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<document id="6920A168C970467BB16EEACFA221FEF1" ID-DOI="10.5281/zenodo.5391395" ID-ISSN="1638-9387" ID-Zenodo-Dep="5391395" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1630314607455" checkinUser="marcus" docAuthor="Newman, William A." docDate="2000" docId="0381B717FFB4D029FD04FF7279B09BEB" docLanguage="en" docName="Zoosystema.2000.22.1.71-84.pdf" docOrigin="Zoosystema 22 (1)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Imbricaverruca yamaguchii Newman 2000, n. sp." docType="treatment" docUuid="6958BBE8-3871-4F1F-9CAA-BCFB88A86D0E" docUuidSource="ZooBank" docVersion="1" lastPageNumber="81" masterDocId="FFB8CF6FFFB7D023FFFDFFA2785A9F69" masterDocTitle="A new genus and species of barnacle (Cirripedia, Verrucomorpha) associated with vents of the Lau Back-Arc Basin: its gross morphology, inferred first juvenile stage and affinities" masterLastPageNumber="84" masterPageNumber="71" pageNumber="74" updateTime="1729878602252" updateUser="juliana" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FEEF0E5F00E2CD6F2908428568994C40">A new genus and species of barnacle (Cirripedia, Verrucomorpha) associated with vents of the Lau Back-Arc Basin: its gross morphology, inferred first juvenile stage and affinities</mods:title>
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<mods:name id="93372867EE5FB6CC3A3E6EE9039D8B63" type="personal">
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<mods:roleTerm id="9ED2E209800D41476B62D7CBD531EBD4">Author</mods:roleTerm>
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<mods:namePart id="FD9CD4FF58D017E5A17580277E104604">Newman, William A.</mods:namePart>
<mods:affiliation id="1D5431326D704F6581380072609819BF">Scripps Institution of Oceanography, University of California San Diego, La Jolla, CA 92093 - 0202 (USA) wnewman @ ucsd. edu</mods:affiliation>
<mods:nameIdentifier id="A44D67DF39F37E540EE70F5C12C0CF7D" type="email">wnewman@ucsd.edu</mods:nameIdentifier>
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<mods:typeOfResource id="BCB05CB818C338B455A06A17F825304B">text</mods:typeOfResource>
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<mods:title id="50DE357D5BE5B94728D2A8717C36D889">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="21B954BCC39E461122F5DD13DBA9EB2A">
<mods:date id="8CB5239DD1C0B4FF2078A454C8D652EE">2000</mods:date>
<mods:detail id="6C63C4825F6558E6184E95C5FC8D7116" type="pubDate">
<mods:number id="E536990633A51D735043D1C9B412E1A7">2000-12-31</mods:number>
</mods:detail>
<mods:detail id="155DEE2DD925ACB03190CA628414271B" type="volume">
<mods:number id="267B2991EE8393DEB4CD07F21F612633">22</mods:number>
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<mods:number id="CD2338EA5A54006DFF5E47E981272C4D">1</mods:number>
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<mods:classification id="1C8C6FD9D23D830EF4B770359BBE9BE5">journal article</mods:classification>
<mods:identifier id="5671348DD0BCBB579107C0C069CC2710" type="DOI">10.5281/zenodo.5391395</mods:identifier>
<mods:identifier id="64BA23919647567E1BA3A00C31C21EA4" type="ISSN">1638-9387</mods:identifier>
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<mods:identifier id="2DDCFBDE244EBA6C0E5E9839E8A93BE3" type="ZooBank">99E068D5-766D-4E67-8C75-8253302C3513</mods:identifier>
</mods:mods>
<treatment id="0381B717FFB4D029FD04FF7279B09BEB" LSID="urn:lsid:zoobank.org:act:6958BBE8-3871-4F1F-9CAA-BCFB88A86D0E" httpUri="http://treatment.plazi.org/id/0381B717FFB4D029FD04FF7279B09BEB" lastPageId="10" lastPageNumber="81" pageId="3" pageNumber="74">
<subSubSection id="C332558AFFB4D020FD04FF727C3C9F83" box="[761,1126,208,234]" pageId="3" pageNumber="74" type="nomenclature">
<paragraph id="8B970601FFB4D020FD04FF727C3C9F83" blockId="3.[761,1126,208,267]" box="[761,1126,208,234]" pageId="3" pageNumber="74">
<taxonomicName id="4C287D82FFB4D020FD04FF727C7E9F83" authority="Newman, 2000" authorityName="Newman" authorityYear="2000" box="[761,1060,208,234]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="yamaguchii" status="sp. nov.">
<emphasis id="B95CDA13FFB4D020FD04FF727C7E9F83" bold="true" box="[761,1060,208,234]" italics="true" pageId="3" pageNumber="74">Imbricaverruca yamaguchii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F6768FFB4D020FBD6FF727C3C9F83" box="[1067,1126,208,234]" pageId="3" pageNumber="74" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C332558AFFB4D020FCD3FF537C6B9E62" box="[814,1073,241,267]" pageId="3" pageNumber="74" type="description">
<paragraph id="8B970601FFB4D020FCD3FF537C6B9E62" blockId="3.[761,1126,208,267]" box="[814,1073,241,267]" pageId="3" pageNumber="74">
(
<figureCitation id="13131A84FFB4D020FCC8FF537BD09E62" box="[821,906,241,267]" captionStart="FIG" captionStartId="2.[114,125,799,816]" captionTargetBox="[262,1053,208,766]" captionTargetId="figure-307@2.[248,1067,202,776]" captionTargetPageId="2" captionText="FIG. 1. — A, Imbricaverruca yamaguchii n. gen. and n. sp. (holotype, MNHN Ci2710); B, Verruca s.l., both viewed from above with the right scutum and tergum forming the operculum.Note that in the former the four plated wall appears to be covered largely by imbricating plates and the operculum includes a large median latus,characters that readily distinguish it from Neoverruca. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5391397" httpUri="https://zenodo.org/record/5391397/files/figure.png" pageId="3" pageNumber="74">Figs 1A</figureCitation>
;
<figureCitation id="13131A84FFB4D020FC6BFF537BFC9E62" box="[918,934,241,267]" captionStart="FIG" captionStartId="4.[114,125,1384,1401]" captionTargetBox="[213,1067,193,1353]" captionTargetId="figure-91@4.[208,1091,186,1354]" captionTargetPageId="4" captionText="FIG. 2. — Plates of Imbricaverruca yamaguchii gen. et n. sp. (paratype, MNHN Ci2711); A, B, viewed from above and from the rostral end, respectively; A, note the operculum, comprising the plates of the right side (MS-L-MT), includes a large median latus; B, note the slightly open aperture to the mantle cavity between the occludent margins of the movable (MS &amp; MT) and the fixed (FS &amp; FT) scutum and tergum, respectively; C-M, variously disarticulated hard parts; C, interior view of wall (R-C-FS-FT) and operculum (MS-L- MT) with all of the imbricating plates, except the uppermost of the three principle lateral tiers of the right side (rl1-l1-cl1), stripped away [an approximation of the rostrum (R), which did not survive dissection, is indicated by the dashed line]; D, E, movable scutum (MS); external and internal views respectively; scutal adductor muscle, inserting from just below to up under the apical concavity in Fig. 2E, extends to its origin on the fixed scutum (FS, Fig. 2G; the relationship of MS to FS can be seen in Fig 2C); F, G, fixed scutum (FS); basal and internal views, respectively. Dashed line in G indicates origin of scutal adductor muscle but no scar is evident. The relationship of the two protuberances (best seen near the tergal margin in Fig. 2G) to the fixed tergum (FT, Fig. 2J, K), was not resolved; H, I, movable tergum (MT); external and internal views respectively (external corrosion in bleach and cleaning inadvertently obliterated growth line ornamentation in H); J, K, fixed tergum (FT); external and internal views respectively; L, M, carina from below and above respectively (the rostrum did not survive dissection and cleaning). Abbreviations: C, carina; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral plates forming three tiers each four plates high, respectively; FS, FT, fixed scutum and tergum; L, median latus; MS, MT, movable scutum and tergum; R, rostrum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391399" httpUri="https://zenodo.org/record/5391399/files/figure.png" pageId="3" pageNumber="74">2</figureCitation>
;
<figureCitation id="13131A84FFB4D020FC4FFF537B9B9E62" box="[946,961,241,267]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="3" pageNumber="74">3</figureCitation>
;
<figureCitation id="13131A84FFB4D020FC33FF537BAB9E62" box="[974,1009,241,267]" captionStart="Fig" captionStartId="7.[113,141,1469,1486]" captionTargetBox="[112,1165,215,1445]" captionTargetId="figure-22@7.[139,1170,219,1510]" captionTargetPageId="7" captionText="Fig. 4. — Schematic plan views and transverse sections of grades of skeletal organization from the most primitive sessile barnacles (A, Brachylepadomorpha) through the asymmetrical sessile barnacles (B-H, Verrucomorpha). Of the eight verrucid genera currently recognized (Young 1998) only the four most representative ones (Altiverruca, Newmaniverruca, Metaverruca and Verruca s.s.) are depicted here. Right sides are to the viewers right and rostral ends are downward in plan views or coming out of the page in transverse sections. The transverse sections are slightly rostral of the rostro-carinal gap or suture. While there are numerous whorls of imbricating plates standing in tiers four plates high in brachylepadomorphs, and on the movable side of neoverrucids, only those of the R-C gap are labeled. Abbreviations: C, R, carina and rostrum; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral tiers of imbricating plates respectively; cl, rl, the pair of imbricating plates in proverrucids; FS, FT, fixed scutum and tergum; hl, hinge line; L, median latus (completely lost in proverrucids and verrucids); m, myophore; MS, MT, movable scutum and tergum; sam, scutal adductor muscle; S, T, normal terga and scuta in brachylepadomorphs; dashed horizontal lines: membranous basis (transverse sections F-H after Young 1998)." figureDoi="http://doi.org/10.5281/zenodo.13988759" httpUri="https://zenodo.org/record/13988759/files/figure.png" pageId="3" pageNumber="74">4C</figureCitation>
; 5; 6)
</paragraph>
</subSubSection>
<subSubSection id="C332558AFFB4D020FD58FE897ABC9DA9" pageId="3" pageNumber="74" type="materials_examined">
<paragraph id="8B970601FFB4D020FD58FE897ABC9DA9" blockId="3.[677,1210,299,865]" pageId="3" pageNumber="74">
<materialsCitation id="3B400C5CFFB4D020FD58FE897AB89DA9" collectionCode="CENTOB" collectorName="A. - M. Alayse-Danet" country="Tonga" county="Hine Hina" determinerName="Newman &amp; Yamaguchi &amp; Southward &amp; Newman &amp; Desbruyeres" elevation="1900" latitude="-22.533333" location="French" longLatPrecision="1259" longitude="-176.71666" municipality="Newman" pageId="3" pageNumber="74" specimenCode="BL-01, BL-03" specimenCount="2" specimenCount-adult="2">
MATERIAL. — Specimens were collected by the
<location id="8EF750DAFFB4D020FB88FE8E7CE09E2D" LSID="urn:lsid:plazi:treatment:0381B717FFB4D029FD04FF7279B09BEB:8EF750DAFFB4D020FB88FE8E7CE09E2D" box="[1141,1210,300,324]" country="Tonga" county="Hine Hina" latitude="-22.533333" longLatPrecision="1259" longitude="-176.71666" municipality="Newman" name="French" pageId="3" pageNumber="74">French</location>
submersible
<emphasis id="B95CDA13FFB4D020FCDFFEE77B309E34" box="[802,874,325,349]" italics="true" pageId="3" pageNumber="74">
<location id="8EF750DAFFB4D020FCDFFEE77B309E34" LSID="urn:lsid:plazi:treatment:0381B717FFB4D029FD04FF7279B09BEB:8EF750DAFFB4D020FCDFFEE77B309E34" box="[802,874,325,349]" country="Tonga" county="Hine Hina" latitude="-22.533333" longLatPrecision="1259" longitude="-176.71666" municipality="Newman" name="Nautile" pageId="3" pageNumber="74">Nautile</location>
</emphasis>
,
<location id="8EF750DAFFB4D020FC84FEE77C449E34" LSID="urn:lsid:plazi:treatment:0381B717FFB4D029FD04FF7279B09BEB:8EF750DAFFB4D020FC84FEE77C449E34" box="[889,1054,325,349]" country="Tonga" county="Hine Hina" latitude="-22.533333" longLatPrecision="1259" longitude="-176.71666" municipality="Newman" name="BIOLAU Cruise" pageId="3" pageNumber="74">BIOLAU Cruise</location>
of 1989 to the
<location id="8EF750DAFFB4D020FD58FEFC7B579E1E" LSID="urn:lsid:plazi:treatment:0381B717FFB4D029FD04FF7279B09BEB:8EF750DAFFB4D020FD58FEFC7B579E1E" box="[677,781,350,375]" country="Tonga" county="Hine Hina" latitude="-22.533333" longLatPrecision="1259" longitude="-176.71666" municipality="Newman" name="Lau Basin" pageId="3" pageNumber="74">Lau Basin</location>
,
<collectingCountry id="F33F4691FFB4D020FCE4FEFC7B009E1F" box="[793,858,350,374]" name="Tonga" pageId="3" pageNumber="74">Tonga</collectingCountry>
(
<collectorName id="26DD63D7FFB4D020FC96FEFD7C619E1E" box="[875,1083,351,375]" pageId="3" pageNumber="74">A.-M. Alayse-Danet</collectorName>
, chief scientist), and they were received from the Centre national de tri d’Océanographie biologique (
<collectionCode id="ED399EC4FFB4D020FBCBFE337CF79EC0" box="[1078,1197,401,425]" pageId="3" pageNumber="74">CENTOB</collectionCode>
), IFREMER, Brest.
<specimenCount id="9D2ECD88FFB4D020FC96FE087B809EAA" box="[875,986,426,451]" count="2" pageId="3" pageNumber="74" type="adult">Two adult</specimenCount>
specimens on round stones partially coated with sulfide deposits, plus several juveniles about to shed their cyprid shells, inferred to belong to this species (Jensen’s x-juveniles of
<bibRefCitation id="EFB97BF0FFB4D020FB7EFE557B0F9D41" author="HOeg J. T. &amp; Newman W. A." bookContentInfo="Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris" pageId="3" pageNumber="74" refId="ref6736" refString="HOeg J. T. &amp; Newman W. A. 1997. - Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga. Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris." title="Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga." type="proceedings" year="1997">HØeg &amp; Newman 1997</bibRefCitation>
; see below), all from station 1 (
<collectingCounty id="62F67E8DFFB4D020FB7BFDB27A809D28" pageId="3" pageNumber="74">Hine Hina</collectingCounty>
,
<geoCoordinate id="EE1C60C6FFB4D020FD18FD8B7B6F9D28" box="[741,821,553,577]" degrees="22" direction="south" minutes="32" orientation="latitude" pageId="3" pageNumber="74" precision="925" value="-22.533333">22°32’S</geoCoordinate>
,
<geoCoordinate id="EE1C60C6FFB4D020FCBDFD8B7BF39D28" box="[832,937,553,577]" degrees="176" direction="west" minutes="43" orientation="longitude" pageId="3" pageNumber="74" precision="925" value="-176.71666">176°43’W</geoCoordinate>
, at
<elevation id="0005E132FFB4D020FC30FD8B7C059D28" box="[973,1119,553,577]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="74" unit="m" value="1900.0">
<quantity id="4CD0ABE4FFB4D020FC30FD8B7C419D29" box="[973,1051,553,577]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="74" unit="m" value="1900.0">1900 m</quantity>
depth
</elevation>
). Two of these samples (
<specimenCode id="DB8EAE7AFFB4D020FCC5FDE17B229D32" box="[824,888,579,603]" pageId="3" pageNumber="74">BL-01</specimenCode>
&amp;
<specimenCode id="DB8EAE7AFFB4D020FC60FDE17B879D32" box="[925,989,579,603]" pageId="3" pageNumber="74">BL-03</specimenCode>
) contained other vent barnacles and other vent organisms (
<bibRefCitation id="EFB97BF0FFB4D020FBC4FDFE7B3B9DE4" author="Newman W. A. &amp; Yamaguchi T." journalOrPublisher="Bulletin du Museum national d'Histoire naturelle" pageId="3" pageNumber="74" pagination="211 - 243" part="17" refId="ref7319" refString="Newman W. A. &amp; Yamaguchi T. 1995. - A new sessile barnacle (Cirripedia, Brachylepadomorpha) from the Lau Back-Arc Basin, Tonga; first record of a living representative since the Miocene. Bulletin du Museum national d'Histoire naturelle, 4 e serie, 17 A (3 - 4): 211 - 243." title="A new sessile barnacle (Cirripedia, Brachylepadomorpha) from the Lau Back-Arc Basin, Tonga; first record of a living representative since the Miocene" type="journal article" year="1995">
<determinerName id="7E4BEAC5FFB4D020FBC4FDFE7B479DE4" pageId="3" pageNumber="74">Newman &amp; Yamaguchi</determinerName>
1995
</bibRefCitation>
;
<bibRefCitation id="EFB97BF0FFB4D020FC92FDD77CEF9DE4" author="Yamaguchi T. &amp; Newman W. A." box="[879,1205,629,653]" journalOrPublisher="Zoosystema" pageId="3" pageNumber="74" pagination="623 - 649" part="19" refId="ref7936" refString="Yamaguchi T. &amp; Newman W. A. 1997. - The hydrothermal vent barnacles Eochionelasmus (Cirripedia, Balanomorpha) from the North Fiji, Lau and Manus Basins, South-West Pacific. Zoosystema 19 (4): 623 - 649." title="The hydrothermal vent barnacles Eochionelasmus (Cirripedia, Balanomorpha) from the North Fiji, Lau and Manus Basins, South-West Pacific" type="journal article" year="1997">Yamaguchi &amp; Newman 1997</bibRefCitation>
;
<bibRefCitation id="EFB97BF0FFB4D020FD58FD2D7B8F9DCE" author="Southward A. J. &amp; Newman W. A." box="[677,981,654,679]" journalOrPublisher="Cahiers de Biologie marine" pageId="3" pageNumber="74" pagination="258 - 262" part="39" refId="ref7585" refString="Southward A. J. &amp; Newman W. A. 1998. - Ectosymbiosis between filamentous sulphur bacteria and a stalked barnacle (Scalpellomorpha, Neolepadinae) from the Lau Back Arc Basin, Tonga. Cahiers de Biologie marine 39 (3 - 4): 258 - 262." title="Ectosymbiosis between filamentous sulphur bacteria and a stalked barnacle (Scalpellomorpha, Neolepadinae) from the Lau Back Arc Basin, Tonga" type="journal article" year="1998">
<determinerName id="7E4BEAC5FFB4D020FD58FD2D7BCF9DCC" box="[677,917,654,679]" pageId="3" pageNumber="74">Southward &amp; Newman</determinerName>
1998
</bibRefCitation>
; cf.
<bibRefCitation id="EFB97BF0FFB4D020FBFEFD2D7A819DA9" author="Desbruyeres D. &amp; Alayse-Danet A. - M. &amp; Ohta S. &amp; the scientific parties of the Biolau and Starmer cruises" journalOrPublisher="Marine Geology" pageId="3" pageNumber="74" pagination="227 - 242" part="116" refId="ref6571" refString="Desbruyeres D., Alayse-Danet A. - M., Ohta S. &amp; the scientific parties of the Biolau and Starmer cruises 1994. - Deep-sea hydrothermal communities in southwestern Pacific back-arc basins (the North Fiji and Lau Basins): Composition, microdistribution and food web. Marine Geology 116: 227 - 242." title="Deep-sea hydrothermal communities in southwestern Pacific back-arc basins (the North Fiji and Lau Basins): Composition, microdistribution and food web" type="journal article" year="1994">
<determinerName id="7E4BEAC5FFB4D020FBFEFD2D7C259DCE" box="[1027,1151,655,679]" pageId="3" pageNumber="74">Desbruyères</determinerName>
<emphasis id="B95CDA13FFB4D020FB75FD2C7CE09DCE" box="[1160,1210,654,679]" italics="true" pageId="3" pageNumber="74">et al.</emphasis>
1994
</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C332558AFFB4D020FD58FD707C5E9C5F" pageId="3" pageNumber="74" type="etymology">
<paragraph id="8B970601FFB4D020FD58FD707C5E9C5F" blockId="3.[677,1210,299,865]" pageId="3" pageNumber="74">ETYMOLOGY. — Named for Toshiyuki Yamaguchi in appreciation of his numerous and exacting works on the systematics and ecology of the living and fossil cirripeds, especially the vent barnacles.</paragraph>
</subSubSection>
<subSubSection id="C332558AFFB4D020FD58FCEA7BBB9C09" box="[677,993,840,865]" pageId="3" pageNumber="74" type="diagnosis">
<paragraph id="8B970601FFB4D020FD58FCEA7BBB9C09" blockId="3.[677,1210,299,865]" box="[677,993,840,865]" pageId="3" pageNumber="74">DIAGNOSIS. — As for the genus.</paragraph>
</subSubSection>
<paragraph id="8B970601FFB4D020FF05FC0B7BD79B57" pageId="3" pageNumber="74">
<table id="F928F4A1FFB42FDCFF05FC0B7C799B57" box="[248,1059,937,1086]" gridcols="5" gridrows="4" pageId="3" pageNumber="74">
<tr id="35180443FFB42FDCFF05FC0B7C799CAA" box="[248,1059,937,963]" gridrow="0" pageId="3" pageNumber="74">
<th id="76C96D3FFFB42FDCFF05FC0B79039CAA" box="[248,345,937,963]" gridcol="0" gridrow="0" pageId="3" pageNumber="74">Sample</th>
<th id="76C96D3FFFB42FDCFE81FC0B79E39CAA" box="[380,441,937,963]" gridcol="1" gridrow="0" pageId="3" pageNumber="74">R-C</th>
<th id="76C96D3FFFB42FDCFE0AFC0B7A4E9CAA" box="[503,532,937,963]" gridcol="2" gridrow="0" pageId="3" pageNumber="74">W</th>
<th id="76C96D3FFFB42FDCFDA0FC0B7B689CAA" box="[605,818,937,963]" gridcol="3" gridrow="0" pageId="3" pageNumber="74">Type designation</th>
<th id="76C96D3FFFB42FDCFC96FC0B7C799CAA" box="[875,1059,937,963]" gridcol="4" gridrow="0" pageId="3" pageNumber="74">Depository</th>
</tr>
<tr id="35180443FFB42FDCFF05FC447C799B69" box="[248,1059,998,1024]" gridrow="1" pageId="3" pageNumber="74">
<th id="76C96D3FFFB42FDCFF05FC4479039B69" box="[248,345,998,1024]" gridcol="0" gridrow="1" pageId="3" pageNumber="74">1. BL-01</th>
<td id="76C96D3FFFB42FDCFE81FC4479E39B69" box="[380,441,998,1024]" gridcol="1" gridrow="1" pageId="3" pageNumber="74">~7</td>
<td id="76C96D3FFFB42FDCFE0AFC447A4E9B69" box="[503,532,998,1024]" gridcol="2" gridrow="1" pageId="3" pageNumber="74">~6</td>
<td id="76C96D3FFFB42FDCFDA0FC447B689B69" box="[605,818,998,1024]" gridcol="3" gridrow="1" pageId="3" pageNumber="74">holotype (intact)</td>
<td id="76C96D3FFFB42FDCFC96FC447C799B69" box="[875,1059,998,1024]" gridcol="4" gridrow="1" pageId="3" pageNumber="74">MNHN-Ci2710</td>
</tr>
<tr id="35180443FFB42FDCFF05FBA67C799B76" box="[248,1059,1028,1055]" gridrow="2" pageId="3" pageNumber="74">
<th id="76C96D3FFFB42FDCFF05FBA679039B76" box="[248,345,1028,1055]" gridcol="0" gridrow="2" pageId="3" pageNumber="74">2. BL-01</th>
<td id="76C96D3FFFB42FDCFE81FBA679E39B76" box="[380,441,1028,1055]" gridcol="1" gridrow="2" pageId="3" pageNumber="74">~7</td>
<td id="76C96D3FFFB42FDCFE0AFBA67A4E9B76" box="[503,532,1028,1055]" gridcol="2" gridrow="2" pageId="3" pageNumber="74">~6</td>
<td id="76C96D3FFFB42FDCFDA0FBA67B689B76" box="[605,818,1028,1055]" gridcol="3" gridrow="2" pageId="3" pageNumber="74">paratype (dissected)</td>
<td id="76C96D3FFFB42FDCFC96FBA67C799B76" box="[875,1059,1028,1055]" gridcol="4" gridrow="2" pageId="3" pageNumber="74">MNHN-Ci2711</td>
</tr>
<tr id="35180443FFB42FDCFF05FB817C799B57" box="[248,1059,1059,1086]" gridrow="3" pageId="3" pageNumber="74">
<th id="76C96D3FFFB42FDCFF05FB8179039B57" box="[248,345,1059,1086]" gridcol="0" gridrow="3" pageId="3" pageNumber="74">3. BL-03</th>
<td id="76C96D3FFFB42FDCFE81FB817C799B57" box="[380,1059,1059,1086]" colspan="4" colspanRight="3" gridcol="1" gridrow="3" pageId="3" pageNumber="74">Jensen’s x-juveniles, care of J. HØeg, Copenhagen</td>
</tr>
</table>
</paragraph>
<tableNote id="76CE078FFFB4D020FF05FBFB79889BEB" pageId="3" pageNumber="74" targetBox="[248,1059,937,1086]" targetPageId="3">
<paragraph id="8B970601FFB4D020FF05FBFB79B29B05" blockId="3.[248,488,1113,1154]" box="[248,488,1113,1132]" pageId="3" pageNumber="74">R-C rostro-carinal diameter</paragraph>
<paragraph id="8B970601FFB4D020FF05FBCD79889BEB" blockId="3.[248,488,1113,1154]" box="[248,466,1135,1154]" pageId="3" pageNumber="74">W width, both in mm.</paragraph>
</tableNote>
<subSubSection id="C332558AFFB4D029FF8CFB6779B09BEB" lastPageId="10" lastPageNumber="81" pageId="3" pageNumber="74" type="description">
<paragraph id="8B970601FFB4D020FF8CFB6779D49BB5" blockId="3.[113,647,1221,1707]" box="[113,398,1221,1247]" pageId="3" pageNumber="74">COMPARATIVEDESCRIPTION</paragraph>
<paragraph id="8B970601FFB4D020FF8CFB4178B89B95" blockId="3.[113,647,1221,1707]" box="[113,226,1251,1277]" pageId="3" pageNumber="74">
<emphasis id="B95CDA13FFB4D020FF8CFB4178B89B95" box="[113,226,1251,1277]" italics="true" pageId="3" pageNumber="74">Hard parts</emphasis>
</paragraph>
<paragraph id="8B970601FFB4D020FF8CFAA07B6E9978" blockId="3.[113,647,1221,1707]" lastBlockId="3.[677,1210,1220,1707]" pageId="3" pageNumber="74">
The low-conic profile of the new genus,
<taxonomicName id="4C287D82FFB4D020FF8CFA83797A9A52" authorityName="Newman" authorityYear="2000" box="[113,288,1313,1339]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB4D020FF8CFA83797A9A52" box="[113,288,1313,1339]" italics="true" pageId="3" pageNumber="74">Imbricaverruca</emphasis>
</taxonomicName>
, is similar to
<taxonomicName id="4C287D82FFB4D020FE25FA83799C9A30" authority="Newman &amp; Yamaguchi, 1995" authorityName="Newman &amp; Yamaguchi" authorityYear="1995" class="Maxillopoda" family="Neobrachylepadidae" genus="Neobrachylepas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB4D020FE25FA837ADC9A52" box="[472,646,1313,1339]" italics="true" pageId="3" pageNumber="74">Neobrachylepas</emphasis>
<bibRefCitation id="EFB97BF0FFB4D020FF8CFA9C799C9A30" author="Newman W. A. &amp; Yamaguchi T." box="[113,454,1342,1369]" journalOrPublisher="Bulletin du Museum national d'Histoire naturelle" pageId="3" pageNumber="74" pagination="211 - 243" part="17" refId="ref7319" refString="Newman W. A. &amp; Yamaguchi T. 1995. - A new sessile barnacle (Cirripedia, Brachylepadomorpha) from the Lau Back-Arc Basin, Tonga; first record of a living representative since the Miocene. Bulletin du Museum national d'Histoire naturelle, 4 e serie, 17 A (3 - 4): 211 - 243." title="A new sessile barnacle (Cirripedia, Brachylepadomorpha) from the Lau Back-Arc Basin, Tonga; first record of a living representative since the Miocene" type="journal article" year="1995">Newman &amp; Yamaguchi, 1995</bibRefCitation>
</taxonomicName>
, whose representatives were relatively young, and to the juveniles of
<taxonomicName id="4C287D82FFB4D020FF11FADE792E9AFC" baseAuthorityName="Newman &amp; Hessler" baseAuthorityYear="1989" box="[236,372,1404,1429]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB4D020FF11FADE792E9AFC" box="[236,372,1404,1429]" italics="true" pageId="3" pageNumber="74">Neoverruca</emphasis>
</taxonomicName>
and
<taxonomicName id="4C287D82FFB4D020FE37FADF79909ADC" authority="Yamaguchi &amp; Newman, 1990" authorityName="Yamaguchi &amp; Newman" authorityYear="1990" class="Maxillopoda" family="Chionelasmatidae" genus="Eochionelasmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB4D020FE37FADF7ADC9AFE" box="[458,646,1405,1431]" italics="true" pageId="3" pageNumber="74">Eochionelasmus</emphasis>
<bibRefCitation id="EFB97BF0FFB4D020FF8CFA3979909ADC" author="Yamaguchi T. &amp; Newman W. A." box="[113,458,1434,1461]" journalOrPublisher="Pacific Science" pageId="3" pageNumber="74" pagination="135 - 155" part="44" refId="ref7889" refString="Yamaguchi T. &amp; Newman W. A. 1990. - A new and primitive barnacle (Cirripedia; Balanomorpha) from the North Fiji Basin abyssal hydrothermal field, and its evolutionary implications. Pacific Science 44 (2): 135 - 155." title="A new and primitive barnacle (Cirripedia; Balanomorpha) from the North Fiji Basin abyssal hydrothermal field, and its evolutionary implications" type="journal article" year="1990">Yamaguchi &amp; Newman, 1990</bibRefCitation>
</taxonomicName>
. The operculum forms a flat lid, as in the
<taxonomicName id="4C287D82FFB4D020FE4FFA187A0A9ABD" authorityName="Newman, 1989" authorityYear="1989" box="[434,592,1466,1492]" class="Maxillopoda" family="Proverrucidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="family">Proverrucidae</taxonomicName>
and
<taxonomicName id="4C287D82FFB4D020FF8CFA7B78B39A9A" authorityName="Darwin" authorityYear="1854" box="[113,233,1497,1523]" class="Maxillopoda" family="Verrucidae" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="family">Verrucidae</taxonomicName>
, but the hinge line is curved rather than essentially straight as in
<taxonomicName id="4C287D82FFB4D020FE44FA557A749979" baseAuthorityName="Newman &amp; Hessler" baseAuthorityYear="1989" box="[441,558,1527,1552]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB4D020FE44FA557A749979" box="[441,558,1527,1552]" italics="true" pageId="3" pageNumber="74">Neoverruca</emphasis>
</taxonomicName>
, and in higher verrucomorphs (cf.
<figureCitation id="13131A84FFB4D020FE2BF9B47A619959" box="[470,571,1558,1584]" captionStart="FIG" captionStartId="2.[114,125,799,816]" captionTargetBox="[262,1053,208,766]" captionTargetId="figure-307@2.[248,1067,202,776]" captionTargetPageId="2" captionText="FIG. 1. — A, Imbricaverruca yamaguchii n. gen. and n. sp. (holotype, MNHN Ci2710); B, Verruca s.l., both viewed from above with the right scutum and tergum forming the operculum.Note that in the former the four plated wall appears to be covered largely by imbricating plates and the operculum includes a large median latus,characters that readily distinguish it from Neoverruca. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5391397" httpUri="https://zenodo.org/record/5391397/files/figure.png" pageId="3" pageNumber="74">Figs 1A</figureCitation>
;
<figureCitation id="13131A84FFB4D020FDACF9B47A209959" box="[593,634,1558,1584]" captionStart="FIG" captionStartId="4.[114,125,1384,1401]" captionTargetBox="[213,1067,193,1353]" captionTargetId="figure-91@4.[208,1091,186,1354]" captionTargetPageId="4" captionText="FIG. 2. — Plates of Imbricaverruca yamaguchii gen. et n. sp. (paratype, MNHN Ci2711); A, B, viewed from above and from the rostral end, respectively; A, note the operculum, comprising the plates of the right side (MS-L-MT), includes a large median latus; B, note the slightly open aperture to the mantle cavity between the occludent margins of the movable (MS &amp; MT) and the fixed (FS &amp; FT) scutum and tergum, respectively; C-M, variously disarticulated hard parts; C, interior view of wall (R-C-FS-FT) and operculum (MS-L- MT) with all of the imbricating plates, except the uppermost of the three principle lateral tiers of the right side (rl1-l1-cl1), stripped away [an approximation of the rostrum (R), which did not survive dissection, is indicated by the dashed line]; D, E, movable scutum (MS); external and internal views respectively; scutal adductor muscle, inserting from just below to up under the apical concavity in Fig. 2E, extends to its origin on the fixed scutum (FS, Fig. 2G; the relationship of MS to FS can be seen in Fig 2C); F, G, fixed scutum (FS); basal and internal views, respectively. Dashed line in G indicates origin of scutal adductor muscle but no scar is evident. The relationship of the two protuberances (best seen near the tergal margin in Fig. 2G) to the fixed tergum (FT, Fig. 2J, K), was not resolved; H, I, movable tergum (MT); external and internal views respectively (external corrosion in bleach and cleaning inadvertently obliterated growth line ornamentation in H); J, K, fixed tergum (FT); external and internal views respectively; L, M, carina from below and above respectively (the rostrum did not survive dissection and cleaning). Abbreviations: C, carina; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral plates forming three tiers each four plates high, respectively; FS, FT, fixed scutum and tergum; L, median latus; MS, MT, movable scutum and tergum; R, rostrum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391399" httpUri="https://zenodo.org/record/5391399/files/figure.png" pageId="3" pageNumber="74">2A</figureCitation>
). Therefore, in order for the operculum to open, there is likely some flexibility to the wall as well as some freedom in the arthrodial membrane forming the hinge, and it must be able to bend along the scuto-tergal articulation, an ability likely facilitated by the basal gap protected by the median latus (
<figureCitation id="13131A84FFB4D020FC82FAA07B849A75" box="[895,990,1282,1308]" captionStart="FIG" captionStartId="4.[114,125,1384,1401]" captionTargetBox="[213,1067,193,1353]" captionTargetId="figure-91@4.[208,1091,186,1354]" captionTargetPageId="4" captionText="FIG. 2. — Plates of Imbricaverruca yamaguchii gen. et n. sp. (paratype, MNHN Ci2711); A, B, viewed from above and from the rostral end, respectively; A, note the operculum, comprising the plates of the right side (MS-L-MT), includes a large median latus; B, note the slightly open aperture to the mantle cavity between the occludent margins of the movable (MS &amp; MT) and the fixed (FS &amp; FT) scutum and tergum, respectively; C-M, variously disarticulated hard parts; C, interior view of wall (R-C-FS-FT) and operculum (MS-L- MT) with all of the imbricating plates, except the uppermost of the three principle lateral tiers of the right side (rl1-l1-cl1), stripped away [an approximation of the rostrum (R), which did not survive dissection, is indicated by the dashed line]; D, E, movable scutum (MS); external and internal views respectively; scutal adductor muscle, inserting from just below to up under the apical concavity in Fig. 2E, extends to its origin on the fixed scutum (FS, Fig. 2G; the relationship of MS to FS can be seen in Fig 2C); F, G, fixed scutum (FS); basal and internal views, respectively. Dashed line in G indicates origin of scutal adductor muscle but no scar is evident. The relationship of the two protuberances (best seen near the tergal margin in Fig. 2G) to the fixed tergum (FT, Fig. 2J, K), was not resolved; H, I, movable tergum (MT); external and internal views respectively (external corrosion in bleach and cleaning inadvertently obliterated growth line ornamentation in H); J, K, fixed tergum (FT); external and internal views respectively; L, M, carina from below and above respectively (the rostrum did not survive dissection and cleaning). Abbreviations: C, carina; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral plates forming three tiers each four plates high, respectively; FS, FT, fixed scutum and tergum; L, median latus; MS, MT, movable scutum and tergum; R, rostrum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391399" httpUri="https://zenodo.org/record/5391399/files/figure.png" pageId="3" pageNumber="74">Fig. 2C</figureCitation>
). Like
<taxonomicName id="4C287D82FFB4D020FBCBFAA37CEB9A73" baseAuthorityName="Newman &amp; Hessler" baseAuthorityYear="1989" box="[1078,1201,1281,1306]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB4D020FBCBFAA37CEB9A73" box="[1078,1201,1281,1306]" italics="true" pageId="3" pageNumber="74">Neoverruca</emphasis>
</taxonomicName>
, the scutal adductor muscle inserts in a pocket below the apex of the movable plate (
<figureCitation id="13131A84FFB4D020FBABFA9D7CF79A30" box="[1110,1197,1343,1369]" captionStart="FIG" captionStartId="4.[114,125,1384,1401]" captionTargetBox="[213,1067,193,1353]" captionTargetId="figure-91@4.[208,1091,186,1354]" captionTargetPageId="4" captionText="FIG. 2. — Plates of Imbricaverruca yamaguchii gen. et n. sp. (paratype, MNHN Ci2711); A, B, viewed from above and from the rostral end, respectively; A, note the operculum, comprising the plates of the right side (MS-L-MT), includes a large median latus; B, note the slightly open aperture to the mantle cavity between the occludent margins of the movable (MS &amp; MT) and the fixed (FS &amp; FT) scutum and tergum, respectively; C-M, variously disarticulated hard parts; C, interior view of wall (R-C-FS-FT) and operculum (MS-L- MT) with all of the imbricating plates, except the uppermost of the three principle lateral tiers of the right side (rl1-l1-cl1), stripped away [an approximation of the rostrum (R), which did not survive dissection, is indicated by the dashed line]; D, E, movable scutum (MS); external and internal views respectively; scutal adductor muscle, inserting from just below to up under the apical concavity in Fig. 2E, extends to its origin on the fixed scutum (FS, Fig. 2G; the relationship of MS to FS can be seen in Fig 2C); F, G, fixed scutum (FS); basal and internal views, respectively. Dashed line in G indicates origin of scutal adductor muscle but no scar is evident. The relationship of the two protuberances (best seen near the tergal margin in Fig. 2G) to the fixed tergum (FT, Fig. 2J, K), was not resolved; H, I, movable tergum (MT); external and internal views respectively (external corrosion in bleach and cleaning inadvertently obliterated growth line ornamentation in H); J, K, fixed tergum (FT); external and internal views respectively; L, M, carina from below and above respectively (the rostrum did not survive dissection and cleaning). Abbreviations: C, carina; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral plates forming three tiers each four plates high, respectively; FS, FT, fixed scutum and tergum; L, median latus; MS, MT, movable scutum and tergum; R, rostrum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391399" httpUri="https://zenodo.org/record/5391399/files/figure.png" pageId="3" pageNumber="74">Fig. 2E</figureCitation>
), and its origin on the fixed scutum, near the centre of the plate (
<figureCitation id="13131A84FFB4D020FC7AFADF7BBE9AFE" box="[903,996,1405,1431]" captionStart="FIG" captionStartId="4.[114,125,1384,1401]" captionTargetBox="[213,1067,193,1353]" captionTargetId="figure-91@4.[208,1091,186,1354]" captionTargetPageId="4" captionText="FIG. 2. — Plates of Imbricaverruca yamaguchii gen. et n. sp. (paratype, MNHN Ci2711); A, B, viewed from above and from the rostral end, respectively; A, note the operculum, comprising the plates of the right side (MS-L-MT), includes a large median latus; B, note the slightly open aperture to the mantle cavity between the occludent margins of the movable (MS &amp; MT) and the fixed (FS &amp; FT) scutum and tergum, respectively; C-M, variously disarticulated hard parts; C, interior view of wall (R-C-FS-FT) and operculum (MS-L- MT) with all of the imbricating plates, except the uppermost of the three principle lateral tiers of the right side (rl1-l1-cl1), stripped away [an approximation of the rostrum (R), which did not survive dissection, is indicated by the dashed line]; D, E, movable scutum (MS); external and internal views respectively; scutal adductor muscle, inserting from just below to up under the apical concavity in Fig. 2E, extends to its origin on the fixed scutum (FS, Fig. 2G; the relationship of MS to FS can be seen in Fig 2C); F, G, fixed scutum (FS); basal and internal views, respectively. Dashed line in G indicates origin of scutal adductor muscle but no scar is evident. The relationship of the two protuberances (best seen near the tergal margin in Fig. 2G) to the fixed tergum (FT, Fig. 2J, K), was not resolved; H, I, movable tergum (MT); external and internal views respectively (external corrosion in bleach and cleaning inadvertently obliterated growth line ornamentation in H); J, K, fixed tergum (FT); external and internal views respectively; L, M, carina from below and above respectively (the rostrum did not survive dissection and cleaning). Abbreviations: C, carina; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral plates forming three tiers each four plates high, respectively; FS, FT, fixed scutum and tergum; L, median latus; MS, MT, movable scutum and tergum; R, rostrum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391399" httpUri="https://zenodo.org/record/5391399/files/figure.png" pageId="3" pageNumber="74">Figs 2G</figureCitation>
;
<figureCitation id="13131A84FFB4D020FC09FADF7C429AFE" box="[1012,1048,1405,1431]" captionStart="Fig" captionStartId="7.[113,141,1469,1486]" captionTargetBox="[112,1165,215,1445]" captionTargetId="figure-22@7.[139,1170,219,1510]" captionTargetPageId="7" captionText="Fig. 4. — Schematic plan views and transverse sections of grades of skeletal organization from the most primitive sessile barnacles (A, Brachylepadomorpha) through the asymmetrical sessile barnacles (B-H, Verrucomorpha). Of the eight verrucid genera currently recognized (Young 1998) only the four most representative ones (Altiverruca, Newmaniverruca, Metaverruca and Verruca s.s.) are depicted here. Right sides are to the viewers right and rostral ends are downward in plan views or coming out of the page in transverse sections. The transverse sections are slightly rostral of the rostro-carinal gap or suture. While there are numerous whorls of imbricating plates standing in tiers four plates high in brachylepadomorphs, and on the movable side of neoverrucids, only those of the R-C gap are labeled. Abbreviations: C, R, carina and rostrum; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral tiers of imbricating plates respectively; cl, rl, the pair of imbricating plates in proverrucids; FS, FT, fixed scutum and tergum; hl, hinge line; L, median latus (completely lost in proverrucids and verrucids); m, myophore; MS, MT, movable scutum and tergum; sam, scutal adductor muscle; S, T, normal terga and scuta in brachylepadomorphs; dashed horizontal lines: membranous basis (transverse sections F-H after Young 1998)." figureDoi="http://doi.org/10.5281/zenodo.13988759" httpUri="https://zenodo.org/record/13988759/files/figure.png" pageId="3" pageNumber="74">4C</figureCitation>
), is supported by an adductor ridge or “myophore” as in
<taxonomicName id="4C287D82FFB4D020FD58FA1B7C5E9ABD" authority="Pilsbry, 1916" authorityName="Pilsbry" authorityYear="1916" box="[677,1028,1465,1492]" class="Maxillopoda" family="Verrucidae" genus="Metaverruca" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB4D020FD58FA1B7B679ABB" box="[677,829,1465,1490]" italics="true" pageId="3" pageNumber="74">Metaverruca</emphasis>
<bibRefCitation id="EFB97BF0FFB4D020FCAFFA187C5E9ABD" author="Pilsbry H. A." box="[850,1028,1466,1492]" journalOrPublisher="Bulletin of the United States National Museum" pageId="3" pageNumber="74" pagination="1 - 366" part="93" refId="ref7445" refString="Pilsbry H. A. 1916. - The sessile barnacles (Cirripedia) contained in the collections of the U. S. National Museum; including a monograph of the American species. Bulletin of the United States National Museum 93: 1 - 366." title="The sessile barnacles (Cirripedia) contained in the collections of the U. S. National Museum; including a monograph of the American species" type="journal article" year="1916">Pilsbry, 1916</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C287D82FFB4D020FBA0FA1B7B2A9A9A" authority="Schumacher, 1817" authorityName="Schumacher" authorityYear="1817" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB4D020FBA0FA1B7CE39ABB" box="[1117,1209,1465,1490]" italics="true" pageId="3" pageNumber="74">Verruca</emphasis>
Schumacher, 1817
</taxonomicName>
(cf.
<bibRefCitation id="EFB97BF0FFB4D020FC58FA7A7C719A9A" author="Young P. S." box="[933,1067,1496,1523]" journalOrPublisher="Zoosystema" pageId="3" pageNumber="74" pagination="31 - 92" part="20" refId="ref7981" refString="Young P. S. 1998. - Cirripedia (Crustacea) from the &quot; Campagne Biacores &quot; in the Azores region, including a generic revision of Verrucidae. Zoosystema 20 (1): 31 - 92." title="Cirripedia (Crustacea) from the &quot; Campagne Biacores &quot; in the Azores region, including a generic revision of Verrucidae" type="journal article" year="1998">Young 1998</bibRefCitation>
for revision) (
<figureCitation id="13131A84FFB4D020FD51FA557AAA9978" box="[684,752,1527,1553]" captionStart="Fig" captionStartId="7.[113,141,1469,1486]" captionTargetBox="[112,1165,215,1445]" captionTargetId="figure-22@7.[139,1170,219,1510]" captionTargetPageId="7" captionText="Fig. 4. — Schematic plan views and transverse sections of grades of skeletal organization from the most primitive sessile barnacles (A, Brachylepadomorpha) through the asymmetrical sessile barnacles (B-H, Verrucomorpha). Of the eight verrucid genera currently recognized (Young 1998) only the four most representative ones (Altiverruca, Newmaniverruca, Metaverruca and Verruca s.s.) are depicted here. Right sides are to the viewers right and rostral ends are downward in plan views or coming out of the page in transverse sections. The transverse sections are slightly rostral of the rostro-carinal gap or suture. While there are numerous whorls of imbricating plates standing in tiers four plates high in brachylepadomorphs, and on the movable side of neoverrucids, only those of the R-C gap are labeled. Abbreviations: C, R, carina and rostrum; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral tiers of imbricating plates respectively; cl, rl, the pair of imbricating plates in proverrucids; FS, FT, fixed scutum and tergum; hl, hinge line; L, median latus (completely lost in proverrucids and verrucids); m, myophore; MS, MT, movable scutum and tergum; sam, scutal adductor muscle; S, T, normal terga and scuta in brachylepadomorphs; dashed horizontal lines: membranous basis (transverse sections F-H after Young 1998)." figureDoi="http://doi.org/10.5281/zenodo.13988759" httpUri="https://zenodo.org/record/13988759/files/figure.png" pageId="3" pageNumber="74">Fig. 4</figureCitation>
G-H).
</paragraph>
<paragraph id="8B970601FFB4D025FD58F9B479E59D74" blockId="3.[677,1210,1220,1707]" lastBlockId="6.[114,647,209,541]" lastPageId="6" lastPageNumber="77" pageId="3" pageNumber="74">
The fixed scutum and tergum are much reduced in height (
<figureCitation id="13131A84FFB4D020FCE8F9977B0E9926" box="[789,852,1589,1615]" captionStart="FIG" captionStartId="4.[114,125,1384,1401]" captionTargetBox="[213,1067,193,1353]" captionTargetId="figure-91@4.[208,1091,186,1354]" captionTargetPageId="4" captionText="FIG. 2. — Plates of Imbricaverruca yamaguchii gen. et n. sp. (paratype, MNHN Ci2711); A, B, viewed from above and from the rostral end, respectively; A, note the operculum, comprising the plates of the right side (MS-L-MT), includes a large median latus; B, note the slightly open aperture to the mantle cavity between the occludent margins of the movable (MS &amp; MT) and the fixed (FS &amp; FT) scutum and tergum, respectively; C-M, variously disarticulated hard parts; C, interior view of wall (R-C-FS-FT) and operculum (MS-L- MT) with all of the imbricating plates, except the uppermost of the three principle lateral tiers of the right side (rl1-l1-cl1), stripped away [an approximation of the rostrum (R), which did not survive dissection, is indicated by the dashed line]; D, E, movable scutum (MS); external and internal views respectively; scutal adductor muscle, inserting from just below to up under the apical concavity in Fig. 2E, extends to its origin on the fixed scutum (FS, Fig. 2G; the relationship of MS to FS can be seen in Fig 2C); F, G, fixed scutum (FS); basal and internal views, respectively. Dashed line in G indicates origin of scutal adductor muscle but no scar is evident. The relationship of the two protuberances (best seen near the tergal margin in Fig. 2G) to the fixed tergum (FT, Fig. 2J, K), was not resolved; H, I, movable tergum (MT); external and internal views respectively (external corrosion in bleach and cleaning inadvertently obliterated growth line ornamentation in H); J, K, fixed tergum (FT); external and internal views respectively; L, M, carina from below and above respectively (the rostrum did not survive dissection and cleaning). Abbreviations: C, carina; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral plates forming three tiers each four plates high, respectively; FS, FT, fixed scutum and tergum; L, median latus; MS, MT, movable scutum and tergum; R, rostrum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391399" httpUri="https://zenodo.org/record/5391399/files/figure.png" pageId="3" pageNumber="74">Fig. 2</figureCitation>
F-G, J-K) and there is a complex articulation between them. Together with the rostrum and carina, which are also much wider than high, they form a ring-like wall supporting the operculum. Therefore, the imbricating plates between this primary wall and the substratum form a substantially greater proportion of the body chamber than in
<taxonomicName id="4C287D82FFB1D025FE90FE8E79B89E2C" baseAuthorityName="Newman &amp; Hessler" baseAuthorityYear="1989" box="[365,482,300,325]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB1D025FE90FE8E79B89E2C" box="[365,482,300,325]" italics="true" pageId="6" pageNumber="77">Neoverruca</emphasis>
</taxonomicName>
. The principal three tiers of lateral imbricating plates, standing four plates high (rl
<superScript id="7C5DAB49FFB1D025FEC4FEC7790F9E11" attach="left" box="[313,341,357,376]" fontSize="8" pageId="6" pageNumber="77">1-4</superScript>
, l
<superScript id="7C5DAB49FFB1D025FE94FEC779DF9E11" attach="left" box="[361,389,357,376]" fontSize="8" pageId="6" pageNumber="77">1-4</superScript>
and cl
<superScript id="7C5DAB49FFB1D025FE31FEC779B29E11" attach="left" box="[460,488,357,376]" fontSize="8" pageId="6" pageNumber="77">1-4</superScript>
), are fully represented in these two apparently adult
<taxonomicName id="4C287D82FFB1D025FDC0FE2B78B99EA9" authorityName="Newman" authorityYear="2000" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB1D025FDC0FE2B78B99EA9" italics="true" pageId="6" pageNumber="77">Imbricaverruca</emphasis>
</taxonomicName>
(
<figureCitation id="13131A84FFB1D025FF0BFE0579039EA8" box="[246,345,423,449]" captionStart="FIG" captionStartId="2.[114,125,799,816]" captionTargetBox="[262,1053,208,766]" captionTargetId="figure-307@2.[248,1067,202,776]" captionTargetPageId="2" captionText="FIG. 1. — A, Imbricaverruca yamaguchii n. gen. and n. sp. (holotype, MNHN Ci2710); B, Verruca s.l., both viewed from above with the right scutum and tergum forming the operculum.Note that in the former the four plated wall appears to be covered largely by imbricating plates and the operculum includes a large median latus,characters that readily distinguish it from Neoverruca. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5391397" httpUri="https://zenodo.org/record/5391397/files/figure.png" pageId="6" pageNumber="77">Figs 1A</figureCitation>
;
<figureCitation id="13131A84FFB1D025FE94FE0579D59EA8" box="[361,399,423,449]" captionStart="FIG" captionStartId="4.[114,125,1384,1401]" captionTargetBox="[213,1067,193,1353]" captionTargetId="figure-91@4.[208,1091,186,1354]" captionTargetPageId="4" captionText="FIG. 2. — Plates of Imbricaverruca yamaguchii gen. et n. sp. (paratype, MNHN Ci2711); A, B, viewed from above and from the rostral end, respectively; A, note the operculum, comprising the plates of the right side (MS-L-MT), includes a large median latus; B, note the slightly open aperture to the mantle cavity between the occludent margins of the movable (MS &amp; MT) and the fixed (FS &amp; FT) scutum and tergum, respectively; C-M, variously disarticulated hard parts; C, interior view of wall (R-C-FS-FT) and operculum (MS-L- MT) with all of the imbricating plates, except the uppermost of the three principle lateral tiers of the right side (rl1-l1-cl1), stripped away [an approximation of the rostrum (R), which did not survive dissection, is indicated by the dashed line]; D, E, movable scutum (MS); external and internal views respectively; scutal adductor muscle, inserting from just below to up under the apical concavity in Fig. 2E, extends to its origin on the fixed scutum (FS, Fig. 2G; the relationship of MS to FS can be seen in Fig 2C); F, G, fixed scutum (FS); basal and internal views, respectively. Dashed line in G indicates origin of scutal adductor muscle but no scar is evident. The relationship of the two protuberances (best seen near the tergal margin in Fig. 2G) to the fixed tergum (FT, Fig. 2J, K), was not resolved; H, I, movable tergum (MT); external and internal views respectively (external corrosion in bleach and cleaning inadvertently obliterated growth line ornamentation in H); J, K, fixed tergum (FT); external and internal views respectively; L, M, carina from below and above respectively (the rostrum did not survive dissection and cleaning). Abbreviations: C, carina; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral plates forming three tiers each four plates high, respectively; FS, FT, fixed scutum and tergum; L, median latus; MS, MT, movable scutum and tergum; R, rostrum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391399" httpUri="https://zenodo.org/record/5391399/files/figure.png" pageId="6" pageNumber="77">2A</figureCitation>
;
<figureCitation id="13131A84FFB1D025FE63FE05799D9EA8" box="[414,455,423,449]" captionStart="Fig" captionStartId="7.[113,141,1469,1486]" captionTargetBox="[112,1165,215,1445]" captionTargetId="figure-22@7.[139,1170,219,1510]" captionTargetPageId="7" captionText="Fig. 4. — Schematic plan views and transverse sections of grades of skeletal organization from the most primitive sessile barnacles (A, Brachylepadomorpha) through the asymmetrical sessile barnacles (B-H, Verrucomorpha). Of the eight verrucid genera currently recognized (Young 1998) only the four most representative ones (Altiverruca, Newmaniverruca, Metaverruca and Verruca s.s.) are depicted here. Right sides are to the viewers right and rostral ends are downward in plan views or coming out of the page in transverse sections. The transverse sections are slightly rostral of the rostro-carinal gap or suture. While there are numerous whorls of imbricating plates standing in tiers four plates high in brachylepadomorphs, and on the movable side of neoverrucids, only those of the R-C gap are labeled. Abbreviations: C, R, carina and rostrum; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral tiers of imbricating plates respectively; cl, rl, the pair of imbricating plates in proverrucids; FS, FT, fixed scutum and tergum; hl, hinge line; L, median latus (completely lost in proverrucids and verrucids); m, myophore; MS, MT, movable scutum and tergum; sam, scutal adductor muscle; S, T, normal terga and scuta in brachylepadomorphs; dashed horizontal lines: membranous basis (transverse sections F-H after Young 1998)." figureDoi="http://doi.org/10.5281/zenodo.13988759" httpUri="https://zenodo.org/record/13988759/files/figure.png" pageId="6" pageNumber="77">4C</figureCitation>
), as they are in brachylepadomorphs (
<bibRefCitation id="EFB97BF0FFB1D025FE8CFE647A7B9E89" author="Newman W. A." box="[369,545,454,480]" editor="Southward A. J." pageId="6" pageNumber="77" pagination="3 - 42" refId="ref7098" refString="Newman W. A. 1987. - Evolution of Cirripedes and their major groups: 3 - 42, in Southward A. J. (ed.), Barnacle Biology, Crustacean Issues 5. A. A. Balkema, Rotterdam." title="Evolution of Cirripedes and their major groups" type="book chapter" volumeTitle="Barnacle Biology, Crustacean Issues 5. A. A. Balkema, Rotterdam" year="1987">Newman 1987</bibRefCitation>
: fig. 4A) and in the juveniles of
<taxonomicName id="4C287D82FFB1D025FE85FE4679AA9E94" baseAuthorityName="Newman &amp; Hessler" baseAuthorityYear="1989" box="[376,496,484,509]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB1D025FE85FE4679AA9E94" box="[376,496,484,509]" italics="true" pageId="6" pageNumber="77">Neoverruca</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB97BF0FFB1D025FDFDFE4779589D74" author="Newman W. A. &amp; Hessler R. R." journalOrPublisher="San Diego Society of Natural History Transactions" pageId="6" pageNumber="77" pagination="259 - 273" part="21" refId="ref7233" refString="Newman W. A. &amp; Hessler R. R. 1989. - A new abyssal hydrothermal verrucomorphan (Cirripedia; Sessilia): The most primitive living sessile barnacle. San Diego Society of Natural History Transactions 21 (16): 259 - 273." title="A new abyssal hydrothermal verrucomorphan (Cirripedia; Sessilia): The most primitive living sessile barnacle" type="journal article" year="1989">Newman &amp; Hessler 1989</bibRefCitation>
;
<bibRefCitation id="EFB97BF0FFB1D025FEF0FDA079E99D74" author="Newman W. A." box="[269,435,514,541]" journalOrPublisher="Bulletin of Marine Science" pageId="6" pageNumber="77" pagination="467 - 477" part="45" refId="ref7144" refString="Newman W. A. 1989. - Juvenile ontogeny and metamorphosis in the most primitive living sessile barnacle, Neoverruca, from an abyssal hydrothermal spring. Bulletin of Marine Science 45 (2): 467 - 477." title="Juvenile ontogeny and metamorphosis in the most primitive living sessile barnacle, Neoverruca, from an abyssal hydrothermal spring" type="journal article" year="1989">Newman 1989</bibRefCitation>
).
</paragraph>
<caption id="DF575689FFB3D027FF8FFACA7B1899CD" ID-DOI="http://doi.org/10.5281/zenodo.5391399" ID-Zenodo-Dep="5391399" httpUri="https://zenodo.org/record/5391399/files/figure.png" pageId="4" pageNumber="75" startId="4.[114,125,1384,1401]" targetBox="[213,1067,193,1353]" targetPageId="4">
<paragraph id="8B970601FFB3D027FF8FFACA7B1899CD" blockId="4.[114,1211,1384,1700]" pageId="4" pageNumber="75">
FIG. 2. — Plates of
<taxonomicName id="4C287D82FFB3D027FEF3FACA79B69A10" authorityName="Newman" authorityYear="2000" box="[270,492,1384,1401]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="4" pageNumber="75" phylum="Arthropoda" rank="species" species="yamaguchii">
<emphasis id="B95CDA13FFB3D027FEF3FACA79B69A10" box="[270,492,1384,1401]" italics="true" pageId="4" pageNumber="75">Imbricaverruca yamaguchii</emphasis>
</taxonomicName>
gen. et n. sp. (paratype, MNHN Ci2711); A, B, viewed from above and from the rostral end, respectively;
<emphasis id="B95CDA13FFB3D027FEF6FADF79429AE7" bold="true" box="[267,280,1405,1422]" pageId="4" pageNumber="75">A</emphasis>
, note the operculum, comprising the plates of the right side (MS-L-MT), includes a large median latus;
<emphasis id="B95CDA13FFB3D027FB81FADF7CD39AE7" bold="true" box="[1148,1161,1405,1422]" pageId="4" pageNumber="75">B</emphasis>
, note the slightly open aperture to the mantle cavity between the occludent margins of the movable (MS &amp; MT) and the fixed (FS &amp; FT) scutum and tergum, respectively; C-M, variously disarticulated hard parts;
<emphasis id="B95CDA13FFB3D027FD31FA0A7A809AD0" bold="true" box="[716,730,1448,1465]" pageId="4" pageNumber="75">C</emphasis>
, interior view of wall (R-C-FS-FT) and operculum (MS-L-MT) with all of the imbricating plates, except the uppermost of the three principle lateral tiers of the right side (rl
<superScript id="7C5DAB49FFB3D027FBD1FA187C6E9AAE" attach="left" box="[1068,1076,1466,1479]" fontSize="5" pageId="4" pageNumber="75">1</superScript>
-l
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-cl
<superScript id="7C5DAB49FFB3D027FBA1FA187C3E9AAE" attach="left" box="[1116,1124,1466,1479]" fontSize="5" pageId="4" pageNumber="75">1</superScript>
), stripped away [an approximation of the rostrum (R), which did not survive dissection, is indicated by the dashed line];
<emphasis id="B95CDA13FFB3D027FBFDFA717C549A8D" bold="true" box="[1024,1038,1491,1508]" pageId="4" pageNumber="75">D</emphasis>
,
<emphasis id="B95CDA13FFB3D027FBE5FA717C7E9A8D" bold="true" box="[1048,1060,1491,1508]" pageId="4" pageNumber="75">E</emphasis>
, movable scutum (MS); external and internal views respectively; scutal adductor muscle, inserting from just below to up under the apical concavity in Fig. 2E, extends to its origin on the fixed scutum (FS, Fig. 2G; the relationship of MS to FS can be seen in Fig 2C);
<emphasis id="B95CDA13FFB3D027FBDCFA5F7C769967" bold="true" box="[1057,1068,1533,1550]" pageId="4" pageNumber="75">F</emphasis>
,
<emphasis id="B95CDA13FFB3D027FBCBFA5F7C1E9967" bold="true" box="[1078,1092,1533,1550]" pageId="4" pageNumber="75">G</emphasis>
, fixed scutum (FS); basal and internal views, respectively. Dashed line in G indicates origin of scutal adductor muscle but no scar is evident. The relationship of the two protuberances (best seen near the tergal margin in Fig. 2G) to the fixed tergum (FT, Fig. 2J, K), was not resolved;
<emphasis id="B95CDA13FFB3D027FF3EF99F788B9927" bold="true" box="[195,209,1597,1614]" pageId="4" pageNumber="75">H</emphasis>
,
<emphasis id="B95CDA13FFB3D027FF26F99F78BB9927" bold="true" box="[219,225,1597,1614]" pageId="4" pageNumber="75">I</emphasis>
, movable tergum (MT); external and internal views respectively (external corrosion in bleach and cleaning inadvertently obliterated growth line ornamentation in H);
<emphasis id="B95CDA13FFB3D027FE02F9F17A53990D" bold="true" box="[511,521,1619,1636]" pageId="4" pageNumber="75">J</emphasis>
,
<emphasis id="B95CDA13FFB3D027FDE8F9F17A78990D" bold="true" box="[533,546,1619,1636]" pageId="4" pageNumber="75">K</emphasis>
, fixed tergum (FT); external and internal views respectively;
<emphasis id="B95CDA13FFB3D027FBD8F9F17C6A990D" bold="true" box="[1061,1072,1619,1636]" pageId="4" pageNumber="75">L</emphasis>
,
<emphasis id="B95CDA13FFB3D027FBC1F9F17C17990D" bold="true" box="[1084,1101,1619,1636]" pageId="4" pageNumber="75">M</emphasis>
, carina from below and above respectively (the rostrum did not survive dissection and cleaning). Abbreviations:
<emphasis id="B95CDA13FFB3D027FC57F9CA7BE29910" bold="true" box="[938,952,1640,1657]" pageId="4" pageNumber="75">C</emphasis>
, carina;
<emphasis id="B95CDA13FFB3D027FC02F9CA7C7E991B" bold="true" box="[1023,1060,1637,1657]" pageId="4" pageNumber="75">
cl
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</emphasis>
,
<emphasis id="B95CDA13FFB3D027FBD2F9CA7C13991B" bold="true" box="[1071,1097,1637,1657]" pageId="4" pageNumber="75">
l
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</emphasis>
,
<emphasis id="B95CDA13FFB3D027FBAEF9CA7C2F991B" bold="true" box="[1107,1141,1637,1657]" pageId="4" pageNumber="75">
rl
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</emphasis>
, carinolateral, lateral and rostrolateral plates forming three tiers each four plates high, respectively;
<emphasis id="B95CDA13FFB3D027FC68F9DF7BF699E7" bold="true" box="[917,940,1661,1678]" pageId="4" pageNumber="75">FS</emphasis>
,
<emphasis id="B95CDA13FFB3D027FC45F9DF7B9599E7" bold="true" box="[952,975,1661,1678]" pageId="4" pageNumber="75">FT</emphasis>
, fixed scutum and tergum;
<emphasis id="B95CDA13FFB3D027FF8FF931782799CD" bold="true" box="[114,125,1683,1700]" pageId="4" pageNumber="75">L</emphasis>
, median latus;
<emphasis id="B95CDA13FFB3D027FF03F931794199CD" bold="true" box="[254,283,1683,1700]" pageId="4" pageNumber="75">MS</emphasis>
,
<emphasis id="B95CDA13FFB3D027FEDBF931791899CD" bold="true" box="[294,322,1683,1700]" pageId="4" pageNumber="75">MT</emphasis>
, movable scutum and tergum;
<emphasis id="B95CDA13FFB3D027FDBBF9317A0999CD" bold="true" box="[582,595,1683,1700]" pageId="4" pageNumber="75">R</emphasis>
, rostrum. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="DF575689FFB2D026FF8CF98A7B2A99CD" ID-DOI="http://doi.org/10.5281/zenodo.5391401" ID-Zenodo-Dep="5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="5" pageNumber="76" startId="5.[113,124,1576,1593]" targetBox="[92,1284,200,1546]" targetPageId="5">
<paragraph id="8B970601FFB2D026FF8CF98A7B2A99CD" blockId="5.[113,1210,1576,1700]" pageId="5" pageNumber="76">
FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of
<taxonomicName id="4C287D82FFB2D026FF8CF99F790A9927" authority="Newman, 2000" authorityName="Newman" authorityYear="2000" box="[113,336,1597,1614]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="76" phylum="Arthropoda" rank="species" species="yamaguchii" status="n. gen. and n. sp.">
<emphasis id="B95CDA13FFB2D026FF8CF99F790A9927" box="[113,336,1597,1614]" italics="true" pageId="5" pageNumber="76">Imbricaverruca yamaguchii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F6768FFB2D026FEA8F99F79BC9927" box="[341,486,1597,1614]" pageId="5" pageNumber="76" rank="species">n. gen. and n. sp.</taxonomicNameLabel>
(paratype, MNHN Ci2711);
<emphasis id="B95CDA13FFB2D026FD34F99F7A8C9927" bold="true" box="[713,726,1597,1614]" pageId="5" pageNumber="76">A</emphasis>
, cirrus I, anterior ramus missing;
<emphasis id="B95CDA13FFB2D026FC17F99F7BAD9927" bold="true" box="[1002,1015,1597,1614]" pageId="5" pageNumber="76">B</emphasis>
, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right);
<emphasis id="B95CDA13FFB2D026FCADF9F17B04990D" bold="true" box="[848,862,1619,1636]" pageId="5" pageNumber="76">C</emphasis>
, cirrus VI (setae of distal half omitted) and penis (annulations omitted);
<emphasis id="B95CDA13FFB2D026FEA6F9CA79339910" bold="true" box="[347,361,1640,1657]" pageId="5" pageNumber="76">D</emphasis>
, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end);
<emphasis id="B95CDA13FFB2D026FB92F9CA7C219910" bold="true" box="[1135,1147,1640,1657]" pageId="5" pageNumber="76">E</emphasis>
, end of penis enlarged;
<emphasis id="B95CDA13FFB2D026FF05F9DF795999E7" bold="true" box="[248,259,1661,1678]" pageId="5" pageNumber="76">F</emphasis>
, labrum and mandibular palps;
<emphasis id="B95CDA13FFB2D026FDEDF9DF7A6299E7" bold="true" box="[528,568,1661,1678]" pageId="5" pageNumber="76">G, H</emphasis>
, right and left mandible;
<emphasis id="B95CDA13FFB2D026FCF6F9DF7B4B99E7" bold="true" box="[779,785,1661,1678]" pageId="5" pageNumber="76">I</emphasis>
, spinous margin of H enlarged;
<emphasis id="B95CDA13FFB2D026FBDCF9DF7C1999E7" bold="true" box="[1057,1091,1661,1678]" pageId="5" pageNumber="76">J, K</emphasis>
, right and left first maxillae;
<emphasis id="B95CDA13FFB2D026FF19F93178B599CD" bold="true" box="[228,239,1683,1700]" pageId="5" pageNumber="76">L</emphasis>
, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B970601FFB1D025FF8FFD9D78889D30" blockId="6.[114,647,575,1247]" box="[114,210,575,601]" pageId="6" pageNumber="77">
<emphasis id="B95CDA13FFB1D025FF8FFD9D78889D30" box="[114,210,575,601]" italics="true" pageId="6" pageNumber="77">Soft parts</emphasis>
</paragraph>
<paragraph id="8B970601FFB1D025FF8FFDFD7A569BB6" blockId="6.[114,647,575,1247]" pageId="6" pageNumber="77">
The trophi and cirri (
<figureCitation id="13131A84FFB1D025FE9CFDFD79F89D10" box="[353,418,607,633]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="6" pageNumber="77">Fig. 3</figureCitation>
) are similar to those of other hydrothermal vent barnacles (
<taxonomicName id="4C287D82FFB1D025FDD1FDDC78C19DBC" authority=", Brachylepadomorpha, Verrucomorpha" authorityName="Brachylepadomorpha, Verrucomorpha" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scalpellomorpha" pageId="6" pageNumber="77" phylum="Arthropoda" rank="order">Scalpellomorpha, Brachylepadomorpha, Verrucomorpha</taxonomicName>
and
<taxonomicName id="4C287D82FFB1D025FF2BFD1979259DBC" box="[214,383,699,725]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Balanomorpha" pageId="6" pageNumber="77" phylum="Arthropoda" rank="order">Balanomorpha</taxonomicName>
), reflecting adaptations to feeding on very fine particles, presumably small, suspended bacterial clumps and filaments (cf.
<bibRefCitation id="EFB97BF0FFB1D025FF62FCB579B19C58" author="Southward A. J. &amp; Newman W. A." box="[159,491,790,817]" journalOrPublisher="Cahiers de Biologie marine" pageId="6" pageNumber="77" pagination="258 - 262" part="39" refId="ref7585" refString="Southward A. J. &amp; Newman W. A. 1998. - Ectosymbiosis between filamentous sulphur bacteria and a stalked barnacle (Scalpellomorpha, Neolepadinae) from the Lau Back Arc Basin, Tonga. Cahiers de Biologie marine 39 (3 - 4): 258 - 262." title="Ectosymbiosis between filamentous sulphur bacteria and a stalked barnacle (Scalpellomorpha, Neolepadinae) from the Lau Back Arc Basin, Tonga" type="journal article" year="1998">Southward &amp; Newman 1998</bibRefCitation>
). The slightly bullate, indented but unnotched labrum of
<taxonomicName id="4C287D82FFB1D025FF8FFCF6795A9C06" authorityName="Newman" authorityYear="2000" box="[114,256,852,879]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="yamaguchii">
<emphasis id="B95CDA13FFB1D025FF8FFCF6795A9C06" box="[114,256,852,879]" italics="true" pageId="6" pageNumber="77">I. yamaguchii</emphasis>
</taxonomicName>
(
<figureCitation id="13131A84FFB1D025FEEDFCF7793A9C06" box="[272,352,853,879]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="6" pageNumber="77">Fig. 3F</figureCitation>
) supports the usual row of fine teeth and relatively large palps provided with relatively fine, stiff setae. The mandible (
<figureCitation id="13131A84FFB1D025FDD6FC307AD89CC5" box="[555,642,914,940]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="6" pageNumber="77">Fig. 3G</figureCitation>
, right exterior, H, I, left interior) is comparable to that of other hydrothermal vent forms, especially that of
<taxonomicName id="4C287D82FFB1D025FF38FC4F7A579B61" authorityName="Newman" authorityYear="1989" box="[197,525,1005,1032]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="brachylepadoformis">
<emphasis id="B95CDA13FFB1D025FF38FC4F7A579B61" box="[197,525,1005,1032]" italics="true" pageId="6" pageNumber="77">Neoverruca brachylepadoformis</emphasis>
</taxonomicName>
, in having one superior incisiform tooth above a setose area, followed by two low teeth each supporting a row of numerous, short, sharp spines along their superior curvatures, several of the uppermost of which roll over onto the inner or posterior surface (
<figureCitation id="13131A84FFB1D025FF87FB0478939BA9" box="[122,201,1190,1216]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="6" pageNumber="77">Fig. 3I</figureCitation>
), and the upper margin of the inferior angle supports a row of similar spines.
</paragraph>
<paragraph id="8B970601FFB1D025FD58FF737CFA9EED" blockId="6.[677,1210,208,603]" pageId="6" pageNumber="77">
Maxillae I and II (
<figureCitation id="13131A84FFB1D025FC91FF737C7F9F83" box="[876,1061,208,235]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="6" pageNumber="77">Fig. 3 J, K and L</figureCitation>
, respectively) each support a relatively uniform comb of slender setae along the cutting edge, but maxillae I of
<taxonomicName id="4C287D82FFB1D025FD58FE8E7B6F9E2E" authorityName="Newman" authorityYear="2000" box="[677,821,300,327]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="yamaguchii">
<emphasis id="B95CDA13FFB1D025FD58FE8E7B6F9E2E" box="[677,821,300,327]" italics="true" pageId="6" pageNumber="77">I. yamaguchii</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C287D82FFB1D025FC3EFE8E7C619E2C" baseAuthorityName="Newman &amp; Hessler" baseAuthorityYear="1989" box="[963,1083,300,325]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB1D025FC3EFE8E7C619E2C" box="[963,1083,300,325]" italics="true" pageId="6" pageNumber="77">Neoverruca</emphasis>
</taxonomicName>
in having a single strong spine among the ordinary setae inside and just below the superior angle (
<figureCitation id="13131A84FFB1D025FBC2FEC87CC99EED" box="[1087,1171,362,388]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="6" pageNumber="77">Fig. 3K</figureCitation>
).
</paragraph>
<paragraph id="8B970601FFB1D025FD58FE2B7C299D32" blockId="6.[677,1210,208,603]" pageId="6" pageNumber="77">
What could be seen of the cirri (
<figureCitation id="13131A84FFB1D025FBF6FE2B7C139ECA" box="[1035,1097,393,419]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="6" pageNumber="77">Fig. 3</figureCitation>
A-D) suggested they were typical of most vent barnacles, but unfortunately those of the
<typeStatus id="5493B8A3FFB1D025FBEAFE677C219EB6" box="[1047,1147,453,479]" pageId="6" pageNumber="77" type="paratype">paratype</typeStatus>
were badly damaged or missing. The right side was in the best condition and the cirral counts are as follows (a, anterior ramus; p, posterior ramus; –, ramus absent; +, terminal articles missing):
</paragraph>
<paragraph id="8B970601FFB1D025FD58FD177CEA9C7E" pageId="6" pageNumber="77">
<table id="F928F4A1FFB12FDCFD58FD177CEA9C76" box="[677,1200,693,799]" gridcols="7" gridrows="3" pageId="6" pageNumber="77">
<tr id="35180443FFB12FDCFD58FD177CEA9DA0" box="[677,1200,693,713]" gridrow="0" pageId="6" pageNumber="77">
<th id="76C96D3FFFB12FDCFD58FD177A879DA0" box="[677,733,693,713]" gridcol="0" gridrow="0" pageId="6" pageNumber="77">
<emphasis id="B95CDA13FFB1D025FD58FD177A879DA0" bold="true" box="[677,733,693,713]" pageId="6" pageNumber="77">Cirrus</emphasis>
</th>
<th id="76C96D3FFFB12FDCFCF5FD177B459DA0" box="[776,799,693,713]" gridcol="1" gridrow="0" pageId="6" pageNumber="77">
<emphasis id="B95CDA13FFB1D025FCF5FD177B549DA0" bold="true" box="[776,782,693,713]" pageId="6" pageNumber="77">I</emphasis>
</th>
<th id="76C96D3FFFB12FDCFCABFD177B379DA0" box="[854,877,693,713]" gridcol="2" gridrow="0" pageId="6" pageNumber="77">
<emphasis id="B95CDA13FFB1D025FCABFD177B3B9DA0" bold="true" box="[854,865,693,713]" pageId="6" pageNumber="77">II</emphasis>
</th>
<th id="76C96D3FFFB12FDCFC5BFD177BED9DA0" box="[934,951,693,713]" gridcol="3" gridrow="0" pageId="6" pageNumber="77">
<emphasis id="B95CDA13FFB1D025FC5BFD177BED9DA0" bold="true" box="[934,951,693,713]" pageId="6" pageNumber="77">III</emphasis>
</th>
<th id="76C96D3FFFB12FDCFC08FD177C4E9DA0" box="[1013,1044,693,713]" gridcol="4" gridrow="0" pageId="6" pageNumber="77">
<emphasis id="B95CDA13FFB1D025FC08FD177C529DA0" bold="true" box="[1013,1032,693,713]" pageId="6" pageNumber="77">IV</emphasis>
</th>
<th id="76C96D3FFFB12FDCFBBEFD177C0A9DA0" box="[1091,1104,693,713]" gridcol="5" gridrow="0" pageId="6" pageNumber="77">
<emphasis id="B95CDA13FFB1D025FBBEFD177C0A9DA0" bold="true" box="[1091,1104,693,713]" pageId="6" pageNumber="77">V</emphasis>
</th>
<th id="76C96D3FFFB12FDCFB6CFD177CEA9DA0" box="[1169,1200,693,713]" gridcol="6" gridrow="0" pageId="6" pageNumber="77">
<emphasis id="B95CDA13FFB1D025FB6CFD177CFE9DA0" bold="true" box="[1169,1188,693,713]" pageId="6" pageNumber="77">VI</emphasis>
</th>
</tr>
<tr id="35180443FFB12FDCFD58FD4A7CEA9D96" box="[677,1200,744,767]" gridrow="1" pageId="6" pageNumber="77">
<th id="76C96D3FFFB12FDCFD58FD4A7A879D96" box="[677,733,744,767]" gridcol="0" gridrow="1" pageId="6" pageNumber="77">a</th>
<td id="76C96D3FFFB12FDCFCF5FD4A7B459D96" box="[776,799,744,767]" gridcol="1" gridrow="1" pageId="6" pageNumber="77">–</td>
<td id="76C96D3FFFB12FDCFCABFD4A7B379D96" box="[854,877,744,767]" gridcol="2" gridrow="1" pageId="6" pageNumber="77">7</td>
<td id="76C96D3FFFB12FDCFC5BFD4A7BED9D96" box="[934,951,744,767]" gridcol="3" gridrow="1" pageId="6" pageNumber="77">–</td>
<td id="76C96D3FFFB12FDCFC08FD4A7C4E9D96" box="[1013,1044,744,767]" gridcol="4" gridrow="1" pageId="6" pageNumber="77">–</td>
<td id="76C96D3FFFB12FDCFBBEFD4A7C0A9D96" box="[1091,1104,744,767]" gridcol="5" gridrow="1" pageId="6" pageNumber="77">–</td>
<td id="76C96D3FFFB12FDCFB6CFD4A7CEA9D96" box="[1169,1200,744,767]" gridcol="6" gridrow="1" pageId="6" pageNumber="77">37+</td>
</tr>
<tr id="35180443FFB12FDCFD58FCAA7CEA9C76" box="[677,1200,776,799]" gridrow="2" pageId="6" pageNumber="77">
<th id="76C96D3FFFB12FDCFD58FCAA7A879C76" box="[677,733,776,799]" gridcol="0" gridrow="2" pageId="6" pageNumber="77">p</th>
<td id="76C96D3FFFB12FDCFCF5FCAA7B459C76" box="[776,799,776,799]" gridcol="1" gridrow="2" pageId="6" pageNumber="77">20</td>
<td id="76C96D3FFFB12FDCFCABFCAA7B379C76" box="[854,877,776,799]" gridcol="2" gridrow="2" pageId="6" pageNumber="77">13</td>
<td id="76C96D3FFFB12FDCFC5BFCAA7BED9C76" box="[934,951,776,799]" gridcol="3" gridrow="2" pageId="6" pageNumber="77">–</td>
<td id="76C96D3FFFB12FDCFC08FCAA7C4E9C76" box="[1013,1044,776,799]" gridcol="4" gridrow="2" pageId="6" pageNumber="77">39+</td>
<td id="76C96D3FFFB12FDCFBBEFCAA7C0A9C76" box="[1091,1104,776,799]" gridcol="5" gridrow="2" pageId="6" pageNumber="77">–</td>
<td id="76C96D3FFFB12FDCFB6CFCAA7CEA9C76" box="[1169,1200,776,799]" gridcol="6" gridrow="2" pageId="6" pageNumber="77">41+</td>
</tr>
</table>
</paragraph>
<paragraph id="8B970601FFB1D025FD58FC307C079BB6" blockId="6.[677,1210,913,1247]" pageId="6" pageNumber="77">
As can be observed (
<figureCitation id="13131A84FFB1D025FC81FC307BB79CC2" box="[892,1005,913,940]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="6" pageNumber="77">Fig. 3A, B</figureCitation>
), the first two pairs of cirri are modified as maxillipeds and at least the posterior ramus of the first as well as of the second are antenniform. The posterior four pairs of cirri are normal (ctenopod), but the lesser curvature of the intermediate articles of cirrus VI each have as many as five pairs of moderately long setae, fewer than
<taxonomicName id="4C287D82FFB1D025FD21FBCA7B089BE8" baseAuthorityName="Newman &amp; Hessler" baseAuthorityYear="1989" box="[732,850,1128,1153]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB1D025FD21FBCA7B089BE8" box="[732,850,1128,1153]" italics="true" pageId="6" pageNumber="77">Neoverruca</emphasis>
</taxonomicName>
, and their s/a ratios are markedly different, that for
<taxonomicName id="4C287D82FFB1D025FC9CFB257BA59BC8" authorityName="Newman" authorityYear="2000" box="[865,1023,1159,1185]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB1D025FC9CFB257BA59BC8" box="[865,1023,1159,1185]" italics="true" pageId="6" pageNumber="77">Imbricaverruca</emphasis>
</taxonomicName>
falling within the range for other vent barnacles while that for
<taxonomicName id="4C287D82FFB1D025FD58FB667B409BB4" baseAuthorityName="Newman &amp; Hessler" baseAuthorityYear="1989" box="[677,794,1220,1245]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB1D025FD58FB667B409BB4" box="[677,794,1220,1245]" italics="true" pageId="6" pageNumber="77">Neoverruca</emphasis>
</taxonomicName>
is the lowest known (
<tableCitation id="C6AA33BAFFB1D025FC00FB677C179BB6" box="[1021,1101,1221,1247]" captionStart="TABLE" captionStartId="6.[114,125,1316,1333]" captionTargetPageId="6" captionText="TABLE 1. — Seta/article ratios for four neolepadines compared to four sessile barnacles also from vents: the ratio (the width of the article of an intermediate segment of cirrus VI as the denominator and the length of longest seta of that article as the numerator), while highest in the bacteria-farming neolepadine (Lau sp. A), is lowest in Neoverruca. That for Imbricaverruca as well as for Neobrachylepas are not only similar to each other but fall in the low end of the range for vent barnacles." pageId="6" pageNumber="77">Table 1</tableCitation>
).
</paragraph>
<caption id="DF575689FFB1D025FF8FFA867B8B9A1C" pageId="6" pageNumber="77" startId="6.[114,125,1316,1333]" targetBox="[114,1177,1432,1687]" targetIsTable="true" targetPageId="6">
<paragraph id="8B970601FFB1D025FF8FFA867B8B9A1C" blockId="6.[114,1211,1316,1397]" pageId="6" pageNumber="77">
TABLE 1. — Seta/article ratios for four neolepadines compared to four sessile barnacles also from vents: the ratio (the width of the article of an intermediate segment of cirrus VI as the denominator and the length of longest seta of that article as the numerator), while highest in the bacteria-farming neolepadine (Lau sp. A), is lowest in
<taxonomicName id="4C287D82FFB1D025FCFCFAED7B3E9A09" baseAuthorityName="Newman &amp; Hessler" baseAuthorityYear="1989" box="[769,868,1359,1376]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB1D025FCFCFAED7B3E9A09" box="[769,868,1359,1376]" italics="true" pageId="6" pageNumber="77">Neoverruca</emphasis>
</taxonomicName>
. That for
<taxonomicName id="4C287D82FFB1D025FC41FAED7C669A09" authorityName="Newman" authorityYear="2000" box="[956,1084,1359,1376]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB1D025FC41FAED7C669A09" box="[956,1084,1359,1376]" italics="true" pageId="6" pageNumber="77">Imbricaverruca</emphasis>
</taxonomicName>
as well as for
<taxonomicName id="4C287D82FFB1D025FF8FFAC678A29A1C" authorityName="Newman &amp; Yamaguchi" authorityYear="1995" box="[114,248,1380,1397]" class="Maxillopoda" family="Neobrachylepadidae" genus="Neobrachylepas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB1D025FF8FFAC678A29A1C" box="[114,248,1380,1397]" italics="true" pageId="6" pageNumber="77">Neobrachylepas</emphasis>
</taxonomicName>
are not only similar to each other but fall in the low end of the range for vent barnacles.
</paragraph>
</caption>
<paragraph id="8B970601FFB1D025FE2BFA3A7CC399FE" pageId="6" pageNumber="77">
<table id="F928F4A1FFB12FDCFF8FFA3A7CC399FE" box="[114,1177,1432,1687]" gridcols="5" gridrows="10" pageId="6" pageNumber="77">
<tr id="35180443FFB12FDCFF8FFA3A7CC39AC5" box="[114,1177,1432,1452]" gridrow="0" pageId="6" pageNumber="77" rowspan-0="1">
<th id="76C96D3FFFB12FDCFE62FA3A7A949AC5" box="[415,718,1432,1452]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="6" pageNumber="77">
<emphasis id="B95CDA13FFB1D025FE2BFA3A7AFD9AC5" bold="true" box="[470,679,1432,1452]" pageId="6" pageNumber="77">Measurement in mm</emphasis>
</th>
<th id="76C96D3FFFB12FDCFCF0FA3A7B199AC5" box="[781,835,1432,1452]" gridcol="3" gridrow="0" pageId="6" pageNumber="77">
<emphasis id="B95CDA13FFB1D025FCF0FA3A7B199AC5" bold="true" box="[781,835,1432,1452]" pageId="6" pageNumber="77">Ratio</emphasis>
</th>
<th id="76C96D3FFFB12FDCFC76FA3A7CC39AC5" box="[907,1177,1432,1452]" gridcol="4" gridrow="0" pageId="6" pageNumber="77">
<emphasis id="B95CDA13FFB1D025FC23FA3A7C1D9AC5" bold="true" box="[990,1095,1432,1452]" pageId="6" pageNumber="77">Reference</emphasis>
</th>
</tr>
<tr id="35180443FFB12FDCFF8FFA127CC39AAD" box="[114,1177,1456,1476]" gridrow="1" pageId="6" pageNumber="77" rowspan-0="1" rowspan-4="1">
<td id="76C96D3FFFB12FDCFE62FA127A469AAD" box="[415,540,1456,1476]" gridcol="1" gridrow="1" pageId="6" pageNumber="77">
<emphasis id="B95CDA13FFB1D025FE62FA127A469AAD" bold="true" box="[415,540,1456,1476]" pageId="6" pageNumber="77">article width</emphasis>
</td>
<td id="76C96D3FFFB12FDCFDAAFA127A949AAD" box="[599,718,1456,1476]" gridcol="2" gridrow="1" pageId="6" pageNumber="77">
<emphasis id="B95CDA13FFB1D025FDAAFA127A949AAD" bold="true" box="[599,718,1456,1476]" pageId="6" pageNumber="77">setal length</emphasis>
</td>
<td id="76C96D3FFFB12FDCFCF0FA127B199AAD" box="[781,835,1456,1476]" gridcol="3" gridrow="1" pageId="6" pageNumber="77">
<emphasis id="B95CDA13FFB1D025FCE4FA127B629AAD" bold="true" box="[793,824,1456,1476]" pageId="6" pageNumber="77">s/a</emphasis>
</td>
</tr>
<tr id="35180443FFB12FDCFF8FFA797CC39A86" box="[114,1177,1499,1519]" gridrow="2" pageId="6" pageNumber="77">
<th id="76C96D3FFFB12FDCFF8FFA7979009A86" box="[114,346,1499,1519]" gridcol="0" gridrow="2" pageId="6" pageNumber="77">Lau sp. A</th>
<td id="76C96D3FFFB12FDCFE62FA797A469A86" box="[415,540,1499,1519]" gridcol="1" gridrow="2" pageId="6" pageNumber="77">0.33</td>
<td id="76C96D3FFFB12FDCFDAAFA797A949A86" box="[599,718,1499,1519]" gridcol="2" gridrow="2" pageId="6" pageNumber="77">4.60</td>
<td id="76C96D3FFFB12FDCFCF0FA797B199A86" box="[781,835,1499,1519]" gridcol="3" gridrow="2" pageId="6" pageNumber="77">14.0</td>
<td id="76C96D3FFFB12FDCFC76FA797CC39A86" box="[907,1177,1499,1519]" gridcol="4" gridrow="2" pageId="6" pageNumber="77">
<bibRefCitation id="EFB97BF0FFB1D025FC70FA797CCD9A86" author="Southward A. J. &amp; Newman W. A." box="[909,1175,1499,1519]" journalOrPublisher="Cahiers de Biologie marine" pageId="6" pageNumber="77" pagination="258 - 262" part="39" refId="ref7585" refString="Southward A. J. &amp; Newman W. A. 1998. - Ectosymbiosis between filamentous sulphur bacteria and a stalked barnacle (Scalpellomorpha, Neolepadinae) from the Lau Back Arc Basin, Tonga. Cahiers de Biologie marine 39 (3 - 4): 258 - 262." title="Ectosymbiosis between filamentous sulphur bacteria and a stalked barnacle (Scalpellomorpha, Neolepadinae) from the Lau Back Arc Basin, Tonga" type="journal article" year="1998">Southward &amp; Newman 1998</bibRefCitation>
</td>
</tr>
<tr id="35180443FFB12FDCFF8FFA517CC3996E" box="[114,1177,1523,1543]" gridrow="3" pageId="6" pageNumber="77">
<th id="76C96D3FFFB12FDCFF8FFA517900996E" box="[114,346,1523,1543]" gridcol="0" gridrow="3" pageId="6" pageNumber="77">Lau sp. B</th>
<td id="76C96D3FFFB12FDCFE62FA517A46996E" box="[415,540,1523,1543]" gridcol="1" gridrow="3" pageId="6" pageNumber="77">0.10</td>
<td id="76C96D3FFFB12FDCFDAAFA517A94996E" box="[599,718,1523,1543]" gridcol="2" gridrow="3" pageId="6" pageNumber="77">0.67</td>
<td id="76C96D3FFFB12FDCFCF0FA517B19996E" box="[781,835,1523,1543]" gridcol="3" gridrow="3" pageId="6" pageNumber="77">6.7</td>
<td id="76C96D3FFFB12FDCFC76FA517CC3996E" box="[907,1177,1523,1543]" gridcol="4" gridrow="3" pageId="6" pageNumber="77">
<emphasis id="B95CDA13FFB1D025FC06FA517C70996E" box="[1019,1066,1523,1543]" italics="true" pageId="6" pageNumber="77">idem</emphasis>
</td>
</tr>
<tr id="35180443FFB12FDCFF8FF9A97CC39976" box="[114,1177,1547,1567]" gridrow="4" pageId="6" pageNumber="77">
<th id="76C96D3FFFB12FDCFF8FF9A979009976" box="[114,346,1547,1567]" gridcol="0" gridrow="4" pageId="6" pageNumber="77">South-East Indian Ridge</th>
<td id="76C96D3FFFB12FDCFE62F9A97A469976" box="[415,540,1547,1567]" gridcol="1" gridrow="4" pageId="6" pageNumber="77">0.18</td>
<td id="76C96D3FFFB12FDCFDAAF9A97A949976" box="[599,718,1547,1567]" gridcol="2" gridrow="4" pageId="6" pageNumber="77">1.70</td>
<td id="76C96D3FFFB12FDCFCF0F9A97B199976" box="[781,835,1547,1567]" gridcol="3" gridrow="4" pageId="6" pageNumber="77">9.4</td>
<td id="76C96D3FFFB12FDCFC76F9A97CC39976" box="[907,1177,1547,1567]" gridcol="4" gridrow="4" pageId="6" pageNumber="77">
<emphasis id="B95CDA13FFB1D025FC06F9A97C709976" box="[1019,1066,1547,1567]" italics="true" pageId="6" pageNumber="77">idem</emphasis>
</td>
</tr>
<tr id="35180443FFB12FDCFF8FF9817CC3995E" box="[114,1177,1571,1591]" gridrow="5" pageId="6" pageNumber="77">
<th id="76C96D3FFFB12FDCFF8FF9817900995E" box="[114,346,1571,1591]" gridcol="0" gridrow="5" pageId="6" pageNumber="77">Lihir volcano</th>
<td id="76C96D3FFFB12FDCFE62F9817A46995E" box="[415,540,1571,1591]" gridcol="1" gridrow="5" pageId="6" pageNumber="77">0.30</td>
<td id="76C96D3FFFB12FDCFDAAF9817A94995E" box="[599,718,1571,1591]" gridcol="2" gridrow="5" pageId="6" pageNumber="77">1.85</td>
<td id="76C96D3FFFB12FDCFCF0F9817B19995E" box="[781,835,1571,1591]" gridcol="3" gridrow="5" pageId="6" pageNumber="77">6.2</td>
<td id="76C96D3FFFB12FDCFC76F9817CC3995E" box="[907,1177,1571,1591]" gridcol="4" gridrow="5" pageId="6" pageNumber="77">
<emphasis id="B95CDA13FFB1D025FC06F9817C70995E" box="[1019,1066,1571,1591]" italics="true" pageId="6" pageNumber="77">idem</emphasis>
</td>
</tr>
<tr id="35180443FFB12FDCFF8FF9997CC39926" box="[114,1177,1595,1615]" gridrow="6" pageId="6" pageNumber="77">
<th id="76C96D3FFFB12FDCFF8FF99979009926" box="[114,346,1595,1615]" gridcol="0" gridrow="6" pageId="6" pageNumber="77">
<taxonomicName id="4C287D82FFB1D025FF8FF99979569926" authorityName="Newman &amp; Yamaguchi" authorityYear="1995" box="[114,268,1595,1615]" class="Maxillopoda" family="Neobrachylepadidae" genus="Neobrachylepas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB1D025FF8FF99979569926" box="[114,268,1595,1615]" italics="true" pageId="6" pageNumber="77">Neobrachylepas</emphasis>
</taxonomicName>
</th>
<td id="76C96D3FFFB12FDCFE62F9997A469926" box="[415,540,1595,1615]" gridcol="1" gridrow="6" pageId="6" pageNumber="77">0.07</td>
<td id="76C96D3FFFB12FDCFDAAF9997A949926" box="[599,718,1595,1615]" gridcol="2" gridrow="6" pageId="6" pageNumber="77">0.31</td>
<td id="76C96D3FFFB12FDCFCF0F9997B199926" box="[781,835,1595,1615]" gridcol="3" gridrow="6" pageId="6" pageNumber="77">4.4</td>
<td id="76C96D3FFFB12FDCFC76F9997CC39926" box="[907,1177,1595,1615]" gridcol="4" gridrow="6" pageId="6" pageNumber="77">
<bibRefCitation id="EFB97BF0FFB1D025FC76F9997CC39926" author="Newman W. A. &amp; Yamaguchi T." box="[907,1177,1595,1615]" journalOrPublisher="Bulletin du Museum national d'Histoire naturelle" pageId="6" pageNumber="77" pagination="211 - 243" part="17" refId="ref7319" refString="Newman W. A. &amp; Yamaguchi T. 1995. - A new sessile barnacle (Cirripedia, Brachylepadomorpha) from the Lau Back-Arc Basin, Tonga; first record of a living representative since the Miocene. Bulletin du Museum national d'Histoire naturelle, 4 e serie, 17 A (3 - 4): 211 - 243." title="A new sessile barnacle (Cirripedia, Brachylepadomorpha) from the Lau Back-Arc Basin, Tonga; first record of a living representative since the Miocene" type="journal article" year="1995">Newman &amp; Yamaguchi 1995</bibRefCitation>
</td>
</tr>
<tr id="35180443FFB12FDCFF8FF9F17CC3990E" box="[114,1177,1619,1639]" gridrow="7" pageId="6" pageNumber="77">
<th id="76C96D3FFFB12FDCFF8FF9F17900990E" box="[114,346,1619,1639]" gridcol="0" gridrow="7" pageId="6" pageNumber="77">
<taxonomicName id="4C287D82FFB1D025FF8FF9F17885990E" baseAuthorityName="Newman &amp; Hessler" baseAuthorityYear="1989" box="[114,223,1619,1639]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB1D025FF8FF9F17885990E" box="[114,223,1619,1639]" italics="true" pageId="6" pageNumber="77">Neoverruca</emphasis>
</taxonomicName>
</th>
<td id="76C96D3FFFB12FDCFE62F9F17A46990E" box="[415,540,1619,1639]" gridcol="1" gridrow="7" pageId="6" pageNumber="77">0.25</td>
<td id="76C96D3FFFB12FDCFDAAF9F17A94990E" box="[599,718,1619,1639]" gridcol="2" gridrow="7" pageId="6" pageNumber="77">0.70</td>
<td id="76C96D3FFFB12FDCFCF0F9F17B19990E" box="[781,835,1619,1639]" gridcol="3" gridrow="7" pageId="6" pageNumber="77">2.8</td>
<td id="76C96D3FFFB12FDCFC76F9F17CC3990E" box="[907,1177,1619,1639]" gridcol="4" gridrow="7" pageId="6" pageNumber="77">
<bibRefCitation id="EFB97BF0FFB1D025FC60F9F17CDD990E" author="Newman W. A. &amp; Hessler R. R." box="[925,1159,1619,1639]" journalOrPublisher="San Diego Society of Natural History Transactions" pageId="6" pageNumber="77" pagination="259 - 273" part="21" refId="ref7233" refString="Newman W. A. &amp; Hessler R. R. 1989. - A new abyssal hydrothermal verrucomorphan (Cirripedia; Sessilia): The most primitive living sessile barnacle. San Diego Society of Natural History Transactions 21 (16): 259 - 273." title="A new abyssal hydrothermal verrucomorphan (Cirripedia; Sessilia): The most primitive living sessile barnacle" type="journal article" year="1989">Newman &amp; Hessler 1989</bibRefCitation>
</td>
</tr>
<tr id="35180443FFB12FDCFF8FF9C97CC39916" box="[114,1177,1643,1663]" gridrow="8" pageId="6" pageNumber="77">
<th id="76C96D3FFFB12FDCFF8FF9C979009916" box="[114,346,1643,1663]" gridcol="0" gridrow="8" pageId="6" pageNumber="77">
<taxonomicName id="4C287D82FFB1D025FF8FF9C978A59916" authorityName="Newman" authorityYear="2000" box="[114,255,1643,1663]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB1D025FF8FF9C978A59916" box="[114,255,1643,1663]" italics="true" pageId="6" pageNumber="77">Imbricaverruca</emphasis>
</taxonomicName>
</th>
<td id="76C96D3FFFB12FDCFE62F9C97A469916" box="[415,540,1643,1663]" gridcol="1" gridrow="8" pageId="6" pageNumber="77">0.15</td>
<td id="76C96D3FFFB12FDCFDAAF9C97A949916" box="[599,718,1643,1663]" gridcol="2" gridrow="8" pageId="6" pageNumber="77">0.68</td>
<td id="76C96D3FFFB12FDCFCF0F9C97B199916" box="[781,835,1643,1663]" gridcol="3" gridrow="8" pageId="6" pageNumber="77">4.5</td>
<td id="76C96D3FFFB12FDCFC76F9C97CC39916" box="[907,1177,1643,1663]" gridcol="4" gridrow="8" pageId="6" pageNumber="77">This paper</td>
</tr>
<tr id="35180443FFB12FDCFF8FF9217CC399FE" box="[114,1177,1667,1687]" gridrow="9" pageId="6" pageNumber="77">
<th id="76C96D3FFFB12FDCFF8FF921790099FE" box="[114,346,1667,1687]" gridcol="0" gridrow="9" pageId="6" pageNumber="77">
<taxonomicName id="4C287D82FFB1D025FF8FF921791299FE" authority="(NFB)" baseAuthorityName="NFB" box="[114,328,1667,1687]" class="Maxillopoda" family="Chionelasmatidae" genus="Eochionelasmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB1D025FF8FF921795799FE" box="[114,269,1667,1687]" italics="true" pageId="6" pageNumber="77">Eochionelasmus</emphasis>
(NFB)
</taxonomicName>
</th>
<td id="76C96D3FFFB12FDCFE62F9217A4699FE" box="[415,540,1667,1687]" gridcol="1" gridrow="9" pageId="6" pageNumber="77">0.25</td>
<td id="76C96D3FFFB12FDCFDAAF9217A9499FE" box="[599,718,1667,1687]" gridcol="2" gridrow="9" pageId="6" pageNumber="77">1.65</td>
<td id="76C96D3FFFB12FDCFCF0F9217B1999FE" box="[781,835,1667,1687]" gridcol="3" gridrow="9" pageId="6" pageNumber="77">6.6</td>
<td id="76C96D3FFFB12FDCFC76F9217CC399FE" box="[907,1177,1667,1687]" gridcol="4" gridrow="9" pageId="6" pageNumber="77">
<bibRefCitation id="EFB97BF0FFB1D025FC76F9217CC399FE" author="Yamaguchi T. &amp; Newman W. A." box="[907,1177,1667,1687]" journalOrPublisher="Pacific Science" pageId="6" pageNumber="77" pagination="135 - 155" part="44" refId="ref7889" refString="Yamaguchi T. &amp; Newman W. A. 1990. - A new and primitive barnacle (Cirripedia; Balanomorpha) from the North Fiji Basin abyssal hydrothermal field, and its evolutionary implications. Pacific Science 44 (2): 135 - 155." title="A new and primitive barnacle (Cirripedia; Balanomorpha) from the North Fiji Basin abyssal hydrothermal field, and its evolutionary implications" type="journal article" year="1990">Yamaguchi &amp; Newman 1990</bibRefCitation>
</td>
</tr>
</table>
</paragraph>
<caption id="DF575689FFB0D024FF8CFA1F790599CD" ID-DOI="http://doi.org/10.5281/zenodo.13988759" ID-Zenodo-Dep="13988759" httpUri="https://zenodo.org/record/13988759/files/figure.png" pageId="7" pageNumber="78" startId="7.[113,141,1469,1486]" subCaptionStartIDs="7.[323,440,1597,1614]" subCaptionStarts="Abbr" targetBox="[112,1165,215,1445]" targetPageId="7">
<paragraph id="8B970601FFB0D024FF8CFA1F790599CD" blockId="7.[113,1210,1469,1700]" pageId="7" pageNumber="78">
Fig. 4. — Schematic plan views and transverse sections of grades of skeletal organization from the most primitive sessile barnacles (
<emphasis id="B95CDA13FFB0D024FF8BFA7178D99A8D" bold="true" box="[118,131,1491,1508]" pageId="7" pageNumber="78">A</emphasis>
, Brachylepadomorpha) through the asymmetrical sessile barnacles (
<emphasis id="B95CDA13FFB0D024FD3FFA717ABE9A8D" bold="true" box="[706,740,1491,1508]" pageId="7" pageNumber="78">B -H</emphasis>
,
<taxonomicName id="4C287D82FFB0D024FD12FA717B2E9A8D" authorityName="Pilsbry" authorityYear="1916" box="[751,884,1491,1508]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="73" phylum="Arthropoda" rank="subOrder" subOrder="Verrucomorpha">Verrucomorpha</taxonomicName>
). Of the eight verrucid genera currently recognized (
<bibRefCitation id="EFB97BF0FFB0D024FF24FA4A791A9A90" author="Young P. S." box="[217,320,1512,1529]" journalOrPublisher="Zoosystema" pageId="7" pageNumber="78" pagination="31 - 92" part="20" refId="ref7981" refString="Young P. S. 1998. - Cirripedia (Crustacea) from the &quot; Campagne Biacores &quot; in the Azores region, including a generic revision of Verrucidae. Zoosystema 20 (1): 31 - 92." title="Cirripedia (Crustacea) from the &quot; Campagne Biacores &quot; in the Azores region, including a generic revision of Verrucidae" type="journal article" year="1998">Young 1998</bibRefCitation>
) only the four most representative ones (
<taxonomicName id="4C287D82FFB0D024FD5DFA4A7AA29A90" authorityName="Pilsbry" authorityYear="1916" box="[672,760,1512,1529]" class="Maxillopoda" family="Verrucidae" genus="Altiverruca" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB0D024FD5DFA4A7AA29A90" box="[672,760,1512,1529]" italics="true" pageId="7" pageNumber="78">Altiverruca</emphasis>
</taxonomicName>
,
<taxonomicName id="4C287D82FFB0D024FCF9FA4A7BD59A90" authorityName="Young" authorityYear="1998" box="[772,911,1512,1529]" class="Maxillopoda" family="Verrucidae" genus="Newmaniverruca" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB0D024FCF9FA4A7BD59A90" box="[772,911,1512,1529]" italics="true" pageId="7" pageNumber="78">Newmaniverruca</emphasis>
</taxonomicName>
,
<taxonomicName id="4C287D82FFB0D024FC66FA4A7C589A90" authorityName="Pilsbry" authorityYear="1916" box="[923,1026,1512,1529]" class="Maxillopoda" family="Verrucidae" genus="Metaverruca" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFB0D024FC66FA4A7C589A90" box="[923,1026,1512,1529]" italics="true" pageId="7" pageNumber="78">Metaverruca</emphasis>
</taxonomicName>
and
<taxonomicName id="4C287D82FFB0D024FBCDFA4A7C2A9A90" authorityName="Schumacher" authorityYear="1817" box="[1072,1136,1512,1529]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="78" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B95CDA13FFB0D024FBCDFA4A7C2A9A90" box="[1072,1136,1512,1529]" italics="true" pageId="7" pageNumber="78">Verruca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F6768FFB0D024FB8BFA4A7CCE9A90" box="[1142,1172,1512,1529]" pageId="7" pageNumber="78" sensu="stricto">s.s.</taxonomicNameLabel>
) are depicted here. Right sides are to the viewers right and rostral ends are downward in plan views or coming out of the page in transverse sections. The transverse sections are slightly rostral of the rostro-carinal gap or suture. While there are numerous whorls of imbricating plates standing in tiers four plates high in brachylepadomorphs, and on the movable side of neoverrucids, only those of the R-C gap are labeled. Abbreviations:
<emphasis id="B95CDA13FFB0D024FE42F99F79979927" bold="true" box="[447,461,1597,1614]" pageId="7" pageNumber="78">C</emphasis>
,
<emphasis id="B95CDA13FFB0D024FE2AF99F79BE9927" bold="true" box="[471,484,1597,1614]" pageId="7" pageNumber="78">R</emphasis>
, carina and rostrum;
<emphasis id="B95CDA13FFB0D024FD6AF99F7AE6992E" bold="true" box="[663,700,1594,1614]" pageId="7" pageNumber="78">
cl
<superScript id="7C5DAB49FFB0D024FD5BF9987AE6992E" attach="left" box="[678,700,1594,1607]" fontSize="5" pageId="7" pageNumber="78">1-4</superScript>
</emphasis>
,
<emphasis id="B95CDA13FFB0D024FD3BF99F7ABA992E" bold="true" box="[710,736,1594,1614]" pageId="7" pageNumber="78">
l
<superScript id="7C5DAB49FFB0D024FD37F9987ABA992E" attach="left" box="[714,736,1594,1607]" fontSize="5" pageId="7" pageNumber="78">1-4</superScript>
</emphasis>
,
<emphasis id="B95CDA13FFB0D024FD17F99F7B56992E" bold="true" box="[746,780,1594,1614]" pageId="7" pageNumber="78">
rl
<superScript id="7C5DAB49FFB0D024FD0BF9987B56992E" attach="left" box="[758,780,1594,1607]" fontSize="5" pageId="7" pageNumber="78">1-4</superScript>
</emphasis>
, carinolateral, lateral and rostrolateral tiers of imbricating plates respectively;
<emphasis id="B95CDA13FFB0D024FEA9F9F1793E990D" bold="true" box="[340,356,1619,1636]" pageId="7" pageNumber="78">cl</emphasis>
,
<emphasis id="B95CDA13FFB0D024FE92F9F17921990D" bold="true" box="[367,379,1619,1636]" pageId="7" pageNumber="78">rl</emphasis>
, the pair of imbricating plates in proverrucids;
<emphasis id="B95CDA13FFB0D024FCF6F9F17B78990D" bold="true" box="[779,802,1619,1636]" pageId="7" pageNumber="78">FS</emphasis>
,
<emphasis id="B95CDA13FFB0D024FCD3F9F17B1E990D" bold="true" box="[814,836,1619,1636]" pageId="7" pageNumber="78">FT</emphasis>
, fixed scutum and tergum;
<emphasis id="B95CDA13FFB0D024FBCCF9F17C1C990D" bold="true" box="[1073,1094,1619,1636]" pageId="7" pageNumber="78">hl,</emphasis>
hinge line;
<emphasis id="B95CDA13FFB0D024FB54F9F17CEE990D" bold="true" box="[1193,1204,1619,1636]" pageId="7" pageNumber="78">L</emphasis>
, median latus (completely lost in proverrucids and verrucids);
<emphasis id="B95CDA13FFB0D024FD7DF9CA7ACB9910" bold="true" box="[640,657,1640,1657]" pageId="7" pageNumber="78">m</emphasis>
, myophore;
<emphasis id="B95CDA13FFB0D024FD02F9CA7B469910" bold="true" box="[767,796,1640,1657]" pageId="7" pageNumber="78">MS</emphasis>
,
<emphasis id="B95CDA13FFB0D024FCD4F9CA7B1F9910" bold="true" box="[809,837,1640,1657]" pageId="7" pageNumber="78">MT</emphasis>
, movable scutum and tergum;
<emphasis id="B95CDA13FFB0D024FBA8F9CA7C219910" bold="true" box="[1109,1147,1640,1657]" pageId="7" pageNumber="78">sam</emphasis>
, scutal adductor muscle;
<emphasis id="B95CDA13FFB0D024FEF5F9DF794E99E7" bold="true" box="[264,276,1661,1678]" pageId="7" pageNumber="78">S</emphasis>
,
<emphasis id="B95CDA13FFB0D024FEE3F9DF797399E7" bold="true" box="[286,297,1661,1678]" pageId="7" pageNumber="78">T</emphasis>
, normal terga and scuta in brachylepadomorphs; dashed horizontal lines: membranous basis (transverse sections F-H after
<bibRefCitation id="EFB97BF0FFB0D024FF13F931790D99CD" author="Young P. S." box="[238,343,1683,1700]" journalOrPublisher="Zoosystema" pageId="7" pageNumber="78" pagination="31 - 92" part="20" refId="ref7981" refString="Young P. S. 1998. - Cirripedia (Crustacea) from the &quot; Campagne Biacores &quot; in the Azores region, including a generic revision of Verrucidae. Zoosystema 20 (1): 31 - 92." title="Cirripedia (Crustacea) from the &quot; Campagne Biacores &quot; in the Azores region, including a generic revision of Verrucidae" type="journal article" year="1998">Young 1998</bibRefCitation>
).
</paragraph>
</caption>
<paragraph id="8B970601FFBFD02BFF8FFF7379699DDE" blockId="8.[114,647,209,695]" pageId="8" pageNumber="79">
The caudal appendages, while multiarticulate in
<taxonomicName id="4C287D82FFBFD02BFF8FFF4D794C9E60" box="[114,278,239,265]" class="Maxillopoda" family="Chionelasmatidae" genus="Eochionelasmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFBFD02BFF8FFF4D794C9E60" box="[114,278,239,265]" italics="true" pageId="8" pageNumber="79">Eochionelasmus</emphasis>
</taxonomicName>
and minute and uniarticulate in
<taxonomicName id="4C287D82FFBFD02BFF8FFEAC79259E41" authority="Newman, 1979" authorityName="Newman" authorityYear="1979" box="[114,383,269,296]" class="Maxillopoda" family="Eolepadidae" genus="Neolepas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="8" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFBFD02BFF8FFEAC78979E41" box="[114,205,270,296]" italics="true" pageId="8" pageNumber="79">Neolepas</emphasis>
<bibRefCitation id="EFB97BF0FFBFD02BFF29FEAF79259E41" author="Newman W. A." box="[212,383,269,296]" journalOrPublisher="San Diego Society of Natural History Transactions" pageId="8" pageNumber="79" pagination="153 - 167" part="19" refId="ref6965" refString="Newman W. A. 1979. - A new scalpellid (Cirripedia); a Mesozoic relic living near an abyssal hydrothermal spring. San Diego Society of Natural History Transactions 19 (11): 153 - 167." title="A new scalpellid (Cirripedia); a Mesozoic relic living near an abyssal hydrothermal spring" type="journal article" year="1979">Newman, 1979</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C287D82FFBFD02BFE44FEAC7A029E41" authorityName="Newman &amp; Yamaguchi" authorityYear="1995" box="[441,600,270,296]" class="Maxillopoda" family="Neobrachylepadidae" genus="Neobrachylepas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFBFD02BFE44FEAC7A029E41" box="[441,600,270,296]" italics="true" pageId="8" pageNumber="79">Neobrachylepas</emphasis>
</taxonomicName>
, are absent in
<taxonomicName id="4C287D82FFBFD02BFF19FE8F79D19E2E" authorityName="Newman" authorityYear="2000" box="[228,395,301,327]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFBFD02BFF19FE8F79D19E2E" box="[228,395,301,327]" italics="true" pageId="8" pageNumber="79">Imbricaverruca</emphasis>
</taxonomicName>
as well as
<taxonomicName id="4C287D82FFBFD02BFDF6FE8E7ADD9E2C" baseAuthorityName="Newman &amp; Hessler" baseAuthorityYear="1989" box="[523,647,300,325]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFBFD02BFDF6FE8E7ADD9E2C" box="[523,647,300,325]" italics="true" pageId="8" pageNumber="79">Neoverruca</emphasis>
</taxonomicName>
(
<figureCitation id="13131A84FFBFD02BFF87FEE978809E0C" box="[122,218,331,357]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="8" pageNumber="79">Fig. 3C</figureCitation>
). Both adults were hermaphroditic; penis long, conspicuously annulated, and sparsely clothed with short, stiff setae (
<figureCitation id="13131A84FFBFD02BFE04FE2B7A239ECB" box="[505,633,392,419]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="8" pageNumber="79">Fig. 3C, E</figureCitation>
). While the generally well-developed penes of vent barnacles have a well-developed pedicel, there is no basidorsal point (
<figureCitation id="13131A84FFBFD02BFE9FFE4779E49E96" box="[354,446,485,511]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="8" pageNumber="79">Fig. 3C</figureCitation>
). Well-developed testes extend into the pedicels of cirrus I &amp; II (
<figureCitation id="13131A84FFBFD02BFF84FD8078A89D52" box="[121,242,545,572]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="8" pageNumber="79">Fig. 3A, B</figureCitation>
), and to a lesser degree into at least those of cirrus III &amp; IV. Ovigerous frena were not seen, but no ovigerous lamellae were being incubated and in their absence frena may be easily overlooked.
</paragraph>
<paragraph id="8B970601FFBFD02BFD58FF727B609F83" blockId="8.[677,1210,208,694]" box="[677,826,208,234]" pageId="8" pageNumber="79">
<emphasis id="B95CDA13FFBFD02BFD58FF727B609F83" box="[677,826,208,234]" italics="true" pageId="8" pageNumber="79">The x-juvenile</emphasis>
</paragraph>
<paragraph id="8B970601FFBFD02BFD58FF4D7CE09DDF" blockId="8.[677,1210,208,694]" pageId="8" pageNumber="79">
A number of attached cyprids found on sulfide deposits associated with hydrothermal activity were included with the collection from the Lau Basin sent by CENTOB. Some were sent to Peter G. Jensen for the study of lattice organs (cf.
<bibRefCitation id="EFB97BF0FFBFD02BFD58FE2A7B3D9ECB" author="Jensen P. G. &amp; HOeg J. T. &amp; Bower S. &amp; Rybakov A." box="[677,871,392,418]" journalOrPublisher="Canadian Journal of Zoology" pageId="8" pageNumber="79" pagination="1018 - 1026" part="72" refId="ref6802" refString="Jensen P. G., HOeg J. T., Bower S. &amp; Rybakov A. 1994. - Scanning electron microscopy of lattice organs in cyprids of the Rhizocephala Akentrogonida (Crustacea, Cirripedia). Canadian Journal of Zoology 72 (6): 1018 - 1026." title="Scanning electron microscopy of lattice organs in cyprids of the Rhizocephala Akentrogonida (Crustacea, Cirripedia)" type="journal article" year="1994">
Jensen
<emphasis id="B95CDA13FFBFD02BFD0FFE2A7B7D9ECB" box="[754,807,392,418]" italics="true" pageId="8" pageNumber="79">et al.</emphasis>
1994
</bibRefCitation>
) who removed the cyprid shell covering the first juvenile stage inside, when preparing them for SEM. Some of these were neolepadines but two were the first juvenile stage of an unusual thoracic cirriped. It was studied and dubbed “Jensen’s x-juvenile” by
<bibRefCitation id="EFB97BF0FFBFD02BFBA4FD807B039D33" author="HOeg J. T. &amp; Newman W. A." bookContentInfo="Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris" pageId="8" pageNumber="79" refId="ref6736" refString="HOeg J. T. &amp; Newman W. A. 1997. - Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga. Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris." title="Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga." type="proceedings" year="1997">HØeg &amp; Newman (1997)</bibRefCitation>
(
<figureCitation id="13131A84FFBFD02BFC95FDE27BF39D33" box="[872,937,576,602]" captionStart="FIG" captionStartId="8.[114,125,1597,1614]" captionTargetBox="[139,1204,771,1569]" captionTargetId="figure-24@8.[127,1202,745,1582]" captionTargetPageId="8" captionText="FIG. 5. — Jensen’s x-juvenile from the Lau Basin, Tonga (Høeg &amp; Newman 1997); A, viewed from the left dorso-lateral; B, left lateral; C, ventral sides, respectively (the shard of chitin extending from the aperture in B and C apparently left over from the last molt). It is a first juvenile of a thoracican cirriped and likely that of Imbricaverruca yamaguchii n. gen. and n. sp.; see Fig. 6 and text for explanation. Abbreviations: A1, first antenna; C, carina; “C”, incipient cirri; LS, left scutum; LT, left tergum; RS, right scutum; P, peduncle. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5391403" httpUri="https://zenodo.org/record/5391403/files/figure.png" pageId="8" pageNumber="79">Fig. 5</figureCitation>
). Its capitulum, supported by a peduncle of comparable length, is armored by the five primordial valves commonly found in juvenile thoracicans, except iblomorphs.
</paragraph>
<caption id="DF575689FFBFD02BFF8FF99F794399CD" ID-DOI="http://doi.org/10.5281/zenodo.5391403" ID-Zenodo-Dep="5391403" httpUri="https://zenodo.org/record/5391403/files/figure.png" pageId="8" pageNumber="79" startId="8.[114,125,1597,1614]" targetBox="[139,1204,771,1569]" targetPageId="8">
<paragraph id="8B970601FFBFD02BFF8FF99F794399CD" blockId="8.[114,1211,1597,1700]" pageId="8" pageNumber="79">
FIG. 5. — Jensen’s x-juvenile from the Lau Basin, Tonga (
<bibRefCitation id="EFB97BF0FFBFD02BFDB1F99F7B509927" author="HOeg J. T. &amp; Newman W. A." bookContentInfo="Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris" box="[588,778,1597,1614]" pageId="8" pageNumber="79" refId="ref6736" refString="HOeg J. T. &amp; Newman W. A. 1997. - Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga. Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris." title="Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga." type="proceedings" year="1997">Høeg &amp; Newman 1997</bibRefCitation>
);
<emphasis id="B95CDA13FFBFD02BFCEAF99F7B7E9927" bold="true" box="[791,804,1597,1614]" pageId="8" pageNumber="79">A</emphasis>
, viewed from the left dorso-lateral;
<emphasis id="B95CDA13FFBFD02BFBB1F99F7C039927" bold="true" box="[1100,1113,1597,1614]" pageId="8" pageNumber="79">B</emphasis>
, left lateral;
<emphasis id="B95CDA13FFBFD02BFF8FF9F178DA990D" bold="true" box="[114,128,1619,1636]" pageId="8" pageNumber="79">C</emphasis>
, ventral sides, respectively (the shard of chitin extending from the aperture in B and C apparently left over from the last molt). It is a first juvenile of a thoracican cirriped and likely that of
<taxonomicName id="4C287D82FFBFD02BFDC7F9CA7B429910" authority="Newman, 2000" authorityName="Newman" authorityYear="2000" box="[570,792,1640,1657]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="yamaguchii" status="n. gen. and n. sp.">
<emphasis id="B95CDA13FFBFD02BFDC7F9CA7B429910" box="[570,792,1640,1657]" italics="true" pageId="8" pageNumber="79">Imbricaverruca yamaguchii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F6768FFBFD02BFCE0F9CA7BEB9910" box="[797,945,1640,1657]" pageId="8" pageNumber="79" rank="species">n. gen. and n. sp.</taxonomicNameLabel>
; see Fig. 6 and text for explanation. Abbreviations:
<emphasis id="B95CDA13FFBFD02BFEE4F9DF797499EE" bold="true" box="[281,302,1658,1678]" pageId="8" pageNumber="79">
A
<superScript id="7C5DAB49FFBFD02BFEDBF9D8797499EE" attach="left" box="[294,302,1658,1671]" fontSize="5" pageId="8" pageNumber="79">1</superScript>
</emphasis>
, first antenna;
<emphasis id="B95CDA13FFBFD02BFE50F9DF79E199E7" bold="true" box="[429,443,1661,1678]" pageId="8" pageNumber="79">C</emphasis>
, carina;
<emphasis id="B95CDA13FFBFD02BFDF9F9DF7A7999E7" bold="true" box="[516,547,1661,1678]" pageId="8" pageNumber="79">“C”</emphasis>
, incipient cirri;
<emphasis id="B95CDA13FFBFD02BFD59F9DF7AE199E7" bold="true" box="[676,699,1661,1678]" pageId="8" pageNumber="79">LS</emphasis>
, left scutum;
<emphasis id="B95CDA13FFBFD02BFCD2F9DF7B1F99E7" bold="true" box="[815,837,1661,1678]" pageId="8" pageNumber="79">LT</emphasis>
, left tergum;
<emphasis id="B95CDA13FFBFD02BFC4AF9DF7B8B99E7" bold="true" box="[951,977,1661,1678]" pageId="8" pageNumber="79">RS</emphasis>
, right scutum;
<emphasis id="B95CDA13FFBFD02BFBADF9DF7C0699E7" bold="true" box="[1104,1116,1661,1678]" pageId="8" pageNumber="79">P</emphasis>
, peduncle. Scale bars: 100 µm.
</paragraph>
</caption>
<caption id="DF575689FFBED02AFF8FFCEC78939CDD" ID-DOI="http://doi.org/10.5281/zenodo.5391405" ID-Zenodo-Dep="5391405" httpUri="https://zenodo.org/record/5391405/files/figure.png" pageId="9" pageNumber="80" startId="9.[114,125,846,863]" targetBox="[297,1063,203,833]" targetPageId="9">
<paragraph id="8B970601FFBED02AFF8FFCEC78939CDD" blockId="9.[114,1212,846,948]" pageId="9" pageNumber="80">
FIG. 6. — Comparison between the primordial plates of the earliest juvenile stages of the principal suborders of thoracic
<taxonomicName id="4C287D82FFBED02AFB99FCEC7CED9C36" authorityName="Burmeister" authorityYear="1834" box="[1124,1207,846,863]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="73" phylum="Arthropoda" rank="subClass" subClass="Cirripedia">Cirripedia</taxonomicName>
;
<emphasis id="B95CDA13FFBED02AFF8FFCC178259C1D" bold="true" box="[114,127,867,884]" pageId="9" pageNumber="80">A</emphasis>
, Lepadomorpha,
<taxonomicName id="4C287D82FFBED02AFEE8FCC1791D9C1D" box="[277,327,867,884]" class="Maxillopoda" family="Lepadidae" genus="Lepas" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFBED02AFEE8FCC1791D9C1D" box="[277,327,867,884]" italics="true" pageId="9" pageNumber="80">Lepas</emphasis>
</taxonomicName>
(from
<bibRefCitation id="EFB97BF0FFBED02AFE82FCC17A719C1D" author="Newman W. A." box="[383,555,867,884]" journalOrPublisher="San Diego Society of Natural History Transactions" pageId="9" pageNumber="80" pagination="153 - 167" part="19" refId="ref6965" refString="Newman W. A. 1979. - A new scalpellid (Cirripedia); a Mesozoic relic living near an abyssal hydrothermal spring. San Diego Society of Natural History Transactions 19 (11): 153 - 167." title="A new scalpellid (Cirripedia); a Mesozoic relic living near an abyssal hydrothermal spring" type="journal article" year="1979">
Newman
<emphasis id="B95CDA13FFBED02AFE2DFCC179A09C1D" box="[464,506,867,884]" italics="true" pageId="9" pageNumber="80">et al.</emphasis>
1979
</bibRefCitation>
: fig. 87);
<emphasis id="B95CDA13FFBED02AFD84FCC17ADC9C1D" bold="true" box="[633,646,867,884]" pageId="9" pageNumber="80">B</emphasis>
,
<taxonomicName id="4C287D82FFBED02AFD6FFCC17B789C1D" box="[658,802,867,884]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scalpellomorpha" pageId="9" pageNumber="80" phylum="Arthropoda" rank="order">Scalpellomorpha</taxonomicName>
, composite of
<taxonomicName id="4C287D82FFBED02AFC58FCC17BA99C1D" authorityName="Newman" authorityYear="1979" box="[933,1011,867,884]" class="Maxillopoda" family="Eolepadidae" genus="Neolepas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFBED02AFC58FCC17BA99C1D" box="[933,1011,867,884]" italics="true" pageId="9" pageNumber="80">Neolepas</emphasis>
</taxonomicName>
and
<taxonomicName id="4C287D82FFBED02AFBE2FCC17C349C1D" authorityName="Leach" authorityYear="1817" box="[1055,1134,867,884]" class="Maxillopoda" family="Pollicipedidae" genus="Pollicipes" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFBED02AFBE2FCC17C349C1D" box="[1055,1134,867,884]" italics="true" pageId="9" pageNumber="80">Pollicipes</emphasis>
</taxonomicName>
(respectively from Newman, in prep. and
<bibRefCitation id="EFB97BF0FFBED02AFE6CFCDB79AF9CE3" author="Broch H." box="[401,501,889,906]" journalOrPublisher="Videnskabelige meddelelser fra Dansk Naturhistorisk Forening i Kobenhavn" pageId="9" pageNumber="80" pagination="215 - 358" part="73" refId="ref6293" refString="Broch H. 1922. - Studies on Pacific cirripeds, in Papers from Dr Th. Mortensen's Pacific Expedition 1914 - 1916. X. Videnskabelige meddelelser fra Dansk Naturhistorisk Forening i Kobenhavn 73: 215 - 358." title="Studies on Pacific cirripeds, in Papers from Dr Th. Mortensen's Pacific Expedition 1914 - 1916. X" type="journal article" year="1922">Broch 1922</bibRefCitation>
);
<emphasis id="B95CDA13FFBED02AFDFEFCDB7A4B9CE3" bold="true" box="[515,529,889,906]" pageId="9" pageNumber="80">C</emphasis>
,
<taxonomicName id="4C287D82FFBED02AFDE1FCDB7A269CE3" baseAuthorityName="Newman &amp; Hessler" baseAuthorityYear="1989" box="[540,636,889,906]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="9" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFBED02AFDE1FCDB7A269CE3" box="[540,636,889,906]" italics="true" pageId="9" pageNumber="80">Neoverruca</emphasis>
</taxonomicName>
(reconstructed from
<bibRefCitation id="EFB97BF0FFBED02AFCD0FCDB7BF09CE3" author="Newman W. A." box="[813,938,889,906]" journalOrPublisher="Bulletin of Marine Science" pageId="9" pageNumber="80" pagination="467 - 477" part="45" refId="ref7144" refString="Newman W. A. 1989. - Juvenile ontogeny and metamorphosis in the most primitive living sessile barnacle, Neoverruca, from an abyssal hydrothermal spring. Bulletin of Marine Science 45 (2): 467 - 477." title="Juvenile ontogeny and metamorphosis in the most primitive living sessile barnacle, Neoverruca, from an abyssal hydrothermal spring" type="journal article" year="1989">Newman 1989</bibRefCitation>
);
<emphasis id="B95CDA13FFBED02AFC4AFCDB7B9F9CE3" bold="true" box="[951,965,889,906]" pageId="9" pageNumber="80">D</emphasis>
, x-juvenile (
<bibRefCitation id="EFB97BF0FFBED02AFBD6FCDB78C69CF6" author="HOeg J. T. &amp; Newman W. A." bookContentInfo="Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris" pageId="9" pageNumber="80" refId="ref6736" refString="HOeg J. T. &amp; Newman W. A. 1997. - Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga. Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris." title="Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga." type="proceedings" year="1997">Høeg &amp; Newman 1997</bibRefCitation>
&amp; herein);
<emphasis id="B95CDA13FFBED02AFF07FC2C795C9CF6" bold="true" box="[250,262,910,927]" pageId="9" pageNumber="80">E</emphasis>
,
<emphasis id="B95CDA13FFBED02AFEEFFC2C79479CF6" bold="true" box="[274,285,910,927]" pageId="9" pageNumber="80">F</emphasis>
,
<taxonomicName id="4C287D82FFBED02AFED4FC2C79339CF6" authorityName="Schumacher" authorityYear="1817" box="[297,361,910,927]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="9" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFBED02AFED4FC2C79339CF6" box="[297,361,910,927]" italics="true" pageId="9" pageNumber="80">Verruca</emphasis>
</taxonomicName>
and
<taxonomicName id="4C287D82FFBED02AFE6AFC2C7A589CF6" authorityName="Pilsbry" authorityYear="1916" box="[407,514,910,927]" class="Maxillopoda" family="Archaeobalanidae" genus="Semibalanus" kingdom="Animalia" order="Sessilia" pageId="9" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFBED02AFE6AFC2C7A589CF6" box="[407,514,910,927]" italics="true" pageId="9" pageNumber="80">Semibalanus</emphasis>
</taxonomicName>
(respectively from
<bibRefCitation id="EFB97BF0FFBED02AFD5BFC2C7B669CF6" author="Runnstrom S." box="[678,828,910,927]" journalOrPublisher="Bergens Museums Arbok" pageId="9" pageNumber="80" pagination="1 - 46" part="5" refId="ref7493" refString="Runnstrom S. 1925. - Zur Biologie und Entwicklung von Balanus balanoides (Linn.). Bergens Museums Arbok 5: 1 - 46." title="Zur Biologie und Entwicklung von Balanus balanoides (Linn.)" type="journal article" year="1925">Runnström, 1925</bibRefCitation>
,
<bibRefCitation id="EFB97BF0FFBED02AFCB8FC2C7B289CF6" author="Runnstrom S." box="[837,882,910,927]" journalOrPublisher="Bergens Museums Arbok" pageId="9" pageNumber="80" pagination="1 - 10" part="7" refId="ref7520" refString="Runnstrom S. 1927. - Uber die Plattenentwicklung von Verruca stroemia. Bergens Museums Arbok 7: 1 - 10." title="Uber die Plattenentwicklung von Verruca stroemia" type="journal article" year="1927">1927</bibRefCitation>
). Abbreviations:
<emphasis id="B95CDA13FFBED02AFC02FC2C7C519CF6" bold="true" box="[1023,1035,910,927]" pageId="9" pageNumber="80">S</emphasis>
, scutum;
<emphasis id="B95CDA13FFBED02AFB9FFC2C7C379CF6" bold="true" box="[1122,1133,910,927]" pageId="9" pageNumber="80">T</emphasis>
, tergum;
<emphasis id="B95CDA13FFBED02AFF8FFC0178DA9CDD" bold="true" box="[114,128,931,948]" pageId="9" pageNumber="80">C</emphasis>
, carina).
</paragraph>
</caption>
<caption id="DF575689FFBED02AFF8CFBDD7AFE9BCC" pageId="9" pageNumber="80" startId="9.[113,124,1151,1168]" targetBox="[113,1171,1240,1689]" targetIsTable="true" targetPageId="9">
<paragraph id="8B970601FFBED02AFF8CFBDD7AFE9BCC" blockId="9.[113,1210,1151,1189]" pageId="9" pageNumber="80">
TABLE 2. — Comparison between the primordial plates of the earliest juvenile stages of the principal suborders of thoracic
<taxonomicName id="4C287D82FFBED02AFB94FBDD7CE09BF9" authorityName="Burmeister" authorityYear="1834" box="[1129,1210,1151,1168]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="73" phylum="Arthropoda" rank="subClass" subClass="Cirripedia">Cirripedia</taxonomicName>
(See Fig. 6 for corresponding figures and text for a full explanation).
</paragraph>
</caption>
<paragraph id="8B970601FFBED02AFF8CFB7A7C7A99E8" pageId="9" pageNumber="80">
<table id="F928F4A1FFBE2FDCFF8CFB7A7CC999F0" box="[113,1171,1240,1689]" gridcols="4" gridrows="8" pageId="9" pageNumber="80">
<tr id="35180443FFBE2FDCFF8CFB7A7CC99A6D" box="[113,1171,1240,1284]" gridrow="0" pageId="9" pageNumber="80">
<th id="76C96D3FFFBE2FDCFF8CFB7A793D9A6D" box="[113,359,1240,1284]" gridcol="0" gridrow="0" pageId="9" pageNumber="80">
<emphasis id="B95CDA13FFBED02AFF8CFB7A79479B85" bold="true" box="[113,285,1240,1260]" pageId="9" pageNumber="80">Taxon/Character</emphasis>
</th>
<th id="76C96D3FFFBE2FDCFE87FB7A7ADD9A6D" box="[378,647,1240,1284]" gridcol="1" gridrow="0" pageId="9" pageNumber="80">
<emphasis id="B95CDA13FFBED02AFE87FB7A7A659B85" bold="true" box="[378,575,1240,1260]" pageId="9" pageNumber="80">Carinal proportions</emphasis>
</th>
<th id="76C96D3FFFBE2FDCFD6FFB7A7BD49A6D" box="[658,910,1240,1284]" gridcol="2" gridrow="0" pageId="9" pageNumber="80">
<emphasis id="B95CDA13FFBED02AFD6FFB7A7B169B85" bold="true" box="[658,844,1240,1260]" pageId="9" pageNumber="80">Carinal of position</emphasis>
</th>
<th id="76C96D3FFFBE2FDCFC54FB7A7CC99A6D" box="[937,1171,1240,1284]" gridcol="3" gridrow="0" pageId="9" pageNumber="80">
<emphasis id="B95CDA13FFBED02AFC54FB7A7C7F9A6D" bold="true" pageId="9" pageNumber="80">Terga and scuta of each side</emphasis>
</th>
</tr>
<tr id="35180443FFBE2FDCFF8CFABC7CC99A23" box="[113,1171,1310,1354]" gridrow="1" pageId="9" pageNumber="80">
<th id="76C96D3FFFBE2FDCFF8CFABC793D9A23" box="[113,359,1310,1354]" gridcol="0" gridrow="1" pageId="9" pageNumber="80">Lepadomorpha (Fig. 6A)</th>
<td id="76C96D3FFFBE2FDCFE87FABC7ADD9A23" box="[378,647,1310,1354]" gridcol="1" gridrow="1" pageId="9" pageNumber="80">Higher than wide and longer than terga</td>
<td id="76C96D3FFFBE2FDCFD6FFABC7BD49A23" box="[658,910,1310,1354]" gridcol="2" gridrow="1" pageId="9" pageNumber="80">Extending up between terga</td>
<td id="76C96D3FFFBE2FDCFC54FABC7CC99A23" box="[937,1171,1310,1354]" gridcol="3" gridrow="1" pageId="9" pageNumber="80">Symmetrical</td>
</tr>
<tr id="35180443FFBE2FDCFF8CFAF07CC99A17" box="[113,1171,1362,1406]" gridrow="2" pageId="9" pageNumber="80">
<th id="76C96D3FFFBE2FDCFF8CFAF0793D9A17" box="[113,359,1362,1406]" gridcol="0" gridrow="2" pageId="9" pageNumber="80">
<taxonomicName id="4C287D82FFBED02AFF8CFAF079489A0F" box="[113,274,1362,1382]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scalpellomorpha" pageId="9" pageNumber="80" phylum="Arthropoda" rank="order">Scalpellomorpha</taxonomicName>
(Fig. 6B)
</th>
<td id="76C96D3FFFBE2FDCFE87FAF07ADD9A17" box="[378,647,1362,1406]" gridcol="1" gridrow="2" pageId="9" pageNumber="80">Higher than wide and nearly as long as terga</td>
<td id="76C96D3FFFBE2FDCFD6FFAF07BD49A17" box="[658,910,1362,1406]" gridcol="2" gridrow="2" pageId="9" pageNumber="80">Extending up between terga</td>
<td id="76C96D3FFFBE2FDCFC54FAF07CC99A17" box="[937,1171,1362,1406]" gridcol="3" gridrow="2" pageId="9" pageNumber="80">Symmetrical</td>
</tr>
<tr id="35180443FFBE2FDCFF8CFA247CC99AF3" box="[113,1171,1414,1434]" gridrow="3" pageId="9" pageNumber="80">
<th id="76C96D3FFFBE2FDCFF8CFA247CC99AF3" box="[113,1171,1414,1434]" colspan="4" colspanRight="3" gridcol="0" gridrow="3" pageId="9" pageNumber="80">
<taxonomicName id="4C287D82FFBED02AFF8CFA24795F9AF3" authorityName="Pilsbry" authorityYear="1916" box="[113,261,1414,1434]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="9" pageNumber="73" phylum="Arthropoda" rank="subOrder" subOrder="Verrucomorpha">Verrucomorpha</taxonomicName>
</th>
</tr>
<tr id="35180443FFBE2FDCFF8CFA007CC99AA7" box="[113,1171,1442,1486]" gridrow="4" pageId="9" pageNumber="80">
<th id="76C96D3FFFBE2FDCFF8CFA00793D9AA7" box="[113,359,1442,1486]" gridcol="0" gridrow="4" pageId="9" pageNumber="80">
<taxonomicName id="4C287D82FFBED02AFF8CFA0078A69ADF" box="[113,252,1442,1462]" class="Maxillopoda" family="Neoverrucidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="9" pageNumber="80" phylum="Arthropoda" rank="family">Neoverrucidae</taxonomicName>
(Fig. 6C)
</th>
<td id="76C96D3FFFBE2FDCFE87FA007ADD9AA7" box="[378,647,1442,1486]" gridcol="1" gridrow="4" pageId="9" pageNumber="80">Higher than wide and shorter than terga</td>
<td id="76C96D3FFFBE2FDCFD6FFA007BD49AA7" box="[658,910,1442,1486]" gridcol="2" gridrow="4" pageId="9" pageNumber="80">Extending up between terga</td>
<td id="76C96D3FFFBE2FDCFC54FA007CC99AA7" box="[937,1171,1442,1486]" gridcol="3" gridrow="4" pageId="9" pageNumber="80">Initially symmetrical</td>
</tr>
<tr id="35180443FFBE2FDCFF8CFA777CC99970" box="[113,1171,1493,1561]" gridrow="5" pageId="9" pageNumber="80">
<th id="76C96D3FFFBE2FDCFF8CFA77793D9970" box="[113,359,1493,1561]" gridcol="0" gridrow="5" pageId="9" pageNumber="80">X-juvenile (Fig. 6D)</th>
<td id="76C96D3FFFBE2FDCFE87FA777ADD9970" box="[378,647,1493,1561]" gridcol="1" gridrow="5" pageId="9" pageNumber="80">Wider than high, shorter than terga and perhaps displaced to one side</td>
<td id="76C96D3FFFBE2FDCFD6FFA777BD49970" box="[658,910,1493,1561]" gridcol="2" gridrow="5" pageId="9" pageNumber="80">Not extending up between terga</td>
<td id="76C96D3FFFBE2FDCFC54FA777CC99970" box="[937,1171,1493,1561]" gridcol="3" gridrow="5" pageId="9" pageNumber="80">Scuta if not terga initially symmetrical</td>
</tr>
<tr id="35180443FFBE2FDCFF8CF9837CC9990C" box="[113,1171,1569,1637]" gridrow="6" pageId="9" pageNumber="80">
<th id="76C96D3FFFBE2FDCFF8CF983793D990C" box="[113,359,1569,1637]" gridcol="0" gridrow="6" pageId="9" pageNumber="80">
<taxonomicName id="4C287D82FFBED02AFF8CF983788D995C" authorityName="Darwin" authorityYear="1854" box="[113,215,1569,1589]" class="Maxillopoda" family="Verrucidae" kingdom="Animalia" order="Sessilia" pageId="9" pageNumber="80" phylum="Arthropoda" rank="family">Verrucidae</taxonomicName>
(Fig. 6E)
</th>
<td id="76C96D3FFFBE2FDCFE87F9837ADD990C" box="[378,647,1569,1637]" gridcol="1" gridrow="6" pageId="9" pageNumber="80">About as wide as high, shorter than terga and displaced to one side</td>
<td id="76C96D3FFFBE2FDCFD6FF9837BD4990C" box="[658,910,1569,1637]" gridcol="2" gridrow="6" pageId="9" pageNumber="80">Not extending up between terga</td>
<td id="76C96D3FFFBE2FDCFC54F9837CC9990C" box="[937,1171,1569,1637]" gridcol="3" gridrow="6" pageId="9" pageNumber="80">Distinctly asymmetrical</td>
</tr>
<tr id="35180443FFBE2FDCFF8CF9CF7CC999F0" box="[113,1171,1645,1689]" gridrow="7" pageId="9" pageNumber="80">
<th id="76C96D3FFFBE2FDCFF8CF9CF793D99F0" box="[113,359,1645,1689]" gridcol="0" gridrow="7" pageId="9" pageNumber="80">
<taxonomicName id="4C287D82FFBED02AFF8CF9CF78A799E8" box="[113,253,1645,1665]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Balanomorpha" pageId="9" pageNumber="80" phylum="Arthropoda" rank="order">Balanomorpha</taxonomicName>
(Fig. 6F)
</th>
<td id="76C96D3FFFBE2FDCFE87F9CF7ADD99F0" box="[378,647,1645,1689]" gridcol="1" gridrow="7" pageId="9" pageNumber="80">Higher than wide</td>
<td id="76C96D3FFFBE2FDCFD6FF9CF7BD499F0" box="[658,910,1645,1689]" gridcol="2" gridrow="7" pageId="9" pageNumber="80">Not extending up between terga</td>
<td id="76C96D3FFFBE2FDCFC54F9CF7CC999F0" box="[937,1171,1645,1689]" gridcol="3" gridrow="7" pageId="9" pageNumber="80">Symmetrical</td>
</tr>
</table>
</paragraph>
<paragraph id="8B970601FFBDD029FF8FFF7279189E2F" blockId="10.[114,647,208,1154]" pageId="10" pageNumber="81">
The position and shape of the carinal primordium and a slight asymmetry between the scuta suggested the juvenile was a verrucomorph (
<bibRefCitation id="EFB97BF0FFBDD029FDB4FEAC796F9E2F" author="HOeg J. T. &amp; Newman W. A." bookContentInfo="Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris" pageId="10" pageNumber="81" refId="ref6736" refString="HOeg J. T. &amp; Newman W. A. 1997. - Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga. Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris." title="Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga." type="proceedings" year="1997">HØeg &amp; Newman 1997</bibRefCitation>
).
</paragraph>
<paragraph id="8B970601FFBDD029FF8FFEE979B09BEB" blockId="10.[114,647,208,1154]" pageId="10" pageNumber="81">
The early juveniles of
<taxonomicName id="4C287D82FFBDD029FE7DFEE878E99EED" authority="(Newman, 1989)" baseAuthorityName="Newman &amp; Hessler" baseAuthorityYear="1989" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="B95CDA13FFBDD029FE7DFEE87A5A9E0A" box="[384,512,330,355]" italics="true" pageId="10" pageNumber="81">Neoverruca</emphasis>
(
<bibRefCitation id="EFB97BF0FFBDD029FDEEFEE978F19EED" author="Newman W. A." journalOrPublisher="Bulletin of Marine Science" pageId="10" pageNumber="81" pagination="467 - 477" part="45" refId="ref7144" refString="Newman W. A. 1989. - Juvenile ontogeny and metamorphosis in the most primitive living sessile barnacle, Neoverruca, from an abyssal hydrothermal spring. Bulletin of Marine Science 45 (2): 467 - 477." title="Juvenile ontogeny and metamorphosis in the most primitive living sessile barnacle, Neoverruca, from an abyssal hydrothermal spring" type="journal article" year="1989">Newman, 1989</bibRefCitation>
)
</taxonomicName>
can be distinguished from the x-juvenile in being more generalized; e.g. in having a carinal primordium that is higher than wide extending up between the terga (
<figureCitation id="13131A84FFBDD029FE8FFE6479ED9E89" box="[370,439,454,480]" captionStart="FIG" captionStartId="9.[114,125,846,863]" captionTargetBox="[297,1063,203,833]" captionTargetId="figure-268@9.[363,983,241,828]" captionTargetPageId="9" captionText="FIG. 6. — Comparison between the primordial plates of the earliest juvenile stages of the principal suborders of thoracic Cirripedia; A, Lepadomorpha, Lepas (from Newman et al. 1979: fig. 87); B, Scalpellomorpha, composite of Neolepas and Pollicipes (respectively from Newman, in prep. and Broch 1922); C, Neoverruca (reconstructed from Newman 1989); D, x-juvenile (Høeg &amp; Newman 1997 &amp; herein); E, F, Verruca and Semibalanus (respectively from Runnström, 1925, 1927). Abbreviations: S, scutum; T, tergum; C, carina)." figureDoi="http://doi.org/10.5281/zenodo.5391405" httpUri="https://zenodo.org/record/5391405/files/figure.png" pageId="10" pageNumber="81">Fig. 6</figureCitation>
; cf.
<tableCitation id="C6AA33BAFFBDD029FE16FE647A189E89" box="[491,578,454,480]" captionStart="TABLE" captionStartId="9.[113,124,1151,1168]" captionTargetPageId="9" captionText="TABLE 2. — Comparison between the primordial plates of the earliest juvenile stages of the principal suborders of thoracic Cirripedia (See Fig. 6 for corresponding figures and text for a full explanation)." pageId="10" pageNumber="81">Table 2</tableCitation>
). The only first juvenile of a verrucid known is that of
<taxonomicName id="4C287D82FFBDD029FF8FFDA079B99D74" authority="Muller, 1776" authorityName="Muller" authorityYear="1776" box="[114,483,514,541]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="species" species="stroemia">
<emphasis id="B95CDA13FFBDD029FF8FFDA079629D72" box="[114,312,514,539]" italics="true" pageId="10" pageNumber="81">Verruca stroemia</emphasis>
Müller, 1776
</taxonomicName>
described by
<bibRefCitation id="EFB97BF0FFBDD029FF8FFD8379659D55" author="Runnstrom S." box="[114,319,545,572]" journalOrPublisher="Bergens Museums Arbok" pageId="10" pageNumber="81" pagination="1 - 10" part="7" refId="ref7520" refString="Runnstrom S. 1927. - Uber die Plattenentwicklung von Verruca stroemia. Bergens Museums Arbok 7: 1 - 10." title="Uber die Plattenentwicklung von Verruca stroemia" type="journal article" year="1927">Runnström (1927)</bibRefCitation>
and it differs from the x-juvenile in the scutal primordia being grossly unequal rather than nearly equal in size. Therefore it was concluded the x-juvenile could represent 1) a verrucid more primitive than
<taxonomicName id="4C287D82FFBDD029FE64FD3E7A569DDC" authorityName="Muller" authorityYear="1776" box="[409,524,668,693]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="species" species="stroemia">
<emphasis id="B95CDA13FFBDD029FE64FD3E7A569DDC" box="[409,524,668,693]" italics="true" pageId="10" pageNumber="81">V. stroemia</emphasis>
</taxonomicName>
such as an
<emphasis id="B95CDA13FFBDD029FF8FFD1978B59DBC" box="[114,239,699,725]" italics="true" pageId="10" pageNumber="81">
<taxonomicName id="4C287D82FFBDD029FF8FFD1978B19DBC" authorityName="Pilsbry" authorityYear="1916" box="[114,235,699,725]" class="Maxillopoda" family="Verrucidae" genus="Altiverruca" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="genus">Altiverruca</taxonomicName>
,
</emphasis>
2) an advanced neoverrucid such as the one described from Lau herein, or 3) a yetto-be discovered population of the presumably extinct proverrucid, and it was thought that, because of these uncertainties, determining the affinities of Jensen’s x-juvenile would require further sampling of the Lau Basin fauna (
<bibRefCitation id="EFB97BF0FFBDD029FDDAFCD179409CC5" author="HOeg J. T. &amp; Newman W. A." bookContentInfo="Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris" pageId="10" pageNumber="81" refId="ref6736" refString="HOeg J. T. &amp; Newman W. A. 1997. - Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga. Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris." title="Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga." type="proceedings" year="1997">HØeg &amp; Newman 1997</bibRefCitation>
). However, the Lau neoverrucid, now known as
<emphasis id="B95CDA13FFBDD029FEDFFC12798B9CA3" box="[290,465,944,970]" italics="true" pageId="10" pageNumber="81">
<taxonomicName id="4C287D82FFBDD029FEDFFC1279979CA3" authorityName="Newman" authorityYear="2000" box="[290,461,944,970]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="genus">Imbricaverruca</taxonomicName>
,
</emphasis>
turned out not only to be more highly evolved than
<emphasis id="B95CDA13FFBDD029FDF7FC6C7ADD9C8E" box="[522,647,974,999]" italics="true" pageId="10" pageNumber="81">
<taxonomicName id="4C287D82FFBDD029FDF7FC6C7AD89C8E" baseAuthorityName="Newman &amp; Hessler" baseAuthorityYear="1989" box="[522,642,974,999]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="genus">Neoverruca</taxonomicName>
,
</emphasis>
but the adult has an autapomorphy, a carina that is much wider than high, which is the most notable characteristic of the x-juvenile. Therefore it seems highly likely that Jensen’s x-juvenile is the first juvenile of
<emphasis id="B95CDA13FFBDD029FEB9FBCA79B09BEB" box="[324,490,1128,1154]" italics="true" pageId="10" pageNumber="81">
<taxonomicName id="4C287D82FFBDD029FEB9FBCA79BC9BEB" authorityName="Newman" authorityYear="2000" box="[324,486,1128,1154]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="genus">Imbricaverruca</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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