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<document id="A4C171757C3B3612B0600F51DDE33252" ID-DOI="10.5281/zenodo.5391395" ID-ISSN="1638-9387" ID-Zenodo-Dep="5391395" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1630314607455" checkinUser="marcus" docAuthor="Newman, William A." docDate="2000" docId="0381B717FFB4D020FF3EFF7278FD9EA8" docLanguage="en" docName="Zoosystema.2000.22.1.71-84.pdf" docOrigin="Zoosystema 22 (1)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Neoverrucidae Newman" docType="treatment" docVersion="2" lastPageNumber="74" masterDocId="FFB8CF6FFFB7D023FFFDFFA2785A9F69" masterDocTitle="A new genus and species of barnacle (Cirripedia, Verrucomorpha) associated with vents of the Lau Back-Arc Basin: its gross morphology, inferred first juvenile stage and affinities" masterLastPageNumber="84" masterPageNumber="71" pageNumber="74" updateTime="1729878602252" updateUser="juliana" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="289EC9424FF69D01C6415F45A9CA1734">A new genus and species of barnacle (Cirripedia, Verrucomorpha) associated with vents of the Lau Back-Arc Basin: its gross morphology, inferred first juvenile stage and affinities</mods:title>
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<mods:namePart id="77DF1F09CC4A3BEE16EAA0B80DFEC05F">Newman, William A.</mods:namePart>
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<mods:affiliation id="2BF80C58D8EDB18C4C4D00A13DEF4565">Scripps Institution of Oceanography, University of California San Diego, La Jolla, CA 92093 - 0202 (USA) wnewman @ ucsd. edu</mods:affiliation>
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<mods:nameIdentifier id="EEFF16C406A0E7C137978233782FB8E4" type="email">wnewman@ucsd.edu</mods:nameIdentifier>
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<mods:title id="4898BAFC3E4B577303A5A9FE6C88DBE6">Zoosystema</mods:title>
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<heading id="D0DFB16DFFB4D020FF3EFF727A139E60" centered="true" fontSize="11" level="2" pageId="3" pageNumber="74" reason="2">
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Family
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<taxonomicName id="4C287D82FFB4D020FEE9FF727A6E9F83" authority="Newman" authorityName="Newman" box="[276,564,208,234]" class="Maxillopoda" family="Neoverrucidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="family">NEOVERRUCIDAE Newman</taxonomicName>
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<emphasis id="B95CDA13FFB4D020FF53FF4C789E9E6E" box="[174,196,238,263]" italics="true" pageId="3" pageNumber="74">in</emphasis>
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<bibRefCitation id="EFB97BF0FFB4D020FF36FF4C79BC9E60" author="Newman W. A. & Hessler R. R." box="[203,486,238,265]" journalOrPublisher="San Diego Society of Natural History Transactions" pageId="3" pageNumber="74" pagination="259 - 273" part="21" refId="ref7233" refString="Newman W. A. & Hessler R. R. 1989. - A new abyssal hydrothermal verrucomorphan (Cirripedia; Sessilia): The most primitive living sessile barnacle. San Diego Society of Natural History Transactions 21 (16): 259 - 273." title="A new abyssal hydrothermal verrucomorphan (Cirripedia; Sessilia): The most primitive living sessile barnacle" type="journal article" year="1989">Newman & Hessler, 1989</bibRefCitation>
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(Recent)
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<paragraph id="8B970601FFB4D020FF8CFE8B78FD9EA8" blockId="3.[113,647,297,448]" pageId="3" pageNumber="74">
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DIAGNOSIS. — Distinguished from the extinct
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<taxonomicName id="4C287D82FFB4D020FF8CFEE178A09E32" authorityName="Newman, 1989" authorityYear="1989" box="[113,250,323,347]" class="Maxillopoda" family="Proverrucidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="family">Proverrucidae</taxonomicName>
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and extant
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<taxonomicName id="4C287D82FFB4D020FE88FEE179BB9E32" authorityName="Darwin" authorityYear="1854" box="[373,481,323,347]" class="Maxillopoda" family="Verrucidae" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="family">Verrucidae</taxonomicName>
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in having a primary wall elevated above the substratum by basal whorls of imbricating plates and an operculum including a median latus at least in juveniles (
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<bibRefCitation id="EFB97BF0FFB4D020FE11FE2D78C19EA8" author="Newman W. A." journalOrPublisher="Bulletin of Marine Science" pageId="3" pageNumber="74" pagination="467 - 477" part="45" refId="ref7144" refString="Newman W. A. 1989. - Juvenile ontogeny and metamorphosis in the most primitive living sessile barnacle, Neoverruca, from an abyssal hydrothermal spring. Bulletin of Marine Science 45 (2): 467 - 477." title="Juvenile ontogeny and metamorphosis in the most primitive living sessile barnacle, Neoverruca, from an abyssal hydrothermal spring" type="journal article" year="1989">Newman 1989: 268</bibRefCitation>
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).
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<mods:affiliation id="1D5431326D704F6581380072609819BF">Scripps Institution of Oceanography, University of California San Diego, La Jolla, CA 92093 - 0202 (USA) wnewman @ ucsd. edu</mods:affiliation>
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<taxonomicName id="4C287D82FFB4D020FD04FF727C7E9F83" authority="Newman, 2000" authorityName="Newman" authorityYear="2000" box="[761,1060,208,234]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="yamaguchii" status="sp. nov.">
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<emphasis id="B95CDA13FFB4D020FD04FF727C7E9F83" bold="true" box="[761,1060,208,234]" italics="true" pageId="3" pageNumber="74">Imbricaverruca yamaguchii</emphasis>
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<taxonomicNameLabel id="A26F6768FFB4D020FBD6FF727C3C9F83" box="[1067,1126,208,234]" pageId="3" pageNumber="74" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8B970601FFB4D020FCD3FF537C6B9E62" blockId="3.[761,1126,208,267]" box="[814,1073,241,267]" pageId="3" pageNumber="74">
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(
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<figureCitation id="13131A84FFB4D020FCC8FF537BD09E62" box="[821,906,241,267]" captionStart="FIG" captionStartId="2.[114,125,799,816]" captionTargetBox="[262,1053,208,766]" captionTargetId="figure-307@2.[248,1067,202,776]" captionTargetPageId="2" captionText="FIG. 1. — A, Imbricaverruca yamaguchii n. gen. and n. sp. (holotype, MNHN Ci2710); B, Verruca s.l., both viewed from above with the right scutum and tergum forming the operculum.Note that in the former the four plated wall appears to be covered largely by imbricating plates and the operculum includes a large median latus,characters that readily distinguish it from Neoverruca. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5391397" httpUri="https://zenodo.org/record/5391397/files/figure.png" pageId="3" pageNumber="74">Figs 1A</figureCitation>
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;
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<figureCitation id="13131A84FFB4D020FC6BFF537BFC9E62" box="[918,934,241,267]" captionStart="FIG" captionStartId="4.[114,125,1384,1401]" captionTargetBox="[213,1067,193,1353]" captionTargetId="figure-91@4.[208,1091,186,1354]" captionTargetPageId="4" captionText="FIG. 2. — Plates of Imbricaverruca yamaguchii gen. et n. sp. (paratype, MNHN Ci2711); A, B, viewed from above and from the rostral end, respectively; A, note the operculum, comprising the plates of the right side (MS-L-MT), includes a large median latus; B, note the slightly open aperture to the mantle cavity between the occludent margins of the movable (MS & MT) and the fixed (FS & FT) scutum and tergum, respectively; C-M, variously disarticulated hard parts; C, interior view of wall (R-C-FS-FT) and operculum (MS-L- MT) with all of the imbricating plates, except the uppermost of the three principle lateral tiers of the right side (rl1-l1-cl1), stripped away [an approximation of the rostrum (R), which did not survive dissection, is indicated by the dashed line]; D, E, movable scutum (MS); external and internal views respectively; scutal adductor muscle, inserting from just below to up under the apical concavity in Fig. 2E, extends to its origin on the fixed scutum (FS, Fig. 2G; the relationship of MS to FS can be seen in Fig 2C); F, G, fixed scutum (FS); basal and internal views, respectively. Dashed line in G indicates origin of scutal adductor muscle but no scar is evident. The relationship of the two protuberances (best seen near the tergal margin in Fig. 2G) to the fixed tergum (FT, Fig. 2J, K), was not resolved; H, I, movable tergum (MT); external and internal views respectively (external corrosion in bleach and cleaning inadvertently obliterated growth line ornamentation in H); J, K, fixed tergum (FT); external and internal views respectively; L, M, carina from below and above respectively (the rostrum did not survive dissection and cleaning). Abbreviations: C, carina; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral plates forming three tiers each four plates high, respectively; FS, FT, fixed scutum and tergum; L, median latus; MS, MT, movable scutum and tergum; R, rostrum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391399" httpUri="https://zenodo.org/record/5391399/files/figure.png" pageId="3" pageNumber="74">2</figureCitation>
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;
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<figureCitation id="13131A84FFB4D020FC4FFF537B9B9E62" box="[946,961,241,267]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="3" pageNumber="74">3</figureCitation>
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;
|
||||
<figureCitation id="13131A84FFB4D020FC33FF537BAB9E62" box="[974,1009,241,267]" captionStart="Fig" captionStartId="7.[113,141,1469,1486]" captionTargetBox="[112,1165,215,1445]" captionTargetId="figure-22@7.[139,1170,219,1510]" captionTargetPageId="7" captionText="Fig. 4. — Schematic plan views and transverse sections of grades of skeletal organization from the most primitive sessile barnacles (A, Brachylepadomorpha) through the asymmetrical sessile barnacles (B-H, Verrucomorpha). Of the eight verrucid genera currently recognized (Young 1998) only the four most representative ones (Altiverruca, Newmaniverruca, Metaverruca and Verruca s.s.) are depicted here. Right sides are to the viewers right and rostral ends are downward in plan views or coming out of the page in transverse sections. The transverse sections are slightly rostral of the rostro-carinal gap or suture. While there are numerous whorls of imbricating plates standing in tiers four plates high in brachylepadomorphs, and on the movable side of neoverrucids, only those of the R-C gap are labeled. Abbreviations: C, R, carina and rostrum; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral tiers of imbricating plates respectively; cl, rl, the pair of imbricating plates in proverrucids; FS, FT, fixed scutum and tergum; hl, hinge line; L, median latus (completely lost in proverrucids and verrucids); m, myophore; MS, MT, movable scutum and tergum; sam, scutal adductor muscle; S, T, normal terga and scuta in brachylepadomorphs; dashed horizontal lines: membranous basis (transverse sections F-H after Young 1998)." figureDoi="http://doi.org/10.5281/zenodo.13988759" httpUri="https://zenodo.org/record/13988759/files/figure.png" pageId="3" pageNumber="74">4C</figureCitation>
|
||||
; 5; 6)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C332558AFFB4D020FD58FE897ABC9DA9" pageId="3" pageNumber="74" type="materials_examined">
|
||||
<paragraph id="8B970601FFB4D020FD58FE897ABC9DA9" blockId="3.[677,1210,299,865]" pageId="3" pageNumber="74">
|
||||
<materialsCitation id="3B400C5CFFB4D020FD58FE897AB89DA9" collectionCode="CENTOB" collectorName="A. - M. Alayse-Danet" country="Tonga" county="Hine Hina" determinerName="Newman & Yamaguchi & Southward & Newman & Desbruyeres" elevation="1900" latitude="-22.533333" location="French" longLatPrecision="1259" longitude="-176.71666" municipality="Newman" pageId="3" pageNumber="74" specimenCode="BL-01, BL-03" specimenCount="2" specimenCount-adult="2">
|
||||
MATERIAL. — Specimens were collected by the
|
||||
<location id="8EF750DAFFB4D020FB88FE8E7CE09E2D" LSID="urn:lsid:plazi:treatment:0381B717FFB4D029FD04FF7279B09BEB:8EF750DAFFB4D020FB88FE8E7CE09E2D" box="[1141,1210,300,324]" country="Tonga" county="Hine Hina" latitude="-22.533333" longLatPrecision="1259" longitude="-176.71666" municipality="Newman" name="French" pageId="3" pageNumber="74">French</location>
|
||||
submersible
|
||||
<emphasis id="B95CDA13FFB4D020FCDFFEE77B309E34" box="[802,874,325,349]" italics="true" pageId="3" pageNumber="74">
|
||||
<location id="8EF750DAFFB4D020FCDFFEE77B309E34" LSID="urn:lsid:plazi:treatment:0381B717FFB4D029FD04FF7279B09BEB:8EF750DAFFB4D020FCDFFEE77B309E34" box="[802,874,325,349]" country="Tonga" county="Hine Hina" latitude="-22.533333" longLatPrecision="1259" longitude="-176.71666" municipality="Newman" name="Nautile" pageId="3" pageNumber="74">Nautile</location>
|
||||
</emphasis>
|
||||
,
|
||||
<location id="8EF750DAFFB4D020FC84FEE77C449E34" LSID="urn:lsid:plazi:treatment:0381B717FFB4D029FD04FF7279B09BEB:8EF750DAFFB4D020FC84FEE77C449E34" box="[889,1054,325,349]" country="Tonga" county="Hine Hina" latitude="-22.533333" longLatPrecision="1259" longitude="-176.71666" municipality="Newman" name="BIOLAU Cruise" pageId="3" pageNumber="74">BIOLAU Cruise</location>
|
||||
of 1989 to the
|
||||
<location id="8EF750DAFFB4D020FD58FEFC7B579E1E" LSID="urn:lsid:plazi:treatment:0381B717FFB4D029FD04FF7279B09BEB:8EF750DAFFB4D020FD58FEFC7B579E1E" box="[677,781,350,375]" country="Tonga" county="Hine Hina" latitude="-22.533333" longLatPrecision="1259" longitude="-176.71666" municipality="Newman" name="Lau Basin" pageId="3" pageNumber="74">Lau Basin</location>
|
||||
,
|
||||
<collectingCountry id="F33F4691FFB4D020FCE4FEFC7B009E1F" box="[793,858,350,374]" name="Tonga" pageId="3" pageNumber="74">Tonga</collectingCountry>
|
||||
(
|
||||
<collectorName id="26DD63D7FFB4D020FC96FEFD7C619E1E" box="[875,1083,351,375]" pageId="3" pageNumber="74">A.-M. Alayse-Danet</collectorName>
|
||||
, chief scientist), and they were received from the Centre national de tri d’Océanographie biologique (
|
||||
<collectionCode id="ED399EC4FFB4D020FBCBFE337CF79EC0" box="[1078,1197,401,425]" pageId="3" pageNumber="74">CENTOB</collectionCode>
|
||||
), IFREMER, Brest.
|
||||
<specimenCount id="9D2ECD88FFB4D020FC96FE087B809EAA" box="[875,986,426,451]" count="2" pageId="3" pageNumber="74" type="adult">Two adult</specimenCount>
|
||||
specimens on round stones partially coated with sulfide deposits, plus several juveniles about to shed their cyprid shells, inferred to belong to this species (Jensen’s x-juveniles of
|
||||
<bibRefCitation id="EFB97BF0FFB4D020FB7EFE557B0F9D41" author="HOeg J. T. & Newman W. A." bookContentInfo="Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris" pageId="3" pageNumber="74" refId="ref6736" refString="HOeg J. T. & Newman W. A. 1997. - Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga. Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris." title="Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga." type="proceedings" year="1997">HØeg & Newman 1997</bibRefCitation>
|
||||
; see below), all from station 1 (
|
||||
<collectingCounty id="62F67E8DFFB4D020FB7BFDB27A809D28" pageId="3" pageNumber="74">Hine Hina</collectingCounty>
|
||||
,
|
||||
<geoCoordinate id="EE1C60C6FFB4D020FD18FD8B7B6F9D28" box="[741,821,553,577]" degrees="22" direction="south" minutes="32" orientation="latitude" pageId="3" pageNumber="74" precision="925" value="-22.533333">22°32’S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE1C60C6FFB4D020FCBDFD8B7BF39D28" box="[832,937,553,577]" degrees="176" direction="west" minutes="43" orientation="longitude" pageId="3" pageNumber="74" precision="925" value="-176.71666">176°43’W</geoCoordinate>
|
||||
, at
|
||||
<elevation id="0005E132FFB4D020FC30FD8B7C059D28" box="[973,1119,553,577]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="74" unit="m" value="1900.0">
|
||||
<quantity id="4CD0ABE4FFB4D020FC30FD8B7C419D29" box="[973,1051,553,577]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="74" unit="m" value="1900.0">1900 m</quantity>
|
||||
depth
|
||||
</elevation>
|
||||
). Two of these samples (
|
||||
<specimenCode id="DB8EAE7AFFB4D020FCC5FDE17B229D32" box="[824,888,579,603]" pageId="3" pageNumber="74">BL-01</specimenCode>
|
||||
&
|
||||
<specimenCode id="DB8EAE7AFFB4D020FC60FDE17B879D32" box="[925,989,579,603]" pageId="3" pageNumber="74">BL-03</specimenCode>
|
||||
) contained other vent barnacles and other vent organisms (
|
||||
<bibRefCitation id="EFB97BF0FFB4D020FBC4FDFE7B3B9DE4" author="Newman W. A. & Yamaguchi T." journalOrPublisher="Bulletin du Museum national d'Histoire naturelle" pageId="3" pageNumber="74" pagination="211 - 243" part="17" refId="ref7319" refString="Newman W. A. & Yamaguchi T. 1995. - A new sessile barnacle (Cirripedia, Brachylepadomorpha) from the Lau Back-Arc Basin, Tonga; first record of a living representative since the Miocene. Bulletin du Museum national d'Histoire naturelle, 4 e serie, 17 A (3 - 4): 211 - 243." title="A new sessile barnacle (Cirripedia, Brachylepadomorpha) from the Lau Back-Arc Basin, Tonga; first record of a living representative since the Miocene" type="journal article" year="1995">
|
||||
<determinerName id="7E4BEAC5FFB4D020FBC4FDFE7B479DE4" pageId="3" pageNumber="74">Newman & Yamaguchi</determinerName>
|
||||
1995
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFB97BF0FFB4D020FC92FDD77CEF9DE4" author="Yamaguchi T. & Newman W. A." box="[879,1205,629,653]" journalOrPublisher="Zoosystema" pageId="3" pageNumber="74" pagination="623 - 649" part="19" refId="ref7936" refString="Yamaguchi T. & Newman W. A. 1997. - The hydrothermal vent barnacles Eochionelasmus (Cirripedia, Balanomorpha) from the North Fiji, Lau and Manus Basins, South-West Pacific. Zoosystema 19 (4): 623 - 649." title="The hydrothermal vent barnacles Eochionelasmus (Cirripedia, Balanomorpha) from the North Fiji, Lau and Manus Basins, South-West Pacific" type="journal article" year="1997">Yamaguchi & Newman 1997</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFB97BF0FFB4D020FD58FD2D7B8F9DCE" author="Southward A. J. & Newman W. A." box="[677,981,654,679]" journalOrPublisher="Cahiers de Biologie marine" pageId="3" pageNumber="74" pagination="258 - 262" part="39" refId="ref7585" refString="Southward A. J. & Newman W. A. 1998. - Ectosymbiosis between filamentous sulphur bacteria and a stalked barnacle (Scalpellomorpha, Neolepadinae) from the Lau Back Arc Basin, Tonga. Cahiers de Biologie marine 39 (3 - 4): 258 - 262." title="Ectosymbiosis between filamentous sulphur bacteria and a stalked barnacle (Scalpellomorpha, Neolepadinae) from the Lau Back Arc Basin, Tonga" type="journal article" year="1998">
|
||||
<determinerName id="7E4BEAC5FFB4D020FD58FD2D7BCF9DCC" box="[677,917,654,679]" pageId="3" pageNumber="74">Southward & Newman</determinerName>
|
||||
1998
|
||||
</bibRefCitation>
|
||||
; cf.
|
||||
<bibRefCitation id="EFB97BF0FFB4D020FBFEFD2D7A819DA9" author="Desbruyeres D. & Alayse-Danet A. - M. & Ohta S. & the scientific parties of the Biolau and Starmer cruises" journalOrPublisher="Marine Geology" pageId="3" pageNumber="74" pagination="227 - 242" part="116" refId="ref6571" refString="Desbruyeres D., Alayse-Danet A. - M., Ohta S. & the scientific parties of the Biolau and Starmer cruises 1994. - Deep-sea hydrothermal communities in southwestern Pacific back-arc basins (the North Fiji and Lau Basins): Composition, microdistribution and food web. Marine Geology 116: 227 - 242." title="Deep-sea hydrothermal communities in southwestern Pacific back-arc basins (the North Fiji and Lau Basins): Composition, microdistribution and food web" type="journal article" year="1994">
|
||||
<determinerName id="7E4BEAC5FFB4D020FBFEFD2D7C259DCE" box="[1027,1151,655,679]" pageId="3" pageNumber="74">Desbruyères</determinerName>
|
||||
<emphasis id="B95CDA13FFB4D020FB75FD2C7CE09DCE" box="[1160,1210,654,679]" italics="true" pageId="3" pageNumber="74">et al.</emphasis>
|
||||
1994
|
||||
</bibRefCitation>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C332558AFFB4D020FD58FD707C5E9C5F" pageId="3" pageNumber="74" type="etymology">
|
||||
<paragraph id="8B970601FFB4D020FD58FD707C5E9C5F" blockId="3.[677,1210,299,865]" pageId="3" pageNumber="74">ETYMOLOGY. — Named for Toshiyuki Yamaguchi in appreciation of his numerous and exacting works on the systematics and ecology of the living and fossil cirripeds, especially the vent barnacles.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C332558AFFB4D020FD58FCEA7BBB9C09" box="[677,993,840,865]" pageId="3" pageNumber="74" type="diagnosis">
|
||||
<paragraph id="8B970601FFB4D020FD58FCEA7BBB9C09" blockId="3.[677,1210,299,865]" box="[677,993,840,865]" pageId="3" pageNumber="74">DIAGNOSIS. — As for the genus.</paragraph>
|
||||
</subSubSection>
|
||||
<paragraph id="8B970601FFB4D020FF05FC0B7BD79B57" pageId="3" pageNumber="74">
|
||||
<table id="F928F4A1FFB42FDCFF05FC0B7C799B57" box="[248,1059,937,1086]" gridcols="5" gridrows="4" pageId="3" pageNumber="74">
|
||||
<tr id="35180443FFB42FDCFF05FC0B7C799CAA" box="[248,1059,937,963]" gridrow="0" pageId="3" pageNumber="74">
|
||||
<th id="76C96D3FFFB42FDCFF05FC0B79039CAA" box="[248,345,937,963]" gridcol="0" gridrow="0" pageId="3" pageNumber="74">Sample</th>
|
||||
<th id="76C96D3FFFB42FDCFE81FC0B79E39CAA" box="[380,441,937,963]" gridcol="1" gridrow="0" pageId="3" pageNumber="74">R-C</th>
|
||||
<th id="76C96D3FFFB42FDCFE0AFC0B7A4E9CAA" box="[503,532,937,963]" gridcol="2" gridrow="0" pageId="3" pageNumber="74">W</th>
|
||||
<th id="76C96D3FFFB42FDCFDA0FC0B7B689CAA" box="[605,818,937,963]" gridcol="3" gridrow="0" pageId="3" pageNumber="74">Type designation</th>
|
||||
<th id="76C96D3FFFB42FDCFC96FC0B7C799CAA" box="[875,1059,937,963]" gridcol="4" gridrow="0" pageId="3" pageNumber="74">Depository</th>
|
||||
</tr>
|
||||
<tr id="35180443FFB42FDCFF05FC447C799B69" box="[248,1059,998,1024]" gridrow="1" pageId="3" pageNumber="74">
|
||||
<th id="76C96D3FFFB42FDCFF05FC4479039B69" box="[248,345,998,1024]" gridcol="0" gridrow="1" pageId="3" pageNumber="74">1. BL-01</th>
|
||||
<td id="76C96D3FFFB42FDCFE81FC4479E39B69" box="[380,441,998,1024]" gridcol="1" gridrow="1" pageId="3" pageNumber="74">~7</td>
|
||||
<td id="76C96D3FFFB42FDCFE0AFC447A4E9B69" box="[503,532,998,1024]" gridcol="2" gridrow="1" pageId="3" pageNumber="74">~6</td>
|
||||
<td id="76C96D3FFFB42FDCFDA0FC447B689B69" box="[605,818,998,1024]" gridcol="3" gridrow="1" pageId="3" pageNumber="74">holotype (intact)</td>
|
||||
<td id="76C96D3FFFB42FDCFC96FC447C799B69" box="[875,1059,998,1024]" gridcol="4" gridrow="1" pageId="3" pageNumber="74">MNHN-Ci2710</td>
|
||||
</tr>
|
||||
<tr id="35180443FFB42FDCFF05FBA67C799B76" box="[248,1059,1028,1055]" gridrow="2" pageId="3" pageNumber="74">
|
||||
<th id="76C96D3FFFB42FDCFF05FBA679039B76" box="[248,345,1028,1055]" gridcol="0" gridrow="2" pageId="3" pageNumber="74">2. BL-01</th>
|
||||
<td id="76C96D3FFFB42FDCFE81FBA679E39B76" box="[380,441,1028,1055]" gridcol="1" gridrow="2" pageId="3" pageNumber="74">~7</td>
|
||||
<td id="76C96D3FFFB42FDCFE0AFBA67A4E9B76" box="[503,532,1028,1055]" gridcol="2" gridrow="2" pageId="3" pageNumber="74">~6</td>
|
||||
<td id="76C96D3FFFB42FDCFDA0FBA67B689B76" box="[605,818,1028,1055]" gridcol="3" gridrow="2" pageId="3" pageNumber="74">paratype (dissected)</td>
|
||||
<td id="76C96D3FFFB42FDCFC96FBA67C799B76" box="[875,1059,1028,1055]" gridcol="4" gridrow="2" pageId="3" pageNumber="74">MNHN-Ci2711</td>
|
||||
</tr>
|
||||
<tr id="35180443FFB42FDCFF05FB817C799B57" box="[248,1059,1059,1086]" gridrow="3" pageId="3" pageNumber="74">
|
||||
<th id="76C96D3FFFB42FDCFF05FB8179039B57" box="[248,345,1059,1086]" gridcol="0" gridrow="3" pageId="3" pageNumber="74">3. BL-03</th>
|
||||
<td id="76C96D3FFFB42FDCFE81FB817C799B57" box="[380,1059,1059,1086]" colspan="4" colspanRight="3" gridcol="1" gridrow="3" pageId="3" pageNumber="74">Jensen’s x-juveniles, care of J. HØeg, Copenhagen</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<tableNote id="76CE078FFFB4D020FF05FBFB79889BEB" pageId="3" pageNumber="74" targetBox="[248,1059,937,1086]" targetPageId="3">
|
||||
<paragraph id="8B970601FFB4D020FF05FBFB79B29B05" blockId="3.[248,488,1113,1154]" box="[248,488,1113,1132]" pageId="3" pageNumber="74">R-C rostro-carinal diameter</paragraph>
|
||||
<paragraph id="8B970601FFB4D020FF05FBCD79889BEB" blockId="3.[248,488,1113,1154]" box="[248,466,1135,1154]" pageId="3" pageNumber="74">W width, both in mm.</paragraph>
|
||||
</tableNote>
|
||||
<subSubSection id="C332558AFFB4D029FF8CFB6779B09BEB" lastPageId="10" lastPageNumber="81" pageId="3" pageNumber="74" type="description">
|
||||
<paragraph id="8B970601FFB4D020FF8CFB6779D49BB5" blockId="3.[113,647,1221,1707]" box="[113,398,1221,1247]" pageId="3" pageNumber="74">COMPARATIVEDESCRIPTION</paragraph>
|
||||
<paragraph id="8B970601FFB4D020FF8CFB4178B89B95" blockId="3.[113,647,1221,1707]" box="[113,226,1251,1277]" pageId="3" pageNumber="74">
|
||||
<emphasis id="B95CDA13FFB4D020FF8CFB4178B89B95" box="[113,226,1251,1277]" italics="true" pageId="3" pageNumber="74">Hard parts</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8B970601FFB4D020FF8CFAA07B6E9978" blockId="3.[113,647,1221,1707]" lastBlockId="3.[677,1210,1220,1707]" pageId="3" pageNumber="74">
|
||||
The low-conic profile of the new genus,
|
||||
<taxonomicName id="4C287D82FFB4D020FF8CFA83797A9A52" authorityName="Newman" authorityYear="2000" box="[113,288,1313,1339]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB4D020FF8CFA83797A9A52" box="[113,288,1313,1339]" italics="true" pageId="3" pageNumber="74">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
, is similar to
|
||||
<taxonomicName id="4C287D82FFB4D020FE25FA83799C9A30" authority="Newman & Yamaguchi, 1995" authorityName="Newman & Yamaguchi" authorityYear="1995" class="Maxillopoda" family="Neobrachylepadidae" genus="Neobrachylepas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB4D020FE25FA837ADC9A52" box="[472,646,1313,1339]" italics="true" pageId="3" pageNumber="74">Neobrachylepas</emphasis>
|
||||
<bibRefCitation id="EFB97BF0FFB4D020FF8CFA9C799C9A30" author="Newman W. A. & Yamaguchi T." box="[113,454,1342,1369]" journalOrPublisher="Bulletin du Museum national d'Histoire naturelle" pageId="3" pageNumber="74" pagination="211 - 243" part="17" refId="ref7319" refString="Newman W. A. & Yamaguchi T. 1995. - A new sessile barnacle (Cirripedia, Brachylepadomorpha) from the Lau Back-Arc Basin, Tonga; first record of a living representative since the Miocene. Bulletin du Museum national d'Histoire naturelle, 4 e serie, 17 A (3 - 4): 211 - 243." title="A new sessile barnacle (Cirripedia, Brachylepadomorpha) from the Lau Back-Arc Basin, Tonga; first record of a living representative since the Miocene" type="journal article" year="1995">Newman & Yamaguchi, 1995</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, whose representatives were relatively young, and to the juveniles of
|
||||
<taxonomicName id="4C287D82FFB4D020FF11FADE792E9AFC" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[236,372,1404,1429]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB4D020FF11FADE792E9AFC" box="[236,372,1404,1429]" italics="true" pageId="3" pageNumber="74">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C287D82FFB4D020FE37FADF79909ADC" authority="Yamaguchi & Newman, 1990" authorityName="Yamaguchi & Newman" authorityYear="1990" class="Maxillopoda" family="Chionelasmatidae" genus="Eochionelasmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB4D020FE37FADF7ADC9AFE" box="[458,646,1405,1431]" italics="true" pageId="3" pageNumber="74">Eochionelasmus</emphasis>
|
||||
<bibRefCitation id="EFB97BF0FFB4D020FF8CFA3979909ADC" author="Yamaguchi T. & Newman W. A." box="[113,458,1434,1461]" journalOrPublisher="Pacific Science" pageId="3" pageNumber="74" pagination="135 - 155" part="44" refId="ref7889" refString="Yamaguchi T. & Newman W. A. 1990. - A new and primitive barnacle (Cirripedia; Balanomorpha) from the North Fiji Basin abyssal hydrothermal field, and its evolutionary implications. Pacific Science 44 (2): 135 - 155." title="A new and primitive barnacle (Cirripedia; Balanomorpha) from the North Fiji Basin abyssal hydrothermal field, and its evolutionary implications" type="journal article" year="1990">Yamaguchi & Newman, 1990</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. The operculum forms a flat lid, as in the
|
||||
<taxonomicName id="4C287D82FFB4D020FE4FFA187A0A9ABD" authorityName="Newman, 1989" authorityYear="1989" box="[434,592,1466,1492]" class="Maxillopoda" family="Proverrucidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="family">Proverrucidae</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C287D82FFB4D020FF8CFA7B78B39A9A" authorityName="Darwin" authorityYear="1854" box="[113,233,1497,1523]" class="Maxillopoda" family="Verrucidae" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="family">Verrucidae</taxonomicName>
|
||||
, but the hinge line is curved rather than essentially straight as in
|
||||
<taxonomicName id="4C287D82FFB4D020FE44FA557A749979" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[441,558,1527,1552]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB4D020FE44FA557A749979" box="[441,558,1527,1552]" italics="true" pageId="3" pageNumber="74">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
, and in higher verrucomorphs (cf.
|
||||
<figureCitation id="13131A84FFB4D020FE2BF9B47A619959" box="[470,571,1558,1584]" captionStart="FIG" captionStartId="2.[114,125,799,816]" captionTargetBox="[262,1053,208,766]" captionTargetId="figure-307@2.[248,1067,202,776]" captionTargetPageId="2" captionText="FIG. 1. — A, Imbricaverruca yamaguchii n. gen. and n. sp. (holotype, MNHN Ci2710); B, Verruca s.l., both viewed from above with the right scutum and tergum forming the operculum.Note that in the former the four plated wall appears to be covered largely by imbricating plates and the operculum includes a large median latus,characters that readily distinguish it from Neoverruca. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5391397" httpUri="https://zenodo.org/record/5391397/files/figure.png" pageId="3" pageNumber="74">Figs 1A</figureCitation>
|
||||
;
|
||||
<figureCitation id="13131A84FFB4D020FDACF9B47A209959" box="[593,634,1558,1584]" captionStart="FIG" captionStartId="4.[114,125,1384,1401]" captionTargetBox="[213,1067,193,1353]" captionTargetId="figure-91@4.[208,1091,186,1354]" captionTargetPageId="4" captionText="FIG. 2. — Plates of Imbricaverruca yamaguchii gen. et n. sp. (paratype, MNHN Ci2711); A, B, viewed from above and from the rostral end, respectively; A, note the operculum, comprising the plates of the right side (MS-L-MT), includes a large median latus; B, note the slightly open aperture to the mantle cavity between the occludent margins of the movable (MS & MT) and the fixed (FS & FT) scutum and tergum, respectively; C-M, variously disarticulated hard parts; C, interior view of wall (R-C-FS-FT) and operculum (MS-L- MT) with all of the imbricating plates, except the uppermost of the three principle lateral tiers of the right side (rl1-l1-cl1), stripped away [an approximation of the rostrum (R), which did not survive dissection, is indicated by the dashed line]; D, E, movable scutum (MS); external and internal views respectively; scutal adductor muscle, inserting from just below to up under the apical concavity in Fig. 2E, extends to its origin on the fixed scutum (FS, Fig. 2G; the relationship of MS to FS can be seen in Fig 2C); F, G, fixed scutum (FS); basal and internal views, respectively. Dashed line in G indicates origin of scutal adductor muscle but no scar is evident. The relationship of the two protuberances (best seen near the tergal margin in Fig. 2G) to the fixed tergum (FT, Fig. 2J, K), was not resolved; H, I, movable tergum (MT); external and internal views respectively (external corrosion in bleach and cleaning inadvertently obliterated growth line ornamentation in H); J, K, fixed tergum (FT); external and internal views respectively; L, M, carina from below and above respectively (the rostrum did not survive dissection and cleaning). Abbreviations: C, carina; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral plates forming three tiers each four plates high, respectively; FS, FT, fixed scutum and tergum; L, median latus; MS, MT, movable scutum and tergum; R, rostrum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391399" httpUri="https://zenodo.org/record/5391399/files/figure.png" pageId="3" pageNumber="74">2A</figureCitation>
|
||||
). Therefore, in order for the operculum to open, there is likely some flexibility to the wall as well as some freedom in the arthrodial membrane forming the hinge, and it must be able to bend along the scuto-tergal articulation, an ability likely facilitated by the basal gap protected by the median latus (
|
||||
<figureCitation id="13131A84FFB4D020FC82FAA07B849A75" box="[895,990,1282,1308]" captionStart="FIG" captionStartId="4.[114,125,1384,1401]" captionTargetBox="[213,1067,193,1353]" captionTargetId="figure-91@4.[208,1091,186,1354]" captionTargetPageId="4" captionText="FIG. 2. — Plates of Imbricaverruca yamaguchii gen. et n. sp. (paratype, MNHN Ci2711); A, B, viewed from above and from the rostral end, respectively; A, note the operculum, comprising the plates of the right side (MS-L-MT), includes a large median latus; B, note the slightly open aperture to the mantle cavity between the occludent margins of the movable (MS & MT) and the fixed (FS & FT) scutum and tergum, respectively; C-M, variously disarticulated hard parts; C, interior view of wall (R-C-FS-FT) and operculum (MS-L- MT) with all of the imbricating plates, except the uppermost of the three principle lateral tiers of the right side (rl1-l1-cl1), stripped away [an approximation of the rostrum (R), which did not survive dissection, is indicated by the dashed line]; D, E, movable scutum (MS); external and internal views respectively; scutal adductor muscle, inserting from just below to up under the apical concavity in Fig. 2E, extends to its origin on the fixed scutum (FS, Fig. 2G; the relationship of MS to FS can be seen in Fig 2C); F, G, fixed scutum (FS); basal and internal views, respectively. Dashed line in G indicates origin of scutal adductor muscle but no scar is evident. The relationship of the two protuberances (best seen near the tergal margin in Fig. 2G) to the fixed tergum (FT, Fig. 2J, K), was not resolved; H, I, movable tergum (MT); external and internal views respectively (external corrosion in bleach and cleaning inadvertently obliterated growth line ornamentation in H); J, K, fixed tergum (FT); external and internal views respectively; L, M, carina from below and above respectively (the rostrum did not survive dissection and cleaning). Abbreviations: C, carina; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral plates forming three tiers each four plates high, respectively; FS, FT, fixed scutum and tergum; L, median latus; MS, MT, movable scutum and tergum; R, rostrum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391399" httpUri="https://zenodo.org/record/5391399/files/figure.png" pageId="3" pageNumber="74">Fig. 2C</figureCitation>
|
||||
). Like
|
||||
<taxonomicName id="4C287D82FFB4D020FBCBFAA37CEB9A73" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[1078,1201,1281,1306]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB4D020FBCBFAA37CEB9A73" box="[1078,1201,1281,1306]" italics="true" pageId="3" pageNumber="74">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
, the scutal adductor muscle inserts in a pocket below the apex of the movable plate (
|
||||
<figureCitation id="13131A84FFB4D020FBABFA9D7CF79A30" box="[1110,1197,1343,1369]" captionStart="FIG" captionStartId="4.[114,125,1384,1401]" captionTargetBox="[213,1067,193,1353]" captionTargetId="figure-91@4.[208,1091,186,1354]" captionTargetPageId="4" captionText="FIG. 2. — Plates of Imbricaverruca yamaguchii gen. et n. sp. (paratype, MNHN Ci2711); A, B, viewed from above and from the rostral end, respectively; A, note the operculum, comprising the plates of the right side (MS-L-MT), includes a large median latus; B, note the slightly open aperture to the mantle cavity between the occludent margins of the movable (MS & MT) and the fixed (FS & FT) scutum and tergum, respectively; C-M, variously disarticulated hard parts; C, interior view of wall (R-C-FS-FT) and operculum (MS-L- MT) with all of the imbricating plates, except the uppermost of the three principle lateral tiers of the right side (rl1-l1-cl1), stripped away [an approximation of the rostrum (R), which did not survive dissection, is indicated by the dashed line]; D, E, movable scutum (MS); external and internal views respectively; scutal adductor muscle, inserting from just below to up under the apical concavity in Fig. 2E, extends to its origin on the fixed scutum (FS, Fig. 2G; the relationship of MS to FS can be seen in Fig 2C); F, G, fixed scutum (FS); basal and internal views, respectively. Dashed line in G indicates origin of scutal adductor muscle but no scar is evident. The relationship of the two protuberances (best seen near the tergal margin in Fig. 2G) to the fixed tergum (FT, Fig. 2J, K), was not resolved; H, I, movable tergum (MT); external and internal views respectively (external corrosion in bleach and cleaning inadvertently obliterated growth line ornamentation in H); J, K, fixed tergum (FT); external and internal views respectively; L, M, carina from below and above respectively (the rostrum did not survive dissection and cleaning). Abbreviations: C, carina; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral plates forming three tiers each four plates high, respectively; FS, FT, fixed scutum and tergum; L, median latus; MS, MT, movable scutum and tergum; R, rostrum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391399" httpUri="https://zenodo.org/record/5391399/files/figure.png" pageId="3" pageNumber="74">Fig. 2E</figureCitation>
|
||||
), and its origin on the fixed scutum, near the centre of the plate (
|
||||
<figureCitation id="13131A84FFB4D020FC7AFADF7BBE9AFE" box="[903,996,1405,1431]" captionStart="FIG" captionStartId="4.[114,125,1384,1401]" captionTargetBox="[213,1067,193,1353]" captionTargetId="figure-91@4.[208,1091,186,1354]" captionTargetPageId="4" captionText="FIG. 2. — Plates of Imbricaverruca yamaguchii gen. et n. sp. (paratype, MNHN Ci2711); A, B, viewed from above and from the rostral end, respectively; A, note the operculum, comprising the plates of the right side (MS-L-MT), includes a large median latus; B, note the slightly open aperture to the mantle cavity between the occludent margins of the movable (MS & MT) and the fixed (FS & FT) scutum and tergum, respectively; C-M, variously disarticulated hard parts; C, interior view of wall (R-C-FS-FT) and operculum (MS-L- MT) with all of the imbricating plates, except the uppermost of the three principle lateral tiers of the right side (rl1-l1-cl1), stripped away [an approximation of the rostrum (R), which did not survive dissection, is indicated by the dashed line]; D, E, movable scutum (MS); external and internal views respectively; scutal adductor muscle, inserting from just below to up under the apical concavity in Fig. 2E, extends to its origin on the fixed scutum (FS, Fig. 2G; the relationship of MS to FS can be seen in Fig 2C); F, G, fixed scutum (FS); basal and internal views, respectively. Dashed line in G indicates origin of scutal adductor muscle but no scar is evident. The relationship of the two protuberances (best seen near the tergal margin in Fig. 2G) to the fixed tergum (FT, Fig. 2J, K), was not resolved; H, I, movable tergum (MT); external and internal views respectively (external corrosion in bleach and cleaning inadvertently obliterated growth line ornamentation in H); J, K, fixed tergum (FT); external and internal views respectively; L, M, carina from below and above respectively (the rostrum did not survive dissection and cleaning). Abbreviations: C, carina; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral plates forming three tiers each four plates high, respectively; FS, FT, fixed scutum and tergum; L, median latus; MS, MT, movable scutum and tergum; R, rostrum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391399" httpUri="https://zenodo.org/record/5391399/files/figure.png" pageId="3" pageNumber="74">Figs 2G</figureCitation>
|
||||
;
|
||||
<figureCitation id="13131A84FFB4D020FC09FADF7C429AFE" box="[1012,1048,1405,1431]" captionStart="Fig" captionStartId="7.[113,141,1469,1486]" captionTargetBox="[112,1165,215,1445]" captionTargetId="figure-22@7.[139,1170,219,1510]" captionTargetPageId="7" captionText="Fig. 4. — Schematic plan views and transverse sections of grades of skeletal organization from the most primitive sessile barnacles (A, Brachylepadomorpha) through the asymmetrical sessile barnacles (B-H, Verrucomorpha). Of the eight verrucid genera currently recognized (Young 1998) only the four most representative ones (Altiverruca, Newmaniverruca, Metaverruca and Verruca s.s.) are depicted here. Right sides are to the viewers right and rostral ends are downward in plan views or coming out of the page in transverse sections. The transverse sections are slightly rostral of the rostro-carinal gap or suture. While there are numerous whorls of imbricating plates standing in tiers four plates high in brachylepadomorphs, and on the movable side of neoverrucids, only those of the R-C gap are labeled. Abbreviations: C, R, carina and rostrum; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral tiers of imbricating plates respectively; cl, rl, the pair of imbricating plates in proverrucids; FS, FT, fixed scutum and tergum; hl, hinge line; L, median latus (completely lost in proverrucids and verrucids); m, myophore; MS, MT, movable scutum and tergum; sam, scutal adductor muscle; S, T, normal terga and scuta in brachylepadomorphs; dashed horizontal lines: membranous basis (transverse sections F-H after Young 1998)." figureDoi="http://doi.org/10.5281/zenodo.13988759" httpUri="https://zenodo.org/record/13988759/files/figure.png" pageId="3" pageNumber="74">4C</figureCitation>
|
||||
), is supported by an adductor ridge or “myophore” as in
|
||||
<taxonomicName id="4C287D82FFB4D020FD58FA1B7C5E9ABD" authority="Pilsbry, 1916" authorityName="Pilsbry" authorityYear="1916" box="[677,1028,1465,1492]" class="Maxillopoda" family="Verrucidae" genus="Metaverruca" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB4D020FD58FA1B7B679ABB" box="[677,829,1465,1490]" italics="true" pageId="3" pageNumber="74">Metaverruca</emphasis>
|
||||
<bibRefCitation id="EFB97BF0FFB4D020FCAFFA187C5E9ABD" author="Pilsbry H. A." box="[850,1028,1466,1492]" journalOrPublisher="Bulletin of the United States National Museum" pageId="3" pageNumber="74" pagination="1 - 366" part="93" refId="ref7445" refString="Pilsbry H. A. 1916. - The sessile barnacles (Cirripedia) contained in the collections of the U. S. National Museum; including a monograph of the American species. Bulletin of the United States National Museum 93: 1 - 366." title="The sessile barnacles (Cirripedia) contained in the collections of the U. S. National Museum; including a monograph of the American species" type="journal article" year="1916">Pilsbry, 1916</bibRefCitation>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C287D82FFB4D020FBA0FA1B7B2A9A9A" authority="Schumacher, 1817" authorityName="Schumacher" authorityYear="1817" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB4D020FBA0FA1B7CE39ABB" box="[1117,1209,1465,1490]" italics="true" pageId="3" pageNumber="74">Verruca</emphasis>
|
||||
Schumacher, 1817
|
||||
</taxonomicName>
|
||||
(cf.
|
||||
<bibRefCitation id="EFB97BF0FFB4D020FC58FA7A7C719A9A" author="Young P. S." box="[933,1067,1496,1523]" journalOrPublisher="Zoosystema" pageId="3" pageNumber="74" pagination="31 - 92" part="20" refId="ref7981" refString="Young P. S. 1998. - Cirripedia (Crustacea) from the " Campagne Biacores " in the Azores region, including a generic revision of Verrucidae. Zoosystema 20 (1): 31 - 92." title="Cirripedia (Crustacea) from the " Campagne Biacores " in the Azores region, including a generic revision of Verrucidae" type="journal article" year="1998">Young 1998</bibRefCitation>
|
||||
for revision) (
|
||||
<figureCitation id="13131A84FFB4D020FD51FA557AAA9978" box="[684,752,1527,1553]" captionStart="Fig" captionStartId="7.[113,141,1469,1486]" captionTargetBox="[112,1165,215,1445]" captionTargetId="figure-22@7.[139,1170,219,1510]" captionTargetPageId="7" captionText="Fig. 4. — Schematic plan views and transverse sections of grades of skeletal organization from the most primitive sessile barnacles (A, Brachylepadomorpha) through the asymmetrical sessile barnacles (B-H, Verrucomorpha). Of the eight verrucid genera currently recognized (Young 1998) only the four most representative ones (Altiverruca, Newmaniverruca, Metaverruca and Verruca s.s.) are depicted here. Right sides are to the viewers right and rostral ends are downward in plan views or coming out of the page in transverse sections. The transverse sections are slightly rostral of the rostro-carinal gap or suture. While there are numerous whorls of imbricating plates standing in tiers four plates high in brachylepadomorphs, and on the movable side of neoverrucids, only those of the R-C gap are labeled. Abbreviations: C, R, carina and rostrum; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral tiers of imbricating plates respectively; cl, rl, the pair of imbricating plates in proverrucids; FS, FT, fixed scutum and tergum; hl, hinge line; L, median latus (completely lost in proverrucids and verrucids); m, myophore; MS, MT, movable scutum and tergum; sam, scutal adductor muscle; S, T, normal terga and scuta in brachylepadomorphs; dashed horizontal lines: membranous basis (transverse sections F-H after Young 1998)." figureDoi="http://doi.org/10.5281/zenodo.13988759" httpUri="https://zenodo.org/record/13988759/files/figure.png" pageId="3" pageNumber="74">Fig. 4</figureCitation>
|
||||
G-H).
|
||||
</paragraph>
|
||||
<paragraph id="8B970601FFB4D025FD58F9B479E59D74" blockId="3.[677,1210,1220,1707]" lastBlockId="6.[114,647,209,541]" lastPageId="6" lastPageNumber="77" pageId="3" pageNumber="74">
|
||||
The fixed scutum and tergum are much reduced in height (
|
||||
<figureCitation id="13131A84FFB4D020FCE8F9977B0E9926" box="[789,852,1589,1615]" captionStart="FIG" captionStartId="4.[114,125,1384,1401]" captionTargetBox="[213,1067,193,1353]" captionTargetId="figure-91@4.[208,1091,186,1354]" captionTargetPageId="4" captionText="FIG. 2. — Plates of Imbricaverruca yamaguchii gen. et n. sp. (paratype, MNHN Ci2711); A, B, viewed from above and from the rostral end, respectively; A, note the operculum, comprising the plates of the right side (MS-L-MT), includes a large median latus; B, note the slightly open aperture to the mantle cavity between the occludent margins of the movable (MS & MT) and the fixed (FS & FT) scutum and tergum, respectively; C-M, variously disarticulated hard parts; C, interior view of wall (R-C-FS-FT) and operculum (MS-L- MT) with all of the imbricating plates, except the uppermost of the three principle lateral tiers of the right side (rl1-l1-cl1), stripped away [an approximation of the rostrum (R), which did not survive dissection, is indicated by the dashed line]; D, E, movable scutum (MS); external and internal views respectively; scutal adductor muscle, inserting from just below to up under the apical concavity in Fig. 2E, extends to its origin on the fixed scutum (FS, Fig. 2G; the relationship of MS to FS can be seen in Fig 2C); F, G, fixed scutum (FS); basal and internal views, respectively. Dashed line in G indicates origin of scutal adductor muscle but no scar is evident. The relationship of the two protuberances (best seen near the tergal margin in Fig. 2G) to the fixed tergum (FT, Fig. 2J, K), was not resolved; H, I, movable tergum (MT); external and internal views respectively (external corrosion in bleach and cleaning inadvertently obliterated growth line ornamentation in H); J, K, fixed tergum (FT); external and internal views respectively; L, M, carina from below and above respectively (the rostrum did not survive dissection and cleaning). Abbreviations: C, carina; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral plates forming three tiers each four plates high, respectively; FS, FT, fixed scutum and tergum; L, median latus; MS, MT, movable scutum and tergum; R, rostrum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391399" httpUri="https://zenodo.org/record/5391399/files/figure.png" pageId="3" pageNumber="74">Fig. 2</figureCitation>
|
||||
F-G, J-K) and there is a complex articulation between them. Together with the rostrum and carina, which are also much wider than high, they form a ring-like wall supporting the operculum. Therefore, the imbricating plates between this primary wall and the substratum form a substantially greater proportion of the body chamber than in
|
||||
<taxonomicName id="4C287D82FFB1D025FE90FE8E79B89E2C" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[365,482,300,325]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB1D025FE90FE8E79B89E2C" box="[365,482,300,325]" italics="true" pageId="6" pageNumber="77">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
. The principal three tiers of lateral imbricating plates, standing four plates high (rl
|
||||
<superScript id="7C5DAB49FFB1D025FEC4FEC7790F9E11" attach="left" box="[313,341,357,376]" fontSize="8" pageId="6" pageNumber="77">1-4</superScript>
|
||||
, l
|
||||
<superScript id="7C5DAB49FFB1D025FE94FEC779DF9E11" attach="left" box="[361,389,357,376]" fontSize="8" pageId="6" pageNumber="77">1-4</superScript>
|
||||
and cl
|
||||
<superScript id="7C5DAB49FFB1D025FE31FEC779B29E11" attach="left" box="[460,488,357,376]" fontSize="8" pageId="6" pageNumber="77">1-4</superScript>
|
||||
), are fully represented in these two apparently adult
|
||||
<taxonomicName id="4C287D82FFB1D025FDC0FE2B78B99EA9" authorityName="Newman" authorityYear="2000" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB1D025FDC0FE2B78B99EA9" italics="true" pageId="6" pageNumber="77">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="13131A84FFB1D025FF0BFE0579039EA8" box="[246,345,423,449]" captionStart="FIG" captionStartId="2.[114,125,799,816]" captionTargetBox="[262,1053,208,766]" captionTargetId="figure-307@2.[248,1067,202,776]" captionTargetPageId="2" captionText="FIG. 1. — A, Imbricaverruca yamaguchii n. gen. and n. sp. (holotype, MNHN Ci2710); B, Verruca s.l., both viewed from above with the right scutum and tergum forming the operculum.Note that in the former the four plated wall appears to be covered largely by imbricating plates and the operculum includes a large median latus,characters that readily distinguish it from Neoverruca. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5391397" httpUri="https://zenodo.org/record/5391397/files/figure.png" pageId="6" pageNumber="77">Figs 1A</figureCitation>
|
||||
;
|
||||
<figureCitation id="13131A84FFB1D025FE94FE0579D59EA8" box="[361,399,423,449]" captionStart="FIG" captionStartId="4.[114,125,1384,1401]" captionTargetBox="[213,1067,193,1353]" captionTargetId="figure-91@4.[208,1091,186,1354]" captionTargetPageId="4" captionText="FIG. 2. — Plates of Imbricaverruca yamaguchii gen. et n. sp. (paratype, MNHN Ci2711); A, B, viewed from above and from the rostral end, respectively; A, note the operculum, comprising the plates of the right side (MS-L-MT), includes a large median latus; B, note the slightly open aperture to the mantle cavity between the occludent margins of the movable (MS & MT) and the fixed (FS & FT) scutum and tergum, respectively; C-M, variously disarticulated hard parts; C, interior view of wall (R-C-FS-FT) and operculum (MS-L- MT) with all of the imbricating plates, except the uppermost of the three principle lateral tiers of the right side (rl1-l1-cl1), stripped away [an approximation of the rostrum (R), which did not survive dissection, is indicated by the dashed line]; D, E, movable scutum (MS); external and internal views respectively; scutal adductor muscle, inserting from just below to up under the apical concavity in Fig. 2E, extends to its origin on the fixed scutum (FS, Fig. 2G; the relationship of MS to FS can be seen in Fig 2C); F, G, fixed scutum (FS); basal and internal views, respectively. Dashed line in G indicates origin of scutal adductor muscle but no scar is evident. The relationship of the two protuberances (best seen near the tergal margin in Fig. 2G) to the fixed tergum (FT, Fig. 2J, K), was not resolved; H, I, movable tergum (MT); external and internal views respectively (external corrosion in bleach and cleaning inadvertently obliterated growth line ornamentation in H); J, K, fixed tergum (FT); external and internal views respectively; L, M, carina from below and above respectively (the rostrum did not survive dissection and cleaning). Abbreviations: C, carina; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral plates forming three tiers each four plates high, respectively; FS, FT, fixed scutum and tergum; L, median latus; MS, MT, movable scutum and tergum; R, rostrum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391399" httpUri="https://zenodo.org/record/5391399/files/figure.png" pageId="6" pageNumber="77">2A</figureCitation>
|
||||
;
|
||||
<figureCitation id="13131A84FFB1D025FE63FE05799D9EA8" box="[414,455,423,449]" captionStart="Fig" captionStartId="7.[113,141,1469,1486]" captionTargetBox="[112,1165,215,1445]" captionTargetId="figure-22@7.[139,1170,219,1510]" captionTargetPageId="7" captionText="Fig. 4. — Schematic plan views and transverse sections of grades of skeletal organization from the most primitive sessile barnacles (A, Brachylepadomorpha) through the asymmetrical sessile barnacles (B-H, Verrucomorpha). Of the eight verrucid genera currently recognized (Young 1998) only the four most representative ones (Altiverruca, Newmaniverruca, Metaverruca and Verruca s.s.) are depicted here. Right sides are to the viewers right and rostral ends are downward in plan views or coming out of the page in transverse sections. The transverse sections are slightly rostral of the rostro-carinal gap or suture. While there are numerous whorls of imbricating plates standing in tiers four plates high in brachylepadomorphs, and on the movable side of neoverrucids, only those of the R-C gap are labeled. Abbreviations: C, R, carina and rostrum; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral tiers of imbricating plates respectively; cl, rl, the pair of imbricating plates in proverrucids; FS, FT, fixed scutum and tergum; hl, hinge line; L, median latus (completely lost in proverrucids and verrucids); m, myophore; MS, MT, movable scutum and tergum; sam, scutal adductor muscle; S, T, normal terga and scuta in brachylepadomorphs; dashed horizontal lines: membranous basis (transverse sections F-H after Young 1998)." figureDoi="http://doi.org/10.5281/zenodo.13988759" httpUri="https://zenodo.org/record/13988759/files/figure.png" pageId="6" pageNumber="77">4C</figureCitation>
|
||||
), as they are in brachylepadomorphs (
|
||||
<bibRefCitation id="EFB97BF0FFB1D025FE8CFE647A7B9E89" author="Newman W. A." box="[369,545,454,480]" editor="Southward A. J." pageId="6" pageNumber="77" pagination="3 - 42" refId="ref7098" refString="Newman W. A. 1987. - Evolution of Cirripedes and their major groups: 3 - 42, in Southward A. J. (ed.), Barnacle Biology, Crustacean Issues 5. A. A. Balkema, Rotterdam." title="Evolution of Cirripedes and their major groups" type="book chapter" volumeTitle="Barnacle Biology, Crustacean Issues 5. A. A. Balkema, Rotterdam" year="1987">Newman 1987</bibRefCitation>
|
||||
: fig. 4A) and in the juveniles of
|
||||
<taxonomicName id="4C287D82FFB1D025FE85FE4679AA9E94" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[376,496,484,509]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB1D025FE85FE4679AA9E94" box="[376,496,484,509]" italics="true" pageId="6" pageNumber="77">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EFB97BF0FFB1D025FDFDFE4779589D74" author="Newman W. A. & Hessler R. R." journalOrPublisher="San Diego Society of Natural History Transactions" pageId="6" pageNumber="77" pagination="259 - 273" part="21" refId="ref7233" refString="Newman W. A. & Hessler R. R. 1989. - A new abyssal hydrothermal verrucomorphan (Cirripedia; Sessilia): The most primitive living sessile barnacle. San Diego Society of Natural History Transactions 21 (16): 259 - 273." title="A new abyssal hydrothermal verrucomorphan (Cirripedia; Sessilia): The most primitive living sessile barnacle" type="journal article" year="1989">Newman & Hessler 1989</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFB97BF0FFB1D025FEF0FDA079E99D74" author="Newman W. A." box="[269,435,514,541]" journalOrPublisher="Bulletin of Marine Science" pageId="6" pageNumber="77" pagination="467 - 477" part="45" refId="ref7144" refString="Newman W. A. 1989. - Juvenile ontogeny and metamorphosis in the most primitive living sessile barnacle, Neoverruca, from an abyssal hydrothermal spring. Bulletin of Marine Science 45 (2): 467 - 477." title="Juvenile ontogeny and metamorphosis in the most primitive living sessile barnacle, Neoverruca, from an abyssal hydrothermal spring" type="journal article" year="1989">Newman 1989</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF575689FFB3D027FF8FFACA7B1899CD" ID-DOI="http://doi.org/10.5281/zenodo.5391399" ID-Zenodo-Dep="5391399" httpUri="https://zenodo.org/record/5391399/files/figure.png" pageId="4" pageNumber="75" startId="4.[114,125,1384,1401]" targetBox="[213,1067,193,1353]" targetPageId="4">
|
||||
<paragraph id="8B970601FFB3D027FF8FFACA7B1899CD" blockId="4.[114,1211,1384,1700]" pageId="4" pageNumber="75">
|
||||
FIG. 2. — Plates of
|
||||
<taxonomicName id="4C287D82FFB3D027FEF3FACA79B69A10" authorityName="Newman" authorityYear="2000" box="[270,492,1384,1401]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="4" pageNumber="75" phylum="Arthropoda" rank="species" species="yamaguchii">
|
||||
<emphasis id="B95CDA13FFB3D027FEF3FACA79B69A10" box="[270,492,1384,1401]" italics="true" pageId="4" pageNumber="75">Imbricaverruca yamaguchii</emphasis>
|
||||
</taxonomicName>
|
||||
gen. et n. sp. (paratype, MNHN Ci2711); A, B, viewed from above and from the rostral end, respectively;
|
||||
<emphasis id="B95CDA13FFB3D027FEF6FADF79429AE7" bold="true" box="[267,280,1405,1422]" pageId="4" pageNumber="75">A</emphasis>
|
||||
, note the operculum, comprising the plates of the right side (MS-L-MT), includes a large median latus;
|
||||
<emphasis id="B95CDA13FFB3D027FB81FADF7CD39AE7" bold="true" box="[1148,1161,1405,1422]" pageId="4" pageNumber="75">B</emphasis>
|
||||
, note the slightly open aperture to the mantle cavity between the occludent margins of the movable (MS & MT) and the fixed (FS & FT) scutum and tergum, respectively; C-M, variously disarticulated hard parts;
|
||||
<emphasis id="B95CDA13FFB3D027FD31FA0A7A809AD0" bold="true" box="[716,730,1448,1465]" pageId="4" pageNumber="75">C</emphasis>
|
||||
, interior view of wall (R-C-FS-FT) and operculum (MS-L-MT) with all of the imbricating plates, except the uppermost of the three principle lateral tiers of the right side (rl
|
||||
<superScript id="7C5DAB49FFB3D027FBD1FA187C6E9AAE" attach="left" box="[1068,1076,1466,1479]" fontSize="5" pageId="4" pageNumber="75">1</superScript>
|
||||
-l
|
||||
<superScript id="7C5DAB49FFB3D027FBC2FA187C1D9AAE" attach="left" box="[1087,1095,1466,1479]" fontSize="5" pageId="4" pageNumber="75">1</superScript>
|
||||
-cl
|
||||
<superScript id="7C5DAB49FFB3D027FBA1FA187C3E9AAE" attach="left" box="[1116,1124,1466,1479]" fontSize="5" pageId="4" pageNumber="75">1</superScript>
|
||||
), stripped away [an approximation of the rostrum (R), which did not survive dissection, is indicated by the dashed line];
|
||||
<emphasis id="B95CDA13FFB3D027FBFDFA717C549A8D" bold="true" box="[1024,1038,1491,1508]" pageId="4" pageNumber="75">D</emphasis>
|
||||
,
|
||||
<emphasis id="B95CDA13FFB3D027FBE5FA717C7E9A8D" bold="true" box="[1048,1060,1491,1508]" pageId="4" pageNumber="75">E</emphasis>
|
||||
, movable scutum (MS); external and internal views respectively; scutal adductor muscle, inserting from just below to up under the apical concavity in Fig. 2E, extends to its origin on the fixed scutum (FS, Fig. 2G; the relationship of MS to FS can be seen in Fig 2C);
|
||||
<emphasis id="B95CDA13FFB3D027FBDCFA5F7C769967" bold="true" box="[1057,1068,1533,1550]" pageId="4" pageNumber="75">F</emphasis>
|
||||
,
|
||||
<emphasis id="B95CDA13FFB3D027FBCBFA5F7C1E9967" bold="true" box="[1078,1092,1533,1550]" pageId="4" pageNumber="75">G</emphasis>
|
||||
, fixed scutum (FS); basal and internal views, respectively. Dashed line in G indicates origin of scutal adductor muscle but no scar is evident. The relationship of the two protuberances (best seen near the tergal margin in Fig. 2G) to the fixed tergum (FT, Fig. 2J, K), was not resolved;
|
||||
<emphasis id="B95CDA13FFB3D027FF3EF99F788B9927" bold="true" box="[195,209,1597,1614]" pageId="4" pageNumber="75">H</emphasis>
|
||||
,
|
||||
<emphasis id="B95CDA13FFB3D027FF26F99F78BB9927" bold="true" box="[219,225,1597,1614]" pageId="4" pageNumber="75">I</emphasis>
|
||||
, movable tergum (MT); external and internal views respectively (external corrosion in bleach and cleaning inadvertently obliterated growth line ornamentation in H);
|
||||
<emphasis id="B95CDA13FFB3D027FE02F9F17A53990D" bold="true" box="[511,521,1619,1636]" pageId="4" pageNumber="75">J</emphasis>
|
||||
,
|
||||
<emphasis id="B95CDA13FFB3D027FDE8F9F17A78990D" bold="true" box="[533,546,1619,1636]" pageId="4" pageNumber="75">K</emphasis>
|
||||
, fixed tergum (FT); external and internal views respectively;
|
||||
<emphasis id="B95CDA13FFB3D027FBD8F9F17C6A990D" bold="true" box="[1061,1072,1619,1636]" pageId="4" pageNumber="75">L</emphasis>
|
||||
,
|
||||
<emphasis id="B95CDA13FFB3D027FBC1F9F17C17990D" bold="true" box="[1084,1101,1619,1636]" pageId="4" pageNumber="75">M</emphasis>
|
||||
, carina from below and above respectively (the rostrum did not survive dissection and cleaning). Abbreviations:
|
||||
<emphasis id="B95CDA13FFB3D027FC57F9CA7BE29910" bold="true" box="[938,952,1640,1657]" pageId="4" pageNumber="75">C</emphasis>
|
||||
, carina;
|
||||
<emphasis id="B95CDA13FFB3D027FC02F9CA7C7E991B" bold="true" box="[1023,1060,1637,1657]" pageId="4" pageNumber="75">
|
||||
cl
|
||||
<superScript id="7C5DAB49FFB3D027FBF2F9C77C7E991B" attach="left" box="[1039,1060,1637,1650]" fontSize="5" pageId="4" pageNumber="75">1-4</superScript>
|
||||
</emphasis>
|
||||
,
|
||||
<emphasis id="B95CDA13FFB3D027FBD2F9CA7C13991B" bold="true" box="[1071,1097,1637,1657]" pageId="4" pageNumber="75">
|
||||
l
|
||||
<superScript id="7C5DAB49FFB3D027FBCEF9C77C13991B" attach="left" box="[1075,1097,1637,1650]" fontSize="5" pageId="4" pageNumber="75">1-4</superScript>
|
||||
</emphasis>
|
||||
,
|
||||
<emphasis id="B95CDA13FFB3D027FBAEF9CA7C2F991B" bold="true" box="[1107,1141,1637,1657]" pageId="4" pageNumber="75">
|
||||
rl
|
||||
<superScript id="7C5DAB49FFB3D027FBA2F9C77C2F991B" attach="left" box="[1119,1141,1637,1650]" fontSize="5" pageId="4" pageNumber="75">1-4</superScript>
|
||||
</emphasis>
|
||||
, carinolateral, lateral and rostrolateral plates forming three tiers each four plates high, respectively;
|
||||
<emphasis id="B95CDA13FFB3D027FC68F9DF7BF699E7" bold="true" box="[917,940,1661,1678]" pageId="4" pageNumber="75">FS</emphasis>
|
||||
,
|
||||
<emphasis id="B95CDA13FFB3D027FC45F9DF7B9599E7" bold="true" box="[952,975,1661,1678]" pageId="4" pageNumber="75">FT</emphasis>
|
||||
, fixed scutum and tergum;
|
||||
<emphasis id="B95CDA13FFB3D027FF8FF931782799CD" bold="true" box="[114,125,1683,1700]" pageId="4" pageNumber="75">L</emphasis>
|
||||
, median latus;
|
||||
<emphasis id="B95CDA13FFB3D027FF03F931794199CD" bold="true" box="[254,283,1683,1700]" pageId="4" pageNumber="75">MS</emphasis>
|
||||
,
|
||||
<emphasis id="B95CDA13FFB3D027FEDBF931791899CD" bold="true" box="[294,322,1683,1700]" pageId="4" pageNumber="75">MT</emphasis>
|
||||
, movable scutum and tergum;
|
||||
<emphasis id="B95CDA13FFB3D027FDBBF9317A0999CD" bold="true" box="[582,595,1683,1700]" pageId="4" pageNumber="75">R</emphasis>
|
||||
, rostrum. Scale bars: 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF575689FFB2D026FF8CF98A7B2A99CD" ID-DOI="http://doi.org/10.5281/zenodo.5391401" ID-Zenodo-Dep="5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="5" pageNumber="76" startId="5.[113,124,1576,1593]" targetBox="[92,1284,200,1546]" targetPageId="5">
|
||||
<paragraph id="8B970601FFB2D026FF8CF98A7B2A99CD" blockId="5.[113,1210,1576,1700]" pageId="5" pageNumber="76">
|
||||
FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of
|
||||
<taxonomicName id="4C287D82FFB2D026FF8CF99F790A9927" authority="Newman, 2000" authorityName="Newman" authorityYear="2000" box="[113,336,1597,1614]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="76" phylum="Arthropoda" rank="species" species="yamaguchii" status="n. gen. and n. sp.">
|
||||
<emphasis id="B95CDA13FFB2D026FF8CF99F790A9927" box="[113,336,1597,1614]" italics="true" pageId="5" pageNumber="76">Imbricaverruca yamaguchii</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A26F6768FFB2D026FEA8F99F79BC9927" box="[341,486,1597,1614]" pageId="5" pageNumber="76" rank="species">n. gen. and n. sp.</taxonomicNameLabel>
|
||||
(paratype, MNHN Ci2711);
|
||||
<emphasis id="B95CDA13FFB2D026FD34F99F7A8C9927" bold="true" box="[713,726,1597,1614]" pageId="5" pageNumber="76">A</emphasis>
|
||||
, cirrus I, anterior ramus missing;
|
||||
<emphasis id="B95CDA13FFB2D026FC17F99F7BAD9927" bold="true" box="[1002,1015,1597,1614]" pageId="5" pageNumber="76">B</emphasis>
|
||||
, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right);
|
||||
<emphasis id="B95CDA13FFB2D026FCADF9F17B04990D" bold="true" box="[848,862,1619,1636]" pageId="5" pageNumber="76">C</emphasis>
|
||||
, cirrus VI (setae of distal half omitted) and penis (annulations omitted);
|
||||
<emphasis id="B95CDA13FFB2D026FEA6F9CA79339910" bold="true" box="[347,361,1640,1657]" pageId="5" pageNumber="76">D</emphasis>
|
||||
, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end);
|
||||
<emphasis id="B95CDA13FFB2D026FB92F9CA7C219910" bold="true" box="[1135,1147,1640,1657]" pageId="5" pageNumber="76">E</emphasis>
|
||||
, end of penis enlarged;
|
||||
<emphasis id="B95CDA13FFB2D026FF05F9DF795999E7" bold="true" box="[248,259,1661,1678]" pageId="5" pageNumber="76">F</emphasis>
|
||||
, labrum and mandibular palps;
|
||||
<emphasis id="B95CDA13FFB2D026FDEDF9DF7A6299E7" bold="true" box="[528,568,1661,1678]" pageId="5" pageNumber="76">G, H</emphasis>
|
||||
, right and left mandible;
|
||||
<emphasis id="B95CDA13FFB2D026FCF6F9DF7B4B99E7" bold="true" box="[779,785,1661,1678]" pageId="5" pageNumber="76">I</emphasis>
|
||||
, spinous margin of H enlarged;
|
||||
<emphasis id="B95CDA13FFB2D026FBDCF9DF7C1999E7" bold="true" box="[1057,1091,1661,1678]" pageId="5" pageNumber="76">J, K</emphasis>
|
||||
, right and left first maxillae;
|
||||
<emphasis id="B95CDA13FFB2D026FF19F93178B599CD" bold="true" box="[228,239,1683,1700]" pageId="5" pageNumber="76">L</emphasis>
|
||||
, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B970601FFB1D025FF8FFD9D78889D30" blockId="6.[114,647,575,1247]" box="[114,210,575,601]" pageId="6" pageNumber="77">
|
||||
<emphasis id="B95CDA13FFB1D025FF8FFD9D78889D30" box="[114,210,575,601]" italics="true" pageId="6" pageNumber="77">Soft parts</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8B970601FFB1D025FF8FFDFD7A569BB6" blockId="6.[114,647,575,1247]" pageId="6" pageNumber="77">
|
||||
The trophi and cirri (
|
||||
<figureCitation id="13131A84FFB1D025FE9CFDFD79F89D10" box="[353,418,607,633]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="6" pageNumber="77">Fig. 3</figureCitation>
|
||||
) are similar to those of other hydrothermal vent barnacles (
|
||||
<taxonomicName id="4C287D82FFB1D025FDD1FDDC78C19DBC" authority=", Brachylepadomorpha, Verrucomorpha" authorityName="Brachylepadomorpha, Verrucomorpha" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scalpellomorpha" pageId="6" pageNumber="77" phylum="Arthropoda" rank="order">Scalpellomorpha, Brachylepadomorpha, Verrucomorpha</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C287D82FFB1D025FF2BFD1979259DBC" box="[214,383,699,725]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Balanomorpha" pageId="6" pageNumber="77" phylum="Arthropoda" rank="order">Balanomorpha</taxonomicName>
|
||||
), reflecting adaptations to feeding on very fine particles, presumably small, suspended bacterial clumps and filaments (cf.
|
||||
<bibRefCitation id="EFB97BF0FFB1D025FF62FCB579B19C58" author="Southward A. J. & Newman W. A." box="[159,491,790,817]" journalOrPublisher="Cahiers de Biologie marine" pageId="6" pageNumber="77" pagination="258 - 262" part="39" refId="ref7585" refString="Southward A. J. & Newman W. A. 1998. - Ectosymbiosis between filamentous sulphur bacteria and a stalked barnacle (Scalpellomorpha, Neolepadinae) from the Lau Back Arc Basin, Tonga. Cahiers de Biologie marine 39 (3 - 4): 258 - 262." title="Ectosymbiosis between filamentous sulphur bacteria and a stalked barnacle (Scalpellomorpha, Neolepadinae) from the Lau Back Arc Basin, Tonga" type="journal article" year="1998">Southward & Newman 1998</bibRefCitation>
|
||||
). The slightly bullate, indented but unnotched labrum of
|
||||
<taxonomicName id="4C287D82FFB1D025FF8FFCF6795A9C06" authorityName="Newman" authorityYear="2000" box="[114,256,852,879]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="yamaguchii">
|
||||
<emphasis id="B95CDA13FFB1D025FF8FFCF6795A9C06" box="[114,256,852,879]" italics="true" pageId="6" pageNumber="77">I. yamaguchii</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="13131A84FFB1D025FEEDFCF7793A9C06" box="[272,352,853,879]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="6" pageNumber="77">Fig. 3F</figureCitation>
|
||||
) supports the usual row of fine teeth and relatively large palps provided with relatively fine, stiff setae. The mandible (
|
||||
<figureCitation id="13131A84FFB1D025FDD6FC307AD89CC5" box="[555,642,914,940]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="6" pageNumber="77">Fig. 3G</figureCitation>
|
||||
, right exterior, H, I, left interior) is comparable to that of other hydrothermal vent forms, especially that of
|
||||
<taxonomicName id="4C287D82FFB1D025FF38FC4F7A579B61" authorityName="Newman" authorityYear="1989" box="[197,525,1005,1032]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="brachylepadoformis">
|
||||
<emphasis id="B95CDA13FFB1D025FF38FC4F7A579B61" box="[197,525,1005,1032]" italics="true" pageId="6" pageNumber="77">Neoverruca brachylepadoformis</emphasis>
|
||||
</taxonomicName>
|
||||
, in having one superior incisiform tooth above a setose area, followed by two low teeth each supporting a row of numerous, short, sharp spines along their superior curvatures, several of the uppermost of which roll over onto the inner or posterior surface (
|
||||
<figureCitation id="13131A84FFB1D025FF87FB0478939BA9" box="[122,201,1190,1216]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="6" pageNumber="77">Fig. 3I</figureCitation>
|
||||
), and the upper margin of the inferior angle supports a row of similar spines.
|
||||
</paragraph>
|
||||
<paragraph id="8B970601FFB1D025FD58FF737CFA9EED" blockId="6.[677,1210,208,603]" pageId="6" pageNumber="77">
|
||||
Maxillae I and II (
|
||||
<figureCitation id="13131A84FFB1D025FC91FF737C7F9F83" box="[876,1061,208,235]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="6" pageNumber="77">Fig. 3 J, K and L</figureCitation>
|
||||
, respectively) each support a relatively uniform comb of slender setae along the cutting edge, but maxillae I of
|
||||
<taxonomicName id="4C287D82FFB1D025FD58FE8E7B6F9E2E" authorityName="Newman" authorityYear="2000" box="[677,821,300,327]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="yamaguchii">
|
||||
<emphasis id="B95CDA13FFB1D025FD58FE8E7B6F9E2E" box="[677,821,300,327]" italics="true" pageId="6" pageNumber="77">I. yamaguchii</emphasis>
|
||||
</taxonomicName>
|
||||
differs from
|
||||
<taxonomicName id="4C287D82FFB1D025FC3EFE8E7C619E2C" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[963,1083,300,325]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB1D025FC3EFE8E7C619E2C" box="[963,1083,300,325]" italics="true" pageId="6" pageNumber="77">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
in having a single strong spine among the ordinary setae inside and just below the superior angle (
|
||||
<figureCitation id="13131A84FFB1D025FBC2FEC87CC99EED" box="[1087,1171,362,388]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="6" pageNumber="77">Fig. 3K</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B970601FFB1D025FD58FE2B7C299D32" blockId="6.[677,1210,208,603]" pageId="6" pageNumber="77">
|
||||
What could be seen of the cirri (
|
||||
<figureCitation id="13131A84FFB1D025FBF6FE2B7C139ECA" box="[1035,1097,393,419]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="6" pageNumber="77">Fig. 3</figureCitation>
|
||||
A-D) suggested they were typical of most vent barnacles, but unfortunately those of the
|
||||
<typeStatus id="5493B8A3FFB1D025FBEAFE677C219EB6" box="[1047,1147,453,479]" pageId="6" pageNumber="77" type="paratype">paratype</typeStatus>
|
||||
were badly damaged or missing. The right side was in the best condition and the cirral counts are as follows (a, anterior ramus; p, posterior ramus; –, ramus absent; +, terminal articles missing):
|
||||
</paragraph>
|
||||
<paragraph id="8B970601FFB1D025FD58FD177CEA9C7E" pageId="6" pageNumber="77">
|
||||
<table id="F928F4A1FFB12FDCFD58FD177CEA9C76" box="[677,1200,693,799]" gridcols="7" gridrows="3" pageId="6" pageNumber="77">
|
||||
<tr id="35180443FFB12FDCFD58FD177CEA9DA0" box="[677,1200,693,713]" gridrow="0" pageId="6" pageNumber="77">
|
||||
<th id="76C96D3FFFB12FDCFD58FD177A879DA0" box="[677,733,693,713]" gridcol="0" gridrow="0" pageId="6" pageNumber="77">
|
||||
<emphasis id="B95CDA13FFB1D025FD58FD177A879DA0" bold="true" box="[677,733,693,713]" pageId="6" pageNumber="77">Cirrus</emphasis>
|
||||
</th>
|
||||
<th id="76C96D3FFFB12FDCFCF5FD177B459DA0" box="[776,799,693,713]" gridcol="1" gridrow="0" pageId="6" pageNumber="77">
|
||||
<emphasis id="B95CDA13FFB1D025FCF5FD177B549DA0" bold="true" box="[776,782,693,713]" pageId="6" pageNumber="77">I</emphasis>
|
||||
</th>
|
||||
<th id="76C96D3FFFB12FDCFCABFD177B379DA0" box="[854,877,693,713]" gridcol="2" gridrow="0" pageId="6" pageNumber="77">
|
||||
<emphasis id="B95CDA13FFB1D025FCABFD177B3B9DA0" bold="true" box="[854,865,693,713]" pageId="6" pageNumber="77">II</emphasis>
|
||||
</th>
|
||||
<th id="76C96D3FFFB12FDCFC5BFD177BED9DA0" box="[934,951,693,713]" gridcol="3" gridrow="0" pageId="6" pageNumber="77">
|
||||
<emphasis id="B95CDA13FFB1D025FC5BFD177BED9DA0" bold="true" box="[934,951,693,713]" pageId="6" pageNumber="77">III</emphasis>
|
||||
</th>
|
||||
<th id="76C96D3FFFB12FDCFC08FD177C4E9DA0" box="[1013,1044,693,713]" gridcol="4" gridrow="0" pageId="6" pageNumber="77">
|
||||
<emphasis id="B95CDA13FFB1D025FC08FD177C529DA0" bold="true" box="[1013,1032,693,713]" pageId="6" pageNumber="77">IV</emphasis>
|
||||
</th>
|
||||
<th id="76C96D3FFFB12FDCFBBEFD177C0A9DA0" box="[1091,1104,693,713]" gridcol="5" gridrow="0" pageId="6" pageNumber="77">
|
||||
<emphasis id="B95CDA13FFB1D025FBBEFD177C0A9DA0" bold="true" box="[1091,1104,693,713]" pageId="6" pageNumber="77">V</emphasis>
|
||||
</th>
|
||||
<th id="76C96D3FFFB12FDCFB6CFD177CEA9DA0" box="[1169,1200,693,713]" gridcol="6" gridrow="0" pageId="6" pageNumber="77">
|
||||
<emphasis id="B95CDA13FFB1D025FB6CFD177CFE9DA0" bold="true" box="[1169,1188,693,713]" pageId="6" pageNumber="77">VI</emphasis>
|
||||
</th>
|
||||
</tr>
|
||||
<tr id="35180443FFB12FDCFD58FD4A7CEA9D96" box="[677,1200,744,767]" gridrow="1" pageId="6" pageNumber="77">
|
||||
<th id="76C96D3FFFB12FDCFD58FD4A7A879D96" box="[677,733,744,767]" gridcol="0" gridrow="1" pageId="6" pageNumber="77">a</th>
|
||||
<td id="76C96D3FFFB12FDCFCF5FD4A7B459D96" box="[776,799,744,767]" gridcol="1" gridrow="1" pageId="6" pageNumber="77">–</td>
|
||||
<td id="76C96D3FFFB12FDCFCABFD4A7B379D96" box="[854,877,744,767]" gridcol="2" gridrow="1" pageId="6" pageNumber="77">7</td>
|
||||
<td id="76C96D3FFFB12FDCFC5BFD4A7BED9D96" box="[934,951,744,767]" gridcol="3" gridrow="1" pageId="6" pageNumber="77">–</td>
|
||||
<td id="76C96D3FFFB12FDCFC08FD4A7C4E9D96" box="[1013,1044,744,767]" gridcol="4" gridrow="1" pageId="6" pageNumber="77">–</td>
|
||||
<td id="76C96D3FFFB12FDCFBBEFD4A7C0A9D96" box="[1091,1104,744,767]" gridcol="5" gridrow="1" pageId="6" pageNumber="77">–</td>
|
||||
<td id="76C96D3FFFB12FDCFB6CFD4A7CEA9D96" box="[1169,1200,744,767]" gridcol="6" gridrow="1" pageId="6" pageNumber="77">37+</td>
|
||||
</tr>
|
||||
<tr id="35180443FFB12FDCFD58FCAA7CEA9C76" box="[677,1200,776,799]" gridrow="2" pageId="6" pageNumber="77">
|
||||
<th id="76C96D3FFFB12FDCFD58FCAA7A879C76" box="[677,733,776,799]" gridcol="0" gridrow="2" pageId="6" pageNumber="77">p</th>
|
||||
<td id="76C96D3FFFB12FDCFCF5FCAA7B459C76" box="[776,799,776,799]" gridcol="1" gridrow="2" pageId="6" pageNumber="77">20</td>
|
||||
<td id="76C96D3FFFB12FDCFCABFCAA7B379C76" box="[854,877,776,799]" gridcol="2" gridrow="2" pageId="6" pageNumber="77">13</td>
|
||||
<td id="76C96D3FFFB12FDCFC5BFCAA7BED9C76" box="[934,951,776,799]" gridcol="3" gridrow="2" pageId="6" pageNumber="77">–</td>
|
||||
<td id="76C96D3FFFB12FDCFC08FCAA7C4E9C76" box="[1013,1044,776,799]" gridcol="4" gridrow="2" pageId="6" pageNumber="77">39+</td>
|
||||
<td id="76C96D3FFFB12FDCFBBEFCAA7C0A9C76" box="[1091,1104,776,799]" gridcol="5" gridrow="2" pageId="6" pageNumber="77">–</td>
|
||||
<td id="76C96D3FFFB12FDCFB6CFCAA7CEA9C76" box="[1169,1200,776,799]" gridcol="6" gridrow="2" pageId="6" pageNumber="77">41+</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<paragraph id="8B970601FFB1D025FD58FC307C079BB6" blockId="6.[677,1210,913,1247]" pageId="6" pageNumber="77">
|
||||
As can be observed (
|
||||
<figureCitation id="13131A84FFB1D025FC81FC307BB79CC2" box="[892,1005,913,940]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="6" pageNumber="77">Fig. 3A, B</figureCitation>
|
||||
), the first two pairs of cirri are modified as maxillipeds and at least the posterior ramus of the first as well as of the second are antenniform. The posterior four pairs of cirri are normal (ctenopod), but the lesser curvature of the intermediate articles of cirrus VI each have as many as five pairs of moderately long setae, fewer than
|
||||
<taxonomicName id="4C287D82FFB1D025FD21FBCA7B089BE8" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[732,850,1128,1153]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB1D025FD21FBCA7B089BE8" box="[732,850,1128,1153]" italics="true" pageId="6" pageNumber="77">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
, and their s/a ratios are markedly different, that for
|
||||
<taxonomicName id="4C287D82FFB1D025FC9CFB257BA59BC8" authorityName="Newman" authorityYear="2000" box="[865,1023,1159,1185]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB1D025FC9CFB257BA59BC8" box="[865,1023,1159,1185]" italics="true" pageId="6" pageNumber="77">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
falling within the range for other vent barnacles while that for
|
||||
<taxonomicName id="4C287D82FFB1D025FD58FB667B409BB4" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[677,794,1220,1245]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB1D025FD58FB667B409BB4" box="[677,794,1220,1245]" italics="true" pageId="6" pageNumber="77">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
is the lowest known (
|
||||
<tableCitation id="C6AA33BAFFB1D025FC00FB677C179BB6" box="[1021,1101,1221,1247]" captionStart="TABLE" captionStartId="6.[114,125,1316,1333]" captionTargetPageId="6" captionText="TABLE 1. — Seta/article ratios for four neolepadines compared to four sessile barnacles also from vents: the ratio (the width of the article of an intermediate segment of cirrus VI as the denominator and the length of longest seta of that article as the numerator), while highest in the bacteria-farming neolepadine (Lau sp. A), is lowest in Neoverruca. That for Imbricaverruca as well as for Neobrachylepas are not only similar to each other but fall in the low end of the range for vent barnacles." pageId="6" pageNumber="77">Table 1</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF575689FFB1D025FF8FFA867B8B9A1C" pageId="6" pageNumber="77" startId="6.[114,125,1316,1333]" targetBox="[114,1177,1432,1687]" targetIsTable="true" targetPageId="6">
|
||||
<paragraph id="8B970601FFB1D025FF8FFA867B8B9A1C" blockId="6.[114,1211,1316,1397]" pageId="6" pageNumber="77">
|
||||
TABLE 1. — Seta/article ratios for four neolepadines compared to four sessile barnacles also from vents: the ratio (the width of the article of an intermediate segment of cirrus VI as the denominator and the length of longest seta of that article as the numerator), while highest in the bacteria-farming neolepadine (Lau sp. A), is lowest in
|
||||
<taxonomicName id="4C287D82FFB1D025FCFCFAED7B3E9A09" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[769,868,1359,1376]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB1D025FCFCFAED7B3E9A09" box="[769,868,1359,1376]" italics="true" pageId="6" pageNumber="77">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
. That for
|
||||
<taxonomicName id="4C287D82FFB1D025FC41FAED7C669A09" authorityName="Newman" authorityYear="2000" box="[956,1084,1359,1376]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB1D025FC41FAED7C669A09" box="[956,1084,1359,1376]" italics="true" pageId="6" pageNumber="77">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
as well as for
|
||||
<taxonomicName id="4C287D82FFB1D025FF8FFAC678A29A1C" authorityName="Newman & Yamaguchi" authorityYear="1995" box="[114,248,1380,1397]" class="Maxillopoda" family="Neobrachylepadidae" genus="Neobrachylepas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB1D025FF8FFAC678A29A1C" box="[114,248,1380,1397]" italics="true" pageId="6" pageNumber="77">Neobrachylepas</emphasis>
|
||||
</taxonomicName>
|
||||
are not only similar to each other but fall in the low end of the range for vent barnacles.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B970601FFB1D025FE2BFA3A7CC399FE" pageId="6" pageNumber="77">
|
||||
<table id="F928F4A1FFB12FDCFF8FFA3A7CC399FE" box="[114,1177,1432,1687]" gridcols="5" gridrows="10" pageId="6" pageNumber="77">
|
||||
<tr id="35180443FFB12FDCFF8FFA3A7CC39AC5" box="[114,1177,1432,1452]" gridrow="0" pageId="6" pageNumber="77" rowspan-0="1">
|
||||
<th id="76C96D3FFFB12FDCFE62FA3A7A949AC5" box="[415,718,1432,1452]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="6" pageNumber="77">
|
||||
<emphasis id="B95CDA13FFB1D025FE2BFA3A7AFD9AC5" bold="true" box="[470,679,1432,1452]" pageId="6" pageNumber="77">Measurement in mm</emphasis>
|
||||
</th>
|
||||
<th id="76C96D3FFFB12FDCFCF0FA3A7B199AC5" box="[781,835,1432,1452]" gridcol="3" gridrow="0" pageId="6" pageNumber="77">
|
||||
<emphasis id="B95CDA13FFB1D025FCF0FA3A7B199AC5" bold="true" box="[781,835,1432,1452]" pageId="6" pageNumber="77">Ratio</emphasis>
|
||||
</th>
|
||||
<th id="76C96D3FFFB12FDCFC76FA3A7CC39AC5" box="[907,1177,1432,1452]" gridcol="4" gridrow="0" pageId="6" pageNumber="77">
|
||||
<emphasis id="B95CDA13FFB1D025FC23FA3A7C1D9AC5" bold="true" box="[990,1095,1432,1452]" pageId="6" pageNumber="77">Reference</emphasis>
|
||||
</th>
|
||||
</tr>
|
||||
<tr id="35180443FFB12FDCFF8FFA127CC39AAD" box="[114,1177,1456,1476]" gridrow="1" pageId="6" pageNumber="77" rowspan-0="1" rowspan-4="1">
|
||||
<td id="76C96D3FFFB12FDCFE62FA127A469AAD" box="[415,540,1456,1476]" gridcol="1" gridrow="1" pageId="6" pageNumber="77">
|
||||
<emphasis id="B95CDA13FFB1D025FE62FA127A469AAD" bold="true" box="[415,540,1456,1476]" pageId="6" pageNumber="77">article width</emphasis>
|
||||
</td>
|
||||
<td id="76C96D3FFFB12FDCFDAAFA127A949AAD" box="[599,718,1456,1476]" gridcol="2" gridrow="1" pageId="6" pageNumber="77">
|
||||
<emphasis id="B95CDA13FFB1D025FDAAFA127A949AAD" bold="true" box="[599,718,1456,1476]" pageId="6" pageNumber="77">setal length</emphasis>
|
||||
</td>
|
||||
<td id="76C96D3FFFB12FDCFCF0FA127B199AAD" box="[781,835,1456,1476]" gridcol="3" gridrow="1" pageId="6" pageNumber="77">
|
||||
<emphasis id="B95CDA13FFB1D025FCE4FA127B629AAD" bold="true" box="[793,824,1456,1476]" pageId="6" pageNumber="77">s/a</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35180443FFB12FDCFF8FFA797CC39A86" box="[114,1177,1499,1519]" gridrow="2" pageId="6" pageNumber="77">
|
||||
<th id="76C96D3FFFB12FDCFF8FFA7979009A86" box="[114,346,1499,1519]" gridcol="0" gridrow="2" pageId="6" pageNumber="77">Lau sp. A</th>
|
||||
<td id="76C96D3FFFB12FDCFE62FA797A469A86" box="[415,540,1499,1519]" gridcol="1" gridrow="2" pageId="6" pageNumber="77">0.33</td>
|
||||
<td id="76C96D3FFFB12FDCFDAAFA797A949A86" box="[599,718,1499,1519]" gridcol="2" gridrow="2" pageId="6" pageNumber="77">4.60</td>
|
||||
<td id="76C96D3FFFB12FDCFCF0FA797B199A86" box="[781,835,1499,1519]" gridcol="3" gridrow="2" pageId="6" pageNumber="77">14.0</td>
|
||||
<td id="76C96D3FFFB12FDCFC76FA797CC39A86" box="[907,1177,1499,1519]" gridcol="4" gridrow="2" pageId="6" pageNumber="77">
|
||||
<bibRefCitation id="EFB97BF0FFB1D025FC70FA797CCD9A86" author="Southward A. J. & Newman W. A." box="[909,1175,1499,1519]" journalOrPublisher="Cahiers de Biologie marine" pageId="6" pageNumber="77" pagination="258 - 262" part="39" refId="ref7585" refString="Southward A. J. & Newman W. A. 1998. - Ectosymbiosis between filamentous sulphur bacteria and a stalked barnacle (Scalpellomorpha, Neolepadinae) from the Lau Back Arc Basin, Tonga. Cahiers de Biologie marine 39 (3 - 4): 258 - 262." title="Ectosymbiosis between filamentous sulphur bacteria and a stalked barnacle (Scalpellomorpha, Neolepadinae) from the Lau Back Arc Basin, Tonga" type="journal article" year="1998">Southward & Newman 1998</bibRefCitation>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35180443FFB12FDCFF8FFA517CC3996E" box="[114,1177,1523,1543]" gridrow="3" pageId="6" pageNumber="77">
|
||||
<th id="76C96D3FFFB12FDCFF8FFA517900996E" box="[114,346,1523,1543]" gridcol="0" gridrow="3" pageId="6" pageNumber="77">Lau sp. B</th>
|
||||
<td id="76C96D3FFFB12FDCFE62FA517A46996E" box="[415,540,1523,1543]" gridcol="1" gridrow="3" pageId="6" pageNumber="77">0.10</td>
|
||||
<td id="76C96D3FFFB12FDCFDAAFA517A94996E" box="[599,718,1523,1543]" gridcol="2" gridrow="3" pageId="6" pageNumber="77">0.67</td>
|
||||
<td id="76C96D3FFFB12FDCFCF0FA517B19996E" box="[781,835,1523,1543]" gridcol="3" gridrow="3" pageId="6" pageNumber="77">6.7</td>
|
||||
<td id="76C96D3FFFB12FDCFC76FA517CC3996E" box="[907,1177,1523,1543]" gridcol="4" gridrow="3" pageId="6" pageNumber="77">
|
||||
<emphasis id="B95CDA13FFB1D025FC06FA517C70996E" box="[1019,1066,1523,1543]" italics="true" pageId="6" pageNumber="77">idem</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35180443FFB12FDCFF8FF9A97CC39976" box="[114,1177,1547,1567]" gridrow="4" pageId="6" pageNumber="77">
|
||||
<th id="76C96D3FFFB12FDCFF8FF9A979009976" box="[114,346,1547,1567]" gridcol="0" gridrow="4" pageId="6" pageNumber="77">South-East Indian Ridge</th>
|
||||
<td id="76C96D3FFFB12FDCFE62F9A97A469976" box="[415,540,1547,1567]" gridcol="1" gridrow="4" pageId="6" pageNumber="77">0.18</td>
|
||||
<td id="76C96D3FFFB12FDCFDAAF9A97A949976" box="[599,718,1547,1567]" gridcol="2" gridrow="4" pageId="6" pageNumber="77">1.70</td>
|
||||
<td id="76C96D3FFFB12FDCFCF0F9A97B199976" box="[781,835,1547,1567]" gridcol="3" gridrow="4" pageId="6" pageNumber="77">9.4</td>
|
||||
<td id="76C96D3FFFB12FDCFC76F9A97CC39976" box="[907,1177,1547,1567]" gridcol="4" gridrow="4" pageId="6" pageNumber="77">
|
||||
<emphasis id="B95CDA13FFB1D025FC06F9A97C709976" box="[1019,1066,1547,1567]" italics="true" pageId="6" pageNumber="77">idem</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35180443FFB12FDCFF8FF9817CC3995E" box="[114,1177,1571,1591]" gridrow="5" pageId="6" pageNumber="77">
|
||||
<th id="76C96D3FFFB12FDCFF8FF9817900995E" box="[114,346,1571,1591]" gridcol="0" gridrow="5" pageId="6" pageNumber="77">Lihir volcano</th>
|
||||
<td id="76C96D3FFFB12FDCFE62F9817A46995E" box="[415,540,1571,1591]" gridcol="1" gridrow="5" pageId="6" pageNumber="77">0.30</td>
|
||||
<td id="76C96D3FFFB12FDCFDAAF9817A94995E" box="[599,718,1571,1591]" gridcol="2" gridrow="5" pageId="6" pageNumber="77">1.85</td>
|
||||
<td id="76C96D3FFFB12FDCFCF0F9817B19995E" box="[781,835,1571,1591]" gridcol="3" gridrow="5" pageId="6" pageNumber="77">6.2</td>
|
||||
<td id="76C96D3FFFB12FDCFC76F9817CC3995E" box="[907,1177,1571,1591]" gridcol="4" gridrow="5" pageId="6" pageNumber="77">
|
||||
<emphasis id="B95CDA13FFB1D025FC06F9817C70995E" box="[1019,1066,1571,1591]" italics="true" pageId="6" pageNumber="77">idem</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35180443FFB12FDCFF8FF9997CC39926" box="[114,1177,1595,1615]" gridrow="6" pageId="6" pageNumber="77">
|
||||
<th id="76C96D3FFFB12FDCFF8FF99979009926" box="[114,346,1595,1615]" gridcol="0" gridrow="6" pageId="6" pageNumber="77">
|
||||
<taxonomicName id="4C287D82FFB1D025FF8FF99979569926" authorityName="Newman & Yamaguchi" authorityYear="1995" box="[114,268,1595,1615]" class="Maxillopoda" family="Neobrachylepadidae" genus="Neobrachylepas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB1D025FF8FF99979569926" box="[114,268,1595,1615]" italics="true" pageId="6" pageNumber="77">Neobrachylepas</emphasis>
|
||||
</taxonomicName>
|
||||
</th>
|
||||
<td id="76C96D3FFFB12FDCFE62F9997A469926" box="[415,540,1595,1615]" gridcol="1" gridrow="6" pageId="6" pageNumber="77">0.07</td>
|
||||
<td id="76C96D3FFFB12FDCFDAAF9997A949926" box="[599,718,1595,1615]" gridcol="2" gridrow="6" pageId="6" pageNumber="77">0.31</td>
|
||||
<td id="76C96D3FFFB12FDCFCF0F9997B199926" box="[781,835,1595,1615]" gridcol="3" gridrow="6" pageId="6" pageNumber="77">4.4</td>
|
||||
<td id="76C96D3FFFB12FDCFC76F9997CC39926" box="[907,1177,1595,1615]" gridcol="4" gridrow="6" pageId="6" pageNumber="77">
|
||||
<bibRefCitation id="EFB97BF0FFB1D025FC76F9997CC39926" author="Newman W. A. & Yamaguchi T." box="[907,1177,1595,1615]" journalOrPublisher="Bulletin du Museum national d'Histoire naturelle" pageId="6" pageNumber="77" pagination="211 - 243" part="17" refId="ref7319" refString="Newman W. A. & Yamaguchi T. 1995. - A new sessile barnacle (Cirripedia, Brachylepadomorpha) from the Lau Back-Arc Basin, Tonga; first record of a living representative since the Miocene. Bulletin du Museum national d'Histoire naturelle, 4 e serie, 17 A (3 - 4): 211 - 243." title="A new sessile barnacle (Cirripedia, Brachylepadomorpha) from the Lau Back-Arc Basin, Tonga; first record of a living representative since the Miocene" type="journal article" year="1995">Newman & Yamaguchi 1995</bibRefCitation>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35180443FFB12FDCFF8FF9F17CC3990E" box="[114,1177,1619,1639]" gridrow="7" pageId="6" pageNumber="77">
|
||||
<th id="76C96D3FFFB12FDCFF8FF9F17900990E" box="[114,346,1619,1639]" gridcol="0" gridrow="7" pageId="6" pageNumber="77">
|
||||
<taxonomicName id="4C287D82FFB1D025FF8FF9F17885990E" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[114,223,1619,1639]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB1D025FF8FF9F17885990E" box="[114,223,1619,1639]" italics="true" pageId="6" pageNumber="77">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
</th>
|
||||
<td id="76C96D3FFFB12FDCFE62F9F17A46990E" box="[415,540,1619,1639]" gridcol="1" gridrow="7" pageId="6" pageNumber="77">0.25</td>
|
||||
<td id="76C96D3FFFB12FDCFDAAF9F17A94990E" box="[599,718,1619,1639]" gridcol="2" gridrow="7" pageId="6" pageNumber="77">0.70</td>
|
||||
<td id="76C96D3FFFB12FDCFCF0F9F17B19990E" box="[781,835,1619,1639]" gridcol="3" gridrow="7" pageId="6" pageNumber="77">2.8</td>
|
||||
<td id="76C96D3FFFB12FDCFC76F9F17CC3990E" box="[907,1177,1619,1639]" gridcol="4" gridrow="7" pageId="6" pageNumber="77">
|
||||
<bibRefCitation id="EFB97BF0FFB1D025FC60F9F17CDD990E" author="Newman W. A. & Hessler R. R." box="[925,1159,1619,1639]" journalOrPublisher="San Diego Society of Natural History Transactions" pageId="6" pageNumber="77" pagination="259 - 273" part="21" refId="ref7233" refString="Newman W. A. & Hessler R. R. 1989. - A new abyssal hydrothermal verrucomorphan (Cirripedia; Sessilia): The most primitive living sessile barnacle. San Diego Society of Natural History Transactions 21 (16): 259 - 273." title="A new abyssal hydrothermal verrucomorphan (Cirripedia; Sessilia): The most primitive living sessile barnacle" type="journal article" year="1989">Newman & Hessler 1989</bibRefCitation>
|
||||
</td>
|
||||
</tr>
|
||||
<tr id="35180443FFB12FDCFF8FF9C97CC39916" box="[114,1177,1643,1663]" gridrow="8" pageId="6" pageNumber="77">
|
||||
<th id="76C96D3FFFB12FDCFF8FF9C979009916" box="[114,346,1643,1663]" gridcol="0" gridrow="8" pageId="6" pageNumber="77">
|
||||
<taxonomicName id="4C287D82FFB1D025FF8FF9C978A59916" authorityName="Newman" authorityYear="2000" box="[114,255,1643,1663]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB1D025FF8FF9C978A59916" box="[114,255,1643,1663]" italics="true" pageId="6" pageNumber="77">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
</th>
|
||||
<td id="76C96D3FFFB12FDCFE62F9C97A469916" box="[415,540,1643,1663]" gridcol="1" gridrow="8" pageId="6" pageNumber="77">0.15</td>
|
||||
<td id="76C96D3FFFB12FDCFDAAF9C97A949916" box="[599,718,1643,1663]" gridcol="2" gridrow="8" pageId="6" pageNumber="77">0.68</td>
|
||||
<td id="76C96D3FFFB12FDCFCF0F9C97B199916" box="[781,835,1643,1663]" gridcol="3" gridrow="8" pageId="6" pageNumber="77">4.5</td>
|
||||
<td id="76C96D3FFFB12FDCFC76F9C97CC39916" box="[907,1177,1643,1663]" gridcol="4" gridrow="8" pageId="6" pageNumber="77">This paper</td>
|
||||
</tr>
|
||||
<tr id="35180443FFB12FDCFF8FF9217CC399FE" box="[114,1177,1667,1687]" gridrow="9" pageId="6" pageNumber="77">
|
||||
<th id="76C96D3FFFB12FDCFF8FF921790099FE" box="[114,346,1667,1687]" gridcol="0" gridrow="9" pageId="6" pageNumber="77">
|
||||
<taxonomicName id="4C287D82FFB1D025FF8FF921791299FE" authority="(NFB)" baseAuthorityName="NFB" box="[114,328,1667,1687]" class="Maxillopoda" family="Chionelasmatidae" genus="Eochionelasmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB1D025FF8FF921795799FE" box="[114,269,1667,1687]" italics="true" pageId="6" pageNumber="77">Eochionelasmus</emphasis>
|
||||
(NFB)
|
||||
</taxonomicName>
|
||||
</th>
|
||||
<td id="76C96D3FFFB12FDCFE62F9217A4699FE" box="[415,540,1667,1687]" gridcol="1" gridrow="9" pageId="6" pageNumber="77">0.25</td>
|
||||
<td id="76C96D3FFFB12FDCFDAAF9217A9499FE" box="[599,718,1667,1687]" gridcol="2" gridrow="9" pageId="6" pageNumber="77">1.65</td>
|
||||
<td id="76C96D3FFFB12FDCFCF0F9217B1999FE" box="[781,835,1667,1687]" gridcol="3" gridrow="9" pageId="6" pageNumber="77">6.6</td>
|
||||
<td id="76C96D3FFFB12FDCFC76F9217CC399FE" box="[907,1177,1667,1687]" gridcol="4" gridrow="9" pageId="6" pageNumber="77">
|
||||
<bibRefCitation id="EFB97BF0FFB1D025FC76F9217CC399FE" author="Yamaguchi T. & Newman W. A." box="[907,1177,1667,1687]" journalOrPublisher="Pacific Science" pageId="6" pageNumber="77" pagination="135 - 155" part="44" refId="ref7889" refString="Yamaguchi T. & Newman W. A. 1990. - A new and primitive barnacle (Cirripedia; Balanomorpha) from the North Fiji Basin abyssal hydrothermal field, and its evolutionary implications. Pacific Science 44 (2): 135 - 155." title="A new and primitive barnacle (Cirripedia; Balanomorpha) from the North Fiji Basin abyssal hydrothermal field, and its evolutionary implications" type="journal article" year="1990">Yamaguchi & Newman 1990</bibRefCitation>
|
||||
</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<caption id="DF575689FFB0D024FF8CFA1F790599CD" ID-DOI="http://doi.org/10.5281/zenodo.13988759" ID-Zenodo-Dep="13988759" httpUri="https://zenodo.org/record/13988759/files/figure.png" pageId="7" pageNumber="78" startId="7.[113,141,1469,1486]" subCaptionStartIDs="7.[323,440,1597,1614]" subCaptionStarts="Abbr" targetBox="[112,1165,215,1445]" targetPageId="7">
|
||||
<paragraph id="8B970601FFB0D024FF8CFA1F790599CD" blockId="7.[113,1210,1469,1700]" pageId="7" pageNumber="78">
|
||||
Fig. 4. — Schematic plan views and transverse sections of grades of skeletal organization from the most primitive sessile barnacles (
|
||||
<emphasis id="B95CDA13FFB0D024FF8BFA7178D99A8D" bold="true" box="[118,131,1491,1508]" pageId="7" pageNumber="78">A</emphasis>
|
||||
, Brachylepadomorpha) through the asymmetrical sessile barnacles (
|
||||
<emphasis id="B95CDA13FFB0D024FD3FFA717ABE9A8D" bold="true" box="[706,740,1491,1508]" pageId="7" pageNumber="78">B -H</emphasis>
|
||||
,
|
||||
<taxonomicName id="4C287D82FFB0D024FD12FA717B2E9A8D" authorityName="Pilsbry" authorityYear="1916" box="[751,884,1491,1508]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="73" phylum="Arthropoda" rank="subOrder" subOrder="Verrucomorpha">Verrucomorpha</taxonomicName>
|
||||
). Of the eight verrucid genera currently recognized (
|
||||
<bibRefCitation id="EFB97BF0FFB0D024FF24FA4A791A9A90" author="Young P. S." box="[217,320,1512,1529]" journalOrPublisher="Zoosystema" pageId="7" pageNumber="78" pagination="31 - 92" part="20" refId="ref7981" refString="Young P. S. 1998. - Cirripedia (Crustacea) from the " Campagne Biacores " in the Azores region, including a generic revision of Verrucidae. Zoosystema 20 (1): 31 - 92." title="Cirripedia (Crustacea) from the " Campagne Biacores " in the Azores region, including a generic revision of Verrucidae" type="journal article" year="1998">Young 1998</bibRefCitation>
|
||||
) only the four most representative ones (
|
||||
<taxonomicName id="4C287D82FFB0D024FD5DFA4A7AA29A90" authorityName="Pilsbry" authorityYear="1916" box="[672,760,1512,1529]" class="Maxillopoda" family="Verrucidae" genus="Altiverruca" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB0D024FD5DFA4A7AA29A90" box="[672,760,1512,1529]" italics="true" pageId="7" pageNumber="78">Altiverruca</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C287D82FFB0D024FCF9FA4A7BD59A90" authorityName="Young" authorityYear="1998" box="[772,911,1512,1529]" class="Maxillopoda" family="Verrucidae" genus="Newmaniverruca" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB0D024FCF9FA4A7BD59A90" box="[772,911,1512,1529]" italics="true" pageId="7" pageNumber="78">Newmaniverruca</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C287D82FFB0D024FC66FA4A7C589A90" authorityName="Pilsbry" authorityYear="1916" box="[923,1026,1512,1529]" class="Maxillopoda" family="Verrucidae" genus="Metaverruca" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFB0D024FC66FA4A7C589A90" box="[923,1026,1512,1529]" italics="true" pageId="7" pageNumber="78">Metaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C287D82FFB0D024FBCDFA4A7C2A9A90" authorityName="Schumacher" authorityYear="1817" box="[1072,1136,1512,1529]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="78" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||||
<emphasis id="B95CDA13FFB0D024FBCDFA4A7C2A9A90" box="[1072,1136,1512,1529]" italics="true" pageId="7" pageNumber="78">Verruca</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A26F6768FFB0D024FB8BFA4A7CCE9A90" box="[1142,1172,1512,1529]" pageId="7" pageNumber="78" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
) are depicted here. Right sides are to the viewers right and rostral ends are downward in plan views or coming out of the page in transverse sections. The transverse sections are slightly rostral of the rostro-carinal gap or suture. While there are numerous whorls of imbricating plates standing in tiers four plates high in brachylepadomorphs, and on the movable side of neoverrucids, only those of the R-C gap are labeled. Abbreviations:
|
||||
<emphasis id="B95CDA13FFB0D024FE42F99F79979927" bold="true" box="[447,461,1597,1614]" pageId="7" pageNumber="78">C</emphasis>
|
||||
,
|
||||
<emphasis id="B95CDA13FFB0D024FE2AF99F79BE9927" bold="true" box="[471,484,1597,1614]" pageId="7" pageNumber="78">R</emphasis>
|
||||
, carina and rostrum;
|
||||
<emphasis id="B95CDA13FFB0D024FD6AF99F7AE6992E" bold="true" box="[663,700,1594,1614]" pageId="7" pageNumber="78">
|
||||
cl
|
||||
<superScript id="7C5DAB49FFB0D024FD5BF9987AE6992E" attach="left" box="[678,700,1594,1607]" fontSize="5" pageId="7" pageNumber="78">1-4</superScript>
|
||||
</emphasis>
|
||||
,
|
||||
<emphasis id="B95CDA13FFB0D024FD3BF99F7ABA992E" bold="true" box="[710,736,1594,1614]" pageId="7" pageNumber="78">
|
||||
l
|
||||
<superScript id="7C5DAB49FFB0D024FD37F9987ABA992E" attach="left" box="[714,736,1594,1607]" fontSize="5" pageId="7" pageNumber="78">1-4</superScript>
|
||||
</emphasis>
|
||||
,
|
||||
<emphasis id="B95CDA13FFB0D024FD17F99F7B56992E" bold="true" box="[746,780,1594,1614]" pageId="7" pageNumber="78">
|
||||
rl
|
||||
<superScript id="7C5DAB49FFB0D024FD0BF9987B56992E" attach="left" box="[758,780,1594,1607]" fontSize="5" pageId="7" pageNumber="78">1-4</superScript>
|
||||
</emphasis>
|
||||
, carinolateral, lateral and rostrolateral tiers of imbricating plates respectively;
|
||||
<emphasis id="B95CDA13FFB0D024FEA9F9F1793E990D" bold="true" box="[340,356,1619,1636]" pageId="7" pageNumber="78">cl</emphasis>
|
||||
,
|
||||
<emphasis id="B95CDA13FFB0D024FE92F9F17921990D" bold="true" box="[367,379,1619,1636]" pageId="7" pageNumber="78">rl</emphasis>
|
||||
, the pair of imbricating plates in proverrucids;
|
||||
<emphasis id="B95CDA13FFB0D024FCF6F9F17B78990D" bold="true" box="[779,802,1619,1636]" pageId="7" pageNumber="78">FS</emphasis>
|
||||
,
|
||||
<emphasis id="B95CDA13FFB0D024FCD3F9F17B1E990D" bold="true" box="[814,836,1619,1636]" pageId="7" pageNumber="78">FT</emphasis>
|
||||
, fixed scutum and tergum;
|
||||
<emphasis id="B95CDA13FFB0D024FBCCF9F17C1C990D" bold="true" box="[1073,1094,1619,1636]" pageId="7" pageNumber="78">hl,</emphasis>
|
||||
hinge line;
|
||||
<emphasis id="B95CDA13FFB0D024FB54F9F17CEE990D" bold="true" box="[1193,1204,1619,1636]" pageId="7" pageNumber="78">L</emphasis>
|
||||
, median latus (completely lost in proverrucids and verrucids);
|
||||
<emphasis id="B95CDA13FFB0D024FD7DF9CA7ACB9910" bold="true" box="[640,657,1640,1657]" pageId="7" pageNumber="78">m</emphasis>
|
||||
, myophore;
|
||||
<emphasis id="B95CDA13FFB0D024FD02F9CA7B469910" bold="true" box="[767,796,1640,1657]" pageId="7" pageNumber="78">MS</emphasis>
|
||||
,
|
||||
<emphasis id="B95CDA13FFB0D024FCD4F9CA7B1F9910" bold="true" box="[809,837,1640,1657]" pageId="7" pageNumber="78">MT</emphasis>
|
||||
, movable scutum and tergum;
|
||||
<emphasis id="B95CDA13FFB0D024FBA8F9CA7C219910" bold="true" box="[1109,1147,1640,1657]" pageId="7" pageNumber="78">sam</emphasis>
|
||||
, scutal adductor muscle;
|
||||
<emphasis id="B95CDA13FFB0D024FEF5F9DF794E99E7" bold="true" box="[264,276,1661,1678]" pageId="7" pageNumber="78">S</emphasis>
|
||||
,
|
||||
<emphasis id="B95CDA13FFB0D024FEE3F9DF797399E7" bold="true" box="[286,297,1661,1678]" pageId="7" pageNumber="78">T</emphasis>
|
||||
, normal terga and scuta in brachylepadomorphs; dashed horizontal lines: membranous basis (transverse sections F-H after
|
||||
<bibRefCitation id="EFB97BF0FFB0D024FF13F931790D99CD" author="Young P. S." box="[238,343,1683,1700]" journalOrPublisher="Zoosystema" pageId="7" pageNumber="78" pagination="31 - 92" part="20" refId="ref7981" refString="Young P. S. 1998. - Cirripedia (Crustacea) from the " Campagne Biacores " in the Azores region, including a generic revision of Verrucidae. Zoosystema 20 (1): 31 - 92." title="Cirripedia (Crustacea) from the " Campagne Biacores " in the Azores region, including a generic revision of Verrucidae" type="journal article" year="1998">Young 1998</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B970601FFBFD02BFF8FFF7379699DDE" blockId="8.[114,647,209,695]" pageId="8" pageNumber="79">
|
||||
The caudal appendages, while multiarticulate in
|
||||
<taxonomicName id="4C287D82FFBFD02BFF8FFF4D794C9E60" box="[114,278,239,265]" class="Maxillopoda" family="Chionelasmatidae" genus="Eochionelasmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBFD02BFF8FFF4D794C9E60" box="[114,278,239,265]" italics="true" pageId="8" pageNumber="79">Eochionelasmus</emphasis>
|
||||
</taxonomicName>
|
||||
and minute and uniarticulate in
|
||||
<taxonomicName id="4C287D82FFBFD02BFF8FFEAC79259E41" authority="Newman, 1979" authorityName="Newman" authorityYear="1979" box="[114,383,269,296]" class="Maxillopoda" family="Eolepadidae" genus="Neolepas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="8" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBFD02BFF8FFEAC78979E41" box="[114,205,270,296]" italics="true" pageId="8" pageNumber="79">Neolepas</emphasis>
|
||||
<bibRefCitation id="EFB97BF0FFBFD02BFF29FEAF79259E41" author="Newman W. A." box="[212,383,269,296]" journalOrPublisher="San Diego Society of Natural History Transactions" pageId="8" pageNumber="79" pagination="153 - 167" part="19" refId="ref6965" refString="Newman W. A. 1979. - A new scalpellid (Cirripedia); a Mesozoic relic living near an abyssal hydrothermal spring. San Diego Society of Natural History Transactions 19 (11): 153 - 167." title="A new scalpellid (Cirripedia); a Mesozoic relic living near an abyssal hydrothermal spring" type="journal article" year="1979">Newman, 1979</bibRefCitation>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C287D82FFBFD02BFE44FEAC7A029E41" authorityName="Newman & Yamaguchi" authorityYear="1995" box="[441,600,270,296]" class="Maxillopoda" family="Neobrachylepadidae" genus="Neobrachylepas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBFD02BFE44FEAC7A029E41" box="[441,600,270,296]" italics="true" pageId="8" pageNumber="79">Neobrachylepas</emphasis>
|
||||
</taxonomicName>
|
||||
, are absent in
|
||||
<taxonomicName id="4C287D82FFBFD02BFF19FE8F79D19E2E" authorityName="Newman" authorityYear="2000" box="[228,395,301,327]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBFD02BFF19FE8F79D19E2E" box="[228,395,301,327]" italics="true" pageId="8" pageNumber="79">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
as well as
|
||||
<taxonomicName id="4C287D82FFBFD02BFDF6FE8E7ADD9E2C" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[523,647,300,325]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBFD02BFDF6FE8E7ADD9E2C" box="[523,647,300,325]" italics="true" pageId="8" pageNumber="79">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="13131A84FFBFD02BFF87FEE978809E0C" box="[122,218,331,357]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="8" pageNumber="79">Fig. 3C</figureCitation>
|
||||
). Both adults were hermaphroditic; penis long, conspicuously annulated, and sparsely clothed with short, stiff setae (
|
||||
<figureCitation id="13131A84FFBFD02BFE04FE2B7A239ECB" box="[505,633,392,419]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="8" pageNumber="79">Fig. 3C, E</figureCitation>
|
||||
). While the generally well-developed penes of vent barnacles have a well-developed pedicel, there is no basidorsal point (
|
||||
<figureCitation id="13131A84FFBFD02BFE9FFE4779E49E96" box="[354,446,485,511]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="8" pageNumber="79">Fig. 3C</figureCitation>
|
||||
). Well-developed testes extend into the pedicels of cirrus I & II (
|
||||
<figureCitation id="13131A84FFBFD02BFF84FD8078A89D52" box="[121,242,545,572]" captionStart="FIG" captionStartId="5.[113,124,1576,1593]" captionTargetBox="[92,1284,200,1546]" captionTargetId="figure-22@5.[61,1325,181,1567]" captionTargetPageId="5" captionText="FIG. 3. — Cirri (right side) and the trophi (appendages from the right side viewed from without, those of the left sides form within) of Imbricaverruca yamaguchii n. gen. and n. sp. (paratype, MNHN Ci2711); A, cirrus I, anterior ramus missing; B, cirrus II, setae omitted from the posterior ramus of the intact cirrus (left) are shown on the excised ramus (right); C, cirrus VI (setae of distal half omitted) and penis (annulations omitted); D, setation of an intermediate segment of outer ramus of cirrus VI (17th article from distal end); E, end of penis enlarged; F, labrum and mandibular palps; G, H, right and left mandible; I, spinous margin of H enlarged; J, K, right and left first maxillae; L, right second maxilla. Scale bars: A-C, 0.5 mm; D, E, 0.15 mm; F-L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391401" httpUri="https://zenodo.org/record/5391401/files/figure.png" pageId="8" pageNumber="79">Fig. 3A, B</figureCitation>
|
||||
), and to a lesser degree into at least those of cirrus III & IV. Ovigerous frena were not seen, but no ovigerous lamellae were being incubated and in their absence frena may be easily overlooked.
|
||||
</paragraph>
|
||||
<paragraph id="8B970601FFBFD02BFD58FF727B609F83" blockId="8.[677,1210,208,694]" box="[677,826,208,234]" pageId="8" pageNumber="79">
|
||||
<emphasis id="B95CDA13FFBFD02BFD58FF727B609F83" box="[677,826,208,234]" italics="true" pageId="8" pageNumber="79">The x-juvenile</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8B970601FFBFD02BFD58FF4D7CE09DDF" blockId="8.[677,1210,208,694]" pageId="8" pageNumber="79">
|
||||
A number of attached cyprids found on sulfide deposits associated with hydrothermal activity were included with the collection from the Lau Basin sent by CENTOB. Some were sent to Peter G. Jensen for the study of lattice organs (cf.
|
||||
<bibRefCitation id="EFB97BF0FFBFD02BFD58FE2A7B3D9ECB" author="Jensen P. G. & HOeg J. T. & Bower S. & Rybakov A." box="[677,871,392,418]" journalOrPublisher="Canadian Journal of Zoology" pageId="8" pageNumber="79" pagination="1018 - 1026" part="72" refId="ref6802" refString="Jensen P. G., HOeg J. T., Bower S. & Rybakov A. 1994. - Scanning electron microscopy of lattice organs in cyprids of the Rhizocephala Akentrogonida (Crustacea, Cirripedia). Canadian Journal of Zoology 72 (6): 1018 - 1026." title="Scanning electron microscopy of lattice organs in cyprids of the Rhizocephala Akentrogonida (Crustacea, Cirripedia)" type="journal article" year="1994">
|
||||
Jensen
|
||||
<emphasis id="B95CDA13FFBFD02BFD0FFE2A7B7D9ECB" box="[754,807,392,418]" italics="true" pageId="8" pageNumber="79">et al.</emphasis>
|
||||
1994
|
||||
</bibRefCitation>
|
||||
) who removed the cyprid shell covering the first juvenile stage inside, when preparing them for SEM. Some of these were neolepadines but two were the first juvenile stage of an unusual thoracic cirriped. It was studied and dubbed “Jensen’s x-juvenile” by
|
||||
<bibRefCitation id="EFB97BF0FFBFD02BFBA4FD807B039D33" author="HOeg J. T. & Newman W. A." bookContentInfo="Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris" pageId="8" pageNumber="79" refId="ref6736" refString="HOeg J. T. & Newman W. A. 1997. - Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga. Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris." title="Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga." type="proceedings" year="1997">HØeg & Newman (1997)</bibRefCitation>
|
||||
(
|
||||
<figureCitation id="13131A84FFBFD02BFC95FDE27BF39D33" box="[872,937,576,602]" captionStart="FIG" captionStartId="8.[114,125,1597,1614]" captionTargetBox="[139,1204,771,1569]" captionTargetId="figure-24@8.[127,1202,745,1582]" captionTargetPageId="8" captionText="FIG. 5. — Jensen’s x-juvenile from the Lau Basin, Tonga (Høeg & Newman 1997); A, viewed from the left dorso-lateral; B, left lateral; C, ventral sides, respectively (the shard of chitin extending from the aperture in B and C apparently left over from the last molt). It is a first juvenile of a thoracican cirriped and likely that of Imbricaverruca yamaguchii n. gen. and n. sp.; see Fig. 6 and text for explanation. Abbreviations: A1, first antenna; C, carina; “C”, incipient cirri; LS, left scutum; LT, left tergum; RS, right scutum; P, peduncle. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5391403" httpUri="https://zenodo.org/record/5391403/files/figure.png" pageId="8" pageNumber="79">Fig. 5</figureCitation>
|
||||
). Its capitulum, supported by a peduncle of comparable length, is armored by the five primordial valves commonly found in juvenile thoracicans, except iblomorphs.
|
||||
</paragraph>
|
||||
<caption id="DF575689FFBFD02BFF8FF99F794399CD" ID-DOI="http://doi.org/10.5281/zenodo.5391403" ID-Zenodo-Dep="5391403" httpUri="https://zenodo.org/record/5391403/files/figure.png" pageId="8" pageNumber="79" startId="8.[114,125,1597,1614]" targetBox="[139,1204,771,1569]" targetPageId="8">
|
||||
<paragraph id="8B970601FFBFD02BFF8FF99F794399CD" blockId="8.[114,1211,1597,1700]" pageId="8" pageNumber="79">
|
||||
FIG. 5. — Jensen’s x-juvenile from the Lau Basin, Tonga (
|
||||
<bibRefCitation id="EFB97BF0FFBFD02BFDB1F99F7B509927" author="HOeg J. T. & Newman W. A." bookContentInfo="Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris" box="[588,778,1597,1614]" pageId="8" pageNumber="79" refId="ref6736" refString="HOeg J. T. & Newman W. A. 1997. - Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga. Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris." title="Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga." type="proceedings" year="1997">Høeg & Newman 1997</bibRefCitation>
|
||||
);
|
||||
<emphasis id="B95CDA13FFBFD02BFCEAF99F7B7E9927" bold="true" box="[791,804,1597,1614]" pageId="8" pageNumber="79">A</emphasis>
|
||||
, viewed from the left dorso-lateral;
|
||||
<emphasis id="B95CDA13FFBFD02BFBB1F99F7C039927" bold="true" box="[1100,1113,1597,1614]" pageId="8" pageNumber="79">B</emphasis>
|
||||
, left lateral;
|
||||
<emphasis id="B95CDA13FFBFD02BFF8FF9F178DA990D" bold="true" box="[114,128,1619,1636]" pageId="8" pageNumber="79">C</emphasis>
|
||||
, ventral sides, respectively (the shard of chitin extending from the aperture in B and C apparently left over from the last molt). It is a first juvenile of a thoracican cirriped and likely that of
|
||||
<taxonomicName id="4C287D82FFBFD02BFDC7F9CA7B429910" authority="Newman, 2000" authorityName="Newman" authorityYear="2000" box="[570,792,1640,1657]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="yamaguchii" status="n. gen. and n. sp.">
|
||||
<emphasis id="B95CDA13FFBFD02BFDC7F9CA7B429910" box="[570,792,1640,1657]" italics="true" pageId="8" pageNumber="79">Imbricaverruca yamaguchii</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A26F6768FFBFD02BFCE0F9CA7BEB9910" box="[797,945,1640,1657]" pageId="8" pageNumber="79" rank="species">n. gen. and n. sp.</taxonomicNameLabel>
|
||||
; see Fig. 6 and text for explanation. Abbreviations:
|
||||
<emphasis id="B95CDA13FFBFD02BFEE4F9DF797499EE" bold="true" box="[281,302,1658,1678]" pageId="8" pageNumber="79">
|
||||
A
|
||||
<superScript id="7C5DAB49FFBFD02BFEDBF9D8797499EE" attach="left" box="[294,302,1658,1671]" fontSize="5" pageId="8" pageNumber="79">1</superScript>
|
||||
</emphasis>
|
||||
, first antenna;
|
||||
<emphasis id="B95CDA13FFBFD02BFE50F9DF79E199E7" bold="true" box="[429,443,1661,1678]" pageId="8" pageNumber="79">C</emphasis>
|
||||
, carina;
|
||||
<emphasis id="B95CDA13FFBFD02BFDF9F9DF7A7999E7" bold="true" box="[516,547,1661,1678]" pageId="8" pageNumber="79">“C”</emphasis>
|
||||
, incipient cirri;
|
||||
<emphasis id="B95CDA13FFBFD02BFD59F9DF7AE199E7" bold="true" box="[676,699,1661,1678]" pageId="8" pageNumber="79">LS</emphasis>
|
||||
, left scutum;
|
||||
<emphasis id="B95CDA13FFBFD02BFCD2F9DF7B1F99E7" bold="true" box="[815,837,1661,1678]" pageId="8" pageNumber="79">LT</emphasis>
|
||||
, left tergum;
|
||||
<emphasis id="B95CDA13FFBFD02BFC4AF9DF7B8B99E7" bold="true" box="[951,977,1661,1678]" pageId="8" pageNumber="79">RS</emphasis>
|
||||
, right scutum;
|
||||
<emphasis id="B95CDA13FFBFD02BFBADF9DF7C0699E7" bold="true" box="[1104,1116,1661,1678]" pageId="8" pageNumber="79">P</emphasis>
|
||||
, peduncle. Scale bars: 100 µm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF575689FFBED02AFF8FFCEC78939CDD" ID-DOI="http://doi.org/10.5281/zenodo.5391405" ID-Zenodo-Dep="5391405" httpUri="https://zenodo.org/record/5391405/files/figure.png" pageId="9" pageNumber="80" startId="9.[114,125,846,863]" targetBox="[297,1063,203,833]" targetPageId="9">
|
||||
<paragraph id="8B970601FFBED02AFF8FFCEC78939CDD" blockId="9.[114,1212,846,948]" pageId="9" pageNumber="80">
|
||||
FIG. 6. — Comparison between the primordial plates of the earliest juvenile stages of the principal suborders of thoracic
|
||||
<taxonomicName id="4C287D82FFBED02AFB99FCEC7CED9C36" authorityName="Burmeister" authorityYear="1834" box="[1124,1207,846,863]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="73" phylum="Arthropoda" rank="subClass" subClass="Cirripedia">Cirripedia</taxonomicName>
|
||||
;
|
||||
<emphasis id="B95CDA13FFBED02AFF8FFCC178259C1D" bold="true" box="[114,127,867,884]" pageId="9" pageNumber="80">A</emphasis>
|
||||
, Lepadomorpha,
|
||||
<taxonomicName id="4C287D82FFBED02AFEE8FCC1791D9C1D" box="[277,327,867,884]" class="Maxillopoda" family="Lepadidae" genus="Lepas" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="80" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBED02AFEE8FCC1791D9C1D" box="[277,327,867,884]" italics="true" pageId="9" pageNumber="80">Lepas</emphasis>
|
||||
</taxonomicName>
|
||||
(from
|
||||
<bibRefCitation id="EFB97BF0FFBED02AFE82FCC17A719C1D" author="Newman W. A." box="[383,555,867,884]" journalOrPublisher="San Diego Society of Natural History Transactions" pageId="9" pageNumber="80" pagination="153 - 167" part="19" refId="ref6965" refString="Newman W. A. 1979. - A new scalpellid (Cirripedia); a Mesozoic relic living near an abyssal hydrothermal spring. San Diego Society of Natural History Transactions 19 (11): 153 - 167." title="A new scalpellid (Cirripedia); a Mesozoic relic living near an abyssal hydrothermal spring" type="journal article" year="1979">
|
||||
Newman
|
||||
<emphasis id="B95CDA13FFBED02AFE2DFCC179A09C1D" box="[464,506,867,884]" italics="true" pageId="9" pageNumber="80">et al.</emphasis>
|
||||
1979
|
||||
</bibRefCitation>
|
||||
: fig. 87);
|
||||
<emphasis id="B95CDA13FFBED02AFD84FCC17ADC9C1D" bold="true" box="[633,646,867,884]" pageId="9" pageNumber="80">B</emphasis>
|
||||
,
|
||||
<taxonomicName id="4C287D82FFBED02AFD6FFCC17B789C1D" box="[658,802,867,884]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scalpellomorpha" pageId="9" pageNumber="80" phylum="Arthropoda" rank="order">Scalpellomorpha</taxonomicName>
|
||||
, composite of
|
||||
<taxonomicName id="4C287D82FFBED02AFC58FCC17BA99C1D" authorityName="Newman" authorityYear="1979" box="[933,1011,867,884]" class="Maxillopoda" family="Eolepadidae" genus="Neolepas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="80" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBED02AFC58FCC17BA99C1D" box="[933,1011,867,884]" italics="true" pageId="9" pageNumber="80">Neolepas</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C287D82FFBED02AFBE2FCC17C349C1D" authorityName="Leach" authorityYear="1817" box="[1055,1134,867,884]" class="Maxillopoda" family="Pollicipedidae" genus="Pollicipes" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="80" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBED02AFBE2FCC17C349C1D" box="[1055,1134,867,884]" italics="true" pageId="9" pageNumber="80">Pollicipes</emphasis>
|
||||
</taxonomicName>
|
||||
(respectively from Newman, in prep. and
|
||||
<bibRefCitation id="EFB97BF0FFBED02AFE6CFCDB79AF9CE3" author="Broch H." box="[401,501,889,906]" journalOrPublisher="Videnskabelige meddelelser fra Dansk Naturhistorisk Forening i Kobenhavn" pageId="9" pageNumber="80" pagination="215 - 358" part="73" refId="ref6293" refString="Broch H. 1922. - Studies on Pacific cirripeds, in Papers from Dr Th. Mortensen's Pacific Expedition 1914 - 1916. X. Videnskabelige meddelelser fra Dansk Naturhistorisk Forening i Kobenhavn 73: 215 - 358." title="Studies on Pacific cirripeds, in Papers from Dr Th. Mortensen's Pacific Expedition 1914 - 1916. X" type="journal article" year="1922">Broch 1922</bibRefCitation>
|
||||
);
|
||||
<emphasis id="B95CDA13FFBED02AFDFEFCDB7A4B9CE3" bold="true" box="[515,529,889,906]" pageId="9" pageNumber="80">C</emphasis>
|
||||
,
|
||||
<taxonomicName id="4C287D82FFBED02AFDE1FCDB7A269CE3" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[540,636,889,906]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="9" pageNumber="80" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBED02AFDE1FCDB7A269CE3" box="[540,636,889,906]" italics="true" pageId="9" pageNumber="80">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
(reconstructed from
|
||||
<bibRefCitation id="EFB97BF0FFBED02AFCD0FCDB7BF09CE3" author="Newman W. A." box="[813,938,889,906]" journalOrPublisher="Bulletin of Marine Science" pageId="9" pageNumber="80" pagination="467 - 477" part="45" refId="ref7144" refString="Newman W. A. 1989. - Juvenile ontogeny and metamorphosis in the most primitive living sessile barnacle, Neoverruca, from an abyssal hydrothermal spring. Bulletin of Marine Science 45 (2): 467 - 477." title="Juvenile ontogeny and metamorphosis in the most primitive living sessile barnacle, Neoverruca, from an abyssal hydrothermal spring" type="journal article" year="1989">Newman 1989</bibRefCitation>
|
||||
);
|
||||
<emphasis id="B95CDA13FFBED02AFC4AFCDB7B9F9CE3" bold="true" box="[951,965,889,906]" pageId="9" pageNumber="80">D</emphasis>
|
||||
, x-juvenile (
|
||||
<bibRefCitation id="EFB97BF0FFBED02AFBD6FCDB78C69CF6" author="HOeg J. T. & Newman W. A." bookContentInfo="Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris" pageId="9" pageNumber="80" refId="ref6736" refString="HOeg J. T. & Newman W. A. 1997. - Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga. Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris." title="Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga." type="proceedings" year="1997">Høeg & Newman 1997</bibRefCitation>
|
||||
& herein);
|
||||
<emphasis id="B95CDA13FFBED02AFF07FC2C795C9CF6" bold="true" box="[250,262,910,927]" pageId="9" pageNumber="80">E</emphasis>
|
||||
,
|
||||
<emphasis id="B95CDA13FFBED02AFEEFFC2C79479CF6" bold="true" box="[274,285,910,927]" pageId="9" pageNumber="80">F</emphasis>
|
||||
,
|
||||
<taxonomicName id="4C287D82FFBED02AFED4FC2C79339CF6" authorityName="Schumacher" authorityYear="1817" box="[297,361,910,927]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="9" pageNumber="80" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBED02AFED4FC2C79339CF6" box="[297,361,910,927]" italics="true" pageId="9" pageNumber="80">Verruca</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C287D82FFBED02AFE6AFC2C7A589CF6" authorityName="Pilsbry" authorityYear="1916" box="[407,514,910,927]" class="Maxillopoda" family="Archaeobalanidae" genus="Semibalanus" kingdom="Animalia" order="Sessilia" pageId="9" pageNumber="80" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBED02AFE6AFC2C7A589CF6" box="[407,514,910,927]" italics="true" pageId="9" pageNumber="80">Semibalanus</emphasis>
|
||||
</taxonomicName>
|
||||
(respectively from
|
||||
<bibRefCitation id="EFB97BF0FFBED02AFD5BFC2C7B669CF6" author="Runnstrom S." box="[678,828,910,927]" journalOrPublisher="Bergens Museums Arbok" pageId="9" pageNumber="80" pagination="1 - 46" part="5" refId="ref7493" refString="Runnstrom S. 1925. - Zur Biologie und Entwicklung von Balanus balanoides (Linn.). Bergens Museums Arbok 5: 1 - 46." title="Zur Biologie und Entwicklung von Balanus balanoides (Linn.)" type="journal article" year="1925">Runnström, 1925</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFB97BF0FFBED02AFCB8FC2C7B289CF6" author="Runnstrom S." box="[837,882,910,927]" journalOrPublisher="Bergens Museums Arbok" pageId="9" pageNumber="80" pagination="1 - 10" part="7" refId="ref7520" refString="Runnstrom S. 1927. - Uber die Plattenentwicklung von Verruca stroemia. Bergens Museums Arbok 7: 1 - 10." title="Uber die Plattenentwicklung von Verruca stroemia" type="journal article" year="1927">1927</bibRefCitation>
|
||||
). Abbreviations:
|
||||
<emphasis id="B95CDA13FFBED02AFC02FC2C7C519CF6" bold="true" box="[1023,1035,910,927]" pageId="9" pageNumber="80">S</emphasis>
|
||||
, scutum;
|
||||
<emphasis id="B95CDA13FFBED02AFB9FFC2C7C379CF6" bold="true" box="[1122,1133,910,927]" pageId="9" pageNumber="80">T</emphasis>
|
||||
, tergum;
|
||||
<emphasis id="B95CDA13FFBED02AFF8FFC0178DA9CDD" bold="true" box="[114,128,931,948]" pageId="9" pageNumber="80">C</emphasis>
|
||||
, carina).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF575689FFBED02AFF8CFBDD7AFE9BCC" pageId="9" pageNumber="80" startId="9.[113,124,1151,1168]" targetBox="[113,1171,1240,1689]" targetIsTable="true" targetPageId="9">
|
||||
<paragraph id="8B970601FFBED02AFF8CFBDD7AFE9BCC" blockId="9.[113,1210,1151,1189]" pageId="9" pageNumber="80">
|
||||
TABLE 2. — Comparison between the primordial plates of the earliest juvenile stages of the principal suborders of thoracic
|
||||
<taxonomicName id="4C287D82FFBED02AFB94FBDD7CE09BF9" authorityName="Burmeister" authorityYear="1834" box="[1129,1210,1151,1168]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="73" phylum="Arthropoda" rank="subClass" subClass="Cirripedia">Cirripedia</taxonomicName>
|
||||
(See Fig. 6 for corresponding figures and text for a full explanation).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B970601FFBED02AFF8CFB7A7C7A99E8" pageId="9" pageNumber="80">
|
||||
<table id="F928F4A1FFBE2FDCFF8CFB7A7CC999F0" box="[113,1171,1240,1689]" gridcols="4" gridrows="8" pageId="9" pageNumber="80">
|
||||
<tr id="35180443FFBE2FDCFF8CFB7A7CC99A6D" box="[113,1171,1240,1284]" gridrow="0" pageId="9" pageNumber="80">
|
||||
<th id="76C96D3FFFBE2FDCFF8CFB7A793D9A6D" box="[113,359,1240,1284]" gridcol="0" gridrow="0" pageId="9" pageNumber="80">
|
||||
<emphasis id="B95CDA13FFBED02AFF8CFB7A79479B85" bold="true" box="[113,285,1240,1260]" pageId="9" pageNumber="80">Taxon/Character</emphasis>
|
||||
</th>
|
||||
<th id="76C96D3FFFBE2FDCFE87FB7A7ADD9A6D" box="[378,647,1240,1284]" gridcol="1" gridrow="0" pageId="9" pageNumber="80">
|
||||
<emphasis id="B95CDA13FFBED02AFE87FB7A7A659B85" bold="true" box="[378,575,1240,1260]" pageId="9" pageNumber="80">Carinal proportions</emphasis>
|
||||
</th>
|
||||
<th id="76C96D3FFFBE2FDCFD6FFB7A7BD49A6D" box="[658,910,1240,1284]" gridcol="2" gridrow="0" pageId="9" pageNumber="80">
|
||||
<emphasis id="B95CDA13FFBED02AFD6FFB7A7B169B85" bold="true" box="[658,844,1240,1260]" pageId="9" pageNumber="80">Carinal of position</emphasis>
|
||||
</th>
|
||||
<th id="76C96D3FFFBE2FDCFC54FB7A7CC99A6D" box="[937,1171,1240,1284]" gridcol="3" gridrow="0" pageId="9" pageNumber="80">
|
||||
<emphasis id="B95CDA13FFBED02AFC54FB7A7C7F9A6D" bold="true" pageId="9" pageNumber="80">Terga and scuta of each side</emphasis>
|
||||
</th>
|
||||
</tr>
|
||||
<tr id="35180443FFBE2FDCFF8CFABC7CC99A23" box="[113,1171,1310,1354]" gridrow="1" pageId="9" pageNumber="80">
|
||||
<th id="76C96D3FFFBE2FDCFF8CFABC793D9A23" box="[113,359,1310,1354]" gridcol="0" gridrow="1" pageId="9" pageNumber="80">Lepadomorpha (Fig. 6A)</th>
|
||||
<td id="76C96D3FFFBE2FDCFE87FABC7ADD9A23" box="[378,647,1310,1354]" gridcol="1" gridrow="1" pageId="9" pageNumber="80">Higher than wide and longer than terga</td>
|
||||
<td id="76C96D3FFFBE2FDCFD6FFABC7BD49A23" box="[658,910,1310,1354]" gridcol="2" gridrow="1" pageId="9" pageNumber="80">Extending up between terga</td>
|
||||
<td id="76C96D3FFFBE2FDCFC54FABC7CC99A23" box="[937,1171,1310,1354]" gridcol="3" gridrow="1" pageId="9" pageNumber="80">Symmetrical</td>
|
||||
</tr>
|
||||
<tr id="35180443FFBE2FDCFF8CFAF07CC99A17" box="[113,1171,1362,1406]" gridrow="2" pageId="9" pageNumber="80">
|
||||
<th id="76C96D3FFFBE2FDCFF8CFAF0793D9A17" box="[113,359,1362,1406]" gridcol="0" gridrow="2" pageId="9" pageNumber="80">
|
||||
<taxonomicName id="4C287D82FFBED02AFF8CFAF079489A0F" box="[113,274,1362,1382]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scalpellomorpha" pageId="9" pageNumber="80" phylum="Arthropoda" rank="order">Scalpellomorpha</taxonomicName>
|
||||
(Fig. 6B)
|
||||
</th>
|
||||
<td id="76C96D3FFFBE2FDCFE87FAF07ADD9A17" box="[378,647,1362,1406]" gridcol="1" gridrow="2" pageId="9" pageNumber="80">Higher than wide and nearly as long as terga</td>
|
||||
<td id="76C96D3FFFBE2FDCFD6FFAF07BD49A17" box="[658,910,1362,1406]" gridcol="2" gridrow="2" pageId="9" pageNumber="80">Extending up between terga</td>
|
||||
<td id="76C96D3FFFBE2FDCFC54FAF07CC99A17" box="[937,1171,1362,1406]" gridcol="3" gridrow="2" pageId="9" pageNumber="80">Symmetrical</td>
|
||||
</tr>
|
||||
<tr id="35180443FFBE2FDCFF8CFA247CC99AF3" box="[113,1171,1414,1434]" gridrow="3" pageId="9" pageNumber="80">
|
||||
<th id="76C96D3FFFBE2FDCFF8CFA247CC99AF3" box="[113,1171,1414,1434]" colspan="4" colspanRight="3" gridcol="0" gridrow="3" pageId="9" pageNumber="80">
|
||||
<taxonomicName id="4C287D82FFBED02AFF8CFA24795F9AF3" authorityName="Pilsbry" authorityYear="1916" box="[113,261,1414,1434]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="9" pageNumber="73" phylum="Arthropoda" rank="subOrder" subOrder="Verrucomorpha">Verrucomorpha</taxonomicName>
|
||||
</th>
|
||||
</tr>
|
||||
<tr id="35180443FFBE2FDCFF8CFA007CC99AA7" box="[113,1171,1442,1486]" gridrow="4" pageId="9" pageNumber="80">
|
||||
<th id="76C96D3FFFBE2FDCFF8CFA00793D9AA7" box="[113,359,1442,1486]" gridcol="0" gridrow="4" pageId="9" pageNumber="80">
|
||||
<taxonomicName id="4C287D82FFBED02AFF8CFA0078A69ADF" box="[113,252,1442,1462]" class="Maxillopoda" family="Neoverrucidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="9" pageNumber="80" phylum="Arthropoda" rank="family">Neoverrucidae</taxonomicName>
|
||||
(Fig. 6C)
|
||||
</th>
|
||||
<td id="76C96D3FFFBE2FDCFE87FA007ADD9AA7" box="[378,647,1442,1486]" gridcol="1" gridrow="4" pageId="9" pageNumber="80">Higher than wide and shorter than terga</td>
|
||||
<td id="76C96D3FFFBE2FDCFD6FFA007BD49AA7" box="[658,910,1442,1486]" gridcol="2" gridrow="4" pageId="9" pageNumber="80">Extending up between terga</td>
|
||||
<td id="76C96D3FFFBE2FDCFC54FA007CC99AA7" box="[937,1171,1442,1486]" gridcol="3" gridrow="4" pageId="9" pageNumber="80">Initially symmetrical</td>
|
||||
</tr>
|
||||
<tr id="35180443FFBE2FDCFF8CFA777CC99970" box="[113,1171,1493,1561]" gridrow="5" pageId="9" pageNumber="80">
|
||||
<th id="76C96D3FFFBE2FDCFF8CFA77793D9970" box="[113,359,1493,1561]" gridcol="0" gridrow="5" pageId="9" pageNumber="80">X-juvenile (Fig. 6D)</th>
|
||||
<td id="76C96D3FFFBE2FDCFE87FA777ADD9970" box="[378,647,1493,1561]" gridcol="1" gridrow="5" pageId="9" pageNumber="80">Wider than high, shorter than terga and perhaps displaced to one side</td>
|
||||
<td id="76C96D3FFFBE2FDCFD6FFA777BD49970" box="[658,910,1493,1561]" gridcol="2" gridrow="5" pageId="9" pageNumber="80">Not extending up between terga</td>
|
||||
<td id="76C96D3FFFBE2FDCFC54FA777CC99970" box="[937,1171,1493,1561]" gridcol="3" gridrow="5" pageId="9" pageNumber="80">Scuta if not terga initially symmetrical</td>
|
||||
</tr>
|
||||
<tr id="35180443FFBE2FDCFF8CF9837CC9990C" box="[113,1171,1569,1637]" gridrow="6" pageId="9" pageNumber="80">
|
||||
<th id="76C96D3FFFBE2FDCFF8CF983793D990C" box="[113,359,1569,1637]" gridcol="0" gridrow="6" pageId="9" pageNumber="80">
|
||||
<taxonomicName id="4C287D82FFBED02AFF8CF983788D995C" authorityName="Darwin" authorityYear="1854" box="[113,215,1569,1589]" class="Maxillopoda" family="Verrucidae" kingdom="Animalia" order="Sessilia" pageId="9" pageNumber="80" phylum="Arthropoda" rank="family">Verrucidae</taxonomicName>
|
||||
(Fig. 6E)
|
||||
</th>
|
||||
<td id="76C96D3FFFBE2FDCFE87F9837ADD990C" box="[378,647,1569,1637]" gridcol="1" gridrow="6" pageId="9" pageNumber="80">About as wide as high, shorter than terga and displaced to one side</td>
|
||||
<td id="76C96D3FFFBE2FDCFD6FF9837BD4990C" box="[658,910,1569,1637]" gridcol="2" gridrow="6" pageId="9" pageNumber="80">Not extending up between terga</td>
|
||||
<td id="76C96D3FFFBE2FDCFC54F9837CC9990C" box="[937,1171,1569,1637]" gridcol="3" gridrow="6" pageId="9" pageNumber="80">Distinctly asymmetrical</td>
|
||||
</tr>
|
||||
<tr id="35180443FFBE2FDCFF8CF9CF7CC999F0" box="[113,1171,1645,1689]" gridrow="7" pageId="9" pageNumber="80">
|
||||
<th id="76C96D3FFFBE2FDCFF8CF9CF793D99F0" box="[113,359,1645,1689]" gridcol="0" gridrow="7" pageId="9" pageNumber="80">
|
||||
<taxonomicName id="4C287D82FFBED02AFF8CF9CF78A799E8" box="[113,253,1645,1665]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Balanomorpha" pageId="9" pageNumber="80" phylum="Arthropoda" rank="order">Balanomorpha</taxonomicName>
|
||||
(Fig. 6F)
|
||||
</th>
|
||||
<td id="76C96D3FFFBE2FDCFE87F9CF7ADD99F0" box="[378,647,1645,1689]" gridcol="1" gridrow="7" pageId="9" pageNumber="80">Higher than wide</td>
|
||||
<td id="76C96D3FFFBE2FDCFD6FF9CF7BD499F0" box="[658,910,1645,1689]" gridcol="2" gridrow="7" pageId="9" pageNumber="80">Not extending up between terga</td>
|
||||
<td id="76C96D3FFFBE2FDCFC54F9CF7CC999F0" box="[937,1171,1645,1689]" gridcol="3" gridrow="7" pageId="9" pageNumber="80">Symmetrical</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<paragraph id="8B970601FFBDD029FF8FFF7279189E2F" blockId="10.[114,647,208,1154]" pageId="10" pageNumber="81">
|
||||
The position and shape of the carinal primordium and a slight asymmetry between the scuta suggested the juvenile was a verrucomorph (
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FDB4FEAC796F9E2F" author="HOeg J. T. & Newman W. A." bookContentInfo="Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris" pageId="10" pageNumber="81" refId="ref6736" refString="HOeg J. T. & Newman W. A. 1997. - Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga. Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris." title="Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga." type="proceedings" year="1997">HØeg & Newman 1997</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B970601FFBDD029FF8FFEE979B09BEB" blockId="10.[114,647,208,1154]" pageId="10" pageNumber="81">
|
||||
The early juveniles of
|
||||
<taxonomicName id="4C287D82FFBDD029FE7DFEE878E99EED" authority="(Newman, 1989)" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBDD029FE7DFEE87A5A9E0A" box="[384,512,330,355]" italics="true" pageId="10" pageNumber="81">Neoverruca</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FDEEFEE978F19EED" author="Newman W. A." journalOrPublisher="Bulletin of Marine Science" pageId="10" pageNumber="81" pagination="467 - 477" part="45" refId="ref7144" refString="Newman W. A. 1989. - Juvenile ontogeny and metamorphosis in the most primitive living sessile barnacle, Neoverruca, from an abyssal hydrothermal spring. Bulletin of Marine Science 45 (2): 467 - 477." title="Juvenile ontogeny and metamorphosis in the most primitive living sessile barnacle, Neoverruca, from an abyssal hydrothermal spring" type="journal article" year="1989">Newman, 1989</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
can be distinguished from the x-juvenile in being more generalized; e.g. in having a carinal primordium that is higher than wide extending up between the terga (
|
||||
<figureCitation id="13131A84FFBDD029FE8FFE6479ED9E89" box="[370,439,454,480]" captionStart="FIG" captionStartId="9.[114,125,846,863]" captionTargetBox="[297,1063,203,833]" captionTargetId="figure-268@9.[363,983,241,828]" captionTargetPageId="9" captionText="FIG. 6. — Comparison between the primordial plates of the earliest juvenile stages of the principal suborders of thoracic Cirripedia; A, Lepadomorpha, Lepas (from Newman et al. 1979: fig. 87); B, Scalpellomorpha, composite of Neolepas and Pollicipes (respectively from Newman, in prep. and Broch 1922); C, Neoverruca (reconstructed from Newman 1989); D, x-juvenile (Høeg & Newman 1997 & herein); E, F, Verruca and Semibalanus (respectively from Runnström, 1925, 1927). Abbreviations: S, scutum; T, tergum; C, carina)." figureDoi="http://doi.org/10.5281/zenodo.5391405" httpUri="https://zenodo.org/record/5391405/files/figure.png" pageId="10" pageNumber="81">Fig. 6</figureCitation>
|
||||
; cf.
|
||||
<tableCitation id="C6AA33BAFFBDD029FE16FE647A189E89" box="[491,578,454,480]" captionStart="TABLE" captionStartId="9.[113,124,1151,1168]" captionTargetPageId="9" captionText="TABLE 2. — Comparison between the primordial plates of the earliest juvenile stages of the principal suborders of thoracic Cirripedia (See Fig. 6 for corresponding figures and text for a full explanation)." pageId="10" pageNumber="81">Table 2</tableCitation>
|
||||
). The only first juvenile of a verrucid known is that of
|
||||
<taxonomicName id="4C287D82FFBDD029FF8FFDA079B99D74" authority="Muller, 1776" authorityName="Muller" authorityYear="1776" box="[114,483,514,541]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="species" species="stroemia">
|
||||
<emphasis id="B95CDA13FFBDD029FF8FFDA079629D72" box="[114,312,514,539]" italics="true" pageId="10" pageNumber="81">Verruca stroemia</emphasis>
|
||||
Müller, 1776
|
||||
</taxonomicName>
|
||||
described by
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FF8FFD8379659D55" author="Runnstrom S." box="[114,319,545,572]" journalOrPublisher="Bergens Museums Arbok" pageId="10" pageNumber="81" pagination="1 - 10" part="7" refId="ref7520" refString="Runnstrom S. 1927. - Uber die Plattenentwicklung von Verruca stroemia. Bergens Museums Arbok 7: 1 - 10." title="Uber die Plattenentwicklung von Verruca stroemia" type="journal article" year="1927">Runnström (1927)</bibRefCitation>
|
||||
and it differs from the x-juvenile in the scutal primordia being grossly unequal rather than nearly equal in size. Therefore it was concluded the x-juvenile could represent 1) a verrucid more primitive than
|
||||
<taxonomicName id="4C287D82FFBDD029FE64FD3E7A569DDC" authorityName="Muller" authorityYear="1776" box="[409,524,668,693]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="species" species="stroemia">
|
||||
<emphasis id="B95CDA13FFBDD029FE64FD3E7A569DDC" box="[409,524,668,693]" italics="true" pageId="10" pageNumber="81">V. stroemia</emphasis>
|
||||
</taxonomicName>
|
||||
such as an
|
||||
<emphasis id="B95CDA13FFBDD029FF8FFD1978B59DBC" box="[114,239,699,725]" italics="true" pageId="10" pageNumber="81">
|
||||
<taxonomicName id="4C287D82FFBDD029FF8FFD1978B19DBC" authorityName="Pilsbry" authorityYear="1916" box="[114,235,699,725]" class="Maxillopoda" family="Verrucidae" genus="Altiverruca" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="genus">Altiverruca</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
2) an advanced neoverrucid such as the one described from Lau herein, or 3) a yetto-be discovered population of the presumably extinct proverrucid, and it was thought that, because of these uncertainties, determining the affinities of Jensen’s x-juvenile would require further sampling of the Lau Basin fauna (
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FDDAFCD179409CC5" author="HOeg J. T. & Newman W. A." bookContentInfo="Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris" pageId="10" pageNumber="81" refId="ref6736" refString="HOeg J. T. & Newman W. A. 1997. - Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga. Abstract: 24, in 1 st International Symposium on Deep-Sea Hydrothermal Vent Biology, 20 - 24 October 1997, Funchal, Madeira. InterRidge, Paris." title="Jensen's xjuvenile, the first juvenile stage of an undescribed verrucomorph (Cirripedia, Thoracica) from the Lau Basin, Tonga." type="proceedings" year="1997">HØeg & Newman 1997</bibRefCitation>
|
||||
). However, the Lau neoverrucid, now known as
|
||||
<emphasis id="B95CDA13FFBDD029FEDFFC12798B9CA3" box="[290,465,944,970]" italics="true" pageId="10" pageNumber="81">
|
||||
<taxonomicName id="4C287D82FFBDD029FEDFFC1279979CA3" authorityName="Newman" authorityYear="2000" box="[290,461,944,970]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="genus">Imbricaverruca</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
turned out not only to be more highly evolved than
|
||||
<emphasis id="B95CDA13FFBDD029FDF7FC6C7ADD9C8E" box="[522,647,974,999]" italics="true" pageId="10" pageNumber="81">
|
||||
<taxonomicName id="4C287D82FFBDD029FDF7FC6C7AD89C8E" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[522,642,974,999]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="genus">Neoverruca</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
but the adult has an autapomorphy, a carina that is much wider than high, which is the most notable characteristic of the x-juvenile. Therefore it seems highly likely that Jensen’s x-juvenile is the first juvenile of
|
||||
<emphasis id="B95CDA13FFBDD029FEB9FBCA79B09BEB" box="[324,490,1128,1154]" italics="true" pageId="10" pageNumber="81">
|
||||
<taxonomicName id="4C287D82FFBDD029FEB9FBCA79BC9BEB" authorityName="Newman" authorityYear="2000" box="[324,486,1128,1154]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="genus">Imbricaverruca</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
335
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335
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<mods:title id="BD7DFF626A3F323CAD21EDCDAB54F4C7">A new genus and species of barnacle (Cirripedia, Verrucomorpha) associated with vents of the Lau Back-Arc Basin: its gross morphology, inferred first juvenile stage and affinities</mods:title>
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Origin and diversification of the
|
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<taxonomicName id="4C287D82FFBDD029FE38FB047A399BA9" authority="Pilsbry, 1916" authorityName="Pilsbry" authorityYear="1916" box="[453,611,1190,1216]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="73" phylum="Arthropoda" rank="subOrder" subOrder="Verrucomorpha">Verrucomorpha</taxonomicName>
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|
||||
Until recently, the affinities of the verrucomorphs have been puzzling. This was in good part due to the fact that while sessile, they appeared to share as many characters with the scalpellomorphs as with the balanomorphs. Darwin was the first to study their anatomy in detail and work out the homologies of their plates, and in his first volume on the fossil barnacles he notes (1851: 5),
|
||||
<taxonomicName id="4C287D82FFBDD029FDCAFA387ADD9ADA" authorityName="Schumacher" authorityYear="1817" box="[567,647,1434,1459]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBDD029FDCAFA387ADD9ADA" box="[567,647,1434,1459]" italics="true" pageId="10" pageNumber="81">Verruca</emphasis>
|
||||
</taxonomicName>
|
||||
“[…] though hitherto included amongst the Sessile Cirripedes, must, when its whole organization is taken into consideration, be ranked in a distinct family of equal value with the
|
||||
<taxonomicName id="4C287D82FFBDD029FDE3F9B47ADD9959" box="[542,647,1558,1584]" class="Maxillopoda" family="Balanidae" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="family">Balanidae</taxonomicName>
|
||||
[balanomorphs] and
|
||||
<taxonomicName id="4C287D82FFBDD029FE76F9967A509927" box="[395,522,1588,1614]" class="Maxillopoda" family="Lepadidae" kingdom="Animalia" order="Pedunculata" pageId="10" pageNumber="81" phylum="Arthropoda" rank="family">Lepadidae</taxonomicName>
|
||||
[scalpellomorphs], but perhaps more nearly related to the latter than to the Sessile Cirripedes.” In his second volume on the fossil barnacles [1855 (1854): 41], he again notes that “Upon the whole, the
|
||||
<taxonomicName id="4C287D82FFBDD029FCDEFF4D7BC39E60" authorityName="Darwin" authorityYear="1854" box="[803,921,239,265]" class="Maxillopoda" family="Verrucidae" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="family">Verrucidae</taxonomicName>
|
||||
are nearly equally related to the
|
||||
<taxonomicName id="4C287D82FFBDD029FD0EFEAC7B329E41" box="[755,872,270,296]" class="Maxillopoda" family="Lepadidae" kingdom="Animalia" order="Pedunculata" pageId="10" pageNumber="81" phylum="Arthropoda" rank="family">Lepadidae</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C287D82FFBDD029FC58FEAC7C429E41" box="[933,1048,270,296]" class="Maxillopoda" family="Balanidae" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="family">Balanidae</taxonomicName>
|
||||
; but certainly nearer to the
|
||||
<taxonomicName id="4C287D82FFBDD029FCC3FE8E7BE89E2F" box="[830,946,300,326]" class="Maxillopoda" family="Lepadidae" kingdom="Animalia" order="Pedunculata" pageId="10" pageNumber="81" phylum="Arthropoda" rank="family">Lepadidae</taxonomicName>
|
||||
, than to the sub-family
|
||||
<taxonomicName id="4C287D82FFBDD029FD58FEE97B4B9E0C" baseAuthorityName="Gistel" baseAuthorityYear="1848" box="[677,785,331,357]" class="Maxillopoda" family="Balanidae" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="subFamily" subFamily="Balaninae">Balaninae</taxonomicName>
|
||||
or typical sessile cirripedes...” But he then goes on to write, “[…] if compelled to place
|
||||
<taxonomicName id="4C287D82FFBDD029FD58FE2A7AA09EC8" authorityName="Schumacher" authorityYear="1817" box="[677,762,392,417]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBDD029FD58FE2A7AA09EC8" box="[677,762,392,417]" italics="true" pageId="10" pageNumber="81">Verruca</emphasis>
|
||||
</taxonomicName>
|
||||
in one of these two families, I should place it amongst the
|
||||
<taxonomicName id="4C287D82FFBDD029FC61FE057C189EA8" authorityName="Darwin" authorityYear="1854" box="[924,1090,423,449]" class="Maxillopoda" family="Balanidae" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="subFamily" subFamily="Chthamalinae">Chthamalinae</taxonomicName>
|
||||
, the other sub-family of the
|
||||
<taxonomicName id="4C287D82FFBDD029FC8FFE647BBE9E89" baseAuthorityName="Gistel" baseAuthorityYear="1848" box="[882,996,454,480]" class="Maxillopoda" family="Balanidae" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="subFamily" subFamily="Balaninae">Balaninae</taxonomicName>
|
||||
”, and on the same page he notes this was written after his Ray Society monograph on the living sessile barnacles (1854: 495) where he gives a more detailed version of the same thing. So, while the verrucomorphs shared characters more or less equally between the scalpellomorphs and balanomorphs, Darwin considered them not just early sessile barnacles but, if compelled to place them within the existing classification, he would include them with the primitive balanomorphs. However, when it came to his classification, Darwin abandoned the
|
||||
<taxonomicName id="4C287D82FFBDD029FCD8FC947BE99C39" box="[805,947,822,848]" class="Maxillopoda" kingdom="Animalia" order="Pedunculata" pageId="10" pageNumber="81" phylum="Arthropoda" rank="order">Pedunculata</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C287D82FFBDD029FC0FFC947C1E9C39" authorityName="Lamarck" authorityYear="1818" box="[1010,1092,822,848]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="81" phylum="Arthropoda" rank="order">Sessilia</taxonomicName>
|
||||
as formal taxa, and subsequent workers (
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FBEAFCF67CEF9C07" author="Pilsbry H. A." box="[1047,1205,852,878]" journalOrPublisher="Bulletin of the United States National Museum" pageId="10" pageNumber="81" pagination="1 - 366" part="93" refId="ref7445" refString="Pilsbry H. A. 1916. - The sessile barnacles (Cirripedia) contained in the collections of the U. S. National Museum; including a monograph of the American species. Bulletin of the United States National Museum 93: 1 - 366." title="The sessile barnacles (Cirripedia) contained in the collections of the U. S. National Museum; including a monograph of the American species" type="journal article" year="1916">Pilsbry 1916</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FD58FCD17B0B9CE4" author="Withers T. H." bookContentInfo="British Museum, Natural History, London, 155 p." box="[677,849,883,909]" pageId="10" pageNumber="81" refId="ref7729" refString="Withers T. H. 1928. - Catalogue of Fossil Cirripedia in the Department of Geology 1 (Triassic and Jurassic). British Museum, Natural History, London, 155 p." title="Catalogue of Fossil Cirripedia in the Department of Geology 1 (Triassic and Jurassic)." type="book" year="1928">Withers 1928</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FC9DFCD07BA49CE4" author="Kruger P." box="[864,1022,882,909]" editor="H. G. Bronns" pageId="10" pageNumber="81" refId="ref6855" refString="Kruger P. 1940. - Cirripedia, in H. G. Bronns Klassen und Ordungen des Tierreichs. 5,1,3,3: 1 - 566. Akademia Verlag, Leipzig." title="Cirripedia" type="book chapter" volumeTitle="Klassen und Ordungen des Tierreichs. 5,1,3,3: 1 - 566. Akademia Verlag, Leipzig" year="1940">Krüger 1940</bibRefCitation>
|
||||
) followed suit. Consequently the sessile barnacles,
|
||||
<taxonomicName id="4C287D82FFBDD029FBAEFC337AA19CA3" authorityName="Pilsbry" authorityYear="1916" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="73" phylum="Arthropoda" rank="subOrder" subOrder="Verrucomorpha">Verrucomorpha</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C287D82FFBDD029FCCBFC127B819CA3" box="[822,987,944,970]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Balanomorpha" pageId="10" pageNumber="81" phylum="Arthropoda" rank="order">Balanomorpha</taxonomicName>
|
||||
, were considered to have sprung independently from the scalpellomorphs (
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FCE1FC4C7C599B61" author="Newman W. A. & Zullo V. A. & Withers T. H." box="[796,1027,1005,1032]" editor="Moore R. C." pageId="10" pageNumber="81" pagination="206 - 295" refId="ref7381" refString="Newman W. A., Zullo V. A. & Withers T. H. 1969. - Cirripedia: R 206 - 295, in Moore R. C. (ed.), Treatise on Invertebrate Paleontology. Part R, Arthropoda 4, Vol. 1. University of Kansas; Geological Society of America, Boulder." title="Cirripedia" type="book chapter" volumeTitle="Treatise on Invertebrate Paleontology. Part R, Arthropoda 4, Vol. 1. University of Kansas; Geological Society of America, Boulder." year="1969">
|
||||
Newman
|
||||
<emphasis id="B95CDA13FFBDD029FC77FC4F7B9A9B61" box="[906,960,1005,1032]" italics="true" pageId="10" pageNumber="81">et al.</emphasis>
|
||||
1969
|
||||
</bibRefCitation>
|
||||
: R266, fig. 133;
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FD58FBAE7BFB9B4F" author="Ghiselin M. T. & Jaffe L." box="[677,929,1036,1062]" journalOrPublisher="Systematic Zoology" pageId="10" pageNumber="81" pagination="132 - 140" part="22" refId="ref6654" refString="Ghiselin M. T. & Jaffe L. 1973. - Phylogenetic classification in Darwin's monograph on the sub-class Cirripedia. Systematic Zoology 22 (2): 132 - 140." title="Phylogenetic classification in Darwin's monograph on the sub-class Cirripedia" type="journal article" year="1973">Ghiselin & Jaffe 1973</bibRefCitation>
|
||||
: fig. 1;
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FC07FBAE7ABA9B2C" author="Newman W. A. & Ross A." journalOrPublisher="San Diego Society of Natural History Memoir" pageId="10" pageNumber="81" pagination="1 - 108" part="9" refId="ref7281" refString="Newman W. A. & Ross A. 1976. - Revision of the balanomorph barnacles; including a catalogue of the species. San Diego Society of Natural History Memoir 9: 1 - 108." title="Revision of the balanomorph barnacles; including a catalogue of the species" type="journal article" year="1976">Newman & Ross 1976</bibRefCitation>
|
||||
: fig. 2).
|
||||
</paragraph>
|
||||
<paragraph id="8B970601FFBDD028FD58FBEB78849E41" blockId="10.[677,1210,208,1706]" lastBlockId="11.[113,647,208,817]" lastPageId="11" lastPageNumber="82" pageId="10" pageNumber="81">
|
||||
In the meantime, however, a third group of sessile barnacles, the presumably extinct Brachylepadomorpha, was discovered (
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FC08FB257CEF9BC8" author="Woodward H." box="[1013,1205,1159,1185]" journalOrPublisher="Geological Magazine, New Series" pageId="10" pageNumber="81" pagination="145 - 152" part="8" refId="ref7803" refString="Woodward H. 1901. - On Pyrgoma cretacea, a cirripede from the Upper Chalk of Norwich and Margate. Geological Magazine, New Series, Decade IV, 8: 145 - 152 + pl. 8 (erratum, p. 240; additional note, p. 528)." title="On Pyrgoma cretacea, a cirripede from the Upper Chalk of Norwich and Margate" type="journal article" year="1901">Woodward 1901</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FD58FB047AB99BA9" author="Woodward H." box="[677,739,1190,1216]" journalOrPublisher="Geology Magazine, New Series" pageId="10" pageNumber="81" pagination="337 - 353" part="3" refId="ref7857" refString="Woodward H. 1906. - Cirripedes from the Trimmingham Chalk and other localities in Norfolk. Geology Magazine, New Series, Decade V, 3: 337 - 353." title="Cirripedes from the Trimmingham Chalk and other localities in Norfolk" type="journal article" year="1906">1906</bibRefCitation>
|
||||
), which Woodward considered central to the origin of the sessile barnacles. However, largely due to the authority of
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FC44FB417CDA9B94" author="Pilsbry H. A." box="[953,1152,1251,1277]" journalOrPublisher="Bulletin of the United States National Museum" pageId="10" pageNumber="81" pagination="1 - 366" part="93" refId="ref7445" refString="Pilsbry H. A. 1916. - The sessile barnacles (Cirripedia) contained in the collections of the U. S. National Museum; including a monograph of the American species. Bulletin of the United States National Museum 93: 1 - 366." title="The sessile barnacles (Cirripedia) contained in the collections of the U. S. National Museum; including a monograph of the American species" type="journal article" year="1916">Pilsbry (1916: 14)</bibRefCitation>
|
||||
, who considered the brachylepadomorphs at least pedunculate barnacles if not simply scalpellomorphs, they came generally accepted as an independent sessile group whereby the sessile barnacles were at least triphyletic (cf.
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FBE1FADE7CEC9AFF" author="Withers T. H." bookContentInfo="British Museum, Natural History, London, 155 p." box="[1052,1206,1404,1430]" pageId="10" pageNumber="81" refId="ref7729" refString="Withers T. H. 1928. - Catalogue of Fossil Cirripedia in the Department of Geology 1 (Triassic and Jurassic). British Museum, Natural History, London, 155 p." title="Catalogue of Fossil Cirripedia in the Department of Geology 1 (Triassic and Jurassic)." type="book" year="1928">Withers 1928</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FD58FA387B6F9ADC" author="Kruger P." box="[677,821,1434,1461]" editor="H. G. Bronns" pageId="10" pageNumber="81" refId="ref6855" refString="Kruger P. 1940. - Cirripedia, in H. G. Bronns Klassen und Ordungen des Tierreichs. 5,1,3,3: 1 - 566. Akademia Verlag, Leipzig." title="Cirripedia" type="book chapter" volumeTitle="Klassen und Ordungen des Tierreichs. 5,1,3,3: 1 - 566. Akademia Verlag, Leipzig" year="1940">Krüger 1940</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FCBFFA387C739ADC" author="Newman W. A. & Zullo V. A. & Withers T. H." box="[834,1065,1434,1461]" editor="Moore R. C." pageId="10" pageNumber="81" pagination="206 - 295" refId="ref7381" refString="Newman W. A., Zullo V. A. & Withers T. H. 1969. - Cirripedia: R 206 - 295, in Moore R. C. (ed.), Treatise on Invertebrate Paleontology. Part R, Arthropoda 4, Vol. 1. University of Kansas; Geological Society of America, Boulder." title="Cirripedia" type="book chapter" volumeTitle="Treatise on Invertebrate Paleontology. Part R, Arthropoda 4, Vol. 1. University of Kansas; Geological Society of America, Boulder." year="1969">
|
||||
Newman
|
||||
<emphasis id="B95CDA13FFBDD029FC52FA387BBF9ADC" box="[943,997,1434,1461]" italics="true" pageId="10" pageNumber="81">et al.</emphasis>
|
||||
1969
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FBCBFA387B7F9ABD" author="Newman W. A. & Ross A." journalOrPublisher="San Diego Society of Natural History Memoir" pageId="10" pageNumber="81" pagination="1 - 108" part="9" refId="ref7281" refString="Newman W. A. & Ross A. 1976. - Revision of the balanomorph barnacles; including a catalogue of the species. San Diego Society of Natural History Memoir 9: 1 - 108." title="Revision of the balanomorph barnacles; including a catalogue of the species" type="journal article" year="1976">Newman & Ross 1976</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FCC9FA1B7BB19ABD" author="Newman W. A." box="[820,1003,1465,1492]" editor="Abele L." pageId="10" pageNumber="81" pagination="197 - 221" refId="ref7006" refString="Newman W. A. 1982. - Cirripedia: 197 - 221, in Abele L. (ed.), The Biology of Crustacea. 1. Academic Press, New York." title="Cirripedia" type="book chapter" volumeTitle="The Biology of Crustacea. 1. Academic Press, New York" year="1982">Newman 1982</bibRefCitation>
|
||||
). However, some authors argued that the brachylepadomorphs were better candidates than the scalpellomorphs for the ancestors of the balanomorphs (
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FB9DF9B47ABA9927" author="Withers T. H." journalOrPublisher="Geology Magazine" pageId="10" pageNumber="81" pagination="321 - 326" part="9" refId="ref7697" refString="Withers T. H. 1912. - The cirripede Brachylepas cretacea H. Woodward. Geology Magazine 9: 321 - 326 & 353 - 359 + pl. 20." title="The cirripede Brachylepas cretacea H. Woodward" type="journal article" year="1912">Withers 1912</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFB97BF0FFBDD029FD17F9967BD59927" author="Newman W. A." box="[746,911,1588,1614]" editor="Southward A. J." pageId="10" pageNumber="81" pagination="3 - 42" refId="ref7098" refString="Newman W. A. 1987. - Evolution of Cirripedes and their major groups: 3 - 42, in Southward A. J. (ed.), Barnacle Biology, Crustacean Issues 5. A. A. Balkema, Rotterdam." title="Evolution of Cirripedes and their major groups" type="book chapter" volumeTitle="Barnacle Biology, Crustacean Issues 5. A. A. Balkema, Rotterdam" year="1987">Newman 1987</bibRefCitation>
|
||||
), and if so the sessile barnacles would be diphyletic rather than triphyletic. Furthermore, it was noted the brachylepadomorphs were also better candidates for the origin of the verrucomorphs, and if so, the sessile barnacles would be monophyletic (
|
||||
<bibRefCitation id="EFB97BF0FFBCD028FE47FF4D7ADB9E60" author="Newman W. A." box="[442,641,239,265]" editor="Southward A. J." pageId="11" pageNumber="82" pagination="3 - 42" refId="ref7098" refString="Newman W. A. 1987. - Evolution of Cirripedes and their major groups: 3 - 42, in Southward A. J. (ed.), Barnacle Biology, Crustacean Issues 5. A. A. Balkema, Rotterdam." title="Evolution of Cirripedes and their major groups" type="book chapter" volumeTitle="Barnacle Biology, Crustacean Issues 5. A. A. Balkema, Rotterdam" year="1987">Newman 1987: 8</bibRefCitation>
|
||||
, 19 & 33).
|
||||
</paragraph>
|
||||
<paragraph id="8B970601FFBCD028FF8CFE8E7C309E0C" blockId="11.[113,647,208,817]" lastBlockId="11.[677,1210,208,817]" pageId="11" pageNumber="82">
|
||||
Compelling evidence for a brachylepadomorph origin of the verrucomorphs appeared with the astonishing discovery of
|
||||
<taxonomicName id="4C287D82FFBCD028FE83FECB79509ECB" authority="(Newman & Hessler 1989)" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBCD028FE83FECB79A99EEB" box="[382,499,361,386]" italics="true" pageId="11" pageNumber="82">Neoverruca</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFB97BF0FFBCD028FDFEFEC879589ECB" author="Newman W. A. & Hessler R. R." journalOrPublisher="San Diego Society of Natural History Transactions" pageId="11" pageNumber="82" pagination="259 - 273" part="21" refId="ref7233" refString="Newman W. A. & Hessler R. R. 1989. - A new abyssal hydrothermal verrucomorphan (Cirripedia; Sessilia): The most primitive living sessile barnacle. San Diego Society of Natural History Transactions 21 (16): 259 - 273." title="A new abyssal hydrothermal verrucomorphan (Cirripedia; Sessilia): The most primitive living sessile barnacle" type="journal article" year="1989">Newman & Hessler 1989</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, whereby monophyly for the sessile barnacles became the favored hypothesis (
|
||||
<bibRefCitation id="EFB97BF0FFBCD028FF87FE64799D9E89" author="Newman W. A. & Hessler R. R." box="[122,455,454,480]" journalOrPublisher="San Diego Society of Natural History Transactions" pageId="11" pageNumber="82" pagination="259 - 273" part="21" refId="ref7233" refString="Newman W. A. & Hessler R. R. 1989. - A new abyssal hydrothermal verrucomorphan (Cirripedia; Sessilia): The most primitive living sessile barnacle. San Diego Society of Natural History Transactions 21 (16): 259 - 273." title="A new abyssal hydrothermal verrucomorphan (Cirripedia; Sessilia): The most primitive living sessile barnacle" type="journal article" year="1989">Newman & Hessler 1989</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFB97BF0FFBCD028FE27FE6479449E97" author="Yamaguchi T. & Newman W. A." journalOrPublisher="Pacific Science" pageId="11" pageNumber="82" pagination="135 - 155" part="44" refId="ref7889" refString="Yamaguchi T. & Newman W. A. 1990. - A new and primitive barnacle (Cirripedia; Balanomorpha) from the North Fiji Basin abyssal hydrothermal field, and its evolutionary implications. Pacific Science 44 (2): 135 - 155." title="A new and primitive barnacle (Cirripedia; Balanomorpha) from the North Fiji Basin abyssal hydrothermal field, and its evolutionary implications" type="journal article" year="1990">Yamaguchi & Newman 1990</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFB97BF0FFBCD028FED1FE467AD89E97" author="Buckeridge J. S. & Newman W. A." box="[300,642,484,510]" journalOrPublisher="Journal of Paleontology" pageId="11" pageNumber="82" pagination="341 - 345" part="66" refId="ref6397" refString="Buckeridge J. S. & Newman W. A. 1992. - A reexamination of Waikalasma (Cirripedia: Thoracica) and its significance in balanomorph phylogeny. Journal of Paleontology 66 (2): 341 - 345." title="A reexamination of Waikalasma (Cirripedia: Thoracica) and its significance in balanomorph phylogeny" type="journal article" year="1992">Buckeridge & Newman 1992</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFB97BF0FFBCD028FF8CFDA179389D74" author="Glenner H. & Grygier M. J. & HOeg J. T. & Jensen P. G. & Schram F. R." box="[113,354,514,541]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="11" pageNumber="82" pagination="365 - 404" part="114" refId="ref6687" refString="Glenner H., Grygier M. J., HOeg J. T., Jensen P. G. & Schram F. R. 1995. - Cladistic analysis of the Cirripedia Thoracica. Zoological Journal of the Linnean Society 114: 365 - 404." title="Cladistic analysis of the Cirripedia Thoracica" type="journal article" year="1995">
|
||||
Glenner
|
||||
<emphasis id="B95CDA13FFBCD028FF20FDA079429D74" box="[221,280,514,541]" italics="true" pageId="11" pageNumber="82">et al.</emphasis>
|
||||
1995
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFB97BF0FFBCD028FE8CFDA07A729D74" author="Newman W. A." box="[369,552,514,541]" editor="Forest J." pageId="11" pageNumber="82" pagination="453 - 540" refId="ref7185" refString="Newman W. A. 1996. - Sous-Classe des Cirripedes, Super-Ordres des Thoraciques et des Acrothoraciques: fascicule 2, 453 - 540, in Forest J. (ed.), Traite de Zoologie. Tome VII: Crustaces. Masson, Paris." title="Sous-Classe des Cirripedes, Super-Ordres des Thoraciques et des Acrothoraciques: fascicule 2" type="book chapter" volumeTitle="Traite de Zoologie. Tome VII: Crustaces. Masson, Paris." year="1996">Newman 1996</bibRefCitation>
|
||||
). When viewed from one side, this barnacle has the same arrangement of opercular, wall and imbricating plates as a brachylepadomorph. The opposite side was peculiar and if there had been but
|
||||
<specimenCount id="9D2ECD88FFBCD028FDE4FDDF78FB9DDF" pageId="11" pageNumber="82" type="generic">one specimen</specimenCount>
|
||||
available it easily could have been considered an imperfectly developed brachylepadomorph, due to crowding, predation damage or the like. But there were numerous specimens each with an “anomalous” right or left side. The basis for the anomaly was the tergum and scutum of that side being immovably integrated into the wall, as in verrucomorphs. Thus, a better “missing link” between the brachylepadomorphs and verrucomorphs could hardly have been imagined.
|
||||
</paragraph>
|
||||
<paragraph id="8B970601FFBCD028FD58FEC87B979C58" blockId="11.[677,1210,208,817]" pageId="11" pageNumber="82">
|
||||
<emphasis id="B95CDA13FFBCD028FD58FEC87B0F9EED" box="[677,853,362,388]" italics="true" pageId="11" pageNumber="82">
|
||||
<taxonomicName id="4C287D82FFBCD028FD58FEC87B0B9EED" authorityName="Newman" authorityYear="2000" box="[677,849,362,388]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">Imbricaverruca</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
while wholly a neoverrucid in organization, differs from
|
||||
<taxonomicName id="4C287D82FFBCD028FC43FE2A7C699EC8" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[958,1075,392,417]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBCD028FC43FE2A7C699EC8" box="[958,1075,392,417]" italics="true" pageId="11" pageNumber="82">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
in outwardly looking more like a verrucid than a brachylepadomorph (
|
||||
<figureCitation id="13131A84FFBCD028FCB9FE647BD89E89" box="[836,898,454,480]" captionStart="FIG" captionStartId="2.[114,125,799,816]" captionTargetBox="[262,1053,208,766]" captionTargetId="figure-307@2.[248,1067,202,776]" captionTargetPageId="2" captionText="FIG. 1. — A, Imbricaverruca yamaguchii n. gen. and n. sp. (holotype, MNHN Ci2710); B, Verruca s.l., both viewed from above with the right scutum and tergum forming the operculum.Note that in the former the four plated wall appears to be covered largely by imbricating plates and the operculum includes a large median latus,characters that readily distinguish it from Neoverruca. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5391397" httpUri="https://zenodo.org/record/5391397/files/figure.png" pageId="11" pageNumber="82">Fig. 1</figureCitation>
|
||||
). Thus the question arises as to whether it represents a grade between
|
||||
<taxonomicName id="4C287D82FFBCD028FD58FDA07B449D72" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[677,798,514,539]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBCD028FD58FDA07B449D72" box="[677,798,514,539]" italics="true" pageId="11" pageNumber="82">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
and the proverrucids. The evidence from both juvenile and adult morphology suggests not. In short,
|
||||
<taxonomicName id="4C287D82FFBCD028FC85FDE27C4C9D33" authorityName="Newman" authorityYear="2000" box="[888,1046,576,602]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBCD028FC85FDE27C4C9D33" box="[888,1046,576,602]" italics="true" pageId="11" pageNumber="82">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
has capitalized on the neoverrucid plan rather than making the reductions and acquiring proportions in the direction of those found in modern forms (
|
||||
<figureCitation id="13131A84FFBCD028FD51FD197AB59DBC" box="[684,751,699,725]" captionStart="Fig" captionStartId="7.[113,141,1469,1486]" captionTargetBox="[112,1165,215,1445]" captionTargetId="figure-22@7.[139,1170,219,1510]" captionTargetPageId="7" captionText="Fig. 4. — Schematic plan views and transverse sections of grades of skeletal organization from the most primitive sessile barnacles (A, Brachylepadomorpha) through the asymmetrical sessile barnacles (B-H, Verrucomorpha). Of the eight verrucid genera currently recognized (Young 1998) only the four most representative ones (Altiverruca, Newmaniverruca, Metaverruca and Verruca s.s.) are depicted here. Right sides are to the viewers right and rostral ends are downward in plan views or coming out of the page in transverse sections. The transverse sections are slightly rostral of the rostro-carinal gap or suture. While there are numerous whorls of imbricating plates standing in tiers four plates high in brachylepadomorphs, and on the movable side of neoverrucids, only those of the R-C gap are labeled. Abbreviations: C, R, carina and rostrum; cl1-4, l1-4, rl1-4, carinolateral, lateral and rostrolateral tiers of imbricating plates respectively; cl, rl, the pair of imbricating plates in proverrucids; FS, FT, fixed scutum and tergum; hl, hinge line; L, median latus (completely lost in proverrucids and verrucids); m, myophore; MS, MT, movable scutum and tergum; sam, scutal adductor muscle; S, T, normal terga and scuta in brachylepadomorphs; dashed horizontal lines: membranous basis (transverse sections F-H after Young 1998)." figureDoi="http://doi.org/10.5281/zenodo.13988759" httpUri="https://zenodo.org/record/13988759/files/figure.png" pageId="11" pageNumber="82">Fig. 4</figureCitation>
|
||||
). Thus the
|
||||
<taxonomicName id="4C287D82FFBCD028FC93FD187BB99DBA" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[878,995,698,723]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBCD028FC93FD187BB99DBA" box="[878,995,698,723]" italics="true" pageId="11" pageNumber="82">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C287D82FFBCD028FBE6FD197CE39DBC" authorityName="Newman" authorityYear="2000" box="[1051,1209,699,725]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBCD028FBE6FD197CE39DBC" box="[1051,1209,699,725]" italics="true" pageId="11" pageNumber="82">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
plans are markedly divergent, the former’s being better suited to the derivation of the proverrucid/verrucid plan (
|
||||
<tableCitation id="C6AA33BAFFBCD028FC96FCB57BE49C58" box="[875,958,791,817]" captionStart="TABLE" captionStartId="11.[113,124,903,920]" captionTargetPageId="11" captionText="TABLE 3. — Comparison of two juvenile and four adult characters in two neoverrucid and one verrucid genus." pageId="11" pageNumber="82">Table 3</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF575689FFBCD028FF8CFC257BA09CF1" box="[113,1018,903,920]" pageId="11" pageNumber="82" startId="11.[113,124,903,920]" targetBox="[113,1207,967,1153]" targetIsTable="true" targetPageId="11">
|
||||
<paragraph id="8B970601FFBCD028FF8CFC257BA09CF1" blockId="11.[113,1018,903,920]" box="[113,1018,903,920]" pageId="11" pageNumber="82">TABLE 3. — Comparison of two juvenile and four adult characters in two neoverrucid and one verrucid genus.</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B970601FFBCD028FE50FC657CFB9BE8" pageId="11" pageNumber="82">
|
||||
<table id="F928F4A1FFBC2FDCFF8CFC657CED9BE8" box="[113,1207,967,1153]" gridcols="4" gridrows="7" pageId="11" pageNumber="82">
|
||||
<tr id="35180443FFBC2FDCFF8CFC657CED9CB2" box="[113,1207,967,987]" gridrow="0" pageId="11" pageNumber="82" rowspan-0="1">
|
||||
<th id="76C96D3FFFBC2FDCFE50FC657A839CB2" box="[429,729,967,987]" gridcol="1" gridrow="0" pageId="11" pageNumber="82">Neoverruca</th>
|
||||
<th id="76C96D3FFFBC2FDCFD11FC657BBC9CB2" box="[748,998,967,987]" gridcol="2" gridrow="0" pageId="11" pageNumber="82">Imbricaverruca</th>
|
||||
<th id="76C96D3FFFBC2FDCFBF6FC657CED9CB2" box="[1035,1207,967,987]" gridcol="3" gridrow="0" pageId="11" pageNumber="82">
|
||||
<emphasis id="B95CDA13FFBCD028FBF6FC657C009CB2" bold="true" box="[1035,1114,967,987]" italics="true" pageId="11" pageNumber="82">Verruca</emphasis>
|
||||
</th>
|
||||
</tr>
|
||||
<tr id="35180443FFBC2FDCFF8CFC577CED9B60" box="[113,1207,1013,1033]" gridrow="1" pageId="11" pageNumber="82">
|
||||
<th id="76C96D3FFFBC2FDCFF8CFC5779C29B60" box="[113,408,1013,1033]" gridcol="0" gridrow="1" pageId="11" pageNumber="82">1) Juvenile pedunculate stages</th>
|
||||
<td id="76C96D3FFFBC2FDCFE50FC577A839B60" box="[429,729,1013,1033]" gridcol="1" gridrow="1" pageId="11" pageNumber="82">Several stages pedunculate</td>
|
||||
<td id="76C96D3FFFBC2FDCFD11FC577BBC9B60" box="[748,998,1013,1033]" gridcol="2" gridrow="1" pageId="11" pageNumber="82">Likely several stages</td>
|
||||
<td id="76C96D3FFFBC2FDCFBF6FC577CED9B60" box="[1035,1207,1013,1033]" gridcol="3" gridrow="1" pageId="11" pageNumber="82">Peduncle vestigial</td>
|
||||
</tr>
|
||||
<tr id="35180443FFBC2FDCFF8CFBAF7CED9B48" box="[113,1207,1037,1057]" gridrow="2" pageId="11" pageNumber="82">
|
||||
<th id="76C96D3FFFBC2FDCFF8CFBAF79C29B48" box="[113,408,1037,1057]" gridcol="0" gridrow="2" pageId="11" pageNumber="82">2) Juvenile carina</th>
|
||||
<td id="76C96D3FFFBC2FDCFE50FBAF7A839B48" box="[429,729,1037,1057]" gridcol="1" gridrow="2" pageId="11" pageNumber="82">Higher than wide</td>
|
||||
<td id="76C96D3FFFBC2FDCFD11FBAF7BBC9B48" box="[748,998,1037,1057]" gridcol="2" gridrow="2" pageId="11" pageNumber="82">Wider than high</td>
|
||||
<td id="76C96D3FFFBC2FDCFBF6FBAF7CED9B48" box="[1035,1207,1037,1057]" gridcol="3" gridrow="2" pageId="11" pageNumber="82">Higher than wide</td>
|
||||
</tr>
|
||||
<tr id="35180443FFBC2FDCFF8CFB877CED9B50" box="[113,1207,1061,1081]" gridrow="3" pageId="11" pageNumber="82">
|
||||
<th id="76C96D3FFFBC2FDCFF8CFB8779C29B50" box="[113,408,1061,1081]" gridcol="0" gridrow="3" pageId="11" pageNumber="82">3) Adult median latus</th>
|
||||
<td id="76C96D3FFFBC2FDCFE50FB877A839B50" box="[429,729,1061,1081]" gridcol="1" gridrow="3" pageId="11" pageNumber="82">Vestigial</td>
|
||||
<td id="76C96D3FFFBC2FDCFD11FB877BBC9B50" box="[748,998,1061,1081]" gridcol="2" gridrow="3" pageId="11" pageNumber="82">Well-developed</td>
|
||||
<td id="76C96D3FFFBC2FDCFBF6FB877CED9B50" box="[1035,1207,1061,1081]" gridcol="3" gridrow="3" pageId="11" pageNumber="82">Lost</td>
|
||||
</tr>
|
||||
<tr id="35180443FFBC2FDCFF8CFB9F7CED9B38" box="[113,1207,1085,1105]" gridrow="4" pageId="11" pageNumber="82">
|
||||
<th id="76C96D3FFFBC2FDCFF8CFB9F79C29B38" box="[113,408,1085,1105]" gridcol="0" gridrow="4" pageId="11" pageNumber="82">4) Imbricating plates</th>
|
||||
<td id="76C96D3FFFBC2FDCFE50FB9F7A839B38" box="[429,729,1085,1105]" gridcol="1" gridrow="4" pageId="11" pageNumber="82">Reduced in number, deciduous</td>
|
||||
<td id="76C96D3FFFBC2FDCFD11FB9F7BBC9B38" box="[748,998,1085,1105]" gridcol="2" gridrow="4" pageId="11" pageNumber="82">Complete, well-developed</td>
|
||||
<td id="76C96D3FFFBC2FDCFBF6FB9F7CED9B38" box="[1035,1207,1085,1105]" gridcol="3" gridrow="4" pageId="11" pageNumber="82">Lost</td>
|
||||
</tr>
|
||||
<tr id="35180443FFBC2FDCFF8CFBF77CED9B00" box="[113,1207,1109,1129]" gridrow="5" pageId="11" pageNumber="82">
|
||||
<th id="76C96D3FFFBC2FDCFF8CFBF779C29B00" box="[113,408,1109,1129]" gridcol="0" gridrow="5" pageId="11" pageNumber="82">5) Fixed scutum & tergum</th>
|
||||
<td id="76C96D3FFFBC2FDCFE50FBF77A839B00" box="[429,729,1109,1129]" gridcol="1" gridrow="5" pageId="11" pageNumber="82">Normal, higher than wide</td>
|
||||
<td id="76C96D3FFFBC2FDCFD11FBF77BBC9B00" box="[748,998,1109,1129]" gridcol="2" gridrow="5" pageId="11" pageNumber="82">Reduced wider than high</td>
|
||||
<td id="76C96D3FFFBC2FDCFBF6FBF77CED9B00" box="[1035,1207,1109,1129]" gridcol="3" gridrow="5" pageId="11" pageNumber="82">As wide as high</td>
|
||||
</tr>
|
||||
<tr id="35180443FFBC2FDCFF8CFBCF7CED9BE8" box="[113,1207,1133,1153]" gridrow="6" pageId="11" pageNumber="82">
|
||||
<th id="76C96D3FFFBC2FDCFF8CFBCF79C29BE8" box="[113,408,1133,1153]" gridcol="0" gridrow="6" pageId="11" pageNumber="82">6) Rostrum & carina</th>
|
||||
<td id="76C96D3FFFBC2FDCFE50FBCF7A839BE8" box="[429,729,1133,1153]" gridcol="1" gridrow="6" pageId="11" pageNumber="82">Normal, higher than wide*</td>
|
||||
<td id="76C96D3FFFBC2FDCFD11FBCF7BBC9BE8" box="[748,998,1133,1153]" gridcol="2" gridrow="6" pageId="11" pageNumber="82">Reduced, wider high</td>
|
||||
<td id="76C96D3FFFBC2FDCFBF6FBCF7CED9BE8" box="[1035,1207,1133,1153]" gridcol="3" gridrow="6" pageId="11" pageNumber="82">As wide as high</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<paragraph id="8B970601FFBCD028FF8CFB3C7A6B9BC6" blockId="11.[113,561,1182,1199]" box="[113,561,1182,1199]" pageId="11" pageNumber="82">
|
||||
<tableNote id="76CE078FFFBCD028FF8CFB3C7A6B9BC6" box="[113,561,1182,1199]" pageId="11" pageNumber="82" targetBox="[113,1207,967,1153]" targetPageId="11">*R-C gap less on movable side, as in all three genera.</tableNote>
|
||||
</paragraph>
|
||||
<paragraph id="8B970601FFBCD02FFF8CFAA179E69C7B" blockId="11.[113,647,1283,1709]" lastBlockId="12.[114,647,208,786]" lastPageId="12" lastPageNumber="83" pageId="11" pageNumber="82">
|
||||
In
|
||||
<tableCitation id="C6AA33BAFFBCD028FF6FFAA678BF9A77" box="[146,229,1284,1310]" captionStart="TABLE" captionStartId="11.[113,124,903,920]" captionTargetPageId="11" captionText="TABLE 3. — Comparison of two juvenile and four adult characters in two neoverrucid and one verrucid genus." pageId="11" pageNumber="82">Table 3</tableCitation>
|
||||
, character 1 concerns the existence of fully pedunculate juveniles during the ontogeny of
|
||||
<taxonomicName id="4C287D82FFBCD028FF6DFAE3795C9A33" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[144,262,1345,1370]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBCD028FF6DFAE3795C9A33" box="[144,262,1345,1370]" italics="true" pageId="11" pageNumber="82">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
missing in the ontogeny of verrucids, possibly in proverrucids, and in balanomorphs (
|
||||
<bibRefCitation id="EFB97BF0FFBCD028FEFEFADD79EF9AF0" author="Newman W. A." box="[259,437,1407,1433]" journalOrPublisher="Bulletin of Marine Science" pageId="11" pageNumber="82" pagination="467 - 477" part="45" refId="ref7144" refString="Newman W. A. 1989. - Juvenile ontogeny and metamorphosis in the most primitive living sessile barnacle, Neoverruca, from an abyssal hydrothermal spring. Bulletin of Marine Science 45 (2): 467 - 477." title="Juvenile ontogeny and metamorphosis in the most primitive living sessile barnacle, Neoverruca, from an abyssal hydrothermal spring" type="journal article" year="1989">Newman 1989</bibRefCitation>
|
||||
). Considering the retention of well-developed imbricating plates and the median latus in
|
||||
<taxonomicName id="4C287D82FFBCD028FE72FA1E7A6C9ABF" authorityName="Newman" authorityYear="2000" box="[399,566,1468,1494]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBCD028FE72FA1E7A6C9ABF" box="[399,566,1468,1494]" italics="true" pageId="11" pageNumber="82">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
, it too likely has several pedunculate stages in its ontogeny. If so, like
|
||||
<taxonomicName id="4C287D82FFBCD028FEFAFA5B7926997B" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[263,380,1529,1554]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBCD028FEFAFA5B7926997B" box="[263,380,1529,1554]" italics="true" pageId="11" pageNumber="82">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
, it is not becoming specialized like verrucids as far as elimination of its pedunculate stages is concerned. Furthermore, it will be noted (character 2), the carina is much wider than high in the presumed juvenile as well as the adult of
|
||||
<taxonomicName id="4C287D82FFBCD028FEEBF93179EE99C4" authorityName="Newman" authorityYear="2000" box="[278,436,1683,1709]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBCD028FEEBF93179EE99C4" box="[278,436,1683,1709]" italics="true" pageId="11" pageNumber="82">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
, and together with the reduced rostrum and fixed scutum and tergum, it forms a ring reinforcing the uppermost imbricating plates rather than forming a major part of the wall. A reduction in the extent of the primary wall in
|
||||
<taxonomicName id="4C287D82FFBCD028FCAAFADD7BAF9AF0" authorityName="Newman" authorityYear="2000" box="[855,1013,1407,1433]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBCD028FCAAFADD7BAF9AF0" box="[855,1013,1407,1433]" italics="true" pageId="11" pageNumber="82">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
is an autapomorphy leading away from rather than towards verrucids. Character 3 concerns the median latus, on the verge of being lost in
|
||||
<taxonomicName id="4C287D82FFBCD028FC4EFA787C729A9A" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[947,1064,1498,1523]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBCD028FC4EFA787C729A9A" box="[947,1064,1498,1523]" italics="true" pageId="11" pageNumber="82">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
but very well developed in
|
||||
<taxonomicName id="4C287D82FFBCD028FCBBFA587BA9997D" authorityName="Newman" authorityYear="2000" box="[838,1011,1530,1556]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBCD028FCBBFA587BA9997D" box="[838,1011,1530,1556]" italics="true" pageId="11" pageNumber="82">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
. It is apparently completely lost in proverrucids, as in verrucids, so
|
||||
<taxonomicName id="4C287D82FFBCD028FD38F9947B669926" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[709,828,1590,1615]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBCD028FD38F9947B669926" box="[709,828,1590,1615]" italics="true" pageId="11" pageNumber="82">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
rather than
|
||||
<taxonomicName id="4C287D82FFBCD028FC36F9957C369938" authorityName="Newman" authorityYear="2000" box="[971,1132,1591,1617]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBCD028FC36F9957C369938" box="[971,1132,1591,1617]" italics="true" pageId="11" pageNumber="82">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
is also closer to them in this regard. Character 4 involves the relative importance of the imbricating plates; their reduction and deciduous nature in
|
||||
<taxonomicName id="4C287D82FFBBD02FFF8FFF7278B09F80" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[114,234,208,233]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBBD02FFF8FFF7278B09F80" box="[114,234,208,233]" italics="true" pageId="12" pageNumber="83">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
in contrast to their further development and importance in
|
||||
<taxonomicName id="4C287D82FFBBD02FFE7EFF4D7A7B9E60" authorityName="Newman" authorityYear="2000" box="[387,545,239,265]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBBD02FFE7EFF4D7A7B9E60" box="[387,545,239,265]" italics="true" pageId="12" pageNumber="83">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
. Character 5 pertains to the relative proportions of the fixed terga and scuta (also note their degree of integration with the rostra and scuta in forming the wall under character 6. In being wider than high, and in their reduced contribution to the wall, those of
|
||||
<taxonomicName id="4C287D82FFBBD02FFEF5FE0579FC9EA8" authorityName="Newman" authorityYear="2000" box="[264,422,423,449]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBBD02FFEF5FE0579FC9EA8" box="[264,422,423,449]" italics="true" pageId="12" pageNumber="83">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
differ conspicuously from the relatively unmodified ones of
|
||||
<taxonomicName id="4C287D82FFBBD02FFF8FFE4678BD9E94" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[114,231,484,509]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBBD02FFF8FFE4678BD9E94" box="[114,231,484,509]" italics="true" pageId="12" pageNumber="83">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
and the squarely proportioned ones of
|
||||
<taxonomicName id="4C287D82FFBBD02FFF8FFDA078989D72" authorityName="Schumacher" authorityYear="1817" box="[114,194,514,539]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBBD02FFF8FFDA078989D72" box="[114,194,514,539]" italics="true" pageId="12" pageNumber="83">Verruca</emphasis>
|
||||
</taxonomicName>
|
||||
. Character 6 pertains to the concomitant changes in the rostrum and carina seen in character 5; e.g. reduction in height and relative importance to the wall which distinguishes
|
||||
<taxonomicName id="4C287D82FFBBD02FFF8FFDDC794F9DF1" authorityName="Newman" authorityYear="2000" box="[114,277,638,664]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBBD02FFF8FFDDC794F9DF1" box="[114,277,638,664]" italics="true" pageId="12" pageNumber="83">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
from
|
||||
<taxonomicName id="4C287D82FFBBD02FFEA3FDDF798D9DFF" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[350,471,637,662]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBBD02FFEA3FDDF798D9DFF" box="[350,471,637,662]" italics="true" pageId="12" pageNumber="83">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C287D82FFBBD02FFDEFFDDF7A3E9DFF" authorityName="Schumacher" authorityYear="1817" box="[530,612,637,662]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBBD02FFDEFFDDF7A3E9DFF" box="[530,612,637,662]" italics="true" pageId="12" pageNumber="83">Verruca</emphasis>
|
||||
</taxonomicName>
|
||||
. It follows, despite
|
||||
<taxonomicName id="4C287D82FFBBD02FFED7FD3E79949DDF" authorityName="Newman" authorityYear="2000" box="[298,462,668,694]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBBD02FFED7FD3E79949DDF" box="[298,462,668,694]" italics="true" pageId="12" pageNumber="83">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
facies similarity with higher verrucomorphs, that
|
||||
<taxonomicName id="4C287D82FFBBD02FFDFBFD187ADD9DBA" baseAuthorityName="Newman & Hessler" baseAuthorityYear="1989" box="[518,647,698,723]" class="Maxillopoda" family="Neoverrucidae" genus="Neoverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBBD02FFDFBFD187ADD9DBA" box="[518,647,698,723]" italics="true" pageId="12" pageNumber="83">Neoverruca</emphasis>
|
||||
</taxonomicName>
|
||||
comes much closer to their ancestral organization plan than does
|
||||
<taxonomicName id="4C287D82FFBBD02FFEEAFD5A79EF9C7B" authorityName="Newman" authorityYear="2000" box="[279,437,760,786]" class="Maxillopoda" family="Neoverrucidae" genus="Imbricaverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95CDA13FFBBD02FFEEAFD5A79EF9C7B" box="[279,437,760,786]" italics="true" pageId="12" pageNumber="83">Imbricaverruca</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
Loading…
Reference in a new issue