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<document id="BE0D3A4EAA02DC97921B3738279F3639" ID-DOI="10.1206/321.1-1" ID-ISSN="0003-0090" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="juliana" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721338539162" checkinUser="felipe" docAuthor="Miller, James S" docDate="2009" docId="03FF87E0FF1B9EEFBCD410B9FC9D4FB1" docLanguage="en" docName="BulAmeMusNatHis.2009.B321p2.675-1022.pdf" docOrigin="Bulletin of the American Museum of Natural History 2009 (321)" docSource="http://dx.doi.org/10.1206/321.1-1" docStyle="DocumentStyle:2B34016C1EBFE58B07368F9517618743.4:BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleId="2B34016C1EBFE58B07368F9517618743" docStyleName="BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleVersion="4" docTitle="Scea grandis, new combination" docType="treatment" docVersion="1" lastPageNumber="849" masterDocId="FFC6FF98FFB59E41BC70127DFF984D13" masterDocTitle="Generic Revision Of The Dioptinae (Lepidoptera: Noctuoidea: Notodontidae) Part 2: Josiini" masterLastPageNumber="1022" masterPageNumber="675" pageNumber="849" updateTime="1722273871849" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8F57B7A242CC6E40EA6CCD94CCA93928">Generic Revision Of The Dioptinae (Lepidoptera: Noctuoidea: Notodontidae) Part 2: Josiini</mods:title>
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<treatment id="03FF87E0FF1B9EEFBCD410B9FC9D4FB1" LSID="urn:lsid:plazi:treatment:03FF87E0FF1B9EEFBCD410B9FC9D4FB1" httpUri="http://treatment.plazi.org/id/03FF87E0FF1B9EEFBCD410B9FC9D4FB1" lastPageNumber="849" pageId="174" pageNumber="849">
<subSubSection id="C34C657DFF1B9EEFBCD410B9FE2E4FE5" pageId="174" pageNumber="849" type="nomenclature">
<paragraph id="8BE936F6FF1B9EEFBCD410B9FE2E4FE5" blockId="174.[164,617,707,759]" pageId="174" pageNumber="849">
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<taxonomicName id="4C564D75FF1B9EEFBCD410B9FE044FCB" ID-CoL="BWX7L" authority="(Druce)" baseAuthorityName="Druce" baseAuthorityYear="1900" box="[164,412,707,730]" class="Insecta" family="Notodontidae" genus="Scea" kingdom="Animalia" order="Lepidoptera" pageId="174" pageNumber="849" phylum="Arthropoda" rank="species" species="grandis" status="comb. nov.">
<emphasis id="B922EAE4FF1B9EEFBCD410B9FEAE4FC9" box="[164,310,707,730]" italics="true" pageId="174" pageNumber="849">Scea grandis</emphasis>
(Druce)
</taxonomicName>
,
<emphasis id="B922EAE4FF1B9EEFBDD910BEFDF14FCA" bold="true" box="[425,617,707,729]" pageId="174" pageNumber="849">
<taxonomicNameLabel id="A211579FFF1B9EEFBDD910BEFDF14FCA" box="[425,617,707,729]" pageId="174" pageNumber="849" rank="species">new combination</taxonomicNameLabel>
</emphasis>
Plate 33
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C657DFF1B9EEFBCF01171FD894E33" box="[128,529,780,801]" pageId="174" pageNumber="849" type="reference_group">
<paragraph id="8BE936F6FF1B9EEFBCF01171FD894E33" blockId="174.[128,654,780,918]" box="[128,529,780,801]" pageId="174" pageNumber="849">
<treatmentCitationGroup id="AB4611D8FF1B9EEFBCF01171FD894E33" box="[128,529,780,801]" pageId="174" pageNumber="849">
<treatmentCitation id="0AF710E7FF1B9EEFBCF01171FD954E33" author="Druce, H." box="[128,525,780,801]" page="510" pageId="174" pageNumber="849" year="1900">
<taxonomicName id="4C564D75FF1B9EEFBCF01171FD954E33" ID-CoL="BXNW5" authority="Druce, 1900: 510" authorityName="Druce" authorityPageNumber="510" authorityYear="1900" box="[128,525,780,801]" class="Insecta" family="Notodontidae" genus="Brachyglene" kingdom="Animalia" order="Lepidoptera" pageId="174" pageNumber="849" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B922EAE4FF1B9EEFBCF01171FED74E33" box="[128,335,780,800]" italics="true" pageId="174" pageNumber="849">Brachyglene grandis</emphasis>
<bibRefCitation id="EFC74B07FF1B9EEFBD271173FD954E33" author="Druce, H." box="[343,525,781,801]" pageId="174" pageNumber="849" pagination="507 - 527" refId="ref139019" refString="Druce, H. 1900. Descriptions of some new genera and species of Heterocera from tropical South America. Annals and Magazine of Natural History 7 (5): 507 - 527." type="journal article" year="1900">Druce, 1900: 510</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34C657DFF1B9EEFBCEC115AFF6C4E85" pageId="174" pageNumber="849" type="materials_examined">
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<collectionCode id="ED47AE33FF1B9EEFBCEC115AFF444E2E" box="[156,220,807,831]" pageId="174" pageNumber="849">TYPE</collectionCode>
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:
<taxonomicName id="4C564D75FF1B9EEFBDF31154FDCF4E2C" authority=", Sierra" authorityName="Sierra" box="[387,599,809,831]" class="Insecta" family="Notodontidae" genus="Erbessa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="174" pageNumber="849" phylum="Arthropoda" rank="species" species="colombia">Colombia, Sierra</taxonomicName>
del Libane,
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:
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, leg. H.H. Smith (
<collectionCode id="ED47AE33FF1B9EEFBCF811FCFF7F4E85" box="[136,231,897,918]" pageId="174" pageNumber="849">BMNH</collectionCode>
).
</paragraph>
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:
<taxonomicName id="4C564D75FF1B9EEFBD4D11C9FE494ED9" baseAuthorityName="Druce" baseAuthorityYear="1900" box="[317,465,947,970]" class="Insecta" family="Notodontidae" genus="Scea" kingdom="Animalia" order="Lepidoptera" pageId="174" pageNumber="849" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B922EAE4FF1B9EEFBD4D11C9FE494ED9" box="[317,465,947,970]" italics="true" pageId="174" pageNumber="849">Scea grandis</emphasis>
</taxonomicName>
resembles both
<taxonomicName id="4C564D75FF1B9EEFBCF011ACFE9C4EF4" authorityName="Warren" authorityYear="1900" baseAuthorityName="Miller and Otero" baseAuthorityYear="1900" box="[128,260,976,999]" class="Insecta" family="Notodontidae" genus="Scea" kingdom="Animalia" order="Lepidoptera" pageId="174" pageNumber="849" phylum="Arthropoda" rank="species" species="discinota">
<emphasis id="B922EAE4FF1B9EEFBCF011ACFE9C4EF4" box="[128,260,976,999]" italics="true" pageId="174" pageNumber="849">S. discinota</emphasis>
</taxonomicName>
and
<taxonomicName id="4C564D75FF1B9EEFBD3011ACFE534EF4" authorityName="Walker" authorityYear="1854" baseAuthorityName="Walker" box="[320,459,976,999]" class="Insecta" family="Notodontidae" genus="Thirmida" kingdom="Animalia" order="Lepidoptera" pageId="174" pageNumber="849" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B922EAE4FF1B9EEFBD3011ACFE534EF4" box="[320,459,976,999]" italics="true" pageId="174" pageNumber="849">S. dimidiata</emphasis>
</taxonomicName>
in size and wing pattern (pl. 33), and genital morphology supports a close relationship between them. The genitalia of
<taxonomicName id="4C564D75FF1B9EEFBD311654FE62492D" authorityName="Hering" authorityYear="1925" baseAuthorityName="Hering" box="[321,506,1065,1086]" class="Insecta" family="Notodontidae" genus="Thirmida" kingdom="Animalia" order="Lepidoptera" pageId="174" pageNumber="849" phylum="Arthropoda" rank="species" species="circumscripta">
<emphasis id="B922EAE4FF1B9EEFBD311654FE62492D" box="[321,506,1065,1086]" italics="true" pageId="174" pageNumber="849">S. circumscripta</emphasis>
</taxonomicName>
(pl. 33) also suggest membership in this group, which seems to form a subclade of
<taxonomicName id="4C564D75FF1B9EEFBE79161EFDA7496B" authorityName="Walker" authorityYear="1854" box="[521,575,1123,1144]" class="Insecta" family="Notodontidae" genus="Scea" kingdom="Animalia" order="Lepidoptera" pageId="174" pageNumber="849" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAE4FF1B9EEFBE79161EFDA7496B" box="[521,575,1123,1144]" italics="true" pageId="174" pageNumber="849">Scea</emphasis>
</taxonomicName>
.
<taxonomicName id="4C564D75FF1B9EEFBE28161EFF4B4985" baseAuthorityName="Druce" baseAuthorityYear="1900" class="Insecta" family="Notodontidae" genus="Scea" kingdom="Animalia" order="Lepidoptera" pageId="174" pageNumber="849" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B922EAE4FF1B9EEFBE28161EFF4B4985" italics="true" pageId="174" pageNumber="849">Scea grandis</emphasis>
</taxonomicName>
is distinctive in exhibiting less orange at the base of the FW and HW than the other three.
</paragraph>
<paragraph id="8BE936F6FF1B9EEFBCED16A5FEC84B5F" blockId="174.[128,654,946,1759]" pageId="174" pageNumber="849">
This species is known from few specimens:
<specimenCount id="9D50FD7FFF1B9EEFBCF01688FE8A4818" box="[128,274,1269,1291]" count="4" pageId="174" pageNumber="849" type="female">four females</specimenCount>
(including the
<typeStatus id="54ED8854FF1B9EEFBDB61688FE614819" box="[454,505,1269,1290]" pageId="174" pageNumber="849">type</typeStatus>
) and a male at the BMNH;
<specimenCount id="9D50FD7FFF1B9EEFBD48176EFE3B483B" box="[312,419,1298,1320]" count="1" pageId="174" pageNumber="849" type="male">one male</specimenCount>
and
<specimenCount id="9D50FD7FFF1B9EEFBD92176EFDF6483B" box="[482,622,1298,1320]" count="2" pageId="174" pageNumber="849" type="female">two females</specimenCount>
at the ZMH; and a single male in the CMNH collection. The specimens of
<taxonomicName id="4C564D75FF1B9EEFBE7B1733FD164870" baseAuthorityName="Druce" baseAuthorityYear="1900" box="[523,654,1356,1379]" class="Insecta" family="Notodontidae" genus="Scea" kingdom="Animalia" order="Lepidoptera" pageId="174" pageNumber="849" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B922EAE4FF1B9EEFBE7B1733FD164870" box="[523,654,1356,1379]" italics="true" pageId="174" pageNumber="849">T. grandis</emphasis>
</taxonomicName>
dissected for this study were collected in the Santa Marta mountain range of northeastern
<collectingCountry id="F3417666FF1B9EEFBCF017D8FF6F48A8" box="[128,247,1445,1467]" name="Colombia" pageId="174" pageNumber="849">Colombia</collectingCountry>
, near the border with
<collectingCountry id="F3417666FF1B9EEFBD8A17D8FDEC48A8" box="[506,628,1445,1467]" name="Venezuela" pageId="174" pageNumber="849">Venezuela</collectingCountry>
, a fascinating region rich in endemic species, but an area that is currently too dangerous for travel. That locality is also near
<collectingRegion id="4992F814FF1B9EEFBE411781FD124B01" box="[561,650,1532,1554]" country="Venezuela" name="Merida" pageId="174" pageNumber="849">Mérida</collectingRegion>
,
<collectingCountry id="F3417666FF1B9EEFBCF01467FF624B23" box="[128,250,1562,1584]" name="Venezuela" pageId="174" pageNumber="849">Venezuela</collectingCountry>
, the home of
<taxonomicName id="4C564D75FF1B9EEFBDD91467FED44B5F" authority="(Miller and Otero, 1994)" baseAuthorityName="Miller and Otero" baseAuthorityYear="1994" class="Insecta" family="Notodontidae" genus="Scea" kingdom="Animalia" order="Lepidoptera" pageId="174" pageNumber="849" phylum="Arthropoda" rank="species" species="discinota">
<emphasis id="B922EAE4FF1B9EEFBDD91467FDAA4B23" box="[425,562,1561,1584]" italics="true" pageId="174" pageNumber="849">S. discinota</emphasis>
(
<bibRefCitation id="EFC74B07FF1B9EEFBE351467FEDB4B5F" author="Miller, J. S. &amp; L. D. Otero" pageId="174" pageNumber="849" pagination="338 - 372" refId="ref146848" refString="Miller, J. S., and L. D. Otero. 1994. Immature stages of Venezuelan Dioptinae (Notodontidae) in Josia and Thirmida. Journal of the Lepidopterists' Society 48: 338 - 372." type="journal article" year="1994">Miller and Otero, 1994</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BE936F6FF1B9EEFBCED1429FB844C51" blockId="174.[128,654,946,1759]" lastBlockId="174.[701,1228,212,674]" pageId="174" pageNumber="849">
The vesica of
<taxonomicName id="4C564D75FF1B9EEFBD311428FE2F4B78" baseAuthorityName="Druce" baseAuthorityYear="1900" box="[321,439,1620,1643]" class="Insecta" family="Notodontidae" genus="Scea" kingdom="Animalia" order="Lepidoptera" pageId="174" pageNumber="849" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B922EAE4FF1B9EEFBD311428FE2F4B78" box="[321,439,1620,1643]" italics="true" pageId="174" pageNumber="849">T. grandis</emphasis>
</taxonomicName>
is extremely long, with a set of robust, spinelike cornuti at its apex. The membrane of the vesica is packed tightly into the aedeagus at rest. The female dissection of
<taxonomicName id="4C564D75FF1B9EEFBD5414B4FE044BCC" baseAuthorityName="Druce" baseAuthorityYear="1900" box="[292,412,1736,1759]" class="Insecta" family="Notodontidae" genus="Scea" kingdom="Animalia" order="Lepidoptera" pageId="174" pageNumber="849" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B922EAE4FF1B9EEFBD5414B4FE044BCC" box="[292,412,1736,1759]" italics="true" pageId="174" pageNumber="849">S. grandis</emphasis>
</taxonomicName>
(JSM-1445) reveals that the large distal spikelike cornuti of the male (e.g., see fig. 351D) can sometimes break off and become lodged in the CB membrane during copulation.
</paragraph>
</subSubSection>
<subSubSection id="C34C657DFF1B9EEFBEAA1335FC9D4FB1" pageId="174" pageNumber="849" type="distribution">
<paragraph id="8BE936F6FF1B9EEFBEAA1335FC964C6F" blockId="174.[701,1228,212,674]" pageId="174" pageNumber="849">
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:
<taxonomicName id="4C564D75FF1B9EEFBFE31337FC924C6F" authority="(BMNH, CMNH, ZMH)" baseAuthorityName="BMNH, CMNH, ZMH" class="Insecta" family="Notodontidae" genus="Erbessa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="174" pageNumber="849" phylum="Arthropoda" rank="species" species="colombia">Colombia (BMNH, CMNH, ZMH)</taxonomicName>
.
</paragraph>
<paragraph id="8BE936F6FF1B9EEFBEAA13FEFC9D4FB1" blockId="174.[701,1228,212,674]" pageId="174" pageNumber="849">
<materialsCitation id="3B3E3CABFF1B9EEFBEAA13FEFB9E4F1C" collectingDate="1974-07-17" collectionCode="BMNH" collectorName="Hotel Solito &amp; C. Gibson &amp; Tilley Lamp, B. M." country="Colombia" elevation="2300" location="Sierra Nevada de Santa Marta" pageId="174" pageNumber="849" specimenCode="JSM-1444" specimenCount="1" stateProvince="Magdalena">
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: ³,
<collectingCountry id="F3417666FF1B9EEFBFC013F8FBBF4C88" box="[944,1063,389,411]" name="Colombia" pageId="174" pageNumber="849">Colombia</collectingCountry>
,
<collectingRegion id="4992F814FF1B9EEFB83113F8FB5F4C88" box="[1089,1223,389,411]" country="Colombia" name="Magdalena" pageId="174" pageNumber="849">Magdalena</collectingRegion>
,
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,
<quantity id="4CAE9B13FF1B9EEFB83713DFFB074CAB" box="[1095,1183,418,440]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="174" pageNumber="849" unit="m" value="2300.0">
<elevation id="007BD1C5FF1B9EEFB83713DFFB074CAB" box="[1095,1183,418,440]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="174" pageNumber="849" unit="m" value="2300.0">2300 m</elevation>
</quantity>
,
<date id="FFE81036FF1B9EEFB8C013DFFCB14CC7" pageId="174" pageNumber="849" value="1974-07-17">
<collectingDate id="EFACE9DEFF1B9EEFB8C013DFFCB14CC7" pageId="174" pageNumber="849" value="1974-07-17">17 Jul 1974</collectingDate>
</date>
,
<collectorName id="26A35320FF1B9EEFBF4C13C2FC4C4CC6" box="[828,980,447,469]" pageId="174" pageNumber="849">Hotel Solito</collectorName>
, leg.
<collectorName id="26A35320FF1B9EEFB86F13C2FB3A4CC6" box="[1055,1186,447,469]" pageId="174" pageNumber="849">C. Gibson</collectorName>
, at
<collectorName id="26A35320FF1B9EEFBECD13A1FC064CE1" box="[701,926,476,498]" pageId="174" pageNumber="849">Tilley Lamp, B.M.</collectorName>
1976-712.,
<collectionCode id="ED47AE33FF1B9EEFB85B13A0FB1D4CE1" box="[1067,1157,477,498]" pageId="174" pageNumber="849">BMNH</collectionCode>
(genitalia slide no.
<specimenCode id="DBF09E8DFF1B9EEFBFF41387FC674F1C" box="[900,1023,506,527]" collectionCode="JSM-" pageId="174" pageNumber="849">JSM-1444</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B3E3CABFF1B9EEFB850138AFC994FB1" collectingDate="1973-07-03" collectingDateMax="1973-09-02" collectingDateMin="1973-07-03" collectionCode="BMNH" collectorName="Oxford Expedn &amp; D.B. MacLean" country="Colombia" elevation="2300" latitude="10.902499" location="Sierra Nevada de Santa Marta" longLatPrecision="21" longitude="-74.0025" pageId="174" pageNumber="849" specimenCode="JSM-1445" specimenCount="1" specimenCount-female="1" stateProvince="Magdalena">
<specimenCount id="9D50FD7FFF1B9EEFB850138AFBAC4F03" box="[1056,1076,503,528]" count="1" pageId="174" pageNumber="849" type="female"></specimenCount>
,
<collectingCountry id="F3417666FF1B9EEFB8211387FB504F03" box="[1105,1224,506,528]" name="Colombia" pageId="174" pageNumber="849">Colombia</collectingCountry>
