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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.960.54053" ID-GBIF-Dataset="cb9000f1-4fb7-4516-8045-609ee2f861de" ID-PMC="PMC7445191" ID-Pensoft-Pub="1313-2970-960-39" ID-Pensoft-UUID="B887F26343D65937AD1404E91BB73583" ID-PubMed="32884397" ID-ZooBank="2444E1F8400240EABA3C4B1D11778DF2" ModsDocID="1313-2970-960-39" checkinTime="1597680771070" checkinUser="pensoft" docAuthor="Tong, Yanfeng, Li, Shuqiang &amp; Bian, Dongju" docDate="2020" docId="928E0B69FD725DC1A0037E8A37BF3908" docLanguage="en" docName="ZooKeys 960: 39-62" docOrigin="ZooKeys 960" docSource="http://dx.doi.org/10.3897/zookeys.960.54053" docTitle="Trilacuna triseta Tong &amp; Li 2020, sp. nov." docType="treatment" docUuid="C04150ED-AD13-41F8-A86D-EE9FDD9B02D0" docUuidSource="ZooBank" docVersion="5" id="B887F26343D65937AD1404E91BB73583" lastPageNumber="39" masterDocId="B887F26343D65937AD1404E91BB73583" masterDocTitle="Taxonomic studies on the genus Trilacuna (Araneae, Oonopidae) from Myanmar" masterLastPageNumber="62" masterPageNumber="39" pageNumber="39" updateTime="1668169272096" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic studies on the genus Trilacuna (Araneae, Oonopidae) from Myanmar</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tong, Yanfeng</mods:namePart>
<mods:affiliation>Life Science College, Shenyang Normal University, Shenyang 110034, China &amp; Southeast Asia Biological Diversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Shuqiang</mods:namePart>
<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-3290-5416</mods:nameIdentifier>
<mods:nameIdentifier type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bian, Dongju</mods:namePart>
<mods:affiliation>CAS Key Laboratory of Forest Ecology and Management, Institute of Applied Ecology, Shenyang 110016, China</mods:affiliation>
<mods:nameIdentifier type="email">biandongju@163.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>960</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>39</mods:start>
<mods:end>62</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.960.54053</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.960.54053</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-960-39</mods:identifier>
<mods:identifier type="ZooBank">2444E1F8400240EABA3C4B1D11778DF2</mods:identifier>
<mods:identifier type="Pensoft-UUID">B887F26343D65937AD1404E91BB73583</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7019551" ID-GBIF-Taxon="166240672" ID-Zenodo-Dep="7019551" LSID="urn:lsid:zoobank.org:act:C04150ED-AD13-41F8-A86D-EE9FDD9B02D0" httpUri="http://treatment.plazi.org/id/928E0B69FD725DC1A0037E8A37BF3908" lastPageNumber="39" pageId="0" pageNumber="39">
<subSubSection pageId="0" pageNumber="39" type="nomenclature">
<paragraph pageId="0" pageNumber="39">
<taxonomicName LSID="928E0B69-FD72-5DC1-A003-7E8A37BF3908" authority="Tong &amp; Li" authorityName="Tong &amp; Li" authorityYear="2020" class="Arachnida" family="Oonopidae" genus="Trilacuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trilacuna triseta" order="Araneae" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="triseta" status="sp. nov.">Trilacuna triseta Tong &amp; Li</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="39">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Trilacuna triseta sp. nov., male holotype A-C habitus in dorsal, lateral, and ventral views D-F, H, I prosoma in dorsal, lateral, ventral, posterior, and anterior views G labium and endites in ventral view (arrow shows the three black, strong setae). Abbreviations: boc = booklung covers; cmp = clypeus median projection; esb = elevated seta base; ldi = labium deep incision. Scale bars: 0.4 mm (A-F, H, I); 0.2 mm (G)." figureDoi="10.3897/zookeys.960.54053.figure11" httpUri="https://binary.pensoft.net/fig/442560" pageId="0" pageNumber="39">Figs 11</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Trilacuna triseta sp. nov., male holotype, left palp, SEM A, B, G prolateral, retrolateral, and dorsal views C, D palpal bulb in prolateral and retrolateral views E, F, H distal part of palpal bulb in prolateral, retrolateral, and dorsal views. Abbreviations: ab = anterior branch; db = dorsal branch; emb = embolus; ml = median lobe; vb = ventral branch." figureDoi="10.3897/zookeys.960.54053.figure12" httpUri="https://binary.pensoft.net/fig/442561" pageId="0" pageNumber="39">, 12</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Trilacuna spp., left male palp A-C T. besucheti D-F T. changzi sp. nov. G-I T. hponkanrazi sp. nov. J-L T. triseta sp. nov. A, D, G, J prolateral views B, E, H, K dorsal views C, F, I, L retrolateral views. Scales: 0.2 mm." figureDoi="10.3897/zookeys.960.54053.figure14" httpUri="https://binary.pensoft.net/fig/442563" pageId="0" pageNumber="39">, 14J-L</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Trilacuna spp., male labium and endites, ventral view A, B T. changzi sp. nov., white arrow shows the long, strongly curved spines C, D T. hponkanrazi sp. nov. E, F T. besucheti G, H T. triseta sp. nov., black arrow shows the three black, strong setae. Abbreviations: bts = bent thick setae; cos = comb-like setae." figureDoi="10.3897/zookeys.960.54053.figure16" httpUri="https://binary.pensoft.net/fig/442565" pageId="0" pageNumber="39">, 16G</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Trilacuna spp., male labium and endites, ventral view A, B T. changzi sp. nov., white arrow shows the long, strongly curved spines C, D T. hponkanrazi sp. nov. E, F T. besucheti G, H T. triseta sp. nov., black arrow shows the three black, strong setae. Abbreviations: bts = bent thick setae; cos = comb-like setae." figureDoi="10.3897/zookeys.960.54053.figure16" httpUri="https://binary.pensoft.net/fig/442565" pageId="0" pageNumber="39">, 16H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="type material">
<paragraph pageId="0" pageNumber="39">Type material.</paragraph>
<paragraph pageId="0" pageNumber="39">
<emphasis bold="true" italics="true" pageId="0" pageNumber="39">Holotype</emphasis>
♂: Myanmar, Kachin State, Putao, Hponkanrazi Wildlife Sanctuary;
<geoCoordinate degrees="27" direction="north" minutes="32.032" orientation="latitude" precision="1" value="27.533867">27°32.032'N</geoCoordinate>
,
<geoCoordinate degrees="97" direction="east" minutes="00.036" orientation="longitude" precision="1" value="97.0006">97°00.036'E</geoCoordinate>
; elevation ca 2010 m; 15.V.2017; Wu J. and Chen Z. leg. (IZCAS AR-25155).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="diagnosis">
<paragraph pageId="0" pageNumber="39">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="39">
The new species is similar to
<taxonomicName lsidName="T. bilingua" pageId="0" pageNumber="39" rank="species" species="bilingua">
<emphasis italics="true" pageId="0" pageNumber="39">T. bilingua</emphasis>
</taxonomicName>
Eichenberger, 2011 but can be distinguished by the three black, thick setae on the endites of the male (Figs
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Trilacuna triseta sp. nov., male holotype A-C habitus in dorsal, lateral, and ventral views D-F, H, I prosoma in dorsal, lateral, ventral, posterior, and anterior views G labium and endites in ventral view (arrow shows the three black, strong setae). Abbreviations: boc = booklung covers; cmp = clypeus median projection; esb = elevated seta base; ldi = labium deep incision. Scale bars: 0.4 mm (A-F, H, I); 0.2 mm (G)." figureDoi="10.3897/zookeys.960.54053.figure11" httpUri="https://binary.pensoft.net/fig/442560" pageId="0" pageNumber="39">11G</figureCitation>
,
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Trilacuna spp., male labium and endites, ventral view A, B T. changzi sp. nov., white arrow shows the long, strongly curved spines C, D T. hponkanrazi sp. nov. E, F T. besucheti G, H T. triseta sp. nov., black arrow shows the three black, strong setae. Abbreviations: bts = bent thick setae; cos = comb-like setae." figureDoi="10.3897/zookeys.960.54053.figure16" httpUri="https://binary.pensoft.net/fig/442565" pageId="0" pageNumber="39">16G</figureCitation>
,
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Trilacuna spp., male labium and endites, ventral view A, B T. changzi sp. nov., white arrow shows the long, strongly curved spines C, D T. hponkanrazi sp. nov. E, F T. besucheti G, H T. triseta sp. nov., black arrow shows the three black, strong setae. Abbreviations: bts = bent thick setae; cos = comb-like setae." figureDoi="10.3897/zookeys.960.54053.figure16" httpUri="https://binary.pensoft.net/fig/442565" pageId="0" pageNumber="39">16H</figureCitation>
) and the slender anterior branch (ab) of the embolus system (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Trilacuna triseta sp. nov., male holotype, left palp, SEM A, B, G prolateral, retrolateral, and dorsal views C, D palpal bulb in prolateral and retrolateral views E, F, H distal part of palpal bulb in prolateral, retrolateral, and dorsal views. Abbreviations: ab = anterior branch; db = dorsal branch; emb = embolus; ml = median lobe; vb = ventral branch." figureDoi="10.3897/zookeys.960.54053.figure12" httpUri="https://binary.pensoft.net/fig/442561" pageId="0" pageNumber="39">12E</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Trilacuna triseta sp. nov., male holotype, left palp, SEM A, B, G prolateral, retrolateral, and dorsal views C, D palpal bulb in prolateral and retrolateral views E, F, H distal part of palpal bulb in prolateral, retrolateral, and dorsal views. Abbreviations: ab = anterior branch; db = dorsal branch; emb = embolus; ml = median lobe; vb = ventral branch." figureDoi="10.3897/zookeys.960.54053.figure12" httpUri="https://binary.pensoft.net/fig/442561" pageId="0" pageNumber="39">12F</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Trilacuna triseta sp. nov., male holotype, left palp, SEM A, B, G prolateral, retrolateral, and dorsal views C, D palpal bulb in prolateral and retrolateral views E, F, H distal part of palpal bulb in prolateral, retrolateral, and dorsal views. Abbreviations: ab = anterior branch; db = dorsal branch; emb = embolus; ml = median lobe; vb = ventral branch." figureDoi="10.3897/zookeys.960.54053.figure12" httpUri="https://binary.pensoft.net/fig/442561" pageId="0" pageNumber="39">12H</figureCitation>
).
<taxonomicName authorityName="Eichenberger" authorityYear="2011" class="Arachnida" family="Oonopidae" genus="Trilacuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trilacuna bilingua" order="Araneae" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="bilingua">
<emphasis italics="true" pageId="0" pageNumber="39">Trilacuna bilingua</emphasis>
</taxonomicName>
lacks black, thick setae on the endites, and the embolus system has two very short lobes basally (Eichenberger et al. 2011: fig. 5-6).
</paragraph>
<caption doi="10.3897/zookeys.960.54053.figure11" httpUri="https://binary.pensoft.net/fig/442560" pageId="0" pageNumber="39" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="39">
<emphasis bold="true" pageId="0" pageNumber="39">Figure 11.</emphasis>
<taxonomicName authorityName="Tong &amp; Li" authorityYear="2020" class="Arachnida" family="Oonopidae" genus="Trilacuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trilacuna triseta" order="Araneae" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="triseta">
<emphasis italics="true" pageId="0" pageNumber="39">Trilacuna triseta</emphasis>
</taxonomicName>
sp. nov., male holotype
<emphasis bold="true" pageId="0" pageNumber="39">A-C</emphasis>
habitus in dorsal, lateral, and ventral views
<emphasis bold="true" pageId="0" pageNumber="39">D-F, H, I</emphasis>
prosoma in dorsal, lateral, ventral, posterior, and anterior views
<emphasis bold="true" pageId="0" pageNumber="39">G</emphasis>
labium and endites in ventral view (arrow shows the three black, strong setae). Abbreviations: boc = booklung covers; cmp = clypeus median projection; esb = elevated seta base; ldi = labium deep incision. Scale bars: 0.4 mm (
<emphasis bold="true" pageId="0" pageNumber="39">A-F, H, I</emphasis>
); 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="39">G</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="description">
<paragraph pageId="0" pageNumber="39">Description.</paragraph>
<paragraph pageId="0" pageNumber="39">
<emphasis bold="true" pageId="0" pageNumber="39">
Male.
<emphasis italics="true" pageId="0" pageNumber="39">Body</emphasis>
</emphasis>
: reddish brown, chelicerae and sternum lighter, legs yellow; habitus as in Figure
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Trilacuna triseta sp. nov., male holotype A-C habitus in dorsal, lateral, and ventral views D-F, H, I prosoma in dorsal, lateral, ventral, posterior, and anterior views G labium and endites in ventral view (arrow shows the three black, strong setae). Abbreviations: boc = booklung covers; cmp = clypeus median projection; esb = elevated seta base; ldi = labium deep incision. Scale bars: 0.4 mm (A-F, H, I); 0.2 mm (G)." figureDoi="10.3897/zookeys.960.54053.figure11" httpUri="https://binary.pensoft.net/fig/442560" pageId="0" pageNumber="39">11A-C</figureCitation>
; body length 1.97.
<emphasis bold="true" italics="true" pageId="0" pageNumber="39">Carapace</emphasis>
: 0.95 long, 0.74 wide; sides smooth; lateral margin rebordered (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Trilacuna triseta sp. nov., male holotype A-C habitus in dorsal, lateral, and ventral views D-F, H, I prosoma in dorsal, lateral, ventral, posterior, and anterior views G labium and endites in ventral view (arrow shows the three black, strong setae). Abbreviations: boc = booklung covers; cmp = clypeus median projection; esb = elevated seta base; ldi = labium deep incision. Scale bars: 0.4 mm (A-F, H, I); 0.2 mm (G)." figureDoi="10.3897/zookeys.960.54053.figure11" httpUri="https://binary.pensoft.net/fig/442560" pageId="0" pageNumber="39">11B</figureCitation>
); posterior surface with several large setal bases (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Trilacuna triseta sp. nov., male holotype A-C habitus in dorsal, lateral, and ventral views D-F, H, I prosoma in dorsal, lateral, ventral, posterior, and anterior views G labium and endites in ventral view (arrow shows the three black, strong setae). Abbreviations: boc = booklung covers; cmp = clypeus median projection; esb = elevated seta base; ldi = labium deep incision. Scale bars: 0.4 mm (A-F, H, I); 0.2 mm (G)." figureDoi="10.3897/zookeys.960.54053.figure11" httpUri="https://binary.pensoft.net/fig/442560" pageId="0" pageNumber="39">11H</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="39">Eyes</emphasis>
: ALE largest; PLE and PME nearly equal in size; ALE-PLE separated by less than ALE radius; PME touching each other; posterior eye row recurved as viewed from above, procurved as viewed from front (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Trilacuna triseta sp. nov., male holotype A-C habitus in dorsal, lateral, and ventral views D-F, H, I prosoma in dorsal, lateral, ventral, posterior, and anterior views G labium and endites in ventral view (arrow shows the three black, strong setae). Abbreviations: boc = booklung covers; cmp = clypeus median projection; esb = elevated seta base; ldi = labium deep incision. Scale bars: 0.4 mm (A-F, H, I); 0.2 mm (G)." figureDoi="10.3897/zookeys.960.54053.figure11" httpUri="https://binary.pensoft.net/fig/442560" pageId="0" pageNumber="39">11D, I</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="39">Clypeus</emphasis>
: height about 0.85 times of ALE diameter, with a triangular, pointed, median projection (cmp).
<emphasis bold="true" italics="true" pageId="0" pageNumber="39">Mouthparts</emphasis>
(Figs
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Trilacuna triseta sp. nov., male holotype A-C habitus in dorsal, lateral, and ventral views D-F, H, I prosoma in dorsal, lateral, ventral, posterior, and anterior views G labium and endites in ventral view (arrow shows the three black, strong setae). Abbreviations: boc = booklung covers; cmp = clypeus median projection; esb = elevated seta base; ldi = labium deep incision. Scale bars: 0.4 mm (A-F, H, I); 0.2 mm (G)." figureDoi="10.3897/zookeys.960.54053.figure11" httpUri="https://binary.pensoft.net/fig/442560" pageId="0" pageNumber="39">11F, I</figureCitation>
,
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Trilacuna spp., male labium and endites, ventral view A, B T. changzi sp. nov., white arrow shows the long, strongly curved spines C, D T. hponkanrazi sp. nov. E, F T. besucheti G, H T. triseta sp. nov., black arrow shows the three black, strong setae. Abbreviations: bts = bent thick setae; cos = comb-like setae." figureDoi="10.3897/zookeys.960.54053.figure16" httpUri="https://binary.pensoft.net/fig/442565" pageId="0" pageNumber="39">16G, H</figureCitation>
): endites with three thick, black setae.
<emphasis bold="true" italics="true" pageId="0" pageNumber="39">Sternum</emphasis>
(Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Trilacuna triseta sp. nov., male holotype A-C habitus in dorsal, lateral, and ventral views D-F, H, I prosoma in dorsal, lateral, ventral, posterior, and anterior views G labium and endites in ventral view (arrow shows the three black, strong setae). Abbreviations: boc = booklung covers; cmp = clypeus median projection; esb = elevated seta base; ldi = labium deep incision. Scale bars: 0.4 mm (A-F, H, I); 0.2 mm (G)." figureDoi="10.3897/zookeys.960.54053.figure11" httpUri="https://binary.pensoft.net/fig/442560" pageId="0" pageNumber="39">11F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="39">Abdomen</emphasis>
: 1.05 long, 0.80 wide; booklung covers ovoid, surface smooth (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Trilacuna triseta sp. nov., male holotype A-C habitus in dorsal, lateral, and ventral views D-F, H, I prosoma in dorsal, lateral, ventral, posterior, and anterior views G labium and endites in ventral view (arrow shows the three black, strong setae). Abbreviations: boc = booklung covers; cmp = clypeus median projection; esb = elevated seta base; ldi = labium deep incision. Scale bars: 0.4 mm (A-F, H, I); 0.2 mm (G)." figureDoi="10.3897/zookeys.960.54053.figure11" httpUri="https://binary.pensoft.net/fig/442560" pageId="0" pageNumber="39">11B</figureCitation>
); dorsal scutum not fused to epigastric scutum; apodemes present; posterior spiracles connected by groove (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Trilacuna triseta sp. nov., male holotype A-C habitus in dorsal, lateral, and ventral views D-F, H, I prosoma in dorsal, lateral, ventral, posterior, and anterior views G labium and endites in ventral view (arrow shows the three black, strong setae). Abbreviations: boc = booklung covers; cmp = clypeus median projection; esb = elevated seta base; ldi = labium deep incision. Scale bars: 0.4 mm (A-F, H, I); 0.2 mm (G)." figureDoi="10.3897/zookeys.960.54053.figure11" httpUri="https://binary.pensoft.net/fig/442560" pageId="0" pageNumber="39">11C</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="39">Palp</emphasis>
(Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Trilacuna triseta sp. nov., male holotype, left palp, SEM A, B, G prolateral, retrolateral, and dorsal views C, D palpal bulb in prolateral and retrolateral views E, F, H distal part of palpal bulb in prolateral, retrolateral, and dorsal views. Abbreviations: ab = anterior branch; db = dorsal branch; emb = embolus; ml = median lobe; vb = ventral branch." figureDoi="10.3897/zookeys.960.54053.figure12" httpUri="https://binary.pensoft.net/fig/442561" pageId="0" pageNumber="39">12</figureCitation>
,
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Trilacuna spp., left male palp A-C T. besucheti D-F T. changzi sp. nov. G-I T. hponkanrazi sp. nov. J-L T. triseta sp. nov. A, D, G, J prolateral views B, E, H, K dorsal views C, F, I, L retrolateral views. Scales: 0.2 mm." figureDoi="10.3897/zookeys.960.54053.figure14" httpUri="https://binary.pensoft.net/fig/442563" pageId="0" pageNumber="39">14J-L</figureCitation>
): orange; 0.72 long (0.17, 0.11, 0.11, 0.33); femur elongated (width/length = 0.53) (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Trilacuna triseta sp. nov., male holotype, left palp, SEM A, B, G prolateral, retrolateral, and dorsal views C, D palpal bulb in prolateral and retrolateral views E, F, H distal part of palpal bulb in prolateral, retrolateral, and dorsal views. Abbreviations: ab = anterior branch; db = dorsal branch; emb = embolus; ml = median lobe; vb = ventral branch." figureDoi="10.3897/zookeys.960.54053.figure12" httpUri="https://binary.pensoft.net/fig/442561" pageId="0" pageNumber="39">12A, B</figureCitation>
); bulb oval, tapering apically; embolus system (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Trilacuna triseta sp. nov., male holotype, left palp, SEM A, B, G prolateral, retrolateral, and dorsal views C, D palpal bulb in prolateral and retrolateral views E, F, H distal part of palpal bulb in prolateral, retrolateral, and dorsal views. Abbreviations: ab = anterior branch; db = dorsal branch; emb = embolus; ml = median lobe; vb = ventral branch." figureDoi="10.3897/zookeys.960.54053.figure12" httpUri="https://binary.pensoft.net/fig/442561" pageId="0" pageNumber="39">12E</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Trilacuna triseta sp. nov., male holotype, left palp, SEM A, B, G prolateral, retrolateral, and dorsal views C, D palpal bulb in prolateral and retrolateral views E, F, H distal part of palpal bulb in prolateral, retrolateral, and dorsal views. Abbreviations: ab = anterior branch; db = dorsal branch; emb = embolus; ml = median lobe; vb = ventral branch." figureDoi="10.3897/zookeys.960.54053.figure12" httpUri="https://binary.pensoft.net/fig/442561" pageId="0" pageNumber="39">12F</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Trilacuna triseta sp. nov., male holotype, left palp, SEM A, B, G prolateral, retrolateral, and dorsal views C, D palpal bulb in prolateral and retrolateral views E, F, H distal part of palpal bulb in prolateral, retrolateral, and dorsal views. Abbreviations: ab = anterior branch; db = dorsal branch; emb = embolus; ml = median lobe; vb = ventral branch." figureDoi="10.3897/zookeys.960.54053.figure12" httpUri="https://binary.pensoft.net/fig/442561" pageId="0" pageNumber="39">12H</figureCitation>
) with an anterior branch (ab), a dorsal branch (db), a ventral branch (vb), and a median lobe (ml) in prolateral view.
</paragraph>
<caption doi="10.3897/zookeys.960.54053.figure12" httpUri="https://binary.pensoft.net/fig/442561" pageId="0" pageNumber="39" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="39">
<emphasis bold="true" pageId="0" pageNumber="39">Figure 12.</emphasis>
<taxonomicName authorityName="Tong &amp; Li" authorityYear="2020" class="Arachnida" family="Oonopidae" genus="Trilacuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trilacuna triseta" order="Araneae" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="triseta">
<emphasis italics="true" pageId="0" pageNumber="39">Trilacuna triseta</emphasis>
</taxonomicName>
sp. nov., male holotype, left palp, SEM
<emphasis bold="true" pageId="0" pageNumber="39">A, B, G</emphasis>
prolateral, retrolateral, and dorsal views
<emphasis bold="true" pageId="0" pageNumber="39">C, D</emphasis>
palpal bulb in prolateral and retrolateral views
<emphasis bold="true" pageId="0" pageNumber="39">E, F, H</emphasis>
distal part of palpal bulb in prolateral, retrolateral, and dorsal views. Abbreviations: ab = anterior branch; db = dorsal branch; emb = embolus; ml = median lobe; vb = ventral branch.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="39">
<emphasis bold="true" pageId="0" pageNumber="39">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="etymology">
<paragraph pageId="0" pageNumber="39">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="39">
The species epithet is a noun in apposition composed of the Latin words
<emphasis italics="true" pageId="0" pageNumber="39">tri</emphasis>
(three) and
<emphasis italics="true" pageId="0" pageNumber="39">seta</emphasis>
and refers to the three black, thick setae on the
<normalizedToken originalValue="male’s">male's</normalizedToken>
endites.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="distribution">
<paragraph pageId="0" pageNumber="39">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="39">Known only from the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="remarks">
<paragraph pageId="0" pageNumber="39">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="39">
There are two species reported from a single female specimen in this study, and one described from a single male:
<taxonomicName lsidName="T. loebli" pageId="0" pageNumber="39" rank="species" species="loebli">
<emphasis italics="true" pageId="0" pageNumber="39">T. loebli</emphasis>
</taxonomicName>
Grismado &amp; Piacentini, 2014,
<taxonomicName lsidName="T. zhigangi" pageId="0" pageNumber="39" rank="species" species="zhigangi">
<emphasis italics="true" pageId="0" pageNumber="39">T. zhigangi</emphasis>
</taxonomicName>
Tong &amp; Li, sp. nov., and
<taxonomicName lsidName="T. triseta" pageId="0" pageNumber="39" rank="species" species="triseta">
<emphasis italics="true" pageId="0" pageNumber="39">T. triseta</emphasis>
</taxonomicName>
Tong &amp; Li, sp. nov., respectively. The following characters indicate that neither of the two females are conspecific with
<taxonomicName lsidName="T. triseta" pageId="0" pageNumber="39" rank="species" species="triseta">
<emphasis italics="true" pageId="0" pageNumber="39">T. triseta</emphasis>
</taxonomicName>
Tong &amp; Li, sp. nov.
<taxonomicName lsidName="T. loebli" pageId="0" pageNumber="39" rank="species" species="loebli">
<emphasis italics="true" pageId="0" pageNumber="39">T. loebli</emphasis>
</taxonomicName>
is dark brown (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Trilacuna loebli, female (IZCAS AR- 25156) A-C habitus in dorsal, ventral, and lateral views D-F, H prosoma in dorsal, ventral, lateral, and anterior views G abdomen in ventral view. Abbreviations: boc = booklung covers; cmp = clypeus median projection; esb = elevated seta base; ldi = labium deep incision; sep = semicircular plate. Scale bars: 0.4 mm." figureDoi="10.3897/zookeys.960.54053.figure10" httpUri="https://binary.pensoft.net/fig/442559" pageId="0" pageNumber="39">10A</figureCitation>
) with a reticulated carapace (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Trilacuna loebli, female (IZCAS AR- 25156) A-C habitus in dorsal, ventral, and lateral views D-F, H prosoma in dorsal, ventral, lateral, and anterior views G abdomen in ventral view. Abbreviations: boc = booklung covers; cmp = clypeus median projection; esb = elevated seta base; ldi = labium deep incision; sep = semicircular plate. Scale bars: 0.4 mm." figureDoi="10.3897/zookeys.960.54053.figure10" httpUri="https://binary.pensoft.net/fig/442559" pageId="0" pageNumber="39">10D</figureCitation>
), and
<taxonomicName lsidName="T. zhigangi" pageId="0" pageNumber="39" rank="species" species="zhigangi">
<emphasis italics="true" pageId="0" pageNumber="39">T. zhigangi</emphasis>
</taxonomicName>
has small eyes (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Trilacuna zhigangi sp. nov., female holotype A-C habitus in dorsal, ventral, and lateral views D-F, H prosoma in dorsal, ventral, lateral, and anterior views G abdomen in ventral view. Abbreviation: esb = elevated seta base; ldi = labium deep incision. Scale bars: 0.4 mm." figureDoi="10.3897/zookeys.960.54053.figure13" httpUri="https://binary.pensoft.net/fig/442562" pageId="0" pageNumber="39">13A</figureCitation>
) and lacks the triangular, pointed, median projection of the clypeus (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Trilacuna zhigangi sp. nov., female holotype A-C habitus in dorsal, ventral, and lateral views D-F, H prosoma in dorsal, ventral, lateral, and anterior views G abdomen in ventral view. Abbreviation: esb = elevated seta base; ldi = labium deep incision. Scale bars: 0.4 mm." figureDoi="10.3897/zookeys.960.54053.figure13" httpUri="https://binary.pensoft.net/fig/442562" pageId="0" pageNumber="39">13H</figureCitation>
). The male,
<taxonomicName lsidName="T. triseta" pageId="0" pageNumber="39" rank="species" species="triseta">
<emphasis italics="true" pageId="0" pageNumber="39">T. triseta</emphasis>
</taxonomicName>
Tong &amp; Li, sp. nov. has a reddish-brown body, a smooth carapace, normal-sized eyes, and a triangular, pointed, median projection (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Trilacuna triseta sp. nov., male holotype A-C habitus in dorsal, lateral, and ventral views D-F, H, I prosoma in dorsal, lateral, ventral, posterior, and anterior views G labium and endites in ventral view (arrow shows the three black, strong setae). Abbreviations: boc = booklung covers; cmp = clypeus median projection; esb = elevated seta base; ldi = labium deep incision. Scale bars: 0.4 mm (A-F, H, I); 0.2 mm (G)." figureDoi="10.3897/zookeys.960.54053.figure11" httpUri="https://binary.pensoft.net/fig/442560" pageId="0" pageNumber="39">11A, D, I</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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