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<document id="F27A80262A0B532D081665189BC5B48C" ID-DOI="10.11646/zootaxa.2048.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5323018" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629617541061" checkinUser="felipe" docAuthor="Hendrich, Lars &amp; Watts, Chris H. S." docDate="2009" docId="03EC87A5FFF1581DB59EF9DBFA35497E" docLanguage="en" docName="zt02048p030.pdf" docOrigin="Zootaxa 2048 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2048.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Carabhydrus Watts 1978" docType="key" docVersion="4" lastPageNumber="29" masterDocId="FFD5FFDDFFEA5801B509FFFDFFA94A11" masterDocTitle="Taxonomic revision of the Australian predaceous water beetle genus Carabhydrus Watts, 1978 (Col. Dytiscidae, Hydroporinae, Hydroporini)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="28" updateTime="1699083959834" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C0093A71F27912E588D69B3F4E7A95CC">Taxonomic revision of the Australian predaceous water beetle genus Carabhydrus Watts, 1978 (Col. Dytiscidae, Hydroporinae, Hydroporini)</mods:title>
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<mods:namePart id="3106852A3760640053BA5F6CA659FD95">Hendrich, Lars</mods:namePart>
<mods:affiliation id="B2129AA73336E6DBC96F5243FFAC7891">Zoologische Staatssammlung, Münchhausenstrasse 21, D- 81247 München, Germany. E-mail: hendrich 1 @ aol. com</mods:affiliation>
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<mods:namePart id="2C16E775D0403026AB235306D6FE972F">Watts, Chris H. S.</mods:namePart>
<mods:affiliation id="121E5BC227715E2B5FDBAE553C6E3F8B">South Australian Museum, North Terrace, Adelaide, South Australia 5000, Australia. E-mail: watts. chris @ saugov. sa. gov. au</mods:affiliation>
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<paragraph id="8BFA36B3FFF1581AB59EF9DBFDB94C51" blockId="27.[151,528,1574,1600]" box="[151,528,1574,1600]" pageId="27" pageNumber="28">
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Key to Species of
<taxonomicName id="4C454D30FFF1581AB47EF9DBFDB94C51" ID-CoL="3H8D" authorityName="Watts" authorityYear="1978" box="[375,528,1574,1600]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAA1FFF1581AB47EF9DBFDB94C51" bold="true" box="[375,528,1574,1600]" italics="true" pageId="27" pageNumber="28">Carabhydrus</emphasis>
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</emphasis>
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</paragraph>
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<paragraph id="8BFA36B3FFF1581AB59EF98DFA354CB6" blockId="27.[151,1436,1648,1991]" pageId="27" pageNumber="28">
<keyLead id="30B49586FFF1581AB59EF98DFA354CB6" pageId="27" pageNumber="28">
1. Species smaller than
<quantity id="4CBD9B56FFF1581AB4A3F98DFE434C96" box="[426,490,1648,1671]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="27" pageNumber="28" unit="mm" value="2.0">2 mm</quantity>
, median lobe in ventral aspect strongly tapered at apex (
<figureCitation id="137E2A36FFF1581AB14BF98DFB3B4C96" box="[1090,1170,1648,1671]" captionStart="FIGURES 2027" captionStartId="7.[151,273,1521,1545]" captionTargetBox="[158,1434,181,1506]" captionTargetId="figure-227@7.[155,1440,166,1506]" captionTargetPageId="7" captionText="FIGURES 2027. Median lobe of aedeagus in ventral and lateral view (scale bar 0.2 mm): 20, 21Carabhydrus mubboonus; 22, 23—C. niger; 24, 25—C. plicatus; 26, 27—C. storeyi sp.n." figureDoi="http://doi.org/10.5281/zenodo.5323028" httpUri="https://zenodo.org/record/5323028/files/figure.png" pageId="27" pageNumber="28">Fig. 26</figureCitation>
); NE QLD ..................... ................................................................................................................................................................
<taxonomicName id="4C454D30FFF1581AB1F0F96DFAC94CB6" ID-CoL="QTZW" authorityName="Hendrich &amp; Watts" authorityYear="2009" box="[1273,1376,1680,1703]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="storeyi" status="sp. nov.">
<emphasis id="B931EAA1FFF1581AB1F0F96DFAA24CB6" box="[1273,1291,1680,1703]" italics="true" pageId="27" pageNumber="28">C</emphasis>
.
<emphasis id="B931EAA1FFF1581AB011F96DFAC94CB6" box="[1304,1376,1680,1703]" italics="true" pageId="27" pageNumber="28">storeyi</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="A20257DAFFF1581AB06FF972FA354CB6" box="[1382,1436,1679,1703]" pageId="27" pageNumber="28" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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1'. Species longer than
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............................................................................................................................................. 2
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</keyStep>
<keyStep id="30B12E16FFF1581AB59EF92DFA354D16" pageId="27" pageNumber="28">
<paragraph id="8BFA36B3FFF1581AB59EF92DFA324CF6" blockId="27.[151,1436,1648,1991]" box="[151,1435,1744,1767]" pageId="27" pageNumber="28">
<keyLead id="30B49586FFF1581AB59EF92DFA324CF6" box="[151,1435,1744,1767]" pageId="27" pageNumber="28">2. Elytron more or less uniformly pale testaceous to reddish black to black, without basal or subbasal spots............... 6</keyLead>
</paragraph>
<paragraph id="8BFA36B3FFF1581AB59EF90DFA354D16" blockId="27.[151,1436,1648,1991]" box="[151,1436,1776,1799]" pageId="27" pageNumber="28">
<keyLead id="30B49586FFF1581AB59EF90DFA354D16" box="[151,1436,1776,1799]" pageId="27" pageNumber="28">2'. Elytron black or brown except a large reddish subbasal spot and on some specimens also the apex .......................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B12E16FFF1581AB59EF8EDFA354D96" pageId="27" pageNumber="28">
<paragraph id="8BFA36B3FFF1581AB59EF8EDFA3C4D76" blockId="27.[151,1436,1648,1991]" pageId="27" pageNumber="28">
<keyLead id="30B49586FFF1581AB59EF8EDFA3C4D76" pageId="27" pageNumber="28">
3. Species larger than
<quantity id="4CBD9B56FFF1581AB495F8EDFE744D36" box="[412,477,1808,1831]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="28" unit="mm" value="3.0">3 mm</quantity>
. Largest species of the genus (
<quantity id="4CBD9B56FFF1581AB62FF8EDFC024D36" box="[806,939,1808,1831]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="3.6" metricValueMin="3.1" pageId="27" pageNumber="28" unit="mm" value="3.35" valueMax="3.6" valueMin="3.1">3.13.6 mm</quantity>
), pronotum more parallel sided (
<figureCitation id="137E2A36FFF1581AB003F8EDFAE24D36" box="[1290,1355,1808,1831]" captionStart="FIGURES 14" captionStartId="3.[151,270,1976,2000]" captionTargetBox="[275,1299,197,1937]" captionTargetId="figure-48@3.[271,1318,197,1942]" captionTargetPageId="3" captionText="FIGURES 14. Habitus of Carabhydrus. 1—Carabhydrus andreas (male); 2—C. innae sp.n. (holotype, male); 3—C. plicatus; 4—C. storeyi sp.n. (paratype, male) (scale bar 1 mm) (Photos: L. Hendrich)." figureDoi="http://doi.org/10.5281/zenodo.5323020" httpUri="https://zenodo.org/record/5323020/files/figure.png" pageId="27" pageNumber="28">Fig. 2</figureCitation>
) and at base broader than head, with three deep longitudinal depressions on elytra, first mesotarsal segment in male formed like a triangular hook (
<figureCitation id="137E2A36FFF1581AB4B3F8ADFDA04D76" box="[442,521,1872,1895]" captionStart="FIGURES 1019" captionStartId="6.[151,273,1904,1928]" captionTargetBox="[178,1412,187,1875]" captionTargetId="figure-83@6.[172,1416,172,1881]" captionTargetPageId="6" captionText="FIGURES 1019. Median lobe of aedeagus in ventral and lateral view (scale bar 0.2 mm): 10, 11—Carabhydrus andreas and 12, 13—C. innae sp.n.; 14, 15—C. innae sp.n. right paramere (14) and first two mesotarsal segments (15) in dorsal view (scale bar 0.2 mm). Median lobe of aedeagus in ventral and lateral view: 16, 17—C. janmillerae sp.n. and 18,19—C. monteithi." figureDoi="http://doi.org/10.5281/zenodo.5323026" httpUri="https://zenodo.org/record/5323026/files/figure.png" pageId="27" pageNumber="28">Fig. 15</figureCitation>
); N NSW, S VIC....................................................................................
<taxonomicName id="4C454D30FFF1581AB1F6F8ADFAF04D76" ID-CoL="QTZP" authorityName="Hendrich &amp; Watts" authorityYear="2009" box="[1279,1369,1872,1895]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="innae" status="sp. nov.">
<emphasis id="B931EAA1FFF1581AB1F6F8ADFAB84D76" box="[1279,1297,1872,1895]" italics="true" pageId="27" pageNumber="28">C</emphasis>
.
<emphasis id="B931EAA1FFF1581AB017F8ADFAF04D76" box="[1310,1369,1872,1895]" italics="true" pageId="27" pageNumber="28">innae</emphasis>
</taxonomicName>
<emphasis id="B931EAA1FFF1581AB069F8B2FA3C4D76" bold="true" box="[1376,1429,1871,1895]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A20257DAFFF1581AB069F8B2FA3C4D76" box="[1376,1429,1871,1895]" pageId="27" pageNumber="28" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BFA36B3FFF1581AB59EF88DFA354D96" blockId="27.[151,1436,1648,1991]" box="[151,1436,1904,1927]" pageId="27" pageNumber="28">
<keyLead id="30B49586FFF1581AB59EF88DFA354D96" box="[151,1436,1904,1927]" pageId="27" pageNumber="28">
3'. Species smaller than
<quantity id="4CBD9B56FFF1581AB4A3F88DFE434D96" box="[426,490,1904,1927]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="28" unit="mm" value="3.0">3 mm</quantity>
, first mesotarsal segment in all known males without a triangular hook.......................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B12E16FFF1581DB59EF86DFA324ADE" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<paragraph id="8BFA36B3FFF1581AB59EF86DFA354DD6" blockId="27.[151,1436,1648,1991]" pageId="27" pageNumber="28">
<keyLead id="30B49586FFF1581AB59EF86DFA354DD6" pageId="27" pageNumber="28">
4. Body flattened, more rounded, elytral grooves well defined and very deep, elytral spots less defined (
<figureCitation id="137E2A36FFF1581AB025F86DFAD94DB6" box="[1324,1392,1936,1959]" captionStart="FIGURES 59" captionStartId="5.[151,275,1938,1962]" captionTargetBox="[201,1382,187,1918]" captionTargetId="figure-53@5.[195,1391,172,1923]" captionTargetPageId="5" captionText="FIGURES 59. Habitus of Carabhydrus. 5—Carabhydrus turaki sp.n. (holotype, female); 6—C. stephanieae (paratype); 7—C. mubboonus (female); 8—C. janmillerae sp.n. (paratype, female); 9—C. niger (female) (scale bar 1 mm) (Photos: L. Hendrich)." figureDoi="http://doi.org/10.5281/zenodo.5323022" httpUri="https://zenodo.org/record/5323022/files/figure.png" pageId="27" pageNumber="28">Fig. 5</figureCitation>
), in female half of elytra covered by a fine microreticulation, male unknown; N NSW
<emphasis id="B931EAA1FFF1581AB139F84DFAC94DD6" box="[1072,1376,1968,1991]" italics="true" pageId="27" pageNumber="28">
...............................
<taxonomicName id="4C454D30FFF1581AB00CF84DFAC94DD6" ID-CoL="QTZX" authorityName="Hendrich &amp; Watts" authorityYear="2009" box="[1285,1376,1968,1991]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="turaki" status="sp. nov.">C. turaki</taxonomicName>
</emphasis>
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<taxonomicNameLabel id="A20257DAFFF1581AB06FF852FA354DD6" box="[1382,1436,1967,1991]" pageId="27" pageNumber="28" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="8BFA36B3FFF6581DB59EFF65FA324ADE" blockId="28.[151,1436,152,879]" pageId="28" pageNumber="29">
<keyLead id="30B49586FFF6581DB59EFF65FA324ADE" pageId="28" pageNumber="29">
4'. Body more convex, elytral grooves less defined, elytral spots well defined (
<figureCitation id="137E2A36FFF6581DB6FBFF65FBF84ABE" box="[1010,1105,152,175]" captionStart="FIGURES 14" captionStartId="3.[151,270,1976,2000]" captionTargetBox="[275,1299,197,1937]" captionTargetId="figure-48@3.[271,1318,197,1942]" captionTargetPageId="3" captionText="FIGURES 14. Habitus of Carabhydrus. 1—Carabhydrus andreas (male); 2—C. innae sp.n. (holotype, male); 3—C. plicatus; 4—C. storeyi sp.n. (paratype, male) (scale bar 1 mm) (Photos: L. Hendrich)." figureDoi="http://doi.org/10.5281/zenodo.5323020" httpUri="https://zenodo.org/record/5323020/files/figure.png" pageId="28" pageNumber="29">Figs 1, 3</figureCitation>
), in female only apex of elytra covered with fine microreticulation .............................................................................................................................. 5
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</paragraph>
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<keyStep id="30B12E16FFF6581DB59EFF25FA3C4B7E" pageId="28" pageNumber="29">
<paragraph id="8BFA36B3FFF6581DB59EFF25FA3C4B1E" blockId="28.[151,1436,152,879]" pageId="28" pageNumber="29">
<keyLead id="30B49586FFF6581DB59EFF25FA3C4B1E" pageId="28" pageNumber="29">
5. Pronotum black (
<figureCitation id="137E2A36FFF6581DB476FF25FE144AFE" box="[383,445,216,239]" captionStart="FIGURES 14" captionStartId="3.[151,270,1976,2000]" captionTargetBox="[275,1299,197,1937]" captionTargetId="figure-48@3.[271,1318,197,1942]" captionTargetPageId="3" captionText="FIGURES 14. Habitus of Carabhydrus. 1—Carabhydrus andreas (male); 2—C. innae sp.n. (holotype, male); 3—C. plicatus; 4—C. storeyi sp.n. (paratype, male) (scale bar 1 mm) (Photos: L. Hendrich)." figureDoi="http://doi.org/10.5281/zenodo.5323020" httpUri="https://zenodo.org/record/5323020/files/figure.png" pageId="28" pageNumber="29">Fig. 1</figureCitation>
), aedeagus in ventral view with apex of each apical lobe sharply pointed and finger-like (
<figureCitation id="137E2A36FFF6581DB078FF25FF4D4B1E" captionStart="FIGURES 1019" captionStartId="6.[151,273,1904,1928]" captionTargetBox="[178,1412,187,1875]" captionTargetId="figure-83@6.[172,1416,172,1881]" captionTargetPageId="6" captionText="FIGURES 1019. Median lobe of aedeagus in ventral and lateral view (scale bar 0.2 mm): 10, 11—Carabhydrus andreas and 12, 13—C. innae sp.n.; 14, 15—C. innae sp.n. right paramere (14) and first two mesotarsal segments (15) in dorsal view (scale bar 0.2 mm). Median lobe of aedeagus in ventral and lateral view: 16, 17—C. janmillerae sp.n. and 18,19—C. monteithi." figureDoi="http://doi.org/10.5281/zenodo.5323026" httpUri="https://zenodo.org/record/5323026/files/figure.png" pageId="28" pageNumber="29">Fig. 10</figureCitation>
); VIC, NSW, ACT ..................................................................................................................................
<taxonomicName id="4C454D30FFF6581DB028FF05FA3C4B1E" ID-CoL="QTZN" authorityName="Zwick" authorityYear="1981" box="[1313,1429,248,271]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="andreas">
<emphasis id="B931EAA1FFF6581DB028FF05FA3C4B1E" box="[1313,1429,248,271]" italics="true" pageId="28" pageNumber="29">C. andreas</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFA36B3FFF6581DB59EFEE5FA3C4B7E" blockId="28.[151,1436,152,879]" pageId="28" pageNumber="29">
<keyLead id="30B49586FFF6581DB59EFEE5FA3C4B7E" pageId="28" pageNumber="29">
5'. Pronotum reddish, only base of pronotum black (
<figureCitation id="137E2A36FFF6581DB7D4FEE5FCB44B3E" box="[733,797,280,303]" captionStart="FIGURES 14" captionStartId="3.[151,270,1976,2000]" captionTargetBox="[275,1299,197,1937]" captionTargetId="figure-48@3.[271,1318,197,1942]" captionTargetPageId="3" captionText="FIGURES 14. Habitus of Carabhydrus. 1—Carabhydrus andreas (male); 2—C. innae sp.n. (holotype, male); 3—C. plicatus; 4—C. storeyi sp.n. (paratype, male) (scale bar 1 mm) (Photos: L. Hendrich)." figureDoi="http://doi.org/10.5281/zenodo.5323020" httpUri="https://zenodo.org/record/5323020/files/figure.png" pageId="28" pageNumber="29">Fig. 3</figureCitation>
), male mesotibia with a well developed subapical tooth on inner edge, aedeagus in ventral view with apical lobes blunt and broadly rounded (
<figureCitation id="137E2A36FFF6581DB13FFEC5FB2F4B5E" box="[1078,1158,312,335]" captionStart="FIGURES 2027" captionStartId="7.[151,273,1521,1545]" captionTargetBox="[158,1434,181,1506]" captionTargetId="figure-227@7.[155,1440,166,1506]" captionTargetPageId="7" captionText="FIGURES 2027. Median lobe of aedeagus in ventral and lateral view (scale bar 0.2 mm): 20, 21Carabhydrus mubboonus; 22, 23—C. niger; 24, 25—C. plicatus; 26, 27—C. storeyi sp.n." figureDoi="http://doi.org/10.5281/zenodo.5323028" httpUri="https://zenodo.org/record/5323028/files/figure.png" pageId="28" pageNumber="29">Fig. 24</figureCitation>
); S QLD, N NSW ..........
<emphasis id="B931EAA1FFF6581DB5C7FEA5FA3C4B7E" box="[206,1429,344,367]" italics="true" pageId="28" pageNumber="29">
......................................................................................................................................................................
<taxonomicName id="4C454D30FFF6581DB02BFEA5FA3C4B7E" ID-CoL="699D6" authorityName="Watts Ribera &amp; Hernando" authorityYear="1978" box="[1314,1429,344,367]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="plicatus">C. plicatus</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B12E16FFF6581DB59EFE85FA324BBE" pageId="28" pageNumber="29">
<paragraph id="8BFA36B3FFF6581DB59EFE85FA324B9E" blockId="28.[151,1436,152,879]" box="[151,1435,376,399]" pageId="28" pageNumber="29">
<keyLead id="30B49586FFF6581DB59EFE85FA324B9E" box="[151,1435,376,399]" pageId="28" pageNumber="29">6. Dorsal side pale testaceous or brown, eyes about half normal size or absent .............................................................. 7</keyLead>
</paragraph>
<paragraph id="8BFA36B3FFF6581DB59EFE65FA324BBE" blockId="28.[151,1436,152,879]" box="[151,1435,408,431]" pageId="28" pageNumber="29">
<keyLead id="30B49586FFF6581DB59EFE65FA324BBE" box="[151,1435,408,431]" pageId="28" pageNumber="29">6' Dorsal side blackish-brown or black, eyes normal ..................................................................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B12E16FFF6581DB59EFE45FA35483E" pageId="28" pageNumber="29">
<paragraph id="8BFA36B3FFF6581DB59EFE45FA354BFE" blockId="28.[151,1436,152,879]" pageId="28" pageNumber="29">
<keyLead id="30B49586FFF6581DB59EFE45FA354BFE" pageId="28" pageNumber="29">
7. Eyes present, ventral side pale to dark brown (
<figureCitation id="137E2A36FFF6581DB7BAFE45FD5B4BDE" box="[691,754,440,463]" captionStart="FIGURES 59" captionStartId="5.[151,275,1938,1962]" captionTargetBox="[201,1382,187,1918]" captionTargetId="figure-53@5.[195,1391,172,1923]" captionTargetPageId="5" captionText="FIGURES 59. Habitus of Carabhydrus. 5—Carabhydrus turaki sp.n. (holotype, female); 6—C. stephanieae (paratype); 7—C. mubboonus (female); 8—C. janmillerae sp.n. (paratype, female); 9—C. niger (female) (scale bar 1 mm) (Photos: L. Hendrich)." figureDoi="http://doi.org/10.5281/zenodo.5323022" httpUri="https://zenodo.org/record/5323022/files/figure.png" pageId="28" pageNumber="29">Fig. 8</figureCitation>
), pronotum narrower than elytra in middle, basal part of median lobe with sclerotized, broad and shell-like coverage (
<figureCitation id="137E2A36FFF6581DB7F2FE25FCDF4BFE" box="[763,886,472,495]" captionStart="FIGURES 1019" captionStartId="6.[151,273,1904,1928]" captionTargetBox="[178,1412,187,1875]" captionTargetId="figure-83@6.[172,1416,172,1881]" captionTargetPageId="6" captionText="FIGURES 1019. Median lobe of aedeagus in ventral and lateral view (scale bar 0.2 mm): 10, 11—Carabhydrus andreas and 12, 13—C. innae sp.n.; 14, 15—C. innae sp.n. right paramere (14) and first two mesotarsal segments (15) in dorsal view (scale bar 0.2 mm). Median lobe of aedeagus in ventral and lateral view: 16, 17—C. janmillerae sp.n. and 18,19—C. monteithi." figureDoi="http://doi.org/10.5281/zenodo.5323026" httpUri="https://zenodo.org/record/5323026/files/figure.png" pageId="28" pageNumber="29">Figs 16, 17</figureCitation>
), S NSW ................................
<taxonomicName id="4C454D30FFF6581DB1CFFE25FAC94BFE" ID-CoL="QTZQ" authorityName="Hendrich &amp; Watts" authorityYear="2009" box="[1222,1376,472,495]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="janmillerae" status="sp. nov.">
<emphasis id="B931EAA1FFF6581DB1CFFE25FB714BFE" box="[1222,1240,472,495]" italics="true" pageId="28" pageNumber="29">C</emphasis>
.
<emphasis id="B931EAA1FFF6581DB1ECFE25FAC94BFE" box="[1253,1376,472,495]" italics="true" pageId="28" pageNumber="29">janmillerae</emphasis>
</taxonomicName>
<emphasis id="B931EAA1FFF6581DB06FFE2AFA354BFE" bold="true" box="[1382,1436,471,495]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A20257DAFFF6581DB06FFE2AFA354BFE" box="[1382,1436,471,495]" pageId="28" pageNumber="29" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BFA36B3FFF6581DB59EFE05FA35483E" blockId="28.[151,1436,152,879]" pageId="28" pageNumber="29">
<keyLead id="30B49586FFF6581DB59EFE05FA35483E" pageId="28" pageNumber="29">
7'. Eyes absent, pale testaceous (
<figureCitation id="137E2A36FFF6581DB704FE05FDF9481E" box="[525,592,504,527]" captionStart="FIGURES 59" captionStartId="5.[151,275,1938,1962]" captionTargetBox="[201,1382,187,1918]" captionTargetId="figure-53@5.[195,1391,172,1923]" captionTargetPageId="5" captionText="FIGURES 59. Habitus of Carabhydrus. 5—Carabhydrus turaki sp.n. (holotype, female); 6—C. stephanieae (paratype); 7—C. mubboonus (female); 8—C. janmillerae sp.n. (paratype, female); 9—C. niger (female) (scale bar 1 mm) (Photos: L. Hendrich)." figureDoi="http://doi.org/10.5281/zenodo.5323022" httpUri="https://zenodo.org/record/5323022/files/figure.png" pageId="28" pageNumber="29">Fig. 6</figureCitation>
), pronotum broader than elytra in middle, male unknown, stygobitic species; NSW
<emphasis id="B931EAA1FFF6581DB40DFDE5FA35483E" box="[260,1436,536,559]" italics="true" pageId="28" pageNumber="29">
........................................................................................................................................................
<taxonomicName id="4C454D30FFF6581DB1F7FDE5FA35483E" ID-CoL="QTZV" box="[1278,1436,536,559]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="stephanieae">C. stephanieae</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B12E16FFF6581DB59EFDC5FA3248BE" pageId="28" pageNumber="29">
<paragraph id="8BFA36B3FFF6581DB59EFDC5FA3C487E" blockId="28.[151,1436,152,879]" pageId="28" pageNumber="29">
<keyLead id="30B49586FFF6581DB59EFDC5FA3C487E" pageId="28" pageNumber="29">
8. Aedeagus in ventral aspect (
<figureCitation id="137E2A36FFF6581DB4F2FDC5FDE2485E" box="[507,587,568,591]" captionStart="FIGURES 2027" captionStartId="7.[151,273,1521,1545]" captionTargetBox="[158,1434,181,1506]" captionTargetId="figure-227@7.[155,1440,166,1506]" captionTargetPageId="7" captionText="FIGURES 2027. Median lobe of aedeagus in ventral and lateral view (scale bar 0.2 mm): 20, 21Carabhydrus mubboonus; 22, 23—C. niger; 24, 25—C. plicatus; 26, 27—C. storeyi sp.n." figureDoi="http://doi.org/10.5281/zenodo.5323028" httpUri="https://zenodo.org/record/5323028/files/figure.png" pageId="28" pageNumber="29">Fig. 22</figureCitation>
) with a deep, broad, subapical notch on each side delimiting a broad, apically bilobed apex; VIC, NSW, TAS
<emphasis id="B931EAA1FFF6581DB71DFDA5FA3C487E" box="[532,1429,600,623]" italics="true" pageId="28" pageNumber="29">
.........................................................................................................................
<taxonomicName id="4C454D30FFF6581DB037FDA5FA3C487E" ID-CoL="QTZT" box="[1342,1429,600,623]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="niger">C. niger</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BFA36B3FFF6581DB59EFD85FA3248BE" blockId="28.[151,1436,152,879]" pageId="28" pageNumber="29">
<keyLead id="30B49586FFF6581DB59EFD85FA3248BE" pageId="28" pageNumber="29">
8'. Aedeagus in ventral aspect (
<figureCitation id="137E2A36FFF6581DB4FCFD85FDEA489E" box="[501,579,632,655]" captionStart="FIGURES 2027" captionStartId="7.[151,273,1521,1545]" captionTargetBox="[158,1434,181,1506]" captionTargetId="figure-227@7.[155,1440,166,1506]" captionTargetPageId="7" captionText="FIGURES 2027. Median lobe of aedeagus in ventral and lateral view (scale bar 0.2 mm): 20, 21Carabhydrus mubboonus; 22, 23—C. niger; 24, 25—C. plicatus; 26, 27—C. storeyi sp.n." figureDoi="http://doi.org/10.5281/zenodo.5323028" httpUri="https://zenodo.org/record/5323028/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
) with a narrow subapical notch on each side, apex subtriangular and not broadly rounded ......................................................................................................................................................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B12E16FFF6581DB59EFD45FA35497E" pageId="28" pageNumber="29">
<paragraph id="8BFA36B3FFF6581DB59EFD45FA3C491E" blockId="28.[151,1436,152,879]" pageId="28" pageNumber="29">
<keyLead id="30B49586FFF6581DB59EFD45FA3C491E" pageId="28" pageNumber="29">
9. Elytron with mesal groove extending almost to apex; pronotum with lateral margin more or less evenly rounded except for short sinuation before posterolateral angle; aedeagus in lateral aspect lacking ventromedial projections (
<figureCitation id="137E2A36FFF6581DB5C6FD05FEB7491E" box="[207,286,760,783]" captionStart="FIGURES 2027" captionStartId="7.[151,273,1521,1545]" captionTargetBox="[158,1434,181,1506]" captionTargetId="figure-227@7.[155,1440,166,1506]" captionTargetPageId="7" captionText="FIGURES 2027. Median lobe of aedeagus in ventral and lateral view (scale bar 0.2 mm): 20, 21Carabhydrus mubboonus; 22, 23—C. niger; 24, 25—C. plicatus; 26, 27—C. storeyi sp.n." figureDoi="http://doi.org/10.5281/zenodo.5323028" httpUri="https://zenodo.org/record/5323028/files/figure.png" pageId="28" pageNumber="29">Fig. 21</figureCitation>
); NE QLD
<emphasis id="B931EAA1FFF6581DB49FFD05FA3C491E" box="[406,1429,760,783]" italics="true" pageId="28" pageNumber="29">
..................................................................................................................................
<taxonomicName id="4C454D30FFF6581DB1F2FD05FA3C491E" ID-CoL="QTZS" authorityName="Larson &amp; Storey" authorityYear="1994" box="[1275,1429,760,783]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="mubboonus">C. mubboonus</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BFA36B3FFF6581DB59EFCE5FA35497E" blockId="28.[151,1436,152,879]" pageId="28" pageNumber="29">
<keyLead id="30B49586FFF6581DB59EFCE5FA35497E" pageId="28" pageNumber="29">
9'. Elytron with mesal groove shallow in apical third and discontinuous before apex; pronotum narrow, sides sinuate basally and more or less straight and parallel before posterolateral angles; aedeagus in lateral aspect (
<figureCitation id="137E2A36FFF6581DB1F7FCC5FAE2495E" box="[1278,1355,824,847]" captionStart="FIGURES 1019" captionStartId="6.[151,273,1904,1928]" captionTargetBox="[178,1412,187,1875]" captionTargetId="figure-83@6.[172,1416,172,1881]" captionTargetPageId="6" captionText="FIGURES 1019. Median lobe of aedeagus in ventral and lateral view (scale bar 0.2 mm): 10, 11—Carabhydrus andreas and 12, 13—C. innae sp.n.; 14, 15—C. innae sp.n. right paramere (14) and first two mesotarsal segments (15) in dorsal view (scale bar 0.2 mm). Median lobe of aedeagus in ventral and lateral view: 16, 17—C. janmillerae sp.n. and 18,19—C. monteithi." figureDoi="http://doi.org/10.5281/zenodo.5323026" httpUri="https://zenodo.org/record/5323026/files/figure.png" pageId="28" pageNumber="29">Fig. 19</figureCitation>
) with a pair of ventromedial projections; N NSW
<emphasis id="B931EAA1FFF6581DB785FCA5FA35497E" box="[652,1436,856,879]" italics="true" pageId="28" pageNumber="29">
.................................................................................................
<taxonomicName id="4C454D30FFF6581DB011FCA5FA35497E" ID-CoL="QTZR" authorityName="Watts" authorityYear="1978" box="[1304,1436,856,879]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="monteithi">C. monteithi</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>