,
<collectingRegion id="4992F814FF1B9EEFBECD106AFCDB4F3E" box="[701,835,535,557]" country="Colombia" name="Magdalena" pageId="174" pageNumber="849">Magdalena</collectingRegion>
,
<location id="8E89602DFF1B9EEFBF19106AFC924F59" LSID="urn:lsid:plazi:treatment:03FF87E0FF1B9EEFBCD410B9FC9D4FB1:8E89602DFF1B9EEFBF19106AFC924F59" country="Colombia" latitude="10.902499" longLatPrecision="21" longitude="-74.0025" name="Sierra Nevada de Santa Marta" pageId="174" pageNumber="849" stateProvince="Magdalena">Sierra Nevada de Santa Marta</location>
,
<geoCoordinate id="EE625031FF1B9EEFBF6D1049FCE64F59" box="[797,894,564,586]" degrees="10" direction="north" minutes="54" orientation="latitude" pageId="174" pageNumber="849" precision="15" seconds="9" value="10.902499">
10
<emphasis id="B922EAE4FF1B9EEFBF481049FCDB4F5A" bold="true" box="[824,835,564,585]" pageId="174" pageNumber="849">°</emphasis>
54
<emphasis id="B922EAE4FF1B9EEFBF2E1048FCFF4F5A" bold="true" box="[862,871,565,585]" italics="true" pageId="174" pageNumber="849">9</emphasis>
N
</geoCoordinate>
<geoCoordinate id="EE625031FF1B9EEFBFFC1049FC6B4F59" box="[908,1011,564,586]" degrees="74" direction="west" minutes="00" orientation="longitude" pageId="174" pageNumber="849" precision="15" seconds="9" value="-74.0025">
74
<emphasis id="B922EAE4FF1B9EEFBFD71049FC2A4F5A" bold="true" box="[935,946,564,585]" pageId="174" pageNumber="849">°</emphasis>
00
<emphasis id="B922EAE4FF1B9EEFBFBC1048FC4D4F5A" bold="true" box="[972,981,565,585]" italics="true" pageId="174" pageNumber="849">9</emphasis>
W
</geoCoordinate>
,
<quantity id="4CAE9B13FF1B9EEFB8761049FBC74F59" box="[1030,1119,564,586]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="174" pageNumber="849" unit="m" value="2300.0">
<elevation id="007BD1C5FF1B9EEFB8761049FBC74F59" box="[1030,1119,564,586]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="174" pageNumber="849" unit="m" value="2300.0">2300 m</elevation>
</quantity>
,
<date id="FFE81036FF1B9EEFB8021049FCB04F74" pageId="174" pageNumber="849" value="1973-07-03" valueMax="1973-09-02" valueMin="1973-07-03">
<collectingDate id="EFACE9DEFF1B9EEFB8021049FCB04F74" pageId="174" pageNumber="849" value="1973-07-03" valueMax="1973-09-02" valueMin="1973-07-03">3 Jul2 Sep 1973</collectingDate>
</date>
,
<collectorName id="26A35320FF1B9EEFBF43102CFC704F74" box="[819,1000,593,615]" pageId="174" pageNumber="849">Oxford Expedn</collectorName>
to
<collectingCountry id="F3417666FF1B9EEFB87E102CFB1D4F74" box="[1038,1157,593,615]" name="Colombia" pageId="174" pageNumber="849">Colombia</collectingCountry>
,
<collectorName id="26A35320FF1B9EEFB8FF102FFB534F74" box="[1167,1227,594,615]" name="D.B. MacLean" pageId="174" pageNumber="849">B.M.</collectorName>
1973-500.,
<collectionCode id="ED47AE33FF1B9EEFBF301012FC024F97" box="[832,922,623,644]" pageId="174" pageNumber="849">BMNH</collectionCode>
(genitalia slide no.
<specimenCode id="DBF09E8DFF1B9EEFB8FC1012FD604FB1" collectionCode="JSM-" pageId="174" pageNumber="849">JSM-1445</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>