468 lines
56 KiB
XML
468 lines
56 KiB
XML
<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra" docDate="2009" docId="1772122E7862FFA1FF013BD7AE77EE60" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Hylekobolus brachiosauroides Wesener & Enghoff & Sierwald 2009, sp. n." docType="treatment" docUuid="40FD10B2-63FB-4627-AAAC-6961BE9102E5" docUuidSource="ZooBank" docVersion="11" lastPageNumber="104" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="102" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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||
<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-09-04</mods:number>
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</mods:detail>
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||
<mods:detail type="volume">
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<mods:number>19</mods:number>
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</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>19</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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||
<mods:start>1</mods:start>
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||
<mods:end>128</mods:end>
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||
</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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||
<mods:classification>journal article</mods:classification>
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||
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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||
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
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||
<mods:identifier type="ISSN">1313–2970</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
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||
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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||
</mods:mods>
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||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791465" ID-GBIF-Taxon="163747613" ID-Zenodo-Dep="3791465" LSID="urn:lsid:zoobank.org:act:40FD10B2-63FB-4627-AAAC-6961BE9102E5" httpUri="http://treatment.plazi.org/id/1772122E7862FFA1FF013BD7AE77EE60" lastPageId="103" lastPageNumber="104" pageId="101" pageNumber="102">
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||
<subSubSection box="[140,679,1187,1214]" pageId="101" pageNumber="102" type="nomenclature">
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||
<paragraph blockId="101.[140,968,1187,1249]" box="[140,679,1187,1214]" pageId="101" pageNumber="102">
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||
<heading box="[140,679,1187,1214]" fontSize="11" level="2" pageId="101" pageNumber="102" reason="3">
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<emphasis bold="true" box="[140,679,1187,1214]" pageId="101" pageNumber="102">
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<taxonomicName authority="Wesener" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,601,1187,1214]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis bold="true" box="[140,493,1187,1214]" italics="true" pageId="101" pageNumber="102">Hylekobolus brachiosauroides</emphasis>
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Wesener
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</taxonomicName>
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,
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<taxonomicNameLabel box="[614,679,1187,1213]" pageId="101" pageNumber="102" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,968,1222,1249]" pageId="101" pageNumber="102" type="description">
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<paragraph blockId="101.[140,968,1187,1249]" box="[140,968,1222,1249]" pageId="101" pageNumber="102">
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<uri box="[140,968,1222,1249]" pageId="101" pageNumber="102">
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urn:lsid:zoobank.org:act:
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<uuid box="[416,968,1222,1249]" pageId="101" pageNumber="102">40FD10B2-63FB-4627-AAAC-6961BE9102E5</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="102" lastPageNumber="103" pageId="101" pageNumber="102" type="materials_examined">
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<paragraph blockId="101.[140,1108,1293,1672]" pageId="101" pageNumber="102">
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<emphasis bold="true" box="[140,363,1293,1319]" pageId="101" pageNumber="102">
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<emphasis box="[140,237,1293,1319]" inLineHeading="true" pageId="101" pageNumber="102" reason="1">Material</emphasis>
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examined:
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</emphasis>
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<materialsCitation ID-GBIF-Occurrence="2608818240" collectingDate="2007-05-23" collectionCode="FMMC" collectorName="Wesener" county="Ivorona" elevation="114" latitude="-24.833767" location="Vohimena Mountains" longLatPrecision="1" longitude="46.94905" municipality="West" pageId="101" pageNumber="102" specimenCode="W089" specimenCount="27" specimenCount-female="21" typeStatus="holotype">
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26 ♂,
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<specimenCount box="[443,501,1293,1320]" pageId="101" pageNumber="102" type="female">21 ♀</specimenCount>
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,
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<specimenCount box="[516,598,1293,1319]" pageId="101" pageNumber="102" type="generic">6 imm.</specimenCount>
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<emphasis box="[605,702,1293,1319]" italics="true" pageId="101" pageNumber="102">
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<typeStatus box="[605,697,1293,1319]" pageId="101" pageNumber="102">Holotype</typeStatus>
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:
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</emphasis>
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1 ♂ (
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<quantity box="[772,854,1293,1320]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="101" pageNumber="102" unit="mm" value="55.0">55 mm</quantity>
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long),
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<collectionCode box="[935,1022,1293,1319]" pageId="101" pageNumber="102">FMMC</collectionCode>
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<specimenCode box="[1030,1103,1293,1319]" collectionCode="FMMC" pageId="101" pageNumber="102">W089</specimenCode>
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,
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<location LSID="urn:lsid:plazi:treatment:1772122E7862FFA1FF013BD7AE77EE60:9A04F5E37862FFA3FF013A44AE22EF1B" box="[140,379,1328,1354]" county="Ivorona" latitude="-24.833767" longLatPrecision="1" longitude="46.94905" municipality="West" name="Vohimena Mountains" pageId="101" pageNumber="102">Vohimena Mountains</location>
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,
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<collectingMunicipality box="[388,442,1329,1355]" pageId="101" pageNumber="102">West</collectingMunicipality>
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slope,
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<collectingCounty box="[514,600,1329,1355]" pageId="101" pageNumber="102">Ivorona</collectingCounty>
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,
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<quantity box="[609,682,1328,1355]" metricMagnitude="2" metricUnit="m" metricValue="1.14" pageId="101" pageNumber="102" unit="m" value="114.0">
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<elevation box="[609,682,1328,1355]" metricMagnitude="2" metricUnit="m" metricValue="1.14" pageId="101" pageNumber="102" unit="m" value="114.0">114 m</elevation>
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</quantity>
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, rainforest,
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<geoCoordinate box="[806,948,1328,1355]" degrees="24" direction="south" minutes="50.026" orientation="latitude" pageId="101" pageNumber="102" precision="1" value="-24.833767">24°50.026’ S</geoCoordinate>
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,
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<geoCoordinate box="[958,1103,1328,1355]" degrees="46" direction="east" minutes="56.943" orientation="longitude" pageId="101" pageNumber="102" precision="1" value="46.94905">46°56.943’ E</geoCoordinate>
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, leg.
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<collectorName box="[183,274,1364,1390]" pageId="101" pageNumber="102">Wesener</collectorName>
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et al.,
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<date box="[345,462,1364,1390]" pageId="101" pageNumber="102" value="2007-05-23">
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<collectingDate box="[345,462,1364,1390]" pageId="101" pageNumber="102" value="2007-05-23">23.V.2007</collectingDate>
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</date>
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</materialsCitation>
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.
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<materialsCitation ID-GBIF-Occurrence="2608818212" box="[472,1103,1363,1390]" collectingDate="2007-05-23" collectionCode="FMMC" collectorName="Wesener" county="Ivorona" elevation="114" latitude="-24.833767" location="Vohimena Mountains" longLatPrecision="1" longitude="46.94905" municipality="West" pageId="101" pageNumber="102" specimenCode="W089" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
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<emphasis box="[472,574,1364,1389]" italics="true" pageId="101" pageNumber="102">
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<typeStatus box="[472,568,1364,1389]" pageId="101" pageNumber="102">Paratypes</typeStatus>
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:
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</emphasis>
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3 ♂,
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<specimenCount box="[635,677,1364,1390]" pageId="101" pageNumber="102" type="female">4 ♀</specimenCount>
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,
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<collectionCode box="[689,776,1364,1390]" pageId="101" pageNumber="102">FMMC</collectionCode>
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<specimenCode box="[783,855,1364,1390]" collectionCode="FMMC" pageId="101" pageNumber="102">W089</specimenCode>
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, same data as holotype
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2608818309" box="[140,652,1399,1425]" collectingDate="2007-05-23" collectionCode="ZMUC" collectorName="Wesener" county="Ivorona" elevation="114" latitude="-24.833767" location="Vohimena Mountains" longLatPrecision="1" longitude="46.94905" municipality="West" pageId="101" pageNumber="102" specimenCode="W089" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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1 ♂,
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<specimenCount box="[192,233,1399,1425]" pageId="101" pageNumber="102" type="female">1 ♀</specimenCount>
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,
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<collectionCode box="[245,331,1399,1425]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="101" pageNumber="102">ZMUC</collectionCode>
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<specimenCode box="[336,408,1399,1425]" collectionCode="ZMUC" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="101" pageNumber="102">W089</specimenCode>
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||
, same data as holotype
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||
</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2608818296" box="[661,1103,1399,1425]" collectingDate="2007-05-23" collectionCode="ZMH" collectorName="Wesener" county="Ivorona" elevation="114" latitude="-24.833767" location="Vohimena Mountains" longLatPrecision="1" longitude="46.94905" municipality="West" pageId="101" pageNumber="102" specimenCode="W089" specimenCount="1" typeStatus="paratype">
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1 ♂,
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<collectionCode box="[714,782,1399,1425]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="101" pageNumber="102" type="Museum">ZMH</collectionCode>
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<specimenCode box="[787,859,1399,1425]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="101" pageNumber="102" type="Museum">W089</specimenCode>
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||
, same data as holotype
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</materialsCitation>
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||
;
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<materialsCitation ID-GBIF-Occurrence="2608818227" box="[140,629,1434,1460]" collectingDate="2007-05-23" collectionCode="CAS" collectorName="Wesener" county="Ivorona" elevation="114" latitude="-24.833767" location="Vohimena Mountains" longLatPrecision="1" longitude="46.94905" municipality="West" pageId="101" pageNumber="102" specimenCode="W089" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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1 ♂,
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<specimenCount box="[195,237,1434,1460]" pageId="101" pageNumber="102" type="female">1 ♀</specimenCount>
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,
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<collectionCode box="[250,301,1434,1460]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="101" pageNumber="102" type="Museum">CAS</collectionCode>
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<specimenCode box="[307,380,1434,1460]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="101" pageNumber="102" type="Museum">W089</specimenCode>
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, same data as holotype
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2608818332" box="[639,1080,1434,1460]" collectingDate="2007-05-23" collectionCode="ZSM" collectorName="Wesener" county="Ivorona" elevation="114" latitude="-24.833767" location="Vohimena Mountains" longLatPrecision="1" longitude="46.94905" municipality="West" pageId="101" pageNumber="102" specimenCode="W089" specimenCount="1" typeStatus="paratype">
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1 ♂,
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<collectionCode box="[694,752,1434,1460]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="101" pageNumber="102" type="Museum">ZSM</collectionCode>
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<specimenCode box="[759,831,1434,1460]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="101" pageNumber="102" type="Museum">W089</specimenCode>
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, same data as holotype
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</materialsCitation>
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.
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</paragraph>
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<paragraph blockId="101.[140,1108,1293,1672]" lastBlockId="102.[140,1108,167,510]" lastPageId="102" lastPageNumber="103" pageId="101" pageNumber="102">
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<materialsCitation ID-GBIF-Occurrence="2608818349" collectingDate="2007-05-23" collectionCode="FMMC" collectorName="University" county="Ivorona" elevation="114" latitude="-24.833767" location="Vohimena Mountains" longLatPrecision="1" longitude="46.94905" municipality="West" pageId="101" pageNumber="102" specimenCount="4" specimenCount-female="4" stateProvince="Antananarivo">
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||
<emphasis bold="true" box="[185,490,1469,1495]" pageId="101" pageNumber="102">
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||
<typeStatus box="[185,485,1469,1495]" isOtherTypeStatus="true" pageId="101" pageNumber="102">
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||
<emphasis box="[185,257,1469,1495]" inLineHeading="true" pageId="101" pageNumber="102" reason="1">Other</emphasis>
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||
material examined
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||
</typeStatus>
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||
:
|
||
</emphasis>
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4 ♂,
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||
<specimenCount box="[551,593,1469,1496]" pageId="101" pageNumber="102" type="female">4 ♀</specimenCount>
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||
, W089, same data as holotype, voucher specimens deposited at the
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<collectorName box="[398,514,1504,1530]" pageId="101" pageNumber="102">University</collectorName>
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||
<collectingRegion box="[523,679,1504,1530]" country="Madagascar" name="Antananarivo" pageId="101" pageNumber="102">Antananarivo</collectingRegion>
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||
</materialsCitation>
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||
;
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||
<materialsCitation ID-GBIF-Occurrence="2608818355" collectingDate="2007-06-08" collectionCode="FMMC" collectorName="Wesener" elevation="46" latitude="-24.828167" location="Isaka-Ivondro Sud" longLatPrecision="1" longitude="46.866184" municipality="Andohahela" pageId="101" pageNumber="102" specimenCode="W049" specimenCount="6" specimenCount-female="4" stateProvince="Antananarivo">
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||
5 ♂,
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||
<specimenCount box="[752,796,1504,1531]" pageId="101" pageNumber="102" type="female">4 ♀</specimenCount>
|
||
,
|
||
<specimenCount box="[812,900,1504,1530]" pageId="101" pageNumber="102" type="generic">2 imm.</specimenCount>
|
||
,
|
||
<collectionCode box="[912,1002,1504,1530]" pageId="101" pageNumber="102">FMMC</collectionCode>
|
||
<specimenCode box="[1010,1084,1504,1530]" collectionCode="FMMC" pageId="101" pageNumber="102">W049</specimenCode>
|
||
B,
|
||
<collectingMunicipality box="[140,279,1539,1565]" pageId="101" pageNumber="102">Andohahela</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:1772122E7862FFA1FF013BD7AE77EE60:9A04F5E37862FFA3FEA83977AEA2EC4C" box="[293,507,1539,1565]" latitude="-24.828167" longLatPrecision="1" longitude="46.866184" municipality="Andohahela" name="Isaka-Ivondro Sud" pageId="101" pageNumber="102" stateProvince="Antananarivo">Isaka-Ivondro Sud</location>
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||
,
|
||
<quantity box="[522,588,1539,1566]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="101" pageNumber="102" unit="m" value="46.0">
|
||
<elevation box="[522,588,1539,1566]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="101" pageNumber="102" unit="m" value="46.0">46 m</elevation>
|
||
</quantity>
|
||
, rainforest,
|
||
<geoCoordinate box="[728,881,1539,1566]" degrees="24" direction="south" minutes="49.690" orientation="latitude" pageId="101" pageNumber="102" precision="1" value="-24.828167">24°49.690’ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[897,1053,1539,1566]" degrees="46" direction="east" minutes="51.971" orientation="longitude" pageId="101" pageNumber="102" precision="1" value="46.866184">46°51.971’ E</geoCoordinate>
|
||
, leg.
|
||
<collectorName box="[140,235,1575,1601]" pageId="101" pageNumber="102">Wesener</collectorName>
|
||
et al.,
|
||
<date box="[309,444,1575,1601]" pageId="101" pageNumber="102" value="2007-06-08">
|
||
<collectingDate box="[309,444,1575,1601]" pageId="101" pageNumber="102" value="2007-06-08">08.VI.2007</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2608818354" collectingDate="2007-06-08" collectionCode="FMMC" collectorName="University" elevation="46" latitude="-24.828167" location="Isaka-Ivondro Sud" longLatPrecision="1" longitude="46.866184" municipality="Andohahela" pageId="101" pageNumber="102" specimenCount="6" specimenCount-female="4" stateProvince="Antananarivo">
|
||
5 ♂,
|
||
<specimenCount box="[510,552,1575,1601]" pageId="101" pageNumber="102" type="female">4 ♀</specimenCount>
|
||
,
|
||
<specimenCount box="[566,651,1575,1601]" pageId="101" pageNumber="102" type="generic">2 imm.</specimenCount>
|
||
, W049B, same data as previous, voucher specimens deposited at the
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<collectorName box="[455,571,1610,1636]" pageId="101" pageNumber="102">University</collectorName>
|
||
<collectingRegion box="[580,736,1610,1636]" country="Madagascar" name="Antananarivo" pageId="101" pageNumber="102">Antananarivo</collectingRegion>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2608818258" collectingDate="2007-06-14" collectionCode="FMMC" collectorName="Wesener" lastPageId="102" lastPageNumber="103" latitude="-25.086176" location="Grand Lavasoa" longLatPrecision="1" longitude="46.74887" municipality="Ambatotsirongorongo Mountain" pageId="101" pageNumber="102" specimenCode="W099" specimenCount="2" specimenCount-female="2" stateProvince="Antananarivo">
|
||
2 ♂,
|
||
<specimenCount box="[810,854,1610,1636]" pageId="101" pageNumber="102" type="female">2 ♀</specimenCount>
|
||
,
|
||
<collectionCode box="[870,960,1610,1636]" pageId="101" pageNumber="102">FMMC</collectionCode>
|
||
<specimenCode box="[968,1044,1610,1636]" collectionCode="FMMC" pageId="101" pageNumber="102">W099</specimenCode>
|
||
,
|
||
<collectingMunicipality pageId="101" pageNumber="102">Ambatotsirongorongo Mountain</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:1772122E7862FFA1FF013BD7AE77EE60:9A04F5E37862FFA3FE6E3919ADD4ECD9" box="[483,653,1645,1672]" latitude="-25.086176" longLatPrecision="1" longitude="46.74887" municipality="Ambatotsirongorongo Mountain" name="Grand Lavasoa" pageId="101" pageNumber="102" stateProvince="Antananarivo">Grand Lavasoa</location>
|
||
, rainforest,
|
||
<geoCoordinate box="[784,944,1645,1672]" degrees="25" direction="south" minutes="5" orientation="latitude" pageId="101" pageNumber="102" precision="1" seconds="10.23" value="-25.086176">25°5’10.23” S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="46" direction="east" lastPageId="102" lastPageNumber="103" minutes="44" orientation="longitude" pageId="101" pageNumber="102" precision="1" seconds="55.93" value="46.74887">46°44’55.93” E</geoCoordinate>
|
||
, rainforest, leg.
|
||
<collectorName box="[341,436,167,193]" pageId="102" pageNumber="103">Wesener</collectorName>
|
||
et al.,
|
||
<date box="[515,650,167,193]" pageId="102" pageNumber="103" value="2007-06-14">
|
||
<collectingDate box="[515,650,167,193]" pageId="102" pageNumber="103" value="2007-06-14">14.VI.2007</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2608818361" collectingDate="2007-06-14" collectionCode="FMMC" collectorName="University" latitude="-25.086176" location="Grand Lavasoa" longLatPrecision="1" longitude="46.74887" municipality="Ambatotsirongorongo Mountain" pageId="102" pageNumber="103" specimenCount="3" specimenCount-female="1" stateProvince="Antananarivo">
|
||
2 ♂,
|
||
<specimenCount box="[722,765,167,193]" pageId="102" pageNumber="103" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount box="[781,867,167,193]" pageId="102" pageNumber="103" type="generic">2 imm.</specimenCount>
|
||
, W099, same data as previous, voucher specimens deposited at the
|
||
<collectorName box="[656,772,202,228]" pageId="102" pageNumber="103">University</collectorName>
|
||
<collectingRegion box="[779,936,202,228]" country="Madagascar" name="Antananarivo" pageId="102" pageNumber="103">Antananarivo</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="102" pageNumber="103" type="diagnosis">
|
||
<paragraph blockId="102.[140,1108,167,510]" pageId="102" pageNumber="103">
|
||
<emphasis bold="true" box="[185,459,237,263]" pageId="102" pageNumber="103">
|
||
<emphasis box="[185,327,237,263]" inLineHeading="true" pageId="102" pageNumber="103" reason="1">Differential</emphasis>
|
||
diagnosis:
|
||
</emphasis>
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[469,691,237,263]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[469,691,237,263]" italics="true" pageId="102" pageNumber="103">H. brachiosauroides</emphasis>
|
||
</taxonomicName>
|
||
differs in size and colour from the sympatric living
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[336,496,272,298]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[336,496,272,298]" italics="true" pageId="102" pageNumber="103">H. albicollaris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[505,569,273,299]" pageId="102" pageNumber="103" rank="species">sp. n.</taxonomicNameLabel>
|
||
The posterior gonopods (
|
||
<figureCitation box="[872,1027,272,299]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="102" pageNumber="103">Figs 55M, O</figureCitation>
|
||
) of
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species">
|
||
<emphasis italics="true" pageId="102" pageNumber="103">H. brachiosauroides</emphasis>
|
||
</taxonomicName>
|
||
differ from other, same-sized
|
||
<taxonomicName box="[668,801,307,333]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[668,801,307,333]" italics="true" pageId="102" pageNumber="103">Hylekobolus</emphasis>
|
||
</taxonomicName>
|
||
species: the lateral margin carries a thick, swollen, well-rounded projection (
|
||
<figureCitation box="[724,839,342,369]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="102" pageNumber="103">Fig. 55M</figureCitation>
|
||
), a feature only shared with
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[201,349,379,404]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[201,349,379,404]" italics="true" pageId="102" pageNumber="103">H. montanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[358,422,378,404]" pageId="102" pageNumber="103" rank="species">sp. n.</taxonomicNameLabel>
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[432,652,378,404]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[432,652,378,404]" italics="true" pageId="102" pageNumber="103">H. brachiosauroides</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[808,956,379,404]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[808,956,379,404]" italics="true" pageId="102" pageNumber="103">H. montanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[965,1029,378,404]" pageId="102" pageNumber="103" rank="species">sp. n.</taxonomicNameLabel>
|
||
in the presence of a protruding apical process (
|
||
<emphasis box="[604,616,414,439]" italics="true" pageId="102" pageNumber="103">x</emphasis>
|
||
) on the posterior gonopods, a weakly protruding lateral retrorse process of the anterior gonopod (
|
||
<figureCitation box="[802,914,448,475]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="102" pageNumber="103">Fig. 55N</figureCitation>
|
||
), and a different colour pattern.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767348" ID-Zenodo-Dep="3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="102" pageNumber="103" startId="102.[140,212,1430,1452]" targetBox="[140,1108,548,1411]" targetPageId="102">
|
||
<paragraph blockId="102.[140,1108,1430,1676]" pageId="102" pageNumber="103">
|
||
<emphasis bold="true" box="[140,254,1430,1452]" pageId="102" pageNumber="103">Figure 55.</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[264,524,1430,1451]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[264,524,1430,1451]" italics="true" pageId="102" pageNumber="103">Hylekobolus brachiosauroides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[532,586,1431,1452]" pageId="102" pageNumber="103" rank="species">sp. n.</taxonomicNameLabel>
|
||
, A–H male paratype, I–K female paratype, L–O male holotype.
|
||
<emphasis bold="true" box="[233,252,1462,1484]" pageId="102" pageNumber="103">A</emphasis>
|
||
head, lateral view
|
||
<emphasis bold="true" box="[422,439,1462,1484]" pageId="102" pageNumber="103">B</emphasis>
|
||
legs 1–7 and gonopods, ventral view
|
||
<emphasis bold="true" box="[784,803,1462,1484]" pageId="102" pageNumber="103">C</emphasis>
|
||
telson
|
||
<emphasis bold="true" box="[869,888,1462,1484]" pageId="102" pageNumber="103">D</emphasis>
|
||
antennal groove
|
||
<emphasis bold="true" box="[1046,1061,1462,1484]" pageId="102" pageNumber="103">E</emphasis>
|
||
gnathochilarium, ventral view
|
||
<emphasis bold="true" box="[392,406,1494,1516]" pageId="102" pageNumber="103">F</emphasis>
|
||
gnathochilarium, mentum
|
||
<emphasis bold="true" box="[665,685,1494,1516]" pageId="102" pageNumber="103">G</emphasis>
|
||
gnathochilarium, stipites
|
||
<emphasis bold="true" box="[927,967,1494,1516]" pageId="102" pageNumber="103">H, I</emphasis>
|
||
right midbody legs
|
||
<emphasis bold="true" box="[179,187,1526,1548]" pageId="102" pageNumber="103">Į</emphasis>
|
||
right vulva, posterior view
|
||
<emphasis bold="true" box="[436,453,1526,1548]" pageId="102" pageNumber="103">K</emphasis>
|
||
right vulva, lateral view
|
||
<emphasis bold="true" box="[676,691,1526,1548]" pageId="102" pageNumber="103">L</emphasis>
|
||
anterior gonopod, anterior view
|
||
<emphasis bold="true" box="[993,1014,1526,1548]" pageId="102" pageNumber="103">M</emphasis>
|
||
right posterior gonopod, anterior view
|
||
<emphasis bold="true" box="[422,442,1558,1580]" pageId="102" pageNumber="103">N</emphasis>
|
||
anterior gonopod, posterior view
|
||
<emphasis bold="true" box="[762,783,1558,1580]" pageId="102" pageNumber="103">O</emphasis>
|
||
right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="103" pageNumber="104" type="description">
|
||
<paragraph blockId="103.[140,1108,166,1073]" pageId="103" pageNumber="104">
|
||
<emphasis bold="true" box="[185,335,166,192]" pageId="103" pageNumber="104">
|
||
<emphasis box="[185,330,166,192]" inLineHeading="true" pageId="103" pageNumber="104" reason="1">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
<emphasis box="[343,495,167,192]" italics="true" pageId="103" pageNumber="104">Measurements</emphasis>
|
||
: males with 50 or 51 rings,
|
||
<quantity box="[818,946,166,193]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="6.2" metricValueMin="5.3" pageId="103" pageNumber="104" unit="mm" value="57.5" valueMax="62.0" valueMin="53.0">53–62 mm</quantity>
|
||
long,
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.2" metricValueMin="4.8" pageId="103" pageNumber="104" unit="mm" value="5.0" valueMax="5.2" valueMin="4.8">4.8–5.2 mm</quantity>
|
||
wide. Females with 50 or 51 rings,
|
||
<quantity box="[591,718,202,229]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="7.2" metricValueMin="5.8" pageId="103" pageNumber="104" unit="mm" value="65.0" valueMax="72.0" valueMin="58.0">58–72 mm</quantity>
|
||
long, 5.1–6.0 mm wide.
|
||
</paragraph>
|
||
<paragraph blockId="103.[140,1108,166,1073]" pageId="103" pageNumber="104">
|
||
<emphasis box="[185,308,237,263]" italics="true" pageId="103" pageNumber="104">Coloration:</emphasis>
|
||
anterior part of head and collum, posterior part of telson and posterior margin of body rings lightly red or white (
|
||
<figureCitation box="[610,714,272,299]" captionStart="Figure 53" captionStartId="97.[140,212,848,870]" captionTargetBox="[140,1108,165,829]" captionTargetId="figure@97.[140,1108,165,829]" captionTargetPageId="97" captionText="Figure 53. Living specimens of A Hylekobolus brachiosauroides sp. n., male B H. griseus sp. n., mating pair C H. albicollaris sp. n." figureDoi="http://doi.org/10.5281/zenodo.3820856" httpUri="https://zenodo.org/record/3820856/files/figure.png" pageId="103" pageNumber="104">Figs 53A</figureCitation>
|
||
,
|
||
<figureCitation box="[725,810,272,299]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="103" pageNumber="104">55A, C</figureCitation>
|
||
). Legs and antennae white (
|
||
<figureCitation box="[148,251,307,334]" captionStart="Figure 53" captionStartId="97.[140,212,848,870]" captionTargetBox="[140,1108,165,829]" captionTargetId="figure@97.[140,1108,165,829]" captionTargetPageId="97" captionText="Figure 53. Living specimens of A Hylekobolus brachiosauroides sp. n., male B H. griseus sp. n., mating pair C H. albicollaris sp. n." figureDoi="http://doi.org/10.5281/zenodo.3820856" httpUri="https://zenodo.org/record/3820856/files/figure.png" pageId="103" pageNumber="104">Fig. 53A</figureCitation>
|
||
). Anterior part of body rings, telson, posterior part of head and collum black (
|
||
<figureCitation box="[147,252,342,369]" captionStart="Figure 53" captionStartId="97.[140,212,848,870]" captionTargetBox="[140,1108,165,829]" captionTargetId="figure@97.[140,1108,165,829]" captionTargetPageId="97" captionText="Figure 53. Living specimens of A Hylekobolus brachiosauroides sp. n., male B H. griseus sp. n., mating pair C H. albicollaris sp. n." figureDoi="http://doi.org/10.5281/zenodo.3820856" httpUri="https://zenodo.org/record/3820856/files/figure.png" pageId="103" pageNumber="104">Figs 53A</figureCitation>
|
||
,
|
||
<figureCitation box="[264,316,342,369]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="103" pageNumber="104">55A</figureCitation>
|
||
).
|
||
<emphasis box="[335,378,343,368]" italics="true" pageId="103" pageNumber="104">Eyes</emphasis>
|
||
with 40–44 ocelli (
|
||
<figureCitation box="[601,708,342,369]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="103" pageNumber="104">Fig. 55D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="103.[140,1108,166,1073]" pageId="103" pageNumber="104">
|
||
<emphasis box="[185,372,378,404]" italics="true" pageId="103" pageNumber="104">Anterior gonopod</emphasis>
|
||
sternite basally wide, apically elevated into a triangular lobe with a triangular, wide, well-rounded tip (
|
||
<figureCitation box="[557,659,413,440]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="103" pageNumber="104">Fig. 55L</figureCitation>
|
||
). Mesal process of coxite wide and only slightly protruding (
|
||
<figureCitation box="[374,479,448,475]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="103" pageNumber="104">Fig. 55L</figureCitation>
|
||
). Apical process of telopodite large and well-rounded. Retrorse margin wide, not projecting above telopodite margin (
|
||
<figureCitation box="[863,971,483,510]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="103" pageNumber="104">Fig. 55N</figureCitation>
|
||
). Margin of apical process basally wide and projecting (
|
||
<figureCitation box="[623,730,518,545]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="103" pageNumber="104">Fig. 55N</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="103.[140,1108,166,1073]" pageId="103" pageNumber="104">
|
||
<emphasis box="[185,370,554,580]" italics="true" pageId="103" pageNumber="104">Posterior gonopods</emphasis>
|
||
telopodite bent 90° (
|
||
<figureCitation box="[596,697,554,581]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="103" pageNumber="104">Fig. 55O</figureCitation>
|
||
). Lateral margin with a thick, swollen, well-rounded projection (
|
||
<emphasis box="[418,437,590,615]" italics="true" pageId="103" pageNumber="104">w</emphasis>
|
||
,
|
||
<figureCitation box="[452,559,589,616]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="103" pageNumber="104">Fig. 55M</figureCitation>
|
||
). Apical process (
|
||
<emphasis box="[751,763,590,615]" italics="true" pageId="103" pageNumber="104">x</emphasis>
|
||
) apically slender, basally with a widely protruding margin (
|
||
<figureCitation box="[426,530,624,651]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="103" pageNumber="104">Fig. 55M</figureCitation>
|
||
). Membranous lobe (
|
||
<emphasis box="[756,766,625,650]" italics="true" pageId="103" pageNumber="104">y</emphasis>
|
||
) rectangular (
|
||
<figureCitation box="[911,1015,624,651]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="103" pageNumber="104">Fig. 55M</figureCitation>
|
||
). Projection of sperm canal (
|
||
<emphasis box="[360,371,660,685]" italics="true" pageId="103" pageNumber="104">z</emphasis>
|
||
) long and slender, slightly curved towards apical process (
|
||
<figureCitation box="[984,1089,659,686]" captionStart="Figure 55" captionStartId="102.[140,212,1430,1452]" captionTargetBox="[140,1108,548,1411]" captionTargetId="figure@102.[140,1109,548,1411]" captionTargetPageId="102" captionText="Figure 55. Hylekobolus brachiosauroides sp. n., A–H male paratype, I–K female paratype, L–O male holotype. A head, lateral view B legs 1–7 and gonopods, ventral view C telson D antennal groove E gnathochilarium, ventral view F gnathochilarium, mentum G gnathochilarium, stipites H, I right midbody legs Į right vulva, posterior view K right vulva, lateral view L anterior gonopod, anterior view M right posterior gonopod, anterior view N anterior gonopod, posterior view O right posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Go = gonopods; h = head; IL = incisura lateralis; L = ledges; Md = mandible; Me = mentum; O = operculum; Pre = preanal ring; pv = posterior valve; St = sternite; St-Cx = sterno-coxite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767348" httpUri="https://zenodo.org/record/3767348/files/figure.png" pageId="103" pageNumber="104">Fig. 55M</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="103.[140,1108,166,1073]" pageId="103" pageNumber="104">
|
||
<emphasis bold="true" box="[185,457,694,720]" pageId="103" pageNumber="104">
|
||
<emphasis box="[185,333,694,720]" inLineHeading="true" pageId="103" pageNumber="104" reason="1">Intraspecific</emphasis>
|
||
variation:
|
||
</emphasis>
|
||
coloration of the posterior margin of body ring varies in adults between white and reddish, while it is always white in imm..
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="103" pageNumber="104" type="distribution">
|
||
<paragraph blockId="103.[140,1108,166,1073]" pageId="103" pageNumber="104">
|
||
<emphasis bold="true" box="[185,501,765,791]" pageId="103" pageNumber="104">
|
||
<emphasis box="[185,336,765,791]" inLineHeading="true" pageId="103" pageNumber="104" reason="1">Distribution</emphasis>
|
||
and ecology
|
||
</emphasis>
|
||
: this species was found in the isolated Ambatotsirongorongo Mountain, the Andohahela Isaka-Ivondro corridor as well as on the western slopes of the Vohimena Mountain (
|
||
<figureCitation box="[655,743,835,862]" captionStart="Figure 54" captionStartId="97.[138,210,1653,1675]" captionTargetBox="[130,1107,1100,1636]" captionTargetId="figure@97.[138,1107,1109,1633]" captionTargetPageId="97" captionText="Figure 54. Distribution map of the genus Hylekobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3767346" httpUri="https://zenodo.org/record/3767346/files/figure.png" pageId="103" pageNumber="104">Fig. 54</figureCitation>
|
||
).
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[767,984,835,861]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[767,984,835,861]" italics="true" pageId="103" pageNumber="104">H. brachiosauroides</emphasis>
|
||
</taxonomicName>
|
||
could not be collected north of the Isaka-Ivondro corridor, where
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[793,939,871,897]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[793,939,871,897]" italics="true" pageId="103" pageNumber="104">H. goodmani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[950,1014,871,897]" pageId="103" pageNumber="104" rank="species">sp. n.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis italics="true" pageId="103" pageNumber="104">H. montanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[253,314,906,932]" pageId="103" pageNumber="104" rank="species">sp. n.</taxonomicNameLabel>
|
||
occur. In the eastern Vohimena slopes,
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[763,855,906,932]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[763,855,906,932]" italics="true" pageId="103" pageNumber="104">H. rufus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[862,922,906,932]" pageId="103" pageNumber="104" rank="species">sp. n.</taxonomicNameLabel>
|
||
replaces
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species">
|
||
<emphasis italics="true" pageId="103" pageNumber="104">H. brachiosauroides</emphasis>
|
||
</taxonomicName>
|
||
. All individuals were collected inside dead logs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="103" pageNumber="104" type="etymology">
|
||
<paragraph blockId="103.[140,1108,166,1073]" pageId="103" pageNumber="104">
|
||
<emphasis bold="true" box="[185,323,976,1002]" pageId="103" pageNumber="104">
|
||
<emphasis box="[185,318,976,1002]" inLineHeading="true" pageId="103" pageNumber="104" reason="1">Etymology</emphasis>
|
||
:
|
||
</emphasis>
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[330,507,976,1002]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[330,507,976,1002]" italics="true" pageId="103" pageNumber="104">brachiosauroides</emphasis>
|
||
</taxonomicName>
|
||
, adjective, refers to the dinosaur genus
|
||
<taxonomicName box="[949,1100,976,1002]" class="Reptilia" family="Brachiosauridae" genus="Brachiosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="103" pageNumber="104" phylum="Chordata" rank="genus">
|
||
<emphasis box="[949,1100,976,1002]" italics="true" pageId="103" pageNumber="104">Brachiosaurus</emphasis>
|
||
</taxonomicName>
|
||
, whose neck and head closely resemble the general shape of the posterior gonopods of this millipede.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |