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<document id="EEA06835077DF0A8EA03D97F8D074C87" ID-DOI="10.5281/zenodo.13158554" ID-ISSN="0068-547X" ID-Zenodo-Dep="13158554" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722539346815" checkinUser="felipe" docAuthor="Griswold, Charles, Ubick, Darrell, Ledford, Joel &amp; Polotow, Daniele" docDate="2022" docId="FB2A53331875FFE9473432B2AFA2EF30" docLanguage="en" docName="ProcCalifAcadSci.67.suppl2.1-193.pdf" docOrigin="Proceedings of the California Academy of Sciences 67" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Uduba irwini Griswold, Ubick, Ledford &amp; Polotow, 2022, new species" docType="treatment" docUuid="9629A679-9F71-4D5F-B36D-F482B0A244D6" docUuidSource="ZooBank" docVersion="2" lastPageNumber="52" masterDocId="07132B4B1844FFDA474F310FAB37E94C" masterDocTitle="A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar" masterLastPageNumber="193" masterPageNumber="1" pageNumber="50" updateTime="1722553396936" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8A9D556F2618C82A7DEF2BDE92B9BD7F">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
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<mods:namePart id="DAAE5C32E44A060BEA8B48D80BE2E521">Griswold, Charles</mods:namePart>
<mods:affiliation id="0C33003D1297C77B0FDF985D6CFE9635">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
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<mods:namePart id="E1E7AEF2BDF2DCD468CC50E386EFEAAE">Ubick, Darrell</mods:namePart>
<mods:affiliation id="5A5A1F217DE7F2D60A0C3B1D4AC92BFA">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
</mods:name>
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<mods:namePart id="BD1F589E2A1A1392E1C7107F355E6831">Ledford, Joel</mods:namePart>
<mods:affiliation id="D74B7CB8FC69B4DF9C3E919CD1CAB509">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
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<mods:namePart id="BDEB101312820ADF7A310910E7320E43">Polotow, Daniele</mods:namePart>
<mods:affiliation id="4240BD6842CFC1E94D6BF99B62444E01">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
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<mods:title id="C4E9085E0B89913A2C311F6759A08BC7">Proceedings of the California Academy of Sciences</mods:title>
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<mods:date id="46C30B87B27524DC8D69B5CA32423BEB">2022</mods:date>
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<mods:number id="811C30DB30EAC77FDC5E7923C89F4984">2022-12-15</mods:number>
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<subSubSection id="3B99B1AE1875FFEB473432B2AA97EA98" box="[123,416,957,980]" pageId="49" pageNumber="50" type="nomenclature">
<paragraph id="733CE2251875FFEB473432B2AA97EA98" blockId="49.[123,493,957,1011]" box="[123,416,957,980]" pageId="49" pageNumber="50">
<heading id="287455491875FFEB473432B2AA97EA98" box="[123,416,957,980]" fontSize="10" level="2" pageId="49" pageNumber="50" reason="4">
<emphasis id="41F73E371875FFEB473432B2AA97EA98" bold="true" box="[123,416,957,980]" pageId="49" pageNumber="50">
<taxonomicName id="B48399A61875FFEB473432B2AA3AEA98" ID-CoL="C8KJJ" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,269,957,980]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="irwini" status="sp. nov.">
<emphasis id="41F73E371875FFEB473432B2AA3AEA98" bold="true" box="[123,269,957,980]" italics="true" pageId="49" pageNumber="50">Uduba irwini</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AC4834C1875FFEB465432B2AA97EA98" box="[283,416,957,980]" pageId="49" pageNumber="50" rank="species">new species</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="3B99B1AE1875FFEB473432D1AADAEABF" box="[123,493,990,1011]" pageId="49" pageNumber="50" type="description">
<paragraph id="733CE2251875FFEB473432D1AADAEABF" blockId="49.[123,493,957,1011]" box="[123,493,990,1011]" pageId="49" pageNumber="50">
<figureCitation id="EBB8FEA01875FFEB473432D1AA21EABF" box="[123,278,990,1011]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="49" pageNumber="50">Figures 46 AC</figureCitation>
,
<figureCitation id="EBB8FEA01875FFEB466F32D1AA58EABF" box="[288,367,990,1011]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="49" pageNumber="50">71 C, D</figureCitation>
,
<figureCitation id="EBB8FEA01875FFEB463532D1AAF4EABF" box="[378,451,990,1011]" captionStart="MAP" captionStartId="177.[155,174,1068,1087]" captionTargetBox="[129,1176,216,1047]" captionTargetId="figure-17@177.[648,1176,216,1059]" captionTargetPageId="177" captionText="MAP 3. Distributions of Uduba rinha and Uduba irwini. MAP 4. Distributions of Uduba madagascariensis, Uduba barbarae, Uduba pseudoevanescens and Uduba orona." figureDoi="http://doi.org/10.5281/zenodo.13158775" httpUri="https://zenodo.org/record/13158775/files/figure.png" pageId="49" pageNumber="50">Maps 3</figureCitation>
, 18.
</paragraph>
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<subSubSection id="3B99B1AE1875FFEB47E4350CAAFFED97" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="733CE2251875FFEB47E4350CAAFFED97" blockId="49.[123,1174,1027,1725]" pageId="49" pageNumber="50">
<emphasis id="41F73E371875FFEB47E4350CAA79ED56" bold="true" box="[171,334,1027,1050]" pageId="49" pageNumber="50">Type material.</emphasis>
<materialsCitation id="C3EBE8781875FFEB461C350CABD9ED36" collectingDate="2009-01-12" collectingDateMax="2009-01-22" collectingDateMin="2009-01-12" collectionCode="CAS" collectorName="M. Irwin &amp; R. Harin'Hala" country="Madagascar" elevation="1700" location="Ankokoy Forest" pageId="49" pageNumber="50" specimenCode="CASENT9062802" specimenCount="1" stateProvince="Antananarivo" typeStatus="holotype">
<typeStatus id="AC385C871875FFEB461C350CAA81ED56" box="[339,438,1027,1050]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
male (
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)
<collectingMethod id="AAC29A321875FFEB459D350CA8C5ED56" box="[722,1010,1027,1050]" pageId="49" pageNumber="50">collected from malaise trap</collectingMethod>
in
<taxonomicName id="B48399A61875FFEB435C350CAF50ED56" box="[1043,1127,1027,1050]" class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" order="Malpighiales" pageId="49" pageNumber="98" phylum="Tracheophyta" rank="genus">
<emphasis id="41F73E371875FFEB435C350CAF50ED56" box="[1043,1127,1027,1050]" italics="true" pageId="49" pageNumber="50">Uapaca</emphasis>
</taxonomicName>
forest at
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<elevation id="F8AE05161875FFEB47F4352CAA3AED76" box="[187,269,1059,1082]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="49" pageNumber="50" unit="m" value="1700.0">1700 m</elevation>
</quantity>
elevation in the
<location id="765CB4FE1875FFEB468F352CA959ED76" LSID="urn:lsid:plazi:treatment:FB2A53331875FFE9473432B2AFA2EF30:765CB4FE1875FFEB468F352CA959ED76" box="[448,622,1059,1082]" country="Madagascar" name="Ankokoy Forest" pageId="49" pageNumber="50" stateProvince="Antananarivo">Ankokoy Forest</location>
(20°04.05ʹS, 46°59.97ʹE),
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<quantity id="B47B4FC01875FFEB44DE352CA8F0ED76" box="[913,967,1059,1082]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="49" pageNumber="50" unit="km" value="3.0">3 km</quantity>
E of Ibity
</locationDeviation>
,
<collectingRegion id="B1472CC71875FFEB4370352CAA10ED16" country="Madagascar" name="Antananarivo" pageId="49" pageNumber="50">Antananarivo Province</collectingRegion>
,
<collectingCountry id="0B94A2B51875FFEB467F354CAA98ED16" box="[304,431,1091,1114]" name="Madagascar" pageId="49" pageNumber="50">Madagascar</collectingCountry>
,
<date id="073DC4E51875FFEB46F4354CA9A2ED16" box="[443,661,1091,1114]" pageId="49" pageNumber="50" value="2009-01-12" valueMax="2009-01-22" valueMin="2009-01-12">
<collectingDate id="17793D0D1875FFEB46F4354CA9A2ED16" box="[443,661,1091,1114]" pageId="49" pageNumber="50" value="2009-01-12" valueMax="2009-01-22" valueMin="2009-01-12">1222 January 2009</collectingDate>
</date>
, collectors
<collectorName id="DE7687F31875FFEB4443354CA85DED16" box="[780,874,1091,1114]" pageId="49" pageNumber="50">M. Irwin</collectorName>
and
<collectorName id="DE7687F31875FFEB44D2354CAF01ED16" box="[925,1078,1091,1114]" pageId="49" pageNumber="50">R. HarinHala</collectorName>
, deposit- ed in
<collectionCode id="15927AE01875FFEB47F8356CABD9ED36" box="[183,238,1123,1146]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="49" pageNumber="50" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3EBE8781875FFEB47B6356CAAF4ED97" collectingDate="2001-11-16" collectingDateMax="2001-11-20" collectingDateMin="2001-11-16" collectionCode="CAS" country="Madagascar" elevation="150" latitude="-19.709446" location="Parc National Tsingy de Bemaraha" longLatPrecision="21" longitude="44.718056" municipality="Fisher-Griswold Arthropod Team" pageId="49" pageNumber="50" specimenCode="CASENT9009383" specimenCount="1" stateProvince="Antananarivo" typeStatus="paratype">
<typeStatus id="AC385C871875FFEB47B6356CAA61ED36" box="[249,342,1123,1146]" pageId="49" pageNumber="50" type="paratype">Paratype</typeStatus>
female (
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) collected at
<quantity id="B47B4FC01875FFEB4442356CA87DED36" box="[781,842,1123,1146]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="49" pageNumber="50" unit="m" value="150.0">
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</quantity>
elevation in tropical dry forest on tsingy from
<location id="765CB4FE1875FFEB4661358BA980EDD7" LSID="urn:lsid:plazi:treatment:FB2A53331875FFE9473432B2AFA2EF30:765CB4FE1875FFEB4661358BA980EDD7" box="[302,695,1156,1179]" country="Madagascar" latitude="-19.709446" longLatPrecision="21" longitude="44.718056" municipality="Fisher-Griswold Arthropod Team" name="Parc National Tsingy de Bemaraha" pageId="49" pageNumber="50" stateProvince="Antananarivo">Parc National Tsingy de Bemaraha</location>
,
<locationDeviation id="628EFB071875FFEB4589358BAF36EDD7" box="[710,1025,1156,1179]" location="Antsalova" pageId="49" pageNumber="50">
<quantity id="B47B4FC01875FFEB4589358BA829EDD7" box="[710,798,1156,1179]" metricMagnitude="4" metricUnit="m" metricValue="1.06" pageId="49" pageNumber="50" unit="km" value="10.6">10.6 km</quantity>
123°ESE Antsalova
</locationDeviation>
(
<geoCoordinate id="16B784E21875FFEB435A358BAFA7EDD7" box="[1045,1168,1156,1179]" degrees="19" direction="south" minutes="42" orientation="latitude" pageId="49" pageNumber="50" precision="15" seconds="34" value="-19.709446">19°42ʹ34ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21875FFEB473435ABABCCEDF7" box="[123,251,1188,1211]" degrees="44" direction="east" minutes="43" orientation="longitude" pageId="49" pageNumber="50" precision="15" seconds="05" value="44.718056">44°43ʹ05ʺE</geoCoordinate>
), collected
<date id="073DC4E51875FFEB46DF35ABA992EDF7" box="[400,677,1188,1211]" pageId="49" pageNumber="50" value="2001-11-16" valueMax="2001-11-20" valueMin="2001-11-16">
<collectingDate id="17793D0D1875FFEB46DF35ABA992EDF7" box="[400,677,1188,1211]" pageId="49" pageNumber="50" value="2001-11-16" valueMax="2001-11-20" valueMin="2001-11-16">1620 November 2001</collectingDate>
</date>
by the
<collectingMunicipality id="9358785F1875FFEB445B35ABAFA2EDF7" box="[788,1173,1188,1211]" pageId="49" pageNumber="50">Fisher-Griswold Arthropod Team</collectingMunicipality>
[BLF4503], deposited in
<collectionCode id="15927AE01875FFEB46C335CBAAF4ED97" box="[396,451,1220,1243]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="49" pageNumber="50" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1875FFEB47E435EBAF43ECF0" pageId="49" pageNumber="50" type="discussion">
<paragraph id="733CE2251875FFEB47E435EBAF43ECF0" blockId="49.[123,1174,1027,1725]" pageId="49" pageNumber="50">
<emphasis id="41F73E371875FFEB47E435EBAA24EDB7" bold="true" box="[171,275,1252,1275]" pageId="49" pageNumber="50">Remarks</emphasis>
. Males of the similar species
<taxonomicName id="B48399A61875FFEB451B35EBA9D4EDB7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[596,739,1252,1275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371875FFEB451B35EBA9D4EDB7" bold="true" box="[596,739,1252,1275]" italics="true" pageId="49" pageNumber="50">Uduba rinha</emphasis>
</taxonomicName>
and
<taxonomicName id="B48399A61875FFEB445735EBA89DEDB7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[792,938,1252,1275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371875FFEB445735EBA89DEDB7" bold="true" box="[792,938,1252,1275]" italics="true" pageId="49" pageNumber="50">Uduba irwini</emphasis>
</taxonomicName>
may be distinguished by the TA2 form. Association of the males and females of
<taxonomicName id="B48399A61875FFEB45B4340BA8BDEC57" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[763,906,1284,1307]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371875FFEB45B4340BA8BDEC57" bold="true" box="[763,906,1284,1307]" italics="true" pageId="49" pageNumber="50">Uduba rinha</emphasis>
</taxonomicName>
is unproblematic in that several were collected together at Parc National de Namoroka. Females associated with
<taxonomicName id="B48399A61875FFEB4304342BAB8FEC17" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371875FFEB4304342BAB8FEC17" bold="true" italics="true" pageId="49" pageNumber="50">Uduba rinha</emphasis>
</taxonomicName>
have the epigynal plate posterior of the ML smooth or grooved and the CO hidden beneath the ML hood (
<figureCitation id="EBB8FEA01875FFEB4659346AAABBEC30" box="[278,396,1381,1404]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="49" pageNumber="50">Figs. 71 G</figureCitation>
,
<figureCitation id="EBB8FEA01875FFEB46D7346AAAFAEC30" box="[408,461,1381,1404]" captionStart="FIGURE" captionStartId="169.[155,167,782,801]" captionTargetBox="[120,1175,217,782]" captionTargetId="figure-19@169.[120,1176,216,770]" captionTargetPageId="169" captionText="FIGURE 81. Uduba spp., ♀ genitalia. A, D. U. fisheri, new species, CASENT9026388 from Anjozorobe. B, E. U. rinha, new species, CASENT9006108 from R.S. Manongarivo. C, F. U. ibonia, new species, CASENT9006033, from Talatakely, Ranomafana National Park. AC. Epigynum, ventral. DF. Vulva, dorsal. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; LL, lateral lobe of epigynum; ML, median lobe of epigynum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13158757" httpUri="https://zenodo.org/record/13158757/files/figure.png" pageId="49" pageNumber="50">81 B</figureCitation>
) whereas a female from Tsingy de Bemeraha (CASENT9009383) differs from these in that it has exposed copulatory openings just behind the ML (
<figureCitation id="EBB8FEA01875FFEB44A6348AAF64ECD0" box="[1001,1107,1413,1436]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="49" pageNumber="50">Fig. 71 C</figureCitation>
). This female is considered a
<typeStatus id="AC385C871875FFEB463B34AAAAE7ECF0" box="[372,464,1445,1468]" pageId="49" pageNumber="50" type="paratype">paratype</typeStatus>
of
<taxonomicName id="B48399A61875FFEB46B934AAA9BEECF0" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[502,649,1445,1468]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371875FFEB46B934AAA9BEECF0" bold="true" box="[502,649,1445,1468]" italics="true" pageId="49" pageNumber="50">Uduba irwini</emphasis>
</taxonomicName>
, which is otherwise known only from males.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1875FFEB47E434CAA92CEF50" pageId="49" pageNumber="50" type="etymology">
<paragraph id="733CE2251875FFEB47E434CAA92CEF50" blockId="49.[123,1174,1027,1725]" pageId="49" pageNumber="50">
<emphasis id="41F73E371875FFEB47E434CAAA1DEC90" bold="true" box="[171,298,1477,1500]" pageId="49" pageNumber="50">Etymology.</emphasis>
The species epithet is a patronym on honor of Dr. Mike Irwin, entomologist and philanthropist and a principal architect of the malaise trap survey of
<collectingCountry id="0B94A2B51875FFEB440134EAA8F8ECB0" box="[846,975,1509,1532]" name="Madagascar" pageId="49" pageNumber="50">Madagascar</collectingCountry>
arthropods, which revealed many new species of spiders.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1875FFE847E4372AA9B5E81F" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="diagnosis">
<paragraph id="733CE2251875FFE847E4372AA9B5E81F" blockId="49.[123,1174,1027,1725]" lastBlockId="50.[122,1175,220,1725]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
<emphasis id="41F73E371875FFEB47E4372AAA8BEF70" bold="true" box="[171,444,1573,1596]" pageId="49" pageNumber="50">
Diagnosis.
<taxonomicName id="B48399A61875FFEB4667372AAA8BEF70" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[296,444,1573,1596]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371875FFEB4667372AAA8BEF70" bold="true" box="[296,444,1573,1596]" italics="true" pageId="49" pageNumber="50">Uduba irwini</emphasis>
</taxonomicName>
</emphasis>
are members of Group IV, the
<taxonomicName id="B48399A61875FFEB445C372AA8ADEF70" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[787,922,1573,1596]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rinha">Uduba rinha</taxonomicName>
group (Map 18). They have a divided cribellum. Males can be distinguished from those of other
<taxonomicName id="B48399A61875FFEB44E63749A8C4EF11" authorityName="Simon" authorityYear="1880" box="[937,1011,1606,1629]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371875FFEB44E63749A8C4EF11" bold="true" box="[937,1011,1606,1629]" italics="true" pageId="49" pageNumber="50">Uduba</emphasis>
</taxonomicName>
except
<taxonomicName id="B48399A61875FFEB43043749AB8FEF31" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371875FFEB43043749AB8FEF31" bold="true" italics="true" pageId="49" pageNumber="50">Uduba rinha</emphasis>
</taxonomicName>
by the simple, trapezoidal MA (
<figureCitation id="EBB8FEA01875FFEB456B3769A984EF31" box="[548,691,1638,1661]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="49" pageNumber="50">Figs 46 AC</figureCitation>
<emphasis id="41F73E371875FFEB45FC3769A98BEF31" bold="true" box="[691,700,1638,1661]" pageId="49" pageNumber="50">)</emphasis>
;
<taxonomicName id="B48399A61875FFEB45823769A854EF31" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[717,867,1638,1661]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371875FFEB45823769A854EF31" bold="true" box="[717,867,1638,1661]" italics="true" pageId="49" pageNumber="50">Uduba irwini</emphasis>
</taxonomicName>
may be distinguished from
<taxonomicName id="B48399A61875FFEB47343789AA3EEFD1" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,265,1670,1693]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371875FFEB47343789AA3EEFD1" bold="true" box="[123,265,1670,1693]" italics="true" pageId="49" pageNumber="50">Uduba rinha</emphasis>
</taxonomicName>
by having the TA2 an erect, rectangular spike (
<figureCitation id="EBB8FEA01875FFEB444B3789A85DEFD1" box="[772,874,1670,1693]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="49" pageNumber="50">Fig. 46 B</figureCitation>
) (
<taxonomicName id="B48399A61875FFEB44CD3789AF26EFD1" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[898,1041,1670,1693]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371875FFEB44CD3789AF26EFD1" bold="true" box="[898,1041,1670,1693]" italics="true" pageId="49" pageNumber="50">Uduba rinha</emphasis>
</taxonomicName>
has the TA2 a flexible hook or curlicue,
<figureCitation id="EBB8FEA01875FFEB46E937A9A939EFF1" box="[422,526,1702,1725]" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="49" pageNumber="50">Fig. 59 B</figureCitation>
). The females (
<figureCitation id="EBB8FEA01875FFEB45FA37A9A82BEFF1" box="[693,796,1702,1725]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="49" pageNumber="50">Fig. 71 C</figureCitation>
<emphasis id="41F73E371875FFEB445437A9A813EFF1" bold="true" box="[795,804,1702,1725]" pageId="49" pageNumber="50">)</emphasis>
and
<taxonomicName id="B48399A61875FFEB441537A9A8DFEFF1" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[858,1000,1702,1725]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371875FFEB441537A9A8DFEFF1" bold="true" box="[858,1000,1702,1725]" italics="true" pageId="49" pageNumber="50">Uduba rinha</emphasis>
</taxonomicName>
(
<figureCitation id="EBB8FEA01875FFEB44B737A9AF57EFF1" box="[1016,1120,1702,1725]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="49" pageNumber="50">Fig. 71 G</figureCitation>
<emphasis id="41F73E371875FFEB432F37A9AF5EEFF1" bold="true" box="[1120,1129,1702,1725]" pageId="49" pageNumber="50">)</emphasis>
can be distinguished from those of other cribellate
<taxonomicName id="B48399A61876FFE8453C31D3A98AE9BF" authorityName="Simon" authorityYear="1880" box="[627,701,220,243]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371876FFE8453C31D3A98AE9BF" bold="true" box="[627,701,220,243]" italics="true" pageId="50" pageNumber="51">Uduba</emphasis>
</taxonomicName>
by having the epigynum lateral lobes fused medially; the lone female of
<taxonomicName id="B48399A61876FFE846FA31F3A97FE85F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[437,584,252,275]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371876FFE846FA31F3A97FE85F" bold="true" box="[437,584,252,275]" italics="true" pageId="50" pageNumber="51">Uduba irwini</emphasis>
</taxonomicName>
has the copulatory openings exposed just posteriad of the ML (
<figureCitation id="EBB8FEA01876FFE8479A3013AA0EE87F" box="[213,313,284,307]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="50" pageNumber="51">Fig. 71 C</figureCitation>
<emphasis id="41F73E371876FFE846763013AA75E87F" bold="true" box="[313,322,284,307]" pageId="50" pageNumber="51">)</emphasis>
whereas the females of
<taxonomicName id="B48399A61876FFE8450F3013A9FAE87F" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[576,717,284,307]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371876FFE8450F3013A9FAE87F" bold="true" box="[576,717,284,307]" italics="true" pageId="50" pageNumber="51">Uduba rinha</emphasis>
</taxonomicName>
have the copulatory openings that are each hidden beneath the sides of the ML (
<figureCitation id="EBB8FEA01876FFE845463033A941E81F" box="[521,630,316,339]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="50" pageNumber="51">Fig. 71 G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1876FFE947E43053A933EF70" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51" type="description">
<paragraph id="733CE2251876FFE947E43053AAEAE898" blockId="50.[122,1175,220,1725]" lastBlockId="51.[123,1175,220,1660]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
<emphasis id="41F73E371876FFE847E43053AA40E83F" bold="true" box="[171,375,348,371]" pageId="50" pageNumber="51">Description. Male</emphasis>
(
<typeStatus id="AC385C871876FFE846C93053AAC7E83F" box="[390,496,348,371]" pageId="50" pageNumber="51" type="holotype">Holotype</typeStatus>
): Total length 9.70. Markings typical of
<taxonomicName id="B48399A61876FFE844EC3053A8DAE83F" authorityName="Simon" authorityYear="1880" box="[931,1005,348,371]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371876FFE844EC3053A8DAE83F" bold="true" box="[931,1005,348,371]" italics="true" pageId="50" pageNumber="51">Uduba</emphasis>
</taxonomicName>
. Carapace 5.50 long, 3.60 wide, 2.10 high; clypeus 0.30 high. Eye diameters: AME 0.14, ALE 0.16, PME 0.18, PLE 0.20. Chelicerae 2.30 long; sternum 2.40 long, 1.80 wide; labium 1.10 long; palpal coxae 1.60 long. Cribellum divided. Spination. (ANJA reserve, CASENT9065405) palpusfemur d1-0-3; leg Ifemur d1-1-1, p0-1-2, r0-1-1-0, tibia d0-1-1, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus d0-1-0, p0- 1-2, v2-2-1, r1-1-1; leg IIfemur d1-1-1, p0-1-2, r0-1-0-0, tibia d0-1-1-0, p0-1-1-0, v2-2-2, r0-1-1- 0, metatarsus p1-1-2, v2-2-1, r1-1-2; leg IIIfemur d1-1-1, p0-1-0-2, r0-1-0-2, tibia d0-1-0, p0-1- 1-0, v2-1-2, r0-1-1-0, metatarsus p1-1-2, v2-2-2, r1-1-2, tarsus v0-1-0; leg IVfemur d1-1-1, p0-1- 1-0, r0-0-0-1, patella r1, tibia d1-0-1-0, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus d0-1-0, p1-1-2, v2- 1-2-2, r1-1-2. Scopulae: cymbium, apicodorsal; tarsi IIV, ventral. Leg measurements (
<typeStatus id="AC385C871876FFE843683371AFBCEBD9" box="[1063,1163,638,661]" pageId="50" pageNumber="51" type="holotype">holotype</typeStatus>
): I: 4.40 + 1.90 + 4.00 + 4.10 + 2.60 = 17.00; II: 3.90 + 1.80 + 3.20 + 3.40 + 2.20 = 14.50; III: 3.40 + 1.50 + 2.10 + 2.70 + 1.70 = 11.40; IV: 4.20 + 1.70 + 3.80 + 4.20 + 2.40 = 16.30; palpus: 2.00 + 0.70 + 0.80 + NA + 2.50 = 6.00. Leg formula 1423.
<emphasis id="41F73E371876FFE8440733D0A8FFEBBA" bold="true" box="[840,968,735,758]" pageId="50" pageNumber="51">Male palp</emphasis>
(ANJA Reserve, CASENT9065405) (
<figureCitation id="EBB8FEA01876FFE8461633F0AAC7EA5A" box="[345,496,767,790]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="50" pageNumber="51">Figs. 46 AC</figureCitation>
): palpal tibia 0.35 cymbial length, RTA long, straight (
<figureCitation id="EBB8FEA01876FFE8430933F0ABB8EA7B" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="50" pageNumber="51">Fig. 46 C</figureCitation>
), RTA length 0.33 times tibia length, length 2.25 times width, with sharply-pointed apex, VTA short, length 1.50 times width, conical, blunt (
<figureCitation id="EBB8FEA01876FFE8453D324FA830EA1B" box="[626,775,832,855]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="50" pageNumber="51">Figs. 46 A, C</figureCitation>
), length 0.52 times tibia width; tibia without stout spines; tegulum convex, without ridge, tegulum length 1.09 times width; TA1 slen- der, extends far distad of tegulum apex, TA1 extends past TA3 apex by 1.38 times tegulum length; TA3 a transverse, blunt hook (
<figureCitation id="EBB8FEA01876FFE846F632AFA929EAFB" box="[441,542,928,951]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="50" pageNumber="51">Fig. 46 B</figureCitation>
<emphasis id="41F73E371876FFE8455232AFA911EAFB" bold="true" box="[541,550,928,951]" pageId="50" pageNumber="51">)</emphasis>
, TA3 extends apicad of TA2 by 1.21 times tegulum length, TA2 a small, erect trapezoid with concave apex, stiff, sclerotized, extends partially in front of TA3 (
<figureCitation id="EBB8FEA01876FFE847CC32EEABD0EAB4" box="[131,231,993,1016]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="50" pageNumber="51">Fig. 46 B</figureCitation>
<emphasis id="41F73E371876FFE847A832EEABC7EAB4" bold="true" box="[231,240,993,1016]" pageId="50" pageNumber="51">)</emphasis>
; MA a broad, flat trapezoid, origin at 0.54 times tegulum length, extends nearly to tegulum apex, tegulum apex at 0.085 tegulum length from MA apex, MA length (including apical processes) 1.22 times width, MA large (but smaller than in
<taxonomicName id="B48399A61876FFE845B7352EA861ED74" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[760,854,1057,1080]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rinha">
<emphasis id="41F73E371876FFE845B7352EA861ED74" bold="true" box="[760,854,1057,1080]" italics="true" pageId="50" pageNumber="51">U. rinha</emphasis>
</taxonomicName>
,
<figureCitation id="EBB8FEA01876FFE8442B352EA8FDED74" box="[868,970,1057,1080]" captionStart="FIGURE" captionStartId="151.[155,167,874,893]" captionTargetBox="[140,1162,236,856]" captionTargetId="figure-16@151.[120,1176,216,871]" captionTargetPageId="151" captionText="FIGURE 59. Uduba rinha, new species, (MRAC174.445) from 25 km W Morarano Chrome, Toamasina Province, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Jenny Speckels. MA, median apophysis; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158705" httpUri="https://zenodo.org/record/13158705/files/figure.png" pageId="50" pageNumber="51">Fig. 59 A</figureCitation>
), length 0.31 tegulum length, width 0.29 tegulum width (
<figureCitation id="EBB8FEA01876FFE84568354DA98DED15" box="[551,698,1090,1113]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="50" pageNumber="51">Figs. 46 AC</figureCitation>
); conductor fan elongate, entire (
<figureCitation id="EBB8FEA01876FFE84350354DAFB1ED15" box="[1055,1158,1090,1113]" captionStart="FIGURE" captionStartId="142.[155,167,954,973]" captionTargetBox="[147,1157,225,932]" captionTargetId="figure-11@142.[120,1176,216,940]" captionTargetPageId="142" captionText="FIGURE 46. Uduba irwini, new species, (CASENT9065405) from ANJA Reserve, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. Illustrations by Rachel Diaz-Bastin. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158677" httpUri="https://zenodo.org/record/13158677/files/figure.png" pageId="50" pageNumber="51">Fig. 46 C</figureCitation>
<emphasis id="41F73E371876FFE843C9354DAFB8ED15" bold="true" box="[1158,1167,1090,1113]" pageId="50" pageNumber="51">)</emphasis>
.
<emphasis id="41F73E371876FFE84734356DABF9ED35" bold="true" box="[123,206,1122,1145]" pageId="50" pageNumber="51">Female</emphasis>
(
<typeStatus id="AC385C871876FFE84795356DAA09ED35" box="[218,318,1122,1145]" pageId="50" pageNumber="51" type="paratype">Paratype</typeStatus>
): Total length 11.56. Markings typical for
<taxonomicName id="B48399A61876FFE845B9356DA877ED35" authorityName="Simon" authorityYear="1880" box="[758,832,1122,1145]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371876FFE845B9356DA877ED35" bold="true" box="[758,832,1122,1145]" italics="true" pageId="50" pageNumber="51">Uduba</emphasis>
</taxonomicName>
. Carapace 5.60 long, 3.80 wide, 2.30 high; clypeus 0.30 high. Eye diameters: AME 0.16, ALE and PME 0.22, PLE 0.24. Chelicerae 2.70 long; sternum 2.40 long, 1.90 wide; labium 1.30 long; palpal coxae 1.90 long.
<emphasis id="41F73E371876FFE8434535ADAF57EDF5" bold="true" box="[1034,1120,1186,1209]" pageId="50" pageNumber="51">Ratios</emphasis>
carapace length / width = 1.47, carapace height / width = 0.61, PER / carapace width = 0.42, PER / OAL = 3.20, PER / AER = 1.36, OAL / OQL = 1.09, OQP / OQA = 1.38, clypeus height / AME diameter width = 1.88, cheliceral length / clypeus height = 9.00, sternum length / width = 1.26, palpal coxa length / width = 2.38, femur I length / carapace width = 1.00, metatarsus I length / carapace width = 0.79, femur IV length / carapace width = 1.03, palpal tarsus length / carapace width = 0.45, palpal tarsus length / palpal femur length = 0.89. Cribellum divided. Spination: Palpusfemur d0-1-2, patella p1, tibia p2-0, tarsus p2-0, v0-0-2; leg Ifemur d1-0-0, p0-0-1, tibia v2-2-2, metatarsus v2-2-3; leg IIfemur d1-0-0, p0-0-1, tibia p0-0-1, v1-1-2, metatarsus v2-2-3; leg IIIfemur p0-1-0-0, r0-1-0-0, tibia d1-0-1-0, p0-1-1-0, v1-0-2, r0-1-1-0, metatarsus p1-2-2, v2-2-2, r1-1-2, tarsus v0-1-0; leg IVfemur d1-1-0-0-0, tibia v2-1-2, r0-1-1-0, metatarsus p0-1-2, v1-2-2, r0-2-2. Scopulae: strong ventral on all tarsi, most of metatarsi I and II, and apices of tarsi III and IV. Leg measurements: I: 3.80 + 1.90 + 3.30 + 3.00 + 1.90 = 13.90; II: 3.20 + 1.60 + 2.60 + 2.30 + 1.80 = 11.50; III: 2.60 + 1.50 + 1.30 + 2.10 + 1.40 = 8.90; IV: 3.90 + 1.80 + 2.90 + 3.40 + 1.70 = 13.70; palpus: 1.90 + 1.00 + 1.00 + NA + 1.70 = 5.69. Leg formula 1423.
<emphasis id="41F73E371876FFE847F23789AA4EEFD1" bold="true" box="[189,377,1670,1693]" pageId="50" pageNumber="51">Female genitalia</emphasis>
(
<figureCitation id="EBB8FEA01876FFE846C83789A92CEFD1" box="[391,539,1670,1693]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="50" pageNumber="51">Figs. 71 C, D</figureCitation>
): epigynal plate without depressed atrium, plate width 1.35 times length; median lobe and lateral lobes differentiated only in anterior half of plate, distance from posterior margin of ML to epigastric furrow 0.35 times epigynum length; median lobe a flat plate with lateral margins extending around to posterior, broad, median lobe width 0.63 times epigynum width, 0.45 times ML length, ML width 2.64 times LL width at sides of ML (
<figureCitation id="EBB8FEA01877FFE943553013AFBFE87F" box="[1050,1160,284,307]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="51" pageNumber="52">Fig. 71 C</figureCitation>
); with copulatory openings exposed on epigynal plate beneath a transverse, crescentic groove on each side just behind ML, distance between copulatory openings 0.26 times epigynum width, CO origin at 0.35 of epigynum length. Vulva (
<figureCitation id="EBB8FEA01877FFE945003072A98FE8D8" box="[591,696,381,404]" captionStart="FIGURE" captionStartId="160.[155,167,808,827]" captionTargetBox="[133,1161,222,781]" captionTargetId="figure-11@160.[120,1176,216,793]" captionTargetPageId="160" captionText="FIGURE 71. Uduba spp., female genitalia drawings. A, B. U. pseudoevanescens, new species, holotype (CASENT9006008) from Périnet. C, D. U. irwini, new species, Paratype (CASENT9009383) from Parc National Tsingy de Bemaraha. E, F. U. evanescens (Dahl, 1901) (CASENT9024032) from Parc National Ranomafana. G, H. U. rinha, new species, paratype (CASENT9018660), Parc National de Namoroka. A, C, E, G. Epigynum, ventral. B, D, F, H. Vulva, dorsal. Illustrations by Charles Griswold. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158733" httpUri="https://zenodo.org/record/13158733/files/figure.png" pageId="51" pageNumber="52">Fig. 71 D</figureCitation>
) with spermathecal ducts forming three longitudinal loops, vulva length 0.64 times width, fertilization ducts widely separated, vulva width 3.57 times distance between FD.
</paragraph>
<paragraph id="733CE2251877FFE947E430D2A9B4EA5A" blockId="51.[123,1175,220,1660]" pageId="51" pageNumber="52">
<emphasis id="41F73E371877FFE947E430D2AA2DE8B8" bold="true" box="[171,282,477,500]" pageId="51" pageNumber="52">Variation.</emphasis>
Male (N= 7): Total length 9.7011.50; carapace length / width = 1.331.53, carapace height / width = 0.400.59, PER / carapace width = 0.340.38, PER / OAL = 2.503.24, PER / AER = 1.371.42, OAL / OQL = 1.051.26, OQP / OQA = 1.181.25, clypeus height / AME 1.11 2.14, cheliceral length / clypeus height = 7.6711.00, sternum length / width = 1.251.39, palpal coxa length / width = 2.252.67, femur I length / carapace width =1.121.24, metatarsus I length / carapace width = 1.091.28, femur IV length / carapace width = 1.151.26, palpal tarsus length / carapace width = 0.640.74, palpal tarsus length / palpal patella length = 3.333.75, palpal tarsus length / palpal tibia length = 3.003.38, palpal tarsus length / palpal femur length = 1.171.33, palpal tibia length / palpal patella length = 1.001.25. Female variation is unknown: only
<specimenCount id="658529AC1877FFE9435633D0AFA6EBBA" box="[1049,1169,735,758]" count="1" pageId="51" pageNumber="52" type="female">one female</specimenCount>
, the
<typeStatus id="AC385C871877FFE947EC33F0ABC8EA5A" box="[163,255,767,790]" pageId="51" pageNumber="52" type="paratype">paratype</typeStatus>
(CASENT9009383), is recognized.
</paragraph>
<paragraph id="733CE2251877FFE947E4322FABEFEC37" blockId="51.[123,1175,220,1660]" pageId="51" pageNumber="52">
M
<emphasis id="41F73E371877FFE9478D322FAAA5EA7B" bold="true" box="[194,402,800,823]" pageId="51" pageNumber="52">aterial examined.</emphasis>
<materialsCitation id="C3EBE8781877FFE946ED322FABDBEADB" collectingDate="2009-01-12" collectingDateMax="2009-01-22" collectingDateMin="2009-01-12" collectionCode="CAS" collectorName="M. Irwin &amp; R. Harin'Hala" country="Madagascar" elevation="1700" location="Ankokoy Forest" pageId="51" pageNumber="52" specimenCode="CASENT9062802" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo" typeStatus="holotype">
<collectingCountry id="0B94A2B51877FFE946ED322FA953EA7B" box="[418,612,800,823]" name="Madagascar" pageId="51" pageNumber="52">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E371877FFE94538322FA84EEA7B" box="[631,889,800,823]" italics="true" pageId="51" pageNumber="52">
<collectingRegion id="B1472CC71877FFE94538322FA84EEA7B" box="[631,889,800,823]" country="Madagascar" name="Antananarivo" pageId="51" pageNumber="52">Antananarivo Province</collectingRegion>
</emphasis>
:
<collectingRegion id="B1472CC71877FFE944C1322FAFA5EA7B" box="[910,1170,800,823]" country="Madagascar" name="Antananarivo" pageId="51" pageNumber="52">Antananarivo Province</collectingRegion>
,
<location id="765CB4FE1877FFE94734324FAA1BEA1B" LSID="urn:lsid:plazi:treatment:FB2A53331875FFE9473432B2AFA2EF30:765CB4FE1877FFE94734324FAA1BEA1B" box="[123,300,832,855]" country="Madagascar" name="Ankokoy Forest" pageId="51" pageNumber="52" stateProvince="Antananarivo">Ankokoy Forest</location>
,
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<quantity id="B47B4FC01877FFE94676324FAA47EA1B" box="[313,368,832,855]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="51" pageNumber="52" unit="km" value="3.0">3 km</quantity>
E of Ibity
</locationDeviation>
, 20°04.05ʹS, 46°59.97ʹE, elev.
<quantity id="B47B4FC01877FFE9447B324FA8B4EA1B" box="[820,899,832,855]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="51" pageNumber="52" unit="m" value="1700.0">
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</quantity>
, malaise, in
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<emphasis id="41F73E371877FFE9435F324FAF53EA1B" box="[1040,1124,832,855]" italics="true" pageId="51" pageNumber="52">Uapaca</emphasis>
</taxonomicName>
forest,
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<collectingDate id="17793D0D1877FFE947E8326FAAB2EA3B" box="[167,389,864,887]" pageId="51" pageNumber="52" value="2009-01-12" valueMax="2009-01-22" valueMin="2009-01-12">1222 January 2009</collectingDate>
</date>
,
<collectorName id="DE7687F31877FFE946DE326FAAC5EA3B" box="[401,498,864,887]" pageId="51" pageNumber="52">M. Irwin</collectorName>
and
<collectorName id="DE7687F31877FFE94567326FA9F6EA3B" box="[552,705,864,887]" pageId="51" pageNumber="52">R. HarinHala</collectorName>
[MG-56-08] (
<typeStatus id="AC385C871877FFE94413326FA889EA3B" box="[860,958,864,887]" pageId="51" pageNumber="52" type="holotype">holotype</typeStatus>
,
<specimenCode id="23254A5E1877FFE94485326FAFA5EA3B" box="[970,1170,864,887]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9062802</specimenCode>
,
<specimenCount id="658529AC1877FFE94734328FABABEADB" box="[123,156,896,919]" count="1" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01877FFE947E3328FABD3EADB" box="[172,228,896,919]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="51" pageNumber="52" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781877FFE947B5328FABE3ED54" collectingDate="1997-12-07" collectingDateMax="1997-12-12" collectingDateMin="1997-12-07" collectionCode="FMNH" collectorName="S. Goodman" country="Madagascar" elevation="1450" location="Reserve Speciale d'Ambohitantely" pageId="51" pageNumber="52" specimenCode="CASENT9016347, CASENT9016348, CASENT9006222, CASENT9016346, CASENT9016349" specimenCount="7" specimenCount-male="7" stateProvince="Antananarivo" typeStatus="holotype">
<location id="765CB4FE1877FFE947B5328FA947EADB" LSID="urn:lsid:plazi:treatment:FB2A53331875FFE9473432B2AFA2EF30:765CB4FE1877FFE947B5328FA947EADB" box="[250,624,896,919]" country="Madagascar" name="Reserve Speciale d'Ambohitantely" pageId="51" pageNumber="52" stateProvince="Antananarivo">Réserve Spéciale dAmbohitantely</location>
,
<locationDeviation id="628EFB071877FFE94530328FA851EADB" box="[639,870,896,919]" location="Ankazobe" pageId="51" pageNumber="52">
<quantity id="B47B4FC01877FFE94530328FA9F4EADB" box="[639,707,896,919]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="51" pageNumber="52" unit="km" value="24.0">24 km</quantity>
NE Ankazobe
</locationDeviation>
, 18°10.1ʹS, 47°16.6ʹE, elev.
<quantity id="B47B4FC01877FFE9473432AEABF0EAF4" box="[123,199,929,952]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="51" pageNumber="52" unit="m" value="1450.0">
<elevation id="F8AE05161877FFE9473432AEABF0EAF4" box="[123,199,929,952]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="51" pageNumber="52" unit="m" value="1450.0">1450m</elevation>
</quantity>
, disturbed transitional montane mossy forest, 30-hectare parcel,
<collectingMethod id="AAC29A321877FFE944DD32AEAF27EAF4" box="[914,1040,929,952]" pageId="51" pageNumber="52">pitfall traps</collectingMethod>
#13,
<date id="073DC4E51877FFE9432F32AEAA10EA94" pageId="51" pageNumber="52" value="1997-12-07" valueMax="1997-12-12" valueMin="1997-12-07">
<collectingDate id="17793D0D1877FFE9432F32AEAA10EA94" pageId="51" pageNumber="52" value="1997-12-07" valueMax="1997-12-12" valueMin="1997-12-07">712 December 1997</collectingDate>
</date>
,
<collectorName id="DE7687F31877FFE9467D32CEAA8FEA94" box="[306,440,961,984]" pageId="51" pageNumber="52">S. Goodman</collectorName>
(
<specimenCode id="23254A5E1877FFE9468832CEA9A7EA94" box="[455,656,961,984]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9016347</specimenCode>
,
<specimenCount id="658529AC1877FFE945D432CEA98BEA94" box="[667,700,961,984]" count="1" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01877FFE9458532CEA82CEA94" box="[714,795,961,984]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="51" pageNumber="52" type="Museum">FMNH</collectionCode>
), (
<specimenCode id="23254A5E1877FFE9447A32CEA8C9EA94" box="[821,1022,961,984]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9016348</specimenCode>
,
<specimenCount id="658529AC1877FFE9434532CEAF1CEA94" box="[1034,1067,961,984]" count="1" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01877FFE9437632CEAFBFEA94" box="[1081,1160,961,984]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="51" pageNumber="52" type="Museum">FMNH</collectionCode>
), (
<specimenCode id="23254A5E1877FFE947CD32EEAA7BEAB4" box="[130,332,993,1016]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9006222</specimenCode>
,
<specimenCount id="658529AC1877FFE9461032EEAAB7EAB4" box="[351,384,993,1016]" count="1" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01877FFE946D932EEAAD0EAB4" box="[406,487,993,1016]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="51" pageNumber="52" type="Museum">FMNH</collectionCode>
), (
<specimenCode id="23254A5E1877FFE9454632EEA9E4EAB4" box="[521,723,993,1016]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9016346</specimenCode>
,
<specimenCount id="658529AC1877FFE945A932EEA830EAB4" box="[742,775,993,1016]" count="1" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01877FFE9445232EEA859EAB4" box="[797,878,993,1016]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="51" pageNumber="52" type="Museum">FMNH</collectionCode>
), (
<specimenCode id="23254A5E1877FFE944DE32EEAF6DEAB4" box="[913,1114,993,1016]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9016349</specimenCode>
,
<specimenCount id="658529AC1877FFE9432232EEAFB8EAB4" box="[1133,1167,993,1016]" count="3" pageId="51" pageNumber="52" type="male">3♂</specimenCount>
,
<collectionCode id="15927AE01877FFE94734350EABFAED54" box="[123,205,1025,1048]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="51" pageNumber="52" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE8781877FFE947AE350EAF58ED15" collectingDate="2014-12-10" collectingDateMax="2014-12-19" collectingDateMin="2014-12-10" collectionCode="CAS" collectorName="M. Irwin &amp; R. Harin'Hala" country="Madagascar" county="Haute Matsiatra Region" elevation="1097" location="Ambalavao" municipality="Reserve" pageId="51" pageNumber="52" specimenCode="CASENT9065491" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="41F73E371877FFE947AE350EAAEDED54" box="[225,474,1025,1048]" italics="true" pageId="51" pageNumber="52">
<collectingRegion id="B1472CC71877FFE947AE350EAAEDED54" box="[225,474,1025,1048]" country="Madagascar" name="Fianarantsoa" pageId="51" pageNumber="52">Fianarantsoa Province</collectingRegion>
</emphasis>
:
<collectingCounty id="9A5D9AA91877FFE946A5350EA9C4ED54" box="[490,755,1025,1048]" pageId="51" pageNumber="52">Haute Matsiatra Region</collectingCounty>
, ANJA
<collectingMunicipality id="9358785F1877FFE94406350EA895ED54" box="[841,930,1025,1048]" pageId="51" pageNumber="52">Reserve</collectingMunicipality>
,
<quantity id="B47B4FC01877FFE944E0350EA8C3ED54" box="[943,1012,1025,1048]" metricMagnitude="4" metricUnit="m" metricValue="6.1" pageId="51" pageNumber="52" unit="km" value="61.0">61 km</quantity>
S
<location id="765CB4FE1877FFE9435B350EAFA5ED54" LSID="urn:lsid:plazi:treatment:FB2A53331875FFE9473432B2AFA2EF30:765CB4FE1877FFE9435B350EAFA5ED54" box="[1044,1170,1025,1048]" country="Madagascar" county="Haute Matsiatra Region" municipality="Reserve" name="Ambalavao" pageId="51" pageNumber="52" stateProvince="Fianarantsoa">Ambalavao</location>
,
<locationDeviation id="628EFB071877FFE94734352EAABFED74" box="[123,392,1057,1080]" location="Fianarantsoa" pageId="51" pageNumber="52">
<quantity id="B47B4FC01877FFE94734352EAB88ED74" box="[123,191,1057,1080]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="51" pageNumber="52" unit="km" value="13.0">13 km</quantity>
S of
<collectingRegion id="B1472CC71877FFE947B2352EAABFED74" box="[253,392,1057,1080]" country="Madagascar" name="Fianarantsoa" pageId="51" pageNumber="52">Fianarantsoa</collectingRegion>
</locationDeviation>
, 22°43.23ʹS, 46°15.47ʹE, elev.
<quantity id="B47B4FC01877FFE94597352EA812ED74" box="[728,805,1057,1080]" metricMagnitude="3" metricUnit="m" metricValue="1.097" pageId="51" pageNumber="52" unit="m" value="1097.0">
<elevation id="F8AE05161877FFE94597352EA812ED74" box="[728,805,1057,1080]" metricMagnitude="3" metricUnit="m" metricValue="1.097" pageId="51" pageNumber="52" unit="m" value="1097.0">1097m</elevation>
</quantity>
, malaise, isleberg vegetation,
<date id="073DC4E51877FFE94322352EAA7EED15" pageId="51" pageNumber="52" value="2014-12-10" valueMax="2014-12-19" valueMin="2014-12-10">
<collectingDate id="17793D0D1877FFE94322352EAA7EED15" pageId="51" pageNumber="52" value="2014-12-10" valueMax="2014-12-19" valueMin="2014-12-10">10 19 December 2014</collectingDate>
</date>
,
<collectorName id="DE7687F31877FFE9461A354DAA83ED15" box="[341,436,1090,1113]" pageId="51" pageNumber="52">M. Irwin</collectorName>
and
<collectorName id="DE7687F31877FFE946A6354DA9B6ED15" box="[489,641,1090,1113]" pageId="51" pageNumber="52">R. HarinHala</collectorName>
[MG-64B-95] (
<specimenCode id="23254A5E1877FFE94463354DA8C2ED15" box="[812,1013,1090,1113]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9065491</specimenCode>
,
<specimenCount id="658529AC1877FFE9434E354DAF15ED15" box="[1025,1058,1090,1113]" count="1" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01877FFE9437F354DAF50ED15" box="[1072,1127,1090,1113]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="51" pageNumber="52" type="Museum">CAS</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="C3EBE8781877FFE94334354DAFA5ED35" collectingDate="2013-12-05" collectingDateMax="2013-12-19" collectingDateMin="2013-12-05" collectionCode="CAS" collectorName="M. Irwin &amp; Harin'Hala" country="Madagascar" pageId="51" pageNumber="52" specimenCode="CASENT9065405" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<date id="073DC4E51877FFE94334354DAA79ED35" pageId="51" pageNumber="52" value="2013-12-05" valueMax="2013-12-19" valueMin="2013-12-05">
<collectingDate id="17793D0D1877FFE94334354DAA79ED35" pageId="51" pageNumber="52" value="2013-12-05" valueMax="2013-12-19" valueMin="2013-12-05">5 19 December 2013</collectingDate>
</date>
,
<collectorName id="DE7687F31877FFE94613356DAA89ED35" box="[348,446,1122,1145]" pageId="51" pageNumber="52">M. Irwin</collectorName>
and R.
<collectorName id="DE7687F31877FFE94556356DA9A6ED35" box="[537,657,1122,1145]" pageId="51" pageNumber="52">HarinHala</collectorName>
{MG-64B-63} (
<specimenCode id="23254A5E1877FFE94406356DAF24ED35" box="[841,1043,1122,1145]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9065405</specimenCode>
,
<specimenCount id="658529AC1877FFE9436F356DAF75ED35" box="[1056,1090,1122,1145]" count="1" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01877FFE9431D356DAFBDED35" box="[1106,1162,1122,1145]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="51" pageNumber="52" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE8781877FFE94734358DAAF0EDB6" collectingDate="2001-11-16" collectingDateMax="2001-11-20" collectingDateMin="2001-11-16" collectionCode="CAS" collectorName="Fisher-Griswold Arthropod Team" country="Madagascar" elevation="150" latitude="-19.709446" location="Antsalova" longLatPrecision="21" longitude="44.718056" municipality="Parc National Tsingy de Bemaraha" pageId="51" pageNumber="52" specimenCode="CASENT9009383" specimenCount="1" specimenCount-female="1" stateProvince="Mahajanga" typeStatus="paratype">
<emphasis id="41F73E371877FFE94734358DAA52EDD5" box="[123,357,1154,1177]" italics="true" pageId="51" pageNumber="52">
<collectingRegion id="B1472CC71877FFE94734358DAA52EDD5" box="[123,357,1154,1177]" country="Madagascar" name="Mahajanga" pageId="51" pageNumber="52">Mahajanga Province</collectingRegion>
</emphasis>
:
<collectingMunicipality id="9358785F1877FFE94632358DA82EEDD5" box="[381,793,1154,1177]" pageId="51" pageNumber="52">Parc National Tsingy de Bemaraha</collectingMunicipality>
,
<quantity id="B47B4FC01877FFE94461358DA8B8EDD5" box="[814,911,1154,1177]" metricMagnitude="4" metricUnit="m" metricValue="1.06" pageId="51" pageNumber="52" unit="km" value="10.6">10.6 km</quantity>
ESE 123°
<location id="765CB4FE1877FFE9436D358DAFA6EDD5" LSID="urn:lsid:plazi:treatment:FB2A53331875FFE9473432B2AFA2EF30:765CB4FE1877FFE9436D358DAFA6EDD5" box="[1058,1169,1154,1177]" country="Madagascar" latitude="-19.709446" longLatPrecision="21" longitude="44.718056" municipality="Parc National Tsingy de Bemaraha" name="Antsalova" pageId="51" pageNumber="52" stateProvince="Mahajanga">Antsalova</location>
,
<geoCoordinate id="16B784E21877FFE9473435ADABC2EDF5" box="[123,245,1186,1209]" degrees="19" direction="south" minutes="42" orientation="latitude" pageId="51" pageNumber="52" precision="15" seconds="34" value="-19.709446">19°42ʹ34ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21877FFE9464B35ADAA43EDF5" box="[260,372,1186,1209]" degrees="44" direction="east" minutes="43" orientation="longitude" pageId="51" pageNumber="52" precision="15" seconds="5" value="44.718056">44°43ʹ5ʺE</geoCoordinate>
, elev.
<quantity id="B47B4FC01877FFE946F335ADAACCEDF5" box="[444,507,1186,1209]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="51" pageNumber="52" unit="m" value="150.0">
<elevation id="F8AE05161877FFE946F335ADAACCEDF5" box="[444,507,1186,1209]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="51" pageNumber="52" unit="m" value="150.0">150m</elevation>
</quantity>
, tropical dry forest on tsingy, general collecting beating and puffing spiders,
<date id="073DC4E51877FFE9466135CCA91BED96" box="[302,556,1219,1242]" pageId="51" pageNumber="52" value="2001-11-16" valueMax="2001-11-20" valueMin="2001-11-16">
<collectingDate id="17793D0D1877FFE9466135CCA91BED96" box="[302,556,1219,1242]" pageId="51" pageNumber="52" value="2001-11-16" valueMax="2001-11-20" valueMin="2001-11-16">1620 November 2001</collectingDate>
</date>
,
<collectorName id="DE7687F31877FFE9457535CCA8A8ED96" box="[570,927,1219,1242]" pageId="51" pageNumber="52">Fisher-Griswold Arthropod Team</collectorName>
[BLF4503] (
<typeStatus id="AC385C871877FFE9437E35CCAFA6ED96" box="[1073,1169,1219,1242]" pageId="51" pageNumber="52" type="paratype">paratype</typeStatus>
,
<specimenCode id="23254A5E1877FFE9473435ECAA75EDB6" box="[123,322,1251,1274]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9009383</specimenCode>
,
<specimenCount id="658529AC1877FFE9461D35ECAA43EDB6" box="[338,372,1251,1274]" count="1" pageId="51" pageNumber="52" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01877FFE946C835ECAA88EDB6" box="[391,447,1251,1274]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="51" pageNumber="52" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3EBE8781877FFE9469935ECABE3EC37" collectingDate="2000-11-02" collectingDateMax="2000-11-08" collectingDateMin="2000-11-02" collectionCode="FMNH" collectorName="S. Goodman" country="Madagascar" elevation="1250" location="Manasy River" municipality="Foret d'Analavelona" pageId="51" pageNumber="52" specimenCode="CASENT9062273, CASENT9062271, CASENT9062272" specimenCount="6" specimenCount-male="6" stateProvince="Toliara" typeStatus="paratype">
<emphasis id="41F73E371877FFE9469935ECA9BCEDB6" box="[470,651,1251,1274]" italics="true" pageId="51" pageNumber="52">
<collectingRegion id="B1472CC71877FFE9469935ECA9BCEDB6" box="[470,651,1251,1274]" country="Madagascar" name="Toliara" pageId="51" pageNumber="52">Toliara Province</collectingRegion>
</emphasis>
:
<collectingMunicipality id="9358785F1877FFE945D035ECA8B4EDB6" box="[671,899,1251,1274]" pageId="51" pageNumber="52">Fôret dAnalavelona</collectingMunicipality>
, near source of
<location id="765CB4FE1877FFE9430E35ECAB83EC56" LSID="urn:lsid:plazi:treatment:FB2A53331875FFE9473432B2AFA2EF30:765CB4FE1877FFE9430E35ECAB83EC56" country="Madagascar" municipality="Foret d'Analavelona" name="Manasy River" pageId="51" pageNumber="52" stateProvince="Toliara">Manasy River</location>
,
<locationDeviation id="628EFB071877FFE9478C340CAAC8EC56" box="[195,511,1283,1306]" location="d'Andranoheza" pageId="51" pageNumber="52">
<quantity id="B47B4FC01877FFE9478C340CAA2CEC56" box="[195,283,1283,1306]" metricMagnitude="4" metricUnit="m" metricValue="1.65" pageId="51" pageNumber="52" unit="km" value="16.5">16.5 km</quantity>
NW dAndranoheza
</locationDeviation>
, 22°38.6ʹS, 044°10.3ʹE, elev.
<quantity id="B47B4FC01877FFE9440D340CA8B8EC56" box="[834,911,1283,1306]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="51" pageNumber="52" unit="m" value="1250.0">
<elevation id="F8AE05161877FFE9440D340CA8B8EC56" box="[834,911,1283,1306]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="51" pageNumber="52" unit="m" value="1250.0">1250m</elevation>
</quantity>
, undisturbed mid-elevation forest w/dry and humid elements,
<collectingMethod id="AAC29A321877FFE94559342CA96FEC76" box="[534,600,1315,1338]" pageId="51" pageNumber="52">pitfall</collectingMethod>
,
<date id="073DC4E51877FFE9452C342CA876EC76" box="[611,833,1315,1338]" pageId="51" pageNumber="52" value="2000-11-02" valueMax="2000-11-08" valueMin="2000-11-02">
<collectingDate id="17793D0D1877FFE9452C342CA876EC76" box="[611,833,1315,1338]" pageId="51" pageNumber="52" value="2000-11-02" valueMax="2000-11-08" valueMin="2000-11-02">28 November 2000</collectingDate>
</date>
,
<collectorName id="DE7687F31877FFE94404342CA8E4EC76" box="[843,979,1315,1338]" pageId="51" pageNumber="52">S. Goodman</collectorName>
, [FMHD#00-224] (
<specimenCode id="23254A5E1877FFE947CD344CAA7CEC16" box="[130,331,1347,1370]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9062273</specimenCode>
,
<specimenCount id="658529AC1877FFE94610344CAAB7EC16" box="[351,384,1347,1370]" count="1" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01877FFE946D9344CAAD0EC16" box="[406,487,1347,1370]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="51" pageNumber="52" type="Museum">FMNH</collectionCode>
), (
<specimenCode id="23254A5E1877FFE94546344CA9E5EC16" box="[521,722,1347,1370]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9062271</specimenCode>
,
<specimenCount id="658529AC1877FFE945A9344CA830EC16" box="[742,775,1347,1370]" count="4" pageId="51" pageNumber="52" type="male">4♂</specimenCount>
,
<collectionCode id="15927AE01877FFE94452344CA859EC16" box="[797,878,1347,1370]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="51" pageNumber="52" type="Museum">FMNH</collectionCode>
), (
<specimenCode id="23254A5E1877FFE944DF344CAF6DEC16" box="[912,1114,1347,1370]" collectionCode="CASENT" pageId="51" pageNumber="52">CASENT9062272</specimenCode>
,
<specimenCount id="658529AC1877FFE94322344CAFB8EC16" box="[1133,1167,1347,1370]" count="1" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectionCode id="15927AE01877FFE94734346BABFAEC37" box="[123,205,1380,1403]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="51" pageNumber="52" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE2251877FFE947E4348BA933EF70" blockId="51.[123,1175,220,1660]" pageId="51" pageNumber="52">
<emphasis id="41F73E371877FFE947E4348BAA54ECD7" bold="true" box="[171,355,1412,1435]" pageId="51" pageNumber="52">Natural history.</emphasis>
Except for a series of males collected in disturbed transitional montane mossy forest at Réserve Spéciale dAmbohitantely, other collections for
<taxonomicName id="B48399A61877FFE9443634ABAF3CECF7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[889,1035,1444,1467]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371877FFE9443634ABAF3CECF7" bold="true" box="[889,1035,1444,1467]" italics="true" pageId="51" pageNumber="52">Uduba irwini</emphasis>
</taxonomicName>
are from dry habitats including in
<taxonomicName id="B48399A61877FFE9462834CBAA8CEC97" box="[359,443,1476,1499]" class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" order="Malpighiales" pageId="51" pageNumber="52" phylum="Tracheophyta" rank="genus">
<emphasis id="41F73E371877FFE9462834CBAA8CEC97" bold="true" box="[359,443,1476,1499]" italics="true" pageId="51" pageNumber="52">Uapaca</emphasis>
</taxonomicName>
forest, on inselbergs and in tsingy. Females of
<taxonomicName id="B48399A61877FFE9449B34CBAF5DEC97" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[980,1130,1476,1499]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371877FFE9449B34CBAF5DEC97" bold="true" box="[980,1130,1476,1499]" italics="true" pageId="51" pageNumber="52">Uduba irwini</emphasis>
</taxonomicName>
are cribellate and may produce sticky silk, though the collecting data that state “general collecting… reveal nothing of burrows or retreats. As is typical of male
<taxonomicName id="B48399A61877FFE945A0370AA80EEF50" authorityName="Simon" authorityYear="1880" box="[751,825,1541,1564]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="41F73E371877FFE945A0370AA80EEF50" bold="true" box="[751,825,1541,1564]" italics="true" pageId="51" pageNumber="52">Uduba</emphasis>
</taxonomicName>
, these were collected wandering on the ground and in the vegetation.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1877FFE947E4374AAFA2EF30" pageId="51" pageNumber="52" type="distribution">
<paragraph id="733CE2251877FFE947E4374AAFA2EF30" blockId="51.[123,1175,220,1660]" pageId="51" pageNumber="52">
<emphasis id="41F73E371877FFE947E4374AAA0AEF10" bold="true" box="[171,317,1605,1628]" pageId="51" pageNumber="52">Distribution.</emphasis>
The species
<taxonomicName id="B48399A61877FFE94685374AA96AEF10" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[458,605,1605,1628]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="41F73E371877FFE94685374AA96AEF10" bold="true" box="[458,605,1605,1628]" italics="true" pageId="51" pageNumber="52">Uduba irwini</emphasis>
</taxonomicName>
is recorded on the west side of the great escarpment across the southern half of
<collectingCountry id="0B94A2B51877FFE946EE376AA915EF30" box="[417,546,1637,1660]" name="Madagascar" pageId="51" pageNumber="52">Madagascar</collectingCountry>
on the high plateau and western dry forests (
<figureCitation id="EBB8FEA01877FFE94344376AAF69EF30" box="[1035,1118,1637,1660]" captionStart="MAP" captionStartId="177.[155,174,1068,1087]" captionTargetBox="[129,1176,216,1047]" captionTargetId="figure-17@177.[648,1176,216,1059]" captionTargetPageId="177" captionText="MAP 3. Distributions of Uduba rinha and Uduba irwini. MAP 4. Distributions of Uduba madagascariensis, Uduba barbarae, Uduba pseudoevanescens and Uduba orona." figureDoi="http://doi.org/10.5281/zenodo.13158775" httpUri="https://zenodo.org/record/13158775/files/figure.png" pageId="51" pageNumber="52">Maps 3</figureCitation>
, 18).
</paragraph>
</subSubSection>
</treatment>
</document>