Contributions To The Taxonomy And Biogeography Of The Genus Dichagyris (Subg. Dichagyris) Lederer, 1867 (Lepidoptera, Noctuidae, Noctuinae) Ii. The Review Of The D. Forficula Species Group
Author
Tang, Jin
Author
Zhang, Gongaote
Author
Guo, Junqi
Author
Luo, Lingxuan
Author
Jiang, Jiamei
Author
Pan, Hongbo
text
Acta Zoologica Academiae Scientiarum Hungaricae
2023
2023-08-25
69
3
265
301
http://dx.doi.org/10.3389/fmicb.2023.1259653
journal article
10.17109/AZH.69.3.265.2023http://zoobank.org/9C2F726C-ECC7-43C4-8DA9-68683652C62D
2064-2474
10272211
Dichagyris forficula akdagestana
subsp. n.
http://zoobank.org/
8258ACB9-4DB4-43F7-9866-D848803B09C7
(
Figs 9–12
,
42
,
69
)
Holotype
: male, “
Russia
, N Caucasus,
Daghestan
, Gubden vill.,
Karabudakhkentsky distr.
,
42°33’N
,
47°25’E
,
1000 m
,
3. VI. 2021
, leg.
V
. Zurilina”, gen. slide GYP
5775m
(coll.
PG
)
.
Paratypes
.
Russia
.
Daghestan
.
3 males
,
1 female
, with same data as holotype
;
1 female
, from the same locality,
27.VII.2020
, leg.
V
.
Zurilina
;
2 males
, from the same locality
21.
V
. and 2.
VI
.2021
, leg.
V
.
Zurilina
;
4 males
,
10 females
, from the same locality,
42°32’N
,
47°25’E
,
1030 m
,
27.
VI
.2022
, leg.
V
.
Zurilina
;
6 males
,
11 females
, from the same locality,
2.VII. and 8.VII.2022
, leg.
V
.
Zurilina
(all in coll.
PG
)
;
1 male
, with same data (coll.
Z. Varga
).
Slides
: GYP5586f, GYP5799f,
VZ
11518m
,
VZ
11973m
.
Ingoushetia
.
1 male
,
2 females
, “ChIASSR Furtoug
1000 m
12. Juli, 1989 leg.
Herczig
,
Uherkovich
,
Horváth
,
Szollát
,
Sárközi
”
;
1 male
, from the same site,
12-13.
VI
.1990
, leg. L. Ábrahám & Á. Uherkovich;
13 males
,
13 females
, with the same locality, “
2.VII.1991
, leg.
B. Herczig
,
V
.
Markó
et
Z. Mészáros
”
;
1 male
,
1 female
, “NE Caucasus, Lezsgi
1300 m
,
8.VII.1991
. leg.
B. Herczig
,
V
. Markó et Z. Mészáros” (coll.
RG
).
Slide Nos
:
VZ
9771m
,
VZ
11513m
,
VZ6753
f,
VZ11473
f,
VZ11479
f,
VZ11519
f,
VZ11523
f
.
Figs 41–43.
Male genitalia of
Dichagyris forficula
sspp. 41 =
D. forficula
(
Eversmann, 1851
)
, NE Turkey, Prov. Van, slide No.: VZ10245m. 42 =
D. forficula akdagestana
ssp. n., paratype, Russia, North Caucasus, Ingoushetia, slide No.: VZ9771m. 43 =
D. forficula hadjina
(
Staudinger, 1892
)
, Turkey, lectotype of
Agrotis hadjina
var.
zeituna
Staudinger, 1899
, slide No.: VZ9548m
Diagnosis. This subspecies belongs to the smaller ones of
D. forficula
, expanse
33–36 mm
. Head, collar, thorax and forewings are nearly concolorous whitish-grey or light ash-grey, without any ochreous or reddish colouration, the forewings are nearly patternless or with faintly marked thin, crenulate crosslines only. Maculation weakly defined with some grey scales and filled with whitish scales, the submarginal field is slightly darker; the cilia are chequered with some darker grey scales. Hindwings whitish grey with diffuse slightly darker margins.
The male and female genitalia (
Figs 42
,
69
) agree with those of the typical subspecies.
Distribution. The new subspecies seems to be confined to certain calcareous hilly areas of the North Caucasus range in
Daghestan
and in the Ingoushia territory,
Russian Federation
. Its range seems to be separated from the much more extended distribution of the nominotypical
race occurring
south from the main ridge of the Great Caucasus massif.
Etymology. The name refers to the
type
locality and the whitish colouration of the subspecies.
IV.
TAXONOMIC SUBDIVISION OF
DICHAGYRIS FORFICULA
IN
CENTRAL ASIA
with description of two new subspecies
The
D. forficula
specimens from the southeastern hilly part of
Afghanistan
(Sarobi, Khurd
Kabul
) and adjacent part of
Pakistan
(
Baluchistan
, near Quetta) are essentially similar to
D. forficula devota
and they were mentioned and figured (
LÖDL
et al.
2012) as
D. devota turana
due to the misidentification of
Agrotis renigera var. turana
(see below). They display nearly the same light ochreous grey colouration of the thorax and forewings as
D. forficula devota
. However, these specimens have a slightly larger wing expanse (
37–43 mm
), the forewings are more elongate and the crosslines are more expressed (
Figs 17–20
) than in
D. forficula devota
(
Figs 13–16
). The male genitalia (
Fig. 49
) are essentially identical with those of the other subspecies of
D. forficula
, however the valvae and the claspers are slightly more elongate. The thin subbasal cornutus is somewhat shorter than in other subspecies, however, the completely, helicoidally recurved vesica is similar to those of
D. forficula forficula
and
D. forficula devota
.