treatments-xml/data/03/9F/EF/039FEF343D09FFC9FF3CF5349471F8A8.xml

<document id="66B8CD06270FB555753E18F03CBD1685" ID-DOI="10.11646/zootaxa.4810.1.12" ID-ISSN="1175-5326" ID-Zenodo-Dep="3937036" ID-ZooBank="424415DC-807E-493A-8883-903D2ADE9427" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594292134250" checkinUser="plazi" docAuthor="Zhu, Wenhui, Yao, Zhiyuan, Zheng, Guo &amp; Li, Shuqiang" docDate="2020" docId="039FEF343D09FFC9FF3CF5349471F8A8" docLanguage="en" docName="zootaxa.4810.1.12.pdf" docOrigin="Zootaxa 4810 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Belisana naling Zhu &amp; Yao &amp; Zheng &amp; Li 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="182" masterDocId="FFA6974C3D0EFFCEFFABF735972AFFDB" masterDocTitle="Six new species of the spider genus Belisana Thorell, 1898 (Araneae: Pholcidae) from southern China" masterLastPageNumber="197" masterPageNumber="175" pageNumber="182" updateTime="1700277344912" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="6FD1CF72734312D1E611FE0AFC039428">Six new species of the spider genus Belisana Thorell, 1898 (Araneae: Pholcidae) from southern China</mods:title>
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<mods:namePart id="9F074456D124B394CDEC7FD1E484E9C2">Zhu, Wenhui</mods:namePart>
<mods:affiliation id="8C60ECEB1EA74B1E585899DA5BCAFF5E">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; zhuwh 1016 @ outlook. com; https: // orcid. org / 0000 - 0001 - 5650 - 1670</mods:affiliation>
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<mods:namePart id="60DE7DF89E65FE74975512EB72AEE2E8">Yao, Zhiyuan</mods:namePart>
<mods:affiliation id="D6BCD3FE4943651477694EA84D26F7E6">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; yaozy @ synu. edu. cn; https: // orcid. org / 0000 - 0002 - 1631 - 0949</mods:affiliation>
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<mods:namePart id="BDFD93ABC3CA1DB66C23968DA1CFFEBE">Zheng, Guo</mods:namePart>
<mods:affiliation id="63D38F01E83013F728340B145DFF2AC8">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; zhengguo @ synu. edu. cn; https: // orcid. org / 0000 - 0002 - 9735 - 9700</mods:affiliation>
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<mods:namePart id="6872878D0DC3D442B7DE40ED306732B1">Li, Shuqiang</mods:namePart>
<mods:affiliation id="19644AF98E824984BB86275B672FCCF5">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; lisq @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 3290 - 5416</mods:affiliation>
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<mods:date id="3F06EF26B90DF02ADE756389CEFDA2E6">2020</mods:date>
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<heading id="D0C1E94E3D09FFC9FF3CF534951EFDC0" bold="true" box="[151,564,513,540]" fontSize="11" level="1" pageId="7" pageNumber="182" reason="1">
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<taxonomicName id="4C3625A13D09FFC9FF3CF53496E5FDC0" ID-CoL="99BJF" authority="Yao &amp; Li" authorityName="Zhu &amp; Yao &amp; Zheng &amp; Li" authorityYear="2020" box="[151,463,513,540]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="182" phylum="Arthropoda" rank="species" species="naling" status="sp. nov.">
<emphasis id="B94282303D09FFC9FF3CF534967CFDC7" bold="true" box="[151,342,513,540]" italics="true" pageId="7" pageNumber="182">Belisana naling</emphasis>
Yao &amp; Li
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<paragraph id="8B895E223D09FFC9FF3CF5109682FDE4" blockId="7.[151,564,513,575]" box="[151,424,548,575]" pageId="7" pageNumber="182">
<figureCitation id="130D42A73D09FFC9FF3CF51097D0FDE4" box="[151,250,548,575]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="8.[151,250,1875,1900]" captionStartId-1="9.[151,250,1910,1935]" captionTargetBox-0="[226,1361,181,1850]" captionTargetBox-1="[214,1373,181,1885]" captionTargetId-0="figure@8.[226,1361,181,1850]" captionTargetId-1="figure@9.[214,1373,181,1885]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 5. Belisana naling sp. nov., holotype male. A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at large proximo-dorsal hump on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large, distal membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-ventral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." captionText-1="FIGURE 6. Belisana naling sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi-0="http://doi.org/10.5281/zenodo.3937046" figureDoi-1="http://doi.org/10.5281/zenodo.3937048" httpUri-0="https://zenodo.org/record/3937046/files/figure.png" httpUri-1="https://zenodo.org/record/3937048/files/figure.png" pageId="7" pageNumber="182">Figs 5–6</figureCitation>
, 
<figureCitation id="130D42A73D09FFC9FEADF5109635FDE4" box="[262,287,549,575]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). A–B. B. guilin sp. nov.; C–D. B. maoer sp. nov.; E–F. B. naling sp. nov.; G–H. B. xiyuan sp. nov.; I–J. B. xuanguan sp. nov.; K–L. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="7" pageNumber="182">13</figureCitation>
E–F, 14E–F
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<paragraph id="8B895E223D09FFC9FF3CF55893A6FD14" blockId="7.[151,1437,620,1907]" pageId="7" pageNumber="182">
<emphasis id="B94282303D09FFC9FF3CF558966FFD5C" bold="true" box="[151,325,620,647]" pageId="7" pageNumber="182">Type material.</emphasis>
<materialsCitation id="3B5E547F3D09FFC9FEE6F558923EFD70" collectingDate="2011-01-04" collectionCode="IZCAS" collectorName="Z. Chen &amp; Z. Zha" country="China" county="Lingyun County" elevation="505" latitude="24.365435" location="Naling Cave" longLatPrecision="1" longitude="106.565186" municipality="Baise" pageId="7" pageNumber="182" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
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<typeStatus id="548DE0803D09FFC9FEE6F5589694FD5C" box="[333,446,621,647]" pageId="7" pageNumber="182" type="holotype">Holotype</typeStatus>
:
</emphasis>
Male (
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Ar39854), 
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(
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, 
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, 
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elevation 
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), 
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, 
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, 
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, 
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<collectingRegion id="49F290C03D09FFC9FD2FF5A595C1FD70" box="[644,747,656,683]" country="China" name="Guangxi" pageId="7" pageNumber="182">Guangxi</collectingRegion>
</emphasis>
, 
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<collectingCountry id="F3211EB23D09FFC9FD53F5A5946BFD70" box="[760,833,656,683]" name="China" pageId="7" pageNumber="182">China</collectingCountry>
</emphasis>
, 
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<collectingDate id="EFCC810A3D09FFC9FCFBF5A494D7FD70" box="[848,1021,657,683]" pageId="7" pageNumber="182" value="2011-01-04">4 January 2011</collectingDate>
</date>
, 
<collectorName id="26C33BF43D09FFC9FBA3F5A49348FD70" box="[1032,1122,657,683]" pageId="7" pageNumber="182">Z. Chen</collectorName>
and 
<collectorName id="26C33BF43D09FFC9FB32F5A493CEFD70" box="[1177,1252,657,683]" pageId="7" pageNumber="182">Z. Zha</collectorName>
leg.
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<emphasis id="B94282303D09FFC9FAB0F5A492B6FD70" bold="true" box="[1307,1436,657,683]" pageId="7" pageNumber="182">
<typeStatus id="548DE0803D09FFC9FAB0F5A492BDFD70" box="[1307,1431,657,683]" pageId="7" pageNumber="182" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D3095AB3D09FFC9FF3CF58097C8FD14" box="[151,226,693,719]" pageId="7" pageNumber="182" type="male">1 male</specimenCount>
(
<collectionCode id="ED27C6E73D09FFC9FF5AF580966EFD14" box="[241,324,693,719]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="7" pageNumber="182">IZCAS</collectionCode>
Ar39855) and
</materialsCitation>
<materialsCitation id="3B5E547F3D09FFC9FE46F58093A3FD14" box="[493,1161,692,719]" collectingDate="2011-01-04" collectionCode="IZCAS" collectorName="Z. Chen &amp; Z. Zha" country="China" county="Lingyun County" elevation="505" latitude="24.365435" location="Naling Cave" longLatPrecision="1" longitude="106.565186" municipality="Baise" pageId="7" pageNumber="182" specimenCount="2" specimenCount-female="2" stateProvince="Guangxi" typeStatus="paratype">
<specimenCount id="9D3095AB3D09FFC9FE46F5809573FD14" box="[493,601,693,719]" pageId="7" pageNumber="182" type="female">2 females</specimenCount>
(
<collectionCode id="ED27C6E73D09FFC9FDCCF5809590FD14" box="[615,698,693,719]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="7" pageNumber="182">IZCAS</collectionCode>
Ar39944–39945), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32C0DA93D09FFC9FF6CF5EC9356FD28" box="[199,1148,729,755]" pageId="7" pageNumber="182" type="etymology">
<paragraph id="8B895E223D09FFC9FF6CF5EC9356FD28" blockId="7.[151,1437,620,1907]" box="[199,1148,729,755]" pageId="7" pageNumber="182">
<emphasis id="B94282303D09FFC9FF6CF5EC9664FD28" bold="true" box="[199,334,729,755]" pageId="7" pageNumber="182">Etymology.</emphasis>
The specific name refers to the 
<typeStatus id="548DE0803D09FFC9FD1EF5EC95CFFD28" box="[693,741,729,755]" pageId="7" pageNumber="182">type</typeStatus>
locality and is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C32C0DA93D09FFC9FF6CF5C99566FC10" pageId="7" pageNumber="182" type="diagnosis">
<paragraph id="8B895E223D09FFC9FF6CF5C99566FC10" blockId="7.[151,1437,620,1907]" pageId="7" pageNumber="182">
<emphasis id="B94282303D09FFC9FF6CF5C99668FCCC" bold="true" box="[199,322,764,791]" pageId="7" pageNumber="182">Diagnosis.</emphasis>
This species resembles 
<taxonomicName id="4C3625A13D09FFC9FDFFF5CB9473FCCC" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" box="[596,857,764,791]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="182" phylum="Arthropoda" rank="species" species="phurua">
<emphasis id="B94282303D09FFC9FDFFF5CB95EFFCCC" box="[596,709,765,791]" italics="true" pageId="7" pageNumber="182">B. phurua</emphasis>
Huber, 2005
</taxonomicName>
(see 
<bibRefCitation id="EFA723D33D09FFC9FC3DF5C8937DFCCC" author="Huber, B. A." box="[918,1111,764,791]" pageId="7" pageNumber="182" pagination="1 - 126" refId="ref8339" refString="Huber, B. A. (2005 a) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica, 155, 1 - 126." type="journal article" year="2005">Huber 2005a: 37</bibRefCitation>
, figs 232–235 and 
<bibRefCitation id="EFA723D33D09FFC9FA84F5C897D1FCE0" author="Yao, Z. &amp; Pham, D. S. &amp; Li, S." pageId="7" pageNumber="182" pagination="1 - 82" refId="ref8985" refString="Yao, Z., Pham, D. S. &amp; Li, S. (2015) Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species. Zootaxa, 3909 (1), 1 - 82. https: // doi. org / 10.11646 / zootaxa. 3909.1.1" type="journal article" year="2015">
Yao 
<emphasis id="B94282303D09FFC9FAC8F5CB92B7FCCC" box="[1379,1437,765,791]" italics="true" pageId="7" pageNumber="182">et al.</emphasis>
2015: 10
</bibRefCitation>
, figs 28A–E) with similar male chelicerae (
<figureCitation id="130D42A73D09FFC9FD4BF4149413FCE7" box="[736,825,801,828]" captionStart="FIGURE 6" captionStartId="9.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@9.[214,1373,181,1885]" captionTargetPageId="9" captionText="FIGURE 6. Belisana naling sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3937048" httpUri="https://zenodo.org/record/3937048/files/figure.png" pageId="7" pageNumber="182">Fig. 6D</figureCitation>
), bulbal apophyses (
<figureCitation id="130D42A73D09FFC9FBB7F4149358FCE7" box="[1052,1138,801,828]" captionStart="FIGURE 6" captionStartId="9.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@9.[214,1373,181,1885]" captionTargetPageId="9" captionText="FIGURE 6. Belisana naling sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3937048" httpUri="https://zenodo.org/record/3937048/files/figure.png" pageId="7" pageNumber="182">Fig. 6C</figureCitation>
) and external female genitalia (
<figureCitation id="130D42A73D09FFC9FF7DF470961EFC84" box="[214,308,837,863]" captionStart="FIGURE 6" captionStartId="9.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@9.[214,1373,181,1885]" captionTargetPageId="9" captionText="FIGURE 6. Belisana naling sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3937048" httpUri="https://zenodo.org/record/3937048/files/figure.png" pageId="7" pageNumber="182">Figs 6A</figureCitation>
, 
<figureCitation id="130D42A73D09FFC9FEEBF470965AFC84" box="[320,368,837,863]" captionStart="FIGURE 14" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[202,1387,196,1913]" captionTargetId="figure@19.[193,1395,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. guilin sp. nov.; C–D. B. maoer sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. naling sp. nov.; G–H. B. xiyuan sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3937064" httpUri="https://zenodo.org/record/3937064/files/figure.png" pageId="7" pageNumber="182">14E</figureCitation>
), but males can be distinguished by absence of spine and disto-prolateral sclerites on procursus (
<figureCitation id="130D42A73D09FFC9FF34F45C97CDFC58" box="[159,231,872,899]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@8.[226,1361,181,1850]" captionTargetPageId="8" captionText="FIGURE 5. Belisana naling sp. nov., holotype male. A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at large proximo-dorsal hump on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large, distal membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-ventral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3937046" httpUri="https://zenodo.org/record/3937046/files/figure.png" pageId="7" pageNumber="182">Figs 5</figureCitation>
A–D, 13E–F), and by presence of small subdisto-dorsal sclerite on procursus (arrow 
<quantity id="4CCEF3C73D09FFC9FB16F45C93C1FC58" box="[1213,1259,873,899]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="7" pageNumber="182" unit="in" value="2.0">2 in</quantity>
<figureCitation id="130D42A73D09FFC9FB5FF45C927BFC58" box="[1268,1361,872,899]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@8.[226,1361,181,1850]" captionTargetPageId="8" captionText="FIGURE 5. Belisana naling sp. nov., holotype male. A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at large proximo-dorsal hump on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large, distal membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-ventral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3937046" httpUri="https://zenodo.org/record/3937046/files/figure.png" pageId="7" pageNumber="182">Figs 5C</figureCitation>
, 
<figureCitation id="130D42A73D09FFC9FAF4F45C92A5FC58" box="[1375,1423,873,899]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). A–B. B. guilin sp. nov.; C–D. B. maoer sp. nov.; E–F. B. naling sp. nov.; G–H. B. xiyuan sp. nov.; I–J. B. xuanguan sp. nov.; K–L. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="7" pageNumber="182">13E</figureCitation>
). Females can be distinguished by having W-shaped vulval anterior arch (
<figureCitation id="130D42A73D09FFC9FC6FF4B8930AFC7C" box="[964,1056,909,935]" captionStart="FIGURE 6" captionStartId="9.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@9.[214,1373,181,1885]" captionTargetPageId="9" captionText="FIGURE 6. Belisana naling sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3937048" httpUri="https://zenodo.org/record/3937048/files/figure.png" pageId="7" pageNumber="182">Figs 6B</figureCitation>
, 
<figureCitation id="130D42A73D09FFC9FB87F4B89373FC7C" box="[1068,1113,909,935]" captionStart="FIGURE 14" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[202,1387,196,1913]" captionTargetId="figure@19.[193,1395,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. guilin sp. nov.; C–D. B. maoer sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. naling sp. nov.; G–H. B. xiyuan sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3937064" httpUri="https://zenodo.org/record/3937064/files/figure.png" pageId="7" pageNumber="182">14F</figureCitation>
) and by small elliptic vulval pore plates (arrow pp in 
<figureCitation id="130D42A73D09FFC9FE01F484952EFC10" box="[426,516,945,971]" captionStart="FIGURE 6" captionStartId="9.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@9.[214,1373,181,1885]" captionTargetPageId="9" captionText="FIGURE 6. Belisana naling sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3937048" httpUri="https://zenodo.org/record/3937048/files/figure.png" pageId="7" pageNumber="182">Figs 6B</figureCitation>
, 
<figureCitation id="130D42A73D09FFC9FDBBF4849515FC10" box="[528,575,945,971]" captionStart="FIGURE 14" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[202,1387,196,1913]" captionTargetId="figure@19.[193,1395,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. guilin sp. nov.; C–D. B. maoer sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. naling sp. nov.; G–H. B. xiyuan sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3937064" httpUri="https://zenodo.org/record/3937064/files/figure.png" pageId="7" pageNumber="182">14F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32C0DA93D09FFC9FF6CF4E193DAF894" pageId="7" pageNumber="182" type="description">
<paragraph id="8B895E223D09FFC9FF6CF4E196F9F9D0" blockId="7.[151,1437,620,1907]" pageId="7" pageNumber="182">
<emphasis id="B94282303D09FFC9FF6CF4E196B5FC34" bold="true" box="[199,415,980,1007]" pageId="7" pageNumber="182">Description. Male</emphasis>
(
<emphasis id="B94282303D09FFC9FE19F4E09533FC34" bold="true" box="[434,537,981,1007]" pageId="7" pageNumber="182">
<typeStatus id="548DE0803D09FFC9FE19F4E09533FC34" box="[434,537,981,1007]" pageId="7" pageNumber="182" type="holotype">holotype</typeStatus>
</emphasis>
, IZCAS Ar39854): Total length 1.14 (1.22 with clypeus), carapace 0.51 long, 0.56 wide, opisthosoma 0.63 long, 0.41 wide. Leg I: 11.08 (3.01 + 0.38 + 2.76 + 3.97 + 0.96), leg II: 9.68 (2.56 + 0.26 + 2.44 + 3.53 + 0.89), leg III: 6.86 (1.92 + 0.19 + 1.60 + 2.44 + 0.71), leg IV: 9.62 (2.88 + 0.26 + 2.44 + 3.21 + 0.83); tibia I L/d: 39. Distance PME-PME 0.19, diameter PME 0.02, distance PME-ALE 0.03, AME absent. Sternum slightly wider than long (0.47/0.38). Habitus as in 
<figureCitation id="130D42A73D09FFC9FCABF350946EFBA4" box="[768,836,1125,1151]" captionStart="FIGURE 6" captionStartId="9.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@9.[214,1373,181,1885]" captionTargetPageId="9" captionText="FIGURE 6. Belisana naling sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3937048" httpUri="https://zenodo.org/record/3937048/files/figure.png" pageId="7" pageNumber="182">Figs 6</figureCitation>
E–F. Carapace and sternum pale, without marks. Legs pale, without darker rings. Opisthosoma pale, without spots. Ocular area not elevated. Eyes without pigment but apparently with small lenses. Thoracic furrow absent. Clypeus unmodified. Chelicerae (
<figureCitation id="130D42A73D09FFC9FBC4F39893ECFB13" box="[1135,1222,1197,1224]" captionStart="FIGURE 6" captionStartId="9.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@9.[214,1373,181,1885]" captionTargetPageId="9" captionText="FIGURE 6. Belisana naling sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3937048" httpUri="https://zenodo.org/record/3937048/files/figure.png" pageId="7" pageNumber="182">Fig. 6D</figureCitation>
) with pair of small proximo-lateral apophyses and pair of long, curved distal apophyses each with hardened and pointed tip (distance between tips of distal apophyses: 0.29). Pedipalps as in 
<figureCitation id="130D42A73D09FFC9FCA2F3C09467FAD4" box="[777,845,1268,1295]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@8.[226,1361,181,1850]" captionTargetPageId="8" captionText="FIGURE 5. Belisana naling sp. nov., holotype male. A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at large proximo-dorsal hump on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large, distal membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-ventral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3937046" httpUri="https://zenodo.org/record/3937046/files/figure.png" pageId="7" pageNumber="182">Figs 5</figureCitation>
A–B; coxa unmodified; trochanter with short ventral apophysis provided with small retrolateral apophysis (arrow 
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<figureCitation id="130D42A73D09FFC9FCDCF22C94E5FAE8" box="[887,975,1304,1331]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@8.[226,1361,181,1850]" captionTargetPageId="8" captionText="FIGURE 5. Belisana naling sp. nov., holotype male. A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at large proximo-dorsal hump on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large, distal membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-ventral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3937046" httpUri="https://zenodo.org/record/3937046/files/figure.png" pageId="7" pageNumber="182">Fig. 5B</figureCitation>
); femur with large proximo-dorsal hump (arrow 
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<figureCitation id="130D42A73D09FFC9FEBCF2089647FA8C" box="[279,365,1340,1367]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@8.[226,1361,181,1850]" captionTargetPageId="8" captionText="FIGURE 5. Belisana naling sp. nov., holotype male. A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at large proximo-dorsal hump on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large, distal membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-ventral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3937046" httpUri="https://zenodo.org/record/3937046/files/figure.png" pageId="7" pageNumber="182">Fig. 5B</figureCitation>
); procursus (
<figureCitation id="130D42A73D09FFC9FE50F208956AFA8C" box="[507,576,1340,1367]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@8.[226,1361,181,1850]" captionTargetPageId="8" captionText="FIGURE 5. Belisana naling sp. nov., holotype male. A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at large proximo-dorsal hump on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large, distal membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-ventral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3937046" httpUri="https://zenodo.org/record/3937046/files/figure.png" pageId="7" pageNumber="182">Figs 5</figureCitation>
A–D, 13E–F) simple proximally but complex distally, with large, distal membranous lamella (arrow 
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<figureCitation id="130D42A73D09FFC9FE7AF2549501FAA0" box="[465,555,1376,1403]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@8.[226,1361,181,1850]" captionTargetPageId="8" captionText="FIGURE 5. Belisana naling sp. nov., holotype male. A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at large proximo-dorsal hump on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large, distal membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-ventral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3937046" httpUri="https://zenodo.org/record/3937046/files/figure.png" pageId="7" pageNumber="182">Figs 5C</figureCitation>
, 
<figureCitation id="130D42A73D09FFC9FD9DF254954CFAA0" box="[566,614,1377,1403]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). A–B. B. guilin sp. nov.; C–D. B. maoer sp. nov.; E–F. B. naling sp. nov.; G–H. B. xiyuan sp. nov.; I–J. B. xuanguan sp. nov.; K–L. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="7" pageNumber="182">13E</figureCitation>
), small, disto-ventral membranous process (arrow in 
<figureCitation id="130D42A73D09FFC9FB1CF2549238FAA0" box="[1207,1298,1376,1403]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@8.[226,1361,181,1850]" captionTargetPageId="8" captionText="FIGURE 5. Belisana naling sp. nov., holotype male. A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at large proximo-dorsal hump on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large, distal membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-ventral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3937046" httpUri="https://zenodo.org/record/3937046/files/figure.png" pageId="7" pageNumber="182">Figs 5D</figureCitation>
, 
<figureCitation id="130D42A73D09FFC9FAB6F2549261FAA0" box="[1309,1355,1377,1403]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). A–B. B. guilin sp. nov.; C–D. B. maoer sp. nov.; E–F. B. naling sp. nov.; G–H. B. xiyuan sp. nov.; I–J. B. xuanguan sp. nov.; K–L. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="7" pageNumber="182">13F</figureCitation>
), small subdisto-dorsal sclerite (arrow 
<quantity id="4CCEF3C73D09FFC9FE5EF2B0950BFA44" box="[501,545,1413,1439]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="7" pageNumber="182" unit="in" value="2.0">2 in</quantity>
<figureCitation id="130D42A73D09FFC9FD82F2B095AEFA44" box="[553,644,1412,1439]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@8.[226,1361,181,1850]" captionTargetPageId="8" captionText="FIGURE 5. Belisana naling sp. nov., holotype male. A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at large proximo-dorsal hump on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large, distal membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-ventral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3937046" httpUri="https://zenodo.org/record/3937046/files/figure.png" pageId="7" pageNumber="182">Figs 5C</figureCitation>
, 
<figureCitation id="130D42A73D09FFC9FD3AF2B09594FA44" box="[657,702,1413,1439]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). A–B. B. guilin sp. nov.; C–D. B. maoer sp. nov.; E–F. B. naling sp. nov.; G–H. B. xiyuan sp. nov.; I–J. B. xuanguan sp. nov.; K–L. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="7" pageNumber="182">13E</figureCitation>
) and large subdisto-prolateral sclerite (arrow 
<quantity id="4CCEF3C73D09FFC9FB68F2B093C5FA44" box="[1219,1263,1413,1439]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="182" unit="in" value="3.0">3 in</quantity>
<figureCitation id="130D42A73D09FFC9FB5CF2B09278FA44" box="[1271,1362,1412,1439]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@8.[226,1361,181,1850]" captionTargetPageId="8" captionText="FIGURE 5. Belisana naling sp. nov., holotype male. A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at large proximo-dorsal hump on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large, distal membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-ventral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3937046" httpUri="https://zenodo.org/record/3937046/files/figure.png" pageId="7" pageNumber="182">Figs 5C</figureCitation>
, 
<figureCitation id="130D42A73D09FFC9FAF4F2B092A5FA44" box="[1375,1423,1413,1439]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). A–B. B. guilin sp. nov.; C–D. B. maoer sp. nov.; E–F. B. naling sp. nov.; G–H. B. xiyuan sp. nov.; I–J. B. xuanguan sp. nov.; K–L. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="7" pageNumber="182">13E</figureCitation>
), without distal spine; bulb (
<figureCitation id="130D42A73D09FFC9FE6EF29C9536FA1F" box="[453,540,1449,1476]" captionStart="FIGURE 6" captionStartId="9.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@9.[214,1373,181,1885]" captionTargetPageId="9" captionText="FIGURE 6. Belisana naling sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3937048" httpUri="https://zenodo.org/record/3937048/files/figure.png" pageId="7" pageNumber="182">Fig. 6C</figureCitation>
) with long hooked apophysis and simple embolus. Retrolateral trichobothria of tibia I at 11% proximally; legs with short vertical setae on metatarsi, without spines and curved setae; tarsus I with 21 distinct pseudosegments.
