287 lines
38 KiB
XML
287 lines
38 KiB
XML
<document id="F31D2980F5E0191E76C54CF8AA7CCFE7" ID-DOI="10.11646/zootaxa.3429.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="10096509" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629801692040" checkinUser="felipe" docAuthor="Jäger, Peter" docDate="2012" docId="C84887861102FF81FF71F8AC543EF950" docLanguage="en" docName="zt03429p063.pdf" docOrigin="Zootaxa 3429 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3429.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Ctenus theodorianum Jäger, 2012, spec. nov." docType="treatment" docVersion="3" lastPageNumber="29" masterDocId="3471FFFE111BFF9DFFE6F83B5749FFC4" masterDocTitle="Asian species of the genera Anahita Karsch 1879, Ctenus Walckenaer 1805 and Amauropelma Raven, Stumkat & Gray 2001 (Arachnida: Araneae: Ctenidae)" masterLastPageNumber="63" masterPageNumber="1" pageNumber="26" updateTime="1699579940834" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="DE60902602FDBFCF1A562037C7C18C37" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E29ACEE664386CB2450F13F5766D5EE3">
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<mods:title id="508C38A331AF51AD7188EF3736FF5C01">Asian species of the genera Anahita Karsch 1879, Ctenus Walckenaer 1805 and Amauropelma Raven, Stumkat & Gray 2001 (Arachnida: Araneae: Ctenidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="EB77BCA1934F3C18787983EA53DFF24A" type="personal">
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<mods:role id="E6673C110547984E0EE4BE2AAD3859C1">
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<mods:roleTerm id="B6209C399B1420884396D6F143428678">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5F95EA233DE7886AE211329F36EC5D76">Jäger, Peter</mods:namePart>
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<mods:typeOfResource id="2941E5DAEB8379F2834B49244522E104">text</mods:typeOfResource>
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<mods:titleInfo id="0CFB4A738413EB4FE19D19758E184296">
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<mods:title id="C55CD590A99253538873DD018E90039E">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="30CC15A0286CC4E2815FAB90C83171EA">
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<mods:date id="EE88145DA6463B5DDDB12F9ABBF4A206">2012</mods:date>
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<mods:detail id="050624696B5FC16A33C537E5CAE4D50E" type="pubDate">
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<mods:number id="73706576F789CB0728336C4495DF2E01">2012-08-20</mods:number>
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</mods:detail>
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<mods:detail id="D8FF4DDBA925BB4F78DEB6AF72838DCC" type="volume">
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<mods:number id="30D062E1ED760FD5AE8E85CF4FAD84A0">3429</mods:number>
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</mods:detail>
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<mods:detail id="2ED45A6F49DA3EBECBEAF31E803CCE4F" type="issue">
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<mods:number id="E0FFD9E42DBB100C8AFC24284714FEC9">1</mods:number>
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<mods:start id="3011EB09355B319BA7700C0111BC5988">1</mods:start>
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<mods:end id="BCB0D23520EF6B21964223EB4E4D6E6A">63</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="5819797F193C727A2007F70671E69313">
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<mods:url id="ABF40BB9614981A795BDEEDF543074C8">http://dx.doi.org/10.11646/zootaxa.3429.1.1</mods:url>
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</mods:location>
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<mods:classification id="969466C1BE72B77F6CE829A9FB155180">journal article</mods:classification>
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<mods:identifier id="5A9FEB9FA764FBC1EF7FB37F56CC5324" type="DOI">10.11646/zootaxa.3429.1.1</mods:identifier>
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<mods:identifier id="0BF157A10357B6A4F47953856FB5A835" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="8D00659A33B4C188A64B43B80F2E302A" type="Zenodo-Dep">10096509</mods:identifier>
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</mods:mods>
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<treatment id="C84887861102FF81FF71F8AC543EF950" LSID="urn:lsid:plazi:treatment:C84887861102FF81FF71F8AC543EF950" httpUri="http://treatment.plazi.org/id/C84887861102FF81FF71F8AC543EF950" lastPageId="28" lastPageNumber="29" pageId="25" pageNumber="26">
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<subSubSection id="08FB651B1102FF84FF71F8AC5554FF75" box="[151,541,151,177]" pageId="25" pageNumber="26" type="nomenclature">
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<paragraph id="405E36901102FF84FF71F8AC5554FF75" blockId="25.[151,541,151,210]" box="[151,541,151,177]" pageId="25" pageNumber="26">
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<heading id="1B1681FC1102FF84FF71F8AC5554FF75" bold="true" box="[151,541,151,177]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
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<emphasis id="7295EA821102FF84FF71F8AC5554FF75" bold="true" box="[151,541,151,177]" pageId="25" pageNumber="26">
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<taxonomicName id="87E14D131102FF84FF71F8AC56D7FF75" ID-CoL="327XX" authorityName="Jager" authorityYear="2012" box="[151,414,151,177]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="theodorianum" status="sp. nov.">
