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<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="13132970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik &amp; Sierwald, Petra" docDate="2009" docId="1772122E7823FFE3FF013DC2AEF9EB7A" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Colossobolus pseudoaculeatus Wesener &amp; Enghoff &amp; Sierwald 2009, sp. n." docType="treatment" docUuid="2C06B20E-C62D-4CAE-9030-0761193A9DE9" docUuidSource="ZooBank" docVersion="11" lastPageNumber="38" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="37" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wesener, Thomas</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. &amp; Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. &amp; B 54 D 3623309</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Enghoff, Henrik</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sierwald, Petra</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-09-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>19</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>19</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>128</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791401" ID-GBIF-Taxon="163747652" ID-Zenodo-Dep="3791401" LSID="urn:lsid:zoobank.org:act:2C06B20E-C62D-4CAE-9030-0761193A9DE9" httpUri="http://treatment.plazi.org/id/1772122E7823FFE3FF013DC2AEF9EB7A" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
<subSubSection box="[140,675,694,721]" pageId="36" pageNumber="37" type="nomenclature">
<paragraph blockId="36.[140,979,694,756]" box="[140,675,694,721]" pageId="36" pageNumber="37">
<heading box="[140,675,694,721]" fontSize="11" level="2" pageId="36" pageNumber="37" reason="3">
<emphasis bold="true" box="[140,675,694,721]" pageId="36" pageNumber="37">
<taxonomicName authority="Wesener" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,598,694,721]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis bold="true" box="[140,490,694,721]" italics="true" pageId="36" pageNumber="37">Colossobolus pseudoaculeatus</emphasis>
Wesener
</taxonomicName>
,
<taxonomicNameLabel box="[610,675,695,721]" pageId="36" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[140,979,730,756]" pageId="36" pageNumber="37" type="description">
<paragraph blockId="36.[140,979,694,756]" box="[140,979,730,756]" pageId="36" pageNumber="37">
<uri box="[140,979,730,756]" pageId="36" pageNumber="37">
urn:lsid:zoobank.org:act:
<uuid box="[417,979,730,756]" pageId="36" pageNumber="37">2C06B20E-C62D-4CAE-9030-0761193A9DE9</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="materials_examined">
<paragraph blockId="36.[140,1108,800,1636]" pageId="36" pageNumber="37">
<emphasis bold="true" box="[140,371,800,826]" pageId="36" pageNumber="37">
<emphasis box="[140,241,800,826]" inLineHeading="true" pageId="36" pageNumber="37" reason="1">Material</emphasis>
examined:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608818273" collectingDate="2008-02-26" collectionCode="ZSM" collectorName="Kohler" country="Madagascar" location="Provate Reserve Le Fontenay" pageId="36" pageNumber="37" specimenCode="FGZC 1847" specimenCount="1" stateProvince="Antsiranana" typeStatus="holotype">
1 ♂
<emphasis box="[428,531,800,826]" italics="true" pageId="36" pageNumber="37">
<typeStatus box="[428,525,800,826]" pageId="36" pageNumber="37">Holotype</typeStatus>
:
</emphasis>
1 ♂ (
<quantity box="[597,695,800,827]" metricMagnitude="-1" metricUnit="m" metricValue="1.19" pageId="36" pageNumber="37" unit="mm" value="119.0">119 mm</quantity>
long),
<collectionCode box="[776,836,800,826]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="36" pageNumber="37" type="Museum">ZSM</collectionCode>
<specimenCode box="[843,987,800,826]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="36" pageNumber="37" type="Museum">FGZC 1847</specimenCode>
,
<collectingCountry name="Madagascar" pageId="36" pageNumber="37">Madagascar</collectingCountry>
, Province
<collectingRegion box="[296,435,835,861]" country="Madagascar" name="Antsiranana" pageId="36" pageNumber="37">Antsiranana</collectingRegion>
, Forêt dAmbre,
<location LSID="urn:lsid:plazi:treatment:1772122E7823FFE3FF013DC2AEF9EB7A:9A04F5E37823FFE2FDF73C30AC9EE90F" box="[634,967,836,862]" country="Madagascar" name="Provate Reserve Le Fontenay" pageId="36" pageNumber="37" stateProvince="Antsiranana">Provate Reserve Le Fontenay</location>
, leg.
<collectorName box="[1030,1107,835,861]" pageId="36" pageNumber="37">Köhler</collectorName>
and Franzen,
<date box="[294,419,871,897]" pageId="36" pageNumber="37" value="2008-02-26">
<collectingDate box="[294,419,871,897]" pageId="36" pageNumber="37" value="2008-02-26">26.II.2008</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="diagnosis">
<paragraph blockId="36.[140,1108,800,1636]" pageId="36" pageNumber="37">
<emphasis bold="true" box="[185,451,905,931]" pageId="36" pageNumber="37">
<emphasis box="[185,324,905,931]" inLineHeading="true" pageId="36" pageNumber="37" reason="1">Differential</emphasis>
diagnosis:
</emphasis>
the anterior and posterior gonopods of
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[904,1108,906,932]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[904,1108,906,932]" italics="true" pageId="36" pageNumber="37">C. pseudoaculeatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[140,203,942,968]" pageId="36" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
can only be confused with those of
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[625,759,941,968]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species">
<emphasis box="[625,759,941,968]" italics="true" pageId="36" pageNumber="37">C. aculeatus</emphasis>
</taxonomicName>
. See the latter for a list of the characters distinguishing these two species.
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="description">
<paragraph blockId="36.[140,1108,800,1636]" pageId="36" pageNumber="37">
<emphasis bold="true" box="[185,335,1011,1037]" pageId="36" pageNumber="37">
<emphasis box="[185,330,1011,1037]" inLineHeading="true" pageId="36" pageNumber="37" reason="1">Description</emphasis>
.
</emphasis>
<emphasis box="[345,497,1012,1037]" italics="true" pageId="36" pageNumber="37">Measurements</emphasis>
: male with 49 body rings, circa
<quantity box="[879,979,1012,1038]" metricMagnitude="-1" metricUnit="m" metricValue="1.19" pageId="36" pageNumber="37" unit="mm" value="119.0">119 mm</quantity>
long,
<quantity metricMagnitude="-3" metricUnit="m" metricValue="9.55" pageId="36" pageNumber="37" unit="mm" value="9.55">9.55 mm</quantity>
wide. Female unknown.
</paragraph>
<paragraph blockId="36.[140,1108,800,1636]" pageId="36" pageNumber="37">
<emphasis box="[185,300,1082,1108]" italics="true" pageId="36" pageNumber="37">Coloration</emphasis>
affected by alcohol. Head, antennae, legs and telson red (
<figureCitation box="[955,1094,1082,1108]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="36" pageNumber="37">Figs 21A, B</figureCitation>
). Median part of collum dark, margins red (
<figureCitation box="[608,710,1117,1143]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="36" pageNumber="37">Fig. 21A</figureCitation>
). Body rings anteriorly dark brown, posteriorly wine-red.
<emphasis box="[384,487,1153,1179]" italics="true" pageId="36" pageNumber="37">Antennae</emphasis>
protruding back to ring 4.
<emphasis box="[803,1018,1152,1178]" italics="true" pageId="36" pageNumber="37">Male coxal processes</emphasis>
present on coxae 3 and 4, on coxae 57 absent.
<emphasis box="[607,769,1187,1213]" italics="true" pageId="36" pageNumber="37">Preanal process</emphasis>
well-rounded, not projecting over anal valves (
<figureCitation box="[329,431,1223,1249]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="36" pageNumber="37">Fig. 21B</figureCitation>
).
</paragraph>
<paragraph blockId="36.[140,1108,800,1636]" pageId="36" pageNumber="37">
<emphasis box="[185,372,1258,1284]" italics="true" pageId="36" pageNumber="37">Anterior gonopod</emphasis>
sternite elongated into a wide, broadly rounded lobe (
<figureCitation box="[987,1094,1258,1284]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="36" pageNumber="37">Fig. 21D</figureCitation>
). Mesal process of coxite weakly developed, protruding into a very short wide lobe with a well-rounded tip; lobe only slightly longer than sternite (
<figureCitation box="[774,880,1328,1354]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="36" pageNumber="37">Fig. 21D</figureCitation>
). Telopodite process long and well-rounded, apically thicker than basally. Mesal margin laterally sharp, not projecting (
<figureCitation box="[270,374,1399,1425]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="36" pageNumber="37">Fig. 21C</figureCitation>
).
</paragraph>
<paragraph blockId="36.[140,1108,800,1636]" pageId="36" pageNumber="37">
<emphasis box="[185,388,1434,1460]" italics="true" pageId="36" pageNumber="37">Posterior gonopods</emphasis>
telopodites positioned parallel to one another, generally Ushaped (
<figureCitation box="[233,370,1469,1496]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="36" pageNumber="37">Figs 21E, F</figureCitation>
). Coxite branch absent (
<figureCitation box="[651,753,1469,1495]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="36" pageNumber="37">Fig. 21E</figureCitation>
), lateral branch basally slender, width apically increasing (
<figureCitation box="[432,566,1504,1531]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="36" pageNumber="37">Figs 21E, F</figureCitation>
); tip swollen, slightly curved mesally (
<figureCitation box="[993,1094,1504,1530]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="36" pageNumber="37">Fig. 21E</figureCitation>
). Central membranous area present as a wide, well-rounded knob (
<figureCitation box="[916,1021,1539,1566]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="36" pageNumber="37">Fig. 21E</figureCitation>
). Main branch slightly shorter than lateral branch, latero-apically towards lateral branch with a large membranous area (
<figureCitation box="[435,569,1610,1636]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="36" pageNumber="37">Figs 21E, F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="38" type="distribution">
<paragraph blockId="37.[140,1107,166,299]" pageId="37" pageNumber="38">
<emphasis bold="true" box="[185,499,166,193]" pageId="37" pageNumber="38">
<emphasis box="[185,336,166,192]" inLineHeading="true" pageId="37" pageNumber="38" reason="1">Distribution</emphasis>
and ecology
</emphasis>
:
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[520,730,167,193]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species">
<emphasis box="[520,730,167,193]" italics="true" pageId="37" pageNumber="38">C. pseudoaculeatus</emphasis>
</taxonomicName>
is only known from the Forêt dAmbre (
<figureCitation box="[254,337,202,228]" captionStart="Figure 10" captionStartId="18.[140,212,1654,1676]" captionTargetBox="[160,1106,1078,1634]" captionTargetId="figure@18.[140,1109,1071,1635]" captionTargetPageId="18" captionText="Figure 10. Distribution of the genus Colossobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3820794" httpUri="https://zenodo.org/record/3820794/files/figure.png" pageId="37" pageNumber="38">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="38" type="etymology">
<paragraph blockId="37.[140,1107,166,299]" pageId="37" pageNumber="38">
<emphasis bold="true" box="[185,323,237,263]" pageId="37" pageNumber="38">
<emphasis box="[185,318,237,263]" inLineHeading="true" pageId="37" pageNumber="38" reason="1">Etymology</emphasis>
:
</emphasis>
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[331,501,237,263]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species">
<emphasis box="[331,501,237,263]" italics="true" pageId="37" pageNumber="38">pseudoaculeatus</emphasis>
</taxonomicName>
, adjective, refers to the close similarity of the species to
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[170,408,272,298]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species">
<emphasis box="[170,408,272,298]" italics="true" pageId="37" pageNumber="38">Colossobolus aculeatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>