</paragraph>
<paragraph id="8B895E223D09FFC9FF6CF12092BDF938" blockId="7.[151,1437,620,1907]" pageId="7" pageNumber="182">
<emphasis id="B94282303D09FFC9FF6CF1209634F9F4" bold="true" box="[199,286,1557,1583]" pageId="7" pageNumber="182">Female</emphasis>
(IZCAS Ar39944): Similar to male, habitus as in 
<figureCitation id="130D42A73D09FFC9FCCBF120948CF9F4" box="[864,934,1557,1583]" captionStart="FIGURE 6" captionStartId="9.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@9.[214,1373,181,1885]" captionTargetPageId="9" captionText="FIGURE 6. Belisana naling sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3937048" httpUri="https://zenodo.org/record/3937048/files/figure.png" pageId="7" pageNumber="182">Figs 6</figureCitation>
G–H. Total length 1.25 (1.38 with clypeus), carapace 0.50 long, 0.63 wide, opisthosoma 0.75 long, 0.53 wide. Tibia I: 3.14; tibia I L/d: 32. Distance PME-PME 0.22, diameter PME 0.03, distance PME-ALE 0.04, AME absent. Sternum slightly longer than wide (0.47/0.31). Eyes with pigment but indistinct. External female genitalia (
<figureCitation id="130D42A73D09FFC9FC85F1B494A1F940" box="[814,907,1665,1691]" captionStart="FIGURE 6" captionStartId="9.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@9.[214,1373,181,1885]" captionTargetPageId="9" captionText="FIGURE 6. Belisana naling sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3937048" httpUri="https://zenodo.org/record/3937048/files/figure.png" pageId="7" pageNumber="182">Figs 6A</figureCitation>
, 
<figureCitation id="130D42A73D09FFC9FC3FF1B494E9F940" box="[916,963,1665,1691]" captionStart="FIGURE 14" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[202,1387,196,1913]" captionTargetId="figure@19.[193,1395,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. guilin sp. nov.; C–D. B. maoer sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. naling sp. nov.; G–H. B. xiyuan sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3937064" httpUri="https://zenodo.org/record/3937064/files/figure.png" pageId="7" pageNumber="182">14E</figureCitation>
) simple and flat, with pair of frontal “epigynal” pockets laterally on sclerotized plate (arrow ep in 
<figureCitation id="130D42A73D09FFC9FCA7F1909478F964" box="[780,850,1701,1727]" captionStart="FIGURE 6" captionStartId="9.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@9.[214,1373,181,1885]" captionTargetPageId="9" captionText="FIGURE 6. Belisana naling sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3937048" httpUri="https://zenodo.org/record/3937048/files/figure.png" pageId="7" pageNumber="182">Figs 6</figureCitation>
A–B, 14E–F), 0.28 apart. Vulva (
<figureCitation id="130D42A73D09FFC9FB6CF1909208F964" box="[1223,1314,1701,1727]" captionStart="FIGURE 6" captionStartId="9.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@9.[214,1373,181,1885]" captionTargetPageId="9" captionText="FIGURE 6. Belisana naling sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3937048" httpUri="https://zenodo.org/record/3937048/files/figure.png" pageId="7" pageNumber="182">Figs 6B</figureCitation>
, 
<figureCitation id="130D42A73D09FFC9FA85F1909271F964" box="[1326,1371,1701,1727]" captionStart="FIGURE 14" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[202,1387,196,1913]" captionTargetId="figure@19.[193,1395,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. guilin sp. nov.; C–D. B. maoer sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. naling sp. nov.; G–H. B. xiyuan sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3937064" httpUri="https://zenodo.org/record/3937064/files/figure.png" pageId="7" pageNumber="182">14F</figureCitation>
) with W-shaped anterior arch and pair of small elliptic pore plates (arrow pp in 
<figureCitation id="130D42A73D09FFC9FC64F1FC9303F938" box="[975,1065,1737,1763]" captionStart="FIGURE 6" captionStartId="9.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@9.[214,1373,181,1885]" captionTargetPageId="9" captionText="FIGURE 6. Belisana naling sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3937048" httpUri="https://zenodo.org/record/3937048/files/figure.png" pageId="7" pageNumber="182">Figs 6B</figureCitation>
, 
<figureCitation id="130D42A73D09FFC9FB9EF1FC9349F938" box="[1077,1123,1737,1763]" captionStart="FIGURE 14" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[202,1387,196,1913]" captionTargetId="figure@19.[193,1395,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. guilin sp. nov.; C–D. B. maoer sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. naling sp. nov.; G–H. B. xiyuan sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3937064" httpUri="https://zenodo.org/record/3937064/files/figure.png" pageId="7" pageNumber="182">14F</figureCitation>
), without serrated sclerites.
</paragraph>
<paragraph id="8B895E223D09FFC9FF6CF1D99661F8F0" blockId="7.[151,1437,620,1907]" pageId="7" pageNumber="182">
<emphasis id="B94282303D09FFC9FF6CF1D9966BF8DC" bold="true" box="[199,321,1772,1799]" pageId="7" pageNumber="182">Variation:</emphasis>
<materialsCitation id="3B5E547F3D09FFC9FEE7F1D89662F8F0" collectionCode="IZCAS" collectorName="Tibia I" pageId="7" pageNumber="182" specimenCount="2" typeStatus="paratype">
<collectorName id="26C33BF43D09FFC9FEE7F1D896B1F8DC" box="[332,411,1773,1799]" pageId="7" pageNumber="182">Tibia I</collectorName>
in male 
<typeStatus id="548DE0803D09FFC9FDA7F1D89546F8DC" box="[524,620,1773,1799]" pageId="7" pageNumber="182" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED27C6E73D09FFC9FD2BF1D895F9F8DC" box="[640,723,1773,1799]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="7" pageNumber="182">IZCAS</collectionCode>
Ar39855): 2.56. 
<collectorName id="26C33BF43D09FFC9FC0AF1D894DAF8DC" box="[929,1008,1773,1799]" pageId="7" pageNumber="182">Tibia I</collectorName>
in another female 
<typeStatus id="548DE0803D09FFC9FB7DF1D8921CF8DC" box="[1238,1334,1773,1799]" pageId="7" pageNumber="182" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED27C6E73D09FFC9FAE1F1D892B7F8DC" box="[1354,1437,1773,1799]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="7" pageNumber="182">IZCAS</collectionCode>
Ar39945): 2.24
</materialsCitation>
.
</paragraph>
<paragraph id="8B895E223D09FFC9FF6CF00093DAF894" blockId="7.[151,1437,620,1907]" box="[199,1264,1844,1871]" pageId="7" pageNumber="182">
<emphasis id="B94282303D09FFC9FF6CF00096AFF894" bold="true" box="[199,389,1844,1871]" pageId="7" pageNumber="182">Natural history.</emphasis>
The species was found underside of rocks in the aphotic zone inside the cave.
</paragraph>
</subSubSection>
<subSubSection id="C32C0DA93D09FFC9FF6CF06D9471F8A8" box="[199,859,1880,1907]" pageId="7" pageNumber="182" type="distribution">
<paragraph id="8B895E223D09FFC9FF6CF06D9471F8A8" blockId="7.[151,1437,620,1907]" box="[199,859,1880,1907]" pageId="7" pageNumber="182">
<emphasis id="B94282303D09FFC9FF6CF06D964BF8A8" bold="true" box="[199,353,1880,1907]" pageId="7" pageNumber="182">Distribution.</emphasis>
<collectingCountry id="F3211EB23D09FFC9FECCF06C9680F8A8" box="[359,426,1881,1907]" name="China" pageId="7" pageNumber="182">China</collectingCountry>
(
<collectingRegion id="49F290C03D09FFC9FE12F06C9534F8A8" box="[441,542,1881,1907]" country="China" name="Guangxi" pageId="7" pageNumber="182">Guangxi</collectingRegion>
, 
<typeStatus id="548DE0803D09FFC9FD82F06C9573F8A8" box="[553,601,1881,1907]" pageId="7" pageNumber="182">type</typeStatus>
locality only; 
<figureCitation id="130D42A73D09FFC9FD57F06C9464F8A8" box="[764,846,1880,1907]" captionStart="FIGURE 15" captionStartId="20.[151,250,1181,1206]" captionTargetBox="[151,1436,181,1156]" captionTargetId="figure@20.[151,1436,181,1156]" captionTargetPageId="20" captionText="FIGURE 15. Distribution records of six new species of the genus Belisana from southern China. 1. B. guilin sp. nov.; 2. B. maoer sp. nov.; 3. B. naling sp. nov.; 4. B. xiyuan sp. nov.; 5. B. xuanguan sp. nov.; 6. B. yuexiu sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3937066" httpUri="https://zenodo.org/record/3937066/files/figure.png" pageId="7" pageNumber="182">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>