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<emphasis id="7295EA821102FF84FF71F8AC56D7FF75" bold="true" box="[151,414,151,177]" italics="true" pageId="25" pageNumber="26">Ctenus theodorianum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="69A657F91102FF84FE43F8AC5554FF75" box="[421,541,151,177]" pageId="25" pageNumber="26" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="08FB651B1102FF84FF71F8815602FF16" box="[151,331,186,210]" pageId="25" pageNumber="26" type="description">
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<paragraph id="405E36901102FF84FF71F8815602FF16" blockId="25.[151,541,151,210]" box="[151,331,186,210]" pageId="25" pageNumber="26">
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<figureCitation id="D8DA2A151102FF84FF71F88157A5FF16" box="[151,236,186,210]" captionStart="FIGURE 39" captionStartId="14.[151,250,1909,1932]" captionTargetBox="[151,1436,193,1888]" captionTargetId="figure-80@14.[151,1436,193,1888]" captionTargetPageId="14" captionText="FIGURE 39. Records of Ctenus spp. in South and Southeast Asia. 1 Between Chitre and Chandrung, Nepal. 2–5 Burma (2" figureDoi="http://doi.org/10.5281/zenodo.10096523" httpUri="https://zenodo.org/record/10096523/files/figure.png" pageId="25" pageNumber="26">Figs 39</figureCitation>
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,
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<figureCitation id="D8DA2A151102FF84FF1EF8815602FF16" box="[248,331,186,210]" captionStart-0="FIGURES 92–95" captionStart-1="FIGURES 96–104" captionStart-2="FIGURES 105–115" captionStartId-0="25.[151,263,1668,1691]" captionStartId-1="26.[151,264,1635,1658]" captionStartId-2="27.[151,264,1933,1956]" captionTargetBox-0="[157,1429,593,1613]" captionTargetBox-1="[153,1434,193,1614]" captionTargetBox-2="[192,1404,203,1909]" captionTargetId-0="figure-266@25.[157,1429,576,1625]" captionTargetId-1="figure-192@26.[153,1434,193,1614]" captionTargetId-2="figure-57@27.[187,1411,193,1919]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionText-0="FIGURES 92–95. Ctenus theodorianum spec. nov., holotype male from Laos (Phou Hiphi, Oudomxai). 92–94 Left male palp (92 prolateral, 93 ventral, 94 retrolateral). 95 Right leg claw I, prolateral." captionText-1="FIGURES 96–104. Ctenus theodorianum spec. nov., paratype females from Laos (Phou Hiphi, Oudomxai). 96, 101 Epigyne, ventral; 97, 102 Epigyne, posterior; 98, 103 Vulva, dorsal; 99, 104 Schematic course of duct system, dorsal. 100 Right leg claw I, retrolateral." captionText-2="FIGURES 105–115. Ctenus theodorianum spec. nov. from Laos (Phou Hiphi, Oudomxai) (105–107, 111–115 holotype male, 108–110 paratype female). 105–110 Habitus (105, 108 dorsal; 106, 109 ventral; 107, 110 lateral). 111 Hump on femur III, ventral. 112–115 Humps on tibia and metatarsus III, different views." figureDoi-0="http://doi.org/10.5281/zenodo.10096543" figureDoi-1="http://doi.org/10.5281/zenodo.10096545" figureDoi-2="http://doi.org/10.5281/zenodo.10096547" httpUri-0="https://zenodo.org/record/10096543/files/figure.png" httpUri-1="https://zenodo.org/record/10096545/files/figure.png" httpUri-2="https://zenodo.org/record/10096547/files/figure.png" pageId="25" pageNumber="26">92–115</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="08FB651B1102FF84FF71F9385208FE42" pageId="25" pageNumber="26" type="materials_examined">
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<paragraph id="405E36901102FF84FF71F9385208FE42" blockId="25.[151,1436,258,499]" pageId="25" pageNumber="26">
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Type material.
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<materialsCitation id="F0893CCD1102FF84FEAAF9385686FEA6" collectingDate="2011-04-18" collectionCode="SMF" collectorName="L. Nophaseud" country="Laos" elevation="850" latitude="20.652695" location="Phou Hiphi" longLatPrecision="1" longitude="102.002945" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Oudomxai" typeStatus="holotype">
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<emphasis id="7295EA821102FF84FEAAF93856B3FEDF" bold="true" box="[332,506,259,283]" pageId="25" pageNumber="26">
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<typeStatus id="9F5A88321102FF84FEAAF93856F0FEDF" box="[332,441,259,283]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
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male
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</emphasis>
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(
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<collectionCode id="26F0AE551102FF84FDEFF939550FFEDE" box="[521,582,258,282]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="25" pageNumber="26">SMF</collectionCode>
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),
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<collectingCountry id="38F676001102FF84FDBCF93855DAFEDE" box="[602,659,259,282]" name="Laos" pageId="25" pageNumber="26">Laos</collectingCountry>
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,
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<collectingRegion id="8225F8721102FF84FD79F93954CBFEDE" box="[671,898,258,282]" country="Laos" name="Oudomxai" pageId="25" pageNumber="26">Oudomxai Province</collectingRegion>
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,
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<location id="453E604B1102FF84FC68F9395346FEDE" LSID="urn:lsid:plazi:treatment:C84887861102FF81FF71F8AC543EF950:453E604B1102FF84FC68F9395346FEDE" box="[910,1039,258,282]" country="Laos" latitude="20.652695" longLatPrecision="1" longitude="102.002945" name="Phou Hiphi" pageId="25" pageNumber="26" stateProvince="Oudomxai">Phou Hiphi</location>
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provincial protected area, ca.
