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<document id="474922F7B8342216CA4094CDA65DD05A" ID="10.11646/zootaxa.4117.4.5" ID-CLB-Dataset="38939" ID-DOI="10.11646/zootaxa.4117.4.5" ID-GBIF-Dataset="d4421d20-2632-4996-9356-e7b6f53dcc59" ID-ISSN="1175-5326" ID-Zenodo-Dep="255732" ID-ZooBank="B05704F2-CF95-4724-9398-2D8F6B01A09B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1464850862829" checkinUser="plazi" docAuthor="Mazuch, Tomáš, Šmíd, Jiří &amp; Bauer, Aaron M." docDate="2016" docId="D5444317FF99FFC1FF3BF9CAFF0BF876" docLanguage="en" docName="zootaxa.4117.4.5.pdf" docOrigin="Zootaxa 4117 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hemidactylus fragilis Calabresi 1915" docType="treatment" docVersion="9" lastPageNumber="538" masterDocId="297D3B6FFF9EFFC8FFACFFE5FFD0FFF2" masterDocTitle="Rediscovery and a new record of Hemidactylus laevis (Reptilia: Gekkonidae) from Somaliland, with notes on and resurrection of Hemidactylus fragilis" masterLastPageNumber="542" masterPageNumber="529" pageNumber="536" updateTime="1698662683761" updateUser="ExternalLinkService">
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<mods:title id="79421AAD8DE799725C3ED6DD2D138498">Rediscovery and a new record of Hemidactylus laevis (Reptilia: Gekkonidae) from Somaliland, with notes on and resurrection of Hemidactylus fragilis</mods:title>
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<mods:namePart id="CB1D33141EC119E1F0C549203A19521D">Mazuch, Tomáš</mods:namePart>
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<mods:namePart id="56E502333154960A343F8DE85E917507">Šmíd, Jiří</mods:namePart>
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<mods:namePart id="5D0EEA1236F4738CDD3EDBE1A3A6D57A">Bauer, Aaron M.</mods:namePart>
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<treatment id="D5444317FF99FFC1FF3BF9CAFF0BF876" ID-DOI="http://doi.org/10.5281/zenodo.5618074" ID-GBIF-Taxon="120735694" ID-Zenodo-Dep="5618074" LSID="urn:lsid:plazi:treatment:D5444317FF99FFC1FF3BF9CAFF0BF876" httpUri="http://treatment.plazi.org/id/D5444317FF99FFC1FF3BF9CAFF0BF876" lastPageId="9" lastPageNumber="538" pageId="7" pageNumber="536">
<subSubSection id="15F7A18AFF99FFCFFF3BF9CAFCFDF9BB" box="[151,813,1583,1609]" pageId="7" pageNumber="536" type="nomenclature">
<paragraph id="5D52F201FF99FFCFFF3BF9CAFCFDF9BB" blockId="7.[151,813,1583,1609]" box="[151,813,1583,1609]" pageId="7" pageNumber="536">
<heading id="061A456DFF99FFCFFF3BF9CAFCFDF9BB" bold="true" box="[151,813,1583,1609]" fontSize="11" level="1" pageId="7" pageNumber="536" reason="1">
<emphasis id="6F992E13FF99FFCFFF3BF9CAFCFDF9BB" bold="true" box="[151,813,1583,1609]" pageId="7" pageNumber="536">
Resurrection of
<taxonomicName id="9AED8982FF99FFCFFECCF9CAFCFDF9BB" authority="Calabresi, 1915" authorityName="Calabresi" authorityYear="1915" box="[352,813,1583,1609]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="536" phylum="Chordata" rank="species" species="fragilis">
<emphasis id="6F992E13FF99FFCFFECCF9CAFDB3F9BB" bold="true" box="[352,611,1583,1609]" italics="true" pageId="7" pageNumber="536">Hemidactylus fragilis</emphasis>
<bibRefCitation id="397C8FF0FF99FFCFFDC7F9CAFCFDF9BB" author="Calabresi" box="[619,813,1583,1609]" pageId="7" pageNumber="536" refString="Calabresi, E. (1915) Contributo alia conoscenza dei Rettili della Somalia. Monitore Zoologico Italiano, 26, 234 - 247." type="journal article" year="1915">Calabresi, 1915</bibRefCitation>
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<subSubSection id="15F7A18AFF99FFC1FF3BF992FF0BF876" lastPageId="9" lastPageNumber="538" pageId="7" pageNumber="536" type="description">
<paragraph id="5D52F201FF99FFC0FF3BF992FAA0FEB2" blockId="7.[151,1436,1653,2005]" lastBlockId="8.[151,1437,151,320]" lastPageId="8" lastPageNumber="537" pageId="7" pageNumber="536">
We examined all specimens previously identified as
<taxonomicName id="9AED8982FF99FFCFFD55F992FCBFF97C" box="[761,879,1654,1678]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="536" phylum="Chordata" rank="species" species="fragilis">
<emphasis id="6F992E13FF99FFCFFD55F992FCBFF97C" box="[761,879,1654,1678]" italics="true" pageId="7" pageNumber="536">H. fragilis</emphasis>
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, which are: (1) the
<typeStatus id="82564CA3FF99FFCFFBF7F990FB6DF97C" box="[1115,1213,1653,1678]" pageId="7" pageNumber="536" type="holotype">holotype</typeStatus>
(MZUF 707; adult male;
<collectingCountry id="25FAB291FF99FFCFFF72F97CFEECF940" box="[222,316,1689,1714]" name="Somalia" pageId="7" pageNumber="536">Somalia</collectingCountry>
, Bur Meldac [=Meel Daaq]); (2) MSNG 53212 (adult female;
<collectingCountry id="25FAB291FF99FFCFFBBCF97CFBA2F940" box="[1040,1138,1689,1714]" name="Ethiopia" pageId="7" pageNumber="536">Ethiopia</collectingCountry>
, Dolo); (3) MSNG 53213 (adult female;
<collectingCountry id="25FAB291FF99FFCFFEEEF958FE71F924" box="[322,417,1725,1750]" name="Somalia" pageId="7" pageNumber="536">Somalia</collectingCountry>
, Rahanuin). Their distribution is summarized in
<figureCitation id="C5D6EE84FF99FFCFFC72F958FBF1F924" box="[990,1057,1725,1750]" captionStart="FIGURE 1" captionStartId="2.[151,250,1622,1644]" captionTargetBox="[182,1411,185,1592]" captionTargetId="figure@2.[170,1417,177,1600]" captionTargetPageId="2" captionText="FIGURE 1. (A) Map of Somaliland and the adjoining part of Somalia showing the distribution of Hemidactylus laevis (red circle—type locality, Gaan Libah; blue circle—new record, Erigavo); (B) Map of the Horn of Africa showing the distribution of H. laticaudatus (green squares) — 1: Harar (type locality); 2: Lalibela; 3: Gondar, Azezo, Fiume Caha; 4: Axum; 5: Adwa; 6: 32 km E of Neghelli; 7: Asmara; 8: Saganeiti (type locality of H. fossatii) and H. fragilis (yellow triangles) — 1: Bur Meldac (type locality); 2: “ Rahanuin; 3: Dolo. Records of H. laticaudatus are from Scortecci (1928), Largen (1997), Largen &amp; Spawls (2006, 2010), and own unpublished data." httpUri="https://zenodo.org/record/255733/files/figure.png" pageId="7" pageNumber="536">Fig. 1</figureCitation>
B.
<taxonomicName id="9AED8982FF99FFCFFBE4F95BFAEEF924" box="[1096,1342,1726,1750]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="536" phylum="Chordata" rank="species" species="fragilis">
<emphasis id="6F992E13FF99FFCFFBE4F95BFAEEF924" box="[1096,1342,1726,1750]" italics="true" pageId="7" pageNumber="536">Hemidactylus fragilis</emphasis>
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has the following combination of morphological characters: (1) small size with maximum recorded SVL
<quantity id="9A155FE4FF99FFCFFB49F904FA9AF90B" box="[1253,1354,1761,1786]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" pageId="7" pageNumber="536" unit="mm" value="31.5">31.5 mm</quantity>
for the male and
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for females (
<figureCitation id="C5D6EE84FF99FFCFFDADF8E0FD94F8EC" box="[513,580,1797,1822]" captionStart="FIGURE 5" captionStartId="8.[151,250,1710,1732]" captionTargetBox="[160,1427,410,1677]" captionTargetId="figure@8.[151,1436,373,1689]" captionTargetPageId="8" captionText="FIGURE 5. (A) A topotype adult male of Hemidactylus laticaudatus from Harar, Ethiopia; (B) The holotype of Hemidactylus fragilis (MZUF 707) from Bur Meldac, Somalia; (C) adult female of H. fragilis (MSNG 53212) from Dolo Odo, Ethiopia, with regenerated tail; (D) female of H. fragilis (MSNG 53213) from Rahanuin, Somalia, with partially regenerated tail; (E) detail of the tail of the previous specimen, note the enlarged tubercles on the original part of tail; (F) detail of mental region of H. frenatus (MZUF 27607) from Mogadishu, Somalia; (G) detail of mental region of H. fragilis (MSNG 53213) from Rahanuin, Somalia; (H) detail of dorsolateral sub-imbricate scales of H. fragilis (MSNG 53213) from Rahanuin region, Somalia." httpUri="https://zenodo.org/record/255737/files/figure.png" pageId="7" pageNumber="536">Fig. 5</figureCitation>
B,C,D); (2) dorsum with subimbricate scales (
<figureCitation id="C5D6EE84FF99FFCFFBE1F8E0FB5FF8EC" box="[1101,1167,1797,1822]" captionStart="FIGURE 5" captionStartId="8.[151,250,1710,1732]" captionTargetBox="[160,1427,410,1677]" captionTargetId="figure@8.[151,1436,373,1689]" captionTargetPageId="8" captionText="FIGURE 5. (A) A topotype adult male of Hemidactylus laticaudatus from Harar, Ethiopia; (B) The holotype of Hemidactylus fragilis (MZUF 707) from Bur Meldac, Somalia; (C) adult female of H. fragilis (MSNG 53212) from Dolo Odo, Ethiopia, with regenerated tail; (D) female of H. fragilis (MSNG 53213) from Rahanuin, Somalia, with partially regenerated tail; (E) detail of the tail of the previous specimen, note the enlarged tubercles on the original part of tail; (F) detail of mental region of H. frenatus (MZUF 27607) from Mogadishu, Somalia; (G) detail of mental region of H. fragilis (MSNG 53213) from Rahanuin, Somalia; (H) detail of dorsolateral sub-imbricate scales of H. fragilis (MSNG 53213) from Rahanuin region, Somalia." httpUri="https://zenodo.org/record/255737/files/figure.png" pageId="7" pageNumber="536">Fig. 5</figureCitation>
H) with a few enlarged, smooth, flat tubercles on flanks of posterior half of body, usually in one or two rows on each side; (3) no enlarged tubercles on postero-dorsal side of thighs; (4) subcaudals slightly enlarged only on first three whorls; (5) six enlarged tubercles on each whorl of the tail (
<figureCitation id="C5D6EE84FF99FFCFFD39F894FD08F878" box="[661,728,1905,1930]" captionStart="FIGURE 5" captionStartId="8.[151,250,1710,1732]" captionTargetBox="[160,1427,410,1677]" captionTargetId="figure@8.[151,1436,373,1689]" captionTargetPageId="8" captionText="FIGURE 5. (A) A topotype adult male of Hemidactylus laticaudatus from Harar, Ethiopia; (B) The holotype of Hemidactylus fragilis (MZUF 707) from Bur Meldac, Somalia; (C) adult female of H. fragilis (MSNG 53212) from Dolo Odo, Ethiopia, with regenerated tail; (D) female of H. fragilis (MSNG 53213) from Rahanuin, Somalia, with partially regenerated tail; (E) detail of the tail of the previous specimen, note the enlarged tubercles on the original part of tail; (F) detail of mental region of H. frenatus (MZUF 27607) from Mogadishu, Somalia; (G) detail of mental region of H. fragilis (MSNG 53213) from Rahanuin, Somalia; (H) detail of dorsolateral sub-imbricate scales of H. fragilis (MSNG 53213) from Rahanuin region, Somalia." httpUri="https://zenodo.org/record/255737/files/figure.png" pageId="7" pageNumber="536">Fig. 5</figureCitation>
E); (6) tail conical without basal constriction (
<figureCitation id="C5D6EE84FF99FFCFFB4AF894FAF9F878" box="[1254,1321,1905,1930]" captionStart="FIGURE 5" captionStartId="8.[151,250,1710,1732]" captionTargetBox="[160,1427,410,1677]" captionTargetId="figure@8.[151,1436,373,1689]" captionTargetPageId="8" captionText="FIGURE 5. (A) A topotype adult male of Hemidactylus laticaudatus from Harar, Ethiopia; (B) The holotype of Hemidactylus fragilis (MZUF 707) from Bur Meldac, Somalia; (C) adult female of H. fragilis (MSNG 53212) from Dolo Odo, Ethiopia, with regenerated tail; (D) female of H. fragilis (MSNG 53213) from Rahanuin, Somalia, with partially regenerated tail; (E) detail of the tail of the previous specimen, note the enlarged tubercles on the original part of tail; (F) detail of mental region of H. frenatus (MZUF 27607) from Mogadishu, Somalia; (G) detail of mental region of H. fragilis (MSNG 53213) from Rahanuin, Somalia; (H) detail of dorsolateral sub-imbricate scales of H. fragilis (MSNG 53213) from Rahanuin region, Somalia." httpUri="https://zenodo.org/record/255737/files/figure.png" pageId="7" pageNumber="536">Fig. 5</figureCitation>
D); (7) 26 preano-femoral pores in the male; (8) three lamellae under the 1st, five under the 3rd, seven under the 4th toe; (9) 78 infralabials and 89 supralabials; (10) snout forming a convex line between eye and nostril in lateral view; (11) anterior postmentals in narrow median contact (
<figureCitation id="C5D6EE84FF96FFC0FD61FF72FCC4FF42" box="[717,788,151,176]" captionStart="FIGURE 5" captionStartId="8.[151,250,1710,1732]" captionTargetBox="[160,1427,410,1677]" captionTargetId="figure@8.[151,1436,373,1689]" captionTargetPageId="8" captionText="FIGURE 5. (A) A topotype adult male of Hemidactylus laticaudatus from Harar, Ethiopia; (B) The holotype of Hemidactylus fragilis (MZUF 707) from Bur Meldac, Somalia; (C) adult female of H. fragilis (MSNG 53212) from Dolo Odo, Ethiopia, with regenerated tail; (D) female of H. fragilis (MSNG 53213) from Rahanuin, Somalia, with partially regenerated tail; (E) detail of the tail of the previous specimen, note the enlarged tubercles on the original part of tail; (F) detail of mental region of H. frenatus (MZUF 27607) from Mogadishu, Somalia; (G) detail of mental region of H. fragilis (MSNG 53213) from Rahanuin, Somalia; (H) detail of dorsolateral sub-imbricate scales of H. fragilis (MSNG 53213) from Rahanuin region, Somalia." httpUri="https://zenodo.org/record/255737/files/figure.png" pageId="8" pageNumber="537">Fig. 5</figureCitation>
G); (12) anterior postmentals only in contact with first infralabials (
<figureCitation id="C5D6EE84FF96FFC0FE89FF59FEB7FF27" box="[293,359,188,213]" captionStart="FIGURE 5" captionStartId="8.[151,250,1710,1732]" captionTargetBox="[160,1427,410,1677]" captionTargetId="figure@8.[151,1436,373,1689]" captionTargetPageId="8" captionText="FIGURE 5. (A) A topotype adult male of Hemidactylus laticaudatus from Harar, Ethiopia; (B) The holotype of Hemidactylus fragilis (MZUF 707) from Bur Meldac, Somalia; (C) adult female of H. fragilis (MSNG 53212) from Dolo Odo, Ethiopia, with regenerated tail; (D) female of H. fragilis (MSNG 53213) from Rahanuin, Somalia, with partially regenerated tail; (E) detail of the tail of the previous specimen, note the enlarged tubercles on the original part of tail; (F) detail of mental region of H. frenatus (MZUF 27607) from Mogadishu, Somalia; (G) detail of mental region of H. fragilis (MSNG 53213) from Rahanuin, Somalia; (H) detail of dorsolateral sub-imbricate scales of H. fragilis (MSNG 53213) from Rahanuin region, Somalia." httpUri="https://zenodo.org/record/255737/files/figure.png" pageId="8" pageNumber="537">Fig. 5</figureCitation>
G); (13) length to width ratio of anterior postmentals 1.351.66; (14) 89 gulars in the first row behind postmentals (
<figureCitation id="C5D6EE84FF96FFC0FE2FFF3AFE17FF0A" box="[387,455,223,248]" captionStart="FIGURE 5" captionStartId="8.[151,250,1710,1732]" captionTargetBox="[160,1427,410,1677]" captionTargetId="figure@8.[151,1436,373,1689]" captionTargetPageId="8" captionText="FIGURE 5. (A) A topotype adult male of Hemidactylus laticaudatus from Harar, Ethiopia; (B) The holotype of Hemidactylus fragilis (MZUF 707) from Bur Meldac, Somalia; (C) adult female of H. fragilis (MSNG 53212) from Dolo Odo, Ethiopia, with regenerated tail; (D) female of H. fragilis (MSNG 53213) from Rahanuin, Somalia, with partially regenerated tail; (E) detail of the tail of the previous specimen, note the enlarged tubercles on the original part of tail; (F) detail of mental region of H. frenatus (MZUF 27607) from Mogadishu, Somalia; (G) detail of mental region of H. fragilis (MSNG 53213) from Rahanuin, Somalia; (H) detail of dorsolateral sub-imbricate scales of H. fragilis (MSNG 53213) from Rahanuin region, Somalia." httpUri="https://zenodo.org/record/255737/files/figure.png" pageId="8" pageNumber="537">Fig. 5</figureCitation>
G); (15) four dark transverse bands on dorsum between axilla and groin, one on neck and one on occiput; distinct dark line from nostrils through eyes and ear openings to shoulder region and continuing diffusely to inguinal region; three dark bands on dorsum of anterior half of the tail (
<figureCitation id="C5D6EE84FF96FFC0FB11FEC2FAD0FEB2" box="[1213,1280,295,320]" captionStart="FIGURE 5" captionStartId="8.[151,250,1710,1732]" captionTargetBox="[160,1427,410,1677]" captionTargetId="figure@8.[151,1436,373,1689]" captionTargetPageId="8" captionText="FIGURE 5. (A) A topotype adult male of Hemidactylus laticaudatus from Harar, Ethiopia; (B) The holotype of Hemidactylus fragilis (MZUF 707) from Bur Meldac, Somalia; (C) adult female of H. fragilis (MSNG 53212) from Dolo Odo, Ethiopia, with regenerated tail; (D) female of H. fragilis (MSNG 53213) from Rahanuin, Somalia, with partially regenerated tail; (E) detail of the tail of the previous specimen, note the enlarged tubercles on the original part of tail; (F) detail of mental region of H. frenatus (MZUF 27607) from Mogadishu, Somalia; (G) detail of mental region of H. fragilis (MSNG 53213) from Rahanuin, Somalia; (H) detail of dorsolateral sub-imbricate scales of H. fragilis (MSNG 53213) from Rahanuin region, Somalia." httpUri="https://zenodo.org/record/255737/files/figure.png" pageId="8" pageNumber="537">Fig. 5</figureCitation>
B,C,D,E).
</paragraph>
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<emphasis id="6F992E13FF96FFC0FF3BF94BFEDDF936" bold="true" box="[151,269,1710,1732]" pageId="8" pageNumber="537">FIGURE 5</emphasis>
. (A) A topotype adult male of
<taxonomicName id="9AED8982FF96FFC0FDE5F94BFC87F936" box="[585,855,1710,1732]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="537" phylum="Chordata" rank="species" species="laticaudatus">
<emphasis id="6F992E13FF96FFC0FDE5F94BFC87F936" box="[585,855,1710,1732]" italics="true" pageId="8" pageNumber="537">Hemidactylus laticaudatus</emphasis>
</taxonomicName>
from Harar, Ethiopia; (B) The holotype of
<taxonomicName id="9AED8982FF96FFC0FABDF94BFF30F911" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="537" phylum="Chordata" rank="species" species="fragilis">
<emphasis id="6F992E13FF96FFC0FABDF94BFF30F911" italics="true" pageId="8" pageNumber="537">Hemidactylus fragilis</emphasis>
</taxonomicName>
(MZUF 707) from Bur Meldac, Somalia; (C) adult female of
<taxonomicName id="9AED8982FF96FFC0FCFEF92BFC69F911" box="[850,953,1741,1763]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="537" phylum="Chordata" rank="species" species="fragilis">
<emphasis id="6F992E13FF96FFC0FCFEF92BFC69F911" box="[850,953,1741,1763]" italics="true" pageId="8" pageNumber="537">H. fragilis</emphasis>
</taxonomicName>
(MSNG 53212) from Dolo Odo, Ethiopia, with regenerated tail; (D) female of
<taxonomicName id="9AED8982FF96FFC0FE7DF908FDE9F8F0" box="[465,569,1772,1794]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="537" phylum="Chordata" rank="species" species="fragilis">
<emphasis id="6F992E13FF96FFC0FE7DF908FDE9F8F0" box="[465,569,1772,1794]" italics="true" pageId="8" pageNumber="537">H. fragilis</emphasis>
</taxonomicName>
(MSNG 53213) from Rahanuin, Somalia, with partially regenerated tail; (E) detail of the tail of the previous specimen, note the enlarged tubercles on the original part of tail; (F) detail of mental region of
<taxonomicName id="9AED8982FF96FFC0FA2FF8EEFF39F8CD" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="537" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="6F992E13FF96FFC0FA2FF8EEFF39F8CD" italics="true" pageId="8" pageNumber="537">H. frenatus</emphasis>
</taxonomicName>
(MZUF 27607) from Mogadishu, Somalia; (G) detail of mental region of
<taxonomicName id="9AED8982FF96FFC0FC48F8CFFB9DF8CD" box="[996,1101,1833,1855]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="537" phylum="Chordata" rank="species" species="fragilis">
<emphasis id="6F992E13FF96FFC0FC48F8CFFB9DF8CD" box="[996,1101,1833,1855]" italics="true" pageId="8" pageNumber="537">H. fragilis</emphasis>
</taxonomicName>
(MSNG 53213) from Rahanuin, Somalia; (H) detail of dorsolateral sub-imbricate scales of
<taxonomicName id="9AED8982FF96FFC0FD44F8ACFC80F8AF" box="[744,848,1863,1886]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="537" phylum="Chordata" rank="species" species="fragilis">
<emphasis id="6F992E13FF96FFC0FD44F8ACFC80F8AF" box="[744,848,1863,1886]" italics="true" pageId="8" pageNumber="537">H. fragilis</emphasis>
</taxonomicName>
(MSNG 53213) from Rahanuin region, Somalia.
</paragraph>
</caption>
<paragraph id="5D52F201FF96FFC1FF6BF86FFDCBFE75" blockId="8.[151,1436,1930,2027]" lastBlockId="9.[151,1436,151,392]" lastPageId="9" lastPageNumber="538" pageId="8" pageNumber="537">
<taxonomicName id="9AED8982FF96FFC0FF6BF86FFE6DF850" box="[199,445,1930,1954]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="537" phylum="Chordata" rank="species" species="fragilis">
<emphasis id="6F992E13FF96FFC0FF6BF86FFE6DF850" box="[199,445,1930,1954]" italics="true" pageId="8" pageNumber="537">Hemidactylus fragilis</emphasis>
</taxonomicName>
can be distinguished from
<emphasis id="6F992E13FF96FFC0FCADF86EFC59F850" box="[769,905,1930,1954]" italics="true" pageId="8" pageNumber="537">
<taxonomicName id="9AED8982FF96FFC0FCADF86EFC55F850" box="[769,901,1930,1954]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="537" phylum="Chordata" rank="species" species="frenatus">H. frenatus</taxonomicName>
,
</emphasis>
with which it has been synonymized, by the following characters (condition in
<taxonomicName id="9AED8982FF96FFC0FDB6F84AFD47F835" box="[538,663,1967,1991]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="537" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="6F992E13FF96FFC0FDB6F84AFD47F835" box="[538,663,1967,1991]" italics="true" pageId="8" pageNumber="537">H. frenatus</emphasis>
</taxonomicName>
in parentheses): smaller size with max. SVL
<quantity id="9A155FE4FF96FFC0FB38F84AFB28F834" box="[1172,1272,1967,1991]" metricMagnitude="-2" metricUnit="m" metricValue="3.72" pageId="8" pageNumber="537" unit="mm" value="37.2">37.2 mm</quantity>
(
<emphasis id="6F992E13FF96FFC0FAABF84AFA9FF834" box="[1287,1359,1967,1990]" italics="true" pageId="8" pageNumber="537">versus</emphasis>
<quantity id="9A155FE4FF96FFC0FAFAF84BFF17F819" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="8" pageNumber="537" unit="mm" value="55.0" valueMax="60.0" valueMin="50.0">5060 mm</quantity>
); subimbricate dorsal scales (
<emphasis id="6F992E13FF96FFC0FDBBF836FD8EF818" box="[535,606,2003,2026]" italics="true" pageId="8" pageNumber="537">versus</emphasis>
juxtaposed); few scattered enlarged tubercles on caudal part of flanks in one or two rows on each side (
<figureCitation id="C5D6EE84FF97FFC1FE5CFF72FDE2FF42" box="[496,562,151,176]" captionStart="FIGURE 5" captionStartId="8.[151,250,1710,1732]" captionTargetBox="[160,1427,410,1677]" captionTargetId="figure@8.[151,1436,373,1689]" captionTargetPageId="8" captionText="FIGURE 5. (A) A topotype adult male of Hemidactylus laticaudatus from Harar, Ethiopia; (B) The holotype of Hemidactylus fragilis (MZUF 707) from Bur Meldac, Somalia; (C) adult female of H. fragilis (MSNG 53212) from Dolo Odo, Ethiopia, with regenerated tail; (D) female of H. fragilis (MSNG 53213) from Rahanuin, Somalia, with partially regenerated tail; (E) detail of the tail of the previous specimen, note the enlarged tubercles on the original part of tail; (F) detail of mental region of H. frenatus (MZUF 27607) from Mogadishu, Somalia; (G) detail of mental region of H. fragilis (MSNG 53213) from Rahanuin, Somalia; (H) detail of dorsolateral sub-imbricate scales of H. fragilis (MSNG 53213) from Rahanuin region, Somalia." httpUri="https://zenodo.org/record/255737/files/figure.png" pageId="9" pageNumber="538">Fig. 5</figureCitation>
C) (
<emphasis id="6F992E13FF97FFC1FDCCFF7CFD78FF42" box="[608,680,153,176]" italics="true" pageId="9" pageNumber="538">versus</emphasis>
enlarged dorsal tubercles in irregular rows, usually well developed posteriorly); larger dorsal scales,
<quantity id="9A155FE4FF97FFC1FDBBFF59FDACFF27" box="[535,636,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.048" metricValueMin="2.794" pageId="9" pageNumber="538" unit="in" value="11.5" valueMax="12.0" valueMin="11.0">1112 in</quantity>
eye diameter (
<emphasis id="6F992E13FF97FFC1FC87FF58FCA3FF26" box="[811,883,189,212]" italics="true" pageId="9" pageNumber="538">versus</emphasis>
smaller,
<quantity id="9A155FE4FF97FFC1FC72FF59FB94FF27" box="[990,1092,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" metricValueMax="4.318" metricValueMin="3.81" pageId="9" pageNumber="538" unit="in" value="16.0" valueMax="17.0" valueMin="15.0">1517 in</quantity>
eye diameter); less elongated anterior postmentals, length to width ratio 1.351.66 (
<emphasis id="6F992E13FF97FFC1FD5AFF04FCEEFF0A" box="[758,830,225,248]" italics="true" pageId="9" pageNumber="538">versus</emphasis>
1.622.6;
<figureCitation id="C5D6EE84FF97FFC1FC1AFF3AFC28FF0A" box="[950,1016,223,248]" captionStart="FIGURE 5" captionStartId="8.[151,250,1710,1732]" captionTargetBox="[160,1427,410,1677]" captionTargetId="figure@8.[151,1436,373,1689]" captionTargetPageId="8" captionText="FIGURE 5. (A) A topotype adult male of Hemidactylus laticaudatus from Harar, Ethiopia; (B) The holotype of Hemidactylus fragilis (MZUF 707) from Bur Meldac, Somalia; (C) adult female of H. fragilis (MSNG 53212) from Dolo Odo, Ethiopia, with regenerated tail; (D) female of H. fragilis (MSNG 53213) from Rahanuin, Somalia, with partially regenerated tail; (E) detail of the tail of the previous specimen, note the enlarged tubercles on the original part of tail; (F) detail of mental region of H. frenatus (MZUF 27607) from Mogadishu, Somalia; (G) detail of mental region of H. fragilis (MSNG 53213) from Rahanuin, Somalia; (H) detail of dorsolateral sub-imbricate scales of H. fragilis (MSNG 53213) from Rahanuin region, Somalia." httpUri="https://zenodo.org/record/255737/files/figure.png" pageId="9" pageNumber="538">Fig. 5</figureCitation>
F,G); 89 gular scales in the first row behind postmentals (
<emphasis id="6F992E13FF97FFC1FE2CFEE0FE18FEEE" box="[384,456,261,284]" italics="true" pageId="9" pageNumber="538">versus</emphasis>
1215;
<figureCitation id="C5D6EE84FF97FFC1FD88FEE1FDB5FEEF" box="[548,613,260,285]" captionStart="FIGURE 5" captionStartId="8.[151,250,1710,1732]" captionTargetBox="[160,1427,410,1677]" captionTargetId="figure@8.[151,1436,373,1689]" captionTargetPageId="8" captionText="FIGURE 5. (A) A topotype adult male of Hemidactylus laticaudatus from Harar, Ethiopia; (B) The holotype of Hemidactylus fragilis (MZUF 707) from Bur Meldac, Somalia; (C) adult female of H. fragilis (MSNG 53212) from Dolo Odo, Ethiopia, with regenerated tail; (D) female of H. fragilis (MSNG 53213) from Rahanuin, Somalia, with partially regenerated tail; (E) detail of the tail of the previous specimen, note the enlarged tubercles on the original part of tail; (F) detail of mental region of H. frenatus (MZUF 27607) from Mogadishu, Somalia; (G) detail of mental region of H. fragilis (MSNG 53213) from Rahanuin, Somalia; (H) detail of dorsolateral sub-imbricate scales of H. fragilis (MSNG 53213) from Rahanuin region, Somalia." httpUri="https://zenodo.org/record/255737/files/figure.png" pageId="9" pageNumber="538">Fig. 5</figureCitation>
F,G); color pattern composed of dark transverse bands on dorsum (
<emphasis id="6F992E13FF97FFC1FAF8FEE0FA4BFEEE" box="[1364,1435,261,284]" italics="true" pageId="9" pageNumber="538">versus</emphasis>
pattern lacking transverse bands, usually longitudinally striped or patternless;
<figureCitation id="C5D6EE84FF97FFC1FB9EFEC2FBA9FEB2" box="[1074,1145,295,320]" captionStart="FIGURE 5" captionStartId="8.[151,250,1710,1732]" captionTargetBox="[160,1427,410,1677]" captionTargetId="figure@8.[151,1436,373,1689]" captionTargetPageId="8" captionText="FIGURE 5. (A) A topotype adult male of Hemidactylus laticaudatus from Harar, Ethiopia; (B) The holotype of Hemidactylus fragilis (MZUF 707) from Bur Meldac, Somalia; (C) adult female of H. fragilis (MSNG 53212) from Dolo Odo, Ethiopia, with regenerated tail; (D) female of H. fragilis (MSNG 53213) from Rahanuin, Somalia, with partially regenerated tail; (E) detail of the tail of the previous specimen, note the enlarged tubercles on the original part of tail; (F) detail of mental region of H. frenatus (MZUF 27607) from Mogadishu, Somalia; (G) detail of mental region of H. fragilis (MSNG 53213) from Rahanuin, Somalia; (H) detail of dorsolateral sub-imbricate scales of H. fragilis (MSNG 53213) from Rahanuin region, Somalia." httpUri="https://zenodo.org/record/255737/files/figure.png" pageId="9" pageNumber="538">Fig. 5</figureCitation>
B,C,D,E). Morphological differences between the two species are summarized in
<tableCitation id="106FC7BAFF97FFC1FCB5FEA9FCA1FE96" box="[793,881,332,357]" captionStart="TABLE 2" captionStartId="9.[151,239,440,463]" captionTargetBox="[159,1423,626,1296]" captionTargetPageId="9" captionText="TABLE 2. Comparison of diagnostic characters of H. laevis, H. laticaudatus, H. fragilis and H. frenatus. Most distinctive characters between H. fragilis and H. frenatus are in bold. (* counted longitudinally; ** measured in the middle of each scale)." httpUri="http://table.plazi.org/id/0992A289FF97FFC1FF3BFE5DFE51FDFD" pageId="9" pageNumber="538" tableUuid="0992A289FF97FFC1FF3BFE5DFE51FDFD">Table 2</tableCitation>
. Based on this evidence we resurrect
<taxonomicName id="9AED8982FF97FFC1FA8AFEA8FA4CFE96" box="[1318,1436,332,356]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="fragilis">
<emphasis id="6F992E13FF97FFC1FA8AFEA8FA4CFE96" box="[1318,1436,332,356]" italics="true" pageId="9" pageNumber="538">H. fragilis</emphasis>
</taxonomicName>
from the synonymy of
<taxonomicName id="9AED8982FF97FFC1FE3BFE94FDC5FE7A" box="[407,533,368,392]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="6F992E13FF97FFC1FE3BFE94FDC5FE7A" box="[407,533,368,392]" italics="true" pageId="9" pageNumber="538">H. frenatus</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="0992A289FF97FFC1FF3BFE5DFE51FDFD" ID-Table-UUID="0992A289FF97FFC1FF3BFE5DFE51FDFD" httpUri="http://table.plazi.org/id/0992A289FF97FFC1FF3BFE5DFE51FDFD" pageId="9" pageNumber="538" targetBox="[159,1423,626,1296]" targetIsTable="true" targetPageId="9">
<paragraph id="5D52F201FF97FFC1FF3BFE5DFE51FDFD" blockId="9.[151,1436,439,527]" pageId="9" pageNumber="538">
<emphasis id="6F992E13FF97FFC1FF3BFE5DFEC0FE3D" bold="true" box="[151,272,440,463]" pageId="9" pageNumber="538">TABLE 2.</emphasis>
Comparison of diagnostic characters of
<emphasis id="6F992E13FF97FFC1FD4BFE5DFB43FE3D" box="[743,1171,439,463]" italics="true" pageId="9" pageNumber="538">
<taxonomicName id="9AED8982FF97FFC1FD4BFE5DFC9FFE3D" box="[743,847,439,463]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="laevis">H. laevis</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FCCCFE5DFBDDFE3D" box="[864,1037,439,463]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="laticaudatus">H. laticaudatus</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FBB3FE5DFB43FE3D" box="[1055,1171,439,463]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="fragilis">H. fragilis</taxonomicName>
</emphasis>
and
<taxonomicName id="9AED8982FF97FFC1FB78FE5DFA81FE3D" box="[1236,1361,439,463]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="6F992E13FF97FFC1FB78FE5DFA81FE3D" box="[1236,1361,439,463]" italics="true" pageId="9" pageNumber="538">H. frenatus</emphasis>
</taxonomicName>
. Most distinctive characters between
<taxonomicName id="9AED8982FF97FFC1FE45FE3DFD8AFE1D" box="[489,602,471,495]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="fragilis">
<emphasis id="6F992E13FF97FFC1FE45FE3DFD8AFE1D" box="[489,602,471,495]" italics="true" pageId="9" pageNumber="538">H. fragilis</emphasis>
</taxonomicName>
and
<taxonomicName id="9AED8982FF97FFC1FD38FE3DFCDEFE1D" box="[660,782,471,495]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="6F992E13FF97FFC1FD38FE3DFCDEFE1D" box="[660,782,471,495]" italics="true" pageId="9" pageNumber="538">H. frenatus</emphasis>
</taxonomicName>
are in bold. (* counted longitudinally; ** measured in the middle of each scale).
</paragraph>
</caption>
<paragraph id="5D52F201FF97FFC1FE52FDCEFCE5FDAD" blockId="9.[510,1408,554,607]" pageId="9" pageNumber="538">
<taxonomicName id="9AED8982FF97FFC1FE52FDCEFD89FDB2" box="[510,601,554,576]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="laevis">
<emphasis id="6F992E13FF97FFC1FE52FDCEFD89FDB2" bold="true" box="[510,601,554,576]" italics="true" pageId="9" pageNumber="538">H. laevis</emphasis>
</taxonomicName>
(n=2)
<emphasis id="6F992E13FF97FFC1FD5DFDCEFB92FDB2" bold="true" box="[753,1090,554,576]" italics="true" pageId="9" pageNumber="538">
<taxonomicName id="9AED8982FF97FFC1FD5DFDCEFC41FDB2" box="[753,913,554,576]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="laticaudatus">H. laticaudatus</taxonomicName>
<taxonomicName id="9AED8982FF97FFC1FC7BFDCEFB92FDB2" box="[983,1090,554,576]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="fragilis">H. fragilis</taxonomicName>
</emphasis>
(n=3)
<taxonomicName id="9AED8982FF97FFC1FB11FDCEFAE4FDB2" box="[1213,1332,554,576]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="6F992E13FF97FFC1FB11FDCEFAE4FDB2" bold="true" box="[1213,1332,554,576]" italics="true" pageId="9" pageNumber="538">H. frenatus</emphasis>
</taxonomicName>
(n=12) (n=22)
</paragraph>
<paragraph id="5D52F201FF97FFC1FF33FD97FB09FB00" pageId="9" pageNumber="538">
<table id="2FED00A1FF970037FF33FD97FA5FFAE2" box="[159,1423,626,1296]" gridcols="5" gridrows="10" pageId="9" pageNumber="538">
<tr id="E3DDF043FF970037FF33FD97FA5FFD7A" box="[159,1423,626,648]" gridrow="0" pageId="9" pageNumber="538">
<th id="A00C993FFF970037FF33FD97FE38FD7A" box="[159,488,626,648]" gridcol="0" gridrow="0" pageId="9" pageNumber="538">Maximum SVL (mm)</th>
<th id="A00C993FFF970037FE52FD97FD1DFD7A" box="[510,717,626,648]" gridcol="1" gridrow="0" pageId="9" pageNumber="538">39</th>
<th id="A00C993FFF970037FD5DFD97FC70FD7A" box="[753,928,626,648]" gridcol="2" gridrow="0" pageId="9" pageNumber="538">62.0</th>
<th id="A00C993FFF970037FC7BFD97FB78FD7A" box="[983,1192,626,648]" gridcol="3" gridrow="0" pageId="9" pageNumber="538">
<emphasis id="6F992E13FF97FFC1FC7BFD97FBD3FD7A" bold="true" box="[983,1027,626,648]" pageId="9" pageNumber="538">37.2</emphasis>
</th>
<th id="A00C993FFF970037FB11FD97FA5FFD7A" box="[1213,1423,626,648]" gridcol="4" gridrow="0" pageId="9" pageNumber="538">
<emphasis id="6F992E13FF97FFC1FB11FD97FB39FD7A" bold="true" box="[1213,1257,626,648]" pageId="9" pageNumber="538">60.0</emphasis>
</th>
</tr>
<tr id="E3DDF043FF970037FF33FD79FA5FFCDE" box="[159,1423,668,812]" gridrow="1" pageId="9" pageNumber="538">
<th id="A00C993FFF970037FF33FD79FE38FCDE" box="[159,488,668,812]" gridcol="0" gridrow="1" pageId="9" pageNumber="538">Enlarged tubercles on dorsum</th>
<td id="A00C993FFF970037FE52FD79FD1DFCDE" box="[510,717,668,812]" gridcol="1" gridrow="1" pageId="9" pageNumber="538">Developed, but smaller and in fewer regular rows</td>
<td id="A00C993FFF970037FD5DFD79FC70FCDE" box="[753,928,668,812]" gridcol="2" gridrow="1" pageId="9" pageNumber="538">Well developed, higher number of regular rows</td>
<td id="A00C993FFF970037FC7BFD79FB78FCDE" box="[983,1192,668,812]" gridcol="3" gridrow="1" pageId="9" pageNumber="538">
<emphasis id="6F992E13FF97FFC1FC7BFD79FBD1FCDE" bold="true" pageId="9" pageNumber="538">Few scattered on flanks in second half of body, in one or two rows on each side</emphasis>
</td>
<td id="A00C993FFF970037FB11FD79FA5FFCDE" box="[1213,1423,668,812]" gridcol="4" gridrow="1" pageId="9" pageNumber="538">
<emphasis id="6F992E13FF97FFC1FB11FD79FAF3FCFC" bold="true" pageId="9" pageNumber="538">Rather irregular rows, usually well developed in second half body</emphasis>
</td>
</tr>
<tr id="E3DDF043FF970037FF33FCA5FA5FFCA4" box="[159,1423,832,854]" gridrow="2" pageId="9" pageNumber="538">
<th id="A00C993FFF970037FF33FCA5FE38FCA4" box="[159,488,832,854]" gridcol="0" gridrow="2" pageId="9" pageNumber="538">Dorsal lepidosis</th>
<td id="A00C993FFF970037FE52FCA5FD1DFCA4" box="[510,717,832,854]" gridcol="1" gridrow="2" pageId="9" pageNumber="538">Juxtaposed</td>
<td id="A00C993FFF970037FD5DFCA5FC70FCA4" box="[753,928,832,854]" gridcol="2" gridrow="2" pageId="9" pageNumber="538">Juxtaposed</td>
<td id="A00C993FFF970037FC7BFCA5FB78FCA4" box="[983,1192,832,854]" gridcol="3" gridrow="2" pageId="9" pageNumber="538">
<emphasis id="6F992E13FF97FFC1FC7BFCA5FBA2FCA4" bold="true" box="[983,1138,832,854]" pageId="9" pageNumber="538">Sub-imbricate</emphasis>
</td>
<td id="A00C993FFF970037FB11FCA5FA5FFCA4" box="[1213,1423,832,854]" gridcol="4" gridrow="2" pageId="9" pageNumber="538">
<emphasis id="6F992E13FF97FFC1FB11FCA5FAE7FCA4" bold="true" box="[1213,1335,832,854]" pageId="9" pageNumber="538">Juxtaposed</emphasis>
</td>
</tr>
<tr id="E3DDF043FF970037FF33FC8CFA5FFC6C" box="[159,1423,873,926]" gridrow="3" pageId="9" pageNumber="538">
<th id="A00C993FFF970037FF33FC8CFE38FC6C" box="[159,488,873,926]" gridcol="0" gridrow="3" pageId="9" pageNumber="538">Number of dorsals in the eye diameter (*)</th>
<td id="A00C993FFF970037FE52FC8CFD1DFC6C" box="[510,717,873,926]" gridcol="1" gridrow="3" pageId="9" pageNumber="538">1112</td>
<td id="A00C993FFF970037FD5DFC8CFC70FC6C" box="[753,928,873,926]" gridcol="2" gridrow="3" pageId="9" pageNumber="538">911</td>
<td id="A00C993FFF970037FC7BFC8CFB78FC6C" box="[983,1192,873,926]" gridcol="3" gridrow="3" pageId="9" pageNumber="538">
<emphasis id="6F992E13FF97FFC1FC7BFC8CFBC5FC8D" bold="true" box="[983,1045,873,895]" pageId="9" pageNumber="538">1112</emphasis>
</td>
<td id="A00C993FFF970037FB11FC8CFA5FFC6C" box="[1213,1423,873,926]" gridcol="4" gridrow="3" pageId="9" pageNumber="538">
<emphasis id="6F992E13FF97FFC1FB11FC8CFB2CFC8D" bold="true" box="[1213,1276,873,895]" pageId="9" pageNumber="538">1517</emphasis>
</td>
</tr>
<tr id="E3DDF043FF970037FF33FC54FA5FFC14" box="[159,1423,945,998]" gridrow="4" pageId="9" pageNumber="538">
<th id="A00C993FFF970037FF33FC54FE38FC14" box="[159,488,945,998]" gridcol="0" gridrow="4" pageId="9" pageNumber="538">Tubercles on postero-dorsal side of thighs</th>
<td id="A00C993FFF970037FE52FC54FD1DFC14" box="[510,717,945,998]" gridcol="1" gridrow="4" pageId="9" pageNumber="538">No</td>
<td id="A00C993FFF970037FD5DFC54FC70FC14" box="[753,928,945,998]" gridcol="2" gridrow="4" pageId="9" pageNumber="538">Yes</td>
<td id="A00C993FFF970037FC7BFC54FB78FC14" box="[983,1192,945,998]" gridcol="3" gridrow="4" pageId="9" pageNumber="538">No</td>
<td id="A00C993FFF970037FB11FC54FA5FFC14" box="[1213,1423,945,998]" gridcol="4" gridrow="4" pageId="9" pageNumber="538">No</td>
</tr>
<tr id="E3DDF043FF970037FF33FC1CFA5FFBFD" box="[159,1423,1017,1039]" gridrow="5" pageId="9" pageNumber="538">
<th id="A00C993FFF970037FF33FC1CFE38FBFD" box="[159,488,1017,1039]" gridcol="0" gridrow="5" pageId="9" pageNumber="538">Enlarged tubercles on tail</th>
<td id="A00C993FFF970037FE52FC1CFD1DFBFD" box="[510,717,1017,1039]" gridcol="1" gridrow="5" pageId="9" pageNumber="538">No</td>
<td id="A00C993FFF970037FD5DFC1CFC70FBFD" box="[753,928,1017,1039]" gridcol="2" gridrow="5" pageId="9" pageNumber="538">Yes</td>
<td id="A00C993FFF970037FC7BFC1CFB78FBFD" box="[983,1192,1017,1039]" gridcol="3" gridrow="5" pageId="9" pageNumber="538">Yes</td>
<td id="A00C993FFF970037FB11FC1CFA5FFBFD" box="[1213,1423,1017,1039]" gridcol="4" gridrow="5" pageId="9" pageNumber="538">Yes</td>
</tr>
<tr id="E3DDF043FF970037FF33FBC7FA5FFBCA" box="[159,1423,1058,1080]" gridrow="6" pageId="9" pageNumber="538">
<th id="A00C993FFF970037FF33FBC7FE38FBCA" box="[159,488,1058,1080]" gridcol="0" gridrow="6" pageId="9" pageNumber="538">Tail with basal constriction</th>
<td id="A00C993FFF970037FE52FBC7FD1DFBCA" box="[510,717,1058,1080]" gridcol="1" gridrow="6" pageId="9" pageNumber="538">Yes</td>
<td id="A00C993FFF970037FD5DFBC7FC70FBCA" box="[753,928,1058,1080]" gridcol="2" gridrow="6" pageId="9" pageNumber="538">Yes</td>
<td id="A00C993FFF970037FC7BFBC7FB78FBCA" box="[983,1192,1058,1080]" gridcol="3" gridrow="6" pageId="9" pageNumber="538">No</td>
<td id="A00C993FFF970037FB11FBC7FA5FFBCA" box="[1213,1423,1058,1080]" gridcol="4" gridrow="6" pageId="9" pageNumber="538">No</td>
</tr>
<tr id="E3DDF043FF970037FF33FBA9FA5FFB72" box="[159,1423,1100,1152]" gridrow="7" pageId="9" pageNumber="538">
<th id="A00C993FFF970037FF33FBA9FE38FB72" box="[159,488,1100,1152]" gridcol="0" gridrow="7" pageId="9" pageNumber="538">Length to width ratio of anterior postmentals (**)</th>
<td id="A00C993FFF970037FE52FBA9FD1DFB72" box="[510,717,1100,1152]" gridcol="1" gridrow="7" pageId="9" pageNumber="538">1.541.77</td>
<td id="A00C993FFF970037FD5DFBA9FC70FB72" box="[753,928,1100,1152]" gridcol="2" gridrow="7" pageId="9" pageNumber="538">1.351.69</td>
<td id="A00C993FFF970037FC7BFBA9FB78FB72" box="[983,1192,1100,1152]" gridcol="3" gridrow="7" pageId="9" pageNumber="538">
<emphasis id="6F992E13FF97FFC1FC7BFBA9FBECFB90" bold="true" box="[983,1084,1100,1122]" pageId="9" pageNumber="538">1.351.66</emphasis>
</td>
<td id="A00C993FFF970037FB11FBA9FA5FFB72" box="[1213,1423,1100,1152]" gridcol="4" gridrow="7" pageId="9" pageNumber="538">
<emphasis id="6F992E13FF97FFC1FB11FBA9FAC6FB90" bold="true" box="[1213,1302,1100,1122]" pageId="9" pageNumber="538">1.622.6</emphasis>
</td>
</tr>
<tr id="E3DDF043FF970037FF33FB71FA5FFB3A" box="[159,1423,1172,1224]" gridrow="8" pageId="9" pageNumber="538">
<th id="A00C993FFF970037FF33FB71FE38FB3A" box="[159,488,1172,1224]" gridcol="0" gridrow="8" pageId="9" pageNumber="538">Number of gulars in first row behind postmentals</th>
<td id="A00C993FFF970037FE52FB71FD1DFB3A" box="[510,717,1172,1224]" gridcol="1" gridrow="8" pageId="9" pageNumber="538">812</td>
<td id="A00C993FFF970037FD5DFB71FC70FB3A" box="[753,928,1172,1224]" gridcol="2" gridrow="8" pageId="9" pageNumber="538">812</td>
<td id="A00C993FFF970037FC7BFB71FB78FB3A" box="[983,1192,1172,1224]" gridcol="3" gridrow="8" pageId="9" pageNumber="538">
<emphasis id="6F992E13FF97FFC1FC7BFB71FC2DFB58" bold="true" box="[983,1021,1172,1194]" pageId="9" pageNumber="538">89</emphasis>
</td>
<td id="A00C993FFF970037FB11FB71FA5FFB3A" box="[1213,1423,1172,1224]" gridcol="4" gridrow="8" pageId="9" pageNumber="538">
<emphasis id="6F992E13FF97FFC1FB11FB71FB2CFB58" bold="true" box="[1213,1276,1172,1194]" pageId="9" pageNumber="538">1215</emphasis>
</td>
</tr>
<tr id="E3DDF043FF970037FF33FB39FA5FFAE2" box="[159,1423,1244,1296]" gridrow="9" pageId="9" pageNumber="538">
<th id="A00C993FFF970037FF33FB39FE38FAE2" box="[159,488,1244,1296]" gridcol="0" gridrow="9" pageId="9" pageNumber="538">Dark transverse bands on dorsum</th>
<td id="A00C993FFF970037FE52FB39FD1DFAE2" box="[510,717,1244,1296]" gridcol="1" gridrow="9" pageId="9" pageNumber="538">Yes</td>
<td id="A00C993FFF970037FD5DFB39FC70FAE2" box="[753,928,1244,1296]" gridcol="2" gridrow="9" pageId="9" pageNumber="538">Yes</td>
<td id="A00C993FFF970037FC7BFB39FB78FAE2" box="[983,1192,1244,1296]" gridcol="3" gridrow="9" pageId="9" pageNumber="538">Yes</td>
<td id="A00C993FFF970037FB11FB39FA5FFAE2" box="[1213,1423,1244,1296]" gridcol="4" gridrow="9" pageId="9" pageNumber="538">No</td>
</tr>
</table>
</paragraph>
<paragraph id="5D52F201FF97FFC1FF6BFAB5FF0BF876" blockId="9.[151,1437,1359,1924]" pageId="9" pageNumber="538">
<taxonomicName id="9AED8982FF97FFC1FF6BFAB5FE68FA9A" box="[199,440,1360,1384]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="fragilis">
<emphasis id="6F992E13FF97FFC1FF6BFAB5FE68FA9A" box="[199,440,1360,1384]" italics="true" pageId="9" pageNumber="538">Hemidactylus fragilis</emphasis>
</taxonomicName>
differs as follows from the species that can be found or are expected to occur in the Horn of Africa:
<emphasis id="6F992E13FF97FFC1FEB8FA91FAEDFA26" italics="true" pageId="9" pageNumber="538">
<taxonomicName id="9AED8982FF97FFC1FEB8FA91FDFAFA7E" box="[276,554,1396,1420]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="angulatus">Hemidactylus angulatus</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FD96FA90FD65FA7E" box="[570,693,1396,1420]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="arnoldi">H. arnoldi</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FD69FA90FCBFFA7E" box="[709,879,1396,1420]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="awashensis">H. awashensis</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FCD2FA90FBDAFA7E" box="[894,1034,1396,1420]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="barbierii">H. barbierii</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FBB6FA90FB6BFA7E" box="[1050,1211,1396,1420]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="barodanus">H. barodanus</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FB66FA90FAA1FA7E" box="[1226,1393,1396,1420]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="bavazzanoi">H. bavazzanoi</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FA2DFA90FF3BFA5D" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="citernii">H. citernii</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FF56FA7CFE52FA42" box="[250,386,1432,1456]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="granchii">H. granchii</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FE3EFA7CFDC7FA42" box="[402,535,1432,1456]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="jubensis">H. jubensis</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FD8AFA7CFD41FA42" box="[550,657,1432,1456]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="laevis">H. laevis</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FD0CFA7CFC84FA42" box="[672,852,1432,1456]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="laticaudatus">H. laticaudatus</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FCCFFA7CFC3FFA42" box="[867,1007,1432,1456]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="mabouia">H. mabouia</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FC52FA7CFB63FA42" box="[1022,1203,1432,1456]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="macropholis">H. macropholis</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FB6EFA7CFAA1FA5D" box="[1218,1393,1432,1456]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="mercatorius">H. mercatorius</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FA2DFA7CFECBFA26" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="mrimaensis">H. mrimaensis</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FE86FA58FE3EFA26" box="[298,494,1468,1492]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="platycephalus">H. platycephalus</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FE51FA58FD5DFA26" box="[509,653,1469,1492]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="puccionii">H. puccionii</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FD30FA58FCF4FA26" box="[668,804,1468,1492]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="robustus">H. robustus</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FC9FFA58FC63FA26" box="[819,947,1468,1492]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="ruspolii">H. ruspolii</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FC6EFA58FB91FA26" box="[962,1089,1469,1492]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="sinaitus">H. sinaitus</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FBFCFA58FB6DFA26" box="[1104,1213,1468,1492]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="smithi">H. smithi</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FB60FA58FAEDFA26" box="[1228,1341,1468,1492]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="taylori">H. taylori</taxonomicName>
</emphasis>
, and
<taxonomicName id="9AED8982FF97FFC1FA2DFA58FE46FA05" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="subSpecies" species="yerburii" subSpecies="pauciporosus">
<emphasis id="6F992E13FF97FFC1FA2DFA58FE46FA05" italics="true" pageId="9" pageNumber="538">H. yerburii pauciporosus</emphasis>
</taxonomicName>
have enlarged dorsal tubercles on the whole dorsum, nape and head.
<taxonomicName id="9AED8982FF97FFC1FAAEFA05FED7F9EE" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="somalicus">
<emphasis id="6F992E13FF97FFC1FAAEFA05FED7F9EE" italics="true" pageId="9" pageNumber="538">Hemidactylus somalicus</emphasis>
</taxonomicName>
does not have depressed body habitus, has very small proximal subdigital lamellae, scarcely larger than the granules on the sole, tail without enlarged tubercles, and first supralabial excluded from nostril.
<emphasis id="6F992E13FF97FFC1FAAEF9CDFAE5F996" italics="true" pageId="9" pageNumber="538">
<taxonomicName id="9AED8982FF97FFC1FAAEF9CDFEE9F996" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="albopunctatus">Hemidactylus albopunctatus</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FEE2F9A8FE0DF996" box="[334,477,1612,1636]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="funaiolii">H. funaiolii</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FE5EF9A8FDA7F996" box="[498,631,1612,1636]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="isolepis">H. isolepis</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FD21F9A8FCC9F996" box="[653,793,1612,1636]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="klauberi">H. klauberi</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FC83F9A8FC1AF996" box="[815,970,1612,1636]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="megalops">H. megalops</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FC4CF9A8FBA7F996" box="[992,1143,1612,1636]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="modestus">H. modestus</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FB21F9A8FAE2F996" box="[1165,1330,1612,1636]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="ophiolepis">H. ophiolepis</taxonomicName>
,
</emphasis>
and
<taxonomicName id="9AED8982FF97FFC1FA2DF9A8FEE4F97A" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="ophiolepoides">
<emphasis id="6F992E13FF97FFC1FA2DF9A8FEE4F97A" italics="true" pageId="9" pageNumber="538">H. ophiolepoides</emphasis>
</taxonomicName>
have homogenous dorsal scales and weakly developed subdigital lamellae, head short and not depressed, and tail without enlarged tubercles.
<emphasis id="6F992E13FF97FFC1FD01F971FBA3F95E" box="[685,1139,1684,1708]" italics="true" pageId="9" pageNumber="538">
<taxonomicName id="9AED8982FF97FFC1FD01F971FC66F95E" box="[685,950,1684,1708]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="barbouri">Hemidactylus barbouri</taxonomicName>
,
<taxonomicName id="9AED8982FF97FFC1FC6FF970FBA3F95E" box="[963,1139,1684,1708]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="squamulatus">H. squamulatus</taxonomicName>
</emphasis>
, and
<taxonomicName id="9AED8982FF97FFC1FB18F970FA8EF95E" box="[1204,1374,1684,1708]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="tropidolepis">
<emphasis id="6F992E13FF97FFC1FB18F970FA8EF95E" box="[1204,1374,1684,1708]" italics="true" pageId="9" pageNumber="538">H. tropidolepis</emphasis>
</taxonomicName>
have heterogeneous dorsal scales without tubercles, some or all scales are keeled, well developed enlarged tubercles on nape and occiput, and head and body not depressed.
<taxonomicName id="9AED8982FF97FFC1FD5CF939FBDBF906" box="[752,1035,1756,1780]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="flaviviridis">
<emphasis id="6F992E13FF97FFC1FD5CF939FBDBF906" box="[752,1035,1756,1780]" italics="true" pageId="9" pageNumber="538">Hemidactylus flaviviridis</emphasis>
</taxonomicName>
is larger (maximum SVL about
<quantity id="9A155FE4FF97FFC1FA2CF939FF17F8EA" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="9" pageNumber="538" unit="mm" value="95.0">95 mm</quantity>
), has swollen, carrot-shaped tail clearly constricted at its base, and dorsal scales small, granular and juxtaposed.
<taxonomicName id="9AED8982FF97FFC1FEB3F8C1FDD2F8CE" box="[287,514,1828,1852]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="538" phylum="Chordata" rank="species" species="curlei">
<emphasis id="6F992E13FF97FFC1FEB3F8C1FDD2F8CE" box="[287,514,1828,1852]" italics="true" pageId="9" pageNumber="538">Hemidactylus curlei</emphasis>
</taxonomicName>
has swollen, carrot-shaped tail clearly constricted at its base, scales on tail large, sub-imbricate and without enlarged tubercles, no enlarged tubercles in sacral region, and males with 4 precloacal pores.
</paragraph>
</subSubSection>
</treatment>
</document>