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<document id="E36FECACA868647F64EC6E38F433BBAB" ID-CLB-Dataset="28060" ID-DOI="10.11646/zootaxa.4506.1.1" ID-GBIF-Dataset="9fce96d4-8e89-4264-944a-4aef2a8046cb" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606718" ID-ZooBank="6F70EE34-FCD0-4426-958E-F734994225F3" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1553585246128" checkinUser="plazi" docAuthor="Qiu, Lu, Che, Yang-Li &amp; Wang, Zong-Qing" docDate="2018" docId="4A1B87C99E0B1958FF3CFCEBFD5BFED8" docLanguage="en" docName="zootaxa.4506.1.1.pdf" docOrigin="Zootaxa 4506 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Eupolyphaga everestiana subsp. reni Qiu &amp; Che &amp; Wang 2018, subsp. nov." docType="treatment" docUuid="95B9ACDD-AA3E-4669-9D51-E53B0D068873" docUuidSource="ZooBank" docVersion="12" lastPageNumber="27" masterDocId="B622FFB19E131942FFABF809FFB6FFCC" masterDocTitle="A taxonomic study of Eupolyphaga Chopard, 1929 (Blattodea: Corydiidae: Corydiinae)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="25" updateTime="1738679641918" updateUser="ExternalLinkService">
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<mods:title id="75843293303E6E106B3974359DF95F84">A taxonomic study of Eupolyphaga Chopard, 1929 (Blattodea: Corydiidae: Corydiinae)</mods:title>
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<mods:namePart id="5F3E95EE0323D34DAECBDBCFFF72B986">Qiu, Lu</mods:namePart>
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<mods:namePart id="8EE8EA8F6F15980368E4C5227CC124E8">Che, Yang-Li</mods:namePart>
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<mods:namePart id="93DDAC44869AB427462AD19F1CD584F5">Wang, Zong-Qing</mods:namePart>
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<mods:title id="F213042764559DB406980783B062BF8C">Zootaxa</mods:title>
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<mods:date id="00686663FF678B654C5E2D55B8E64ACC">2018</mods:date>
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<mods:number id="7B8B16BDFB85A89BC389A024F001EA34">2018-10-29</mods:number>
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<treatment id="4A1B87C99E0B1958FF3CFCEBFD5BFED8" ID-DOI="http://doi.org/10.5281/zenodo.5978907" ID-GBIF-Taxon="154495644" ID-Zenodo-Dep="5978907" LSID="urn:lsid:zoobank.org:act:95B9ACDD-AA3E-4669-9D51-E53B0D068873" httpUri="http://treatment.plazi.org/id/4A1B87C99E0B1958FF3CFCEBFD5BFED8" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" scope_family="Corydiidae" scope_order="Blattodea" scope_subFamily="Corydiinae">
<subSubSection id="8AA865549E0B195AFF3CFCEBFD27FB31" box="[151,657,1250,1277]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="C20D36DF9E0B195AFF3CFCEBFD27FB31" blockId="24.[151,657,1250,1310]" box="[151,657,1250,1277]" pageId="24" pageNumber="25">
<heading id="994581B39E0B195AFF3CFCEBFD27FB31" bold="true" box="[151,657,1250,1277]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<emphasis id="F0C6EACD9E0B195AFF3CFCEBFD27FB31" bold="true" box="[151,657,1250,1277]" pageId="24" pageNumber="25">
<taxonomicName id="05B24D5C9E0B195AFF3CFCEBFDB6FB30" authority="Qiu &amp; Che &amp; Wang, 2018" authorityName="Qiu &amp; Che &amp; Wang" authorityYear="2018" box="[151,512,1250,1276]" class="Insecta" family="Corydiidae" genus="Eupolyphaga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="24" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="everestiana" status="subsp. nov." subSpecies="reni">
<emphasis id="F0C6EACD9E0B195AFF3CFCEBFDB6FB30" bold="true" box="[151,512,1250,1276]" italics="true" pageId="24" pageNumber="25">Eupolyphaga everestiana reni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EBF557B69E0B195AFDACFCEAFD27FB31" box="[519,657,1251,1277]" pageId="24" pageNumber="25" rank="subSpecies">
<taxonomicName id="05B24D5C9E0B195AFDACFCEAFD3AFB31" box="[519,652,1251,1277]" pageId="24" pageNumber="25" rank="subSpecies" subSpecies="nov">subsp. nov</taxonomicName>
.
</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8AA865549E0B195AFF3CFD0CFE42FAD2" box="[151,500,1285,1310]" pageId="24" pageNumber="25" type="description">
<paragraph id="C20D36DF9E0B195AFF3CFD0CFE42FAD2" blockId="24.[151,657,1250,1310]" box="[151,500,1285,1310]" pageId="24" pageNumber="25">
(
<figureCitation id="5A892A5A9E0B195AFF34FD0CFF5AFAD1" box="[159,236,1285,1310]" captionStart="FIGURE 7" captionStartId="25.[151,250,1695,1717]" captionTargetBox="[178,1413,645,1669]" captionTargetId="figure@25.[151,1436,625,1674]" captionTargetPageId="25" captionText="FIGURE 7. Eupolyphaga species from Tibet, China: AC. Polyphaga thibetana Chopard, male, holotype (with labels), from Gyantse, Shigatse. DI. Polyphaga everestiana Chopard: DF. Male, syntype (with labels), from Mt. Everest; GI. Female, syntype (with labels), from Mt. Everest. AF photographed by Zong-Qing Wang, GI provided by George Beccaloni, copyright The Natural History Museum, United Kingdom (NHM). JM. Eupolyphaga everestiana reni subsp. nov., from Lhünzê, Lhoka: JK. Male, holotype; LM. Female, paratype. Scale bar 1 cm for all." httpUri="https://zenodo.org/record/2606734/files/figure.png" pageId="24" pageNumber="25">Figs. 7</figureCitation>
JM; 14 GI; 24 AD)
</paragraph>
</subSubSection>
<subSubSection id="8AA865549E0B195AFF3CFD58FC5AFA31" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="C20D36DF9E0B195AFF3CFD58FEB6FA15" blockId="24.[151,1437,1360,2037]" pageId="24" pageNumber="25">
<emphasis id="F0C6EACD9E0B195AFF3CFD58FDC8FAA6" bold="true" box="[151,638,1361,1386]" pageId="24" pageNumber="25">
Type material.
<materialsCitation id="72DA3C829E0B195AFEE0FD58FDC8FAA6" ID-GBIF-Occurrence="2028138476" box="[331,638,1361,1386]" collectingDate="2014-08-09" collectionCode="MHBU" collectorName="Guo-Dong Ren &amp; Long Bai &amp; Jun-Sheng Shan" country="China" county="Lhunze County" elevation="4190" latitude="28.400116" location="Ridang Town" longLatPrecision="1" longitude="92.15562" municipality="Shaqiong Village" pageId="24" pageNumber="25" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<typeStatus id="1D09887D9E0B195AFEE0FD58FE0AFAA6" box="[331,444,1361,1386]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="BAA5764F9E0B195AFE63FD58FD9CFAA6" box="[456,554,1361,1386]" name="China" pageId="24" pageNumber="25">CHINA</collectingCountry>
:
<collectingRegion id="0076F83D9E0B195AFD9DFD58FDCFFAA6" box="[566,633,1361,1386]" country="China" name="Xizang" pageId="24" pageNumber="25">Tibet</collectingRegion>
:
</materialsCitation>
</emphasis>
male (MHBU),
<collectingMunicipality id="2269ACA59E0B195AFC9CFD58FC4FFAA6" box="[823,1017,1361,1386]" pageId="24" pageNumber="25">Shaqiong Village</collectingMunicipality>
[沙琼村],
<location id="C76D60049E0B195AFB29FD58FAAFFAA5" LSID="urn:lsid:plazi:treatment:4A1B87C99E0B1958FF3CFCEBFD5BFED8:C76D60049E0B195AFB29FD58FAAFFAA5" box="[1154,1305,1361,1386]" country="China" county="Lhunze County" latitude="28.400116" longLatPrecision="1" longitude="92.15562" municipality="Shaqiong Village" name="Ridang Town" pageId="24" pageNumber="25" stateProvince="Xizang">Ridang Town</location>
[日当镇],
<collectingCounty id="2B6C4E539E0B195AFF3CFD71FEF3FA5D" box="[151,325,1400,1425]" pageId="24" pageNumber="25">Lhünzê County</collectingCounty>
[隆子县],
<location id="C76D60049E0B195AFE76FD71FDEBFA5D" LSID="urn:lsid:plazi:treatment:4A1B87C99E0B1958FF3CFCEBFD5BFED8:C76D60049E0B195AFE76FD71FDEBFA5D" box="[477,605,1400,1425]" country="China" county="Lhunze County" latitude="28.400116" longLatPrecision="1" longitude="92.15562" municipality="Shaqiong Village" name="Lhoka City" pageId="24" pageNumber="25" stateProvince="Xizang">Lhoka City</location>
,
<geoCoordinate id="A78650189E0B195AFDC2FD71FCB4FA5C" box="[617,770,1400,1425]" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="1" value="28.400116">28°24.007 N</geoCoordinate>
,
<geoCoordinate id="A78650189E0B195AFCA5FD71FC15FA5C" box="[782,931,1400,1425]" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="1" value="92.15562">92°09.337 E</geoCoordinate>
,
<quantity id="054A9B3A9E0B195AFC1BFD70FBB7FA5D" box="[944,1025,1401,1425]" metricMagnitude="3" metricUnit="m" metricValue="4.1899999999999995" pageId="24" pageNumber="25" unit="m" value="4190.0">
<elevation id="499FD1EC9E0B195AFC1BFD70FBB7FA5D" box="[944,1025,1401,1425]" metricMagnitude="3" metricUnit="m" metricValue="4.1899999999999995" pageId="24" pageNumber="25" unit="m" value="4190.0">4190m</elevation>
</quantity>
,
<date id="B60C101F9E0B195AFBA6FD70FB22FA5D" box="[1037,1172,1401,1425]" pageId="24" pageNumber="25" value="2014-08-09">
<collectingDate id="A648E9F79E0B195AFBA6FD70FB22FA5D" box="[1037,1172,1401,1425]" pageId="24" pageNumber="25" value="2014-08-09">9.VIII.2014</collectingDate>
</date>
,
<collectorName id="6F4753099E0B195AFB0BFD70FAF9FA5C" box="[1184,1359,1401,1425]" pageId="24" pageNumber="25">Guo-Dong Ren</collectorName>
, Xing-
<collectorName id="6F4753099E0B195AFF3CFD94FEB2FA79" box="[151,260,1436,1461]" pageId="24" pageNumber="25">Long Bai</collectorName>
&amp;
<collectorName id="6F4753099E0B195AFE9BFD95FE5FFA79" box="[304,489,1436,1461]" pageId="24" pageNumber="25">Jun-Sheng Shan</collectorName>
leg.
<materialsCitation id="72DA3C829E0B195AFD83FD95FF4BFA15" ID-GBIF-Occurrence="2028138473" collectingDate="2014-08-09" collectionCode="MHBU" collectorName="Guo-Dong Ren &amp; Long Bai &amp; Jun-Sheng Shan" country="China" county="Lhunze County" elevation="4190" latitude="28.400116" location="Ridang Town" longLatPrecision="1" longitude="92.15562" municipality="Shaqiong Village" pageId="24" pageNumber="25" specimenCount="9" specimenCount-female="5" specimenCount-male="4" stateProvince="Xizang" typeStatus="paratype">
<emphasis id="F0C6EACD9E0B195AFD83FD95FCD8FA79" bold="true" box="[552,878,1436,1461]" pageId="24" pageNumber="25">
<typeStatus id="1D09887D9E0B195AFD83FD95FD12FA79" box="[552,676,1436,1461]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="BAA5764F9E0B195AFD1FFD95FCA0FA79" box="[692,790,1436,1461]" name="China" pageId="24" pageNumber="25">CHINA</collectingCountry>
:
<collectingRegion id="0076F83D9E0B195AFC8DFD95FCDFFA79" box="[806,873,1436,1461]" country="China" name="Xizang" pageId="24" pageNumber="25">Tibet</collectingRegion>
:
</emphasis>
<specimenCount id="D4B4FD569E0B195AFCD2FD94FC60FA79" box="[889,982,1436,1461]" pageId="24" pageNumber="25" type="male">4 males</specimenCount>
,
<specimenCount id="D4B4FD569E0B195AFC4EFD95FBE3FA79" box="[997,1109,1436,1461]" pageId="24" pageNumber="25" type="female">5 females</specimenCount>
(
<collectionCode id="A4A3AE1A9E0B195AFBC2FD94FB74FA79" box="[1129,1218,1437,1461]" pageId="24" pageNumber="25">MHBU</collectionCode>
), same data as the holotype
</materialsCitation>
.
</paragraph>
<paragraph id="C20D36DF9E0B195AFF6CFDEDFC5AFA31" blockId="24.[151,1437,1360,2037]" box="[199,1004,1508,1533]" pageId="24" pageNumber="25">
<emphasis id="F0C6EACD9E0B195AFF6CFDEDFE4AFA31" bold="true" box="[199,508,1508,1533]" pageId="24" pageNumber="25">Other material examined.</emphasis>
4 nymphs (MHBU), same data as the
<typeStatus id="1D09887D9E0B195AFC01FDECFC5EFA31" box="[938,1000,1509,1533]" pageId="24" pageNumber="25">types</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="8AA865549E0B195AFF6CFE01FCC7F988" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="C20D36DF9E0B195AFF6CFE01FCC7F988" blockId="24.[151,1437,1360,2037]" pageId="24" pageNumber="25">
<emphasis id="F0C6EACD9E0B195AFF6CFE01FEF7F9ED" bold="true" box="[199,321,1544,1569]" pageId="24" pageNumber="25">Diagnosis.</emphasis>
<taxonomicName id="05B24D5C9E0B195AFEE1FE00FD96F9EC" authorityName="Qiu &amp; Che &amp; Wang" authorityYear="2018" box="[330,544,1545,1568]" class="Insecta" family="Corydiidae" genus="Eupolyphaga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="24" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="everestiana" subSpecies="reni">
<emphasis id="F0C6EACD9E0B195AFEE1FE00FD96F9EC" box="[330,544,1545,1568]" italics="true" pageId="24" pageNumber="25">E. everestiana reni</emphasis>
</taxonomicName>
<emphasis id="F0C6EACD9E0B195AFD81FE01FD19F9ED" bold="true" box="[554,687,1544,1569]" pageId="24" pageNumber="25">
<taxonomicName id="05B24D5C9E0B195AFD81FE01FD1CF9ED" box="[554,682,1544,1569]" pageId="24" pageNumber="25" rank="subSpecies" subSpecies="nov">subsp. nov</taxonomicName>
.
</emphasis>
is distinguished from the nominotypical subspecies by yellowish abdomen, more broad and shorter tegmina and shorter pronotum.
</paragraph>
</subSubSection>
<subSubSection id="8AA865549E0B1958FF6CFE59FE19FF1E" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" type="description">
<paragraph id="C20D36DF9E0B195BFF6CFE59FD9DFDF0" blockId="24.[151,1437,1360,2037]" lastBlockId="25.[151,1437,151,573]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<emphasis id="F0C6EACD9E0B195AFF6CFE59FD90F9A5" bold="true" box="[199,550,1616,1641]" pageId="24" pageNumber="25">Description. Male. General:</emphasis>
measurements (mm): body length: 18.018.5, overall length: 22.324.2, pronotum length×width: 4.75.2×8.39.3, tegmen length: 17.920.5. Size stout, yellowish brown, with brown maculae on tegmina (
<figureCitation id="5A892A5A9E0B195AFE39FE91FE61F97C" box="[402,471,1688,1713]" captionStart="FIGURE 7" captionStartId="25.[151,250,1695,1717]" captionTargetBox="[178,1413,645,1669]" captionTargetId="figure@25.[151,1436,625,1674]" captionTargetPageId="25" captionText="FIGURE 7. Eupolyphaga species from Tibet, China: AC. Polyphaga thibetana Chopard, male, holotype (with labels), from Gyantse, Shigatse. DI. Polyphaga everestiana Chopard: DF. Male, syntype (with labels), from Mt. Everest; GI. Female, syntype (with labels), from Mt. Everest. AF photographed by Zong-Qing Wang, GI provided by George Beccaloni, copyright The Natural History Museum, United Kingdom (NHM). JM. Eupolyphaga everestiana reni subsp. nov., from Lhünzê, Lhoka: JK. Male, holotype; LM. Female, paratype. Scale bar 1 cm for all." httpUri="https://zenodo.org/record/2606734/files/figure.png" pageId="24" pageNumber="25">Fig. 7</figureCitation>
JK).
<emphasis id="F0C6EACD9E0B195AFD82FE91FDC4F97D" bold="true" box="[553,626,1688,1713]" pageId="24" pageNumber="25">Head:</emphasis>
vertex and the space between ocelli dark brownish black. Interocular space wide, as wide as the distance between ocelli. Ocelli large, robust drop-shaped, ocelli ridge not very distinct, straight, anterior edge with many yellowish long pubescence. Frons raised, posterior completely yellowish, slightly pubescent. Antennae brownish yellow. Clypeus distinct; ante-clypeus widely occupies the lateral and hind margins of the clypeus, light brownish yellow; post-clypeus brown. Labrum wide, yellowish brown, pubescent, lateral corners of distal part round, median of apex emarginated. Maxillary palpi and labial palpi brownish (
<figureCitation id="5A892A5A9E0B195AFAA3FF45FAEFF8A9" box="[1288,1369,1868,1893]" captionStart="FIGURE 14" captionStartId="33.[151,250,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@33.[151,1436,193,1884]" captionText="FIGURE 14. Pronotum (A, D, G, J, M, P), vertex (B, E, H, K, N, Q) and face (C, F, I, L, O, R) features of Eupolyphaga species: AC. Eupolyphaga yunnanensis (Chopard); DF. Eupolyphaga thibetana (Chopard); GI. Eupolyphaga everestiana reni subsp. nov.; JL. Eupolyphaga pilosa sp. nov.; MO. Eupolyphaga xuorum sp. nov.; PR. Eupolyphaga hengduana (Woo &amp; Feng). Scale bar 1 mm for all." figureDoi="http://doi.org/10.5281/zenodo.14802514" httpUri="https://zenodo.org/record/14802514/files/figure.png" pageId="24" pageNumber="25">Fig. 14</figureCitation>
HI).
<emphasis id="F0C6EACD9E0B195AFF3CFF79FEAFF845" bold="true" box="[151,281,1904,1929]" pageId="24" pageNumber="25">Pronotum:</emphasis>
anterior widely whitish yellow marginated, disc and hind margin dark brown, lateral parts light brown, the edge between the whitish yellow and brown parts light orange. Surface generally with small yellowish pubescence, lateral fore margins and apex with many additional long yellowish setae. Pronotum widest on the hind border near the middle, apex slightly convex, straight, lateral fore margins oblique, slightly curved, lateral parts subangular, posterolateral angles truncated, hind margin slightly arched (
<figureCitation id="5A892A5A9E0A195BFC7CF89EFBFCFF7C" box="[983,1098,151,176]" captionStart="FIGURE 14" captionStartId="33.[151,250,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure@33.[151,1436,193,1884]" captionText="FIGURE 14. Pronotum (A, D, G, J, M, P), vertex (B, E, H, K, N, Q) and face (C, F, I, L, O, R) features of Eupolyphaga species: AC. Eupolyphaga yunnanensis (Chopard); DF. Eupolyphaga thibetana (Chopard); GI. Eupolyphaga everestiana reni subsp. nov.; JL. Eupolyphaga pilosa sp. nov.; MO. Eupolyphaga xuorum sp. nov.; PR. Eupolyphaga hengduana (Woo &amp; Feng). Scale bar 1 mm for all." figureDoi="http://doi.org/10.5281/zenodo.14802514" httpUri="https://zenodo.org/record/14802514/files/figure.png" pageId="25" pageNumber="26">Fig. 14 G</figureCitation>
).
<emphasis id="F0C6EACD9E0A195BFBCBF89EFAE5FF7C" bold="true" box="[1120,1363,151,176]" pageId="25" pageNumber="26">Tegmina and wings:</emphasis>
short, extended the end of abdomen about
<quantity id="054A9B3A9E0A195BFD9AF8B5FD0DFF18" box="[561,699,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="6.4" metricValueMin="3.4" pageId="25" pageNumber="26" unit="mm" value="4.9" valueMax="6.4" valueMin="3.4">3.46.4 mm</quantity>
. Tegmina short and broad, hyaline, slightly brownish, evenly and incompactly with irregular dark brown speckles. Wings hyaline, with distinct brown speckles at distal half.
<emphasis id="F0C6EACD9E0A195BFAF0F8D6FA2AFF34" bold="true" box="[1371,1436,223,248]" pageId="25" pageNumber="26">Legs:</emphasis>
light brownish yellow, pubescent, spines on the legs reddish brown, distal darker.
<emphasis id="F0C6EACD9E0A195BFB85F90DFB1AFED1" bold="true" box="[1070,1196,260,285]" pageId="25" pageNumber="26">Abdomen:</emphasis>
in ventral view, light yellow; each segment slightly brownish laterally and medially, with two brown dimples laterally except the last two; the last three segments much more brownish. Supra-anal plate pubescent, transverse; two median sclerites small to large; cerci short (
<figureCitation id="5A892A5A9E0A195BFE66F966FDF7FE44" box="[461,577,367,392]" captionStart="FIGURE 24" captionStartId="40.[151,250,867,889]" captionTargetBox="[357,1223,212,829]" captionTargetId="figure@40.[340,1247,193,845]" captionTargetPageId="40" captionText="FIGURE 24. Eupolyphaga everestiana reni subsp. nov., male: A. Supra-anal plate, ventral; B. Subgenital plate, ventral; C. Genitalia, dorsal; D. Right phallomere, right-ventral. Scale bar 1 mm." httpUri="https://zenodo.org/record/2606764/files/figure.png" pageId="25" pageNumber="26">Fig. 24 A</figureCitation>
). Subgenital plate asymmetrical, brown but with muddy yellow coloration, hind margin setose; styli similar, robust (
<figureCitation id="5A892A5A9E0A195BFDCCF99DFD63FE60" box="[615,725,404,429]" captionStart="FIGURE 24" captionStartId="40.[151,250,867,889]" captionTargetBox="[357,1223,212,829]" captionTargetId="figure@40.[340,1247,193,845]" captionTargetPageId="40" captionText="FIGURE 24. Eupolyphaga everestiana reni subsp. nov., male: A. Supra-anal plate, ventral; B. Subgenital plate, ventral; C. Genitalia, dorsal; D. Right phallomere, right-ventral. Scale bar 1 mm." httpUri="https://zenodo.org/record/2606764/files/figure.png" pageId="25" pageNumber="26">Fig. 24 B</figureCitation>
).
<emphasis id="F0C6EACD9E0A195BFD40F99DFCD5FE61" bold="true" box="[747,867,404,429]" pageId="25" pageNumber="26">Genitalia:</emphasis>
well sclerited.
<emphasis id="F0C6EACD9E0A195BFBB9F99DFB6AFE61" bold="true" box="[1042,1244,404,429]" pageId="25" pageNumber="26">Left phallomere:</emphasis>
L1 with anterior much protruded, round, left with a wide process, posterior with two angled lobes; L2 much curved, right end with two well sclerotized process, the distal one elongate; L3 straight, apex curved; pda well developed, paa bud shaped.
<emphasis id="F0C6EACD9E0A195BFF3CF9F6FEC6FDD4" bold="true" box="[151,368,511,536]" pageId="25" pageNumber="26">Right phallomere:</emphasis>
large. R1M stout, posterior hollow, transparent; R1L thin, slender; R3 much concaved; R2 well sclerotized, irregular (
<figureCitation id="5A892A5A9E0A195BFE25FA2DFE68FDF0" box="[398,478,548,573]" captionStart="FIGURE 24" captionStartId="40.[151,250,867,889]" captionTargetBox="[357,1223,212,829]" captionTargetId="figure@40.[340,1247,193,845]" captionTargetPageId="40" captionText="FIGURE 24. Eupolyphaga everestiana reni subsp. nov., male: A. Supra-anal plate, ventral; B. Subgenital plate, ventral; C. Genitalia, dorsal; D. Right phallomere, right-ventral. Scale bar 1 mm." httpUri="https://zenodo.org/record/2606764/files/figure.png" pageId="25" pageNumber="26">Fig. 24</figureCitation>
CD).
</paragraph>
<caption id="96CD66579E0A195BFF3CFE96FCFBF8FC" httpUri="https://zenodo.org/record/2606734/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,250,1695,1717]" targetBox="[178,1413,645,1669]" targetPageId="25">
<paragraph id="C20D36DF9E0A195BFF3CFE96FCFBF8FC" blockId="25.[151,1437,1695,1840]" pageId="25" pageNumber="26">
<emphasis id="F0C6EACD9E0A195BFF3CFE96FEA5F979" bold="true" box="[151,275,1695,1717]" pageId="25" pageNumber="26">FIGURE 7.</emphasis>
<taxonomicName id="05B24D5C9E0A195BFEB0FE96FE14F979" authorityName="Chopard" authorityYear="1929" box="[283,418,1695,1717]" class="Insecta" family="Corydiidae" genus="Eupolyphaga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="F0C6EACD9E0A195BFEB0FE96FE14F979" box="[283,418,1695,1717]" italics="true" pageId="25" pageNumber="26">Eupolyphaga</emphasis>
</taxonomicName>
species from Tibet, China:
<emphasis id="F0C6EACD9E0A195BFD14FE96FD41F979" bold="true" box="[703,759,1695,1717]" pageId="25" pageNumber="26">AC.</emphasis>
<taxonomicName id="05B24D5C9E0A195BFD56FE96FB82F979" authority="Chopard" authorityName="Chopard" authorityYear="1922" box="[765,1076,1695,1717]" class="Insecta" family="Corydiidae" genus="Polyphaga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="thibetana">
<emphasis id="F0C6EACD9E0A195BFD56FE96FC65F979" box="[765,979,1695,1717]" italics="true" pageId="25" pageNumber="26">Polyphaga thibetana</emphasis>
Chopard
</taxonomicName>
, male, holotype (with labels), from Gyantse, Shigatse. DI.
<taxonomicName id="05B24D5C9E0A195BFE3FFEB7FD54F918" authority="Chopard" authorityName="Chopard" authorityYear="1922" box="[404,738,1726,1748]" class="Insecta" family="Corydiidae" genus="Polyphaga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="everestiana">
<emphasis id="F0C6EACD9E0A195BFE3FFEB7FDC8F918" box="[404,638,1726,1748]" italics="true" pageId="25" pageNumber="26">Polyphaga everestiana</emphasis>
Chopard
</taxonomicName>
: DF. Male, syntype (with labels), from Mt. Everest; GI. Female, syntype (with labels), from Mt. Everest. AF photographed by Zong-Qing Wang, GI provided by George Beccaloni, copyright The Natural History Museum, United Kingdom (NHM). JM.
<taxonomicName id="05B24D5C9E0A195BFCA5FEF2FB8AF8DD" authority="Qiu &amp; Che &amp; Wang, 2018" authorityName="Qiu &amp; Che &amp; Wang" authorityYear="2018" box="[782,1084,1787,1809]" class="Insecta" family="Corydiidae" genus="Eupolyphaga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="25" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="everestiana" status="subsp. nov." subSpecies="reni">
<emphasis id="F0C6EACD9E0A195BFCA5FEF2FB8AF8DD" box="[782,1084,1787,1809]" italics="true" pageId="25" pageNumber="26">Eupolyphaga everestiana reni</emphasis>
</taxonomicName>
<emphasis id="F0C6EACD9E0A195BFBE9FEF5FB0EF8DD" bold="true" box="[1090,1208,1788,1809]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="EBF557B69E0A195BFBE9FEF5FB0EF8DD" box="[1090,1208,1788,1809]" pageId="25" pageNumber="26" rank="subSpecies">
<taxonomicName id="05B24D5C9E0A195BFBE9FEF5FB02F8DD" box="[1090,1204,1788,1809]" pageId="25" pageNumber="26" rank="subSpecies" subSpecies="nov">subsp. nov</taxonomicName>
.
</taxonomicNameLabel>
</emphasis>
, from Lhünzê, Lhoka: JK. Male, holotype; LM. Female, paratype. Scale bar 1 cm for all.
</paragraph>
</caption>
<paragraph id="C20D36DF9E0A195BFF6CFF55FD7FF815" blockId="25.[151,1437,1884,2010]" pageId="25" pageNumber="26">
<emphasis id="F0C6EACD9E0A195BFF6CFF55FE93F8B9" bold="true" box="[199,293,1884,1909]" pageId="25" pageNumber="26">Female.</emphasis>
Measurements (mm): body length: 20.121.1, body width: 11.813.3. Unicolored, reddish brown to deep brown depending on specimen, pubescent; in dorsal view, segments from 3th to 5th of abdomen each with two dark spots laterally; supra-anal plate protruded
<typeStatus id="1D09887D9E0A195BFD75FFA9FCA7F874" box="[734,785,1952,1976]" pageId="25" pageNumber="26">type</typeStatus>
, median with a distinct longitudinal line which extending into the other segments of abdomen (
<figureCitation id="5A892A5A9E0A195BFD9CFFC8FDCFF815" box="[567,633,1985,2010]" captionStart="FIGURE 7" captionStartId="25.[151,250,1695,1717]" captionTargetBox="[178,1413,645,1669]" captionTargetId="figure@25.[151,1436,625,1674]" captionTargetPageId="25" captionText="FIGURE 7. Eupolyphaga species from Tibet, China: AC. Polyphaga thibetana Chopard, male, holotype (with labels), from Gyantse, Shigatse. DI. Polyphaga everestiana Chopard: DF. Male, syntype (with labels), from Mt. Everest; GI. Female, syntype (with labels), from Mt. Everest. AF photographed by Zong-Qing Wang, GI provided by George Beccaloni, copyright The Natural History Museum, United Kingdom (NHM). JM. Eupolyphaga everestiana reni subsp. nov., from Lhünzê, Lhoka: JK. Male, holotype; LM. Female, paratype. Scale bar 1 cm for all." httpUri="https://zenodo.org/record/2606734/files/figure.png" pageId="25" pageNumber="26">Fig. 7</figureCitation>
LM).
</paragraph>
<paragraph id="C20D36DF9E091958FF6CF89EFCB3FF7C" blockId="26.[199,1425,151,277]" box="[199,773,151,176]" pageId="26" pageNumber="27">
<emphasis id="F0C6EACD9E091958FF6CF89EFE90FF7C" bold="true" box="[199,294,151,176]" pageId="26" pageNumber="27">Nymph.</emphasis>
Similar to the female, but with light color.
</paragraph>
<paragraph id="C20D36DF9E091958FF6CF8B0FE19FF1E" blockId="26.[199,1425,151,277]" box="[199,431,185,210]" pageId="26" pageNumber="27">
<emphasis id="F0C6EACD9E091958FF6CF8B0FE87FF1E" bold="true" box="[199,305,185,210]" pageId="26" pageNumber="27">Ootheca.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="8AA865549E091958FF6CF8D3FA27FF3F" box="[199,1425,218,243]" pageId="26" pageNumber="27" type="etymology">
<paragraph id="C20D36DF9E091958FF6CF8D3FA27FF3F" blockId="26.[199,1425,151,277]" box="[199,1425,218,243]" pageId="26" pageNumber="27">
<emphasis id="F0C6EACD9E091958FF6CF8D3FEF8FF3F" bold="true" box="[199,334,218,243]" pageId="26" pageNumber="27">Etymology.</emphasis>
This subspecies is named in honour of Prof. Guo-Dong Ren (MHBU), one of the
<typeStatus id="1D09887D9E091958FB4EF8D2FAA3FF3F" box="[1253,1301,219,243]" pageId="26" pageNumber="27">type</typeStatus>
collectors.
</paragraph>
</subSubSection>
<subSubSection id="8AA865549E091958FF6CF8F5FD5BFED8" box="[199,749,252,277]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="C20D36DF9E091958FF6CF8F5FD5BFED8" blockId="26.[199,1425,151,277]" box="[199,749,252,277]" pageId="26" pageNumber="27">
<emphasis id="F0C6EACD9E091958FF6CF8F5FED6FED9" bold="true" box="[199,352,252,277]" pageId="26" pageNumber="27">Distribution.</emphasis>
<collectingCountry id="BAA5764F9E091958FECCF8F5FE1CFED9" box="[359,426,252,277]" name="China" pageId="26" pageNumber="27">China</collectingCountry>
(
<collectingRegion id="0076F83D9E091958FE12F8F5FE4FFED9" box="[441,505,252,277]" country="China" name="Xizang" pageId="26" pageNumber="27">Tibet</collectingRegion>
: Lhoka City) (
<figureCitation id="5A892A5A9E091958FD30F8F5FD56FED8" box="[667,736,252,277]" captionStart="FIGURE 2" captionStartId="6.[151,250,954,976]" captionTargetBox="[340,1247,193,933]" captionTargetId="figure@6.[340,1247,193,933]" captionTargetPageId="6" captionText="FIGURE 2. Distribution map of Eupolyphaga species in China (except the species only found in Yunnan. Larger circles indicate the type localities)." figureDoi="http://doi.org/10.5281/zenodo.2606724" httpUri="https://zenodo.org/record/2606724/files/figure.png" pageId="26" pageNumber="27">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>