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<quantity id="87199B751102FF84FA85F93952D5FEDE" box="[1379,1436,258,282]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="26" unit="km" value="4.0">4 km</quantity>
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S of
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<collectingRegion id="8225F8721102FF84FF2BF91D560FFEFA" box="[205,326,294,318]" country="Laos" name="Oudomxai" pageId="25" pageNumber="26">Oudomxai</collectingRegion>
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,
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<geoCoordinate id="25D550571102FF84FEB4F91C56ACFEFA" box="[338,485,294,318]" degrees="20" direction="north" minutes="39" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="9.7" value="20.652695">N 20°39'9.7''</geoCoordinate>
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,
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<geoCoordinate id="25D550571102FF84FE14F91C55D7FEFA" box="[498,670,294,318]" degrees="102" direction="east" minutes="0" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="10.59" value="102.002945">E 102°0'10.59''</geoCoordinate>
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||
,
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<quantity id="87199B751102FF84FD4CF91D55BBFEFA" box="[682,754,294,318]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="25" pageNumber="26" unit="m" value="850.0">
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||
<elevation id="CBCCD1A31102FF84FD4CF91D55BBFEFA" box="[682,754,294,318]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="25" pageNumber="26" unit="m" value="850.0">850 m</elevation>
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</quantity>
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altitude, ground, vegetation, tree bark, by hand, by night,
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<collectorName id="ED1453461102FF84FA62F91C565CFEA6" pageId="25" pageNumber="26">L. Nophaseud</collectorName>
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||
leg.
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||
<date id="345F10501102FF84FEB7F9715686FEA6" box="[337,463,330,354]" pageId="25" pageNumber="26" value="2011-04-18">
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||
<collectingDate id="241BE9B81102FF84FEB7F9715686FEA6" box="[337,463,330,354]" pageId="25" pageNumber="26" value="2011-04-18">18.IV.2011</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="F0893CCD1102FF84FE3AF97053DBFEA6" box="[476,1170,330,355]" collectingDate="2011-04-18" collectionCode="SMF" collectorName="L. Nophaseud" country="Laos" elevation="850" latitude="20.652695" location="Phou Hiphi" longLatPrecision="1" longitude="102.002945" pageId="25" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="Oudomxai" typeStatus="paratype">
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||
<emphasis id="7295EA821102FF84FE3AF9705584FEA7" bold="true" box="[476,717,330,355]" pageId="25" pageNumber="26">
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<typeStatus id="9F5A88321102FF84FE3AF9705511FEA7" box="[476,600,331,355]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
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||
:
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<specimenCount id="56E7FD191102FF84FD81F9715584FEA7" box="[615,717,330,355]" pageId="25" pageNumber="26" type="female">1 female</specimenCount>
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||
</emphasis>
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||
(
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<collectionCode id="26F0AE551102FF84FD39F9715453FEA6" box="[735,794,330,354]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="25" pageNumber="26">SMF</collectionCode>
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) with same data as for holotype
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||
</materialsCitation>
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.
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<materialsCitation id="F0893CCD1102FF84FB79F97156FCFE42" collectingDate="2011-04-18" collectionCode="SMF" collectorName="L. Nophaseud" country="Laos" elevation="850" latitude="20.652695" location="Phou Hiphi" longLatPrecision="1" longitude="102.002945" pageId="25" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="Oudomxai" typeStatus="paratype">
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||
<emphasis id="7295EA821102FF84FB79F971524CFEA7" bold="true" box="[1183,1285,330,355]" pageId="25" pageNumber="26">
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||
<specimenCount id="56E7FD191102FF84FB79F971524CFEA7" box="[1183,1285,330,355]" pageId="25" pageNumber="26" type="female">1 female</specimenCount>
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||
</emphasis>
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||
(
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<collectionCode id="26F0AE551102FF84FAF1F971521DFEA6" box="[1303,1364,330,354]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="25" pageNumber="26">SMF</collectionCode>
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||
), with same data as for holotype
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||
</materialsCitation>
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<materialsCitation id="F0893CCD1102FF84FE5AF9545274FE42" box="[444,1341,366,390]" country="Laos" elevation="830" latitude="20.667866" location="Oudomxai" longLatPrecision="1" longitude="102.04037" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Oudomxai" typeStatus="paratype">
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||
except for:
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||
<quantity id="87199B751102FF84FDDDF955553DFE42" box="[571,628,366,390]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="26" unit="km" value="4.0">4 km</quantity>
|
||
S of
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<collectingRegion id="8225F8721102FF84FD56F9555460FE42" box="[688,809,366,390]" country="Laos" name="Oudomxai" pageId="25" pageNumber="26">Oudomxai</collectingRegion>
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||
,
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||
<geoCoordinate id="25D550571102FF84FCD2F954549CFE42" box="[820,981,366,390]" degrees="20" direction="north" minutes="40" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="4.32" value="20.667866">N 20°40'4.32''</geoCoordinate>
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||
,
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||
<geoCoordinate id="25D550571102FF84FC07F95453C4FE42" box="[993,1165,366,390]" degrees="102" direction="east" minutes="2" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="25.32" value="102.04037">E 102°2'25.32''</geoCoordinate>
|
||
,
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||
<quantity id="87199B751102FF84FB7EF9555396FE42" box="[1176,1247,366,390]" metricMagnitude="2" metricUnit="m" metricValue="8.3" pageId="25" pageNumber="26" unit="m" value="830.0">
|
||
<elevation id="CBCCD1A31102FF84FB7EF9555396FE42" box="[1176,1247,366,390]" metricMagnitude="2" metricUnit="m" metricValue="8.3" pageId="25" pageNumber="26" unit="m" value="830.0">830 m</elevation>
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||
</quantity>
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||
altitude
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||
</materialsCitation>
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||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="08FB651B1102FF84FF21F9A855B3FE36" pageId="25" pageNumber="26" type="etymology">
|
||
<paragraph id="405E36901102FF84FF21F9A855B3FE36" blockId="25.[151,1436,258,499]" pageId="25" pageNumber="26">
|
||
<emphasis id="7295EA821102FF84FF21F9A85607FE6F" bold="true" box="[199,334,403,427]" inLineHeading="true" pageId="25" pageNumber="26" reason="1">Etymology.</emphasis>
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||
This species is named in honour of my secondary school “Gymnasium Theodorianum” in Paderborn,
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<collectingCountry id="38F676001102FF84FEFCF98D56CAFE0A" box="[282,387,438,462]" name="Germany" pageId="25" pageNumber="26">Germany</collectingCountry>
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||
, on the occasion of its 400th anniversary for providing an excellent education and training, therefore a good start into my life; name in apposition.
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||
</paragraph>
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||
</subSubSection>
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||
<caption id="149E66181102FF84FF71FEBF5433F97A" ID-DOI="http://doi.org/10.5281/zenodo.10096543" ID-Zenodo-Dep="10096543" httpUri="https://zenodo.org/record/10096543/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,263,1668,1691]" targetBox="[157,1429,593,1613]" targetPageId="25">
|
||
<paragraph id="405E36901102FF84FF71FEBF5433F97A" blockId="25.[151,1436,1668,1726]" pageId="25" pageNumber="26">
|
||
<emphasis id="7295EA821102FF84FF71FEBF55E0F95E" bold="true" box="[151,681,1668,1691]" pageId="25" pageNumber="26">
|
||
FIGURES 92–95.
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<taxonomicName id="87E14D131102FF84FEBFFEBE5575F95F" authorityName="Jager" authorityYear="2012" box="[345,572,1669,1691]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="theodorianum" status="sp. nov.">
|
||
<emphasis id="7295EA821102FF84FEBFFEBE5575F95F" bold="true" box="[345,572,1669,1691]" italics="true" pageId="25" pageNumber="26">Ctenus theodorianum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="69A657F91102FF84FDA4FEBE55E0F95E" box="[578,681,1669,1690]" pageId="25" pageNumber="26" rank="species">spec. nov.</taxonomicNameLabel>
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||
</emphasis>
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, holotype male from Laos (Phou Hiphi, Oudomxai). 92–94 Left male palp (92 prolateral, 93 ventral, 94 retrolateral). 95 Right leg claw I, prolateral.
|
||
</paragraph>
|
||
</caption>
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||
<subSubSection id="08FB651B1102FF84FF21FED552D5F852" pageId="25" pageNumber="26" type="diagnosis">
|
||
<paragraph id="405E36901102FF84FF21FED552D5F852" blockId="25.[151,1436,1774,1943]" pageId="25" pageNumber="26">
|
||
<emphasis id="7295EA821102FF84FF21FED55608F8C3" bold="true" box="[199,321,1774,1799]" inLineHeading="true" pageId="25" pageNumber="26" reason="1">Diagnosis.</emphasis>
|
||
Medium-sized
|
||
<taxonomicName id="87E14D131102FF84FE1BFED55529F8C2" authorityName="Keyserling" authorityYear="1877" box="[509,608,1774,1798]" class="Arachnida" family="Ctenidae" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="family">Ctenidae</taxonomicName>
|
||
(total length male 10.8, female 12.3–13.8). Distinguished from other
|
||
<taxonomicName id="87E14D131102FF84FF71FF295655F8EE" box="[151,284,1810,1834]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="7295EA821102FF84FF71FF2957ADF8EE" box="[151,228,1810,1834]" italics="true" pageId="25" pageNumber="26">Ctenus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, but not
|
||
<taxonomicName id="87E14D131102FF84FE9AFF2956B4F8EE" authorityName="Thorell" authorityYear="1897" box="[380,509,1810,1834]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="robustus">
|
||
<emphasis id="7295EA821102FF84FE9AFF2956B4F8EE" box="[380,509,1810,1834]" italics="true" pageId="25" pageNumber="26">C. robustus</emphasis>
|
||
</taxonomicName>
|
||
, by the modified leg III in males with ventral hump on proximal femur covered by small stout spines (
|
||
<figureCitation id="D8DA2A151102FF84FE8BFF0D5683F88A" box="[365,458,1846,1870]" captionStart="FIGURES 105–115" captionStartId="27.[151,264,1933,1956]" captionTargetBox="[192,1404,203,1909]" captionTargetId="figure-57@27.[187,1411,193,1919]" captionTargetPageId="27" captionText="FIGURES 105–115. Ctenus theodorianum spec. nov. from Laos (Phou Hiphi, Oudomxai) (105–107, 111–115 holotype male, 108–110 paratype female). 105–110 Habitus (105, 108 dorsal; 106, 109 ventral; 107, 110 lateral). 111 Hump on femur III, ventral. 112–115 Humps on tibia and metatarsus III, different views." figureDoi="http://doi.org/10.5281/zenodo.10096547" httpUri="https://zenodo.org/record/10096547/files/figure.png" pageId="25" pageNumber="26">Fig. 111</figureCitation>
|
||
) and two ventral cone-shaped humps each with one spine on proximal metatarsus (
|
||
<figureCitation id="D8DA2A151102FF84FA8CFF0D57B2F8B6" captionStart="FIGURES 105–115" captionStartId="27.[151,264,1933,1956]" captionTargetBox="[192,1404,203,1909]" captionTargetId="figure-57@27.[187,1411,193,1919]" captionTargetPageId="27" captionText="FIGURES 105–115. Ctenus theodorianum spec. nov. from Laos (Phou Hiphi, Oudomxai) (105–107, 111–115 holotype male, 108–110 paratype female). 105–110 Habitus (105, 108 dorsal; 106, 109 ventral; 107, 110 lateral). 111 Hump on femur III, ventral. 112–115 Humps on tibia and metatarsus III, different views." figureDoi="http://doi.org/10.5281/zenodo.10096547" httpUri="https://zenodo.org/record/10096547/files/figure.png" pageId="25" pageNumber="26">Figs 112–115</figureCitation>
|
||
). For differential diagnosis see
|
||
<taxonomicName id="87E14D131102FF84FD86FF6155A9F8B6" authorityName="Thorell" authorityYear="1897" box="[608,736,1882,1906]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="robustus">
|
||
<emphasis id="7295EA821102FF84FD86FF6155A9F8B6" box="[608,736,1882,1906]" italics="true" pageId="25" pageNumber="26">C. robustus</emphasis>
|
||
</taxonomicName>
|
||
. Females show typical characters of other Asian
|
||
<taxonomicName id="87E14D131102FF84FAF6FF6152D1F8B6" box="[1296,1432,1882,1906]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="7295EA821102FF84FAF6FF615217F8B6" box="[1296,1374,1882,1906]" italics="true" pageId="25" pageNumber="26">Ctenus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
: broad median epigynal plate with two lateral teeth, internal duct system consisting of spherical spermathecae, short
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="149E66181101FF87FF71FE58566BF97A" ID-DOI="http://doi.org/10.5281/zenodo.10096545" ID-Zenodo-Dep="10096545" httpUri="https://zenodo.org/record/10096545/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,264,1635,1658]" targetBox="[153,1434,193,1614]" targetPageId="26">
|
||
<paragraph id="405E36901101FF87FF71FE58566BF97A" blockId="26.[151,1436,1635,1726]" pageId="26" pageNumber="27">
|
||
<emphasis id="7295EA821101FF87FF71FE5855FFF9BD" bold="true" box="[151,694,1635,1658]" pageId="26" pageNumber="27">
|
||
FIGURES 96–104.
|
||
<taxonomicName id="87E14D131101FF87FE80FE585500F9BD" authorityName="Jager" authorityYear="2012" box="[358,585,1635,1657]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="theodorianum" status="sp. nov.">
|
||
<emphasis id="7295EA821101FF87FE80FE585500F9BD" bold="true" box="[358,585,1635,1657]" italics="true" pageId="26" pageNumber="27">Ctenus theodorianum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="69A657F91101FF87FDB6FE5F55FFF9BD" box="[592,694,1636,1657]" pageId="26" pageNumber="27" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype females from Laos (Phou Hiphi, Oudomxai). 96, 101 Epigyne, ventral; 97, 102 Epigyne, posterior; 98, 103 Vulva, dorsal; 99, 104 Schematic course of duct system, dorsal. 100 Right leg claw I, retrolateral.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="08FB651B1101FF81FF21FED654ABF9AB" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="description">
|
||
<paragraph id="405E36901101FF87FF21FED652D5F852" blockId="26.[151,1437,1773,1943]" pageId="26" pageNumber="27">
|
||
<emphasis id="7295EA821101FF87FF21FED656D4F8C2" bold="true" box="[199,413,1773,1798]" pageId="26" pageNumber="27">
|
||
<emphasis id="7295EA821101FF87FF21FED65611F8C2" box="[199,344,1773,1798]" inLineHeading="true" pageId="26" pageNumber="27" reason="1">Description.</emphasis>
|
||
Male
|
||
</emphasis>
|
||
(
|
||
<typeStatus id="9F5A88321101FF87FE48FED5555EF8C2" box="[430,535,1774,1798]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
|
||
). PL 6.2, PW 4.8, AW 2.5, OL 4.6, OW 2.9. Eye diameters and interdistances: AME 0.32, ALE 0.21, PME 0.40, PLE 0.39, AME–AME 0.12, AME–ALE 0.33, PME–PME 0.11, PME–PLE 0.31, AME–PME 0.08, ALE–PLE 0.12, clypeus AME 0.14, clypeus ALE 0.61. Palp and leg measurements: palp 6.55 (2.4, 1.1, 1.25, -, 1.5), I 22.2 (5.8, 2.5, 6.0, 5.7, 2.2), II 19.4 (5.3, 2.3, 5.0, 5.0, 1.8), III 16.1 (4.6, 2.0, 3.4, 4.5, 1.6), IV 23.8 (6.2, 2.2, 5.7, 7.6, 2.1). Leg formula 4123. Spination of palp and legs: palp 141, 100, 1010; femora I p121,
|
||
</paragraph>
|
||
<caption id="149E66181100FF86FF71FFB65405F82C" ID-DOI="http://doi.org/10.5281/zenodo.10096547" ID-Zenodo-Dep="10096547" httpUri="https://zenodo.org/record/10096547/files/figure.png" pageId="27" pageNumber="28" startId="27.[151,264,1933,1956]" targetBox="[192,1404,203,1909]" targetPageId="27">
|
||
<paragraph id="405E36901100FF86FF71FFB65405F82C" blockId="27.[151,1436,1933,2024]" pageId="27" pageNumber="28">
|
||
<emphasis id="7295EA821100FF86FF71FFB6558BF867" bold="true" box="[151,706,1933,1956]" pageId="27" pageNumber="28">
|
||
FIGURES 105–115.
|
||
<taxonomicName id="87E14D131100FF86FE94FFB6551CF867" authorityName="Jager" authorityYear="2012" box="[370,597,1933,1955]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="theodorianum" status="sp. nov.">
|
||
<emphasis id="7295EA821100FF86FE94FFB6551CF867" bold="true" box="[370,597,1933,1955]" italics="true" pageId="27" pageNumber="28">Ctenus theodorianum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="69A657F91100FF86FDBDFFB5558BF867" box="[603,706,1934,1955]" pageId="27" pageNumber="28" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from Laos (Phou Hiphi, Oudomxai) (105–107, 111–115 holotype male, 108–110 paratype female). 105–110 Habitus (105, 108 dorsal; 106, 109 ventral; 107, 110 lateral). 111 Hump on femur III, ventral. 112–115 Humps on tibia and metatarsus III, different views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="405E36901107FF81FF71F8AC54F7FEDA" blockId="28.[151,1437,151,1685]" pageId="28" pageNumber="29">
|
||
Chelicerae with 3 promarginal, 4 retromarginal teeth, and with elongated patch of many tiny denticles along entire cheliceral furrow. Retromargin of chelicerae close to fang base with several short and thin bristles. Sparse scopula restricted almost entirely to tarsi, only metatarsi I–II with sparse scopula hairs. Right leg claw I with 5 secondary teeth (
|
||
<figureCitation id="D8DA2A151107FF81FF3AF93D5679FEDA" box="[220,304,262,286]" captionStart="FIGURES 92–95" captionStartId="25.[151,263,1668,1691]" captionTargetBox="[157,1429,593,1613]" captionTargetId="figure-266@25.[157,1429,576,1625]" captionTargetPageId="25" captionText="FIGURES 92–95. Ctenus theodorianum spec. nov., holotype male from Laos (Phou Hiphi, Oudomxai). 92–94 Left male palp (92 prolateral, 93 ventral, 94 retrolateral). 95 Right leg claw I, prolateral." figureDoi="http://doi.org/10.5281/zenodo.10096543" httpUri="https://zenodo.org/record/10096543/files/figure.png" pageId="28" pageNumber="29">Fig. 95</figureCitation>
|
||
). Position of tarsal organ: I 0.90, II 0.90, III 0.80, IV 0.85.
|
||
</paragraph>
|
||
<paragraph id="405E36901107FF81FF21F9105581FE12" blockId="28.[151,1437,151,1685]" pageId="28" pageNumber="29">
|
||
Palp as in diagnosis (see also
|
||
<taxonomicName id="87E14D131107FF81FDC4F91055EDFE87" authorityName="Thorell" authorityYear="1897" box="[546,676,299,323]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="robustus">
|
||
<emphasis id="7295EA821107FF81FDC4F91055EDFE87" box="[546,676,299,323]" italics="true" pageId="28" pageNumber="29">C. robustus</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation id="D8DA2A151107FF81FD59F910540DFE87" box="[703,836,299,323]" captionStart="FIGURES 92–95" captionStartId="25.[151,263,1668,1691]" captionTargetBox="[157,1429,593,1613]" captionTargetId="figure-266@25.[157,1429,576,1625]" captionTargetPageId="25" captionText="FIGURES 92–95. Ctenus theodorianum spec. nov., holotype male from Laos (Phou Hiphi, Oudomxai). 92–94 Left male palp (92 prolateral, 93 ventral, 94 retrolateral). 95 Right leg claw I, prolateral." figureDoi="http://doi.org/10.5281/zenodo.10096543" httpUri="https://zenodo.org/record/10096543/files/figure.png" pageId="28" pageNumber="29">Figs 92–94</figureCitation>
|
||
). Palpal tibia with strong RTA, with broad base and blunt tip. Cymbium tip slightly conical, retro-proximally with distinct pointed extension. Embolus arising at 7.30- o’clock-position, short, its tip situated almost centrally. Conductor arising at 12-o’clock-position subdistally, partly fused with tegulum. Tegular apophysis arising at 6- to 6.30-o’clock-position from tegulum, in ventral view divided into two parts, distinctly excavated on dorsal side.
|
||
</paragraph>
|
||
<paragraph id="405E36901107FF81FF21F9D85340FD38" blockId="28.[151,1437,151,1685]" pageId="28" pageNumber="29">
|
||
Colour (
|
||
<figureCitation id="D8DA2A151107FF81FECFF9D85684FE3F" box="[297,461,483,507]" captionStart="FIGURES 105–115" captionStartId="27.[151,264,1933,1956]" captionTargetBox="[192,1404,203,1909]" captionTargetId="figure-57@27.[187,1411,193,1919]" captionTargetPageId="27" captionText="FIGURES 105–115. Ctenus theodorianum spec. nov. from Laos (Phou Hiphi, Oudomxai) (105–107, 111–115 holotype male, 108–110 paratype female). 105–110 Habitus (105, 108 dorsal; 106, 109 ventral; 107, 110 lateral). 111 Hump on femur III, ventral. 112–115 Humps on tibia and metatarsus III, different views." figureDoi="http://doi.org/10.5281/zenodo.10096547" httpUri="https://zenodo.org/record/10096547/files/figure.png" pageId="28" pageNumber="29">Figs 105–107</figureCitation>
|
||
). Brown partly with dark-brown pattern. Dorsal prosoma with broad lateral bands marginally and dark bands around distinctly marked fovea. Sternum, ventral coxae III + IV, labium and gnathocoxae dark, partly with lighter patches, coxae I + II yellowish-brown without pattern. Chelicerae as dorsal prosoma with longitudinal lines in proximal half and with dark distal half. Leg femora dark with irregular darker patches, patellae to tarsi yellowish-brown. Dorsal opisthosoma greyish-brown, mottled with small dark spots, anterior half with lighter patches. Lateral opisthosoma with spots in longitudinal rows. Ventral opisthosoma black with two pairs of white patches consisting of white hairs; epiandrium and muscle sigilla light. Anterior lateral spinnerets dark, posterior lateral and median spinnerets and anal tubercle light.
|
||
</paragraph>
|
||
<paragraph id="405E36901107FF81FF21FB32561FFB72" blockId="28.[151,1437,151,1685]" pageId="28" pageNumber="29">
|
||
<emphasis id="7295EA821107FF81FF21FB3255FBFCE5" box="[199,690,777,801]" inLineHeading="true" pageId="28" pageNumber="29" reason="1">
|
||
<emphasis id="7295EA821107FF81FF21FB325657FCE5" bold="true" box="[199,286,777,801]" pageId="28" pageNumber="29">Female</emphasis>
|
||
(
|
||
<typeStatus id="9F5A88321107FF81FED6FB3256DBFCE5" box="[304,402,777,801]" pageId="28" pageNumber="29" type="paratype">paratype</typeStatus>
|
||
together with
|
||
<typeStatus id="9F5A88321107FF81FDA6FB3255EEFCE5" box="[576,679,777,801]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
|
||
).
|
||
</emphasis>
|
||
PL 6.9, PW 5.6, AW 3.6, OL 7.0, OW 4.6. Eye diameters and interdistances: AME 0.37, ALE 0.28, PME 0.47, PLE 0.39, AME–AME 0.18, AME–ALE 0.40, PME–PME 0.24, PME–PLE 0.41, AME–PME 0.11, ALE–PLE 0.15, clypeus AME 0.22, clypeus ALE 0.70. Palp and leg measurements: palp 7.0 (2.4, 1.3, 1.5, -, 1.8), I 18.15 (5.2, 2.9, 4.7, 3.9, 1.45), II 17.0 (4.9, 2.7, 4.2, 3.7, 1.5), III 14.9 (4.3, 2.2, 3.3, 3.6, 1.5), IV 20.8 (5.6, 2.5, 4.9, 6.0, 1.8). Leg formula 4123. Spination of palp and legs: palp 131, 100, 1211, 1013; femora I p021, d111, r111, II–III p112, d111, r 111, IV p111, d111, r011; patellae I–II 000, III–IV 101; tibiae I–II v22222, III–IV p11, d111, r11, v222; metatarsi I v22(1)2, II v222, III p112, d010, r112, v222, IV p112, d010, r112,
|
||
<date id="345F10501107FF81FE29FC305551FBE7" box="[463,536,1035,1059]" pageId="28" pageNumber="29" value="2122-05">v2122</date>
|
||
. Chelicerae with 3 promarginal, 4 retromarginal teeth, and with elongated patch of many tiny denticles along entire cheliceral furrow. Retromargin of chelicerae close to fang base with 13 thin bristles. Ventral tarsi and entire metatarsi with sparse scopula. Claw tufts with concave distal surface. Palpal claw with 5 secondary teeth. Right leg claw I with 3 secondary teeth (
|
||
<figureCitation id="D8DA2A151107FF81FC88FC425498FB55" box="[878,977,1145,1169]" captionStart="FIGURES 96–104" captionStartId="26.[151,264,1635,1658]" captionTargetBox="[153,1434,193,1614]" captionTargetId="figure-192@26.[153,1434,193,1614]" captionTargetPageId="26" captionText="FIGURES 96–104. Ctenus theodorianum spec. nov., paratype females from Laos (Phou Hiphi, Oudomxai). 96, 101 Epigyne, ventral; 97, 102 Epigyne, posterior; 98, 103 Vulva, dorsal; 99, 104 Schematic course of duct system, dorsal. 100 Right leg claw I, retrolateral." figureDoi="http://doi.org/10.5281/zenodo.10096545" httpUri="https://zenodo.org/record/10096545/files/figure.png" pageId="28" pageNumber="29">Fig. 100</figureCitation>
|
||
). Position of tarsal organ: I 0.90, II 0.85, III 0.76, IV 0.83.
|
||
</paragraph>
|
||
<paragraph id="405E36901107FF81FF21FCF85258FA57" blockId="28.[151,1437,151,1685]" pageId="28" pageNumber="29">
|
||
Copulatory organ as in diagnosis (see also
|
||
<taxonomicName id="87E14D131107FF81FD2CFCF85406FB1F" authorityName="Thorell" authorityYear="1897" box="[714,847,1219,1243]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="robustus">
|
||
<emphasis id="7295EA821107FF81FD2CFCF85406FB1F" box="[714,847,1219,1243]" italics="true" pageId="28" pageNumber="29">C. robustus</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation id="D8DA2A151107FF81FC8BFCF8534AFB1F" box="[877,1027,1219,1243]" captionStart="FIGURES 96–104" captionStartId="26.[151,264,1635,1658]" captionTargetBox="[153,1434,193,1614]" captionTargetId="figure-192@26.[153,1434,193,1614]" captionTargetPageId="26" captionText="FIGURES 96–104. Ctenus theodorianum spec. nov., paratype females from Laos (Phou Hiphi, Oudomxai). 96, 101 Epigyne, ventral; 97, 102 Epigyne, posterior; 98, 103 Vulva, dorsal; 99, 104 Schematic course of duct system, dorsal. 100 Right leg claw I, retrolateral." figureDoi="http://doi.org/10.5281/zenodo.10096545" httpUri="https://zenodo.org/record/10096545/files/figure.png" pageId="28" pageNumber="29">Figs 96–104</figureCitation>
|
||
). Epigynal field with two separate anterio-lateral patches, close to these patches two slit sense organs. Median plate narrow anteriorly, widening medially and narrowing posteriorly again, posteriorly with subparallel margins, lateral teeth situated at widest part. Posterior epigyne with distinct lateral furrows. Internal duct system with two large lateral folds running diagonally from medially to laterally. Spherical spermathecae separated by their diameter or more than that from each other, with a small round chamber between spermathecae and fertilisation ducts, the latter pointing medially.
|
||
</paragraph>
|
||
<paragraph id="405E36901107FF81FF21FDA4527BF9C5" blockId="28.[151,1437,151,1685]" pageId="28" pageNumber="29">
|
||
Colour (
|
||
<figureCitation id="D8DA2A151107FF81FEC2FDA4568BFA73" box="[292,450,1439,1463]" captionStart="FIGURES 105–115" captionStartId="27.[151,264,1933,1956]" captionTargetBox="[192,1404,203,1909]" captionTargetId="figure-57@27.[187,1411,193,1919]" captionTargetPageId="27" captionText="FIGURES 105–115. Ctenus theodorianum spec. nov. from Laos (Phou Hiphi, Oudomxai) (105–107, 111–115 holotype male, 108–110 paratype female). 105–110 Habitus (105, 108 dorsal; 106, 109 ventral; 107, 110 lateral). 111 Hump on femur III, ventral. 112–115 Humps on tibia and metatarsus III, different views." figureDoi="http://doi.org/10.5281/zenodo.10096547" httpUri="https://zenodo.org/record/10096547/files/figure.png" pageId="28" pageNumber="29">Figs 108–110</figureCitation>
|
||
). As in male, but generally darker. Dorsal prosoma with two posteriorly diverging lines starting at PLE. Distal leg segments with indistinct patches, especially in legs III–IV. Dorsal opisthosoma with pairs of indistinct small light patches. Ventral opisthosoma with ca. 8 pairs of white patches, partly fused.
|
||
</paragraph>
|
||
<paragraph id="405E36901107FF81FF21FE3654ABF9AB" blockId="28.[151,1437,151,1685]" pageId="28" pageNumber="29">
|
||
<emphasis id="7295EA821107FF81FF21FE365608F9E2" bold="true" box="[199,321,1549,1574]" inLineHeading="true" pageId="28" pageNumber="29" reason="1">Variation.</emphasis>
|
||
Second female
|
||
<typeStatus id="9F5A88321107FF81FDE7FE34552CF9E3" box="[513,613,1551,1575]" pageId="28" pageNumber="29" type="paratype">paratype</typeStatus>
|
||
: PL 6.3, OL 6.3. Epigyne smaller and relatively slightly longer, with posterior parallel margins indistinct. Lateral teeth not as pointed as in syntopic
|
||
<typeStatus id="9F5A88321107FF81FBE1FE085322F98F" box="[1031,1131,1587,1611]" pageId="28" pageNumber="29" type="paratype">paratype</typeStatus>
|
||
. In the internal duct system glandular pores could be recognised and are added in the schematic course.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="08FB651B1107FF81FF21FE47543EF950" box="[199,887,1660,1685]" pageId="28" pageNumber="29" type="distribution">
|
||
<paragraph id="405E36901107FF81FF21FE47543EF950" blockId="28.[151,1437,151,1685]" box="[199,887,1660,1685]" pageId="28" pageNumber="29">
|
||
<emphasis id="7295EA821107FF81FF21FE475629F951" bold="true" box="[199,352,1660,1685]" inLineHeading="true" pageId="28" pageNumber="29" reason="1">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="9F5A88321107FF81FD86FE4655D9F951" box="[608,656,1661,1685]" pageId="28" pageNumber="29">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="D8DA2A151107FF81FD1CFE475406F950" box="[762,847,1660,1684]" captionStart="FIGURE 39" captionStartId="14.[151,250,1909,1932]" captionTargetBox="[151,1436,193,1888]" captionTargetId="figure-80@14.[151,1436,193,1888]" captionTargetPageId="14" captionText="FIGURE 39. Records of Ctenus spp. in South and Southeast Asia. 1 Between Chitre and Chandrung, Nepal. 2–5 Burma (2" figureDoi="http://doi.org/10.5281/zenodo.10096523" httpUri="https://zenodo.org/record/10096523/files/figure.png" pageId="28" pageNumber="29">Fig. 39</figureCitation>
|
||
: 8).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |