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<document id="D00C5D50120E85A15F25C577941286A9" ID-DOI="10.17109/AZH.69.3.265.2023http://zoobank.org/9C2F726C-ECC7-43C4-8DA9-68683652C62D" ID-ISSN="2064-2474" ID-Zenodo-Dep="10272211" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701821093587" checkinUser="felipe" docAuthor="Tang, Jin, Zhang, Gongaote, Guo, Junqi, Luo, Lingxuan, Jiang, Jiamei &amp; Pan, Hongbo" docDate="2023" docId="03B86709FF917904FEACC20EFEBA6788" docLanguage="en" docName="ActaZoolAcadSciH.69.3.265-301.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 69 (3)" docSource="http://dx.doi.org/10.3389/fmicb.2023.1259653" docStyle="DocumentStyle:183DC955716D8EEDE76F04077FE39880.4:ActaZoolAcadSciH.2015-.journal_article" docStyleId="183DC955716D8EEDE76F04077FE39880" docStyleName="ActaZoolAcadSciH.2015-.journal_article" docStyleVersion="4" docTitle="Dichagyris forficula subsp. akdagestana Tang &amp; Zhang &amp; Guo &amp; Luo &amp; Jiang &amp; Pan 2023, subsp. n." docType="treatment" docVersion="2" lastPageNumber="277" masterDocId="FF811F71FF9B790BFFE5C767FFD96608" masterDocTitle="Contributions To The Taxonomy And Biogeography Of The Genus Dichagyris (Subg. Dichagyris) Lederer, 1867 (Lepidoptera, Noctuidae, Noctuinae) Ii. The Review Of The D. Forficula Species Group" masterLastPageNumber="301" masterPageNumber="265" pageNumber="275" updateTime="1701821906243" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="1521A5766740E848A847A256C89A352F">Contributions To The Taxonomy And Biogeography Of The Genus Dichagyris (Subg. Dichagyris) Lederer, 1867 (Lepidoptera, Noctuidae, Noctuinae) Ii. The Review Of The D. Forficula Species Group</mods:title>
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<emphasis id="B9650A0DFF917901FEACC20EFCDD638B" bold="true" box="[329,772,1385,1411]" pageId="10" pageNumber="275">Dichagyris forficula akdagestana</emphasis>
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http://zoobank.org/
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(
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,
<figureCitation id="132ACA9AFF917901FD66C2D7FD7A63C2" box="[643,675,1456,1482]" captionStart="Figs 4143" captionStartId="11.[174,219,1495,1517]" captionTargetBox="[183,1092,192,1463]" captionTargetId="figure-18@11.[174,1095,189,1469]" captionTargetPageId="11" captionText="Figs 4143. Male genitalia ofDichagyris forficula sspp. 41 =D. forficula (Eversmann, 1851), NE Turkey, Prov. Van, slide No.: VZ10245m. 42 = D. forficula akdagestana ssp. n., paratype, Russia, North Caucasus, Ingoushetia, slide No.: VZ9771m. 43 = D. forficula hadjina (Staudinger, 1892), Turkey, lectotype of Agrotis hadjina var.zeituna Staudinger, 1899, slide No.: VZ9548m" figureDoi="http://doi.org/10.5281/zenodo.10272225" httpUri="https://zenodo.org/record/10272225/files/figure.png" pageId="10" pageNumber="275">42</figureCitation>
,
<figureCitation id="132ACA9AFF917901FD4AC2D7FD1763C2" box="[687,718,1456,1482]" captionStart="Figs 6870" captionStartId="29.[174,219,1467,1489]" captionTargetBox="[193,1112,504,1429]" captionTargetId="figure-79@29.[174,1117,500,1436]" captionTargetPageId="29" captionText="Figs 6870. Genitalia of Dichagyris spp. 68 = Male genitalia of D. erubescens (Staudinger, 1892), Turkey, Prov. Malatya, slide No.: VZ11515m. 69 = Female genitalia of D. forficula akdagestana ssp. n., paratype, Russia, North Caucasus, Ingoushetia, slide No.: VZ6753f. 70 = female genitalia of D. forficula devota (Christoph, 1884), Turkmenistan, Kopet-Dagh, slide No.: VZ11483f" figureDoi="http://doi.org/10.5281/zenodo.10272251" httpUri="https://zenodo.org/record/10272251/files/figure.png" pageId="10" pageNumber="275">69</figureCitation>
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: male, “
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, N Caucasus,
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, leg.
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. Zurilina”, gen. slide GYP
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(coll.
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, with same data as holotype
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;
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, from the same locality,
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, leg.
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;
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, from the same locality
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21.
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. and 2.
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.2021
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, leg.
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.
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;
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,
<specimenCount id="9D171D96FF977907FD8BC79FFD386706" box="[622,737,248,270]" count="10" pageId="12" pageNumber="277" type="female">10 females</specimenCount>
, from the same locality,
<geoCoordinate id="EE25B0D8FF977907FC02C79FFBE76705" box="[999,1086,248,269]" degrees="42" direction="north" minutes="32" orientation="latitude" pageId="12" pageNumber="277" precision="925" value="42.533333">42°32N</geoCoordinate>
,
<geoCoordinate id="EE25B0D8FF977907FF6CC673FF036722" box="[137,218,276,298]" degrees="47" direction="east" minutes="25" orientation="longitude" pageId="12" pageNumber="277" precision="925" value="47.416668">47°25E</geoCoordinate>
,
<quantity id="4CE97BFAFF977907FF03C672FEEC6722" box="[230,309,277,298]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="12" pageNumber="277" unit="m" value="1030.0">
<elevation id="003C312CFF977907FF03C672FEEC6722" box="[230,309,277,298]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="12" pageNumber="277" unit="m" value="1030.0">1030 m</elevation>
</quantity>
,
<date id="FFAFF0DFFF977907FEA4C672FE696722" box="[321,432,277,298]" pageId="12" pageNumber="277" value="2022-06-27">
<collectingDate id="EFEB0937FF977907FEA4C672FE696722" box="[321,432,277,298]" pageId="12" pageNumber="277" value="2022-06-27">
27.
<collectionCode id="ED004EDAFF977907FEB8C672FEA16722" box="[349,376,277,298]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="12" pageNumber="277">VI</collectionCode>
.2022
</collectingDate>
</date>
, leg.
<collectionCode id="ED004EDAFF977907FE0CC672FE256722" box="[489,508,277,298]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="277" type="Museum">V</collectionCode>
.
<location id="8ECE80C4FF977907FDE2C672FDB96723" LSID="urn:lsid:plazi:treatment:03B86709FF917904FEACC20EFEBA6788:8ECE80C4FF977907FDE2C672FDB96723" box="[519,608,277,299]" country="Russia" latitude="42.533333" longLatPrecision="1149" longitude="47.416668" name="Zurilina" pageId="12" pageNumber="277" stateProvince="Daghestan">Zurilina</location>
</materialsCitation>
;
<materialsCitation id="3B79DC42FF977907FD89C672FD6E674F" collectingDate="2022-07-02" collectingDateMax="2022-07-08" collectingDateMin="2022-07-02" collectionCode="V, PG" collectorName="Zurilina" country="Russia" elevation="1030" latitude="42.533333" location="Zurilina" longLatPrecision="1149" longitude="47.416668" pageId="12" pageNumber="277" specimenCount="17" specimenCount-female="11" specimenCount-male="6" stateProvince="Daghestan" typeStatus="paratype">
<specimenCount id="9D171D96FF977907FD89C672FD196723" box="[620,704,277,299]" count="6" pageId="12" pageNumber="277" type="male">6 males</specimenCount>
,
<specimenCount id="9D171D96FF977907FD2EC672FCE66723" box="[715,831,277,299]" count="11" pageId="12" pageNumber="277" type="female">11 females</specimenCount>
, from the same locality,
<date id="FFAFF0DFFF977907FF6FC655FEBC674F" box="[138,357,305,327]" pageId="12" pageNumber="277" value="2022-07-02" valueMax="2022-07-08" valueMin="2022-07-02">
<collectingDate id="EFEB0937FF977907FF6FC655FEBC674F" box="[138,357,305,327]" pageId="12" pageNumber="277" value="2022-07-02" valueMax="2022-07-08" valueMin="2022-07-02">2.VII. and 8.VII.2022</collectingDate>
</date>
, leg.
<collectionCode id="ED004EDAFF977907FE79C655FE76674F" box="[412,431,306,327]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="277" type="Museum">V</collectionCode>
.
<collectorName id="26E4B3C9FF977907FE5DC656FDD6674F" box="[440,527,305,327]" pageId="12" pageNumber="277">Zurilina</collectorName>
(all in coll.
<collectionCode id="ED004EDAFF977907FD69C655FD69674F" box="[652,688,306,327]" pageId="12" pageNumber="277">PG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B79DC42FF977907FD27C655FCAD676C" collectingDate="2022-07-02" collectingDateMax="2022-07-08" collectingDateMin="2022-07-02" collectionCode="VZ" collectorName="Z. Varga &amp; Slides" country="Russia" elevation="11518" latitude="42.533333" location="Ingoushetia" longLatPrecision="1149" longitude="47.416668" pageId="12" pageNumber="277" specimenCount="1" specimenCount-male="1" stateProvince="Daghestan" typeStatus="paratype">
<specimenCount id="9D171D96FF977907FD27C655FCD3674F" box="[706,778,305,327]" count="1" pageId="12" pageNumber="277" type="male">1 male</specimenCount>
, with same data (coll.
<collectorName id="26E4B3C9FF977907FC1CC655FF7D676B" pageId="12" pageNumber="277">Z. Varga</collectorName>
).
<collectorName id="26E4B3C9FF977907FF5DC629FF23676C" box="[184,250,334,356]" pageId="12" pageNumber="277">Slides</collectorName>
: GYP5586f, GYP5799f,
<collectionCode id="ED004EDAFF977907FE17C629FDCD676C" box="[498,532,334,356]" pageId="12" pageNumber="277">VZ</collectionCode>
<quantity id="4CE97BFAFF977907FDF0C629FDBE676C" box="[533,615,334,356]" metricMagnitude="4" metricUnit="m" metricValue="1.1518" pageId="12" pageNumber="277" unit="m" value="11518.0">
<elevation id="003C312CFF977907FDF0C629FDBE676C" box="[533,615,334,356]" metricMagnitude="4" metricUnit="m" metricValue="1.1518" pageId="12" pageNumber="277" unit="m" value="11518.0">11518m</elevation>
</quantity>
,
<collectionCode id="ED004EDAFF977907FD96C629FD4F676B" box="[627,662,334,355]" pageId="12" pageNumber="277">VZ</collectionCode>
<quantity id="4CE97BFAFF977907FD73C629FD31676B" box="[662,744,334,355]" metricMagnitude="4" metricUnit="m" metricValue="1.1972999999999998" pageId="12" pageNumber="277" unit="m" value="11973.0">
<elevation id="003C312CFF977907FD73C629FD31676B" box="[662,744,334,355]" metricMagnitude="4" metricUnit="m" metricValue="1.1972999999999998" pageId="12" pageNumber="277" unit="m" value="11973.0">11973m</elevation>
</quantity>
.
<location id="8ECE80C4FF977907FD11C629FCAD676C" LSID="urn:lsid:plazi:treatment:03B86709FF917904FEACC20EFEBA6788:8ECE80C4FF977907FD11C629FCAD676C" box="[756,884,334,356]" country="Russia" name="Ingoushetia" pageId="12" pageNumber="277" stateProvince="Daghestan">Ingoushetia</location>
</materialsCitation>
.
<materialsCitation id="3B79DC42FF977907FC9AC629FF706796" collectorName="Herczig &amp; Uherkovich &amp; Horvath &amp; Szollat &amp; Sarkozi" country="Russia" elevation="1000" pageId="12" pageNumber="277" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Daghestan" typeStatus="paratype">
<specimenCount id="9D171D96FF977907FC9AC629FC10676C" box="[895,969,334,356]" count="1" pageId="12" pageNumber="277" type="male">1 male</specimenCount>
,
<specimenCount id="9D171D96FF977907FC30C629FBE4676C" box="[981,1085,334,356]" count="2" pageId="12" pageNumber="277" type="female">2 females</specimenCount>
, “ChIASSR Furtoug
<quantity id="4CE97BFAFF977907FEBDC60CFE7B6788" box="[344,418,363,384]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="277" unit="m" value="1000.0">
<elevation id="003C312CFF977907FEBDC60CFE7B6788" box="[344,418,363,384]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="277" unit="m" value="1000.0">1000 m</elevation>
</quantity>
12. Juli, 1989 leg.
<collectorName id="26E4B3C9FF977907FDB9C60CFD6B6789" box="[604,690,363,385]" pageId="12" pageNumber="277">Herczig</collectorName>
,
<collectorName id="26E4B3C9FF977907FD5EC60CFCE36789" box="[699,826,363,385]" pageId="12" pageNumber="277">Uherkovich</collectorName>
,
<collectorName id="26E4B3C9FF977907FCA1C60CFC796789" box="[836,928,363,385]" pageId="12" pageNumber="277">Horváth</collectorName>
,
<collectorName id="26E4B3C9FF977907FC4CC60CFC2A6789" box="[937,1011,363,385]" pageId="12" pageNumber="277">Szollát</collectorName>
,
<collectorName id="26E4B3C9FF977907FC19C60CFF476796" pageId="12" pageNumber="277">Sárközi</collectorName>
</materialsCitation>
;
<specimenCount id="9D171D96FF977907FF56C6EFFF226796" box="[179,251,392,414]" count="1" pageId="12" pageNumber="277" type="male">1 male</specimenCount>
, from the same site,
<date id="FFAFF0DFFF977907FE36C6EFFDBB6795" box="[467,610,392,413]" pageId="12" pageNumber="277" value="1990-06-12" valueMax="1990-06-13" valueMin="1990-06-12">
12-13.
<collectionCode id="ED004EDAFF977907FDEBC6EFFDF26795" box="[526,555,392,413]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="12" pageNumber="277">VI</collectionCode>
.1990
</date>
, leg. L. Ábrahám &amp; Á. Uherkovich;
<materialsCitation id="3B79DC42FF977907FC3AC6EFFBE467B2" collectingDate="1991-07-02" collectionCode="V" collectorName="B. Herczig &amp; Z. Meszaros" country="Russia" elevation="1000" location="Marko" pageId="12" pageNumber="277" specimenCount="26" specimenCount-female="13" specimenCount-male="13" stateProvince="Daghestan" typeStatus="paratype">
<specimenCount id="9D171D96FF977907FC3AC6EFFBE46796" box="[991,1085,392,414]" count="13" pageId="12" pageNumber="277" type="male">13 males</specimenCount>
,
<specimenCount id="9D171D96FF977907FF6DC6C2FF2267B3" box="[136,251,421,443]" count="13" pageId="12" pageNumber="277" type="female">13 females</specimenCount>
, with the same locality, “
<date id="FFAFF0DFFF977907FDE2C6C2FDAC67B2" box="[519,629,421,442]" pageId="12" pageNumber="277" value="1991-07-02">
<collectingDate id="EFEB0937FF977907FDE2C6C2FDAC67B2" box="[519,629,421,442]" pageId="12" pageNumber="277" value="1991-07-02">2.VII.1991</collectingDate>
</date>
, leg.
<collectorName id="26E4B3C9FF977907FD49C6C2FCC467B3" box="[684,797,421,443]" pageId="12" pageNumber="277">B. Herczig</collectorName>
,
<collectionCode id="ED004EDAFF977907FCCDC6C2FCE267B2" box="[808,827,421,442]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="277" type="Museum">V</collectionCode>
.
<location id="8ECE80C4FF977907FCA0C6C2FC5567B3" LSID="urn:lsid:plazi:treatment:03B86709FF917904FEACC20EFEBA6788:8ECE80C4FF977907FCA0C6C2FC5567B3" box="[837,908,421,443]" country="Russia" name="Marko" pageId="12" pageNumber="277" stateProvince="Daghestan">Markó</location>
et
<collectorName id="26E4B3C9FF977907FC48C6C2FBEA67B2" box="[941,1075,421,442]" pageId="12" pageNumber="277">Z. Mészáros</collectorName>
</materialsCitation>
;
<materialsCitation id="3B79DC42FF977907FF6CC6A5FEBD6419" collectingDate="1991-07-08" collectionCode="V, RG, VZ" collectorName="B. Herczig &amp; Slide Nos" country="Russia" elevation="1300" pageId="12" pageNumber="277" specimenCode="VZ6753, VZ11473, VZ11479, VZ11519, VZ11523" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Daghestan" typeStatus="paratype">
<specimenCount id="9D171D96FF977907FF6CC6A5FF0A67DF" box="[137,211,449,471]" count="1" pageId="12" pageNumber="277" type="male">1 male</specimenCount>
,
<specimenCount id="9D171D96FF977907FF3AC6A5FEE567DF" box="[223,316,449,471]" count="1" pageId="12" pageNumber="277" type="female">1 female</specimenCount>
, “NE Caucasus, Lezsgi
<quantity id="4CE97BFAFF977907FDA5C6A5FD5667DF" box="[576,655,450,471]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="277" unit="m" value="1300.0">
<elevation id="003C312CFF977907FDA5C6A5FD5667DF" box="[576,655,450,471]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="277" unit="m" value="1300.0">1300 m</elevation>
</quantity>
,
<date id="FFAFF0DFFF977907FD79C6A5FCD167DF" box="[668,776,450,471]" pageId="12" pageNumber="277" value="1991-07-08">
<collectingDate id="EFEB0937FF977907FD79C6A5FCD167DF" box="[668,776,450,471]" pageId="12" pageNumber="277" value="1991-07-08">8.VII.1991</collectingDate>
</date>
. leg.
<collectorName id="26E4B3C9FF977907FCA7C6A5FC6D67DF" box="[834,948,449,471]" pageId="12" pageNumber="277">B. Herczig</collectorName>
,
<collectionCode id="ED004EDAFF977907FC24C6A5FC0D67DF" box="[961,980,450,471]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="277" type="Museum">V</collectionCode>
. Markó et Z. Mészáros” (coll.
<collectionCode id="ED004EDAFF977907FEBFC6B9FE5967FB" box="[346,384,478,499]" pageId="12" pageNumber="277">RG</collectionCode>
).
<collectorName id="26E4B3C9FF977907FE77C6B9FE2567FC" box="[402,508,478,500]" pageId="12" pageNumber="277">Slide Nos</collectorName>
:
<collectionCode id="ED004EDAFF977907FDE2C6B9FDF367FB" box="[519,554,478,499]" pageId="12" pageNumber="277">VZ</collectionCode>
<quantity id="4CE97BFAFF977907FDCCC6B9FDB667FB" box="[553,623,478,499]" metricMagnitude="3" metricUnit="m" metricValue="9.771" pageId="12" pageNumber="277" unit="m" value="9771.0">
<elevation id="003C312CFF977907FDCCC6B9FDB667FB" box="[553,623,478,499]" metricMagnitude="3" metricUnit="m" metricValue="9.771" pageId="12" pageNumber="277" unit="m" value="9771.0">9771m</elevation>
</quantity>
,
<collectionCode id="ED004EDAFF977907FD9EC6B9FD4467FC" box="[635,669,478,500]" pageId="12" pageNumber="277">VZ</collectionCode>
<quantity id="4CE97BFAFF977907FD7BC6B9FD2967FC" box="[670,752,478,500]" metricMagnitude="4" metricUnit="m" metricValue="1.1513" pageId="12" pageNumber="277" unit="m" value="11513.0">
<elevation id="003C312CFF977907FD7BC6B9FD2967FC" box="[670,752,478,500]" metricMagnitude="4" metricUnit="m" metricValue="1.1513" pageId="12" pageNumber="277" unit="m" value="11513.0">11513m</elevation>
</quantity>
,
<specimenCode id="DBB77E64FF977907FD1EC6B9FC9767FC" box="[763,846,478,500]" collectionCode="VZ" pageId="12" pageNumber="277">VZ6753</specimenCode>
f,
<specimenCode id="DBB77E64FF977907FC87C6B9FC1867FC" box="[866,961,478,500]" collectionCode="VZ" pageId="12" pageNumber="277">VZ11473</specimenCode>
f,
<specimenCode id="DBB77E64FF977907FC30C6B9FBED67FC" box="[981,1076,478,500]" collectionCode="VZ" pageId="12" pageNumber="277">VZ11479</specimenCode>
f,
<specimenCode id="DBB77E64FF977907FF6FC69CFF316419" box="[138,232,507,529]" collectionCode="VZ" pageId="12" pageNumber="277">VZ11519</specimenCode>
f,
<specimenCode id="DBB77E64FF977907FF1EC69CFE836419" box="[251,346,507,529]" collectionCode="VZ" pageId="12" pageNumber="277">VZ11523</specimenCode>
f
</materialsCitation>
.
</paragraph>
<caption id="DF6E8697FF907900FF4BC2B0FBBE604C" ID-DOI="http://doi.org/10.5281/zenodo.10272225" ID-Zenodo-Dep="10272225" httpUri="https://zenodo.org/record/10272225/files/figure.png" pageId="11" pageNumber="276" startId="11.[174,219,1495,1517]" targetBox="[183,1092,192,1463]" targetPageId="11" targetType="figure">
<paragraph id="8BAED61FFF907900FF4BC2B0FBBE604C" blockId="11.[172,1127,1495,1604]" pageId="11" pageNumber="276">
<emphasis id="B9650A0DFF907900FF4BC2B0FEF863E4" bold="true" box="[174,289,1495,1517]" pageId="11" pageNumber="276">Figs 4143.</emphasis>
Male genitalia of
<taxonomicName id="4C11AD9CFF907900FE32C2BFFD4E63E5" box="[471,663,1496,1517]" class="Insecta" family="Noctuidae" genus="Dichagyris" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="276" phylum="Arthropoda" rank="species" species="forficula">
<emphasis id="B9650A0DFF907900FE32C2BFFD4E63E5" box="[471,663,1496,1517]" italics="true" pageId="11" pageNumber="276">Dichagyris forficula</emphasis>
</taxonomicName>
sspp. 41 =
<taxonomicName id="4C11AD9CFF907900FCE1C2BFFBE263E5" authority="(Eversmann, 1851)" baseAuthorityName="Eversmann" baseAuthorityYear="1851" box="[772,1083,1495,1517]" class="Insecta" family="Noctuidae" genus="Dichagyris" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="276" phylum="Arthropoda" rank="species" species="forficula">
<emphasis id="B9650A0DFF907900FCE1C2BFFCA863E5" box="[772,881,1496,1517]" italics="true" pageId="11" pageNumber="276">D. forficula</emphasis>
(
<bibRefCitation id="EF80ABEEFF907900FC9BC2BFFBED63E5" author="EVERSMANN, E. F." box="[894,1076,1495,1517]" pageId="11" pageNumber="276" pagination="610 - 644" refId="ref12022" refString="EVERSMANN, E. F. (1851): Description de quelques nouvelles espece de Lepidopteres de la Russie - Bulletin de la Societe Imperiale des Naturalistes de Moscou 24 (2): 610 - 644." type="journal article" year="1851">Eversmann, 1851</bibRefCitation>
)
</taxonomicName>
, NE Turkey, Prov. Van, slide No.: VZ10245m. 42 =
<taxonomicName id="4C11AD9CFF907900FD6BC292FCAE6001" authorityName="Tang &amp; Zhang &amp; Guo &amp; Luo &amp; Jiang &amp; Pan" authorityYear="2023" box="[654,887,1524,1546]" class="Insecta" family="Noctuidae" genus="Dichagyris" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="276" phylum="Arthropoda" rank="subSpecies" species="forficula" subSpecies="akdagestana">
<emphasis id="B9650A0DFF907900FD6BC292FCAE6001" box="[654,887,1524,1546]" italics="true" pageId="11" pageNumber="276">D. forficula akdagestana</emphasis>
</taxonomicName>
ssp. n., paratype, Russia, North Caucasus, Ingoushetia, slide No.: VZ9771m. 43 =
<taxonomicName id="4C11AD9CFF907900FCC4C175FF31604B" authority="(Staudinger, 1892)" baseAuthorityName="Staudinger" baseAuthorityYear="1892" class="Insecta" family="Noctuidae" genus="Dichagyris" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="276" phylum="Arthropoda" rank="subSpecies" species="forficula" subSpecies="hadjina">
<emphasis id="B9650A0DFF907900FCC4C175FC04602E" box="[801,989,1553,1575]" italics="true" pageId="11" pageNumber="276">D. forficula hadjina</emphasis>
(
<bibRefCitation id="EF80ABEEFF907900FC0FC176FF38604B" author="STAUDINGER, O." pageId="11" pageNumber="276" pagination="224 - 339" refId="ref13158" refString="STAUDINGER, O. (1892): Neue Arten und Varietaten von Lepidopteren des palaarktischen Faunengebiets. - Deutsche Entomologische Zeitung Iris 4 (2): 224 - 339, pl. 3 - 4." type="journal article" year="1892">Staudinger, 1892</bibRefCitation>
)
</taxonomicName>
, Turkey, lectotype of
<taxonomicName id="4C11AD9CFF907900FE23C149FC57604B" authority="Staudinger, 1899" authorityName="Staudinger" authorityYear="1899" box="[454,910,1582,1604]" class="Insecta" family="Noctuidae" genus="Agrotis" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="276" phylum="Arthropoda" rank="variety" species="hadjina" variety="zeituna">
<emphasis id="B9650A0DFF907900FE23C149FD85604B" box="[454,604,1582,1603]" italics="true" pageId="11" pageNumber="276">Agrotis hadjina</emphasis>
var.
<emphasis id="B9650A0DFF907900FD6BC148FD0F604C" box="[654,726,1583,1604]" italics="true" pageId="11" pageNumber="276">zeituna</emphasis>
Staudinger, 1899
</taxonomicName>
, slide No.: VZ9548m
</paragraph>
</caption>
<paragraph id="8BAED61FFF977907FF20C551FE6B6533" blockId="12.[134,1090,566,1062]" pageId="12" pageNumber="277">
Diagnosis. This subspecies belongs to the smaller ones of
<taxonomicName id="4C11AD9CFF977907FC69C550FBD46458" baseAuthorityName="Eversmann" baseAuthorityYear="1851" box="[908,1037,566,592]" class="Insecta" family="Noctuidae" genus="Dichagyris" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="277" phylum="Arthropoda" rank="species" species="forficula">
<emphasis id="B9650A0DFF977907FC69C550FBD46458" box="[908,1037,566,592]" italics="true" pageId="12" pageNumber="277">D. forficula</emphasis>
</taxonomicName>
, expanse
<quantity id="4CE97BFAFF977907FF3CC530FE85647A" box="[217,348,599,626]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="3.6" metricValueMin="3.3" pageId="12" pageNumber="277" unit="mm" value="34.5" valueMax="36.0" valueMin="33.0">3336 mm</quantity>
. Head, collar, thorax and forewings are nearly concolorous whitish-grey or light ash-grey, without any ochreous or reddish colouration, the forewings are nearly patternless or with faintly marked thin, crenulate crosslines only. Maculation weakly defined with some grey scales and filled with whitish scales, the submarginal field is slightly darker; the cilia are chequered with some darker grey scales. Hindwings whitish grey with diffuse slightly darker margins.
</paragraph>
<paragraph id="8BAED61FFF977907FF20C424FECA6576" blockId="12.[134,1090,566,1062]" pageId="12" pageNumber="277">
The male and female genitalia (
<figureCitation id="132ACA9AFF977907FDA6C424FD456555" box="[579,668,835,861]" captionStart="Figs 4143" captionStartId="11.[174,219,1495,1517]" captionTargetBox="[183,1092,192,1463]" captionTargetId="figure-18@11.[174,1095,189,1469]" captionTargetPageId="11" captionText="Figs 4143. Male genitalia ofDichagyris forficula sspp. 41 =D. forficula (Eversmann, 1851), NE Turkey, Prov. Van, slide No.: VZ10245m. 42 = D. forficula akdagestana ssp. n., paratype, Russia, North Caucasus, Ingoushetia, slide No.: VZ9771m. 43 = D. forficula hadjina (Staudinger, 1892), Turkey, lectotype of Agrotis hadjina var.zeituna Staudinger, 1899, slide No.: VZ9548m" figureDoi="http://doi.org/10.5281/zenodo.10272225" httpUri="https://zenodo.org/record/10272225/files/figure.png" pageId="12" pageNumber="277">Figs 42</figureCitation>
,
<figureCitation id="132ACA9AFF977907FD43C425FD1A6554" box="[678,707,834,860]" captionStart="Figs 6870" captionStartId="29.[174,219,1467,1489]" captionTargetBox="[193,1112,504,1429]" captionTargetId="figure-79@29.[174,1117,500,1436]" captionTargetPageId="29" captionText="Figs 6870. Genitalia of Dichagyris spp. 68 = Male genitalia of D. erubescens (Staudinger, 1892), Turkey, Prov. Malatya, slide No.: VZ11515m. 69 = Female genitalia of D. forficula akdagestana ssp. n., paratype, Russia, North Caucasus, Ingoushetia, slide No.: VZ6753f. 70 = female genitalia of D. forficula devota (Christoph, 1884), Turkmenistan, Kopet-Dagh, slide No.: VZ11483f" figureDoi="http://doi.org/10.5281/zenodo.10272251" httpUri="https://zenodo.org/record/10272251/files/figure.png" pageId="12" pageNumber="277">69</figureCitation>
) agree with those of the typical subspecies.
</paragraph>
<paragraph id="8BAED61FFF977907FF20C4E1FD6D622E" blockId="12.[134,1090,566,1062]" pageId="12" pageNumber="277">
Distribution. The new subspecies seems to be confined to certain calcareous hilly areas of the North Caucasus range in
<collectingRegion id="49D518FDFF977907FD59C4C0FC9965C9" box="[700,832,935,961]" country="Russia" name="Dagestan" pageId="12" pageNumber="277">Daghestan</collectingRegion>
and in the Ingoushia territory,
<collectingCountry id="F306968FFF977907FF18C4AEFE3665EB" box="[253,495,969,995]" name="Russia" pageId="12" pageNumber="277">Russian Federation</collectingCountry>
. Its range seems to be separated from the much more extended distribution of the nominotypical
<taxonomicName id="4C11AD9CFF977907FD1CC48CFC74620D" box="[761,941,1003,1029]" pageId="12" pageNumber="277" rank="subSpecies" subSpecies="occurring">race occurring</taxonomicName>
south from the main ridge of the Great Caucasus massif.
</paragraph>
<paragraph id="8BAED61FFF977907FF20C32EFEDB6274" blockId="12.[138,1088,1097,1148]" pageId="12" pageNumber="277">
Etymology. The name refers to the
<typeStatus id="54AA68BDFF977907FDA6C32DFDAB6257" box="[579,626,1098,1119]" pageId="12" pageNumber="277">type</typeStatus>
locality and the whitish colouration of the subspecies.
</paragraph>
<paragraph id="8BAED61FFF977907FF25C3D2FCBC631D" blockId="12.[192,1033,1205,1302]" pageId="12" pageNumber="277">
IV.
<collectionCode id="ED004EDAFF977907FF09C3D2FD4762C7" box="[236,670,1205,1231]" pageId="12" pageNumber="277">TAXONOMIC SUBDIVISION OF</collectionCode>
<taxonomicName id="4C11AD9CFF977907FD43C3D2FDDD62FA" authority="IN" authorityName="IN" class="Insecta" family="Noctuidae" genus="Dichagyris" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="277" phylum="Arthropoda" rank="species" species="forficula">
<emphasis id="B9650A0DFF977907FD43C3D2FBD062C7" box="[678,1033,1205,1231]" italics="true" pageId="12" pageNumber="277">DICHAGYRIS FORFICULA</emphasis>
IN
</taxonomicName>
<collectionCode id="ED004EDAFF977907FDE9C3BFFD3F62FA" box="[524,742,1240,1266]" pageId="12" pageNumber="277">CENTRAL ASIA</collectionCode>
with description of two new subspecies
</paragraph>
<paragraph id="8BAED61FFF977904FF20C253FEBA6788" blockId="12.[136,1090,1332,1594]" lastBlockId="15.[173,1128,189,384]" lastPageId="15" lastPageNumber="280" pageId="12" pageNumber="277">
The
<taxonomicName id="4C11AD9CFF977907FF18C252FE586346" box="[253,385,1332,1358]" class="Insecta" family="Noctuidae" genus="Dichagyris" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="277" phylum="Arthropoda" rank="species" species="forficula">
<emphasis id="B9650A0DFF977907FF18C252FE586346" box="[253,385,1332,1358]" italics="true" pageId="12" pageNumber="277">D. forficula</emphasis>
</taxonomicName>
specimens from the southeastern hilly part of
<collectingCountry id="F306968FFF977907FC37C253FF626378" name="Afghanistan" pageId="12" pageNumber="277">Afghanistan</collectingCountry>
(Sarobi, Khurd
<collectingRegion id="49D518FDFF977907FE6DC231FE0D6378" box="[392,468,1366,1392]" country="Afghanistan" name="Kabul" pageId="12" pageNumber="277">Kabul</collectingRegion>
) and adjacent part of
<collectingCountry id="F306968FFF977907FD08C231FC8F6378" box="[749,854,1366,1392]" name="Pakistan" pageId="12" pageNumber="277">Pakistan</collectingCountry>
(
<collectingRegion id="49D518FDFF977907FC8DC231FC256378" box="[872,1020,1366,1392]" country="Pakistan" name="Balochistan" pageId="12" pageNumber="277">Baluchistan</collectingRegion>
, near Quetta) are essentially similar to
<taxonomicName id="4C11AD9CFF977907FDC3C21FFD206399" authorityName="Tang &amp; Zhang &amp; Guo &amp; Luo &amp; Jiang &amp; Pan" authorityYear="2023" baseAuthorityName="Christoph" baseAuthorityYear="1884" box="[550,761,1399,1425]" class="Insecta" family="Noctuidae" genus="Dichagyris" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="277" phylum="Arthropoda" rank="subSpecies" species="forficula" subSpecies="devota">
<emphasis id="B9650A0DFF977907FDC3C21FFD206399" box="[550,761,1399,1425]" italics="true" pageId="12" pageNumber="277">D. forficula devota</emphasis>
</taxonomicName>
and they were mentioned and figured (
<collectionCode id="ED004EDAFF977907FEC9C2FEFEB263BD" box="[300,363,1433,1461]" pageId="12" pageNumber="277">LÖDL</collectionCode>
<emphasis id="B9650A0DFF977907FE96C2FDFE7263BB" box="[371,427,1433,1459]" italics="true" pageId="12" pageNumber="277">et al.</emphasis>
2012) as
<taxonomicName id="4C11AD9CFF977907FDFAC2FDFD3863BB" box="[543,737,1433,1459]" class="Insecta" family="Noctuidae" genus="Agrotis" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="277" phylum="Arthropoda" rank="subSpecies" species="devota" subSpecies="turana">
<emphasis id="B9650A0DFF977907FDFAC2FDFD3863BB" box="[543,737,1433,1459]" italics="true" pageId="12" pageNumber="277">D. devota turana</emphasis>
</taxonomicName>
due to the misidentification of
<taxonomicName id="4C11AD9CFF977907FF4CC2DCFE3263DC" authorityName="Staudinger" authorityYear="1892" box="[169,491,1467,1493]" class="Insecta" family="Noctuidae" genus="Agrotis" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="277" phylum="Arthropoda" rank="variety" species="renigera" variety="turana">
<emphasis id="B9650A0DFF977907FF4CC2DCFE3263DC" box="[169,491,1467,1493]" italics="true" pageId="12" pageNumber="277">Agrotis renigera var. turana</emphasis>
</taxonomicName>
(see below). They display nearly the same light ochreous grey colouration of the thorax and forewings as
<taxonomicName id="4C11AD9CFF977907FC83C2BAFBE063FE" authorityName="Tang &amp; Zhang &amp; Guo &amp; Luo &amp; Jiang &amp; Pan" authorityYear="2023" baseAuthorityName="Christoph" baseAuthorityYear="1884" box="[870,1081,1500,1526]" class="Insecta" family="Noctuidae" genus="Dichagyris" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="277" phylum="Arthropoda" rank="subSpecies" species="forficula" subSpecies="devota">
<emphasis id="B9650A0DFF977907FC83C2BAFBE063FE" box="[870,1081,1500,1526]" italics="true" pageId="12" pageNumber="277">D. forficula devota</emphasis>
</taxonomicName>
. However, these specimens have a slightly larger wing expanse (
<quantity id="4CE97BFAFF977907FC4BC299FBEA6010" box="[942,1075,1534,1560]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.3" metricValueMin="3.7" pageId="12" pageNumber="277" unit="mm" value="40.0" valueMax="43.0" valueMin="37.0">3743 mm</quantity>
), the forewings are more elongate and the crosslines are more expressed (
<figureCitation id="132ACA9AFF977904FBEBC178FF2F66D0" captionStart="Figs 1724" captionStartId="6.[136,181,1450,1472]" captionTargetBox="[141,1081,193,1423]" captionTargetId="figure-19@6.[136,1088,189,1424]" captionTargetPageId="6" captionText="Figs 1724. Dichagyris forficula sspp. 1720 = D. forficula pseudoturana ssp. n.: 1718 = paratype, females, Afghanistan, Paghman Mts; 19 = paratype, male, Pakistan, Quetta; 20 = paratype, female, Pakistan Quetta. 2124 = D. forficula chitralensis ssp. n., paratypes, Pakistan: 2122 = males, 2324 = females" figureDoi="http://doi.org/10.5281/zenodo.10272219" httpUri="https://zenodo.org/record/10272219/files/figure.png" lastPageId="15" lastPageNumber="280" pageId="12" pageNumber="277">Figs 1720</figureCitation>
) than in
<taxonomicName id="4C11AD9CFF947904FE82C7D9FDE366DF" authorityName="Tang &amp; Zhang &amp; Guo &amp; Luo &amp; Jiang &amp; Pan" authorityYear="2023" baseAuthorityName="Christoph" baseAuthorityYear="1884" box="[359,570,189,215]" class="Insecta" family="Noctuidae" genus="Dichagyris" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="280" phylum="Arthropoda" rank="subSpecies" species="forficula" subSpecies="devota">
<emphasis id="B9650A0DFF947904FE82C7D9FDE366DF" box="[359,570,189,215]" italics="true" pageId="15" pageNumber="280">D. forficula devota</emphasis>
</taxonomicName>
(
<figureCitation id="132ACA9AFF947904FDAFC7DAFD0966DF" box="[586,720,189,215]" captionStart="Figs 916" captionStartId="5.[174,219,1406,1428]" captionTargetBox="[178,1123,192,1367]" captionTargetId="figure-19@5.[174,1126,189,1369]" captionTargetPageId="5" captionText="Figs 916. Dichagyris forficula sspp. 912 = D. forficula akdagestana ssp. n., paratypes, Russia, Chechnya, Furtoug: 910 = male; 1112 = females. 1316 = D. forficula devota (Christoph, 1884): 13 = male, Turkmenistan, Kopet-Dagh, 14 = female, Turkmenistan, Kopet-Dagh, 15 = male, Iran, Shah Kuh, 16 = female, paratype of Agrotis devota eremica Amsel, 1935, Palestine" figureDoi="http://doi.org/10.5281/zenodo.10272217" httpUri="https://zenodo.org/record/10272217/files/figure.png" pageId="15" pageNumber="280">Figs 1316</figureCitation>
). The male genitalia (
<figureCitation id="132ACA9AFF947904FC3EC7DAFBF666D0" box="[987,1071,189,216]" captionStart="Figs 4749" captionStartId="14.[136,181,1473,1495]" captionTargetBox="[138,1081,193,1448]" captionTargetId="figure-19@14.[136,1088,189,1454]" captionTargetPageId="14" captionText="Figs 4749. Male genitalia of Dichagyris forficula sspp. 4748 = D. forficula devota (Christoph, 1884): 47 = Turkmenistan, Kopet-Dagh, slide No.: VZ9961m; 48 = Iran, Prov. Khorassan, slide No.: VZ11518m. 49 = D. forficula pseudoturana ssp. n., paratype, Afghanistan, Paghman Mts, slide No.: VZ10134m" figureDoi="http://doi.org/10.5281/zenodo.10272231" httpUri="https://zenodo.org/record/10272231/files/figure.png" pageId="15" pageNumber="280">Fig. 49</figureCitation>
) are essentially identical with those of the other subspecies of
<taxonomicName id="4C11AD9CFF947904FC89C787FC3466F1" box="[876,1005,223,249]" class="Insecta" family="Noctuidae" genus="Dichagyris" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="280" phylum="Arthropoda" rank="species" species="forficula">
<emphasis id="B9650A0DFF947904FC89C787FC3466F1" box="[876,1005,223,249]" italics="true" pageId="15" pageNumber="280">D. forficula</emphasis>
</taxonomicName>
, however the valvae and the claspers are slightly more elongate. The thin subbasal cornutus is somewhat shorter than in other subspecies, however, the completely, helicoidally recurved vesica is similar to those of
<taxonomicName id="4C11AD9CFF947904FCC5C622FBD36756" baseAuthorityName="Eversmann" baseAuthorityYear="1851" box="[800,1034,324,350]" class="Insecta" family="Noctuidae" genus="Dichagyris" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="280" phylum="Arthropoda" rank="subSpecies" species="forficula" subSpecies="forficula">
<emphasis id="B9650A0DFF947904FCC5C622FBD36756" box="[800,1034,324,350]" italics="true" pageId="15" pageNumber="280">D. forficula forficula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C11AD9CFF947904FBA9C622FE856777" authorityName="Tang &amp; Zhang &amp; Guo &amp; Luo &amp; Jiang &amp; Pan" authorityYear="2023" baseAuthorityName="Christoph" baseAuthorityYear="1884" class="Insecta" family="Noctuidae" genus="Dichagyris" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="280" phylum="Arthropoda" rank="subSpecies" species="forficula" subSpecies="devota">
<emphasis id="B9650A0DFF947904FBA9C622FE856777" italics="true" pageId="15" pageNumber="280">D. forficula devota</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>