176 lines
27 KiB
XML
176 lines
27 KiB
XML
<document ID-CLB-Dataset="6710" ID-DOI="http://dx.doi.org/10.3897/dez.63.10241" ID-GBIF-Dataset="0a6965a7-b124-4871-95d4-9e35c49073de" ID-Pensoft-Pub="1860-1324-2-211" ID-Pensoft-UUID="51243BBBE9F158F58BBC9BF3EA964017" ID-Zenodo-Dep="167458" ID-ZooBank="45552C4EC6AE4F9499980C2D492333B3" ModsDocID="1860-1324-63-2-211" checkinTime="1621372567443" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2016" docId="B7A28488AE3B3393CE79E53DE5E95048" docLanguage="en" docName="DeutEntomolZeit 63(2): 211-270" docOrigin="Deutsche Entomologische Zeitschrift 63 (2)" docPubDate="2016-11-18" docSource="http://dx.doi.org/10.3897/dez.63.10241" docTitle="Cyphocoleus heterogenus Chaudoir 1877" docType="treatment" docVersion="5" id="51243BBBE9F158F58BBC9BF3EA964017" lastPageNumber="229" masterDocId="51243BBBE9F158F58BBC9BF3EA964017" masterDocTitle="Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia" masterLastPageNumber="270" masterPageNumber="211" pageNumber="229" updateTime="1691866435949" updateUser="ExternalLinkService">
|
||
<mods:mods id="3EE16F94681BD0775610611A96D402B5" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="EEBCD983FEC14203F7A0232501D45F91">
|
||
<mods:title id="B107DF86B74C2B59A3A8C5DF61F5D01F">Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="C41F9BFC8D79081D0502052DFD67FCAF" type="personal">
|
||
<mods:role id="7A52E1E6D15F394F496BBBAD0E224C2A">
|
||
<mods:roleTerm id="55D7B7F822B112116694EA6B3E1FFB64">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2E1F7F5B0DBEF1CA92B202D42713AD4C">Liebherr, James K.</mods:namePart>
|
||
<mods:nameIdentifier id="011E2B4B4C84E12CC4E8B93A72A87F9E" type="ORCID">https://orcid.org/0000-0001-9831-884X</mods:nameIdentifier>
|
||
<mods:affiliation id="F093A303ED6B95DF831D5E752B3AC1E6">Cornell University Insect Collection, John H. and Anna B. Comstock Hall, Cornell University, Ithaca, NY 14853 - 2601, USA</mods:affiliation>
|
||
<mods:nameIdentifier id="308863A28856B6D29C1F3F71E8AF870C" type="email">jkl5@cornell.edu</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="5980BC0D3A35B971AE2DD1C6C7EB6BBE">text</mods:typeOfResource>
|
||
<mods:relatedItem id="A61388EF5646B651833F849E86E5165F" type="host">
|
||
<mods:titleInfo id="20E72B4D9BA021F75DAAE58CB27D2759">
|
||
<mods:title id="AD51906A44CF6CE148EE690F8E778AFC">Deutsche Entomologische Zeitschrift</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="220385DA76812BEC63E08EE94358603D">
|
||
<mods:date id="629B410B69A090C0E26F4A91E8F6F20C">2016</mods:date>
|
||
<mods:detail id="81EDF5A92C4EA7E036B1905837FC930F" type="pubDate">
|
||
<mods:number id="7C64928EFCA3087B6A06A80207BA85F1">2016-11-18</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="06D741380F22603030786E0540F97E9F" type="volume">
|
||
<mods:number id="439C0FF50A3F8B51A9A2FC9A99B4B31E">63</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="ECD4432E9F77B85D13C64E5740E0E4D0" type="issue">
|
||
<mods:number id="A820269471C96BC1EEE14C009CB54556">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="C53CDB0BE892170D2EE75FC6611D9C68" unit="page">
|
||
<mods:start id="9E478A9EA5275D3B256618208F2B9E4B">211</mods:start>
|
||
<mods:end id="2D9B9FD710A00BADBC4B9CF561D5F4C3">270</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="32821A39DEB145F67485B9C7C74D16A5">
|
||
<mods:url id="6B3CDDD3718F60B25CB6080DC9A9AC6A">http://dx.doi.org/10.3897/dez.63.10241</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="000BE80EC2386DE1DD4054FE34C1EA69">journal article</mods:classification>
|
||
<mods:identifier id="3846A836CF948B8DC84BB65EDA75FB8F" type="DOI">http://dx.doi.org/10.3897/dez.63.10241</mods:identifier>
|
||
<mods:identifier id="F7AA56689E4F9BC6959EE23011B351C8" type="Pensoft-Pub">1860-1324-2-211</mods:identifier>
|
||
<mods:identifier id="4F6607E182672B2D136745457002736A" type="ZooBank">45552C4EC6AE4F9499980C2D492333B3</mods:identifier>
|
||
<mods:identifier id="A3936BCEC2BDF1891E4B05114860A74F" type="Pensoft-UUID">51243BBBE9F158F58BBC9BF3EA964017</mods:identifier>
|
||
<mods:identifier id="831B9026A634863B245491502DAEB37D" type="Zenodo-Dep">167458</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="B7A28488AE3B3393CE79E53DE5E95048" ID-GBIF-Taxon="182223714" LSID="urn:lsid:plazi:treatment:B7A28488AE3B3393CE79E53DE5E95048" httpUri="http://treatment.plazi.org/id/B7A28488AE3B3393CE79E53DE5E95048" lastPageNumber="229" pageId="18" pageNumber="229">
|
||
<subSubSection id="AFD202E809D2D91146CBE7AC80E99751" pageId="18" pageNumber="229" type="nomenclature">
|
||
<paragraph id="D2C0A1A920D5BFC1DAE09342500235AF" pageId="18" pageNumber="229">
|
||
3.
|
||
<taxonomicName id="50D954180AF63808BBC0B9E3FDD7CC65" ID-CoL="6C8YZ" authority="Chaudoir, 1877" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus heterogenus" order="Coleoptera" pageId="18" pageNumber="229" phylum="Arthropoda" rank="species" species="heterogenus">Cyphocoleus heterogenus Chaudoir, 1877</taxonomicName>
|
||
<figureCitation id="7F69390DA156A3823E2136D6407841F8" captionStart="Figures 28–33" captionStartId="F5" captionText="Figures 28 - 33. 28 - 32 Male aedeagus with associated abdominal segments VIII and IX, dorsal view, Cyphocoleus spp.: 28, Cyphocoleus heterogenus; 29, Cyphocoleus angustatus; 30, Cyphocoleus mirabilis; 31, Cyphocoleus cordatus; 32, Cyphocoleus subulatus. 33. Abdominal lateroventrites VIII (left) and laterotergites VIII (right), showing membranous division of laterotergites, male Cyphocoleus latipennis. Dorsal gland reservoir is globose without dorsal lobe. For abbreviations see Table 1." figureDoi="10.3897/dez.63.10241.figures28-33" httpUri="https://binary.pensoft.net/fig/111541" pageId="18" pageNumber="229">Figures 28</figureCitation>
|
||
<figureCitation id="67F348BE6CAB58D5EF566B41F358839D" captionStart="Figures 47–50" captionStartId="F10" captionText="Figures 47 - 50. Dorsal habitus photographs of Cyphocoleus spp.: 47, Cyphocoleus lissus female, Mt. Humboldt; 48, Cyphocoleus prolixus male, Mt. Humboldt; 49, Cyphocoleus heterogenus male, Yahoue; 50, Cyphocoleus parovicollis male, Pic d'Amoa." figureDoi="10.3897/dez.63.10241.figures47-50" httpUri="https://binary.pensoft.net/fig/111547" pageId="18" pageNumber="229">, 49</figureCitation>
|
||
<figureCitation id="82DAAA561B5D069998E88D425D37AE19" captionStart="Figures 51–59" captionStartId="F11" captionText="Figures 51 - 59. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 51, Cyphocoleus lissus, Mt. Humboldt; 52, Cyphocoleus prolixus, Mt. Humboldt; 53, Cyphocoleus heterogenus, Yahoue; 54, Cyphocoleus heterogenus, internal sac everted, La Foa; 55, Cyphocoleus parovicollis, Me Maoya; 56, Cyphocoleus parovicollis, internal sac everted, Me Maoya; 57, Cyphocoleus ovicollis, Ningua Res.; 58 - 59, Cyphocoleus ovicollis, internal sac everted, Mt. Do; 58, laevoventral view; 59, dextrodorsal view. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures51-59" httpUri="https://binary.pensoft.net/fig/111548" pageId="18" pageNumber="229">, 53-54</figureCitation>
|
||
<figureCitation id="E3575FFB63220B85EAC11A6F721AAD1E" captionStart="Figures 60–65" captionStartId="F12" captionText="Figures 60 - 65. Female reproductive tract, ventral view, for Cyphocoleus spp.; 60, Cyphocoleus lissus, Mt. Humboldt; 61, Cyphocoleus prolixus, Mt. Humboldt; 62, Cyphocoleus heterogenus, Yahoue; 63, Cyphocoleus parovicollis, Mandjelia; 64, Cyphocoleus monteithi, Mt. Mou; 65, Cyphocoleus miricollis, Foret Nord. For abbreviations see Table 1." figureDoi="10.3897/dez.63.10241.figures60-65" httpUri="https://binary.pensoft.net/fig/111549" pageId="18" pageNumber="229">, 62</figureCitation>
|
||
<figureCitation id="25FE4C052A64A85906BD37B7CA51C6EB" captionStart="Figures 66–75" captionStartId="F13" captionText="Figures 66 - 75. Female right gonocoxae, ventral view, for Cyphocoleus spp. illustrating apical fringe setae of basal gonocoxite 1, and two or three lateral ensiform setae, one dorsal ensiform seta, and two apical nematiform setae on apical gonocoxite 2; 66, Cyphocoleus lissus, Mt. Humboldt; 67, Cyphocoleus prolixus, Mt. Humboldt; 68, Cyphocoleus heterogenus, Yahoue; 69, Cyphocoleus parovicollis, Mandjelia; 70, Cyphocoleus ovicollis, Ningua Res.; 71, Cyphocoleus burwelli, Mt. Humboldt; 72, Cyphocoleus monteithi, Mt. Mou; 73, Cyphocoleus miricollis, Foret Nord; 74, Cyphocoleus mirabilis, Mt. Koghi; 75, Cyphocoleus lescheni, apical gonocoxite damaged, Aoupinie. Scale bar, 0.10 mm." figureDoi="10.3897/dez.63.10241.figures66-75" httpUri="https://binary.pensoft.net/fig/111550" pageId="18" pageNumber="229">, 68</figureCitation>
|
||
<figureCitation id="2E0A82EF0616DD6B51EF8184FEAF1D2A" captionStart="Figures 76–81" captionStartId="F14" captionText="Figures 76 - 81. Distribution maps of Cyphocoleus spp.: 76, Cyphocoleus lissus; 77, Cyphocoleus heterogenus, Cyphocoleus prolixus; 78, Cyphocoleus parovicollis, Cyphocoleus ovicollis; 79, Cyphocoleus burwelli, Cyphocoleus angustatus, Cyphocoleus monteithi; 80, Cyphocoleus miricollis, Cyphocoleus fasciatus; 81, Cyphocoleus mirabilis, Cyphocoleus moorei, Cyphocoleus lescheni." figureDoi="10.3897/dez.63.10241.figures76-81" httpUri="https://binary.pensoft.net/fig/111551" pageId="18" pageNumber="229">, 77</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9D573B81AD677A744CC3F0F124D0C2B0" pageId="18" pageNumber="229" type="diagnosis">
|
||
<paragraph id="9166BBB9A0298B34F18153437242605E" pageId="18" pageNumber="229">Diagnosis.</paragraph>
|
||
<paragraph id="4C36285052E444CBB9078C0C47ADB097" pageId="18" pageNumber="229">
|
||
Beetles as large as
|
||
<taxonomicName id="3FB7416EF27A5CEF25C27EEC655A6FE2" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus prolixus" order="Diptera" pageId="18" pageNumber="229" phylum="Arthropoda" rank="species" species="prolixus">
|
||
<emphasis id="D3CE4BC9D1D978E0CDDABDA62775CE11" italics="true" pageId="18" pageNumber="229">Cyphocoleus prolixus</emphasis>
|
||
</taxonomicName>
|
||
, standardized body length 10.8-13.1 mm, but head and prothorax much broader, the head much more developed at the expense of pronotal length (Figs
|
||
<figureCitation id="B24EF7B22171E3D0F5A9BF53335EC5EE" captionStart="Figures 47–50" captionStartId="F10" captionText="Figures 47 - 50. Dorsal habitus photographs of Cyphocoleus spp.: 47, Cyphocoleus lissus female, Mt. Humboldt; 48, Cyphocoleus prolixus male, Mt. Humboldt; 49, Cyphocoleus heterogenus male, Yahoue; 50, Cyphocoleus parovicollis male, Pic d'Amoa." figureDoi="10.3897/dez.63.10241.figures47-50" httpUri="https://binary.pensoft.net/fig/111547" pageId="18" pageNumber="229">48</figureCitation>
|
||
,
|
||
<figureCitation id="DE8C863A0FDFA6EDCA5ECF2277903BDC" captionStart="Figures 47–50" captionStartId="F10" captionText="Figures 47 - 50. Dorsal habitus photographs of Cyphocoleus spp.: 47, Cyphocoleus lissus female, Mt. Humboldt; 48, Cyphocoleus prolixus male, Mt. Humboldt; 49, Cyphocoleus heterogenus male, Yahoue; 50, Cyphocoleus parovicollis male, Pic d'Amoa." figureDoi="10.3897/dez.63.10241.figures47-50" httpUri="https://binary.pensoft.net/fig/111547" pageId="18" pageNumber="229">49</figureCitation>
|
||
).Though the mandibles are shorter than in
|
||
<taxonomicName id="312F8205093080C9D5C4B647CC083DBE" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus prolixus" order="Diptera" pageId="18" pageNumber="229" phylum="Arthropoda" rank="species" species="prolixus">
|
||
<emphasis id="D89749E28E02F80345CD1E070BE5E51A" italics="true" pageId="18" pageNumber="229">Cyphocoleus prolixus</emphasis>
|
||
</taxonomicName>
|
||
, with mandibular length 2.0
|
||
<normalizedToken id="8EE5D6F0ED60E3CFD87E83E7D7BBAF2C" originalValue="×">x</normalizedToken>
|
||
distance from antennal socket to anterolateral margin of labrum, the antennae are longer, with scape length 4.1
|
||
<normalizedToken id="8537CF9CFEB918DC3D65F46B4964EC41" originalValue="×">x</normalizedToken>
|
||
maximal breadth. The pronotum is unique among
|
||
<taxonomicName id="BFF2D4F33484D68547CF4CB289B6F0BA" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus" order="Diptera" pageId="18" pageNumber="229" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9539ABDF4C29683ABF1CC221044BEA9B" italics="true" pageId="18" pageNumber="229">Cyphocoleus</emphasis>
|
||
</taxonomicName>
|
||
due to the extremely elongate, acutely protruded front angles. The presence of three dorsal elytral setae is shared only with
|
||
<taxonomicName id="D47133AE07E140C62E1AF7A063CD554D" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus ovicollis" order="Diptera" pageId="18" pageNumber="229" phylum="Arthropoda" rank="species" species="ovicollis">
|
||
<emphasis id="4B94A9AE728D3B1FEF6CFEAF412E8BB8" italics="true" pageId="18" pageNumber="229">Cyphocoleus ovicollis</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="D7F5A2FF40828303C89D8419E9EF537E" captionStart="Figures 82–85" captionStartId="F15" captionText="Figures 82 - 85. Dorsal habitus photographs of Cyphocoleus spp.: 82, Cyphocoleus ovicollis female, Ningua Res.; 83, Cyphocoleus burwelli female, Mt. Canala; 84, Cyphocoleus angustatus male, Mt. Panie; 85, Cyphocoleus monteithi male, Dzumac Rd. jct." figureDoi="10.3897/dez.63.10241.figures82-85" httpUri="https://binary.pensoft.net/fig/111552" pageId="18" pageNumber="229">82</figureCitation>
|
||
), all other species exhibiting more reduced numbers of dorsal elytral setae or none at all.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D0E191859FA14616ED7F8EEDB7451089" pageId="18" pageNumber="229" type="description">
|
||
<paragraph id="0493B47E62AB689C832578B51FB4E303" pageId="18" pageNumber="229">Description</paragraph>
|
||
<paragraph id="7D7324928C9B3383B61C7B062D3B18D1" pageId="18" pageNumber="229">
|
||
(n = 5). Head capsule broad, robust, distance from clypeal-labral suture subequal to width across eyes; frons broadly convex, with small chevron-shaped indentation medially between front margins of eyes; eyes moderately large, convex, 28 ommatidia across horizontal diameter; eyes moderately small, convex, ocular ratio 1.35-1.42; only posterior supraorbital seta present; neck distinctly impressed just before juncture with prothorax; gena glabrous, subgenal seta absent. Pronotum distinctly cordate, margin sinuately constricted basally; lateral marginal bead present from acute front angles to just mesad tightly rounded hind angles; median base margin by fine bead; proepisternum slightly bulging, visible in dorsal view along middle half of prothorax; median longitudinal impression shallow, finely incised; anterior transverse impression broad, shallow, obliquely meeting median impression; anterior callosity slightly convex; prosternal process with fine median ridge anteriorly between procoxae, medially depressed on ventral and posterior face. Elytra broadly ovoid, disc moderately convex with side only moderately depressed relative to disc; elytral striae impunctate; parascutellar seta present, articulatory socket not upraised above elytral surface; elytra narrow basally, humeri obtuse-angulate at base of fifth stria; lateral margin straight to slightly concave lateral anterior series of lateral elytral setae; lateral elytral setae arranged as 6 + 1 + 7-8; subapical and apical elytral setae present; subapical sinuation distinctly invaginated, curvature greatest just mesad obtusely rounded juncture of sinuation and lateral margin. Mesepisternum impunctate; metepisternal dorsal length 0.86
|
||
<normalizedToken id="49FE5D703E0BAAA02C9DE28851B8263C" originalValue="×">x</normalizedToken>
|
||
diagonal width. Abdomen of males with apical margin of apical ventrite distinctly notched, the invagination smoothly rounded; apical ventrite of male with one seta each side, of female with two setae each side. Legs elongate, gracile, mt1 length/tibial length = 0.26; metacoxae bisetose; tarsomeres with short sparse setae dorsally; mt4 outer lobe 1.7
|
||
<normalizedToken id="0ECB5D9DE8D12D8A70F3D7745C01BBEC" originalValue="×">x</normalizedToken>
|
||
median length, 4-5 ventrolateral setae each side. Microsculpture of frons and vertex a stretched isodiametric mesh; pronotal disc and discal elytral intervals covered with fine transverse lines; dorsal surface of body apparently glabrous except for standard macrosetae; pro-, meso-, and metasternum and visible abdominal ventrites with sparsely distributed pelage of very short setae. Coloration of dorsum rufopiceous with cyanotic to cupreous iridescence due to microsculpture; antennae dark rufous, paler apically, palps brunneous; basal abdominal ventrites rufopiceous, ventrite 6 rufobrunneous, margin narrowly brunneous; femora rufopiceous, tibiae rufobrunneous, tarsi slightly paler, dark rufous.
|
||
</paragraph>
|
||
<paragraph id="730DBA925AA0B801AF0DCF3ACDD23B47" pageId="18" pageNumber="229">
|
||
<emphasis id="58C70A970802BF1344063B53F6ED5CDB" italics="true" pageId="18" pageNumber="229">Male genitalia</emphasis>
|
||
(n = 8). Male aedeagal median lobe robust, broadly parallel sided in basal 3/4 of length, then evenly narrowed to projected, narrow tip, dorsoventral breadth at midlength 0.25
|
||
<normalizedToken id="C2D1736C114FA2891A8F467FF9707D36" originalValue="×">x</normalizedToken>
|
||
distance from tip to base of closed basal bulb (Fig.
|
||
<figureCitation id="D40D8B9432F4ED1F56A4626CCF40B7DB" captionStart="Figures 51–59" captionStartId="F11" captionText="Figures 51 - 59. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 51, Cyphocoleus lissus, Mt. Humboldt; 52, Cyphocoleus prolixus, Mt. Humboldt; 53, Cyphocoleus heterogenus, Yahoue; 54, Cyphocoleus heterogenus, internal sac everted, La Foa; 55, Cyphocoleus parovicollis, Me Maoya; 56, Cyphocoleus parovicollis, internal sac everted, Me Maoya; 57, Cyphocoleus ovicollis, Ningua Res.; 58 - 59, Cyphocoleus ovicollis, internal sac everted, Mt. Do; 58, laevoventral view; 59, dextrodorsal view. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures51-59" httpUri="https://binary.pensoft.net/fig/111548" pageId="18" pageNumber="229">53</figureCitation>
|
||
); lobe shaft slightly melanized, the internal sac visible through lobe wall in uneverted position; lobe apex tapered to tightly rounded tip, length distad ostial opening 2.5
|
||
<normalizedToken id="316DEDB19B900780C4FEBE14880FD0D0" originalValue="×">x</normalizedToken>
|
||
dorsoventral breadth; internal sac evenly covered with short melanized spicules in distal half, of sinuous shape but sides parallel in extension (Fig.
|
||
<figureCitation id="6AE54E7B09711CCD8D432279F80E78E7" captionStart="Figures 51–59" captionStartId="F11" captionText="Figures 51 - 59. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 51, Cyphocoleus lissus, Mt. Humboldt; 52, Cyphocoleus prolixus, Mt. Humboldt; 53, Cyphocoleus heterogenus, Yahoue; 54, Cyphocoleus heterogenus, internal sac everted, La Foa; 55, Cyphocoleus parovicollis, Me Maoya; 56, Cyphocoleus parovicollis, internal sac everted, Me Maoya; 57, Cyphocoleus ovicollis, Ningua Res.; 58 - 59, Cyphocoleus ovicollis, internal sac everted, Mt. Do; 58, laevoventral view; 59, dextrodorsal view. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures51-59" httpUri="https://binary.pensoft.net/fig/111548" pageId="18" pageNumber="229">54</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="28233F314BD57D81056E143419BE6D28" pageId="18" pageNumber="229">
|
||
<emphasis id="26AD28E0A0C472FF563E3568E989917C" italics="true" pageId="18" pageNumber="229">Female reproductive tract</emphasis>
|
||
(n = 2). Bursa copulatrix vase-shaped, basally stalked, distance from base of gonocoxites to spermathecal basal sclerite 2.4
|
||
<normalizedToken id="DF0DFC282B78C658BC118C64D022C322" originalValue="×">x</normalizedToken>
|
||
maximum breadth (dissection compressed under cover slip) (Fig.
|
||
<figureCitation id="F1769E6DAC838D1DB95BE0F5F86D2D14" captionStart="Figures 60–65" captionStartId="F12" captionText="Figures 60 - 65. Female reproductive tract, ventral view, for Cyphocoleus spp.; 60, Cyphocoleus lissus, Mt. Humboldt; 61, Cyphocoleus prolixus, Mt. Humboldt; 62, Cyphocoleus heterogenus, Yahoue; 63, Cyphocoleus parovicollis, Mandjelia; 64, Cyphocoleus monteithi, Mt. Mou; 65, Cyphocoleus miricollis, Foret Nord. For abbreviations see Table 1." figureDoi="10.3897/dez.63.10241.figures60-65" httpUri="https://binary.pensoft.net/fig/111549" pageId="18" pageNumber="229">62</figureCitation>
|
||
); bursal walls thin, translucent, with broad band of spicules lining lumen near midlength; basal gonocoxite with apical fringe of seven to eight setae (Fig.
|
||
<figureCitation id="EA357720DBFB2F9879CA7B4FBDB8373C" captionStart="Figures 66–75" captionStartId="F13" captionText="Figures 66 - 75. Female right gonocoxae, ventral view, for Cyphocoleus spp. illustrating apical fringe setae of basal gonocoxite 1, and two or three lateral ensiform setae, one dorsal ensiform seta, and two apical nematiform setae on apical gonocoxite 2; 66, Cyphocoleus lissus, Mt. Humboldt; 67, Cyphocoleus prolixus, Mt. Humboldt; 68, Cyphocoleus heterogenus, Yahoue; 69, Cyphocoleus parovicollis, Mandjelia; 70, Cyphocoleus ovicollis, Ningua Res.; 71, Cyphocoleus burwelli, Mt. Humboldt; 72, Cyphocoleus monteithi, Mt. Mou; 73, Cyphocoleus miricollis, Foret Nord; 74, Cyphocoleus mirabilis, Mt. Koghi; 75, Cyphocoleus lescheni, apical gonocoxite damaged, Aoupinie. Scale bar, 0.10 mm." figureDoi="10.3897/dez.63.10241.figures66-75" httpUri="https://binary.pensoft.net/fig/111550" pageId="18" pageNumber="229">68</figureCitation>
|
||
); apical gonocoxite broadly rounded at apex with three lateral ensiform setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="74FAC6CAD9E0AB6ED7CAA54C79E34542" pageId="18" pageNumber="229" type="materials_examined">
|
||
<paragraph id="7B1A17ED32C99C4B695E5A74C08322C8" pageId="18" pageNumber="229">
|
||
<typeStatus id="4E9A993B48F6A93CF6B4F368412257A5">Type</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="F42835B0C33555EC9F481B47B29283DE" pageId="18" pageNumber="229">
|
||
<typeStatus id="BA6233B4AFC366C13E2FE3DF95C0A4A7">Holotype</typeStatus>
|
||
male (MNHN): Ex
|
||
<normalizedToken id="BD82DB074D481B0239EB589AB7A6023A" originalValue="Musæo">Musaeo</normalizedToken>
|
||
/ Chaudoir (red-inked white label with red border) //
|
||
<typeStatus id="9E569C3FC8CF79AB7D2AC3043CED1D12">HOLOTYPE</typeStatus>
|
||
/
|
||
<taxonomicName id="083FB0FE960C3CCFCF19D412B49B5003" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus" order="Diptera" pageId="18" pageNumber="229" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="87DF02B328BF0E907E9B27063457F5BD" italics="true" pageId="18" pageNumber="229">Cyphocoleus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="0EC79E854F511F38B3609D4CA9C0B696" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="heterogenus" order="Diptera" pageId="18" pageNumber="229" phylum="Arthropoda" rank="species" species="heterogenus">
|
||
<emphasis id="8803C1D98A37872B7D8133165E03E2B0" italics="true" pageId="18" pageNumber="229">heterogenus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<bibRefCitation id="B8D5261606FEDCD6125FF9A02E55C872" author="Chaudoir, M de" journalOrPublisher="Bulletin de la Societe Imperiale des naturalistes de Moscou" pageId="48" pageNumber="259" pagination="1 - 61" refId="B12" refString="Chaudoir, M de, 1877. Genres nouveaux et especes inedites de la famille des Carabiques. Bulletin de la Societe Imperiale des naturalistes de Moscou 52: 1 - 61" title="Genres nouveaux et especes inedites de la famille des Carabiques." volume="52" year="1877">Chaudoir 1877</bibRefCitation>
|
||
(black-bordered red label).
|
||
<typeStatus id="96C40818FF9A4BA22DBAFA67DAE86874">Holotype</typeStatus>
|
||
status is based on
|
||
<normalizedToken id="054E5A45106F7A1830D0A4D1BDD69021" originalValue="Chaudoir’s">Chaudoir's</normalizedToken>
|
||
(1887: 193) statement "Cet insecte
|
||
<normalizedToken id="4052800C777E88C9D745B3AD817BB79C" originalValue="m’a">m'a</normalizedToken>
|
||
<normalizedToken id="162381DE8F24C9EFFAD2BFE8ED40F4E6" originalValue="été">ete</normalizedToken>
|
||
<normalizedToken id="D62EFD7F7CBC38E5B5D7D46914DD2A87" originalValue="cédé">cede</normalizedToken>
|
||
par M.
|
||
<normalizedToken id="D6C4D33AA3B529F594CE758B390FCA34" originalValue="Sallé.”">Salle."</normalizedToken>
|
||
<normalizedToken id="5C8CE1104118D4C2AD3DA830458336BF" originalValue="Yahoué">Yahoue</normalizedToken>
|
||
hereby designated type locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0D5D61464249F0DCB742E18714723AFF" pageId="18" pageNumber="229" type="distribution">
|
||
<paragraph id="746DBE52BECDD14481D9F3EF5E2BFF28" pageId="18" pageNumber="229">Distribution and habitat.</paragraph>
|
||
<paragraph id="8F1B06697B54902F6D7B1F66264B18EF" pageId="18" pageNumber="229">
|
||
Found throughout Grande Terre, from Mt. Ignambi and Mt.
|
||
<normalizedToken id="23BEB84AFC7244B246B82666C5F97E17" originalValue="Panié">Panie</normalizedToken>
|
||
on the north, to
|
||
<normalizedToken id="92E98012EA0F43BC4D2A130B83779076" originalValue="Forêt">Foret</normalizedToken>
|
||
de Thi on the south (Fig.
|
||
<figureCitation id="B6707D3E14C898969D525694D8805793" captionStart="Figures 76–81" captionStartId="F14" captionText="Figures 76 - 81. Distribution maps of Cyphocoleus spp.: 76, Cyphocoleus lissus; 77, Cyphocoleus heterogenus, Cyphocoleus prolixus; 78, Cyphocoleus parovicollis, Cyphocoleus ovicollis; 79, Cyphocoleus burwelli, Cyphocoleus angustatus, Cyphocoleus monteithi; 80, Cyphocoleus miricollis, Cyphocoleus fasciatus; 81, Cyphocoleus mirabilis, Cyphocoleus moorei, Cyphocoleus lescheni." figureDoi="10.3897/dez.63.10241.figures76-81" httpUri="https://binary.pensoft.net/fig/111551" pageId="18" pageNumber="229">77</figureCitation>
|
||
, Suppl. material 3). Beetles have been collected from pyrethrin spray samples from trees and logs, with and without epiphytes, from under a rock near a dead tree, from
|
||
<taxonomicName id="C61F868A335E9AF5F42AE3C3D7CC0007" class="Monocotyledoneae" family="Pandanaceae" genus="Freycinetia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Freycinetia" order="Pandanales" pageId="18" pageNumber="229" phylum="Angiospermae" rank="genus">
|
||
<emphasis id="19F5333CB101667DFC00B0CFF24F93FE" italics="true" pageId="18" pageNumber="229">Freycinetia</emphasis>
|
||
</taxonomicName>
|
||
axils, and by hand during night.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |