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<document id="76FB3F351B3071C8CBA1D5FC19F45F62" ID-CLB-Dataset="22089" ID-DOI="10.5852/ejt.2017.330" ID-GBIF-Dataset="14d5ad89-9e69-45f8-bc61-e14ca9deafbd" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829744" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1589572621568" checkinUser="carolina" docAuthor="Inkhavilay, Khamla, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2017" docId="03DAD35F6904EF2BFEDDAB97FAAC26E1" docLanguage="en" docName="ejt-2017-330.pdf.imf" docOrigin="European Journal of Taxonomy 330" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Amphidromus (Syndromus) xiengkhaungensis Inkhavilay &amp; Sutcharit &amp; Panha 2017, sp. nov." docType="treatment" docUuid="7BA4B989-3003-4F09-94E5-F21B392BEA28" docUuidSource="ZooBank" docVersion="9" lastPageNumber="36" masterDocId="FFE3AB276926EF08FF8BAF3EFFC82563" masterDocTitle="Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species" masterLastPageNumber="40" masterPageNumber="1" pageNumber="35" updateTime="1698839926330" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="664576FAB04D98BA60B0DECF8ED30331">Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species</mods:title>
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<mods:namePart id="471A7E67E28D52025B06CA133FB68DED">Inkhavilay, Khamla</mods:namePart>
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<mods:affiliation id="9C1862E6B732A9BFAA2D4262005D67F5">Email: kinkhavilay @ gmail. com &amp; urn: lsid: zoobank. org: author: 2 E 620 B 9 C- 48 BA- 48 E 6 - A 017 - FE 8 ED 2 AB 0 E 59</mods:affiliation>
<mods:nameIdentifier id="9F4678BE6B15F761533F51568A9CF7A3" type="email">kinkhavilay@gmail.com</mods:nameIdentifier>
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<mods:namePart id="CFFEAD1004DB5CE9BBF3B1D3E8F087A7">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="7E6AE2D1D922448AD4E9AE9E86E6CC54">urn: lsid: zoobank. org: author: C 2 E 2 FA 6 B-A 3 F 9 - 4 F 33 - B 447 - B 59 B 1 BD 322 D 4 &amp; Corresponding authors: jirasak 4 @ yahoo. com; somsak. pan @ chula. ac. th</mods:affiliation>
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<mods:namePart id="D3A31AB4593106E9C93A2C364425EBFF">Panha, Somsak</mods:namePart>
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<mods:affiliation id="C0CE4711866D058C20472D77D36F7872">urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79 &amp; Corresponding authors: jirasak 4 @ yahoo. com; somsak. pan @ chula. ac. th</mods:affiliation>
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<mods:title id="C7EB90791AC48A6B4A5EF36E075AF102">European Journal of Taxonomy</mods:title>
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<treatment id="03DAD35F6904EF2BFEDDAB97FAAC26E1" ID-DOI="http://doi.org/10.5281/zenodo.3848189" ID-GBIF-Taxon="164161070" ID-Zenodo-Dep="3848189" LSID="urn:lsid:zoobank.org:act:7BA4B989-3003-4F09-94E5-F21B392BEA28" httpUri="http://treatment.plazi.org/id/03DAD35F6904EF2BFEDDAB97FAAC26E1" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<subSubSection id="C36931C26904EF2AFEDDAB97FB1621A7" box="[342,1246,1193,1220]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="8BCC62496904EF2AFEDDAB97FB1621A7" blockId="34.[342,1246,1193,1291]" box="[342,1246,1193,1220]" pageId="34" pageNumber="35">
<heading id="D084D5256904EF2AFEDDAB97FB1621A7" box="[342,1246,1193,1220]" centered="true" fontSize="11" level="2" pageId="34" pageNumber="35" reason="2">
<taxonomicName id="4C7319CA6904EF2AFEDDAB97FBB421A7" ID-CoL="7Q27F" authority="Inkhavilay &amp; Panha" authorityName="Inkhavilay &amp; Sutcharit &amp; Panha" authorityYear="2017" box="[342,1148,1193,1220]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="xiengkhaungensis" status="sp. nov." subGenus="Syndromus">
<emphasis id="B907BE5B6904EF2AFEDDAB97FC4221A7" bold="true" box="[342,906,1193,1220]" pageId="34" pageNumber="35">
<emphasis id="B907BE5B6904EF2AFEDDAB97FDCB21A7" bold="true" box="[342,515,1193,1220]" italics="true" pageId="34" pageNumber="35">Amphidromus</emphasis>
(
<emphasis id="B907BE5B6904EF2AFD9FAB97FD5521A7" bold="true" box="[532,669,1193,1220]" italics="true" pageId="34" pageNumber="35">Syndromus</emphasis>
)
<emphasis id="B907BE5B6904EF2AFD25AB97FC4221A7" bold="true" box="[686,906,1193,1220]" italics="true" pageId="34" pageNumber="35">xiengkhaungensis</emphasis>
</emphasis>
Inkhavilay &amp; Panha
</taxonomicName>
<taxonomicNameLabel id="A23403206904EF2AFB08AB94FB1621A7" box="[1155,1246,1194,1220]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C36931C26904EF2AFEFFABF3FC572068" pageId="34" pageNumber="35" type="description">
<paragraph id="8BCC62496904EF2AFEFFABF3FB08218B" blockId="34.[342,1246,1193,1291]" box="[372,1216,1229,1256]" pageId="34" pageNumber="35">
<uri id="FFE26E4B6904EF2AFEFFABF3FB08218B" box="[372,1216,1229,1256]" pageId="34" pageNumber="35">
urn:lsid:zoobank.org:act:
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</paragraph>
<paragraph id="8BCC62496904EF2AFD1EABCEFC572068" blockId="34.[342,1246,1193,1291]" box="[661,927,1264,1291]" pageId="34" pageNumber="35">
<figureCitation id="13487ECC6904EF2AFD1EABCEFD162068" box="[661,734,1264,1291]" captionStart="Fig" captionStartId="4.[189,232,1991,2017]" captionTargetBox="[416,1172,981,1949]" captionTargetId="figure@4.[416,1172,981,1949]" captionTargetPageId="4" captionText="Fig. 1. Distribution map of Amphidromus spp. in Laos recognized in this study." figureDoi="http://doi.org/10.5281/zenodo.3829746" httpUri="https://zenodo.org/record/3829746/files/figure.png" pageId="34" pageNumber="35">Figs 1</figureCitation>
,
<figureCitation id="13487ECC6904EF2AFD67ABCFFCCF2068" box="[748,775,1265,1291]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="34" pageNumber="35">13</figureCitation>
S–T;
<tableCitation id="C6F157F26904EF2AFCC3ABCEFC572068" box="[840,927,1264,1291]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionTargetId="graphics@3.[189,1398,1154,1193]" captionTargetPageId="3" captionText="Table 1 (continued on next page). Shell measurements of nine species of Amphidromus from Laos. Specimen catalogue numbers are indicated in parentheses." httpUri="http://table.plazi.org/id/DF0C32C16925EF0BFF36AE32FCA1242B" pageId="34" pageNumber="35" tableUuid="DF0C32C16925EF0BFF36AE32FCA1242B">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36931C26904EF2AFF36AA13FD6B23F2" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="8BCC62496904EF2AFF36AA13FEFE2024" blockId="34.[189,310,1325,1351]" box="[189,310,1325,1351]" pageId="34" pageNumber="35">
<heading id="D084D5256904EF2AFF36AA13FEFE2024" bold="true" box="[189,310,1325,1351]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B907BE5B6904EF2AFF36AA13FEFE2024" bold="true" box="[189,310,1325,1351]" pageId="34" pageNumber="35">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC62496904EF2AFF36AA62FD6B23F2" blockId="34.[189,1399,1372,1681]" pageId="34" pageNumber="35">
The new species is superficially similar to
<taxonomicName id="4C7319CA6904EF2AFD3EAA62FCFA2015" baseAuthorityName="Pfeiffer" baseAuthorityYear="1863" box="[693,818,1372,1398]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="roemeri">
<emphasis id="B907BE5B6904EF2AFD3EAA62FCFA2015" box="[693,818,1372,1398]" italics="true" pageId="34" pageNumber="35">A. roemeri</emphasis>
</taxonomicName>
, but the distinguishing characters are a relatively large and thick shell, dark to brownish spire, Bands 2 and 3 merged and become tinted pink stained, with a pale tinted pink lip. In contrast,
<taxonomicName id="4C7319CA6904EF2AFDF7AA9DFD3020DE" baseAuthorityName="Pfeiffer" baseAuthorityYear="1863" box="[636,760,1443,1469]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="roemeri">
<emphasis id="B907BE5B6904EF2AFDF7AA9DFD3020DE" box="[636,760,1443,1469]" italics="true" pageId="34" pageNumber="35">A. roemeri</emphasis>
</taxonomicName>
shows a relatively small and thin shell, whitish ground color, Bands 2 and 3 absent, and has a white lip.
<taxonomicName id="4C7319CA6904EF2AFC89AAF8FBB72083" authority="Inkhavilay &amp; Sutcharit &amp; Panha, 2017" authorityName="Inkhavilay &amp; Sutcharit &amp; Panha" authorityYear="2017" box="[770,1151,1478,1504]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="xiengkhaungensis" status="sp. nov.">
<emphasis id="B907BE5B6904EF2AFC89AAF8FBB72083" box="[770,1151,1478,1504]" italics="true" pageId="34" pageNumber="35">Amphidromus xiengkhaungensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23403206904EF2AFB0DAAF9FB2A2082" box="[1158,1250,1479,1505]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C7319CA6904EF2AFF36AAD4FEF32360" baseAuthorityName="Benson" baseAuthorityYear="1851" box="[189,315,1514,1540]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="sinensis">
<emphasis id="B907BE5B6904EF2AFF36AAD4FEF32360" box="[189,315,1514,1540]" italics="true" pageId="34" pageNumber="35">A. sinensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7319CA6904EF2AFEFFAAD4FE112367" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[372,473,1514,1540]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="flavus">
<emphasis id="B907BE5B6904EF2AFEFFAAD4FE112367" box="[372,473,1514,1540]" italics="true" pageId="34" pageNumber="35">A. flavus</emphasis>
</taxonomicName>
by having an ovated conic shell, Bands 2 and 3 stained with tinted pink, ovate aperture and slightly angular last whorl. In comparison,
<taxonomicName id="4C7319CA6904EF2AFCFAA933FC3B2344" baseAuthorityName="Benson" baseAuthorityYear="1851" box="[881,1011,1549,1575]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="sinensis">
<emphasis id="B907BE5B6904EF2AFCFAA933FC3B2344" box="[881,1011,1549,1575]" italics="true" pageId="34" pageNumber="35">A. sinensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7319CA6904EF2AFBBFA933FB552344" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[1076,1181,1549,1575]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="flavus">
<emphasis id="B907BE5B6904EF2AFBBFA933FB552344" box="[1076,1181,1549,1575]" italics="true" pageId="34" pageNumber="35">A. flavus</emphasis>
</taxonomicName>
have an elongate conical, relatively small and thin shell, with a yellowish color and rounded last whorl. In addition,
<taxonomicName id="4C7319CA6904EF2AFF36A96AFEA0230D" authorityName="Sutcharit &amp; Panha" authorityYear="2015" box="[189,360,1620,1646]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="globonevilli">
<emphasis id="B907BE5B6904EF2AFF36A96AFEA0230D" box="[189,360,1620,1646]" italics="true" pageId="34" pageNumber="35">A. globonevilli</emphasis>
</taxonomicName>
differs from this new species by having a small and thin shell, with a yellowish color, last whorl rounded and Bands 2 and 3 absent.
</paragraph>
</subSubSection>
<subSubSection id="C36931C26904EF2AFF36A98AFB68239E" box="[189,1184,1716,1789]" pageId="34" pageNumber="35" type="etymology">
<paragraph id="8BCC62496904EF2AFF36A98AFE8C23AD" blockId="34.[189,324,1716,1742]" box="[189,324,1716,1742]" pageId="34" pageNumber="35">
<heading id="D084D5256904EF2AFF36A98AFE8C23AD" bold="true" box="[189,324,1716,1742]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B907BE5B6904EF2AFF36A98AFE8C23AD" bold="true" box="[189,324,1716,1742]" pageId="34" pageNumber="35">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC62496904EF2AFF36A9DDFB68239E" blockId="34.[189,1184,1763,1789]" box="[189,1184,1763,1789]" pageId="34" pageNumber="35">
The specific epithet ‘
<taxonomicName id="4C7319CA6904EF2AFE3FA9DDFD4D239E" authorityName="Inkhavilay &amp; Sutcharit &amp; Panha" authorityYear="2017" box="[436,645,1763,1789]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="xiengkhaungensis">
<emphasis id="B907BE5B6904EF2AFE3FA9DDFD4D239E" box="[436,645,1763,1789]" italics="true" pageId="34" pageNumber="35">xiengkhaungensis</emphasis>
</taxonomicName>
’ refers to the
<typeStatus id="54C8DCEB6904EF2AFCACA9DDFC92239E" box="[807,858,1763,1789]" pageId="34" pageNumber="35">type</typeStatus>
locality of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C36931C26904EF2AFF36A81EFC08228F" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="8BCC62496904EF2AFF36A81EFEA4225A" blockId="34.[189,364,1823,1850]" box="[189,364,1823,1850]" pageId="34" pageNumber="35">
<heading id="D084D5256904EF2AFF36A81EFEA4225A" bold="true" box="[189,364,1823,1850]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B907BE5B6904EF2AFF36A81EFEA4225A" bold="true" box="[189,364,1823,1850]" pageId="34" pageNumber="35">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B1B68146904EF2AFF61A870FB2322EF" ID-GBIF-Occurrence="2620312303" collectionCode="CUMZ" country="Laos" location="LAOS" pageId="34" pageNumber="35" specimenCode="CUMZ 7045" specimenCount="1" typeStatus="holotype">
<paragraph id="8BCC62496904EF2AFF61A870FE94220B" blockId="34.[189,1259,1870,1932]" box="[234,348,1870,1896]" pageId="34" pageNumber="35">
<emphasis id="B907BE5B6904EF2AFF61A870FE94220B" bold="true" box="[234,348,1870,1896]" pageId="34" pageNumber="35">
<typeStatus id="54C8DCEB6904EF2AFF61A870FE94220B" box="[234,348,1870,1896]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCC62496904EF2AFF36A84CFB2322EF" blockId="34.[189,1259,1870,1932]" box="[189,1259,1905,1932]" pageId="34" pageNumber="35">
<collectingCountry id="F36422D96904EF2AFF36A84CFEC622EF" box="[189,270,1906,1932]" name="Laos" pageId="34" pageNumber="35">LAOS</collectingCountry>
: shell height
<quantity id="4C8BCFAC6904EF2AFE23A84FFDDC22EF" box="[424,532,1905,1932]" metricMagnitude="-2" metricUnit="m" metricValue="2.56" pageId="34" pageNumber="35" unit="mm" value="25.6">25.6 mm</quantity>
, shell width
<quantity id="4C8BCFAC6904EF2AFD23A84FFCDC22EF" box="[680,788,1905,1932]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="34" pageNumber="35" unit="mm" value="15.6">15.6 mm</quantity>
, with 6 whorls (
<specimenCode id="DBD5CA326904EF2AFC5BA84CFBA422EF" box="[976,1132,1905,1932]" collectionCode="CUMZ" country="0" name="Cameroon University, Museum of Zoology" pageId="34" pageNumber="35">CUMZ 7045</specimenCode>
,
<figureCitation id="13487ECC6904EF2AFBFCA84FFB1522EF" box="[1143,1245,1905,1932]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="34" pageNumber="35">Fig. 13S</figureCitation>
).
</paragraph>
</materialsCitation>
<paragraph id="8BCC62496904EF2AFF61A891FEAF22AA" blockId="34.[189,960,1967,2028]" box="[234,359,1967,1993]" pageId="34" pageNumber="35">
<emphasis id="B907BE5B6904EF2AFF61A891FEAF22AA" bold="true" box="[234,359,1967,1993]" pageId="34" pageNumber="35">
<typeStatus id="54C8DCEB6904EF2AFF61A891FEAF22AA" box="[234,359,1967,1993]" pageId="34" pageNumber="35" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCC62496904EF2AFF36A8ECFC08228F" blockId="34.[189,960,1967,2028]" box="[189,960,2002,2028]" pageId="34" pageNumber="35">
<collectingCountry id="F36422D96904EF2AFF36A8ECFEC6228F" box="[189,270,2002,2028]" name="Laos" pageId="34" pageNumber="35">LAOS</collectingCountry>
: 10S shells (
<collectionCode id="ED62FA8C6904EF2AFE2BA8ECFE3F228F" box="[416,503,2002,2028]" country="0" name="Cameroon University, Museum of Zoology" pageId="34" pageNumber="35">CUMZ</collectionCode>
7046,
<figureCitation id="13487ECC6904EF2AFDCCA8ECFD78228F" box="[583,688,2002,2028]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="34" pageNumber="35">Fig. 13T</figureCitation>
); 2S shells (
<collectionCode id="ED62FA8C6904EF2AFCB5A8ECFC7C228F" box="[830,948,2002,2028]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="34" pageNumber="35" type="Museum">NHMUK</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C36931C26905EF2BFF36AE35FDD6241A" pageId="35" pageNumber="36" type="distribution">
<paragraph id="8BCC62496905EF2BFF36AE35FE952446" blockId="35.[189,349,267,293]" box="[189,349,267,293]" pageId="35" pageNumber="36">
<heading id="D084D5256905EF2BFF36AE35FE952446" bold="true" box="[189,349,267,293]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B907BE5B6905EF2BFF36AE35FE952446" bold="true" box="[189,349,267,293]" pageId="35" pageNumber="36">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC62496905EF2BFF36AE05FDD6241A" blockId="35.[189,1399,313,378]" pageId="35" pageNumber="36">
<collectingCountry id="F36422D96905EF2BFF36AE05FEC62436" box="[189,270,315,341]" name="Laos" pageId="35" pageNumber="36">LAOS</collectingCountry>
: limestone outcrop at Ban Nong Tang, Phou Kood District, Xieng Khaung,
<geoCoordinate id="EE47048E6905EF2BFB4AAE05FABA2436" box="[1217,1394,313,341]" degrees="19" direction="north" minutes="30" orientation="latitude" pageId="35" pageNumber="36" precision="1" seconds="59.2" value="19.516445">19°30′59.2″ N</geoCoordinate>
,
<geoCoordinate id="EE47048E6905EF2BFF36AE61FEBB2419" box="[189,371,349,378]" degrees="102" direction="east" minutes="53" orientation="longitude" pageId="35" pageNumber="36" precision="1" seconds="37.6" value="102.893776">102°53′37.6″ E</geoCoordinate>
,
<quantity id="4C8BCFAC6905EF2BFE0BAE61FE102419" box="[384,472,351,378]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="35" pageNumber="36" unit="m" value="1140.0">1140 m</quantity>
amsl.
</paragraph>
</subSubSection>
<subSubSection id="C36931C26905EF2BFF36AE9CFD4627A4" pageId="35" pageNumber="36" type="description">
<paragraph id="8BCC62496905EF2BFF36AE9CFE8624DF" blockId="35.[189,334,418,444]" box="[189,334,418,444]" pageId="35" pageNumber="36">
<heading id="D084D5256905EF2BFF36AE9CFE8624DF" bold="true" box="[189,334,418,444]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B907BE5B6905EF2BFF36AE9CFE8624DF" bold="true" box="[189,334,418,444]" pageId="35" pageNumber="36">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC62496905EF2BFF36AEECFD4627A4" blockId="35.[189,1399,466,711]" pageId="35" pageNumber="36">Shell sinistral, ovate conical, rather thick and glossy. Apex acute, color tinted pink; spire conical with dark color; suture wide and depressed. Whorls 5 and 6 nearly smooth; last whorl slightly angular. Periostracum thin, corneous and transparent. Earlier whorls with darker color; varix absent. Shell banding without Band 1; Bands 2 and 3 merged and becoming tinted pink stained; Bands 4 and 5 dark brown; Band 6 absent. Parietal callus thin and transparent to slightly thick and white. Aperture ovate; peristome expanded and weakly thickened; lip pale tinted pink to white. Columella straight, thickend and white. Umbilicus narrowly opened.</paragraph>
</subSubSection>
<subSubSection id="C36931C26905EF2BFF36ADCEFAAC26E1" pageId="35" pageNumber="36" type="discussion">
<paragraph id="8BCC62496905EF2BFF36ADCEFEE72669" blockId="35.[189,303,752,778]" box="[189,303,752,778]" pageId="35" pageNumber="36">
<heading id="D084D5256905EF2BFF36ADCEFEE72669" bold="true" box="[189,303,752,778]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B907BE5B6905EF2BFF36ADCEFEE72669" bold="true" box="[189,303,752,778]" pageId="35" pageNumber="36">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC62496905EF2BFF36AC21FAAC26E1" blockId="35.[189,1399,799,899]" pageId="35" pageNumber="36">
<taxonomicName id="4C7319CA6905EF2BFF36AC21FDF4265A" authority="Inkhavilay &amp; Sutcharit &amp; Panha, 2017" authorityName="Inkhavilay &amp; Sutcharit &amp; Panha" authorityYear="2017" box="[189,572,799,825]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="xiengkhaungensis" status="sp. nov.">
<emphasis id="B907BE5B6905EF2BFF36AC21FDF4265A" box="[189,572,799,825]" italics="true" pageId="35" pageNumber="36">Amphidromus xiengkhaungensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23403206905EF2BFDCEAC1EFD6B2659" box="[581,675,800,826]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
is currently known from the
<typeStatus id="54C8DCEB6905EF2BFB81AC1EFBF52659" box="[1034,1085,800,826]" pageId="35" pageNumber="36">type</typeStatus>
locality only, a dry green forest and isolated limestone forest surrounded by a reservoir and agricultural areas. The empty shells were found on the floor among the leaf litter. Unfortunately, no living specimens have been collected.
</paragraph>
</subSubSection>
</treatment>
</document>
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