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<document ID-CLB-Dataset="57144" ID-DOI="http://dx.doi.org/10.3897/zookeys.1168.104844" ID-GBIF-Dataset="67f9e0f5-21e5-478f-8ee2-1924430bb454" ID-PMC="PMC10843360" ID-Pensoft-Pub="1313-2970-1168-231" ID-Pensoft-UUID="6928794CB97051F9AAE3B64050E5FAF3" ID-PubMed="38318436" ID-ZooBank="28A21391A4334E808845E5C0996EFAF0" ModsDocID="1313-2970-1168-231" checkinTime="1688439534387" checkinUser="pensoft" docAuthor="Kaltenbach, Thomas, Kluge, Nikita J. &amp; Gattolliat, Jean-Luc" docDate="2023" docId="6E4FA03D5D32522DA54FBD02C733CE43" docLanguage="en" docName="ZooKeys 1168: 231-266" docOrigin="ZooKeys 1168" docPubDate="2023-07-03" docSource="http://dx.doi.org/10.3897/zookeys.1168.104844" docTitle="Arcobaetis sumbawensis Kaltenbach &amp; Kluge &amp; Gattolliat 2023, sp. nov." docType="treatment" docUuid="7A575C67-8D0D-4A93-BB2B-BF7669B229D5" docUuidSource="ZooBank" docVersion="5" id="6928794CB97051F9AAE3B64050E5FAF3" lastPageNumber="231" masterDocId="6928794CB97051F9AAE3B64050E5FAF3" masterDocTitle="A new, widespread genus of Baetidae from South Asia (Insecta, Ephemeroptera)" masterLastPageNumber="266" masterPageNumber="231" pageNumber="231" updateTime="1732691564844" updateUser="ExternalLinkService">
<mods:mods id="9A911A2488A0EDA01233D30010F3FCE9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8CCDFF1E2CC922F20CC678E0F7F93625">
<mods:title id="1B610550D6C2375D50FADF5FF0EF89EF">A new, widespread genus of Baetidae from South Asia (Insecta, Ephemeroptera)</mods:title>
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<mods:name id="1C96C03A43FD4EE189A211FEF72AA9C6" type="personal">
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<mods:namePart id="B74B9BE72FB4CCD4016C227D7E05ECD4">Kaltenbach, Thomas</mods:namePart>
<mods:nameIdentifier id="AB0272850E2C82882CA717A97BBAE17F" type="ORCID">https://orcid.org/0000-0001-8052-0388</mods:nameIdentifier>
<mods:affiliation id="0CEB0911A3ED4DCB66181E87EEA89271">Museum cantonal des Sciences Naturelles, Departement de zoologie, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
<mods:nameIdentifier id="152C15C7BD91B6D5D01C02BE59325C31" type="email">thomas.kaltenbach@bluewin.ch</mods:nameIdentifier>
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<mods:namePart id="9CA5D54844C3ACC14EC275D3D87C3EA7">Kluge, Nikita J.</mods:namePart>
<mods:nameIdentifier id="25D319BD524ADB338A54F045B1848A95" type="ORCID">https://orcid.org/0000-0001-9741-7790</mods:nameIdentifier>
<mods:affiliation id="A43A29AE5D3FF37FAE934708DDD8DDB2">Department of Entomology, Biological Faculty, Saint-Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia</mods:affiliation>
</mods:name>
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<mods:namePart id="3078DAFEA4B5CD22584F2E05B2DCFC01">Gattolliat, Jean-Luc</mods:namePart>
<mods:nameIdentifier id="0D40A19C3ADC16C1ABA929CD31D10AEB" type="ORCID">https://orcid.org/0000-0001-5873-5083</mods:nameIdentifier>
<mods:affiliation id="EA1047EF114EC3AA8C00061FA7240BA6">Museum cantonal des Sciences Naturelles, Departement de zoologie, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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<mods:title id="9182C11AF0033A2986661D148AF21EE2">ZooKeys</mods:title>
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<mods:part id="07FD751633E6190BDE690C5491F8B44A">
<mods:date id="2B144EE0FC4E69D7258F33233449AD76">2023</mods:date>
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<mods:number id="F2CA6EE7B45550723D7825B35CE695AC">2023-07-03</mods:number>
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<mods:number id="98060A2AD5A1D420FE3271A5CEF93045">1168</mods:number>
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<mods:classification id="39F2D57F5F2470F47FD93A8931CDC2C0">journal article</mods:classification>
<mods:identifier id="7FC5E6A9F0B0E768B00463E15FEA9AF1" type="DOI">http://dx.doi.org/10.3897/zookeys.1168.104844</mods:identifier>
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<treatment id="6E4FA03D5D32522DA54FBD02C733CE43" ID-GBIF-Taxon="210808360" LSID="urn:lsid:zoobank.org:act:7A575C67-8D0D-4A93-BB2B-BF7669B229D5" httpUri="http://treatment.plazi.org/id/6E4FA03D5D32522DA54FBD02C733CE43" lastPageNumber="231" pageId="0" pageNumber="231" scope_class="Insecta" scope_family="Baetidae" scope_order="Ephemeroptera">
<subSubSection id="F8174D0AE70781DEF217790966D9B184" pageId="0" pageNumber="231" type="nomenclature">
<paragraph id="A485BE271CD183A53C37DC3B706E7594" pageId="0" pageNumber="231">
<taxonomicName id="7DD52BA5BCCD572BEC4F6FD7A1AA9904" LSID="https://zoobank.org/7A575C67-8D0D-4A93-BB2B-BF7669B229D5" authority="Kaltenbach &amp; Kluge &amp; Gattolliat, 2023" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Arcobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arcobaetis sumbawensis" order="Ephemeroptera" pageId="0" pageNumber="231" phylum="Arthropoda" rank="species" species="sumbawensis" status="sp. nov.">Arcobaetis sumbawensis</taxonomicName>
<taxonomicNameLabel id="4B4C44610698AB94C7C876DADA2CE349" pageId="0" pageNumber="231">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="0D188BFA5368F36EC329B1A4012BA8CF" pageId="0" pageNumber="231" type="description">
<paragraph id="4A8C06565C60530858DFE26A44C5FDA1" pageId="0" pageNumber="231">
<figureCitation id="2F790E2520745B2DC9EF81ACA8F6E4F1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Arcobaetis sumbawensis sp. nov., larva, habitus (holotype) a dorsal view b lateral view c ventral view. Scale bars: 1 mm" figureDoi="10.3897/zookeys.1168.104844.figure1" httpUri="https://binary.pensoft.net/fig/872203" pageId="0" pageNumber="231">Figs 1</figureCitation>
<figureCitation id="8DC6FF1EE96419EA38610563A7F37475" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">, 2</figureCitation>
<figureCitation id="B89B9E27ED219787F31DB991B2D302E1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct." figureDoi="10.3897/zookeys.1168.104844.figure3" httpUri="https://binary.pensoft.net/fig/872205" pageId="0" pageNumber="231">, 3</figureCitation>
<figureCitation id="3F3FFF0EAF3B16C8ACC3A4F282ECC2D4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Arcobaetis sumbawensis sp. nov., larva a foreleg b middle leg c hind leg d subimaginal gonostylus developing under cuticle of last instar male larva e head. Scale bars: 0.05 mm (e); 0.1 mm (d)." figureDoi="10.3897/zookeys.1168.104844.figure4" httpUri="https://binary.pensoft.net/fig/872206" pageId="0" pageNumber="231">, 4</figureCitation>
<figureCitation id="10E372FF434D0955E1DD519BDFCAC385" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Arcobaetis sumbawensis sp. nov., larva, SEM pictures a fore claw, lateroventral view b middle claw, anterior view c middle claw, posterior view. Arrows: subapical setae. Scale bars: 0.01 mm." figureDoi="10.3897/zookeys.1168.104844.figure5" httpUri="https://binary.pensoft.net/fig/872207" pageId="0" pageNumber="231">, 5</figureCitation>
<figureCitation id="2683387F4F59C37BD4D2AE9F0712DCF7" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Arcobaetis gen. nov., distribution." figureDoi="10.3897/zookeys.1168.104844.figure19" httpUri="https://binary.pensoft.net/fig/872221" pageId="0" pageNumber="231">, 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4A4C8986907466D77BEE179F70222182" pageId="0" pageNumber="231" type="type material">
<paragraph id="BDC136121323E428C3DB0B1C730D20C0" pageId="0" pageNumber="231">Type material.</paragraph>
<paragraph id="B40CB8B63F46E600C11B8BD2ECD20D13" pageId="0" pageNumber="231">
<emphasis id="122E19ED4D8C4F166B4DC46C94F5022F" bold="true" pageId="0" pageNumber="231">
<emphasis id="AB070641BAA6B865455A484B5AD8407E" italics="true" pageId="0" pageNumber="231">Holotype</emphasis>
.
</emphasis>
Indonesia • male larva (last instar); Sumbawa, Batu Dulang, Mt. Batu Pasak, forest streams;
<geoCoordinate id="90A3ED3EF1034F8AF4CB98A3211ED865" degrees="08" direction="south" minutes="37" orientation="latitude" precision="15" seconds="42" value="-8.628333">08°37'42&quot;S</geoCoordinate>
,
<geoCoordinate id="1A018B2EB1A8ACAE7E145199BC6101F6" degrees="117" direction="east" minutes="15" orientation="longitude" precision="15" seconds="27" value="117.2575">117°15'27&quot;E</geoCoordinate>
, SUMB09; 1380 m; 17.ix.2011; leg. M. Balke; on slide; GBIFCH00975680; MZB.
<emphasis id="AE4688D37E5BAE09AB2A9D8532785496" bold="true" pageId="0" pageNumber="231">
<emphasis id="06FD9FB50E50F332074DA3F0E3765C45" italics="true" pageId="0" pageNumber="231">Paratypes</emphasis>
.
</emphasis>
9 larvae; same data as holotype; 4 on slides; GBIFCH00692615, GBIFCH00975688, GBIFCH00592652, GBIFCH00592653; 5 in alcohol; GenBank OQ699910, OQ699911; GBIFCH00975695, GBIFCH00975689, GBIFCH00975690, GBIFCH00975694; MZL.
</paragraph>
</subSubSection>
<subSubSection id="C073AD18E65B5BF9B73988CDDB564256" pageId="0" pageNumber="231" type="diagnosis">
<paragraph id="2C6F7CCC878E68E3EA4EE386B2D69B1B" pageId="0" pageNumber="231">Diagnosis.</paragraph>
<paragraph id="5F5AF262ADBA63BDD64C69ACAE7CBE90" pageId="0" pageNumber="231">
<emphasis id="2A714520E75667F974AC46BD91F5268F" bold="true" pageId="0" pageNumber="231">Larva</emphasis>
(Table
<tableCitation id="9A6A74154E47B89A2E6B1B3BBF411CAD" captionStart="Table 1" captionStartId="T1" captionText="Table 1. States of selected larval characters of Arcobaetis gen. nov. Figure numbers refer to those in this paper, while those of A. gracilentus comb. nov. refer to Kang et al. (1994)." httpUri="http://table.plazi.org/id/2503F8B5E92CE0301DEF57DAFBE238D1" pageId="0" pageNumber="231" tableUuid="2503F8B5E92CE0301DEF57DAFBE238D1">1</tableCitation>
). The following combination of characters distinguish
<taxonomicName id="113ED4BCB1C1FFB394545673EC11A586" lsidName="A. sumbawensis" pageId="0" pageNumber="231" rank="species" species="sumbawensis">
<emphasis id="BE9D6798CE0163150B9DE313145412A3" italics="true" pageId="0" pageNumber="231">A. sumbawensis</emphasis>
</taxonomicName>
sp. nov. from other species of
<taxonomicName id="D8C57751C30296AB4C7E579DAF000379" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Arcobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arcobaetis" order="Ephemeroptera" pageId="0" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="490CE0970CE99133007D316BD09F17BB" italics="true" pageId="0" pageNumber="231">Arcobaetis</emphasis>
</taxonomicName>
gen. nov.: A) distal margins of segments in middle part of flagellum without enlarged spines; B) labial palp segment III sub-rectangular, at base approx. as wide as distal margin of segment II (Fig.
<figureCitation id="5084A1CEBA00867C826BFAF0530BE002" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">2h</figureCitation>
); C) hind protoptera absent; D) claw with single row of denticles, four or five distalmost denticles larger, ca. ten basal denticles small to minute (Figs
<figureCitation id="FBF24DEBC4E21FC0468EDE7F07BA862A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct." figureDoi="10.3897/zookeys.1168.104844.figure3" httpUri="https://binary.pensoft.net/fig/872205" pageId="0" pageNumber="231">3b</figureCitation>
,
<figureCitation id="B74ECE7534487466C63975F4D7CB205E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Arcobaetis sumbawensis sp. nov., larva, SEM pictures a fore claw, lateroventral view b middle claw, anterior view c middle claw, posterior view. Arrows: subapical setae. Scale bars: 0.01 mm." figureDoi="10.3897/zookeys.1168.104844.figure5" httpUri="https://binary.pensoft.net/fig/872207" pageId="0" pageNumber="231">5a</figureCitation>
); E) tergalii present on abdominal segments I-VII; F) posterior margin of tergum IV with triangular spines, slightly wider than long (Fig.
<figureCitation id="C32912D30562A4677E6855AA3CCBA6F4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct." figureDoi="10.3897/zookeys.1168.104844.figure3" httpUri="https://binary.pensoft.net/fig/872205" pageId="0" pageNumber="231">3c</figureCitation>
).
</paragraph>
<caption id="2503F8B5E92CE0301DEF57DAFBE238D1" ID-Table-UUID="2503F8B5E92CE0301DEF57DAFBE238D1" httpUri="http://table.plazi.org/id/2503F8B5E92CE0301DEF57DAFBE238D1" pageId="0" pageNumber="231" start="Table 1" startId="T1">
<paragraph id="9616A3A03D213933C310B0E4A9371BCA" pageId="0" pageNumber="231">
<emphasis id="7D7CED256194AB7E450153B000D0059B" bold="true" pageId="0" pageNumber="231">Table 1.</emphasis>
States of selected larval characters of
<taxonomicName id="58727F5BF924E798285832F309AE5C9C" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Arcobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arcobaetis" order="Ephemeroptera" pageId="0" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="296B33EAD739C1D5A97B25C38994B071" italics="true" pageId="0" pageNumber="231">Arcobaetis</emphasis>
</taxonomicName>
gen. nov. Figure numbers refer to those in this paper, while those of
<taxonomicName id="AC7DC458D1F71AA88F198166D24735F4" lsidName="A. gracilentus" pageId="0" pageNumber="231" rank="species" species="gracilentus">
<emphasis id="EEE223D348249A41F5282DE254673681" italics="true" pageId="0" pageNumber="231">A. gracilentus</emphasis>
</taxonomicName>
comb. nov. refer to
<bibRefCitation id="A6B94C34476FE48F29B3DADE1C0A2DC3" author="Kang, S-C" journalOrPublisher="Journal of Taiwan Museum" pageId="0" pageNumber="231" pagination="9 - 44" refId="B13" refString="Kang, S-C, Chang, H-C, Yang, C-T, 1994. A revision of the genus Baetis in Taiwan (Ephemeroptera, Baetidae). Journal of Taiwan Museum 47: 9 - 44" title="A revision of the genus Baetis in Taiwan (Ephemeroptera, Baetidae)." volume="47" year="1994">Kang et al. (1994)</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="C388832C69C9EC5276C6E18D1E5FD129" pageId="0" pageNumber="231">
<table id="54EA6BB4D09C73F5944D1F09FB5C5D66" pageId="0" pageNumber="231">
<tr id="3D92C2038B2C62ADEB2045A388701FF3" pageId="0" pageNumber="231">
<th id="C5A695BC9B058DAB84E1B1411E887C03" colspan="1" pageId="0" pageNumber="231" rowspan="2">Characters</th>
<th id="27D2533D863F617191F0F32BCC4E4E60" colspan="1" pageId="0" pageNumber="231" rowspan="1">
<taxonomicName id="300588CF9E5485235743E64A32B682F9" lsidName="A. sumbawensis" pageId="0" pageNumber="231" rank="species" species="sumbawensis">
<emphasis id="BDD25C74C0909393DDD5189BDF2F284C" italics="true" pageId="0" pageNumber="231">A. sumbawensis</emphasis>
</taxonomicName>
sp. nov.
</th>
<th id="4C581468A1EF2374CC748E706BEB8D84" colspan="1" pageId="0" pageNumber="231" rowspan="2">Fig.</th>
<th id="06651EC5A720AD995210B768D67BDA91" colspan="1" pageId="0" pageNumber="231" rowspan="1">
<taxonomicName id="5D5BFBDCED7EA2B69179BCCB09BA9638" lsidName="A. sumatrensis" pageId="0" pageNumber="231" rank="species" species="sumatrensis">
<emphasis id="69BA95EC545E4D6E179648C539031B1E" italics="true" pageId="0" pageNumber="231">A. sumatrensis</emphasis>
</taxonomicName>
sp. nov.
</th>
<th id="20F8CBE2A6386F76F2DAEAEE1E140023" colspan="1" pageId="0" pageNumber="231" rowspan="2">Fig.</th>
<th id="F1E82F4D4D982E70EDE7E32841CA4EC2" colspan="1" pageId="0" pageNumber="231" rowspan="1">
<taxonomicName id="700BC1B541E414BFBE168D6D91686957" lsidName="A. bornensis" pageId="0" pageNumber="231" rank="species" species="bornensis">
<emphasis id="1D66C422417130BE57AD733ED6022E9D" italics="true" pageId="0" pageNumber="231">A. bornensis</emphasis>
</taxonomicName>
sp. nov.
</th>
<th id="9E1D99BBF9EE5D5ECA4E9FF7DDBE1D3A" colspan="1" pageId="0" pageNumber="231" rowspan="2">Fig.</th>
<th id="41E00F3682AF51F47B4B8ABF7598E360" colspan="1" pageId="0" pageNumber="231" rowspan="1">
<taxonomicName id="D62C72E1CF1EFE3A51A73B23C4D01A6E" lsidName="A. sripadai" pageId="0" pageNumber="231" rank="species" species="sripadai">
<emphasis id="6FE90A7D7B259849F5D0BD4F759A433A" italics="true" pageId="0" pageNumber="231">A. sripadai</emphasis>
</taxonomicName>
sp. nov.
</th>
<th id="B5EF431CECD41EDB4D374929274B7548" colspan="1" pageId="0" pageNumber="231" rowspan="2">Fig.</th>
<th id="FF735704CF7AF5F909D7D9D9A19D747C" colspan="1" pageId="0" pageNumber="231" rowspan="1">
<taxonomicName id="453AA0B510CACE886573EA281DB2ED15" lsidName="A. gracilentus" pageId="0" pageNumber="231" rank="species" species="gracilentus">
<emphasis id="D6E08A0CD8F3CA42E3F5F4553886C2A1" italics="true" pageId="0" pageNumber="231">A. gracilentus</emphasis>
</taxonomicName>
comb. nov.
</th>
<th id="1B66E3560AC6E58FC1710BBD85A2CFA9" colspan="1" pageId="0" pageNumber="231" rowspan="2">Fig.</th>
</tr>
<tr id="B8754939D9B1C3138659CEDD44566464" pageId="0" pageNumber="231">
<th id="55FD467161F9D5F3F0A7551B0F99B2EB" colspan="1" pageId="0" pageNumber="231" rowspan="1">Sumbawa</th>
<th id="10E4D7066DC9FB9D0F7822FA605BC3FC" colspan="1" pageId="0" pageNumber="231" rowspan="1">Sumatra</th>
<th id="156A5F302D5D1FF7EC2A80527EB8C0A0" colspan="1" pageId="0" pageNumber="231" rowspan="1">Borneo</th>
<th id="6ABAB1A4F1BDE9CB9458150B21859F5E" colspan="1" pageId="0" pageNumber="231" rowspan="1">Sri Lanka</th>
<th id="3F666A7B2A91FD3D89B5DB12A1D4C9C8" colspan="1" pageId="0" pageNumber="231" rowspan="1">Taiwan</th>
</tr>
<tr id="3EC7ECBFDD433A456CE14A2ADAC6282C" pageId="0" pageNumber="231">
<td id="C37447D7920946A48879CD7FB84D3E4D" colspan="1" pageId="0" pageNumber="231" rowspan="1">Flagellum, enlarged spines on distal margins of segments</td>
<td id="4F3AE975018383E37A04E63969CAA974" colspan="1" pageId="0" pageNumber="231" rowspan="1">absent</td>
<td id="8D4C47BAA3F74B29968DD95827A35D44" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
<td id="47A89EA4A176EA1FD0ED6AA87D277212" colspan="1" pageId="0" pageNumber="231" rowspan="1">absent</td>
<td id="24C67313885A651DD82BF195127E34BE" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
<td id="B723CCE69DEB7452DC790825046C6D35" colspan="1" pageId="0" pageNumber="231" rowspan="1">present (middle part)</td>
<td id="6449010293328AB2E9A22F71F2127E33" colspan="1" pageId="0" pageNumber="231" rowspan="1">11f</td>
<td id="ED467347E628F6CA0DF8084A0B1B15E8" colspan="1" pageId="0" pageNumber="231" rowspan="1">absent</td>
<td id="EEC8EFC53DDB56D7951B29F692E2A47E" colspan="1" pageId="0" pageNumber="231" rowspan="1">14a</td>
<td id="835219AF9E37AAB2762734357F84B2C3" colspan="1" pageId="0" pageNumber="231" rowspan="1">absent</td>
<td id="4E36F5E166854770B51974EA15F68069" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
</tr>
<tr id="9A48C5FDABD3D1D1DF7C5D1FA686069A" pageId="0" pageNumber="231">
<td id="E383406FA6D72DD6FCBF0C43D76798D4" colspan="1" pageId="0" pageNumber="231" rowspan="1">Labial palp segment III</td>
<td id="1706B0D69AFD031742A96BDC3D2CE352" colspan="1" pageId="0" pageNumber="231" rowspan="1">sub-rectangular, base approx. as wide as distal margin of segment II</td>
<td id="1122B3ADD2B40EB79EDDB1015B53F36D" colspan="1" pageId="0" pageNumber="231" rowspan="1">2h</td>
<td id="656E0F2C0B2EFD07AE689C59893B4BC0" colspan="1" pageId="0" pageNumber="231" rowspan="1">sub-rectangular, base narrower than distal margin of segment II</td>
<td id="490BA2BBF01EEE420CBB7C82A238C2F9" colspan="1" pageId="0" pageNumber="231" rowspan="1">7h</td>
<td id="8B8E857516321AEAFB92CA06D997119F" colspan="1" pageId="0" pageNumber="231" rowspan="1">sub-quadrangular, base narrower than distal margin of segment II</td>
<td id="E745338C394745AB45B02F4D85B48038" colspan="1" pageId="0" pageNumber="231" rowspan="1">10h</td>
<td id="95C6199661A8C9E0E66DF23937971C9A" colspan="1" pageId="0" pageNumber="231" rowspan="1">sub-quadrangular, base approx. as wide as distal margin of segment II</td>
<td id="01F5E40CF9C01F0470750049FEA03A54" colspan="1" pageId="0" pageNumber="231" rowspan="1">14i</td>
<td id="A4FFB886CC239AC0664651824A37C76B" colspan="1" pageId="0" pageNumber="231" rowspan="1">sub-quadrangular, base narrower than distal margin of segment II</td>
<td id="97EFE57D57CB1461B740F16E75992BED" colspan="1" pageId="0" pageNumber="231" rowspan="1">7E</td>
</tr>
<tr id="27DAA4A0E6EFCDCC5C52C0F6AC2A0009" pageId="0" pageNumber="231">
<td id="C6CA43015AD5AF5F9FA4EEE97EE8E1B2" colspan="1" pageId="0" pageNumber="231" rowspan="1">Hind protoptera</td>
<td id="07E761ACAD7E753AF2C6386B35DF17F4" colspan="1" pageId="0" pageNumber="231" rowspan="1">absent</td>
<td id="F17C0863C013E5F157333DB0A2CF7F92" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
<td id="D3E4F39D2B718B9245469AAAFC547188" colspan="1" pageId="0" pageNumber="231" rowspan="1">absent</td>
<td id="0076DE046E3DB18AADE5FE95133BF182" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
<td id="2FF984FE02509ADB2B84309405E57771" colspan="1" pageId="0" pageNumber="231" rowspan="1">absent</td>
<td id="2445CF0F997AFD96B740CABB6FD814CE" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
<td id="14FFCA1AFFDDD1A53F33783AF5E88B28" colspan="1" pageId="0" pageNumber="231" rowspan="1">absent</td>
<td id="C3B92ED0E8616A4A4B4AAE91F98A270F" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
<td id="A7C5B5D1360BA2E076342CEF3043D78B" colspan="1" pageId="0" pageNumber="231" rowspan="1">present</td>
<td id="7E90D24B2B29A0C6E03A0837D64C079F" colspan="1" pageId="0" pageNumber="231" rowspan="1">7G</td>
</tr>
<tr id="B15D552645A44D85EA0EAD4235D13487" pageId="0" pageNumber="231">
<td id="6DA012FF75F7358D75EE0E58466C18B8" colspan="1" pageId="0" pageNumber="231" rowspan="1">Tergalii on abdominal segments</td>
<td id="0168E680CC83E91FA5A223FA3ABB88D6" colspan="1" pageId="0" pageNumber="231" rowspan="1">I-VII</td>
<td id="09894FE2AF53564D8C28047BEC37CD15" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
<td id="4371819C287ED415922B18FC37AE56CE" colspan="1" pageId="0" pageNumber="231" rowspan="1">II-VII</td>
<td id="60EEE190E3C0C0015C8861C686FD2F2E" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
<td id="103DE538FFF2A0DDDC7E4F28F6F5614B" colspan="1" pageId="0" pageNumber="231" rowspan="1">I-VII</td>
<td id="872BB6ABEF7ABA64D1536F36C281EF0B" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
<td id="442A3F6E40EBDEF46B8D142CDE7B95A6" colspan="1" pageId="0" pageNumber="231" rowspan="1">II-VII</td>
<td id="64940D744F5215B3F1F92246FBC8C3A7" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
<td id="E38E05C287FC5CAC4132674977A7A7C4" colspan="1" pageId="0" pageNumber="231" rowspan="1">I-VII</td>
<td id="5138FF9229B367CEBE5C39161C4D2CE6" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
</tr>
<tr id="161D31ED1FB2BED9DE3A9C0AF62D0D38" pageId="0" pageNumber="231">
<td id="2322D516E0CF05A3B1E3DCF7EC9FE321" colspan="1" pageId="0" pageNumber="231" rowspan="2">Abdominal tergite IV, spines on posterior margin</td>
<td id="024C784D09AE8B4102B45DF73EB14476" colspan="1" pageId="0" pageNumber="231" rowspan="1">triangular</td>
<td id="992953A6BEE575F2674850777F5A8148" colspan="1" pageId="0" pageNumber="231" rowspan="1">3c</td>
<td id="91BF4D16C26702EE638A3A2D2E5A9521" colspan="1" pageId="0" pageNumber="231" rowspan="1">triangular</td>
<td id="4B387EE88DB25A6316366DD995873694" colspan="1" pageId="0" pageNumber="231" rowspan="1">8i</td>
<td id="67BACD34B084694E2E5613FE2443B667" colspan="1" pageId="0" pageNumber="231" rowspan="1">triangular</td>
<td id="89217F1D2A0A0BF3E399FB071D658CCF" colspan="1" pageId="0" pageNumber="231" rowspan="1">11c</td>
<td id="9290DFDB70EA9527747F29720DD50872" colspan="1" pageId="0" pageNumber="231" rowspan="1">triangular</td>
<td id="D0F6A2EA648A67B03DC617F52C919031" colspan="1" pageId="0" pageNumber="231" rowspan="1">15f</td>
<td id="C12025D17064E860D69A8CC876FD4DEE" colspan="1" pageId="0" pageNumber="231" rowspan="1">triangular</td>
<td id="C5DEB161C07688CCBD594C771690E5CD" colspan="1" pageId="0" pageNumber="231" rowspan="1">7K</td>
</tr>
<tr id="054F8B6A83EE0B9F090F37E85F5212A4" pageId="0" pageNumber="231">
<td id="D0740133E1BB6D0743DDFA8B80C39D99" colspan="1" pageId="0" pageNumber="231" rowspan="1">slightly wider than long</td>
<td id="6C84923C70A90AB70B011EC5D5122C27" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
<td id="D1AA0A1CBB13C06751333A1C94BDCD02" colspan="1" pageId="0" pageNumber="231" rowspan="1">wider than long</td>
<td id="F72FA77C9473AEEF287BE5BA970B0196" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
<td id="912D6580CD484D48C8C76335B1F73F19" colspan="1" pageId="0" pageNumber="231" rowspan="1">wider than long</td>
<td id="B8511C1E201D341C20E020D53E698771" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
<td id="D04F733E08A8CDBC9F7FCBA43FF3C756" colspan="1" pageId="0" pageNumber="231" rowspan="1">slightly wider than long</td>
<td id="2606939B944EB58B0AA14118B15F1C85" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
<td id="2A7617B9218D1B482B6E7D1A81BA03E5" colspan="1" pageId="0" pageNumber="231" rowspan="1">longer than wide</td>
<td id="E413FE767201EFC3CD3394B2DFC0B4E7" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
</tr>
<tr id="2C0A480B6917F9E70E57E8B78BC29C94" pageId="0" pageNumber="231">
<td id="917F756E4FAB341A4A4EFF240A730008" colspan="1" pageId="0" pageNumber="231" rowspan="1">Cerci, spines on outer sides</td>
<td id="48EE597D7B03FB6FF3E3DD10BD354B4D" colspan="1" pageId="0" pageNumber="231" rowspan="1">unknown</td>
<td id="62238A66C409AB1E8F6B4E5E16CF76DA" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
<td id="9B78492151A438419CAA43A45DA23B24" colspan="1" pageId="0" pageNumber="231" rowspan="1">two somewhat longer on each 2nd segment</td>
<td id="8D4DE402D78B42D0A88B51D6AED87F08" colspan="1" pageId="0" pageNumber="231" rowspan="1">9f</td>
<td id="94EF70AD9106DA50EEF46AC9400A8AFD" colspan="1" pageId="0" pageNumber="231" rowspan="1">unknown</td>
<td id="DA5B38F8ED65E5B9ED4118872F661049" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
<td id="088E5D841A30B77E064AE10B8C5700C3" colspan="1" pageId="0" pageNumber="231" rowspan="1">several strongly enlarged on each 2nd segment</td>
<td id="D45E00155549D7CAE0F8FAEEA52888BC" colspan="1" pageId="0" pageNumber="231" rowspan="1">15o, p</td>
<td id="3A04BEF5379078477ABE26FF10353A33" colspan="1" pageId="0" pageNumber="231" rowspan="1">unknown</td>
<td id="6DC769F0F6BE5617FF8B55F459939D61" colspan="1" isEmpty="true" pageId="0" pageNumber="231" rowspan="1">-</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="D886A0AAF38207C65903BC8AB851E83E" pageId="0" pageNumber="231" type="description">
<paragraph id="1D0B795EC71ED6640F151BD5B0AE93A1" pageId="0" pageNumber="231">Description.</paragraph>
<paragraph id="0F5666546A034A0D843F4E9C12F4E1FB" pageId="0" pageNumber="231">
<emphasis id="BB594F8EFB398387D00C9FEF58C430EE" bold="true" pageId="0" pageNumber="231">Larva</emphasis>
(Figs
<figureCitation id="14D63D81268455CD7AA90A61FD0F1C2F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Arcobaetis sumbawensis sp. nov., larva, habitus (holotype) a dorsal view b lateral view c ventral view. Scale bars: 1 mm" figureDoi="10.3897/zookeys.1168.104844.figure1" httpUri="https://binary.pensoft.net/fig/872203" pageId="0" pageNumber="231">1</figureCitation>
-
<figureCitation id="AE4A00DC259BD399249B42900D010D74" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Arcobaetis sumbawensis sp. nov., larva, SEM pictures a fore claw, lateroventral view b middle claw, anterior view c middle claw, posterior view. Arrows: subapical setae. Scale bars: 0.01 mm." figureDoi="10.3897/zookeys.1168.104844.figure5" httpUri="https://binary.pensoft.net/fig/872207" pageId="0" pageNumber="231">5</figureCitation>
). Body length 5.7-7.0 mm.
</paragraph>
<paragraph id="E120FAAADEE3AE56B5E75AE3DAA1D268" pageId="0" pageNumber="231">
<emphasis id="259A875CB27955030845E1AF6BE636DD" bold="true" italics="true" pageId="0" pageNumber="231">Cuticular colouration</emphasis>
(Fig.
<figureCitation id="75E2A81872370F57F0C833A3A2AD502D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Arcobaetis sumbawensis sp. nov., larva, habitus (holotype) a dorsal view b lateral view c ventral view. Scale bars: 1 mm" figureDoi="10.3897/zookeys.1168.104844.figure1" httpUri="https://binary.pensoft.net/fig/872203" pageId="0" pageNumber="231">1a-c</figureCitation>
). Head, thorax, and abdomen dorsally brown, thorax darker. Head, thorax, and abdomen ventrally ochre. Legs ochre, femur distally with brown streak; tibia basally along patella-tibial suture darker. Caudalii ochre.
</paragraph>
<paragraph id="5B24C040A4A0F86C57295E9C0D9AFCCF" pageId="0" pageNumber="231">
<emphasis id="D95ADFFA23A21506714870C2EE889BB1" bold="true" italics="true" pageId="0" pageNumber="231">Hypodermal colouration.</emphasis>
Abdomen dorsally with narrow reddish transverse stripes on intersegmental membranes (Fig.
<figureCitation id="1F9AAD762EC422DD38B7F0FE34CE9BFB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Arcobaetis sumbawensis sp. nov., larva, habitus (holotype) a dorsal view b lateral view c ventral view. Scale bars: 1 mm" figureDoi="10.3897/zookeys.1168.104844.figure1" httpUri="https://binary.pensoft.net/fig/872203" pageId="0" pageNumber="231">1a</figureCitation>
).
</paragraph>
<paragraph id="1416F750474FCFC35D11403DE62372FE" pageId="0" pageNumber="231">
<emphasis id="1565CBE1015B6A67C48BA27433003900" bold="true" italics="true" pageId="0" pageNumber="231">Antenna.</emphasis>
Flagellum in middle part without enlarged spines at distal margin of segments.
</paragraph>
<paragraph id="AD6F357DBDFCE41360AD7DE907BA963A" pageId="0" pageNumber="231">
<emphasis id="3A17B493FF76649B2BD351144B440B7C" bold="true" italics="true" pageId="0" pageNumber="231">Labrum</emphasis>
(Fig.
<figureCitation id="3B5FD8443D3D2E60FC5B6E12023101BE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">2a</figureCitation>
). Length 0.7
<normalizedToken id="4654BEB0B6587901D9D3BF5C62706B28" originalValue="×">x</normalizedToken>
maximum width. Dorsally with sub-median seta, short sub-marginal arc of simple setae, and two setae in between.
</paragraph>
<paragraph id="10C3711E2CFCFDE4266EDFE9809BCD6E" pageId="0" pageNumber="231">
<emphasis id="601FC6C869C4F994895CC17202ADAABC" bold="true" italics="true" pageId="0" pageNumber="231">Right mandible</emphasis>
(Fig.
<figureCitation id="8738A1F2FD87F60F704F008881209067" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">2b, c</figureCitation>
). Incisor with five denticles; kinetodontium with three denticles.
</paragraph>
<paragraph id="448FB103E5CABD99EC8EDCAD3E16C240" pageId="0" pageNumber="231">
<emphasis id="EC73293F47DD424A40E871D37B98B15D" bold="true" italics="true" pageId="0" pageNumber="231">Left mandible</emphasis>
(Fig.
<figureCitation id="A3C6DC1A8C78E2F8C0E2879C717E255F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">2d, e</figureCitation>
). Incisor with five denticles; kinetodontium with three denticles.
</paragraph>
<paragraph id="6895E5D5CAD4953316C2F4F929EBF575" pageId="0" pageNumber="231">
<emphasis id="A8178362CA1116F2F78EA10BB2AD0F5F" bold="true" italics="true" pageId="0" pageNumber="231">Hypopharynx and superlinguae</emphasis>
(Fig.
<figureCitation id="66865717BB34C4807A2476ABE6EF6A1E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">2f</figureCitation>
). Lingua as long as superlinguae, longer than broad. Superlinguae distally straight; lateral margins rounded; fine, long, simple setae along distal margin.
</paragraph>
<paragraph id="5D960D873B74D2C5F7341C8E039A7AE7" pageId="0" pageNumber="231">
<emphasis id="EF3E3543E22AFCECB1D9F76BAEE17C61" bold="true" italics="true" pageId="0" pageNumber="231">Maxilla</emphasis>
(Fig.
<figureCitation id="34C09F10B116D36D14BD9B087F3301B7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">2g</figureCitation>
). Galea-lacinia ventrally with two simple setae just proximad of canines. Medially with one pectinate, spine-like seta and ca. six short to long, simple setae. Maxillary palp approx. 1.5
<normalizedToken id="BFDD2A4AE281D10A9399BCF1E8C1DBDE" originalValue="×">x</normalizedToken>
as long as galea-lacinia; palp segment II approx. 1.3
<normalizedToken id="68B0E0EB0B80521B068B67F962CA04E8" originalValue="×">x</normalizedToken>
as long as segment I; setae on maxillary palp fine, simple, scattered over surface of segments I and II.
</paragraph>
<paragraph id="627E0DA381710CCCCCD3AF15E4009184" pageId="0" pageNumber="231">
<emphasis id="02BE711F1F2404678A556EF2175854D8" bold="true" italics="true" pageId="0" pageNumber="231">Labium</emphasis>
(Fig.
<figureCitation id="95748C003BED29FB750162A5348C2891" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">2h-j</figureCitation>
). Inner margin of glossa with ca. 16 spine-like setae, increasing in length distally; apex with two long and one medium robust setae; outer margin with ca. 12 spine-like setae; paraglossa ventrally with ca. four medium, simple setae in anteromedial area; dorsally with arc of ca. 10 long, spine-like setae in distal area. Labial palp with segment I 0.7
<normalizedToken id="C95CB926D9A61E55142ABFCC1975A35D" originalValue="×">x</normalizedToken>
length of segments II and III combined. Segment II dorsally with row of ca. four spine-like setae. Segment III sub-rectangular; at base approx. as wide as distal margin of segment II; ventral surface with short, spine-like, simple setae and short, fine, simple setae.
</paragraph>
<paragraph id="32997A8F2782C38D52A9220939FCC043" pageId="0" pageNumber="231">
<emphasis id="FA83E1EF91F37AF219219871624A9C4C" bold="true" italics="true" pageId="0" pageNumber="231">Hind protoptera</emphasis>
absent.
</paragraph>
<paragraph id="A4539EBD694550F65B02BFBE091FB922" pageId="0" pageNumber="231">
<emphasis id="24C6CBD9B8CFB7696025F39EC012F820" bold="true" italics="true" pageId="0" pageNumber="231">Legs</emphasis>
(Figs
<figureCitation id="3FC98BE081A981CE1BF5DD99370CC953" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct." figureDoi="10.3897/zookeys.1168.104844.figure3" httpUri="https://binary.pensoft.net/fig/872205" pageId="0" pageNumber="231">3a, b</figureCitation>
,
<figureCitation id="E5AAAB09E8A161D643A129800B5615A7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Arcobaetis sumbawensis sp. nov., larva a foreleg b middle leg c hind leg d subimaginal gonostylus developing under cuticle of last instar male larva e head. Scale bars: 0.05 mm (e); 0.1 mm (d)." figureDoi="10.3897/zookeys.1168.104844.figure4" httpUri="https://binary.pensoft.net/fig/872206" pageId="0" pageNumber="231">4a-c</figureCitation>
,
<figureCitation id="CBA77D846974743E1A7756B2EB0A3B15" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Arcobaetis sumbawensis sp. nov., larva, SEM pictures a fore claw, lateroventral view b middle claw, anterior view c middle claw, posterior view. Arrows: subapical setae. Scale bars: 0.01 mm." figureDoi="10.3897/zookeys.1168.104844.figure5" httpUri="https://binary.pensoft.net/fig/872207" pageId="0" pageNumber="231">5a-c</figureCitation>
). Long and slender, middle, and hind legs slenderer than fore legs. Ratio of leg segments: fore leg 1.3:1.0:0.7:0.2, middle and hind legs 1.2:1.0:0.6:0.1.
<emphasis id="3F922C22824A2111728460BECC23D9F7" bold="true" italics="true" pageId="0" pageNumber="231">Femur</emphasis>
. Length of fore femur ca. 5
<normalizedToken id="021B2E3BC3BC731B7D7291A6E30006BE" originalValue="×">x</normalizedToken>
maximum width, outer and inner margins almost parallel; length of middle and hind femora&gt;6
<normalizedToken id="DBDBC2BD5CD1A4F6EABF195227865D36" originalValue="×">x</normalizedToken>
maximum width, outer margins slightly concave. Outer margin with row of ca. 12 short, curved, spine-like setae, on fore leg larger than on middle and hind legs. On ventral side with short, spine-like, pectinate, pointed setae; larger and denser on foreleg, smaller and less dense on middle and hind legs.
<emphasis id="FB5A4E94B6C23DD83AA039B8334376E1" bold="true" italics="true" pageId="0" pageNumber="231">Tibia</emphasis>
. Short, stout, pointed, pectinate setae near inner margin. Inner margin with row of short, curved, spine-like, pectinate setae, on apex two longer, curved, spine-like, pectinate setae. Outer margin with one apical, spine-like seta; on fore leg without other spine-like setae, on middle and hind legs with row of few medium, spine-like setae. Patella-tibial suture on proximal ¹/3 of tibia on all legs.
<emphasis id="441847EC34650C24336397CEE6C5349D" bold="true" italics="true" pageId="0" pageNumber="231">Tarsus</emphasis>
. Inner margin with row of short, curved, pectinate, spine-like setae; outer margin bare.
<emphasis id="BAC5B50E91E95A171DB7677898FF1FCB" bold="true" italics="true" pageId="0" pageNumber="231">Claw</emphasis>
with one row of denticles; four or five distalmost denticles larger and directed distad; basally ca. 10 small to minute denticles; two vestigial subapical setae on anterior side, one vestigial subapical seta on posterior side (Fig.
<figureCitation id="E73D458B8DFF157D3D4BC6AE681CCE91" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Arcobaetis sumbawensis sp. nov., larva, SEM pictures a fore claw, lateroventral view b middle claw, anterior view c middle claw, posterior view. Arrows: subapical setae. Scale bars: 0.01 mm." figureDoi="10.3897/zookeys.1168.104844.figure5" httpUri="https://binary.pensoft.net/fig/872207" pageId="0" pageNumber="231">5a, b</figureCitation>
).
</paragraph>
<paragraph id="8F7FBA778FA530913896E2C4D35D7E19" pageId="0" pageNumber="231">
<emphasis id="23C3292BB717A0AAD09F5EC58411F659" bold="true" italics="true" pageId="0" pageNumber="231">Abdominal terga</emphasis>
(Fig.
<figureCitation id="AC0034AACC4453A7E0D134CA7BF91F17" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct." figureDoi="10.3897/zookeys.1168.104844.figure3" httpUri="https://binary.pensoft.net/fig/872205" pageId="0" pageNumber="231">3c, d</figureCitation>
). Posterior margin of terga: I smooth, without spines; II-IX with triangular spines, increasing in length toward IX; slightly wider than long on tergum IV; row of spines on tergum IX interrupted by smaller spines in middle part behind bases of submedian setae (similar to Fig.
<figureCitation id="39D24D70F6E2E2375520D5FFCA7DD89E" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Arcobaetis sripadai sp. nov., larval exuviae a, b fore and middle legs (triangles show points where patella-tibial suture crosses inner margin c claw d-k posterior margins of abdominal terga II-VI and VIII-X l-m posterior margin of abdominal sterna VIII and IX of male larva n paraproct. o-p fragments of cerci (lateral view)." figureDoi="10.3897/zookeys.1168.104844.figure15" httpUri="https://binary.pensoft.net/fig/872217" pageId="0" pageNumber="231">15j</figureCitation>
). Posterior margin of tergum X with median concavity with smaller spines (similar to Fig.
<figureCitation id="B894F2AD684FE6B7E22E753FD3CC012D" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Arcobaetis sripadai sp. nov., larval exuviae a, b fore and middle legs (triangles show points where patella-tibial suture crosses inner margin c claw d-k posterior margins of abdominal terga II-VI and VIII-X l-m posterior margin of abdominal sterna VIII and IX of male larva n paraproct. o-p fragments of cerci (lateral view)." figureDoi="10.3897/zookeys.1168.104844.figure15" httpUri="https://binary.pensoft.net/fig/872217" pageId="0" pageNumber="231">15k</figureCitation>
).
</paragraph>
<paragraph id="758D5FEBE150DFD2AA4ADCF786675B04" pageId="0" pageNumber="231">
<emphasis id="D01E15E7C7E2C32DDE8B005FC3885ADF" bold="true" italics="true" pageId="0" pageNumber="231">Abdominal sterna.</emphasis>
Posterior margins of sterna: I-V smooth, without spines; VI-IX with triangular spines. On sternum IX of male mature larva row of narrow pointed spines between protogonostyli, smaller spines laterad of protogono-styli and larger, pointed spines laterad of them (similar to Fig.
<figureCitation id="B6D10173BBDC7795225EBBC840B14A56" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Arcobaetis sripadai sp. nov., larval exuviae a, b fore and middle legs (triangles show points where patella-tibial suture crosses inner margin c claw d-k posterior margins of abdominal terga II-VI and VIII-X l-m posterior margin of abdominal sterna VIII and IX of male larva n paraproct. o-p fragments of cerci (lateral view)." figureDoi="10.3897/zookeys.1168.104844.figure15" httpUri="https://binary.pensoft.net/fig/872217" pageId="0" pageNumber="231">15m</figureCitation>
).
</paragraph>
<paragraph id="B2DF9373171ABDAB6D95CD090761645C" pageId="0" pageNumber="231">
<emphasis id="A5885C6218CCCC34A5A0EEA3B33A7942" bold="true" italics="true" pageId="0" pageNumber="231">Tergalii</emphasis>
(Fig.
<figureCitation id="B15EE57F348C34DFA9692D36F6D3A705" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct." figureDoi="10.3897/zookeys.1168.104844.figure3" httpUri="https://binary.pensoft.net/fig/872205" pageId="0" pageNumber="231">3e</figureCitation>
). Present on abdominal segments I-VII. Tracheation in tergalius I limited to basal part of main trunk. In nine well preserved larvae, only a few tergalii were found, but insertions were always well developed: two tergalii I in one specimen; one tergalius IV and one tergalius V in another specimen, both very small, looking like replacements of naturally broken tergalii.
</paragraph>
<paragraph id="84B15AF80B47EE514525F288F318E645" pageId="0" pageNumber="231">
<emphasis id="8CD0FC4242FF9D18435E98F3899D5E79" bold="true" italics="true" pageId="0" pageNumber="231">Paraproct</emphasis>
(Fig.
<figureCitation id="0B5C9400A2F9CB92CFD479F60939E449" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct." figureDoi="10.3897/zookeys.1168.104844.figure3" httpUri="https://binary.pensoft.net/fig/872205" pageId="0" pageNumber="231">3f</figureCitation>
). Posterior margin with ca. 16 stout spines. Surface scattered with scale bases and micropores.
</paragraph>
<paragraph id="F976B9B672D42FCC6F424270A3F5410B" pageId="0" pageNumber="231">
<emphasis id="10A486426C41D9FE83C776F75F47AADA" bold="true" pageId="0" pageNumber="231">
<emphasis id="A4873A21AF28BF9E5220494CFDD272C5" italics="true" pageId="0" pageNumber="231">Caudalii</emphasis>
.
</emphasis>
Cerci and paracercus with broad triangular spines on posterior margin of each segment.
</paragraph>
</subSubSection>
<subSubSection id="D41B795ECFC57E289785F0B63E3ED8CC" pageId="0" pageNumber="231" type="etymology">
<paragraph id="80F815AAB3BEDA6D668459B3A7B6D1C6" pageId="0" pageNumber="231">Etymology.</paragraph>
<paragraph id="A234D0391AE32177C32845D1CFB2AF0E" pageId="0" pageNumber="231">The specific epithet refers to the type locality in Sumbawa (Indonesia).</paragraph>
</subSubSection>
<subSubSection id="A4580D080F4C289E14FFA3E80ECF6519" pageId="0" pageNumber="231" type="distribution">
<paragraph id="840E58436EE203AE05D20415B4F4DB87" pageId="0" pageNumber="231">Distribution.</paragraph>
<paragraph id="95832DDD3E8AC92AAC1E4E22EF11C65A" pageId="0" pageNumber="231">
Indonesia: Sumbawa (Fig.
<figureCitation id="76AFAAA57D85E614F0A74C44BB200550" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Arcobaetis gen. nov., distribution." figureDoi="10.3897/zookeys.1168.104844.figure19" httpUri="https://binary.pensoft.net/fig/872221" pageId="0" pageNumber="231">19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7EF2AD5EBDB593CBF6AB2F6479556B9E" pageId="0" pageNumber="231" type="biological aspects">
<paragraph id="1ABA75A4DBCEFBE0EE95ED48E3871F83" pageId="0" pageNumber="231">Biological aspects.</paragraph>
<paragraph id="E61EFDB0D0C276F523343C577654766B" pageId="0" pageNumber="231">The species was found at an altitude of 1380 m.</paragraph>
</subSubSection>
</treatment>
</document>
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