treatments-xml/data/07/DA/91/07DA9195C1BF585ABE8F3D4C5B91BD08.xml

<document ID-CLB-Dataset="21450" ID-DOI="http://dx.doi.org/10.3897/zookeys.950.51298" ID-GBIF-Dataset="21975627-96b7-45c7-9c80-0a73a850169a" ID-PMC="PMC7387366" ID-Pensoft-Pub="1313-2970-950-51" ID-Pensoft-UUID="7F08092C01585A70A7948EA57A2A8E2E" ID-PubMed="32774101" ID-ZooBank="3E805885D3354FB3AE8DFA443FAD82AE" ModsDocID="1313-2970-950-51" checkinTime="1595285263451" checkinUser="pensoft" docAuthor="Takaoka, Hiroyuki, Srisuka, Wichai, Fukuda, Masako &amp; Saeung, Atiporn" docDate="2020" docId="07DA9195C1BF585ABE8F3D4C5B91BD08" docLanguage="en" docName="ZooKeys 950: 51-152" docOrigin="ZooKeys 950" docSource="http://dx.doi.org/10.3897/zookeys.950.51298" docTitle="Simulium (Gomphostilbia) maewongense Takaoka, Srisuka &amp; Saeung 2020, sp. nov." docType="treatment" docUuid="232C34A2-71F4-4F44-B1C9-E0E8D405219F" docUuidSource="ZooBank" docVersion="6" id="7F08092C01585A70A7948EA57A2A8E2E" lastPageNumber="51" masterDocId="7F08092C01585A70A7948EA57A2A8E2E" masterDocTitle="Twenty-one new species of the Simulium (Gomphostilbia) asakoae species group (Diptera, Simuliidae) in Thailand, with their genetic relationships" masterLastPageNumber="152" masterPageNumber="51" pageNumber="51" updateTime="1732680320779" updateUser="ExternalLinkService">
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<mods:title id="FE236B7A8E6B3ADDE1095D4C6B7708FA">Twenty-one new species of the Simulium (Gomphostilbia) asakoae species group (Diptera, Simuliidae) in Thailand, with their genetic relationships</mods:title>
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<mods:namePart id="777C372EE516016C1D16266D2367E340">Takaoka, Hiroyuki</mods:namePart>
<mods:affiliation id="A32FEF5E7580ABC419116F2A264CA463">Tropical Infectious Diseases Research and Education Centre (TIDREC), University of Malaya, 50603, Kuala Lumpur, Malaysia</mods:affiliation>
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<mods:namePart id="0D20663D63FD117732ABB4233EAECE29">Srisuka, Wichai</mods:namePart>
<mods:affiliation id="84AE44BE92C70A50B3E7007A76717239">Entomology Section, Queen Sirikit Botanic Garden, P. O. Box 7, Maerim, Chiang Mai 50180, Thailand</mods:affiliation>
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<mods:namePart id="1EF055D25DC9BB6C0BC531C8D2A34F19">Fukuda, Masako</mods:namePart>
<mods:affiliation id="CAD07F9EDD4D9DB83ADC25445696A6FE">Institute for Research Promotion, Oita University, Idaigaoka 1 - 1, Hasama, Yufu City, Oita, 879 - 5593, Japan</mods:affiliation>
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<mods:namePart id="06446650F5BFB883591D8911D82B6420">Saeung, Atiporn</mods:namePart>
<mods:affiliation id="7330BA5A6E2F83E2E34D350747745FE1">Center of Insect Vector Study, Department of Parasitology, Faculty of Medicine, Chiang Mai University, Chiang Mai 50200, Thailand</mods:affiliation>
<mods:nameIdentifier id="A43761E63E4C0C139C398A60DA92C7E4" type="orcid">https://orcid.org/0000-0003-3550-5992</mods:nameIdentifier>
<mods:nameIdentifier id="A428D2649C925FD64083D5420DE2F4A9" type="email">atisaeung.noi@gmail.com</mods:nameIdentifier>
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<treatment id="07DA9195C1BF585ABE8F3D4C5B91BD08" ID-GBIF-Taxon="165843379" LSID="urn:lsid:zoobank.org:act:232C34A2-71F4-4F44-B1C9-E0E8D405219F" httpUri="http://treatment.plazi.org/id/07DA9195C1BF585ABE8F3D4C5B91BD08" lastPageNumber="51" pageId="0" pageNumber="51" scope_family="Simuliidae" scope_order="Diptera" scope_subGenus="Gomphostilbia">
<subSubSection id="69FDD0B5B56F1F50BFBBE7C8D4BC79AE" pageId="0" pageNumber="51" type="nomenclature">
<paragraph id="53DCDB116E006B2EBCDB6C49ACB3E105" pageId="0" pageNumber="51">
<taxonomicName id="4AB0CA8CA6BE4643F7172F562C294012" ID-CoL="8DXHF" LSID="07DA9195-C1BF-585A-BE8F-3D4C5B91BD08" authority="Takaoka, Srisuka &amp; Saeung" authorityName="Takaoka, Srisuka &amp; Saeung" authorityYear="2020" class="Insecta" family="Simuliidae" genus="Simulium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Simulium (Gomphostilbia) maewongense" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="maewongense" status="sp. nov." subGenus="Gomphostilbia">Simulium (Gomphostilbia) maewongense Takaoka, Srisuka &amp; Saeung</taxonomicName>
<taxonomicNameLabel id="3976464618C07FD58447CFD81F0495D4" pageId="0" pageNumber="51">sp. nov.</taxonomicNameLabel>
<figureCitation id="FF2E75C8DCCCBA0F6EAE7972F9070E6F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male and pupa of S. maewongense sp. nov. A-D male E-G pupa. A hind basitarsus and second tarsomere (left side; lateral view) B coxites, styles and ventral plate (ventral view) C ventral plate and median sclerite (lateral view) D ventral plate (caudal view) E gill filaments (right side; lateral view) F terminal hooks (caudal view) G cocoon (dorsal view). Scale bars: 1.0 mm (G); 0.1 mm (A, E); 0.02 mm (B-D); 0.01 mm (F)." figureDoi="10.3897/zookeys.950.51298.figure7" httpUri="https://binary.pensoft.net/fig/433871" pageId="0" pageNumber="51">Figs 7</figureCitation>
<figureCitation id="C49B9D2FDBC8C1EE26D9EC996A9ABE0D" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Schematic illustrations of larval body color patterns (dorsal view). A S. teerachanense sp. nov. B S. pitasawatae sp. nov. C S. sutheppuiense sp. nov. D S. asakoae E S. klonglanense sp. nov. F S. namdanense sp. nov. G S. chaowaense sp. nov. H S. loeiense sp. nov. I S. huaimorense sp. nov. J S. nanoiense sp. nov. K S. junkumae sp. nov. L S. myanmarense M S. maewongense sp. nov. N S. phapeungense sp. nov. O S. muangpanense sp. nov. P S. maehongsonense sp. nov. Q S. thungchangense sp. nov. R S. chiangdaoense S S. inthanonense (morph 1) T S. inthanonense (morph 2)." figureDoi="10.3897/zookeys.950.51298.figure25" httpUri="https://binary.pensoft.net/fig/433889" pageId="0" pageNumber="51">, 25M</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="FE73F44975FDBB3624A12CF658530F4D" pageId="0" pageNumber="51" type="materials_examined">
<paragraph id="579F0356F7F33E3733F7E8DA42A256E4" pageId="0" pageNumber="51">Material examined.</paragraph>
<paragraph id="D595850F46D73F5CE7AE44FB72B72938" pageId="0" pageNumber="51">
<emphasis id="8891B8D0D89ECC64D018098BF8B9A9DF" bold="true" italics="true" pageId="0" pageNumber="51">Holotype</emphasis>
: Male (with its associated pupal exuviae and cocoon) (in 80% ethanol) labeled as &quot;Holotype: 
<taxonomicName id="B4A97DA9FD05E9E08797F6A52987E91D" authorityName="Takaoka, Srisuka &amp; Saeung" authorityYear="2020" class="Insecta" family="Simuliidae" genus="Simulium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Simulium maewongense" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="maewongense">
<emphasis id="180D6228B175BD4E16683D8B7E6B0E08" italics="true" pageId="0" pageNumber="51">Simulium maewongense</emphasis>
</taxonomicName>
male, QSBG col. no. 80, Thailand, 12-VII-2018, by W. Srisuka&quot;, collected from a small stream (width 20 cm, depth 2 cm bed sandy, slow flow, 19.5 °C, partially shaded, elevation 1,322 m, 
<geoCoordinate id="69DB7A1497265AD49B4EB497BC75E284" degrees="16" direction="north" minutes="06" orientation="latitude" precision="1" seconds="02.9" value="16.100805">16°06'02.9&quot;N</geoCoordinate>
, 
<geoCoordinate id="EA11CB64BDAA7C94D31BC4158459A42D" degrees="99" direction="east" minutes="06" orientation="longitude" precision="1" seconds="23.8" value="99.10661">99°06'23.8&quot;E</geoCoordinate>
, 98°30'53.0&quot;E), at Chon Yen, Mae Wong National Park, Klong Lan District, Kham Phaeng Phet Province, Thailand, 12-VII-2018, by W. Srisuka (Coll. No. 80).
</paragraph>
<paragraph id="F93FF1C980355522E07C1B2464E2536E" pageId="0" pageNumber="51">
<emphasis id="13C74496076D75ECE6467C04384A645A" bold="true" italics="true" pageId="0" pageNumber="51">Paratypes</emphasis>
: Three females, four males (thorax of one male for DNA analysis) (with their associated pupal exuviae and cocoons), and 14 mature larvae (two mature larvae for DNA analysis) (in 80% ethanol), same data as for holotype; one female (with its associated pupal exuviae and cocoon). (in 80% ethanol), collected from a stream of Klong Nam Lai (width 1.4 m, depth 13 cm, bed sandy, moderate flow, pH 6.23, 25.8 °C, exposed to the sun, elevation 196 m, 
<geoCoordinate id="ACD1CEB4A63D1FA81958209F4C36D463" degrees="16" direction="north" minutes="12" orientation="latitude" precision="1" seconds="28.3" value="16.20786">16°12'28.3&quot;N</geoCoordinate>
, 
<geoCoordinate id="F2503AD4A7224D158D0F3FEA60C90D18" degrees="99" direction="east" minutes="15" orientation="longitude" precision="1" seconds="47.8" value="99.263275">99°15'47.8&quot;E</geoCoordinate>
), at Klong Lan District, Kham Phaeng Phet Province, Thailand, 27-VI-2013, by W. Srisuka (Coll. No. 144).
</paragraph>
</subSubSection>
<subSubSection id="69508BA3767C776858B0A06B7B41561D" pageId="0" pageNumber="51" type="diagnosis">
<paragraph id="D05C1FA0C02147ED1B4C77E03E7C9636" pageId="0" pageNumber="51">Diagnosis.</paragraph>
<paragraph id="153BA75D65C7E14338BFE5025AF78976" pageId="0" pageNumber="51">
Female: mandible lacking teeth on the outer margin. Male: upper-eye (large) facets in eleven vertical columns and 13 or 14 horizontal rows. Pupa: thorax bare except the anterior two-fifths or half and small area of the dorsal surface near the posterior margin moderately covered with tubercles, and cocoon with a short anterodorsal projection or bulge (Fig. 
<figureCitation id="043FBD58FF9EE4EB35EE327C8C8F94C3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male and pupa of S. maewongense sp. nov. A-D male E-G pupa. A hind basitarsus and second tarsomere (left side; lateral view) B coxites, styles and ventral plate (ventral view) C ventral plate and median sclerite (lateral view) D ventral plate (caudal view) E gill filaments (right side; lateral view) F terminal hooks (caudal view) G cocoon (dorsal view). Scale bars: 1.0 mm (G); 0.1 mm (A, E); 0.02 mm (B-D); 0.01 mm (F)." figureDoi="10.3897/zookeys.950.51298.figure7" httpUri="https://binary.pensoft.net/fig/433871" pageId="0" pageNumber="51">7G</figureCitation>
). Larva: abdominal segments 1 and 2 greyish (Fig. 
<figureCitation id="13F906DF4CAC6582D063B441C256C59B" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Schematic illustrations of larval body color patterns (dorsal view). A S. teerachanense sp. nov. B S. pitasawatae sp. nov. C S. sutheppuiense sp. nov. D S. asakoae E S. klonglanense sp. nov. F S. namdanense sp. nov. G S. chaowaense sp. nov. H S. loeiense sp. nov. I S. huaimorense sp. nov. J S. nanoiense sp. nov. K S. junkumae sp. nov. L S. myanmarense M S. maewongense sp. nov. N S. phapeungense sp. nov. O S. muangpanense sp. nov. P S. maehongsonense sp. nov. Q S. thungchangense sp. nov. R S. chiangdaoense S S. inthanonense (morph 1) T S. inthanonense (morph 2)." figureDoi="10.3897/zookeys.950.51298.figure25" httpUri="https://binary.pensoft.net/fig/433889" pageId="0" pageNumber="51">25M</figureCitation>
), and the postgenal cleft medium-long, 0.9-1.2 times as long as the postgenal bridge.
</paragraph>
<caption id="B28DAC508196EABD09E91208ACBC8380" doi="10.3897/zookeys.950.51298.figure7" httpUri="https://binary.pensoft.net/fig/433871" pageId="0" pageNumber="51" start="Figure 7" startId="F7">
<paragraph id="D483C02A5C98D1D3B0513CB0AB2FB3FF" pageId="0" pageNumber="51">
<emphasis id="49A658213392392667201048B4384ADA" bold="true" pageId="0" pageNumber="51">Figure 7.</emphasis>
Male and pupa of 
<taxonomicName id="F6C43569C7FCD0C31AA2415895777692" lsidName="S. maewongense" pageId="0" pageNumber="51" rank="species" species="maewongense">
<emphasis id="B66FCB0864E21F871C3553D2DC39F3BA" italics="true" pageId="0" pageNumber="51">S. maewongense</emphasis>
</taxonomicName>
sp. nov. 
<emphasis id="25FFA6FD520CE697D4D80C3FE1905F92" bold="true" pageId="0" pageNumber="51">A-D</emphasis>
male 
<emphasis id="BC0448EA82357CC2B7CABB79330BF2EF" bold="true" pageId="0" pageNumber="51">E-G</emphasis>
pupa. 
<emphasis id="30322930CCE615B852DEF64210B16337" bold="true" pageId="0" pageNumber="51">A</emphasis>
hind basitarsus and second tarsomere (left side; lateral view) 
<emphasis id="126228A4116B75E9CA4CD1CE4E529CF6" bold="true" pageId="0" pageNumber="51">B</emphasis>
coxites, styles and ventral plate (ventral view) 
<emphasis id="3F012D6AA0769B9DFBA36C33E8B9FE5F" bold="true" pageId="0" pageNumber="51">C</emphasis>
ventral plate and median sclerite (lateral view) 
<emphasis id="6BC0374A2FB03DBEAB68F14F5AAB5D5E" bold="true" pageId="0" pageNumber="51">D</emphasis>
ventral plate (caudal view) 
<emphasis id="EC3853CC6063910291C08170DEA989D7" bold="true" pageId="0" pageNumber="51">E</emphasis>
gill filaments (right side; lateral view) 
<emphasis id="38FB0401695FA3C8BE3E78B0E4201E92" bold="true" pageId="0" pageNumber="51">F</emphasis>
terminal hooks (caudal view) 
<emphasis id="D8D93231AB53A715F431A67169F59D68" bold="true" pageId="0" pageNumber="51">G</emphasis>
cocoon (dorsal view). Scale bars: 1.0 mm (
<emphasis id="C941A9E3D62E0988B20EDC79838D0591" bold="true" pageId="0" pageNumber="51">G</emphasis>
); 0.1 mm (
<emphasis id="680420E5CD4C5F338FC9DD8908C34D44" bold="true" pageId="0" pageNumber="51">A, E</emphasis>
); 0.02 mm (
<emphasis id="2DEC4459A783AFAB813C1B85547569F2" bold="true" pageId="0" pageNumber="51">B-D</emphasis>
); 0.01 mm (
<emphasis id="3EEDE8E3FF48E9C852F8E56C008FDCDA" bold="true" pageId="0" pageNumber="51">F</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="F1DB717E468AEADA69179DD0C1C40BC3" pageId="0" pageNumber="51" type="description">
<paragraph id="F6ECBC9AF901BD5560E7527CE0510808" pageId="0" pageNumber="51">Description.</paragraph>
<paragraph id="B1736B8156FA7132B78896804E8CC853" pageId="0" pageNumber="51">
<emphasis id="AE9E242EC9F2369300E4CFD4090B43B3" bold="true" pageId="0" pageNumber="51">Female</emphasis>
(
<emphasis id="E024820A3387315A8B8342383FAD4EDB" italics="true" pageId="0" pageNumber="51">N</emphasis>
= 4). Body length 2.1-2.2 mm.
</paragraph>
<paragraph id="671DC6442E83D2C5872A011EC1D296C8" pageId="0" pageNumber="51">
<emphasis id="488EFCABB9146446E5589AAA4B7D9610" bold="true" italics="true" pageId="0" pageNumber="51">Head.</emphasis>
Frontal ratio 1.8-2.0:1.0:2.3-3.3. Frons: head ratio 1.0:4.3-5.4. Labrum 0.68 times length of clypeus. Maxillary palpus: proportional length of third, fourth and fifth palpal segments 1.0:1.0-1.1:2.2-2.3; sensory vesicle medium-long, 0.36 times as long as third palpal segment. Lacinia with 8-11 inner and 12 or 13 outer teeth. Mandible with 21-23 teeth on inner margin and lacking teeth on outer margin (though outer margin undulated).
</paragraph>
<paragraph id="1430AF4A7593CA97A3D15873B4581391" pageId="0" pageNumber="51">
<emphasis id="B71C3164517B687645E3A13E2E9701AB" bold="true" italics="true" pageId="0" pageNumber="51">Legs.</emphasis>
Fore basitarsus 6.3-6.5 times as long as its greatest width. Hind basitarsus 5.9-6.1 times as long as its greatest width and 0.7 and 0.6 times as wide as greatest width of hind tibia and femur, respectively. Calcipala nearly as long as wide and 0.56 times as wide as greatest width of hind basitarsus.
</paragraph>
<paragraph id="7132F28A97058DC2BCCA214B67BB4BCC" pageId="0" pageNumber="51">
<emphasis id="E0CCAA86FB9A4413B2D7D7F156F1E48A" bold="true" italics="true" pageId="0" pageNumber="51">Wing.</emphasis>
Length 2.2-2.3 mm.
</paragraph>
<paragraph id="8296C0C5D4039E49F723D1BF3E25FF69" pageId="0" pageNumber="51">
<emphasis id="AC934F91BB6DBE0B29549A15E27E8DF3" bold="true" italics="true" pageId="0" pageNumber="51">Terminalia.</emphasis>
Sternite 8 with 17-22 medium to long stout hairs and three to five short slender hairs. Ovipositor valve with one to four short hairs. Paraproct 0.5 times as long as wide, and with 22-25 short to medium-long hairs on outer surface. Cercus 0.4 times as long as wide. Spermatheca ellipsoidal, 1.4 times as long as its greatest width.
</paragraph>
<paragraph id="86090B87B704AADAE5B793834C604D35" pageId="0" pageNumber="51">
<emphasis id="80CCBCA4DBE3A9178B6A04C1549CE102" bold="true" pageId="0" pageNumber="51">Male</emphasis>
(
<emphasis id="E1842B00D49BF6B53C5B41E35F931010" italics="true" pageId="0" pageNumber="51">N</emphasis>
= 5). Body length 2.6 mm.
</paragraph>
<paragraph id="FECDDCE55F5AD875A4429BCDA2A70367" pageId="0" pageNumber="51">
<emphasis id="1AE93CDEBD9580AC258C24C8352EEA16" bold="true" italics="true" pageId="0" pageNumber="51">Head.</emphasis>
Upper eye dark brown, consisting of large facets in eleven vertical columns and 13 or 14 horizontal rows on each side. Antenna: first flagellomere 1.6 times as long as second. Maxillary palpus: proportional length of third, fourth and fifth palpal segments 1.0:1.2:2.8; sensory vesicle small, globular or ellipsoidal, 0.19-0.21 times as long as palpal segment 3.
</paragraph>
<paragraph id="17FFADCC852BA2553A1EA42C7D017058" pageId="0" pageNumber="51">
<emphasis id="D069437E78607E141BB965DA9F66C37D" bold="true" italics="true" pageId="0" pageNumber="51">Legs.</emphasis>
Foreleg: basitarsus 7.7 times as long as its greatest width. Hind leg: basitarsus (Fig. 
<figureCitation id="88A6AA222D16E2AE72897FC2D73A1969" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male and pupa of S. maewongense sp. nov. A-D male E-G pupa. A hind basitarsus and second tarsomere (left side; lateral view) B coxites, styles and ventral plate (ventral view) C ventral plate and median sclerite (lateral view) D ventral plate (caudal view) E gill filaments (right side; lateral view) F terminal hooks (caudal view) G cocoon (dorsal view). Scale bars: 1.0 mm (G); 0.1 mm (A, E); 0.02 mm (B-D); 0.01 mm (F)." figureDoi="10.3897/zookeys.950.51298.figure7" httpUri="https://binary.pensoft.net/fig/433871" pageId="0" pageNumber="51">7A</figureCitation>
) dark brown except little less than basal half yellowish white, and enlarged, 3.5 times as long as its greatest width, and 1.0 and 1.2-1.3 times as wide as greatest width of hind tibia and femur, respectively. Calcipala (Fig. 
<figureCitation id="A6D4D1B1BEF0351B89F1E9127002B81B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male and pupa of S. maewongense sp. nov. A-D male E-G pupa. A hind basitarsus and second tarsomere (left side; lateral view) B coxites, styles and ventral plate (ventral view) C ventral plate and median sclerite (lateral view) D ventral plate (caudal view) E gill filaments (right side; lateral view) F terminal hooks (caudal view) G cocoon (dorsal view). Scale bars: 1.0 mm (G); 0.1 mm (A, E); 0.02 mm (B-D); 0.01 mm (F)." figureDoi="10.3897/zookeys.950.51298.figure7" httpUri="https://binary.pensoft.net/fig/433871" pageId="0" pageNumber="51">7A</figureCitation>
) as long as wide and 0.3 times as wide as greatest width of hind basitarsus.
</paragraph>
<paragraph id="58739965E88FBC68B4C4A928C32BC383" pageId="0" pageNumber="51">
<emphasis id="52E4960911E49516198390B7A3BD603E" bold="true" italics="true" pageId="0" pageNumber="51">Wing.</emphasis>
Length 2.2 mm. Subcosta haired except near apex bare.
</paragraph>
<paragraph id="D4A887AB8D7C659A2CD3B0F0AE2C1EA1" pageId="0" pageNumber="51">
<emphasis id="1CB1A3F0D20679B405BA01E21FE6D8D1" bold="true" italics="true" pageId="0" pageNumber="51">Genitalia.</emphasis>
Ventral plate in ventral view (Fig. 
<figureCitation id="F958D4EA9629A2D39E01ABB00E5AECCE" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male and pupa of S. maewongense sp. nov. A-D male E-G pupa. A hind basitarsus and second tarsomere (left side; lateral view) B coxites, styles and ventral plate (ventral view) C ventral plate and median sclerite (lateral view) D ventral plate (caudal view) E gill filaments (right side; lateral view) F terminal hooks (caudal view) G cocoon (dorsal view). Scale bars: 1.0 mm (G); 0.1 mm (A, E); 0.02 mm (B-D); 0.01 mm (F)." figureDoi="10.3897/zookeys.950.51298.figure7" httpUri="https://binary.pensoft.net/fig/433871" pageId="0" pageNumber="51">7B</figureCitation>
) somewhat emarginated on each lateral margin; ventral plate in caudal view (Fig. 
<figureCitation id="7DECFF60EDAFEE83890920F3739CE50E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male and pupa of S. maewongense sp. nov. A-D male E-G pupa. A hind basitarsus and second tarsomere (left side; lateral view) B coxites, styles and ventral plate (ventral view) C ventral plate and median sclerite (lateral view) D ventral plate (caudal view) E gill filaments (right side; lateral view) F terminal hooks (caudal view) G cocoon (dorsal view). Scale bars: 1.0 mm (G); 0.1 mm (A, E); 0.02 mm (B-D); 0.01 mm (F)." figureDoi="10.3897/zookeys.950.51298.figure7" httpUri="https://binary.pensoft.net/fig/433871" pageId="0" pageNumber="51">7D</figureCitation>
) rounded ventrally. Cercus with 15-17 hairs.
</paragraph>
<paragraph id="CDFC4D2E431A253A1D6982CC3D230C7C" pageId="0" pageNumber="51">
<emphasis id="173AC9413671383F88627527064E471A" bold="true" pageId="0" pageNumber="51">Pupa</emphasis>
(
<emphasis id="74BE1C9E99DA1A43CA7B059CF53114EE" italics="true" pageId="0" pageNumber="51">N</emphasis>
= 9). Body length 3.0 mm.
</paragraph>
<paragraph id="3FD88F9B323C596E0AB9A0A8A61ECB46" pageId="0" pageNumber="51">
<emphasis id="187C688DE21E5835DCEBD317766E0A77" bold="true" italics="true" pageId="0" pageNumber="51">Thorax.</emphasis>
Integument almost bare except anterior two-fifths to half moderately covered with round tubercles and small dorsal area near posterior margin sparsely covered with tubercles. Gill (Fig. 
<figureCitation id="025AB36F8522DE8B49C6D023C98D093D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male and pupa of S. maewongense sp. nov. A-D male E-G pupa. A hind basitarsus and second tarsomere (left side; lateral view) B coxites, styles and ventral plate (ventral view) C ventral plate and median sclerite (lateral view) D ventral plate (caudal view) E gill filaments (right side; lateral view) F terminal hooks (caudal view) G cocoon (dorsal view). Scale bars: 1.0 mm (G); 0.1 mm (A, E); 0.02 mm (B-D); 0.01 mm (F)." figureDoi="10.3897/zookeys.950.51298.figure7" httpUri="https://binary.pensoft.net/fig/433871" pageId="0" pageNumber="51">7E</figureCitation>
) composed of eight slender thread-like filaments, arranged as [3+(1+2)]+2or [(2+1)+(1+2)]+2 or [(2+1)+3]+2 from dorsal to ventral, with medium-long common basal stalk having somewhat swollen transparent basal fenestra; common basal stalk 0.6-0.7 times length of interspiracular trunk; dorsal and middle triplets sharing short stalk, and dorsal triplet mostly composed of three individual filaments arising at same level, middle triplet mostly composed of one individual and two paired filaments with extremely short secondary stalk; stalk of ventral pair of filaments, 0.9-1.3 times length of common basal stalk, and 0.6-0.9 times length of interspiracular trunk, and 0.9 times as thick as common stalk of middle and dorsal triplets; primary stalk of dorsal triplet lying against that of lower pair at angle of 60-90° when viewed laterally; filaments of dorsal and middle triplets subequal in length (1.8-2.2 mm) and thickness to one another; two filaments of ventral pair subequal in length (2.8-3.2 mm) and thickness to each other and 1.6 times as thick as six other filaments of dorsal and middle triplets when compared basally.
</paragraph>
<paragraph id="A864BFB5753452971980C88E95589DCB" pageId="0" pageNumber="51">
<emphasis id="18DEB89546311FDF46849D34D6C1C9B8" bold="true" italics="true" pageId="0" pageNumber="51">Abdomen.</emphasis>
Dorsally, segments 1 and 2 bare. Terminal hooks (Fig. 
<figureCitation id="9DA1E4E3EDE1DA77DFDBF533E6757A0C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male and pupa of S. maewongense sp. nov. A-D male E-G pupa. A hind basitarsus and second tarsomere (left side; lateral view) B coxites, styles and ventral plate (ventral view) C ventral plate and median sclerite (lateral view) D ventral plate (caudal view) E gill filaments (right side; lateral view) F terminal hooks (caudal view) G cocoon (dorsal view). Scale bars: 1.0 mm (G); 0.1 mm (A, E); 0.02 mm (B-D); 0.01 mm (F)." figureDoi="10.3897/zookeys.950.51298.figure7" httpUri="https://binary.pensoft.net/fig/433871" pageId="0" pageNumber="51">7F</figureCitation>
) with outer margin 2.0 times as long as inner margin.
</paragraph>
<paragraph id="EB46F20E969405A4E31301A717F55CA2" pageId="0" pageNumber="51">
<emphasis id="F691B5967D047075417ACEA4F0EDED54" bold="true" italics="true" pageId="0" pageNumber="51">Cocoon</emphasis>
(Fig. 
<figureCitation id="8FCB86013C4CF0B28B84581461DE61C1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male and pupa of S. maewongense sp. nov. A-D male E-G pupa. A hind basitarsus and second tarsomere (left side; lateral view) B coxites, styles and ventral plate (ventral view) C ventral plate and median sclerite (lateral view) D ventral plate (caudal view) E gill filaments (right side; lateral view) F terminal hooks (caudal view) G cocoon (dorsal view). Scale bars: 1.0 mm (G); 0.1 mm (A, E); 0.02 mm (B-D); 0.01 mm (F)." figureDoi="10.3897/zookeys.950.51298.figure7" httpUri="https://binary.pensoft.net/fig/433871" pageId="0" pageNumber="51">7G</figureCitation>
). Slipper-shaped, moderately woven, widely extended ventrolaterally; anterior margin thickly woven medially, with bulge or short projection; 3.0-4.0 mm long by 2.5-2.8 mm wide.
</paragraph>
<paragraph id="9BE42F7C47C903637AD3560E85CE7B36" pageId="0" pageNumber="51">
<emphasis id="2301955B3DA4413352F128CB35025EA8" bold="true" pageId="0" pageNumber="51">Mature larva</emphasis>
(
<emphasis id="5D8B0AF5B16271324924D9A577F380F0" italics="true" pageId="0" pageNumber="51">N</emphasis>
= 12). Body length 5.5-6.5 mm. Body creamy white to light ochreous with following color markings: thoracic segment 1 encircled with ochreous or reddish brown band (though disconnected ventromedially), thoracic segments 2 and 3 ochreous on ventral surface; abdominal segments 1 and 2 entirely grey, dorsal and dorsolateral surface of abdominal segment 4 with faint reddish brown band or spot (though completely faded in some larvae), abdominal segments 5 and 6 each with reddish brown, W-shaped, transverse band near posterior margin of dorsal and dorsolateral surface, which is always distinct on abdominal segment 5 but is faded to varying extent on abdominal segment 6 leaving one small round dorsomedial spot and two distinct lateral areas, abdominal segments 7 and 8 distinctly covered with reddish brown pigment on each dorsolateral surface (Fig. 
<figureCitation id="F1844D85243C1C4B29B43EBC90916B74" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Schematic illustrations of larval body color patterns (dorsal view). A S. teerachanense sp. nov. B S. pitasawatae sp. nov. C S. sutheppuiense sp. nov. D S. asakoae E S. klonglanense sp. nov. F S. namdanense sp. nov. G S. chaowaense sp. nov. H S. loeiense sp. nov. I S. huaimorense sp. nov. J S. nanoiense sp. nov. K S. junkumae sp. nov. L S. myanmarense M S. maewongense sp. nov. N S. phapeungense sp. nov. O S. muangpanense sp. nov. P S. maehongsonense sp. nov. Q S. thungchangense sp. nov. R S. chiangdaoense S S. inthanonense (morph 1) T S. inthanonense (morph 2)." figureDoi="10.3897/zookeys.950.51298.figure25" httpUri="https://binary.pensoft.net/fig/433889" pageId="0" pageNumber="51">25M</figureCitation>
), and ventral surface of abdominal segments 6 and 7 each with two light reddish brown spots (though absent in some larvae).
</paragraph>
<paragraph id="EC5693BCA43F308FCE6AF2860F16B934" pageId="0" pageNumber="51">
<emphasis id="BDCE13DC4848D15BCBC2A2C342BCA076" bold="true" italics="true" pageId="0" pageNumber="51">Head.</emphasis>
Head spots faintly (or rarely moderately) positive or indistinct. Antenna: proportional lengths of first, second, and third articles 1.0:0.8:0.7-0.8. Labral fan with 27-29 primary rays. Hypostoma: median tooth little longer than each corner tooth. Postgenal cleft small, rounded, 0.9-1.2 times length of postgenal bridge.
</paragraph>
<paragraph id="91AE2D0E8B77FA38CDE1CC1DC716529B" pageId="0" pageNumber="51">
<emphasis id="F15CDAE6D6E83D8FEF63D90BF07E1475" bold="true" italics="true" pageId="0" pageNumber="51">Abdomen.</emphasis>
Rectal organ compound, each of three lobes with 6-8 finger-like secondary lobules. Anal sclerite with anterior arms 1.1-1.2 times as long as posterior ones. Posterior circlet with 91-96 rows of hooklets with up to 14 or 15 hooklets per row.
</paragraph>
</subSubSection>
<subSubSection id="77B8C504E636D2822189414F38E396DC" pageId="0" pageNumber="51" type="etymology">
<paragraph id="C1162B555CF93BD54F600B7AD7A0BB9F" pageId="0" pageNumber="51">Etymology.</paragraph>
<paragraph id="6B5A8FDE4CC7273D31C54467046753DB" pageId="0" pageNumber="51">
The species name, 
<taxonomicName id="A20940CBF31A3E87F513C38B112AFC3C" lsidName="maewongense" pageId="0" pageNumber="51" rank="species" species="maewongense">
<emphasis id="806376FAE3F752147785E10001E3A156" italics="true" pageId="0" pageNumber="51">maewongense</emphasis>
</taxonomicName>
, refers to the name of the national park, Mae Wong, where this species was collected.
</paragraph>
</subSubSection>
<subSubSection id="944F5BC7F47940A39FAF3CC23106AB33" pageId="0" pageNumber="51" type="distribution">
<paragraph id="DA810EA9F74BD44F49657E639CD66F1F" pageId="0" pageNumber="51">Distribution.</paragraph>
<paragraph id="0D28CCC4CAA50A1DA85955311C0FDBDB" pageId="0" pageNumber="51">Thailand (Kham Phaeng Phet).</paragraph>
</subSubSection>
<subSubSection id="A3CBACA3738E239EDE1391AD625B949D" pageId="0" pageNumber="51" type="discussion">
<paragraph id="72D1F6730AC245BE41EDDD353392F819" pageId="0" pageNumber="51">Discussion.</paragraph>
<paragraph id="D318839953CFE2504F2F68776451E015" pageId="0" pageNumber="51">
Among 36 species of the 
<taxonomicName id="31D960530C4257194B6DCC7BB87C1601" lsidName="S. asakoae" pageId="0" pageNumber="51" rank="species" species="asakoae">
<emphasis id="15A44F97686F2297A84C47393A86A1E0" italics="true" pageId="0" pageNumber="51">S. asakoae</emphasis>
</taxonomicName>
species group, 
<taxonomicName id="DF1BCF7AD8164CE7E41AAED9C7563C5C" lsidName="S. gyorkosae" pageId="0" pageNumber="51" rank="species" species="gyorkosae">
<emphasis id="927BED23C966633BB4D2A9C57BA55673" italics="true" pageId="0" pageNumber="51">S. gyorkosae</emphasis>
</taxonomicName>
Takaoka &amp; Davies from Indonesia (
<bibRefCitation id="E493397708D4EE892DE27C37C04A0922" author="Takaoka, H" journalOrPublisher="Medical Entomology and Zoology" pageId="0" pageNumber="51" refId="B18" refString="Takaoka, H, Davies, DM, 1996. The Black Flies (Diptera: Simuliidae) of Java, Indonesia. Bishop Museum Bulletin in Entomology 6, Bishop Museum Press, Honolulu, viii + 81 pp." title="The Black Flies (Diptera: Simuliidae) of Java, Indonesia. Bishop Museum Bulletin in Entomology 6, Bishop Museum Press, Honolulu, viii + 81 pp." year="1996">Takaoka and Davies 1996</bibRefCitation>
), 
<taxonomicName id="E8E3FEF80C42DF71E7A6EB211253F5D6" lsidName="S. jianfengense" pageId="0" pageNumber="51" rank="species" species="jianfengense">
<emphasis id="C4E1D633A46AAD06DA064A650D227577" italics="true" pageId="0" pageNumber="51">S. jianfengense</emphasis>
</taxonomicName>
Long et al. from Hainan Island, China (
<bibRefCitation id="0DC417E0A895FFE8CC1AC1FEFA3E18E2" author="Long, ZM" journalOrPublisher="Chinese Journal of Vector Biology and Control" pageId="0" pageNumber="51" pagination="405 - 409" publicationUrl="https://www.biotaxa.org/Zootaxa/article/view/zootaxa.3838.3.6" refId="B8" refString="Long, ZM, An, JY, Hao, BS, Yan, G, Yuan, WH, 1994. Two new blackflies of Simulium (Gomphostilbia) from Hainan Island, China (Diptera, Simuliidae). Chinese Journal of Vector Biology and Control 5: 405 - 409, https://www.biotaxa.org/Zootaxa/article/view/zootaxa.3838.3.6" title="Two new blackflies of Simulium (Gomphostilbia) from Hainan Island, China (Diptera, Simuliidae)." url="https://www.biotaxa.org/Zootaxa/article/view/zootaxa.3838.3.6" volume="5" year="1994">Long et al. 1994</bibRefCitation>
), 
<taxonomicName id="946806B3E9150C3EB8B890D439E04E95" lsidName="S. myanmarense" pageId="0" pageNumber="51" rank="species" species="myanmarense">
<emphasis id="4B7540492645A6E31E13F0A7BB70D00D" italics="true" pageId="0" pageNumber="51">S. myanmarense</emphasis>
</taxonomicName>
from Myanmar (
<bibRefCitation id="3789A966F7B732D2CC392ACEBA480FD8" DOI="https://doi.org/10.1016/j.actatropica.2017.07.024" author="Takaoka, H" journalOrPublisher="Acta Tropica" pageId="0" pageNumber="51" pagination="39 - 50" refId="B27" refString="Takaoka, H, Srisuka, W, Low, VL, Saeung, A, 2017b. Two new species of the Simulium asakoae species-group of Simulium (Gomphostilbia) (Diptera: Simuliidae) from Myanmar and their phylogenetic relationships within the S. asakoae species-group. Acta Tropica 176: 39 - 50, DOI: https://doi.org/10.1016/j.actatropica.2017.07.024" title="Two new species of the Simulium asakoae species-group of Simulium (Gomphostilbia) (Diptera: Simuliidae) from Myanmar and their phylogenetic relationships within the S. asakoae species-group." url="https://doi.org/10.1016/j.actatropica.2017.07.024" volume="176" year="2017 b">Takaoka et al. 2017b</bibRefCitation>
), 
<taxonomicName id="88EB5C2FA42F243CE421DEC64FCAAB23" lsidName="S. yunnanense" pageId="0" pageNumber="51" rank="species" species="yunnanense">
<emphasis id="371C7659305EEE66B2249255F16E0DB6" italics="true" pageId="0" pageNumber="51">S. yunnanense</emphasis>
</taxonomicName>
Chen &amp; Zhang from Yunnan, China (
<bibRefCitation id="6ABB33FC3BD952332DFCB0025FBCD22D" author="Chen, HB" journalOrPublisher="Acta Parasitology et Medica Entomologica Sinica" pageId="0" pageNumber="51" pagination="87 - 90" refId="B3" refString="Chen, HB, Zhang, CL, 2004. A new species of Simulium (Gomphostilbia) from Yunnan Province, China (Diptera: Simuliidae). Acta Parasitology et Medica Entomologica Sinica 11: 87 - 90" title="A new species of Simulium (Gomphostilbia) from Yunnan Province, China (Diptera: Simuliidae)." volume="11" year="2004">Chen and Zhang 2004</bibRefCitation>
), and 
<taxonomicName id="E0934D7EC9AB330B1D5357F1C796AE83" lsidName="S. phulocense" pageId="0" pageNumber="51" rank="species" species="phulocense">
<emphasis id="F4C61ECE9E20EFB472EC65D7872DABDC" italics="true" pageId="0" pageNumber="51">S. phulocense</emphasis>
</taxonomicName>
Takaoka &amp; Chen and 
<taxonomicName id="05F7CCE7C28A703E9DE0AB42B99B8264" lsidName="S. unii" pageId="0" pageNumber="51" rank="species" species="unii">
<emphasis id="2AA08439CCE008B5F7A7D6C1BF766BA1" italics="true" pageId="0" pageNumber="51">S. unii</emphasis>
</taxonomicName>
Takaoka &amp; Pham, both from Vietnam (
<bibRefCitation id="23CF27683BBE877F7DD175D0892D6D5C" DOI="https://doi.org/10.11646/zootaxa.3961.1.1" author="Takaoka, H" journalOrPublisher="Zootaxa" pageId="0" pageNumber="51" pagination="1 - 96" refId="B24" refString="Takaoka, H, Sofian-Azirun, M, Ya'cob, Z, Chen, CD, Lau, KW, Pham, XD, 2015. The black flies (Diptera: Simuliidae) from Thua Thien Hue and Lam Dong Provinces, Vietnam. Zootaxa 3961: 1 - 96, DOI: https://doi.org/10.11646/zootaxa.3961.1.1" title="The black flies (Diptera: Simuliidae) from Thua Thien Hue and Lam Dong Provinces, Vietnam." url="https://doi.org/10.11646/zootaxa.3961.1.1" volume="3961" year="2015">Takaoka et al. 2015</bibRefCitation>
, 
<bibRefCitation id="391FD1F9D83870FB63092FA923FF9F1F" DOI="https://doi.org/10.11646/zootaxa.4261.1.1" author="Takaoka, H" journalOrPublisher="ZooKeys" pageId="0" pageNumber="51" refId="B22" refString="Takaoka, H, Sofian-Azirun, M, Ya'cob, Z, Chen, CD, Lau, KW, Low, VL, Pham, XD, Adler, PH, 2017a. The black flies (Diptera: Simuliidae) of Vietnam. Zootaxa 4261, 1-165. https://doi.org/10.11646/zootaxa.4261.1.1" title="The black flies (Diptera: Simuliidae) of Vietnam. Zootaxa 4261, 1 - 165." url="https://doi.org/10.11646/zootaxa.4261.1.1" year="2017 a">2017a</bibRefCitation>
), have a similar cocoon with a short anterodorsal projection. However, 
<taxonomicName id="4BA29AEF7131CBAC498668CBF9B9EE08" lsidName="S. maewongense" pageId="0" pageNumber="51" rank="species" species="maewongense">
<emphasis id="D3D388AA4FC15D456169AF3F61755879" italics="true" pageId="0" pageNumber="51">S. maewongense</emphasis>
</taxonomicName>
sp. nov. is distinguished from these six known species by the following characters (those of each related species in parentheses): from 
<taxonomicName id="2003B102067715D3509AD9EEA16F60CA" lsidName="S. gyorkosae" pageId="0" pageNumber="51" rank="species" species="gyorkosae">
<emphasis id="601575225D03727CD2A1FA5B6257D3CC" italics="true" pageId="0" pageNumber="51">S. gyorkosae</emphasis>
</taxonomicName>
by the female mandible lacking outer teeth (four or five outer teeth); from 
<taxonomicName id="64305B1DC7F8F79F486D1C09D4605BD5" lsidName="myanmarense" pageId="0" pageNumber="51" rank="species" species="myanmarense">
<emphasis id="E98B8839F1EF3BF1FB764BBAE6715702" italics="true" pageId="0" pageNumber="51">myanmarense</emphasis>
</taxonomicName>
by the number of male upper-eye facets in eleven vertical columns (15 or rarely 14 vertical columns); from 
<taxonomicName id="F20AD8C14310642C4FE4003861C84E3C" lsidName="S. jianfengense" pageId="0" pageNumber="51" rank="species" species="jianfengense">
<emphasis id="9F3F3129C71A2B162B0F890F7B88E4E5" italics="true" pageId="0" pageNumber="51">S. jianfengense</emphasis>
</taxonomicName>
by the ventral plate produced anteromedially when viewed ventrally (Fig. 
<figureCitation id="F647B3D65E039FB7CC230E2471EE500A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male and pupa of S. maewongense sp. nov. A-D male E-G pupa. A hind basitarsus and second tarsomere (left side; lateral view) B coxites, styles and ventral plate (ventral view) C ventral plate and median sclerite (lateral view) D ventral plate (caudal view) E gill filaments (right side; lateral view) F terminal hooks (caudal view) G cocoon (dorsal view). Scale bars: 1.0 mm (G); 0.1 mm (A, E); 0.02 mm (B-D); 0.01 mm (F)." figureDoi="10.3897/zookeys.950.51298.figure7" httpUri="https://binary.pensoft.net/fig/433871" pageId="0" pageNumber="51">7B</figureCitation>
) (not produced), and with its ventral margin rounded when viewed posteriorly (Fig. 
<figureCitation id="C5B17D07D518A58429687B09FF83C9E6" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male and pupa of S. maewongense sp. nov. A-D male E-G pupa. A hind basitarsus and second tarsomere (left side; lateral view) B coxites, styles and ventral plate (ventral view) C ventral plate and median sclerite (lateral view) D ventral plate (caudal view) E gill filaments (right side; lateral view) F terminal hooks (caudal view) G cocoon (dorsal view). Scale bars: 1.0 mm (G); 0.1 mm (A, E); 0.02 mm (B-D); 0.01 mm (F)." figureDoi="10.3897/zookeys.950.51298.figure7" httpUri="https://binary.pensoft.net/fig/433871" pageId="0" pageNumber="51">7D</figureCitation>
) (pointed ventrally); from 
<taxonomicName id="D6277E25A500C549CED4D0CE2346295B" lsidName="S. yunnanense" pageId="0" pageNumber="51" rank="species" species="yunnanense">
<emphasis id="F616D3BC3235C2AAAECA18B58487E760" italics="true" pageId="0" pageNumber="51">S. yunnanense</emphasis>
</taxonomicName>
by the yellow hair tuft of the base of the radial vein (black hair tuft); from 
<taxonomicName id="984F4CDD76D615959FC83CDFD97A247D" lsidName="S. phulocense" pageId="0" pageNumber="51" rank="species" species="phulocense">
<emphasis id="FEC07096906A1E8E81E6B4015CB3C274" italics="true" pageId="0" pageNumber="51">S. phulocense</emphasis>
</taxonomicName>
by the relative length of the female sensory vesicle against the third palpal segment 0.36 (0.26-0.27) and male upper-eye facets in eleven vertical columns (12 or 13 vertical columns); from 
<taxonomicName id="2E9E574A5E1F629BF823E799A02A1F43" lsidName="S. unii" pageId="0" pageNumber="51" rank="species" species="unii">
<emphasis id="8E9134894660EE66D3975A5E2BDB9E3E" italics="true" pageId="0" pageNumber="51">S. unii</emphasis>
</taxonomicName>
by upper-eye facets in eleven vertical columns (13 or 14 vertical columns), and the ventral plate with its lateral margins emarginated medially (Fig. 
<figureCitation id="ACE67D109316D16DCFCF374E5A058F6A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male and pupa of S. maewongense sp. nov. A-D male E-G pupa. A hind basitarsus and second tarsomere (left side; lateral view) B coxites, styles and ventral plate (ventral view) C ventral plate and median sclerite (lateral view) D ventral plate (caudal view) E gill filaments (right side; lateral view) F terminal hooks (caudal view) G cocoon (dorsal view). Scale bars: 1.0 mm (G); 0.1 mm (A, E); 0.02 mm (B-D); 0.01 mm (F)." figureDoi="10.3897/zookeys.950.51298.figure7" httpUri="https://binary.pensoft.net/fig/433871" pageId="0" pageNumber="51">7B</figureCitation>
) (ventral plate narrowed posteriorly).
</paragraph>
<paragraph id="CDAF6936786500098876CD1C61EF4BD7" pageId="0" pageNumber="51">
The larval body color pattern (Fig. 
<figureCitation id="405B26552384031E9CC2785AAB5C6167" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Schematic illustrations of larval body color patterns (dorsal view). A S. teerachanense sp. nov. B S. pitasawatae sp. nov. C S. sutheppuiense sp. nov. D S. asakoae E S. klonglanense sp. nov. F S. namdanense sp. nov. G S. chaowaense sp. nov. H S. loeiense sp. nov. I S. huaimorense sp. nov. J S. nanoiense sp. nov. K S. junkumae sp. nov. L S. myanmarense M S. maewongense sp. nov. N S. phapeungense sp. nov. O S. muangpanense sp. nov. P S. maehongsonense sp. nov. Q S. thungchangense sp. nov. R S. chiangdaoense S S. inthanonense (morph 1) T S. inthanonense (morph 2)." figureDoi="10.3897/zookeys.950.51298.figure25" httpUri="https://binary.pensoft.net/fig/433889" pageId="0" pageNumber="51">25M</figureCitation>
) of this new species is similar to that of 
<taxonomicName id="2B154B6CB546D7114417C9410C9584D3" lsidName="S. tuenense" pageId="0" pageNumber="51" rank="species" species="tuenense">
<emphasis id="A0BD4421F395045F9E84680CA437406D" italics="true" pageId="0" pageNumber="51">S. tuenense</emphasis>
</taxonomicName>
Takaoka from Taiwan (
<bibRefCitation id="762F12750F91648D86A7C2EB6852901A" author="Takaoka, H" journalOrPublisher="Pacific Insects" pageId="0" pageNumber="51" pagination="365 - 403" publicationUrl="http://hbs.bishopmuseum.org/pi/pdf/20%284%29-365.pdf" refId="B12" refString="Takaoka, H, 1979. The black flies of Taiwan (Diptera: Simuliidae). Pacific Insects 20: 365 - 403, http://hbs.bishopmuseum.org/pi/pdf/20%284%29-365.pdf" title="The black flies of Taiwan (Diptera: Simuliidae)." url="http://hbs.bishopmuseum.org/pi/pdf/20%284%29-365.pdf" volume="20" year="1979">Takaoka 1979</bibRefCitation>
), although 
<taxonomicName id="4A4B8B6D3AD5068F9F551286F8833118" lsidName="S. tuenense" pageId="0" pageNumber="51" rank="species" species="tuenense">
<emphasis id="B6A72358A92AC9EE5B6BB96AB6B200A7" italics="true" pageId="0" pageNumber="51">S. tuenense</emphasis>
</taxonomicName>
differs from this new species by the number of male upper-eye facets in 15 vertical columns and 15 or 16 horizontal rows and cocoon without an anterodorsal projection (
<bibRefCitation id="908C2AB9934E23D34F15048C26B80A8A" author="Huang, YT" journalOrPublisher="Tropical Biomedicine" pageId="0" pageNumber="51" pagination="577 - 588" publicationUrl="https://pubmed.ncbi.nlm.nih.gov/22433887" refId="B5" refString="Huang, YT, Takaoka, H, Otsuka, Y, Aoki, C, 2011. Simulium (Gomphostilbia) taitungense, a new species of black fly (Diptera: Simuliidae) from Taiwan, with description of the male of S. (G.) tuenense Takaoka. Tropical Biomedicine 28: 577 - 588, https://pubmed.ncbi.nlm.nih.gov/22433887" title="Simulium (Gomphostilbia) taitungense, a new species of black fly (Diptera: Simuliidae) from Taiwan, with description of the male of S. (G.) tuenense Takaoka." url="https://pubmed.ncbi.nlm.nih.gov/22433887" volume="28" year="2011">Huang et. al. 2011</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>