272 lines
50 KiB
XML
272 lines
50 KiB
XML
<document id="22427C3F781BA94B74B0D84214D52E1E" ID-DOI="10.5281/zenodo.13390843" ID-ISSN="1732-2421" ID-Zenodo-Dep="13390843" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="karina" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1723062031241" checkinUser="felipe" docAuthor="Schröder, Stefan & Soto, Francisco" docDate="2003" docId="F5521A73FF8A36013723F81BFD50E89D" docLanguage="en" docName="ActaPalaeontolPol.48.4.547-558.pdf" docOrigin="Acta Palaeontologica Polonica 48 (4)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Cantabriastraea cantabrica Schröder & Soto, 2003, gen. et" docType="treatment" docVersion="4" lastPageNumber="556" masterDocId="096B620BFF8C3608340FFF9EFFC9EE73" masterDocTitle="Lower Devonian (Emsian) rugose corals from the Cantabrian Mountains, northern Spain" masterLastPageNumber="558" masterPageNumber="547" pageNumber="553" updateTime="1724879454672" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="7E6A1F4CBCFDDDD809B2DE9919ADE028" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8889CC28141B4B4E5FA5CC95B3F202C6">
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<mods:title id="4348A667C4DC431A0F363B411E97A807">Lower Devonian (Emsian) rugose corals from the Cantabrian Mountains, northern Spain</mods:title>
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</mods:titleInfo>
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<mods:name id="6DB44C659A259E5CFE00EC35E5DF2252" type="personal">
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<mods:roleTerm id="1F0BD12F7B9E8A8D316E91E550A746FD">Author</mods:roleTerm>
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<mods:namePart id="526A320BF6FC2BF048C33B81DCC72EC0">Schröder, Stefan</mods:namePart>
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</mods:name>
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<mods:name id="079BD76197C49E0A87EAC2DB27B92CF5" type="personal">
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<mods:role id="F928F91F2A63F4426A4D28F9F67963C8">
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<mods:roleTerm id="1F81115CB8A99ACB64BC9F1C396C26B5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0BA8FF6EE0F6398A706B73114E30E12C">Soto, Francisco</mods:namePart>
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<mods:typeOfResource id="FF072D288A242935B69ADB54A1B5C086">text</mods:typeOfResource>
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<mods:titleInfo id="3292ACDE6FB9B099A7D0AD3ED7128421">
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<mods:title id="E68C9CBDD22A14BFECCA7C32E72DDBAC">Acta Palaeontologica Polonica</mods:title>
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</mods:titleInfo>
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<mods:part id="AC6AEA57DE6F96A0F584D38506A82D7A">
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<mods:date id="55326B06DEF7E0CDCB781B09FCFC5327">2003</mods:date>
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<mods:detail id="71519F181F9E6696DB0DA22EF494E1AC" type="pubDate">
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<mods:number id="AC8E1D37622714EA4183392CD7C969D4">2003-09-30</mods:number>
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<mods:detail id="DCF08989A1B3A196BE593C8AD5743906" type="volume">
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<mods:number id="09060EDBB231B4793833DC773EA1AE44">48</mods:number>
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</mods:detail>
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<mods:detail id="1B18086F3BE0389D0A7165A1D61F2D42" type="issue">
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<mods:number id="50280B3BAE5C3D4DFDFE957A3A0C37EC">4</mods:number>
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<mods:start id="1B883D21C83F6E784E84E2923075376A">547</mods:start>
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<mods:classification id="58F5CB5BB4BE9372911DC8022D466D76">journal article</mods:classification>
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<mods:identifier id="BACB8D37BB561AAC68A0C74EC69F4FE8" type="DOI">10.5281/zenodo.13390843</mods:identifier>
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<mods:identifier id="9CFDDAFD896C34AF1B01E61567B77975" type="ISSN">1732-2421</mods:identifier>
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<mods:identifier id="E2A0ED1AD8C17808A39024A07936222E" type="Zenodo-Dep">13390843</mods:identifier>
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<treatment id="F5521A73FF8A36013723F81BFD50E89D" ID-DOI="http://doi.org/10.5281/zenodo.13390903" ID-Zenodo-Dep="13390903" LSID="urn:lsid:plazi:treatment:F5521A73FF8A36013723F81BFD50E89D" httpUri="http://treatment.plazi.org/id/F5521A73FF8A36013723F81BFD50E89D" lastPageId="9" lastPageNumber="556" pageId="6" pageNumber="553">
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<subSubSection id="35E1F8EEFF8A360E3723F81BFA95E9D1" box="[812,1372,1925,1954]" pageId="6" pageNumber="553" type="nomenclature">
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<paragraph id="7D44AB65FF8A360E3723F81BFA95E9D1" blockId="6.[812,1372,1925,1987]" box="[812,1372,1925,1954]" pageId="6" pageNumber="553">
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<heading id="260C1C09FF8A360E3723F81BFA95E9D1" box="[812,1372,1925,1954]" fontSize="12" level="2" pageId="6" pageNumber="553" reason="9">
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<taxonomicName id="BAFBD0E6FF8A360E3723F81BFA95E9D1" authorityName="gen. et" authorityYear="2003" box="[812,1372,1925,1954]" class="Anthozoa" family="Disphyllidae" genus="Cantabriastraea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="6" pageNumber="555" phylum="Cnidaria" rank="species" species="cantabrica" status="gen. et">
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<emphasis id="4F8F7777FF8A360E3723F81BFB5AE9D1" box="[812,1171,1925,1954]" italics="true" pageId="6" pageNumber="553">Cantabriastraea cantabrica</emphasis>
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<taxonomicNameLabel id="54BCCA0CFF8A360E3095F818FB26E9D1" box="[1178,1263,1926,1954]" pageId="6" pageNumber="553" rank="species">gen. et</taxonomicNameLabel>
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sp. nov.
