumber="42">Vaejovis maculosus</emphasis>
</taxonomicName>
(♂):
<emphasis id="B8FAEAFAFF8D196AFD123B5BFD1FF8FF" bold="true" box="[659,696,1870,1892]" pageId="41" pageNumber="42">a—</emphasis>
dorsal view;
<emphasis id="B8FAEAFAFF8D196AFCBD3B5BFCC4F8FF" bold="true" box="[828,867,1870,1892]" pageId="41" pageNumber="42">b—</emphasis>
ventral view. Scale bar = 1 cm.
</paragraph>
</caption>
<paragraph id="8A3136E8FF8D196AFF463B9EFC5BF877" blockId="41.[151,1436,1931,2029]" pageId="41" pageNumber="42">
Hemispermatophore (Fig:
<date id="FE301028FF8D196AFE7B3B9EFD8CF83F" box="[506,555,1931,1956]" bridgedPair="-" pageId="41" pageNumber="42" value="1919-01">19-i</date>
): Lamelliform (TL 2.3; LL 1.5; LW
<quantity id="4D769B0DFF8D196AFC663B99FBE1F83F" box="[999,1094,1932,1956]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="41" pageNumber="42" unit="mm" value="0.3">0.3 mm</quantity>
); lamella simple, with hooks located basally, lobed, the accessory hook merged to principal lobe and not well differentiated; notch present basally on the lamella, at level where hooks are located. Median lobe present.
</paragraph>
<paragraph id="8A3136E8FF8E1969FF463C82FB5DFF2B" blockId="42.[151,1437,151,357]" box="[199,1274,151,176]" pageId="42" pageNumber="43">Full morphometric variation is given in tables 13 and 14. Setae variation is given in tables 9–12.</paragraph>
</subSubSection>
<subSubSection id="C2946563FF8E1969FF463CA9FB7EFEFE" pageId="42" pageNumber="43" type="distribution">
<paragraph id="8A3136E8FF8E1969FF463CA9FCAFFF4F" blockId="42.[151,1437,151,357]" box="[199,776,188,213]" pageId="42" pageNumber="43">
<emphasis id="B8FAEAFAFF8E1969FF463CA9FEC7FF4E" bold="true" box="[199,352,188,213]" pageId="42" pageNumber="43">Distribution.</emphasis>
<collectingCountry id="F2997678FF8E1969FEE63CA9FE66FF4E" box="[359,449,188,213]" name="Mexico" pageId="42" pageNumber="43">Mexico</collectingCountry>
:
<collectingRegion id="484AF80AFF8E1969FE4A3CA9FDBDFF4E" box="[459,538,188,213]" country="Mexico" name="Puebla" pageId="42" pageNumber="43">Puebla</collectingRegion>
,
<collectingRegion id="484AF80AFF8E1969FDA73CA8FD2DFF4F" box="[550,650,189,212]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="42" pageNumber="43">Veracruz</collectingRegion>
(
<figureCitation id="12B52A6DFF8E1969FD193CA9FD5DFF4F" box="[664,762,188,213]" captionStart="FIGURE 4" captionStartId="12.[151,250,1698,1720]" captionTargetBox="[162,1430,957,1668]" captionTargetId="figure@12.[150,1436,949,1676]" captionTargetPageId="12" captionText="FIGURE 4. Known distribution of all the species of Vaejovis, included in the “mexicanus” group." httpUri="https://zenodo.org/record/2659101/files/figure.png" pageId="42" pageNumber="43">Figure 4</figureCitation>
).
</paragraph>
<paragraph id="8A3136E8FF8E1969FF463CCAFB7EFEFE" blockId="42.[151,1437,151,357]" pageId="42" pageNumber="43">
<emphasis id="B8FAEAFAFF8E1969FF463CCAFE2FFF63" bold="true" box="[199,392,223,248]" pageId="42" pageNumber="43">Natural history.</emphasis>
This species inhabits the highlands of part of the Mexican states of
<collectingRegion id="484AF80AFF8E1969FB283CCAFB52FF63" box="[1193,1269,223,248]" country="Mexico" name="Puebla" pageId="42" pageNumber="43">Puebla</collectingRegion>
and
<collectingRegion id="484AF80AFF8E1969FAB03CF5FA3FFF6C" box="[1329,1432,224,247]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="42" pageNumber="43">Veracruz</collectingRegion>
. Some specimens were collected in re-planted pine forest, but also can be found under and between the leaves of dead
<emphasis id="B8FAEAFAFF8E1969FF563D3DFEEEFEDB" box="[215,329,296,320]" italics="true" pageId="42" pageNumber="43">
<taxonomicName id="4D8E4D6BFF8E1969FF563D3DFEBAFEA4" box="[215,285,296,319]" class="Liliopsida" family="Asparagaceae" genus="Agave" kingdom="Plantae" order="Asparagales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="genus">Agave</taxonomicName>
sp.
</emphasis>
in a very humid microhabitat. Additionally, this species has a litter size of approximately
<specimenCount id="9C88FD61FF8E1969FA013D3DFEA7FEFE" pageId="42" pageNumber="43" type="juvenile">20 juveniles</specimenCount>
; females with their broods and adult males can be found in the months of June and July.
</paragraph>
</subSubSection>
</treatment>
</document>
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