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<document id="CC61FC31B546F293E1232C2466759573" ID-CLB-Dataset="306039" ID-DOI="10.11646/zootaxa.5537.1.8" ID-GBIF-Dataset="3cc04113-3025-4a16-b612-576f862c0ca0" ID-ISSN="1175-5326" ID-Zenodo-Dep="14239399" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1732800215542" checkinUser="plazi" docAuthor="Zhang, Haibin, Yu, Kun &amp; Zhang, Feng" docDate="2024" docId="035B87AC76355F28FF11F8C67404CB64" docLanguage="en" docName="zootaxa.5537.1.8.pdf" docOrigin="Zootaxa 5537 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5537.1.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Shinobius yulong Zhang, Yu &amp; Zhang 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="146" masterDocId="FF62FFD476355F2DFF86FF897608C95F" masterDocTitle="The third species of the genus Shinobius Yaginuma, 1991 from Asia (Araneae, Trechaleidae)" masterLastPageNumber="146" masterPageNumber="141" pageNumber="141" updateTime="1733409733185" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="CD2AA827BCB161905A8E4D45CAE6ADCC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="3EC3FC3B319EAB78B48A5BAA52829899">The third species of the genus Shinobius Yaginuma, 1991 from Asia (Araneae, Trechaleidae)</mods:title>
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<mods:namePart id="1BA1C00855C70E41A34FC99E814EF049">Zhang, Haibin</mods:namePart>
<mods:nameIdentifier id="84ADC58BD22208A161EF441E95F2C067" type="ORCID">0009-0007-6642-9175</mods:nameIdentifier>
<mods:affiliation id="775E7B41BC59DBA0C04E73D6403A43AA">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China &amp; Hebei Basic Science Center for Biotic Interaction, Hebei University, Baoding, Hebei 071002, P. R. China</mods:affiliation>
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<mods:namePart id="A2F95275431DECBE627F90DAB54BFEFC">Yu, Kun</mods:namePart>
<mods:nameIdentifier id="EB8B57056278084DB611D499B5A4325E" type="ORCID">0000-0002-5517-4484</mods:nameIdentifier>
<mods:affiliation id="0B24739A9953AD0F13CAC053F1DF5E6F">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China &amp; Hebei Basic Science Center for Biotic Interaction, Hebei University, Baoding, Hebei 071002, P. R. China</mods:affiliation>
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<mods:namePart id="69E1794E95C62318D6F08C70196DF691">Zhang, Feng</mods:namePart>
<mods:nameIdentifier id="B57BF13371490289D76EEA901C380B20" type="ORCID">0000-0002-3347-1031</mods:nameIdentifier>
<mods:affiliation id="4C1618A05920DE9CB6C50DABF9FEB12B">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China &amp; Hebei Basic Science Center for Biotic Interaction, Hebei University, Baoding, Hebei 071002, P. R. China</mods:affiliation>
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<mods:title id="6BBA6B61F214F2C8D997922E89859598">Zootaxa</mods:title>
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<mods:date id="EC9D3CB4620962F77AE18AF1EEE92A01">2024</mods:date>
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<mods:number id="3FFFBFAD9343A255A67516DD9732CEA0">2024-11-07</mods:number>
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<treatment id="035B87AC76355F28FF11F8C67404CB64" ID-DOI="http://doi.org/10.5281/zenodo.14248069" ID-Zenodo-Dep="14248069" LSID="urn:lsid:plazi:treatment:035B87AC76355F28FF11F8C67404CB64" httpUri="http://treatment.plazi.org/id/035B87AC76355F28FF11F8C67404CB64" lastPageId="5" lastPageNumber="146" pageId="0" pageNumber="141">
<subSubSection id="C3E8653176355F2DFF11F8C6756BCE36" box="[151,867,1871,1898]" pageId="0" pageNumber="141" type="nomenclature">
<paragraph id="8B4D36BA76355F2DFF11F8C6756BCE36" blockId="0.[151,1437,1763,2041]" box="[151,867,1871,1898]" pageId="0" pageNumber="141">
<heading id="D00581D676355F2DFF11F8C6756BCE36" bold="true" box="[151,867,1871,1898]" fontSize="11" level="1" pageId="0" pageNumber="141" reason="1">
<emphasis id="B986EAA876355F2DFF11F8C6756BCE36" bold="true" box="[151,867,1871,1898]" pageId="0" pageNumber="141">
<taxonomicName id="4CF24D3976355F2DFF11F8C67466CE36" authority="Zhang, Yu &amp; Zhang" authorityName="Zhang, Yu &amp; Zhang" authorityYear="2024" box="[151,622,1871,1898]" class="Arachnida" family="Trechaleidae" genus="Shinobius" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="yulong" status="sp. nov.">
<emphasis id="B986EAA876355F2DFF11F8C67764CE35" bold="true" box="[151,364,1871,1898]" italics="true" pageId="0" pageNumber="141">Shinobius yulong</emphasis>
Zhang, Yu &amp; Zhang
</taxonomicName>
<taxonomicNameLabel id="A2B557D376355F2DFDF3F8C674DBCE36" box="[629,723,1871,1897]" pageId="0" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
(ṮŽ侵Ʀ)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3E8653176355F2DFF11F8FC76FCCED2" box="[151,244,1909,1933]" pageId="0" pageNumber="141" type="description">
<paragraph id="8B4D36BA76355F2DFF11F8FC76FCCED2" blockId="0.[151,1437,1763,2041]" box="[151,244,1909,1933]" pageId="0" pageNumber="141">
<figureCitation id="13C92A3F76355F2DFF11F8FC76FCCED2" box="[151,244,1909,1933]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="1.[152,255,921,945]" captionStartId-1="2.[152,255,1810,1834]" captionStartId-2="3.[152,255,1943,1967]" captionStartId-3="4.[152,255,1031,1055]" captionStartId-4="4.[152,255,1958,1982]" captionTargetBox-0="[151,1436,181,898]" captionTargetBox-1="[151,1436,181,1787]" captionTargetBox-2="[208,1380,184,1914]" captionTargetBox-3="[155,1428,208,1002]" captionTargetBox-4="[151,1436,1187,1934]" captionTargetId-0="figure-19@1.[151,1436,181,898]" captionTargetId-1="figure-22@2.[151,1436,181,1787]" captionTargetId-2="figure-19@3.[203,1384,181,1919]" captionTargetId-3="figure-22@4.[151,1436,199,1007]" captionTargetId-4="figure-81@4.[151,1436,1187,1934]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="4" captionText-0="FIGURE 1. Shinobius yulong sp. nov., male holotype (A, D, F) and female paratype (B, C, E). A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Female habitus, ventral view; D. Male chelicerae, ventral view; E. Female chelicerae, ventral view; F. Male endites, labium and sternum, ventral view." captionText-1="FIGURE 2. Shinobius yulong sp. nov., male left palp in ventral (A) and retrolateral (B) views.Abbreviations: C, conductor; E, embolus; H, hood; MA, median apophysis; SD, sperm duct; ST, subtegulum; T, tegulum." captionText-2="FIGURE 3. Shinobius yulong sp. nov., palpal bulb in retrolateral (A), dorsal (B), ventral (C) and prolateral (D) views. Abbreviations: C, conductor; E, embolus; MA, median apophysis; SD, sperm duct; ST, subtegulum; T, tegulum." captionText-3="FIGURE 4. Shinobius yulong sp. nov., female epigyne (A, C, D), and vulva (B), in ventral (A), dorsal (B), posterior (C) and anterior (D) views. Abbreviations: AS, accessory spermatheca; CO, copulatory opening; FD, fertilization duct; HS, head of spermatheca; MP, median plate; SS, stalk of spermatheca." captionText-4="FIGURE 5. Distribution records of Shinobius cona and S. Yulong sp. nov. in China." figureDoi-0="http://doi.org/10.5281/zenodo.14239401" figureDoi-1="http://doi.org/10.5281/zenodo.14239403" figureDoi-2="http://doi.org/10.5281/zenodo.14239405" figureDoi-3="http://doi.org/10.5281/zenodo.14239407" figureDoi-4="http://doi.org/10.5281/zenodo.14239409" httpUri-0="https://zenodo.org/record/14239401/files/figure.png" httpUri-1="https://zenodo.org/record/14239403/files/figure.png" httpUri-2="https://zenodo.org/record/14239405/files/figure.png" httpUri-3="https://zenodo.org/record/14239407/files/figure.png" httpUri-4="https://zenodo.org/record/14239409/files/figure.png" pageId="0" pageNumber="141">Figs 1–5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3E8653176355F2CFF11F81075CCCD39" lastPageId="1" lastPageNumber="142" pageId="0" pageNumber="141" type="materials_examined">
<paragraph id="8B4D36BA76355F2DFF11F81076EBCEA6" blockId="0.[151,1437,1763,2041]" pageId="0" pageNumber="141">
Type material.
<materialsCitation id="3B9A3CE776355F2DFEC0F81076EBCEA6" collectingDate="2023-07-16" collectorName="Xiangbo Guo &amp; Haibin Zhang." country="China" county="Dingkye County" elevation="2272" latitude="27.874842" location="Zhentang Town" longLatPrecision="1" longitude="87.415215" municipality="Autonomous Region" pageId="0" pageNumber="141" specimenCode="MHBU-ARA-2023-814-1" specimenCount="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="holotype">
<emphasis id="B986EAA876355F2DFEC0F81077A6CEEE" bold="true" box="[326,430,1945,1969]" pageId="0" pageNumber="141">
<typeStatus id="5449881876355F2DFEC0F81077A6CEEE" box="[326,430,1945,1969]" pageId="0" pageNumber="141" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9DF4FD3376355F2DFE30F81377C2CEEE" box="[438,458,1946,1969]" count="1" pageId="0" pageNumber="141" type="male">♂</specimenCount>
(
<specimenCode id="DB549EC176355F2DFE5CF81074FDCEEE" box="[474,757,1945,1969]" pageId="0" pageNumber="141">MHBU-ARA-2023-814-1</specimenCode>
),
<emphasis id="B986EAA876355F2DFC8FF810756ACEEE" bold="true" box="[777,866,1945,1969]" pageId="0" pageNumber="141">
<collectingCountry id="F3E5762A76355F2DFC8FF810756ACEEE" box="[777,866,1945,1969]" name="China" pageId="0" pageNumber="141">CHINA</collectingCountry>
</emphasis>
:
<collectingRegion id="4936F85876355F2DFCF4F81075B7CEEE" box="[882,959,1945,1969]" country="China" name="Xizang" pageId="0" pageNumber="141">Xizang</collectingRegion>
<collectingMunicipality id="6B29ACC076355F2DFC43F81072AFCEEE" box="[965,1191,1945,1969]" pageId="0" pageNumber="141">Autonomous Region</collectingMunicipality>
,
<collectingMunicipality id="6B29ACC076355F2DFB35F810733CCEEE" box="[1203,1332,1945,1969]" pageId="0" pageNumber="141">Rigaze City</collectingMunicipality>
,
<collectingCounty id="622C4E3676355F2DFAC6F81076EFCE8A" pageId="0" pageNumber="141">Dingkye County</collectingCounty>
,
<location id="8E2D606176355F2DFF72F8347795CE8A" LSID="urn:lsid:plazi:treatment:035B87AC76355F28FF11F8C67404CB64:8E2D606176355F2DFF72F8347795CE8A" box="[244,413,1981,2005]" country="China" county="Dingkye County" latitude="27.874842" longLatPrecision="1" longitude="87.415215" municipality="Autonomous Region" name="Zhentang Town" pageId="0" pageNumber="141" stateProvince="Xizang">Zhentang Town</location>
(
<geoCoordinate id="EEC6507D76355F2DFE28F835745BCE8A" box="[430,595,1980,2005]" degrees="27" direction="north" minutes="52" orientation="latitude" pageId="0" pageNumber="141" precision="1" seconds="29.43" value="27.874842">27°52′29.43′′N</geoCoordinate>
,
<geoCoordinate id="EEC6507D76355F2DFDE6F8357509CE8A" box="[608,769,1980,2005]" degrees="87" direction="east" minutes="24" orientation="longitude" pageId="0" pageNumber="141" precision="1" seconds="54.78" value="87.415215">87°24′54.78′′E</geoCoordinate>
, alt.
<quantity id="4C0A9B5F76355F2DFCBFF83475ABCE8A" box="[825,931,1981,2005]" metricMagnitude="3" metricUnit="m" metricValue="2.2717" pageId="0" pageNumber="141" unit="m" value="2271.7">
<elevation id="00DFD18976355F2DFCBFF83475ABCE8A" box="[825,931,1981,2005]" metricMagnitude="3" metricUnit="m" metricValue="2.2717" pageId="0" pageNumber="141" unit="m" value="2271.7">2271.7 m</elevation>
</quantity>
),
<date id="FF4C107A76355F2DFC3FF8347242CE8A" box="[953,1098,1981,2005]" pageId="0" pageNumber="141" value="2023-07-16">
<collectingDate id="EF08E99276355F2DFC3FF8347242CE8A" box="[953,1098,1981,2005]" pageId="0" pageNumber="141" value="2023-07-16">16 July 2023</collectingDate>
</date>
, leg.
<collectorName id="2607536C76355F2DFB0EF8347313CE8A" box="[1160,1307,1981,2005]" pageId="0" pageNumber="141">Xiangbo Guo</collectorName>
and
<collectorName id="2607536C76355F2DFAD2F83476EBCEA6" pageId="0" pageNumber="141">Haibin Zhang.</collectorName>
</materialsCitation>
</paragraph>
<caption id="DF8D663276345F2CFF1EFC1074DFCAA5" ID-DOI="http://doi.org/10.5281/zenodo.14239401" ID-Zenodo-Dep="14239401" httpUri="https://zenodo.org/record/14239401/files/figure.png" pageId="1" pageNumber="142" startId="1.[152,255,921,945]" targetBox="[151,1436,181,898]" targetPageId="1" targetType="figure">
<paragraph id="8B4D36BA76345F2CFF1EFC1074DFCAA5" blockId="1.[151,1436,921,1018]" pageId="1" pageNumber="142">
<emphasis id="B986EAA876345F2CFF1EFC107711CAEE" bold="true" box="[152,281,921,945]" pageId="1" pageNumber="142">FIGURE 1.</emphasis>
<taxonomicName id="4CF24D3976345F2CFEA6FC1377C7CAED" authorityName="Zhang &amp; Yu &amp; Zhang" authorityYear="2024" box="[288,463,922,946]" class="Arachnida" family="Trechaleidae" genus="Shinobius" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="yulong" status="sp. nov.">
<emphasis id="B986EAA876345F2CFEA6FC1377C7CAED" box="[288,463,922,946]" italics="true" pageId="1" pageNumber="142">Shinobius yulong</emphasis>
</taxonomicName>
<emphasis id="B986EAA876345F2CFE50FC13742FCAED" bold="true" box="[470,551,922,946]" pageId="1" pageNumber="142">
<taxonomicNameLabel id="A2B557D376345F2CFE50FC13742FCAED" box="[470,551,922,946]" pageId="1" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (A, D, F) and female paratype (B, C, E). A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Female habitus, ventral view; D. Male chelicerae, ventral view; E. Female chelicerae, ventral view; F. Male endites, labium and sternum, ventral view.
</paragraph>
</caption>
<paragraph id="8B4D36BA76345F2CFF41FBC775CCCD39" blockId="1.[151,1437,1102,2026]" box="[199,964,1102,1126]" pageId="1" pageNumber="142">
<materialsCitation id="3B9A3CE776345F2CFF41FBC775C8CD39" box="[199,960,1102,1126]" collectingDate="2023-07-16" collectorName="Xiangbo Guo &amp; Haibin Zhang." country="China" county="Dingkye County" elevation="2272" latitude="27.874842" location="Rigaze City" longLatPrecision="1" longitude="87.415215" municipality="Autonomous Region" pageId="1" pageNumber="142" specimenCode="MHBU-ARA-2023-814-2" specimenCount="1" specimenCount-female="1" stateProvince="Xizang" typeStatus="paratype">
<emphasis id="B986EAA876345F2CFF41FBC77730CD39" bold="true" box="[199,312,1102,1126]" pageId="1" pageNumber="142">
<typeStatus id="5449881876345F2CFF41FBC7773BCD39" box="[199,307,1102,1126]" pageId="1" pageNumber="142" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9DF4FD3376345F2CFEB8FBC77768CD39" box="[318,352,1102,1126]" count="1" pageId="1" pageNumber="142" type="female">1♀</specimenCount>
(
<specimenCode id="DB549EC176345F2CFEE8FBC77482CD39" box="[366,650,1102,1126]" pageId="1" pageNumber="142">MHBU-ARA-2023-814-2</specimenCode>
), with same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3E8653176345F2CFF41FBFB74F8CDF1" pageId="1" pageNumber="142" type="etymology">
<paragraph id="8B4D36BA76345F2CFF41FBFB74F8CDF1" blockId="1.[151,1437,1102,2026]" pageId="1" pageNumber="142">
<emphasis id="B986EAA876345F2CFF41FBFB7740CDD5" bold="true" box="[199,328,1138,1162]" inLineHeading="true" pageId="1" pageNumber="142" reason="1">Etymology.</emphasis>
The specific name refers to Hongshan Jade Dragon, a national cultural relic of
<collectingCountry id="F3E5762A76345F2CFB22FBFB72EECDD5" box="[1188,1254,1138,1162]" name="China" pageId="1" pageNumber="142">China</collectingCountry>
, whose C- shape resembles this species spermatheca. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3E8653176345F2CFF41FB307366CCAD" pageId="1" pageNumber="142" type="diagnosis">
<paragraph id="8B4D36BA76345F2CFF41FB307366CCAD" blockId="1.[151,1437,1102,2026]" pageId="1" pageNumber="142">
<emphasis id="B986EAA876345F2CFF41FB307734CD8E" bold="true" box="[199,316,1209,1233]" inLineHeading="true" pageId="1" pageNumber="142" reason="1">Diagnosis.</emphasis>
The new species resembles
<taxonomicName id="4CF24D3976345F2CFDE4FB3374E9CD8D" baseAuthorityName="Yaginuma" baseAuthorityYear="1967" box="[610,737,1210,1234]" class="Arachnida" family="Trechaleidae" genus="Shinobius" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B986EAA876345F2CFDE4FB3374E9CD8D" box="[610,737,1210,1234]" italics="true" pageId="1" pageNumber="142">S. orientalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4CF24D3976345F2CFC94FB337557CD8E" authorityName="Wang, Mu, Zhang, Marusik &amp; Zhang" authorityYear="2024" box="[786,863,1210,1233]" class="Arachnida" family="Trechaleidae" genus="Shinobius" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="cona">
<emphasis id="B986EAA876345F2CFC94FB337557CD8E" box="[786,863,1210,1233]" italics="true" pageId="1" pageNumber="142">S. cona</emphasis>
</taxonomicName>
in the shape of palpal bulb and conductor, and absence of RTA (see
<bibRefCitation id="EF634B4B76345F2CFE9FFB5777BECDA9" author="Sierwald, P." box="[281,438,1246,1270]" pageId="1" pageNumber="142" pagination="53 - 74" refId="ref2223" refString="Sierwald, P. (1993) Revision of the spider genus Paradossenus, with notes on the family Trechaleidae and the subfamily Rhoicininae (Araneae, Lycosoidea). Revue Arachnologique, 10, 53 - 74." type="journal article" year="1993">Sierwald 1993</bibRefCitation>
;
<bibRefCitation id="EF634B4B76345F2CFE46FB57747ECDA9" author="Wang, L. Y. &amp; Mu, Y. N. &amp; Zhang, F. &amp; Marusik Y. M. &amp; Zhang, Z. S." box="[448,630,1246,1270]" pageId="1" pageNumber="142" pagination="189 - 195" refId="ref2262" refString="Wang, L. Y., Mu, Y. N., Zhang, F., Marusik Y. M. &amp; Zhang, Z. S. (2024) First record of the spider family Trechaleidae Simon, 1890 (Araneae) from China. Zookeys, 1203, 189 - 195. https: // doi. org / 10.3897 / zookeys. 1203.124808" type="journal article" year="2024">
Wang
<emphasis id="B986EAA876345F2CFD85FB577430CDA9" box="[515,568,1246,1270]" italics="true" pageId="1" pageNumber="142">et al.</emphasis>
2024
</bibRefCitation>
), but it can be distinguished from
<taxonomicName id="4CF24D3976345F2CFC5CFB577252CDA9" baseAuthorityName="Yaginuma" baseAuthorityYear="1967" box="[986,1114,1246,1270]" class="Arachnida" family="Trechaleidae" genus="Shinobius" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B986EAA876345F2CFC5CFB577252CDA9" box="[986,1114,1246,1270]" italics="true" pageId="1" pageNumber="142">S. orientalis</emphasis>
</taxonomicName>
by: 1) the width of the median apophysis over half that of the bulb in ventral view (
<figureCitation id="13C92A3F76345F2CFD40FA8B7513CC45" box="[710,795,1282,1306]" captionStart="FIGURE 2" captionStartId="2.[152,255,1810,1834]" captionTargetBox="[151,1436,181,1787]" captionTargetId="figure-22@2.[151,1436,181,1787]" captionTargetPageId="2" captionText="FIGURE 2. Shinobius yulong sp. nov., male left palp in ventral (A) and retrolateral (B) views.Abbreviations: C, conductor; E, embolus; H, hood; MA, median apophysis; SD, sperm duct; ST, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14239403" httpUri="https://zenodo.org/record/14239403/files/figure.png" pageId="1" pageNumber="142">Fig. 2A</figureCitation>
) vs. less than half the width (
<bibRefCitation id="EF634B4B76345F2CFBD5FA8B72FBCC45" author="Sierwald, P." box="[1107,1267,1282,1306]" pageId="1" pageNumber="142" pagination="53 - 74" refId="ref2223" refString="Sierwald, P. (1993) Revision of the spider genus Paradossenus, with notes on the family Trechaleidae and the subfamily Rhoicininae (Araneae, Lycosoidea). Revue Arachnologique, 10, 53 - 74." type="journal article" year="1993">Sierwald 1993</bibRefCitation>
: figs 20, 22); 2) copulatory openings semicircular, located at posterior edge of epigyne (
<figureCitation id="13C92A3F76345F2CFC13FAAF7205CC61" box="[917,1037,1318,1342]" captionStart="FIGURE 4" captionStartId="4.[152,255,1031,1055]" captionTargetBox="[155,1428,208,1002]" captionTargetId="figure-22@4.[151,1436,199,1007]" captionTargetPageId="4" captionText="FIGURE 4. Shinobius yulong sp. nov., female epigyne (A, C, D), and vulva (B), in ventral (A), dorsal (B), posterior (C) and anterior (D) views. Abbreviations: AS, accessory spermatheca; CO, copulatory opening; FD, fertilization duct; HS, head of spermatheca; MP, median plate; SS, stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14239407" httpUri="https://zenodo.org/record/14239407/files/figure.png" pageId="1" pageNumber="142">Figs 4A–C</figureCitation>
) vs. circular, located in the middle of epigyne (
<bibRefCitation id="EF634B4B76345F2CFF7DFAC37794CC3D" author="Sierwald, P." box="[251,412,1354,1378]" pageId="1" pageNumber="142" pagination="53 - 74" refId="ref2223" refString="Sierwald, P. (1993) Revision of the spider genus Paradossenus, with notes on the family Trechaleidae and the subfamily Rhoicininae (Araneae, Lycosoidea). Revue Arachnologique, 10, 53 - 74." type="journal article" year="1993">Sierwald 1993</bibRefCitation>
: fig. 15); 3) stalk of spermatheca bent inwards (
<figureCitation id="13C92A3F76345F2CFC25FAC375FECC3D" box="[931,1014,1354,1378]" captionStart="FIGURE 4" captionStartId="4.[152,255,1031,1055]" captionTargetBox="[155,1428,208,1002]" captionTargetId="figure-22@4.[151,1436,199,1007]" captionTargetPageId="4" captionText="FIGURE 4. Shinobius yulong sp. nov., female epigyne (A, C, D), and vulva (B), in ventral (A), dorsal (B), posterior (C) and anterior (D) views. Abbreviations: AS, accessory spermatheca; CO, copulatory opening; FD, fertilization duct; HS, head of spermatheca; MP, median plate; SS, stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14239407" httpUri="https://zenodo.org/record/14239407/files/figure.png" pageId="1" pageNumber="142">Fig. 4B</figureCitation>
) vs. bent upwards (
<bibRefCitation id="EF634B4B76345F2CFB4DFAC37364CC3D" author="Sierwald, P." box="[1227,1388,1354,1378]" pageId="1" pageNumber="142" pagination="53 - 74" refId="ref2223" refString="Sierwald, P. (1993) Revision of the spider genus Paradossenus, with notes on the family Trechaleidae and the subfamily Rhoicininae (Araneae, Lycosoidea). Revue Arachnologique, 10, 53 - 74." type="journal article" year="1993">Sierwald 1993</bibRefCitation>
: fig. 16). It differs from
<taxonomicName id="4CF24D3976345F2CFEE2FAE777BACCDA" authorityName="Wang, Mu, Zhang, Marusik &amp; Zhang" authorityYear="2024" box="[356,434,1390,1413]" class="Arachnida" family="Trechaleidae" genus="Shinobius" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="cona">
<emphasis id="B986EAA876345F2CFEE2FAE777BACCDA" box="[356,434,1390,1413]" italics="true" pageId="1" pageNumber="142">S. cona</emphasis>
</taxonomicName>
in: 1) the median apophysis divided into two branches (
<figureCitation id="13C92A3F76345F2CFB83FAE77242CCD9" box="[1029,1098,1390,1414]" captionStart="FIGURE 2" captionStartId="2.[152,255,1810,1834]" captionTargetBox="[151,1436,181,1787]" captionTargetId="figure-22@2.[151,1436,181,1787]" captionTargetPageId="2" captionText="FIGURE 2. Shinobius yulong sp. nov., male left palp in ventral (A) and retrolateral (B) views.Abbreviations: C, conductor; E, embolus; H, hood; MA, median apophysis; SD, sperm duct; ST, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14239403" httpUri="https://zenodo.org/record/14239403/files/figure.png" pageId="1" pageNumber="142">Figs 2</figureCitation>
,
<figureCitation id="13C92A3F76345F2CFBD3FAE7729ECCD9" box="[1109,1174,1390,1414]" captionStart="FIGURE 3" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[208,1380,184,1914]" captionTargetId="figure-19@3.[203,1384,181,1919]" captionTargetPageId="3" captionText="FIGURE 3. Shinobius yulong sp. nov., palpal bulb in retrolateral (A), dorsal (B), ventral (C) and prolateral (D) views. Abbreviations: C, conductor; E, embolus; MA, median apophysis; SD, sperm duct; ST, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14239405" httpUri="https://zenodo.org/record/14239405/files/figure.png" pageId="1" pageNumber="142">3A, C</figureCitation>
) vs. unbifurcated (
<bibRefCitation id="EF634B4B76345F2CFADBFAE77718CCF5" author="Wang, L. Y. &amp; Mu, Y. N. &amp; Zhang, F. &amp; Marusik Y. M. &amp; Zhang, Z. S." pageId="1" pageNumber="142" pagination="189 - 195" refId="ref2262" refString="Wang, L. Y., Mu, Y. N., Zhang, F., Marusik Y. M. &amp; Zhang, Z. S. (2024) First record of the spider family Trechaleidae Simon, 1890 (Araneae) from China. Zookeys, 1203, 189 - 195. https: // doi. org / 10.3897 / zookeys. 1203.124808" type="journal article" year="2024">
Wang
<emphasis id="B986EAA876345F2CFF11FA1B76C7CCF5" box="[151,207,1426,1450]" italics="true" pageId="1" pageNumber="142">et al.</emphasis>
2024
</bibRefCitation>
: figs 2A, 3A); 2) the outer margin of membrane between tegulum and median apophysis relatively smooth (
<figureCitation id="13C92A3F76345F2CFF19FA3F76F0CC91" box="[159,248,1462,1486]" captionStart="FIGURE 2" captionStartId="2.[152,255,1810,1834]" captionTargetBox="[151,1436,181,1787]" captionTargetId="figure-22@2.[151,1436,181,1787]" captionTargetPageId="2" captionText="FIGURE 2. Shinobius yulong sp. nov., male left palp in ventral (A) and retrolateral (B) views.Abbreviations: C, conductor; E, embolus; H, hood; MA, median apophysis; SD, sperm duct; ST, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14239403" httpUri="https://zenodo.org/record/14239403/files/figure.png" pageId="1" pageNumber="142">Figs 2A</figureCitation>
,
<figureCitation id="13C92A3F76345F2CFE82FA3F772CCC91" box="[260,292,1462,1486]" captionStart="FIGURE 3" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[208,1380,184,1914]" captionTargetId="figure-19@3.[203,1384,181,1919]" captionTargetPageId="3" captionText="FIGURE 3. Shinobius yulong sp. nov., palpal bulb in retrolateral (A), dorsal (B), ventral (C) and prolateral (D) views. Abbreviations: C, conductor; E, embolus; MA, median apophysis; SD, sperm duct; ST, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14239405" httpUri="https://zenodo.org/record/14239405/files/figure.png" pageId="1" pageNumber="142">3C</figureCitation>
) vs. arched (
<bibRefCitation id="EF634B4B76345F2CFE29FA3F7465CC91" author="Wang, L. Y. &amp; Mu, Y. N. &amp; Zhang, F. &amp; Marusik Y. M. &amp; Zhang, Z. S." box="[431,621,1462,1486]" pageId="1" pageNumber="142" pagination="189 - 195" refId="ref2262" refString="Wang, L. Y., Mu, Y. N., Zhang, F., Marusik Y. M. &amp; Zhang, Z. S. (2024) First record of the spider family Trechaleidae Simon, 1890 (Araneae) from China. Zookeys, 1203, 189 - 195. https: // doi. org / 10.3897 / zookeys. 1203.124808" type="journal article" year="2024">
Wang
<emphasis id="B986EAA876345F2CFE70FA3F7425CC91" box="[502,557,1462,1486]" italics="true" pageId="1" pageNumber="142">et al.</emphasis>
2024
</bibRefCitation>
: figs 2A, 3A, C); 3) the head of spermatheca spherical, visible in the dorsal view of vulva (
<figureCitation id="13C92A3F76345F2CFEBEFA537783CCAD" box="[312,395,1498,1522]" captionStart="FIGURE 4" captionStartId="4.[152,255,1031,1055]" captionTargetBox="[155,1428,208,1002]" captionTargetId="figure-22@4.[151,1436,199,1007]" captionTargetPageId="4" captionText="FIGURE 4. Shinobius yulong sp. nov., female epigyne (A, C, D), and vulva (B), in ventral (A), dorsal (B), posterior (C) and anterior (D) views. Abbreviations: AS, accessory spermatheca; CO, copulatory opening; FD, fertilization duct; HS, head of spermatheca; MP, median plate; SS, stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14239407" httpUri="https://zenodo.org/record/14239407/files/figure.png" pageId="1" pageNumber="142">Fig. 4B</figureCitation>
) vs. cylindrical, half hidden behind the stalk of spermatheca (
<bibRefCitation id="EF634B4B76345F2CFB9AFA5372DFCCAD" author="Wang, L. Y. &amp; Mu, Y. N. &amp; Zhang, F. &amp; Marusik Y. M. &amp; Zhang, Z. S." box="[1052,1239,1498,1522]" pageId="1" pageNumber="142" pagination="189 - 195" refId="ref2262" refString="Wang, L. Y., Mu, Y. N., Zhang, F., Marusik Y. M. &amp; Zhang, Z. S. (2024) First record of the spider family Trechaleidae Simon, 1890 (Araneae) from China. Zookeys, 1203, 189 - 195. https: // doi. org / 10.3897 / zookeys. 1203.124808" type="journal article" year="2024">
Wang
<emphasis id="B986EAA876345F2CFBE4FA537290CCAD" box="[1122,1176,1498,1522]" italics="true" pageId="1" pageNumber="142">et al.</emphasis>
2024
</bibRefCitation>
: figs 2D, 3G).
</paragraph>
</subSubSection>
<subSubSection id="C3E8653176345F28FF41FA747734CB48" lastPageId="5" lastPageNumber="146" pageId="1" pageNumber="142" type="description">
<paragraph id="8B4D36BA76345F2CFF41FA7477C5CE21" blockId="1.[151,1437,1102,2026]" pageId="1" pageNumber="142">
<emphasis id="B986EAA876345F2CFF41FA747799CF49" bold="true" box="[199,401,1533,1558]" pageId="1" pageNumber="142">
<emphasis id="B986EAA876345F2CFF41FA74775ACF4A" box="[199,338,1533,1557]" inLineHeading="true" pageId="1" pageNumber="142" reason="1">Description.</emphasis>
Male
</emphasis>
<typeStatus id="5449881876345F2CFE11FA7777FCCF49" box="[407,500,1534,1558]" pageId="1" pageNumber="142" type="holotype">holotype</typeStatus>
(
<figureCitation id="13C92A3F76345F2CFD87FA77749ACF4A" box="[513,658,1534,1558]" captionStart="FIGURE 1" captionStartId="1.[152,255,921,945]" captionTargetBox="[151,1436,181,898]" captionTargetId="figure-19@1.[151,1436,181,898]" captionTargetPageId="1" captionText="FIGURE 1. Shinobius yulong sp. nov., male holotype (A, D, F) and female paratype (B, C, E). A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Female habitus, ventral view; D. Male chelicerae, ventral view; E. Female chelicerae, ventral view; F. Male endites, labium and sternum, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14239401" httpUri="https://zenodo.org/record/14239401/files/figure.png" pageId="1" pageNumber="142">Figs 1A, D, F</figureCitation>
,
<figureCitation id="13C92A3F76345F2CFD1AFA7774A3CF49" box="[668,683,1534,1558]" captionStart="FIGURE 2" captionStartId="2.[152,255,1810,1834]" captionTargetBox="[151,1436,181,1787]" captionTargetId="figure-22@2.[151,1436,181,1787]" captionTargetPageId="2" captionText="FIGURE 2. Shinobius yulong sp. nov., male left palp in ventral (A) and retrolateral (B) views.Abbreviations: C, conductor; E, embolus; H, hood; MA, median apophysis; SD, sperm duct; ST, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14239403" httpUri="https://zenodo.org/record/14239403/files/figure.png" pageId="1" pageNumber="142">2</figureCitation>
,
<figureCitation id="13C92A3F76345F2CFD30FA7774CDCF49" box="[694,709,1534,1558]" captionStart="FIGURE 3" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[208,1380,184,1914]" captionTargetId="figure-19@3.[203,1384,181,1919]" captionTargetPageId="3" captionText="FIGURE 3. Shinobius yulong sp. nov., palpal bulb in retrolateral (A), dorsal (B), ventral (C) and prolateral (D) views. Abbreviations: C, conductor; E, embolus; MA, median apophysis; SD, sperm duct; ST, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14239405" httpUri="https://zenodo.org/record/14239405/files/figure.png" pageId="1" pageNumber="142">3</figureCitation>
). Total length 4.28; carapace 2.13 long, 1.75 wide; opisthosoma 2.37 long, 1.67 wide. Eye sizes and interdistances: AME 0.11, ALE 0.10, PME 0.14, PLE 0.15; AME‒AME 0.09, AME‒ALE 0.05, ALE‒ALE 0.40, PME‒PME 0.09, PME‒PLE 0.16, PLE‒PLE 0.63, ALE‒PLE 0.24. OAW 0.84, CRW 0.99, OAW/ CRW 0.85, MOA 0.36 long, anterior width 0.29, posterior width 0.40.
<collectingCountry id="F3E5762A76345F2CFC0BF9E375BACFDD" box="[909,946,1642,1666]" name="Switzerland" pageId="1" pageNumber="142">CH</collectingCountry>
0.13,
<collectingCountry id="F3E5762A76345F2CFC73F9E37211CFDD" box="[1013,1049,1642,1666]" name="Switzerland" pageId="1" pageNumber="142">CH</collectingCountry>
/AME 1.18. Labium 0.35 long, 0.38 wide. Sternum 1.11 long, 1.12 wide. Leg measurements: Ⅰ 7.80 (2.16, 0.62, 2.06, 1.91, 1.05), II 7.79 (2.23, 0.71, 1.97, 1.97, 0.91), III 6.87 (2.03, 0.62, 1.69, 1.70, 0.83), IV 8.08 (2.27, 0.70, 1.90, 2.31, 0.90); leg formula 4123. Leg spination: Ⅰ: Fe d3, Ti pv3 rv3, Mt pv3 rv3; II: Fe d3, Ti pv3 rv3, Mt pv3 rv3; III: Fe d3, Ti pv3 rv3, Mt pv3 rv3; IV: Fe d3, Ti pv3 rv3, Mt pv3 rv3. Chelicerae promargin with three teeth, retromargin with three teeth (
<figureCitation id="13C92A3F76345F2CFBC4F9737290CE4D" box="[1090,1176,1786,1810]" captionStart="FIGURE 1" captionStartId="1.[152,255,921,945]" captionTargetBox="[151,1436,181,898]" captionTargetId="figure-19@1.[151,1436,181,898]" captionTargetPageId="1" captionText="FIGURE 1. Shinobius yulong sp. nov., male holotype (A, D, F) and female paratype (B, C, E). A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Female habitus, ventral view; D. Male chelicerae, ventral view; E. Female chelicerae, ventral view; F. Male endites, labium and sternum, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14239401" httpUri="https://zenodo.org/record/14239401/files/figure.png" pageId="1" pageNumber="142">Fig. 1D</figureCitation>
). Color in ethanol (
<figureCitation id="13C92A3F76345F2CFAF7F97376B2CE69" captionStart="FIGURE 1" captionStartId="1.[152,255,921,945]" captionTargetBox="[151,1436,181,898]" captionTargetId="figure-19@1.[151,1436,181,898]" captionTargetPageId="1" captionText="FIGURE 1. Shinobius yulong sp. nov., male holotype (A, D, F) and female paratype (B, C, E). A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Female habitus, ventral view; D. Male chelicerae, ventral view; E. Female chelicerae, ventral view; F. Male endites, labium and sternum, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14239401" httpUri="https://zenodo.org/record/14239401/files/figure.png" pageId="1" pageNumber="142">Fig. 1A</figureCitation>
): carapace brownish-yellow, radial furrows blackish-brown and median fovea reddish-brown. Legs pale-yellow with brown rings on femur, tibia and metatarsus. Opisthosoma black-grey with irregular white marking and several chevron-like white stripes posteriorly.
</paragraph>
<paragraph id="8B4D36BA76345F2CFF41F80375FFCEB5" blockId="1.[151,1437,1102,2026]" pageId="1" pageNumber="142">
Palp (
<figureCitation id="13C92A3F76345F2CFE81F8037743CEFD" box="[263,331,1930,1954]" captionStart="FIGURE 2" captionStartId="2.[152,255,1810,1834]" captionTargetBox="[151,1436,181,1787]" captionTargetId="figure-22@2.[151,1436,181,1787]" captionTargetPageId="2" captionText="FIGURE 2. Shinobius yulong sp. nov., male left palp in ventral (A) and retrolateral (B) views.Abbreviations: C, conductor; E, embolus; H, hood; MA, median apophysis; SD, sperm duct; ST, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14239403" httpUri="https://zenodo.org/record/14239403/files/figure.png" pageId="1" pageNumber="142">Fig. 2</figureCitation>
): RTA absent, replaced by a sclerotized hood; median apophysis heavily sclerotized, with broad base and two tips, dorsal division hook-shaped; conductor finger-shaped, membranous; sperm duct meandering; embolus arcshaped, behind the median apophysis, tip slightly curved towards the dorsal side.
</paragraph>
<caption id="DF8D663276375F2FFF1EF89B722FCE11" ID-DOI="http://doi.org/10.5281/zenodo.14239403" ID-Zenodo-Dep="14239403" httpUri="https://zenodo.org/record/14239403/files/figure.png" pageId="2" pageNumber="143" startId="2.[152,255,1810,1834]" targetBox="[151,1436,181,1787]" targetPageId="2" targetType="figure">
<paragraph id="8B4D36BA76375F2FFF1EF89B722FCE11" blockId="2.[151,1436,1810,1871]" pageId="2" pageNumber="143">
<emphasis id="B986EAA876375F2FFF1EF89B7710CE75" bold="true" box="[152,280,1810,1834]" pageId="2" pageNumber="143">FIGURE 2.</emphasis>
<taxonomicName id="4CF24D3976375F2FFE9BF89B77C4CE75" authorityName="Zhang &amp; Yu &amp; Zhang" authorityYear="2024" box="[285,460,1810,1834]" class="Arachnida" family="Trechaleidae" genus="Shinobius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="yulong" status="sp. nov.">
<emphasis id="B986EAA876375F2FFE9BF89B77C4CE75" box="[285,460,1810,1834]" italics="true" pageId="2" pageNumber="143">Shinobius yulong</emphasis>
</taxonomicName>
<emphasis id="B986EAA876375F2FFE54F89A742ACE74" bold="true" box="[466,546,1811,1835]" pageId="2" pageNumber="143">
<taxonomicNameLabel id="A2B557D376375F2FFE54F89A742ACE74" box="[466,546,1811,1835]" pageId="2" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male left palp in ventral (A) and retrolateral (B) views. Abbreviations:
<emphasis id="B986EAA876375F2FFB73F89B730FCE75" bold="true" box="[1269,1287,1810,1834]" pageId="2" pageNumber="143">C</emphasis>
, conductor;
<emphasis id="B986EAA876375F2FFA03F89A739ECE74" bold="true" box="[1413,1430,1811,1835]" pageId="2" pageNumber="143">E</emphasis>
, embolus;
<emphasis id="B986EAA876375F2FFF7CF8BE7706CE10" bold="true" box="[250,270,1847,1871]" pageId="2" pageNumber="143">H</emphasis>
, hood;
<emphasis id="B986EAA876375F2FFEDDF8BF778DCE11" bold="true" box="[347,389,1846,1870]" pageId="2" pageNumber="143">MA</emphasis>
, median apophysis;
<emphasis id="B986EAA876375F2FFDD3F8BF747DCE11" bold="true" box="[597,629,1846,1870]" pageId="2" pageNumber="143">SD</emphasis>
, sperm duct;
<emphasis id="B986EAA876375F2FFC86F8BF7517CE11" bold="true" box="[768,799,1846,1870]" pageId="2" pageNumber="143">ST</emphasis>
, subtegulum;
<emphasis id="B986EAA876375F2FFC29F8BE75C8CE10" bold="true" box="[943,960,1847,1871]" pageId="2" pageNumber="143">T</emphasis>
, tegulum.
</paragraph>
</caption>
<caption id="DF8D663276365F2EFF1EF81E731ACE8C" ID-DOI="http://doi.org/10.5281/zenodo.14239405" ID-Zenodo-Dep="14239405" httpUri="https://zenodo.org/record/14239405/files/figure.png" pageId="3" pageNumber="144" startId="3.[152,255,1943,1967]" targetBox="[208,1380,184,1914]" targetPageId="3" targetType="figure">
<paragraph id="8B4D36BA76365F2EFF1EF81E731ACE8C" blockId="3.[151,1437,1943,2003]" pageId="3" pageNumber="144">
<emphasis id="B986EAA876365F2EFF1EF81E7715CEF0" bold="true" box="[152,285,1943,1967]" pageId="3" pageNumber="144">FIGURE 3.</emphasis>
<taxonomicName id="4CF24D3976365F2EFEA1F81E77D3CEF0" authorityName="Zhang &amp; Yu &amp; Zhang" authorityYear="2024" box="[295,475,1943,1967]" class="Arachnida" family="Trechaleidae" genus="Shinobius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="yulong" status="sp. nov.">
<emphasis id="B986EAA876365F2EFEA1F81E77D3CEF0" box="[295,475,1943,1967]" italics="true" pageId="3" pageNumber="144">Shinobius yulong</emphasis>
</taxonomicName>
<emphasis id="B986EAA876365F2EFE60F81E7433CEF0" bold="true" box="[486,571,1943,1967]" pageId="3" pageNumber="144">
<taxonomicNameLabel id="A2B557D376365F2EFE60F81E7433CEF0" box="[486,571,1943,1967]" pageId="3" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, palpal bulb in retrolateral (A), dorsal (B), ventral (C) and prolateral (D) views. Abbreviations:
<emphasis id="B986EAA876365F2EFEB2F832774ECE8C" bold="true" box="[308,326,1979,2003]" pageId="3" pageNumber="144">C</emphasis>
, conductor;
<emphasis id="B986EAA876365F2EFE40F83277DFCE8C" bold="true" box="[454,471,1979,2003]" pageId="3" pageNumber="144">E</emphasis>
, embolus;
<emphasis id="B986EAA876365F2EFDC0F8327478CE8C" bold="true" box="[582,624,1979,2003]" pageId="3" pageNumber="144">MA</emphasis>
, median apophysis;
<emphasis id="B986EAA876365F2EFCC7F8327569CE8C" bold="true" box="[833,865,1979,2003]" pageId="3" pageNumber="144">SD</emphasis>
, sperm duct;
<emphasis id="B986EAA876365F2EFC6DF8327202CE8C" bold="true" box="[1003,1034,1979,2003]" pageId="3" pageNumber="144">ST</emphasis>
, subtegulum;
<emphasis id="B986EAA876365F2EFB1CF83272A3CE8C" bold="true" box="[1178,1195,1979,2003]" pageId="3" pageNumber="144">T</emphasis>
, tegulum.
</paragraph>
</caption>
<caption id="DF8D663276315F29FF1EFB8E74EFCD38" ID-DOI="http://doi.org/10.5281/zenodo.14239407" ID-Zenodo-Dep="14239407" httpUri="https://zenodo.org/record/14239407/files/figure.png" pageId="4" pageNumber="145" startId="4.[152,255,1031,1055]" targetBox="[155,1428,208,1002]" targetPageId="4" targetType="figure">
<paragraph id="8B4D36BA76315F29FF1EFB8E74EFCD38" blockId="4.[151,1437,1031,1127]" pageId="4" pageNumber="145">
<emphasis id="B986EAA876315F29FF1EFB8E7711CD40" bold="true" box="[152,281,1031,1055]" pageId="4" pageNumber="145">FIGURE 4.</emphasis>
<taxonomicName id="4CF24D3976315F29FEA6FB8E77D8CD40" authorityName="Zhang &amp; Yu &amp; Zhang" authorityYear="2024" box="[288,464,1031,1055]" class="Arachnida" family="Trechaleidae" genus="Shinobius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="145" phylum="Arthropoda" rank="species" species="yulong" status="sp. nov.">
<emphasis id="B986EAA876315F29FEA6FB8E77D8CD40" box="[288,464,1031,1055]" italics="true" pageId="4" pageNumber="145">Shinobius yulong</emphasis>
</taxonomicName>
<emphasis id="B986EAA876315F29FE51FB8E7420CD40" bold="true" box="[471,552,1031,1055]" pageId="4" pageNumber="145">
<taxonomicNameLabel id="A2B557D376315F29FE51FB8E7420CD40" box="[471,552,1031,1055]" pageId="4" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female epigyne (A, C, D), and vulva (B), in ventral (A), dorsal (B), posterior (C) and anterior (D) views. Abbreviations:
<emphasis id="B986EAA876315F29FD87FBA27420CD1C" bold="true" box="[513,552,1067,1091]" pageId="4" pageNumber="145">AS,</emphasis>
accessory spermatheca;
<emphasis id="B986EAA876315F29FCACFBA27558CD1C" bold="true" box="[810,848,1067,1091]" pageId="4" pageNumber="145">CO</emphasis>
, copulatory opening;
<emphasis id="B986EAA876315F29FBB2FBA2725ECD1C" bold="true" box="[1076,1110,1067,1091]" pageId="4" pageNumber="145">FD</emphasis>
, fertilization duct;
<emphasis id="B986EAA876315F29FA98FBA27348CD1C" bold="true" box="[1310,1344,1067,1091]" pageId="4" pageNumber="145">HS</emphasis>
, head of spermatheca;
<emphasis id="B986EAA876315F29FEA5FBC67742CD38" bold="true" box="[291,330,1103,1127]" pageId="4" pageNumber="145">MP</emphasis>
, median plate;
<emphasis id="B986EAA876315F29FE61FBC6740BCD38" bold="true" box="[487,515,1103,1127]" pageId="4" pageNumber="145">SS</emphasis>
, stalk of spermatheca.
</paragraph>
</caption>
<caption id="DF8D663276315F29FF1EF82F75FBCEE1" ID-DOI="http://doi.org/10.5281/zenodo.14239409" ID-Zenodo-Dep="14239409" box="[152,1011,1958,1982]" httpUri="https://zenodo.org/record/14239409/files/figure.png" pageId="4" pageNumber="145" startId="4.[152,255,1958,1982]" targetBox="[151,1436,1187,1934]" targetPageId="4" targetType="figure">
<paragraph id="8B4D36BA76315F29FF1EF82F75FBCEE1" blockId="4.[152,1011,1958,1982]" box="[152,1011,1958,1982]" pageId="4" pageNumber="145">
<emphasis id="B986EAA876315F29FF1EF82F7710CEE1" bold="true" box="[152,280,1958,1982]" pageId="4" pageNumber="145">FIGURE 5.</emphasis>
Distribution records of
<taxonomicName id="4CF24D3976315F29FD8AF82F74AFCEE1" authorityName="Wang, Mu, Zhang, Marusik &amp; Zhang" authorityYear="2024" box="[524,679,1958,1982]" class="Arachnida" family="Trechaleidae" genus="Shinobius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="145" phylum="Arthropoda" rank="species" species="cona">
<emphasis id="B986EAA876315F29FD8AF82F74AFCEE1" box="[524,679,1958,1982]" italics="true" pageId="4" pageNumber="145">Shinobius cona</emphasis>
</taxonomicName>
and
<emphasis id="B986EAA876315F29FD5FF82F753FCEE1" box="[729,823,1958,1982]" italics="true" pageId="4" pageNumber="145">S. Yulong</emphasis>
<emphasis id="B986EAA876315F29FCB8F82F7586CEE1" bold="true" box="[830,910,1958,1982]" pageId="4" pageNumber="145">
<taxonomicNameLabel id="A2B557D376315F29FCB8F82F7586CEE1" box="[830,910,1958,1982]" pageId="4" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in China.
</paragraph>
</caption>
<paragraph id="8B4D36BA76305F28FF41FF117404C8F4" blockId="5.[151,1437,151,571]" pageId="5" pageNumber="146">
<emphasis id="B986EAA876305F28FF41FF117712C9EF" bold="true" box="[199,282,152,176]" pageId="5" pageNumber="146">Female</emphasis>
<typeStatus id="5449881876305F28FEA4FF117776C9EF" box="[290,382,152,176]" pageId="5" pageNumber="146" type="paratype">paratype</typeStatus>
(
<figureCitation id="13C92A3F76305F28FE0BFF1E742BC9F0" box="[397,547,151,176]" captionStart="FIGURE 1" captionStartId="1.[152,255,921,945]" captionTargetBox="[151,1436,181,898]" captionTargetId="figure-19@1.[151,1436,181,898]" captionTargetPageId="1" captionText="FIGURE 1. Shinobius yulong sp. nov., male holotype (A, D, F) and female paratype (B, C, E). A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Female habitus, ventral view; D. Male chelicerae, ventral view; E. Female chelicerae, ventral view; F. Male endites, labium and sternum, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14239401" httpUri="https://zenodo.org/record/14239401/files/figure.png" pageId="5" pageNumber="146">Figs 1B–C, E</figureCitation>
,
<figureCitation id="13C92A3F76305F28FDA9FF117437C9EF" box="[559,575,152,176]" captionStart="FIGURE 4" captionStartId="4.[152,255,1031,1055]" captionTargetBox="[155,1428,208,1002]" captionTargetId="figure-22@4.[151,1436,199,1007]" captionTargetPageId="4" captionText="FIGURE 4. Shinobius yulong sp. nov., female epigyne (A, C, D), and vulva (B), in ventral (A), dorsal (B), posterior (C) and anterior (D) views. Abbreviations: AS, accessory spermatheca; CO, copulatory opening; FD, fertilization duct; HS, head of spermatheca; MP, median plate; SS, stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14239407" httpUri="https://zenodo.org/record/14239407/files/figure.png" pageId="5" pageNumber="146">4</figureCitation>
). Total length 5.35; carapace 2.38 long, 1.97 wide; opisthosoma 3.27 long, 2.45 wide. Eye sizes and inter-distances:
<collectionCode id="EDE3AE7F76305F28FDA3FF357468C98B" box="[549,608,188,212]" pageId="5" pageNumber="146">AME</collectionCode>
0.12,
<collectionCode id="EDE3AE7F76305F28FD21FF3574D3C98B" box="[679,731,188,212]" pageId="5" pageNumber="146">ALE</collectionCode>
0.11,
<collectionCode id="EDE3AE7F76305F28FCA7FF357550C98C" box="[801,856,188,211]" pageId="5" pageNumber="146">PME</collectionCode>
0.15,
<collectionCode id="EDE3AE7F76305F28FC26FF3575C7C98C" box="[928,975,188,211]" pageId="5" pageNumber="146">PLE</collectionCode>
0.14;
<collectionCode id="EDE3AE7F76305F28FB91FF35725AC98B" box="[1047,1106,188,212]" pageId="5" pageNumber="146">AME</collectionCode>
‒
<collectionCode id="EDE3AE7F76305F28FBE6FF357293C98B" box="[1120,1179,188,212]" pageId="5" pageNumber="146">AME</collectionCode>
0.10,
<collectionCode id="EDE3AE7F76305F28FB67FF357314C98B" box="[1249,1308,188,212]" pageId="5" pageNumber="146">AME</collectionCode>
‒
<collectionCode id="EDE3AE7F76305F28FAACFF357356C98B" box="[1322,1374,188,212]" pageId="5" pageNumber="146">ALE</collectionCode>
0.10,
<collectionCode id="EDE3AE7F76305F28FF11FF6976C3C9A7" box="[151,203,224,248]" pageId="5" pageNumber="146">ALE</collectionCode>
‒
<collectionCode id="EDE3AE7F76305F28FF5EFF697704C9A7" box="[216,268,224,248]" pageId="5" pageNumber="146">ALE</collectionCode>
0.46,
<collectionCode id="EDE3AE7F76305F28FEC8FF69778DC9A8" box="[334,389,224,247]" pageId="5" pageNumber="146">PME</collectionCode>
‒
<collectionCode id="EDE3AE7F76305F28FE14FF6977C1C9A8" box="[402,457,224,247]" pageId="5" pageNumber="146">PME</collectionCode>
0.13,
<collectionCode id="EDE3AE7F76305F28FD8DFF69744AC9A8" box="[523,578,224,247]" pageId="5" pageNumber="146">PME</collectionCode>
‒
<collectionCode id="EDE3AE7F76305F28FDC9FF697476C9A8" box="[591,638,224,247]" pageId="5" pageNumber="146">PLE</collectionCode>
0.24,
<collectionCode id="EDE3AE7F76305F28FD47FF6974F8C9A8" box="[705,752,224,247]" pageId="5" pageNumber="146">PLE</collectionCode>
‒
<collectionCode id="EDE3AE7F76305F28FD78FF697525C9A8" box="[766,813,224,247]" pageId="5" pageNumber="146">PLE</collectionCode>
0.76,
<collectionCode id="EDE3AE7F76305F28FCE8FF6975AAC9A7" box="[878,930,224,248]" pageId="5" pageNumber="146">ALE</collectionCode>
‒
<collectionCode id="EDE3AE7F76305F28FC29FF6975D7C9A7" box="[943,991,224,248]" pageId="5" pageNumber="146">PLE</collectionCode>
0.29. OAW 1.01, CRW 1.22, OAW/
<collectionCode id="EDE3AE7F76305F28FAE4FF697395C9A7" box="[1378,1437,224,248]" pageId="5" pageNumber="146">CRW</collectionCode>
0.83,
<collectionCode id="EDE3AE7F76305F28FF52FE8D7719C843" box="[212,273,260,284]" pageId="5" pageNumber="146">MOA</collectionCode>
0.38 long, anterior width 0.33, posterior width 0.45.
<collectingCountry id="F3E5762A76305F28FCCDFE8D7578C843" box="[843,880,260,284]" name="Switzerland" pageId="5" pageNumber="146">CH</collectingCountry>
0.13,
<collectingCountry id="F3E5762A76305F28FC32FE8D75D0C843" box="[948,984,260,284]" name="Switzerland" pageId="5" pageNumber="146">CH</collectingCountry>
/AME 1.18.
<location id="8E2D606176305F28FBD9FE8A72BAC844" LSID="urn:lsid:plazi:treatment:035B87AC76355F28FF11F8C67404CB64:8E2D606176305F28FBD9FE8A72BAC844" box="[1119,1202,259,283]" country="Switzerland" county="Coloration" municipality="Leg" name="Labium" pageId="5" pageNumber="146">Labium</location>
0.43 long, 0.46 wide.
<location id="8E2D606176305F28FF11FEA176F9C81F" LSID="urn:lsid:plazi:treatment:035B87AC76355F28FF11F8C67404CB64:8E2D606176305F28FF11FEA176F9C81F" box="[151,241,296,320]" country="Switzerland" county="Coloration" municipality="Leg" name="Sternum" pageId="5" pageNumber="146">Sternum</location>
1.19 long, 1.24 wide.
<location id="8E2D606176305F28FE6CFEA1741BC81F" LSID="urn:lsid:plazi:treatment:035B87AC76355F28FF11F8C67404CB64:8E2D606176305F28FE6CFEA1741BC81F" box="[490,531,296,320]" country="Switzerland" county="Coloration" municipality="Leg" name="Leg" pageId="5" pageNumber="146">Leg</location>
measurements: I 8.07 (2.31, 0.78, 2.09, 1.91, 0.98), II 8.16 (2.37, 0.75, 2.18, 1.98, 0.88), III 7.49 (2.25, 0.76, 1.78, 1.84, 0.86), IV 8.60 (2.46, 0.71, 1.97, 2.45, 1.01), leg formula 4213. Leg spination and chelicerae teeth on each of retro- and pro-margin as in male (
<figureCitation id="13C92A3F76305F28FC9FFEE67562C8D7" box="[793,874,367,392]" captionStart="FIGURE 1" captionStartId="1.[152,255,921,945]" captionTargetBox="[151,1436,181,898]" captionTargetId="figure-19@1.[151,1436,181,898]" captionTargetPageId="1" captionText="FIGURE 1. Shinobius yulong sp. nov., male holotype (A, D, F) and female paratype (B, C, E). A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Female habitus, ventral view; D. Male chelicerae, ventral view; E. Female chelicerae, ventral view; F. Male endites, labium and sternum, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14239401" httpUri="https://zenodo.org/record/14239401/files/figure.png" pageId="5" pageNumber="146">Fig. 1E</figureCitation>
).
<collectingCounty id="622C4E3676305F28FCF8FEE675F8C8D8" box="[894,1008,367,391]" pageId="5" pageNumber="146">Coloration</collectingCounty>
(
<figureCitation id="13C92A3F76305F28FC7BFEE67279C8D7" box="[1021,1137,367,392]" captionStart="FIGURE 1" captionStartId="1.[152,255,921,945]" captionTargetBox="[151,1436,181,898]" captionTargetId="figure-19@1.[151,1436,181,898]" captionTargetPageId="1" captionText="FIGURE 1. Shinobius yulong sp. nov., male holotype (A, D, F) and female paratype (B, C, E). A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Female habitus, ventral view; D. Male chelicerae, ventral view; E. Female chelicerae, ventral view; F. Male endites, labium and sternum, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14239401" httpUri="https://zenodo.org/record/14239401/files/figure.png" pageId="5" pageNumber="146">Figs 1B–C</figureCitation>
) like male, but opisthosoma without chevron-like white stripes.
</paragraph>
<paragraph id="8B4D36BA76305F28FF41FE3E7734CB48" blockId="5.[151,1437,151,571]" pageId="5" pageNumber="146">
Epigyne (
<figureCitation id="13C92A3F76305F28FEA8FE3E77C2C88F" box="[302,458,439,464]" captionStart="FIGURE 4" captionStartId="4.[152,255,1031,1055]" captionTargetBox="[155,1428,208,1002]" captionTargetId="figure-22@4.[151,1436,199,1007]" captionTargetPageId="4" captionText="FIGURE 4. Shinobius yulong sp. nov., female epigyne (A, C, D), and vulva (B), in ventral (A), dorsal (B), posterior (C) and anterior (D) views. Abbreviations: AS, accessory spermatheca; CO, copulatory opening; FD, fertilization duct; HS, head of spermatheca; MP, median plate; SS, stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14239407" httpUri="https://zenodo.org/record/14239407/files/figure.png" pageId="5" pageNumber="146">Figs 4A, C–D</figureCitation>
): epigynal plate trapezoid, with a pair of lateral folds. Vulva (
<figureCitation id="13C92A3F76305F28FBDFFE3E72A5C88F" box="[1113,1197,439,464]" captionStart="FIGURE 4" captionStartId="4.[152,255,1031,1055]" captionTargetBox="[155,1428,208,1002]" captionTargetId="figure-22@4.[151,1436,199,1007]" captionTargetPageId="4" captionText="FIGURE 4. Shinobius yulong sp. nov., female epigyne (A, C, D), and vulva (B), in ventral (A), dorsal (B), posterior (C) and anterior (D) views. Abbreviations: AS, accessory spermatheca; CO, copulatory opening; FD, fertilization duct; HS, head of spermatheca; MP, median plate; SS, stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14239407" httpUri="https://zenodo.org/record/14239407/files/figure.png" pageId="5" pageNumber="146">Fig. 4B</figureCitation>
): head of spermatheca spherical, significantly enlarged; accessory spermathecae finger-shaped; stalk of spermatheca C-shaped; copulatory ducts short and wide.
</paragraph>
</subSubSection>
<subSubSection id="C3E8653176305F28FF41FDAA7404CB64" box="[199,524,547,571]" pageId="5" pageNumber="146" type="distribution">
<paragraph id="8B4D36BA76305F28FF41FDAA7404CB64" blockId="5.[151,1437,151,571]" box="[199,524,547,571]" pageId="5" pageNumber="146">
<emphasis id="B986EAA876305F28FF41FDAA7752CB64" bold="true" box="[199,346,547,571]" inLineHeading="true" pageId="5" pageNumber="146" reason="1">Distribution.</emphasis>
<collectingCountry id="F3E5762A76305F28FEE6FDAA77A8CB64" box="[352,416,547,571]" name="China" pageId="5" pageNumber="146">China</collectingCountry>
(
<collectingRegion id="4936F85876305F28FE28FDAA7409CB64" box="[430,513,547,571]" country="China" name="Xizang" pageId="5" pageNumber="146">Xizang</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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