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="35E1F8EEFF8A360E3723F833FBBAE9B0" box="[812,1139,1965,1987]" pageId="6" pageNumber="553" type="description">
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<paragraph id="7D44AB65FF8A360E3723F833FBBAE9B0" blockId="6.[812,1372,1925,1987]" box="[812,1139,1965,1987]" pageId="6" pageNumber="553">
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<figureCitation id="E5C0B7E0FF8A360E3723F833FC54E9B0" box="[812,925,1965,1987]" captionStart="Fig" captionStartId="5.[102,135,1320,1339]" captionTargetBox="[102,775,228,1277]" captionTargetId="figure-576@5.[102,776,228,1277]" captionTargetPageId="5" captionText="Fig. 6. Cantabriastraea cantabrica gen. et sp. nov. A. Cross section of holotype DPO 14780. B. Longitudinal section of the same. Both × 2. Specimen from the Upper Emsian of Colle." figureDoi="http://doi.org/10.5281/zenodo.13390885" httpUri="https://zenodo.org/record/13390885/files/figure.png" pageId="6" pageNumber="553">Figs. 6A, B</figureCitation>
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,
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<figureCitation id="E5C0B7E0FF8A360E37A8F833FC2DE9B0" box="[935,996,1965,1987]" captionStart="Fig" captionStartId="6.[812,845,1576,1595]" captionTargetBox="[813,1484,234,1546]" captionTargetId="figure-543@6.[813,1485,234,1546]" captionTargetPageId="6" captionText="Fig. 7. Cantabriastraea cantabrica gen. et sp. nov. A. Colony DPO 14811 consisting of a large protocorallite disrupted by nonparricidal, marginal increase in distal view (A1); septa appear as broad and flat ridges which are stronglyretiformtowardstheperiphery.Undersideofthesamecolony(A2), partly encrusted by Aulopora sp.; × 2. B. Colony DPO 14813. B1. Colony underside showing a single protocorallite turned to the left. B2. Upper surface of the same colony; × 1. All specimens from the Upper Emsian of Colle." figureDoi="http://doi.org/10.5281/zenodo.13390893" httpUri="https://zenodo.org/record/13390893/files/figure.png" pageId="6" pageNumber="553">7A–D</figureCitation>
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,
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<figureCitation id="E5C0B7E0FF8A360E37E1F833FBEAE9B0" box="[1006,1059,1965,1987]" captionStart="Fig" captionStartId="8.[102,135,1813,1832]" captionText="Fig. 8. Cantabriastraea cantabrica gen. et sp. nov. A. Cross section of colony DPO 14789 with medium sized corallites. B. Cross section of colony DPO 1790. Corallites are very large and show highly variable development of carinate septa. C. Protocorallite of colony DPO 14788. D. Cross section of small colony DPO 14783. Septa are strongly thickened especially at dissepimentarium/tabularium boundary. E. Longitudinal section DPO 14786. Note strongly arched series of dissepiment−layers and convex tabulae. F. Cross section of colony−fragment DPO 14830 with development of juvenile corallites by marginal increase on lower right side of large protocorallite. G. Longitudinal section DPO 14790. H. Cross section of colony DPO 14787, consisting of a large protocorallite and four juvenile offsets developed by marginal increase. Undilated major septa do not reach to corallite axis. I. Longitudinal section of colony DPO 14787. J. Corallites of colony DPO 14790 (compare Fig. 8B), with strongly carinate septa breaking down into discrete trabeculae, becoming retiform. A–C, E–I, × 2; D × 3; J × 2.5. All specimens from the Upper Emsian of Colle." pageId="6" pageNumber="553">8A–J</figureCitation>
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,
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<figureCitation id="E5C0B7E0FF8A360E3023F833FBA7E9B0" box="[1068,1134,1965,1987]" captionStart="Fig" captionStartId="8.[102,135,731,750]" captionTargetBox="[556,1486,229,1340]" captionTargetId="figure-389@8.[556,1486,229,1340]" captionTargetPageId="8" captionText="Fig. 9. A. Cantabriastraea cantabrica gen. et sp. nov., specimen DPO 14789. A1. Cross sec− tion of tabularium and inner dissepimentarium. Trabeculae are oriented perpendicular to the rows of dissepiments; × 15. A2. Detail of A1; trabeculae intercalate between existing ones or bifurcate; × 20. A3. Septa in cross section with strong dilations in inner dissepimentarium caused by thickening of trabeculae. Towards the periphery (left side) septa become irregular carinate and may split into discrete trabeculae; × 24. B, C. Tabulophyllum bonarense sp. nov. B. Wall of corallite DPO 14673 in cross sec− tion. Thickenings of wall and lonsdaleoid dis− sepiments show a fibrous structure caused by thickened septal bases; × 24. C. Thin septa of corallite DPO 14686 (compare with Fig. 4G) inserting on lonsdaleoid dissepiments (dark lines in lower part of picture). Central part to− wards the axis (top of picture) is strongly af− fected by cement; × 24. All specimens from the Upper Emsian of Colle." figureDoi="http://doi.org/10.5281/zenodo.13390897" httpUri="https://zenodo.org/record/13390897/files/figure.png" pageId="6" pageNumber="553">9 A–C</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="35E1F8EEFF8A36013723F84DFD50E89D" lastPageId="9" lastPageNumber="556" pageId="6" pageNumber="553" type="materials_examined">
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<paragraph id="7D44AB65FF8A36003723F84DFBB8EBDD" blockId="6.[812,1111,2003,2025]" lastBlockId="8.[59,1485,1395,1795]" lastPageId="8" lastPageNumber="555" pageId="6" pageNumber="553">
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<materialsCitation id="CD93A138FF8A36003723F84DFBB8EBDD" county="Bonar" lastPageId="8" lastPageNumber="555" latitude="42.844997" location="Colle" longLatPrecision="19" longitude="1.5602777" municipality="Tabulae" pageId="6" pageNumber="553" specimenCount="1" typeStatus="holotype">
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<emphasis id="4F8F7777FF8A360E3723F84DFC4DE99A" box="[812,900,2003,2025]" italics="true" pageId="6" pageNumber="553">
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<typeStatus id="A24015C7FF8A360E3723F84DFC4DE99A" box="[812,900,2003,2025]" pageId="6" pageNumber="553" type="holotype">Holotype</typeStatus>
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</emphasis>
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: DPO 14780 (
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<figureCitation id="E5C0B7E0FF8A360E301FF84DFB82E99A" box="[1040,1099,2003,2025]" captionStart="Fig" captionStartId="5.[102,135,1320,1339]" captionTargetBox="[102,775,228,1277]" captionTargetId="figure-576@5.[102,776,228,1277]" captionTargetPageId="5" captionText="Fig. 6. Cantabriastraea cantabrica gen. et sp. nov. A. Cross section of holotype DPO 14780. B. Longitudinal section of the same. Both × 2. Specimen from the Upper Emsian of Colle." figureDoi="http://doi.org/10.5281/zenodo.13390885" httpUri="https://zenodo.org/record/13390885/files/figure.png" pageId="6" pageNumber="553">Fig. 6</figureCitation>
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).
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<emphasis id="4F8F7777FF8436003469FAEAFF2AEBFA" box="[102,227,1395,1417]" italics="true" pageId="8" pageNumber="555">Type locality</emphasis>
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:
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<location id="7824FDBEFF84360034FEFAEDFEEEEBFA" LSID="urn:lsid:plazi:treatment:F5521A73FF8A36013723F81BFD50E89D:7824FDBEFF84360034FEFAEDFEEEEBFA" box="[241,295,1395,1417]" county="Bonar" latitude="42.844997" longLatPrecision="19" longitude="1.5602777" municipality="Tabulae" name="Colle" pageId="8" pageNumber="555">Colle</location>
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, ca.
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<locationDeviation id="6CF6B247FF843600355CFAEDFDC1EBFA" box="[339,520,1395,1417]" location="Bonar" pageId="8" pageNumber="555">
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<quantity id="BA030680FF843600355CFAEDFE4CEBFA" box="[339,389,1395,1417]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="555" unit="km" value="5.0">5 km</quantity>
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SE of Boñar
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</locationDeviation>
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,
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<location id="7824FDBEFF843600361DFAEDFD6BEBFA" LSID="urn:lsid:plazi:treatment:F5521A73FF8A36013723F81BFD50E89D:7824FDBEFF843600361DFAEDFD6BEBFA" box="[530,674,1395,1417]" county="Bonar" latitude="42.844997" longLatPrecision="19" longitude="1.5602777" municipality="Tabulae" name="Province Leon" pageId="8" pageNumber="555">Province León</location>
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;
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<location id="7824FDBEFF84360036A2FAEDFD15EBFA" LSID="urn:lsid:plazi:treatment:F5521A73FF8A36013723F81BFD50E89D:7824FDBEFF84360036A2FAEDFD15EBFA" box="[685,732,1395,1417]" county="Bonar" latitude="42.844997" longLatPrecision="19" longitude="1.5602777" municipality="Tabulae" name="Hoja" pageId="8" pageNumber="555">Hoja</location>
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104 3.5–
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<quantity id="BA030680FF8436003750FAEBFC6EEBFE" box="[863,935,1397,1421]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="555" unit="mm" value="5.0">5 mm</quantity>
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.
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<collectingMunicipality id="9D20311FFF84360037B8FAEBFBD8EBFE" box="[951,1041,1397,1421]" pageId="8" pageNumber="555">Tabulae</collectingMunicipality>
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complete and slightly arched (13–18 (
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<collectingCounty id="9425D3E9FF8436003462FA11FF62EBD6" box="[109,171,1423,1445]" pageId="8" pageNumber="555">Boñar</collectingCounty>
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),
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<geoCoordinate id="18CFCDA2FF84360034B2FA11FEE8EBD6" box="[189,289,1423,1445]" degrees="1" direction="east" minutes="33" orientation="longitude" pageId="8" pageNumber="555" precision="15" seconds="37" value="1.5602777">1°33’37’’E</geoCoordinate>
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/
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<geoCoordinate id="18CFCDA2FF8436003525FA11FE61EBD6" box="[298,424,1423,1445]" degrees="42" direction="north" minutes="50" orientation="latitude" pageId="8" pageNumber="555" precision="15" seconds="42" value="42.844997">42°50’42’’N</geoCoordinate>
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. tabulae per
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<quantity id="BA030680FF84360037A0FA08FC39EBDD" box="[943,1008,1430,1454]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="555" unit="mm" value="5.0">5 mm</quantity>
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vertically).
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</materialsCitation>
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</paragraph>
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<caption id="2984FBEDFF8436003469FD45FEF7EB49" ID-DOI="http://doi.org/10.5281/zenodo.13390897" ID-Zenodo-Dep="13390897" httpUri="https://zenodo.org/record/13390897/files/figure.png" pageId="8" pageNumber="555" startId="8.[102,135,731,750]" targetBox="[556,1486,229,1340]" targetPageId="8" targetType="figure">
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<paragraph id="7D44AB65FF8436003469FD45FDC3EC9C" blockId="8.[102,524,731,1339]" box="[102,522,731,751]" pageId="8" pageNumber="555">
|
||
Fig. 9.
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<emphasis id="4F8F7777FF84360034A6FD45FF70EC9D" bold="true" box="[169,185,731,750]" pageId="8" pageNumber="555">A</emphasis>
|
||
.
|
||
<taxonomicName id="BAFBD0E6FF84360034C8FD45FDC3EC9C" authorityName="gen. et" authorityYear="2003" box="[199,522,731,751]" class="Anthozoa" family="Disphyllidae" genus="Cantabriastraea" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="555" phylum="Cnidaria" rank="species" species="cantabrica" status="gen. et">
|
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<emphasis id="4F8F7777FF84360034C8FD45FE0CEC9D" box="[199,453,731,750]" italics="true" pageId="8" pageNumber="555">Cantabriastraea cantabrica</emphasis>
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gen. et
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||
</taxonomicName>
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||
</paragraph>
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<paragraph id="7D44AB65FF8436003469FD66FDC5ED55" blockId="8.[102,524,731,1339]" pageId="8" pageNumber="555">
|
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sp. nov., specimen DPO 14789. A
|
||
<subScript id="E17FA920FF8436003595FC9EFE6BED7D" attach="left" box="[410,418,768,782]" fontSize="6" pageId="8" pageNumber="555">1</subScript>
|
||
. Cross section of tabularium and inner dissepimentarium.
|
||
</paragraph>
|
||
<paragraph id="7D44AB65FF8436003469FCB1FDC3ED2C" blockId="8.[102,524,731,1339]" pageId="8" pageNumber="555">
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Trabeculae are oriented perpendicular to the rows of dissepiments; × 15. A
|
||
<subScript id="E17FA920FF8436003571FCCAFE4FED11" attach="left" box="[382,390,852,866]" fontSize="6" pageId="8" pageNumber="555">2</subScript>
|
||
. Detail of A
|
||
<subScript id="E17FA920FF84360035F3FCCAFDCDED11" attach="left" box="[508,516,852,866]" fontSize="6" pageId="8" pageNumber="555">1</subScript>
|
||
;
|
||
</paragraph>
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||
<paragraph id="7D44AB65FF8436003469FCF9FDC5EA75" blockId="8.[102,524,731,1339]" pageId="8" pageNumber="555">
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||
trabeculae intercalate between existing ones or bifurcate; × 20. A
|
||
<subScript id="E17FA920FF8436003508FC12FEC6EDE9" attach="right" box="[263,271,908,922]" fontSize="6" pageId="8" pageNumber="555">3</subScript>
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||
. Septa in cross section with strong dilations in inner dissepimentarium caused by thickening of trabeculae. Towards the periphery (left side) septa become irregular carinate and may split into discrete trabeculae;
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||
</paragraph>
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||
<paragraph id="7D44AB65FF8436003469FB8EFDC5EA50" blockId="8.[102,524,731,1339]" box="[102,524,1039,1059]" pageId="8" pageNumber="555">
|
||
× 24.
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<emphasis id="4F8F7777FF8436003495FB91FF60EA51" bold="true" box="[154,169,1039,1058]" pageId="8" pageNumber="555">B</emphasis>
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||
,
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<emphasis id="4F8F7777FF84360034BAFB91FF0CEA51" bold="true" box="[181,197,1039,1058]" pageId="8" pageNumber="555">C</emphasis>
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.
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<taxonomicName id="BAFBD0E6FF84360034DEFB91FE74EA51" authorityName="Schröder & Soto" authorityYear="2003" box="[209,445,1039,1058]" class="Anthozoa" family="Chonophyllidae" genus="Tabulophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="555" phylum="Cnidaria" rank="species" species="bonarense" status="sp. nov.">
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<emphasis id="4F8F7777FF84360034DEFB91FE74EA51" box="[209,445,1039,1058]" italics="true" pageId="8" pageNumber="555">Tabulophyllum bonarense</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="54BCCA0CFF84360035CCFB8EFDC5EA50" box="[451,524,1040,1059]" pageId="8" pageNumber="555" rank="species">sp. nov.</taxonomicNameLabel>
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||
</paragraph>
|
||
<paragraph id="7D44AB65FF8436003469FBB5FDC3EADD" blockId="8.[102,524,731,1339]" pageId="8" pageNumber="555">
|
||
<emphasis id="4F8F7777FF8436003469FBB5FFBCEA4D" bold="true" box="[102,117,1067,1086]" pageId="8" pageNumber="555">B</emphasis>
|
||
. Wall of corallite DPO 14673 in cross section. Thickenings of wall and lonsdaleoid dissepiments show a fibrous structure caused by thickened septal bases; × 24.
|
||
<emphasis id="4F8F7777FF843600357BFBE1FE4DEAE1" bold="true" box="[372,388,1151,1170]" pageId="8" pageNumber="555">C</emphasis>
|
||
. Thin septa of corallite DPO 14686 (compare with Fig. 4G)
|
||
</paragraph>
|
||
<paragraph id="7D44AB65FF8436003469FB29FDC2EB6D" blockId="8.[102,524,731,1339]" pageId="8" pageNumber="555">inserting on lonsdaleoid dissepiments (dark lines in lower part of picture). Central part towards the axis (top of picture) is strongly affected by cement; × 24. All specimens from the</paragraph>
|
||
<paragraph id="7D44AB65FF8436003469FAB6FEF7EB49" blockId="8.[102,524,731,1339]" box="[102,318,1319,1339]" pageId="8" pageNumber="555">Upper Emsian of Colle.</paragraph>
|
||
</caption>
|
||
<paragraph id="7D44AB65FF8436003469FA29FCCFEBBF" blockId="8.[59,1485,1395,1795]" box="[102,774,1462,1484]" pageId="8" pageNumber="555">
|
||
<emphasis id="4F8F7777FF8436003469FA29FF2DEBBF" box="[102,228,1462,1484]" italics="true" pageId="8" pageNumber="555">Type horizon</emphasis>
|
||
: Lower Devonian, Emsian, La Vid Group, upper part of
|
||
</paragraph>
|
||
<paragraph id="7D44AB65FF8436003723FA58FCCFEB9B" blockId="8.[59,1485,1395,1795]" pageId="8" pageNumber="555">
|
||
<emphasis id="4F8F7777FF8436003723FA58FC64EBAE" box="[812,941,1478,1501]" italics="true" pageId="8" pageNumber="555">Description</emphasis>
|
||
.—Coralla are typically small, with a diameter of the Valporquero Formation (COL A−1) and lowermost part of the
|
||
</paragraph>
|
||
<paragraph id="7D44AB65FF8436013469FA70FD62E8B2" blockId="8.[59,1485,1395,1795]" lastBlockId="9.[102,775,232,1729]" lastPageId="9" lastPageNumber="556" pageId="8" pageNumber="555">
|
||
Coladilla Formation (COL B−3).
|
||
<quantity id="BA030680FF8436003723FA78FC49EB8E" box="[812,896,1510,1534]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="8" pageNumber="555" unit="cm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 cm</quantity>
|
||
at a maximum height of
|
||
<quantity id="BA030680FF8436003094FA78FB1AEB8E" box="[1179,1235,1510,1534]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="555" unit="cm" value="5.0">5 cm</quantity>
|
||
and have a distinctive
|
||
<emphasis id="4F8F7777FF8436003469F98FFE8AE855" box="[102,323,1552,1574]" italics="true" pageId="8" pageNumber="555">Derivation of the name</emphasis>
|
||
: The name derives from the Cantabrian Moun− growth form of a flat cake. Small colonies are mushroom tains. shaped and comprise only a few corallites (
|
||
<figureCitation id="E5C0B7E0FF8436003106F9B6FA97E833" box="[1289,1374,1576,1600]" captionStart="Fig" captionStartId="6.[812,845,1576,1595]" captionTargetBox="[813,1484,234,1546]" captionTargetId="figure-543@6.[813,1485,234,1546]" captionTargetPageId="6" captionText="Fig. 7. Cantabriastraea cantabrica gen. et sp. nov. A. Colony DPO 14811 consisting of a large protocorallite disrupted by nonparricidal, marginal increase in distal view (A1); septa appear as broad and flat ridges which are stronglyretiformtowardstheperiphery.Undersideofthesamecolony(A2), partly encrusted by Aulopora sp.; × 2. B. Colony DPO 14813. B1. Colony underside showing a single protocorallite turned to the left. B2. Upper surface of the same colony; × 1. All specimens from the Upper Emsian of Colle." figureDoi="http://doi.org/10.5281/zenodo.13390893" httpUri="https://zenodo.org/record/13390893/files/figure.png" pageId="8" pageNumber="555">Fig. 7C</figureCitation>
|
||
), whereas
|
||
<emphasis id="4F8F7777FF8436003469F9D0FF0EE810" box="[102,199,1614,1635]" italics="true" pageId="8" pageNumber="555">
|
||
<typeStatus id="A24015C7FF8436003469F9D0FF0EE810" box="[102,199,1614,1635]" pageId="8" pageNumber="555" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="6BFD60ECFF84360034DAF9D0FE93E817" box="[213,346,1614,1636]" count="34" pageId="8" pageNumber="555" type="generic">34 specimens</specimenCount>
|
||
, DPO 14781–14804, 14809–14814, 14828– adultcoloniestypicallyhaveaslightlydomedsurface.Growth 14830. starts with the development of a single, cylindrical
|
||
<emphasis id="4F8F7777FF8436003469F910FF11E8D6" box="[102,216,1678,1701]" italics="true" pageId="8" pageNumber="555">protoDiagnosis</emphasis>
|
||
.—Type species of
|
||
<taxonomicName id="BAFBD0E6FF84360035BDF910FDA2E8D5" authorityName="Schroeder & Soto" authorityYear="2003" box="[434,619,1678,1702]" class="Anthozoa" family="Disphyllidae" genus="Cantabriastraea" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="555" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="4F8F7777FF84360035BDF910FDA2E8D5" box="[434,619,1678,1702]" italics="true" pageId="8" pageNumber="555">Cantabriastraea</emphasis>
|
||
</taxonomicName>
|
||
with corallite corallite, reaching a maximum diameter of
|
||
<quantity id="BA030680FF8436003115F915FA9DE8D0" box="[1306,1364,1675,1699]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="555" unit="cm" value="2.0">2 cm</quantity>
|
||
(
|
||
<figureCitation id="E5C0B7E0FF8436003169F915FA09E8D0" box="[1382,1472,1675,1699]" captionStart="Fig" captionStartId="8.[102,135,1813,1832]" captionText="Fig. 8. Cantabriastraea cantabrica gen. et sp. nov. A. Cross section of colony DPO 14789 with medium sized corallites. B. Cross section of colony DPO 1790. Corallites are very large and show highly variable development of carinate septa. C. Protocorallite of colony DPO 14788. D. Cross section of small colony DPO 14783. Septa are strongly thickened especially at dissepimentarium/tabularium boundary. E. Longitudinal section DPO 14786. Note strongly arched series of dissepiment−layers and convex tabulae. F. Cross section of colony−fragment DPO 14830 with development of juvenile corallites by marginal increase on lower right side of large protocorallite. G. Longitudinal section DPO 14790. H. Cross section of colony DPO 14787, consisting of a large protocorallite and four juvenile offsets developed by marginal increase. Undilated major septa do not reach to corallite axis. I. Longitudinal section of colony DPO 14787. J. Corallites of colony DPO 14790 (compare Fig. 8B), with strongly carinate septa breaking down into discrete trabeculae, becoming retiform. A–C, E–I, × 2; D × 3; J × 2.5. All specimens from the Upper Emsian of Colle." pageId="8" pageNumber="555">Fig. 8C</figureCitation>
|
||
). diameter of
|
||
<quantity id="BA030680FF84360034E0F930FE92E8B6" box="[239,347,1710,1734]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="8" pageNumber="555" unit="mm" value="10.5" valueMax="13.0" valueMin="8.0">8–13 mm</quantity>
|
||
and 19–24 major septa, which do not This corallite is increasingly replaced by offsets developed by reach the corallite axis. Diameter of tabularium ranges from nonparricidal, marginal increase (
|
||
<figureCitation id="E5C0B7E0FF8436003094F953FAC5E895" box="[1179,1292,1741,1766]" captionStart="Fig" captionStartId="8.[102,135,1813,1832]" captionText="Fig. 8. Cantabriastraea cantabrica gen. et sp. nov. A. Cross section of colony DPO 14789 with medium sized corallites. B. Cross section of colony DPO 1790. Corallites are very large and show highly variable development of carinate septa. C. Protocorallite of colony DPO 14788. D. Cross section of small colony DPO 14783. Septa are strongly thickened especially at dissepimentarium/tabularium boundary. E. Longitudinal section DPO 14786. Note strongly arched series of dissepiment−layers and convex tabulae. F. Cross section of colony−fragment DPO 14830 with development of juvenile corallites by marginal increase on lower right side of large protocorallite. G. Longitudinal section DPO 14790. H. Cross section of colony DPO 14787, consisting of a large protocorallite and four juvenile offsets developed by marginal increase. Undilated major septa do not reach to corallite axis. I. Longitudinal section of colony DPO 14787. J. Corallites of colony DPO 14790 (compare Fig. 8B), with strongly carinate septa breaking down into discrete trabeculae, becoming retiform. A–C, E–I, × 2; D × 3; J × 2.5. All specimens from the Upper Emsian of Colle." pageId="8" pageNumber="555">Fig. 8F, H</figureCitation>
|
||
) and transformed
|
||
<emphasis id="4F8F7777FF8436003434F974FF9EE970" box="[59,87,1770,1795]" italics="true" pageId="8" pageNumber="555">Ą</emphasis>
|
||
to colonial growth. Apical portions of those protocorallites are not preserved. Some fragments show an attachment to corallite fragments of
|
||
<taxonomicName id="BAFBD0E6FF853601352AFEB6FD7EEF32" authority="Soto" authorityName="Soto" box="[293,695,296,321]" class="Magnoliopsida" family="Aizoaceae" genus="Synaptophyllum" kingdom="Plantae" order="Caryophyllales" pageId="9" pageNumber="556" phylum="Tracheophyta" rank="species" species="multiseptatum">
|
||
<emphasis id="4F8F7777FF853601352AFEB6FDB3EF33" box="[293,634,296,320]" italics="true" pageId="9" pageNumber="556">Synaptophyllum multiseptatum</emphasis>
|
||
Soto
|
||
</taxonomicName>
|
||
, which has obviously been used as hardground. Colony surface is usually well preserved and calices show an everted calicular plane with a deeply depressed and flat central part (
|
||
<figureCitation id="E5C0B7E0FF85360136A6FE16FFB4EFB2" captionStart="Fig" captionStartId="6.[812,845,1576,1595]" captionTargetBox="[813,1484,234,1546]" captionTargetId="figure-543@6.[813,1485,234,1546]" captionTargetPageId="6" captionText="Fig. 7. Cantabriastraea cantabrica gen. et sp. nov. A. Colony DPO 14811 consisting of a large protocorallite disrupted by nonparricidal, marginal increase in distal view (A1); septa appear as broad and flat ridges which are stronglyretiformtowardstheperiphery.Undersideofthesamecolony(A2), partly encrusted by Aulopora sp.; × 2. B. Colony DPO 14813. B1. Colony underside showing a single protocorallite turned to the left. B2. Upper surface of the same colony; × 1. All specimens from the Upper Emsian of Colle." figureDoi="http://doi.org/10.5281/zenodo.13390893" httpUri="https://zenodo.org/record/13390893/files/figure.png" pageId="9" pageNumber="556">Fig. 7A, D</figureCitation>
|
||
). Growth form is astreoid−thamnasterioid, but septa are rarely confluent and limitations of corallites are usually marked by break down of septa. There are no true corallite walls. Calical centres are
|
||
<quantity id="BA030680FF8536013582FD97FE33EC53" box="[397,506,521,545]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="9" pageNumber="556" unit="mm" value="10.5" valueMax="13.0" valueMin="8.0">8–13 mm</quantity>
|
||
apart, but can reach to
|
||
<quantity id="BA030680FF8536013469FDB7FF7CEC33" box="[102,181,553,577]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="556" unit="mm" value="17.0">17 mm</quantity>
|
||
in juvenile coralla consisting of only a protocorallite and few smalloffsets(
|
||
<figureCitation id="E5C0B7E0FF8536013555FDD6FE66EC12" box="[346,431,584,609]" captionStart="Fig" captionStartId="8.[102,135,1813,1832]" captionText="Fig. 8. Cantabriastraea cantabrica gen. et sp. nov. A. Cross section of colony DPO 14789 with medium sized corallites. B. Cross section of colony DPO 1790. Corallites are very large and show highly variable development of carinate septa. C. Protocorallite of colony DPO 14788. D. Cross section of small colony DPO 14783. Septa are strongly thickened especially at dissepimentarium/tabularium boundary. E. Longitudinal section DPO 14786. Note strongly arched series of dissepiment−layers and convex tabulae. F. Cross section of colony−fragment DPO 14830 with development of juvenile corallites by marginal increase on lower right side of large protocorallite. G. Longitudinal section DPO 14790. H. Cross section of colony DPO 14787, consisting of a large protocorallite and four juvenile offsets developed by marginal increase. Undilated major septa do not reach to corallite axis. I. Longitudinal section of colony DPO 14787. J. Corallites of colony DPO 14790 (compare Fig. 8B), with strongly carinate septa breaking down into discrete trabeculae, becoming retiform. A–C, E–I, × 2; D × 3; J × 2.5. All specimens from the Upper Emsian of Colle." pageId="9" pageNumber="556">Fig. 8H</figureCitation>
|
||
). Morphology ofsepta ishighly variable within a colony as well as in individual corallites.
|
||
<collectingRegion id="BF3F6587FF8536013469FD17FF6AECD2" box="[102,163,649,673]" country="Spain" name="Ceuta" pageId="9" pageNumber="556">Septa</collectingRegion>
|
||
are usually thin near the corallite boundary, but moderately thickened and may bear numerous irregular carinae or lateralspines(vepreculae)intheirmiddlepart(
|
||
<figureCitation id="E5C0B7E0FF8536013662FD56FD71EC93" box="[621,696,712,736]" captionStart="Fig" captionStartId="8.[102,135,731,750]" captionTargetBox="[556,1486,229,1340]" captionTargetId="figure-389@8.[556,1486,229,1340]" captionTargetPageId="8" captionText="Fig. 9. A. Cantabriastraea cantabrica gen. et sp. nov., specimen DPO 14789. A1. Cross sec− tion of tabularium and inner dissepimentarium. Trabeculae are oriented perpendicular to the rows of dissepiments; × 15. A2. Detail of A1; trabeculae intercalate between existing ones or bifurcate; × 20. A3. Septa in cross section with strong dilations in inner dissepimentarium caused by thickening of trabeculae. Towards the periphery (left side) septa become irregular carinate and may split into discrete trabeculae; × 24. B, C. Tabulophyllum bonarense sp. nov. B. Wall of corallite DPO 14673 in cross sec− tion. Thickenings of wall and lonsdaleoid dis− sepiments show a fibrous structure caused by thickened septal bases; × 24. C. Thin septa of corallite DPO 14686 (compare with Fig. 4G) inserting on lonsdaleoid dissepiments (dark lines in lower part of picture). Central part to− wards the axis (top of picture) is strongly af− fected by cement; × 24. All specimens from the Upper Emsian of Colle." figureDoi="http://doi.org/10.5281/zenodo.13390897" httpUri="https://zenodo.org/record/13390897/files/figure.png" pageId="9" pageNumber="556">Fig.9C</figureCitation>
|
||
).Close to the dissepimentarium boundary they are usually strongly thickened and may be in lateral contact (
|
||
<figureCitation id="E5C0B7E0FF8536013637FC96FD5AED52" box="[568,659,776,801]" captionStart="Fig" captionStartId="8.[102,135,1813,1832]" captionText="Fig. 8. Cantabriastraea cantabrica gen. et sp. nov. A. Cross section of colony DPO 14789 with medium sized corallites. B. Cross section of colony DPO 1790. Corallites are very large and show highly variable development of carinate septa. C. Protocorallite of colony DPO 14788. D. Cross section of small colony DPO 14783. Septa are strongly thickened especially at dissepimentarium/tabularium boundary. E. Longitudinal section DPO 14786. Note strongly arched series of dissepiment−layers and convex tabulae. F. Cross section of colony−fragment DPO 14830 with development of juvenile corallites by marginal increase on lower right side of large protocorallite. G. Longitudinal section DPO 14790. H. Cross section of colony DPO 14787, consisting of a large protocorallite and four juvenile offsets developed by marginal increase. Undilated major septa do not reach to corallite axis. I. Longitudinal section of colony DPO 14787. J. Corallites of colony DPO 14790 (compare Fig. 8B), with strongly carinate septa breaking down into discrete trabeculae, becoming retiform. A–C, E–I, × 2; D × 3; J × 2.5. All specimens from the Upper Emsian of Colle." pageId="9" pageNumber="556">Fig. 8D</figureCitation>
|
||
). Carinate septa show a strong tendency to break down into discrete trabeculae giving septa a spongy, retiform morphology. Isolated groups of trabeculae are usually connected by numerous sections of lateral dissepiments, which gives the impression of very broad septa (
|
||
<figureCitation id="E5C0B7E0FF853601352AFC36FE4CEDB2" box="[293,389,936,961]" captionStart="Fig" captionStartId="6.[812,845,1576,1595]" captionTargetBox="[813,1484,234,1546]" captionTargetId="figure-543@6.[813,1485,234,1546]" captionTargetPageId="6" captionText="Fig. 7. Cantabriastraea cantabrica gen. et sp. nov. A. Colony DPO 14811 consisting of a large protocorallite disrupted by nonparricidal, marginal increase in distal view (A1); septa appear as broad and flat ridges which are stronglyretiformtowardstheperiphery.Undersideofthesamecolony(A2), partly encrusted by Aulopora sp.; × 2. B. Colony DPO 14813. B1. Colony underside showing a single protocorallite turned to the left. B2. Upper surface of the same colony; × 1. All specimens from the Upper Emsian of Colle." figureDoi="http://doi.org/10.5281/zenodo.13390893" httpUri="https://zenodo.org/record/13390893/files/figure.png" pageId="9" pageNumber="556">Figs. 7A</figureCitation>
|
||
,
|
||
<figureCitation id="E5C0B7E0FF8536013580FC37FE63EDB2" box="[399,426,937,961]" captionStart="Fig" captionStartId="8.[102,135,1813,1832]" captionText="Fig. 8. Cantabriastraea cantabrica gen. et sp. nov. A. Cross section of colony DPO 14789 with medium sized corallites. B. Cross section of colony DPO 1790. Corallites are very large and show highly variable development of carinate septa. C. Protocorallite of colony DPO 14788. D. Cross section of small colony DPO 14783. Septa are strongly thickened especially at dissepimentarium/tabularium boundary. E. Longitudinal section DPO 14786. Note strongly arched series of dissepiment−layers and convex tabulae. F. Cross section of colony−fragment DPO 14830 with development of juvenile corallites by marginal increase on lower right side of large protocorallite. G. Longitudinal section DPO 14790. H. Cross section of colony DPO 14787, consisting of a large protocorallite and four juvenile offsets developed by marginal increase. Undilated major septa do not reach to corallite axis. I. Longitudinal section of colony DPO 14787. J. Corallites of colony DPO 14790 (compare Fig. 8B), with strongly carinate septa breaking down into discrete trabeculae, becoming retiform. A–C, E–I, × 2; D × 3; J × 2.5. All specimens from the Upper Emsian of Colle." pageId="9" pageNumber="556">8J</figureCitation>
|
||
). Trabecularthickening ofsepta (
|
||
<figureCitation id="E5C0B7E0FF8536013461FC56FF0AED92" box="[110,195,968,993]" captionStart="Fig" captionStartId="8.[102,135,731,750]" captionTargetBox="[556,1486,229,1340]" captionTargetId="figure-389@8.[556,1486,229,1340]" captionTargetPageId="8" captionText="Fig. 9. A. Cantabriastraea cantabrica gen. et sp. nov., specimen DPO 14789. A1. Cross sec− tion of tabularium and inner dissepimentarium. Trabeculae are oriented perpendicular to the rows of dissepiments; × 15. A2. Detail of A1; trabeculae intercalate between existing ones or bifurcate; × 20. A3. Septa in cross section with strong dilations in inner dissepimentarium caused by thickening of trabeculae. Towards the periphery (left side) septa become irregular carinate and may split into discrete trabeculae; × 24. B, C. Tabulophyllum bonarense sp. nov. B. Wall of corallite DPO 14673 in cross sec− tion. Thickenings of wall and lonsdaleoid dis− sepiments show a fibrous structure caused by thickened septal bases; × 24. C. Thin septa of corallite DPO 14686 (compare with Fig. 4G) inserting on lonsdaleoid dissepiments (dark lines in lower part of picture). Central part to− wards the axis (top of picture) is strongly af− fected by cement; × 24. All specimens from the Upper Emsian of Colle." figureDoi="http://doi.org/10.5281/zenodo.13390897" httpUri="https://zenodo.org/record/13390897/files/figure.png" pageId="9" pageNumber="556">Fig. 9C</figureCitation>
|
||
) increases towards the tabularium border where they are flanked by numerous coarse carinae. However, there are colonies or corallites with very weak or even smooth septa, which are not affected by any septal degeneration (
|
||
<figureCitation id="E5C0B7E0FF85360136AFFBB6FD33EA32" box="[672,762,1064,1089]" captionStart="Fig" captionStartId="8.[102,135,1813,1832]" captionText="Fig. 8. Cantabriastraea cantabrica gen. et sp. nov. A. Cross section of colony DPO 14789 with medium sized corallites. B. Cross section of colony DPO 1790. Corallites are very large and show highly variable development of carinate septa. C. Protocorallite of colony DPO 14788. D. Cross section of small colony DPO 14783. Septa are strongly thickened especially at dissepimentarium/tabularium boundary. E. Longitudinal section DPO 14786. Note strongly arched series of dissepiment−layers and convex tabulae. F. Cross section of colony−fragment DPO 14830 with development of juvenile corallites by marginal increase on lower right side of large protocorallite. G. Longitudinal section DPO 14790. H. Cross section of colony DPO 14787, consisting of a large protocorallite and four juvenile offsets developed by marginal increase. Undilated major septa do not reach to corallite axis. I. Longitudinal section of colony DPO 14787. J. Corallites of colony DPO 14790 (compare Fig. 8B), with strongly carinate septa breaking down into discrete trabeculae, becoming retiform. A–C, E–I, × 2; D × 3; J × 2.5. All specimens from the Upper Emsian of Colle." pageId="9" pageNumber="556">Fig. 8H</figureCitation>
|
||
). Particularly septa in corallites of small colonies and protocorallites(
|
||
<figureCitation id="E5C0B7E0FF85360134D6FBF6FEE0EAF2" box="[217,297,1128,1153]" captionStart="Fig" captionStartId="8.[102,135,1813,1832]" captionText="Fig. 8. Cantabriastraea cantabrica gen. et sp. nov. A. Cross section of colony DPO 14789 with medium sized corallites. B. Cross section of colony DPO 1790. Corallites are very large and show highly variable development of carinate septa. C. Protocorallite of colony DPO 14788. D. Cross section of small colony DPO 14783. Septa are strongly thickened especially at dissepimentarium/tabularium boundary. E. Longitudinal section DPO 14786. Note strongly arched series of dissepiment−layers and convex tabulae. F. Cross section of colony−fragment DPO 14830 with development of juvenile corallites by marginal increase on lower right side of large protocorallite. G. Longitudinal section DPO 14790. H. Cross section of colony DPO 14787, consisting of a large protocorallite and four juvenile offsets developed by marginal increase. Undilated major septa do not reach to corallite axis. I. Longitudinal section of colony DPO 14787. J. Corallites of colony DPO 14790 (compare Fig. 8B), with strongly carinate septa breaking down into discrete trabeculae, becoming retiform. A–C, E–I, × 2; D × 3; J × 2.5. All specimens from the Upper Emsian of Colle." pageId="9" pageNumber="556">Fig. 8C</figureCitation>
|
||
)are weakly carinate. Majorsepta vary from
|
||
<quantity id="BA030680FF8536013469FB17FF03EAD3" box="[102,202,1160,1185]" metricMagnitude="-1" metricUnit="m" metricValue="5.461" metricValueMax="6.096" metricValueMin="4.826" pageId="9" pageNumber="556" unit="in" value="21.5" valueMax="24.0" valueMin="19.0">19–24 in</quantity>
|
||
number, but can reach up to
|
||
<quantity id="BA030680FF8536013615FB17FD9CEAD3" box="[538,597,1160,1185]" metricMagnitude="-1" metricUnit="m" metricValue="7.874" pageId="9" pageNumber="556" unit="in" value="31.0">31 in</quantity>
|
||
a protocorallite (
|
||
<figureCitation id="E5C0B7E0FF8536013461FB36FF0FEAB2" box="[110,198,1192,1217]" captionStart="Fig" captionStartId="8.[102,135,1813,1832]" captionText="Fig. 8. Cantabriastraea cantabrica gen. et sp. nov. A. Cross section of colony DPO 14789 with medium sized corallites. B. Cross section of colony DPO 1790. Corallites are very large and show highly variable development of carinate septa. C. Protocorallite of colony DPO 14788. D. Cross section of small colony DPO 14783. Septa are strongly thickened especially at dissepimentarium/tabularium boundary. E. Longitudinal section DPO 14786. Note strongly arched series of dissepiment−layers and convex tabulae. F. Cross section of colony−fragment DPO 14830 with development of juvenile corallites by marginal increase on lower right side of large protocorallite. G. Longitudinal section DPO 14790. H. Cross section of colony DPO 14787, consisting of a large protocorallite and four juvenile offsets developed by marginal increase. Undilated major septa do not reach to corallite axis. I. Longitudinal section of colony DPO 14787. J. Corallites of colony DPO 14790 (compare Fig. 8B), with strongly carinate septa breaking down into discrete trabeculae, becoming retiform. A–C, E–I, × 2; D × 3; J × 2.5. All specimens from the Upper Emsian of Colle." pageId="9" pageNumber="556">Fig. 8C</figureCitation>
|
||
). Although there are rhopaloid septa in some corallites, they are generally very slender within the tabularium and do not reach the corallite axis, so that a small axial space of 3.5–
|
||
<quantity id="BA030680FF8536013498FA96FF1CEB53" box="[151,213,1288,1312]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="556" unit="mm" value="5.0">5 mm</quantity>
|
||
remains free, which is traversed by rare sections of tabulae. Minor septa barely extend into the tabularium. Sections of axially convex dissepiments are numerous, but are increasingly replaced by irregular or lateral dissepiments towards the periphery. Diameter of tabularia ranges from 3.5–5 mmand includespredominantly complete and slightly convex tabulae which are arranged above one another or are supplemented with peripheral tabellae. There are 13–30 tabulae/cm corallite length. The boundary to the dissepimentarium is characterized by steeply inclined globular dissepiments and accordingwiththeevertedcalice,thedissepimentallayersarestrongly arched (
|
||
<figureCitation id="E5C0B7E0FF85360134B2F9F6FEFBE8F2" box="[189,306,1640,1665]" captionStart="Fig" captionStartId="8.[102,135,1813,1832]" captionText="Fig. 8. Cantabriastraea cantabrica gen. et sp. nov. A. Cross section of colony DPO 14789 with medium sized corallites. B. Cross section of colony DPO 1790. Corallites are very large and show highly variable development of carinate septa. C. Protocorallite of colony DPO 14788. D. Cross section of small colony DPO 14783. Septa are strongly thickened especially at dissepimentarium/tabularium boundary. E. Longitudinal section DPO 14786. Note strongly arched series of dissepiment−layers and convex tabulae. F. Cross section of colony−fragment DPO 14830 with development of juvenile corallites by marginal increase on lower right side of large protocorallite. G. Longitudinal section DPO 14790. H. Cross section of colony DPO 14787, consisting of a large protocorallite and four juvenile offsets developed by marginal increase. Undilated major septa do not reach to corallite axis. I. Longitudinal section of colony DPO 14787. J. Corallites of colony DPO 14790 (compare Fig. 8B), with strongly carinate septa breaking down into discrete trabeculae, becoming retiform. A–C, E–I, × 2; D × 3; J × 2.5. All specimens from the Upper Emsian of Colle." pageId="9" pageNumber="556">Fig. 8E, G</figureCitation>
|
||
). They are commonly traversed by fanning monacanth trabeculae, which are arranged perpendicular to the dissepiments and may show bifurcations (
|
||
<figureCitation id="E5C0B7E0FF8536013626F936FD57E8B2" box="[553,670,1704,1729]" captionStart="Fig" captionStartId="8.[102,135,731,750]" captionTargetBox="[556,1486,229,1340]" captionTargetId="figure-389@8.[556,1486,229,1340]" captionTargetPageId="8" captionText="Fig. 9. A. Cantabriastraea cantabrica gen. et sp. nov., specimen DPO 14789. A1. Cross sec− tion of tabularium and inner dissepimentarium. Trabeculae are oriented perpendicular to the rows of dissepiments; × 15. A2. Detail of A1; trabeculae intercalate between existing ones or bifurcate; × 20. A3. Septa in cross section with strong dilations in inner dissepimentarium caused by thickening of trabeculae. Towards the periphery (left side) septa become irregular carinate and may split into discrete trabeculae; × 24. B, C. Tabulophyllum bonarense sp. nov. B. Wall of corallite DPO 14673 in cross sec− tion. Thickenings of wall and lonsdaleoid dis− sepiments show a fibrous structure caused by thickened septal bases; × 24. C. Thin septa of corallite DPO 14686 (compare with Fig. 4G) inserting on lonsdaleoid dissepiments (dark lines in lower part of picture). Central part to− wards the axis (top of picture) is strongly af− fected by cement; × 24. All specimens from the Upper Emsian of Colle." figureDoi="http://doi.org/10.5281/zenodo.13390897" httpUri="https://zenodo.org/record/13390897/files/figure.png" pageId="9" pageNumber="556">Fig. 9A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="2984FBEDFF8436003469F88BFC48E99B" pageId="8" pageNumber="555" startId="8.[102,135,1813,1832]" targetType="figure">
|
||
<paragraph id="7D44AB65FF8436003469F88BFC48E99B" blockId="8.[102,1485,1813,2025]" pageId="8" pageNumber="555">
|
||
Fig. 8.
|
||
<taxonomicName id="BAFBD0E6FF84360034AAF88BFE2DE95A" authorityName="gen. et" authorityYear="2003" box="[165,484,1813,1833]" class="Anthozoa" family="Disphyllidae" genus="Cantabriastraea" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="555" phylum="Cnidaria" rank="species" species="cantabrica" status="gen. et">
|
||
<emphasis id="4F8F7777FF84360034AAF88BFE6BE95B" box="[165,418,1813,1832]" italics="true" pageId="8" pageNumber="555">Cantabriastraea cantabrica</emphasis>
|
||
<taxonomicNameLabel id="54BCCA0CFF84360035A7F888FE2DE95A" box="[424,484,1814,1833]" pageId="8" pageNumber="555" rank="species">gen. et</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="4F8F7777FF8436003639F88BFD8FE95B" bold="true" box="[566,582,1813,1832]" pageId="8" pageNumber="555">A</emphasis>
|
||
. Cross section of colony DPO 14789 with medium sized corallites.
|
||
<emphasis id="4F8F7777FF84360030A4F88BFB73E95B" bold="true" box="[1195,1210,1813,1832]" pageId="8" pageNumber="555">B</emphasis>
|
||
. Cross section of colony DPO 1790. Corallites are very large and show highly variable development of carinate septa.
|
||
<emphasis id="4F8F7777FF8436003772F8AEFC44E930" bold="true" box="[893,909,1840,1859]" pageId="8" pageNumber="555">C</emphasis>
|
||
. Protocorallite of colony DPO 14788.
|
||
<emphasis id="4F8F7777FF84360030E5F8AFFB33E937" bold="true" box="[1258,1274,1841,1860]" pageId="8" pageNumber="555">D</emphasis>
|
||
. Cross section of small colony DPO 14783. Septa are strongly thickened especially at dissepimentarium/tabularium boundary.
|
||
<emphasis id="4F8F7777FF843600300BF8D2FBDAE92C" bold="true" box="[1028,1043,1868,1887]" pageId="8" pageNumber="555">E</emphasis>
|
||
. Longitudinal section DPO 14786. Note strongly arched series of dissepiment−layers and convex tabulae.
|
||
<emphasis id="4F8F7777FF843600366EF8F6FDA6E908" bold="true" box="[609,623,1896,1915]" pageId="8" pageNumber="555">F</emphasis>
|
||
. Cross section of colony−fragment DPO 14830 with development of juvenile corallites by marginal increase on lower right side of large protocorallite.
|
||
<emphasis id="4F8F7777FF843600366FF81DFDBBE9E5" bold="true" box="[608,626,1923,1942]" pageId="8" pageNumber="555">G</emphasis>
|
||
. Longitudinal section DPO 14790.
|
||
<emphasis id="4F8F7777FF84360037BFF81DFC0BE9E5" bold="true" box="[944,962,1923,1942]" pageId="8" pageNumber="555">H</emphasis>
|
||
. Cross section of colony DPO 14787, consisting of a large protocorallite and four juvenile offsets developed by marginal increase. Undilated major septa do not reach to corallite axis.
|
||
<emphasis id="4F8F7777FF84360030B0F801FB01E9C1" bold="true" box="[1215,1224,1951,1970]" pageId="8" pageNumber="555">I</emphasis>
|
||
. Longitudinal section of colony DPO 14787.
|
||
<emphasis id="4F8F7777FF8436003508F824FEDBE9BE" bold="true" box="[263,274,1978,1997]" pageId="8" pageNumber="555">J</emphasis>
|
||
. Corallites of colony DPO 14790 (compare Fig. 8B), with strongly carinate septa breaking down into discrete trabeculae, becoming retiform. A–C, E–I, × 2; D × 3; J × 2.5. All specimens from the Upper Emsian of Colle.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7D44AB65FF8536013469F949FD50E89D" blockId="9.[102,665,1750,1775]" box="[102,665,1750,1775]" pageId="9" pageNumber="556">
|
||
<emphasis id="4F8F7777FF8536013469F949FF23E89D" box="[102,234,1751,1774]" italics="true" pageId="9" pageNumber="556">Occurrence</emphasis>
|
||
.—Only known from the
|
||
<typeStatus id="A24015C7FF8536013609F949FDFFE89C" box="[518,566,1751,1775]" pageId="9" pageNumber="556">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |