<documentID-CLB-Dataset="57144"ID-DOI="http://dx.doi.org/10.3897/zookeys.1168.104844"ID-GBIF-Dataset="67f9e0f5-21e5-478f-8ee2-1924430bb454"ID-PMC="PMC10843360"ID-Pensoft-Pub="1313-2970-1168-231"ID-Pensoft-UUID="6928794CB97051F9AAE3B64050E5FAF3"ID-PubMed="38318436"ID-ZooBank="28A21391A4334E808845E5C0996EFAF0"ModsDocID="1313-2970-1168-231"checkinTime="1688439534387"checkinUser="pensoft"docAuthor="Kaltenbach, Thomas, Kluge, Nikita J. & Gattolliat, Jean-Luc"docDate="2023"docId="6E4FA03D5D32522DA54FBD02C733CE43"docLanguage="en"docName="ZooKeys 1168: 231-266"docOrigin="ZooKeys 1168"docPubDate="2023-07-03"docSource="http://dx.doi.org/10.3897/zookeys.1168.104844"docTitle="Arcobaetis sumbawensis Kaltenbach & Kluge & Gattolliat 2023, sp. nov."docType="treatment"docUuid="7A575C67-8D0D-4A93-BB2B-BF7669B229D5"docUuidSource="ZooBank"docVersion="5"id="6928794CB97051F9AAE3B64050E5FAF3"lastPageNumber="231"masterDocId="6928794CB97051F9AAE3B64050E5FAF3"masterDocTitle="A new, widespread genus of Baetidae from South Asia (Insecta, Ephemeroptera)"masterLastPageNumber="266"masterPageNumber="231"pageNumber="231"updateTime="1732691564844"updateUser="ExternalLinkService">
<mods:affiliationid="0CEB0911A3ED4DCB66181E87EEA89271">Museum cantonal des Sciences Naturelles, Departement de zoologie, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland & University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
<mods:affiliationid="EA1047EF114EC3AA8C00061FA7240BA6">Museum cantonal des Sciences Naturelles, Departement de zoologie, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland & University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
<taxonomicNameid="7DD52BA5BCCD572BEC4F6FD7A1AA9904"LSID="https://zoobank.org/7A575C67-8D0D-4A93-BB2B-BF7669B229D5"authority="Kaltenbach & Kluge & Gattolliat, 2023"authorityName="Kaltenbach & Kluge & Gattolliat"authorityYear="2023"class="Insecta"family="Baetidae"genus="Arcobaetis"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Arcobaetis sumbawensis"order="Ephemeroptera"pageId="0"pageNumber="231"phylum="Arthropoda"rank="species"species="sumbawensis"status="sp. nov.">Arcobaetis sumbawensis</taxonomicName>
<figureCitationid="2F790E2520745B2DC9EF81ACA8F6E4F1"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Arcobaetis sumbawensis sp. nov., larva, habitus (holotype) a dorsal view b lateral view c ventral view. Scale bars: 1 mm"figureDoi="10.3897/zookeys.1168.104844.figure1"httpUri="https://binary.pensoft.net/fig/872203"pageId="0"pageNumber="231">Figs 1</figureCitation>
<figureCitationid="8DC6FF1EE96419EA38610563A7F37475"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)."figureDoi="10.3897/zookeys.1168.104844.figure2"httpUri="https://binary.pensoft.net/fig/872204"pageId="0"pageNumber="231">, 2</figureCitation>
<figureCitationid="B89B9E27ED219787F31DB991B2D302E1"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct."figureDoi="10.3897/zookeys.1168.104844.figure3"httpUri="https://binary.pensoft.net/fig/872205"pageId="0"pageNumber="231">, 3</figureCitation>
<figureCitationid="3F3FFF0EAF3B16C8ACC3A4F282ECC2D4"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Arcobaetis sumbawensis sp. nov., larva a foreleg b middle leg c hind leg d subimaginal gonostylus developing under cuticle of last instar male larva e head. Scale bars: 0.05 mm (e); 0.1 mm (d)."figureDoi="10.3897/zookeys.1168.104844.figure4"httpUri="https://binary.pensoft.net/fig/872206"pageId="0"pageNumber="231">, 4</figureCitation>
<figureCitationid="10E372FF434D0955E1DD519BDFCAC385"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Arcobaetis sumbawensis sp. nov., larva, SEM pictures a fore claw, lateroventral view b middle claw, anterior view c middle claw, posterior view. Arrows: subapical setae. Scale bars: 0.01 mm."figureDoi="10.3897/zookeys.1168.104844.figure5"httpUri="https://binary.pensoft.net/fig/872207"pageId="0"pageNumber="231">, 5</figureCitation>
<figureCitationid="2683387F4F59C37BD4D2AE9F0712DCF7"captionStart="Figure 19"captionStartId="F19"captionText="Figure 19. Arcobaetis gen. nov., distribution."figureDoi="10.3897/zookeys.1168.104844.figure19"httpUri="https://binary.pensoft.net/fig/872221"pageId="0"pageNumber="231">, 19</figureCitation>
9 larvae; same data as holotype; 4 on slides; GBIFCH00692615, GBIFCH00975688, GBIFCH00592652, GBIFCH00592653; 5 in alcohol; GenBank OQ699910, OQ699911; GBIFCH00975695, GBIFCH00975689, GBIFCH00975690, GBIFCH00975694; MZL.
<tableCitationid="9A6A74154E47B89A2E6B1B3BBF411CAD"captionStart="Table 1"captionStartId="T1"captionText="Table 1. States of selected larval characters of Arcobaetis gen. nov. Figure numbers refer to those in this paper, while those of A. gracilentus comb. nov. refer to Kang et al. (1994)."httpUri="http://table.plazi.org/id/2503F8B5E92CE0301DEF57DAFBE238D1"pageId="0"pageNumber="231"tableUuid="2503F8B5E92CE0301DEF57DAFBE238D1">1</tableCitation>
). The following combination of characters distinguish
<taxonomicNameid="D8C57751C30296AB4C7E579DAF000379"authorityName="Kaltenbach & Kluge & Gattolliat"authorityYear="2023"class="Insecta"family="Baetidae"genus="Arcobaetis"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Arcobaetis"order="Ephemeroptera"pageId="0"pageNumber="231"phylum="Arthropoda"rank="genus">
gen. nov.: A) distal margins of segments in middle part of flagellum without enlarged spines; B) labial palp segment III sub-rectangular, at base approx. as wide as distal margin of segment II (Fig.
<figureCitationid="5084A1CEBA00867C826BFAF0530BE002"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)."figureDoi="10.3897/zookeys.1168.104844.figure2"httpUri="https://binary.pensoft.net/fig/872204"pageId="0"pageNumber="231">2h</figureCitation>
); C) hind protoptera absent; D) claw with single row of denticles, four or five distalmost denticles larger, ca. ten basal denticles small to minute (Figs
<figureCitationid="FBF24DEBC4E21FC0468EDE7F07BA862A"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct."figureDoi="10.3897/zookeys.1168.104844.figure3"httpUri="https://binary.pensoft.net/fig/872205"pageId="0"pageNumber="231">3b</figureCitation>
,
<figureCitationid="B74ECE7534487466C63975F4D7CB205E"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Arcobaetis sumbawensis sp. nov., larva, SEM pictures a fore claw, lateroventral view b middle claw, anterior view c middle claw, posterior view. Arrows: subapical setae. Scale bars: 0.01 mm."figureDoi="10.3897/zookeys.1168.104844.figure5"httpUri="https://binary.pensoft.net/fig/872207"pageId="0"pageNumber="231">5a</figureCitation>
); E) tergalii present on abdominal segments I-VII; F) posterior margin of tergum IV with triangular spines, slightly wider than long (Fig.
<figureCitationid="C32912D30562A4677E6855AA3CCBA6F4"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct."figureDoi="10.3897/zookeys.1168.104844.figure3"httpUri="https://binary.pensoft.net/fig/872205"pageId="0"pageNumber="231">3c</figureCitation>
<taxonomicNameid="58727F5BF924E798285832F309AE5C9C"authorityName="Kaltenbach & Kluge & Gattolliat"authorityYear="2023"class="Insecta"family="Baetidae"genus="Arcobaetis"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Arcobaetis"order="Ephemeroptera"pageId="0"pageNumber="231"phylum="Arthropoda"rank="genus">
<bibRefCitationid="A6B94C34476FE48F29B3DADE1C0A2DC3"author="Kang, S-C"journalOrPublisher="Journal of Taiwan Museum"pageId="0"pageNumber="231"pagination="9 - 44"refId="B13"refString="Kang, S-C, Chang, H-C, Yang, C-T, 1994. A revision of the genus Baetis in Taiwan (Ephemeroptera, Baetidae). Journal of Taiwan Museum 47: 9 - 44"title="A revision of the genus Baetis in Taiwan (Ephemeroptera, Baetidae)."volume="47"year="1994">Kang et al. (1994)</bibRefCitation>
<tdid="1706B0D69AFD031742A96BDC3D2CE352"colspan="1"pageId="0"pageNumber="231"rowspan="1">sub-rectangular, base approx. as wide as distal margin of segment II</td>
<tdid="656E0F2C0B2EFD07AE689C59893B4BC0"colspan="1"pageId="0"pageNumber="231"rowspan="1">sub-rectangular, base narrower than distal margin of segment II</td>
<tdid="8B8E857516321AEAFB92CA06D997119F"colspan="1"pageId="0"pageNumber="231"rowspan="1">sub-quadrangular, base narrower than distal margin of segment II</td>
<tdid="95C6199661A8C9E0E66DF23937971C9A"colspan="1"pageId="0"pageNumber="231"rowspan="1">sub-quadrangular, base approx. as wide as distal margin of segment II</td>
<tdid="A4FFB886CC239AC0664651824A37C76B"colspan="1"pageId="0"pageNumber="231"rowspan="1">sub-quadrangular, base narrower than distal margin of segment II</td>
<figureCitationid="75E2A81872370F57F0C833A3A2AD502D"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Arcobaetis sumbawensis sp. nov., larva, habitus (holotype) a dorsal view b lateral view c ventral view. Scale bars: 1 mm"figureDoi="10.3897/zookeys.1168.104844.figure1"httpUri="https://binary.pensoft.net/fig/872203"pageId="0"pageNumber="231">1a-c</figureCitation>
). Head, thorax, and abdomen dorsally brown, thorax darker. Head, thorax, and abdomen ventrally ochre. Legs ochre, femur distally with brown streak; tibia basally along patella-tibial suture darker. Caudalii ochre.
<figureCitationid="3B5FD8443D3D2E60FC5B6E12023101BE"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)."figureDoi="10.3897/zookeys.1168.104844.figure2"httpUri="https://binary.pensoft.net/fig/872204"pageId="0"pageNumber="231">2a</figureCitation>
<figureCitationid="8738A1F2FD87F60F704F008881209067"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)."figureDoi="10.3897/zookeys.1168.104844.figure2"httpUri="https://binary.pensoft.net/fig/872204"pageId="0"pageNumber="231">2b, c</figureCitation>
). Incisor with five denticles; kinetodontium with three denticles.
<figureCitationid="A3C6DC1A8C78E2F8C0E2879C717E255F"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)."figureDoi="10.3897/zookeys.1168.104844.figure2"httpUri="https://binary.pensoft.net/fig/872204"pageId="0"pageNumber="231">2d, e</figureCitation>
). Incisor with five denticles; kinetodontium with three denticles.
<emphasisid="A8178362CA1116F2F78EA10BB2AD0F5F"bold="true"italics="true"pageId="0"pageNumber="231">Hypopharynx and superlinguae</emphasis>
(Fig.
<figureCitationid="66865717BB34C4807A2476ABE6EF6A1E"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)."figureDoi="10.3897/zookeys.1168.104844.figure2"httpUri="https://binary.pensoft.net/fig/872204"pageId="0"pageNumber="231">2f</figureCitation>
). Lingua as long as superlinguae, longer than broad. Superlinguae distally straight; lateral margins rounded; fine, long, simple setae along distal margin.
<figureCitationid="34C09F10B116D36D14BD9B087F3301B7"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)."figureDoi="10.3897/zookeys.1168.104844.figure2"httpUri="https://binary.pensoft.net/fig/872204"pageId="0"pageNumber="231">2g</figureCitation>
). Galea-lacinia ventrally with two simple setae just proximad of canines. Medially with one pectinate, spine-like seta and ca. six short to long, simple setae. Maxillary palp approx. 1.5
<figureCitationid="95748C003BED29FB750162A5348C2891"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)."figureDoi="10.3897/zookeys.1168.104844.figure2"httpUri="https://binary.pensoft.net/fig/872204"pageId="0"pageNumber="231">2h-j</figureCitation>
). Inner margin of glossa with ca. 16 spine-like setae, increasing in length distally; apex with two long and one medium robust setae; outer margin with ca. 12 spine-like setae; paraglossa ventrally with ca. four medium, simple setae in anteromedial area; dorsally with arc of ca. 10 long, spine-like setae in distal area. Labial palp with segment I 0.7
length of segments II and III combined. Segment II dorsally with row of ca. four spine-like setae. Segment III sub-rectangular; at base approx. as wide as distal margin of segment II; ventral surface with short, spine-like, simple setae and short, fine, simple setae.
<figureCitationid="3FC98BE081A981CE1BF5DD99370CC953"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct."figureDoi="10.3897/zookeys.1168.104844.figure3"httpUri="https://binary.pensoft.net/fig/872205"pageId="0"pageNumber="231">3a, b</figureCitation>
,
<figureCitationid="E5AAAB09E8A161D643A129800B5615A7"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Arcobaetis sumbawensis sp. nov., larva a foreleg b middle leg c hind leg d subimaginal gonostylus developing under cuticle of last instar male larva e head. Scale bars: 0.05 mm (e); 0.1 mm (d)."figureDoi="10.3897/zookeys.1168.104844.figure4"httpUri="https://binary.pensoft.net/fig/872206"pageId="0"pageNumber="231">4a-c</figureCitation>
,
<figureCitationid="CBA77D846974743E1A7756B2EB0A3B15"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Arcobaetis sumbawensis sp. nov., larva, SEM pictures a fore claw, lateroventral view b middle claw, anterior view c middle claw, posterior view. Arrows: subapical setae. Scale bars: 0.01 mm."figureDoi="10.3897/zookeys.1168.104844.figure5"httpUri="https://binary.pensoft.net/fig/872207"pageId="0"pageNumber="231">5a-c</figureCitation>
). Long and slender, middle, and hind legs slenderer than fore legs. Ratio of leg segments: fore leg 1.3:1.0:0.7:0.2, middle and hind legs 1.2:1.0:0.6:0.1.
maximum width, outer margins slightly concave. Outer margin with row of ca. 12 short, curved, spine-like setae, on fore leg larger than on middle and hind legs. On ventral side with short, spine-like, pectinate, pointed setae; larger and denser on foreleg, smaller and less dense on middle and hind legs.
. Short, stout, pointed, pectinate setae near inner margin. Inner margin with row of short, curved, spine-like, pectinate setae, on apex two longer, curved, spine-like, pectinate setae. Outer margin with one apical, spine-like seta; on fore leg without other spine-like setae, on middle and hind legs with row of few medium, spine-like setae. Patella-tibial suture on proximal ¹/3 of tibia on all legs.
with one row of denticles; four or five distalmost denticles larger and directed distad; basally ca. 10 small to minute denticles; two vestigial subapical setae on anterior side, one vestigial subapical seta on posterior side (Fig.
<figureCitationid="E73D458B8DFF157D3D4BC6AE681CCE91"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Arcobaetis sumbawensis sp. nov., larva, SEM pictures a fore claw, lateroventral view b middle claw, anterior view c middle claw, posterior view. Arrows: subapical setae. Scale bars: 0.01 mm."figureDoi="10.3897/zookeys.1168.104844.figure5"httpUri="https://binary.pensoft.net/fig/872207"pageId="0"pageNumber="231">5a, b</figureCitation>
<figureCitationid="AC0034AACC4453A7E0D134CA7BF91F17"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct."figureDoi="10.3897/zookeys.1168.104844.figure3"httpUri="https://binary.pensoft.net/fig/872205"pageId="0"pageNumber="231">3c, d</figureCitation>
). Posterior margin of terga: I smooth, without spines; II-IX with triangular spines, increasing in length toward IX; slightly wider than long on tergum IV; row of spines on tergum IX interrupted by smaller spines in middle part behind bases of submedian setae (similar to Fig.
<figureCitationid="39D24D70F6E2E2375520D5FFCA7DD89E"captionStart="Figure 15"captionStartId="F15"captionText="Figure 15. Arcobaetis sripadai sp. nov., larval exuviae a, b fore and middle legs (triangles show points where patella-tibial suture crosses inner margin c claw d-k posterior margins of abdominal terga II-VI and VIII-X l-m posterior margin of abdominal sterna VIII and IX of male larva n paraproct. o-p fragments of cerci (lateral view)."figureDoi="10.3897/zookeys.1168.104844.figure15"httpUri="https://binary.pensoft.net/fig/872217"pageId="0"pageNumber="231">15j</figureCitation>
). Posterior margin of tergum X with median concavity with smaller spines (similar to Fig.
<figureCitationid="B894F2AD684FE6B7E22E753FD3CC012D"captionStart="Figure 15"captionStartId="F15"captionText="Figure 15. Arcobaetis sripadai sp. nov., larval exuviae a, b fore and middle legs (triangles show points where patella-tibial suture crosses inner margin c claw d-k posterior margins of abdominal terga II-VI and VIII-X l-m posterior margin of abdominal sterna VIII and IX of male larva n paraproct. o-p fragments of cerci (lateral view)."figureDoi="10.3897/zookeys.1168.104844.figure15"httpUri="https://binary.pensoft.net/fig/872217"pageId="0"pageNumber="231">15k</figureCitation>
Posterior margins of sterna: I-V smooth, without spines; VI-IX with triangular spines. On sternum IX of male mature larva row of narrow pointed spines between protogonostyli, smaller spines laterad of protogono-styli and larger, pointed spines laterad of them (similar to Fig.
<figureCitationid="B6D10173BBDC7795225EBBC840B14A56"captionStart="Figure 15"captionStartId="F15"captionText="Figure 15. Arcobaetis sripadai sp. nov., larval exuviae a, b fore and middle legs (triangles show points where patella-tibial suture crosses inner margin c claw d-k posterior margins of abdominal terga II-VI and VIII-X l-m posterior margin of abdominal sterna VIII and IX of male larva n paraproct. o-p fragments of cerci (lateral view)."figureDoi="10.3897/zookeys.1168.104844.figure15"httpUri="https://binary.pensoft.net/fig/872217"pageId="0"pageNumber="231">15m</figureCitation>
<figureCitationid="B15EE57F348C34DFA9692D36F6D3A705"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct."figureDoi="10.3897/zookeys.1168.104844.figure3"httpUri="https://binary.pensoft.net/fig/872205"pageId="0"pageNumber="231">3e</figureCitation>
). Present on abdominal segments I-VII. Tracheation in tergalius I limited to basal part of main trunk. In nine well preserved larvae, only a few tergalii were found, but insertions were always well developed: two tergalii I in one specimen; one tergalius IV and one tergalius V in another specimen, both very small, looking like replacements of naturally broken tergalii.
<figureCitationid="0B5C9400A2F9CB92CFD479F60939E449"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct."figureDoi="10.3897/zookeys.1168.104844.figure3"httpUri="https://binary.pensoft.net/fig/872205"pageId="0"pageNumber="231">3f</figureCitation>
). Posterior margin with ca. 16 stout spines. Surface scattered with scale bases and micropores.
<paragraphid="A234D0391AE32177C32845D1CFB2AF0E"pageId="0"pageNumber="231">The specific epithet refers to the type locality in Sumbawa (Indonesia).</paragraph>
