Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda) Author Wesener, Thomas urn:lsid:zoobank.org:author: Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309 Author Enghoff, Henrik urn:lsid:zoobank.org:author: Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark Author Sierwald, Petra urn:lsid:zoobank.org:author: Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. text ZooKeys 2009 2009-09-04 19 19 1 128 journal article 10.3897/zookeys.19.221 e6ea8eea-7156-4785-8313-f4bd88dadad1 1313–2970 576503 C473F9F6-1AE7-4B3F-B17F-CA1C2709010C Colossobolus pseudoaculeatus Wesener , sp. n. urn:lsid:zoobank.org:act: 2C06B20E-C62D-4CAE-9030-0761193A9DE9 Material examined: 1 ♂ Holotype : 1 ♂ ( 119 mm long), ZSM FGZC 1847 , Madagascar , Province Antsiranana , Forêt d’Ambre, Provate Reserve Le Fontenay , leg. Köhler and Franzen, 26.II.2008 . Differential diagnosis: the anterior and posterior gonopods of C. pseudoaculeatus sp. n. can only be confused with those of C. aculeatus . See the latter for a list of the characters distinguishing these two species. Description . Measurements : male with 49 body rings, circa 119 mm long, 9.55 mm wide. Female unknown. Coloration affected by alcohol. Head, antennae, legs and telson red ( Figs 21A, B ). Median part of collum dark, margins red ( Fig. 21A ). Body rings anteriorly dark brown, posteriorly wine-red. Antennae protruding back to ring 4. Male coxal processes present on coxae 3 and 4, on coxae 5–7 absent. Preanal process well-rounded, not projecting over anal valves ( Fig. 21B ). Anterior gonopod sternite elongated into a wide, broadly rounded lobe ( Fig. 21D ). Mesal process of coxite weakly developed, protruding into a very short wide lobe with a well-rounded tip; lobe only slightly longer than sternite ( Fig. 21D ). Telopodite process long and well-rounded, apically thicker than basally. Mesal margin laterally sharp, not projecting ( Fig. 21C ). Posterior gonopods telopodites positioned parallel to one another, generally Ushaped ( Figs 21E, F ). Coxite branch absent ( Fig. 21E ), lateral branch basally slender, width apically increasing ( Figs 21E, F ); tip swollen, slightly curved mesally ( Fig. 21E ). Central membranous area present as a wide, well-rounded knob ( Fig. 21E ). Main branch slightly shorter than lateral branch, latero-apically towards lateral branch with a large membranous area ( Figs 21E, F ). Distribution and ecology : C. pseudoaculeatus is only known from the Forêt d’Ambre ( Fig. 10 ). Etymology : pseudoaculeatus , adjective, refers to the close similarity of the species to Colossobolus aculeatus .