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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.661.10677" ID-GBIF-Dataset="9bee49e3-7184-47ae-8c81-8087c21739a6" ID-PMC="PMC5539366" ID-Pensoft-Pub="1313-2970-661-15" ID-PubMed="28769602" ID-ZBK="7ADEBBF244A04276AB7D1EB2BBFD8953" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-661-15" ModsDocOrigin="ZooKeys 661" ModsDocTitle="Taxonomic notes on the armored spiders of the families Pacullidae and Tetrablemmidae (Arachnida, Araneae) from Singapore" checkinTime="1489572793026" checkinUser="pensoft" docAuthor="Lin, Yucheng, Koh, Joseph K. H., Koponen, Seppo &amp; Li, Shuqiang" docDate="2017" docId="A4265D80A53F3EEADF848C5861F6E866" docLanguage="en" docName="ZooKeys 661: 15-60" docOrigin="ZooKeys 661" docSource="http://dx.doi.org/10.3897/zookeys.661.10677" docTitle="Paculla globosa Lin &amp; Li, sp. n." docType="treatment" docUuid="CAD62F40-4F41-4D91-8E0E-7E9AAAA1632C" docUuidSource="ZooBank" docVersion="5" lastPageNumber="19" masterDocId="D064C572FFEFFFD88F28FFD23F2FFFDE" masterDocTitle="Taxonomic notes on the armored spiders of the families Pacullidae and Tetrablemmidae (Arachnida, Araneae) from Singapore" masterLastPageNumber="60" masterPageNumber="15" pageNumber="17" updateTime="1668164295527" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic notes on the armored spiders of the families Pacullidae and Tetrablemmidae (Arachnida, Araneae) from Singapore</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lin, Yucheng</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Koh, Joseph K. H.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Koponen, Seppo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Shuqiang</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>661</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>15</mods:start>
<mods:end>60</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.661.10677</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.661.10677</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-661-15</mods:identifier>
<mods:identifier type="ZBK">7ADEBBF244A04276AB7D1EB2BBFD8953</mods:identifier>
<mods:identifier type="ZooBank">7ADEBBF244A04276AB7D1EB2BBFD8953</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127875378" LSID="urn:lsid:zoobank.org:act:CAD62F40-4F41-4D91-8E0E-7E9AAAA1632C" httpUri="http://treatment.plazi.org/id/A4265D80A53F3EEADF848C5861F6E866" lastPageId="4" lastPageNumber="19" pageId="2" pageNumber="17">
<subSubSection pageId="2" pageNumber="17" type="nomenclature">
<paragraph pageId="2" pageNumber="17">
<taxonomicName LSID="http://zoobank.org/CAD62F40-4F41-4D91-8E0E-7E9AAAA1632C" authority="Lin &amp; Li" class="Arachnida" family="Pacullidae" genus="Paculla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paculla globosa" order="Araneae" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="globosa">Paculla globosa Lin &amp; Li</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="17">sp. n.</taxonomicNameLabel>
Figs 4, 5, 6
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="18" type="type material">
<paragraph pageId="3" pageNumber="18">
<pageBreakToken pageId="3" pageNumber="18" start="start">Type</pageBreakToken>
material.
</paragraph>
<paragraph pageId="3" pageNumber="18">
Holotype ♂ (LKCNHM), SINGAPORE: Bukit Timah Nature Reserve, Jungle Fall Stream, altitude 118 m,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="1.3570555">1°21'25.4&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="103.7737">103°46'25.3&quot;E</geoCoordinate>
, 21 August 2015, S. Li and Y. Tong leg. Paratype 1♀ (LKCNHM), SINGAPORE: Bukit Timah Nature Reserve, Bukit Timah Summit, altitude 163 m,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="1.3546389">1°21'16.7&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="103.77639">103°46'35.0&quot;E</geoCoordinate>
, 19 August 2015, S. Li and Y. Tong leg.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="18" type="etymology">
<paragraph pageId="3" pageNumber="18">Etymology.</paragraph>
<paragraph pageId="3" pageNumber="18">
The specific epithet derives from the Latin word
<normalizedToken originalValue="“globosus”">&quot;globosus&quot;</normalizedToken>
= globular, drawing attention to the shape of palpal bulb; adjective.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="18" type="diagnosis">
<paragraph pageId="3" pageNumber="18">Diagnosis.</paragraph>
<paragraph pageId="3" pageNumber="18">
This new species can be distinguished from all congeners with the exception of
<taxonomicName class="Arachnida" family="Pacullidae" genus="Paculla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paculla sulaimani" order="Araneae" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="sulaimani">Paculla sulaimani</taxonomicName>
Lehtinen, 1981 by the globular bulb (Fig. 5
<normalizedToken originalValue="CD">C-D</normalizedToken>
), the long, finger-like embolus (Fig. 5
<normalizedToken originalValue="AD">A-D</normalizedToken>
) and the presence of three bridge fragments of MV (Fig. 6B). Its male differs from
<taxonomicName class="Arachnida" family="Pacullidae" genus="Paculla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paculla sulaimani" order="Araneae" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="sulaimani">Paculla sulaimani</taxonomicName>
(see Lehtinen, 1981: 18, fig. 13) by the narrower embolus with weakly lobed end (Fig. 5A,
<normalizedToken originalValue="CD">C-D</normalizedToken>
). The female differs from
<taxonomicName class="Arachnida" family="Pacullidae" genus="Paculla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paculla bukittimahensis" order="Araneae" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="bukittimahensis">Paculla bukittimahensis</taxonomicName>
sp. n. (Fig. 3B) by the anterior median part of the atrium protruding broadly and the larger, bridge fragments of MV almost touching each other (Fig. 6B).
</paragraph>
<caption pageId="3" pageNumber="18">
<paragraph pageId="3" pageNumber="18">
Figure 4.
<taxonomicName class="Arachnida" family="Pacullidae" genus="Paculla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paculla globosa" order="Araneae" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="globosa">Paculla globosa</taxonomicName>
sp. n., male holotype (
<normalizedToken originalValue="AB">A-B</normalizedToken>
, E) and female paratype (
<normalizedToken originalValue="CD">C-D</normalizedToken>
,
<normalizedToken originalValue="FG">F-G</normalizedToken>
).
<normalizedToken originalValue="AF">A-F</normalizedToken>
habitus G genital area (untreated). A, C dorsal B, D, G ventral
<normalizedToken originalValue="EF">E-F</normalizedToken>
lateral. Abbreviations: A = anal plate; AV = anterior ventrolateral plate; L = lateral plate; MV = median ventrolateral plate; P = pulmonary plate; PA = preanal plate; PG = postgenital plate. Scale bars: 0.50 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="19" pageId="3" pageNumber="18" type="description">
<paragraph pageId="3" pageNumber="18">Description.</paragraph>
<paragraph pageId="3" pageNumber="18">Male (holotype). Coloration: body dark reddish brown; legs reddish-brown.</paragraph>
<paragraph pageId="3" pageNumber="18">Measurements: total length 4.40; carapace 1.95 long, 1.40 wide, 1.20 high; abdomen 2.50 long, 1.90 wide, 2.30 high; clypeus 0.49 high; sternum 1.30 long, 1.00 wide. Length of legs: I 8.28 (2.53, 0.65, 2.35, 1.75, 1.00); II 6.95 (2.10, 0.60, 1.85, 1.50, 0.90); III 5.75 (1.70, 0.54, 1.41, 1.30, 0.80); IV 8.16 (2.50, 0.62, 2.30, 1.91, 0.83).</paragraph>
<paragraph lastPageId="4" lastPageNumber="19" pageId="3" pageNumber="18">
Prosoma (Fig. 4
<normalizedToken originalValue="AB">A-B</normalizedToken>
, E): carapace low, sparsely granulated over whole surface, but surface between the granules smooth; eyes white, ALE=AME=PLE; cephalic part moderately raised; cervical groove distinct; thoracic fovea shallow; clypeus vertical an
<pageBreakToken pageId="4" pageNumber="19" start="start">teriorly</pageBreakToken>
; labium triangular, distally obtuse; sternum rough, covered with thin setae, posterior corner protruded. Legs: cuticle striated, weakly granular.
</paragraph>
<paragraph pageId="4" pageNumber="19">
Opisthosoma (Fig. 4
<normalizedToken originalValue="AB">A-B</normalizedToken>
, E): dorsal scutum long, oval, covered with thin setae, margin rugose, center modified by sparse pits; rows of small sclerites between lateral scuta, lateral scutum I short; ventral scutum smooth, margin rugose; perigenital scutum broad, triangular.
</paragraph>
<paragraph pageId="4" pageNumber="19">
Palp (Fig. 5
<normalizedToken originalValue="AD">A-D</normalizedToken>
): femoral cuticle smooth, approximately two times as long as patella; patella proximally narrow, distally wide; tibia broad, swollen, 1.3 times as wide as femur; cymbium compressed, slightly shorter than tibia, distally obtuse bifurcate; bulb globular, surface smooth (Fig. 5
<normalizedToken originalValue="CD">C-D</normalizedToken>
); embolus long, clavate, starting from subdistal-ventral 1/3 position of bulbous surface (Fig. 5B), proximally sclerotized and bent, distally shallow furcated; embolic tip faint, with split ends (Fig. 5A).
</paragraph>
<caption pageId="4" pageNumber="19">
<paragraph pageId="4" pageNumber="19">
Figure 5.
<taxonomicName class="Arachnida" family="Pacullidae" genus="Paculla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paculla globosa" order="Araneae" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="globosa">Paculla globosa</taxonomicName>
sp. n., male holotype. A,
<normalizedToken originalValue="CD">C-D</normalizedToken>
left palp B palpal bulb. A anterior B ventral C prolateral D retrolateral. Abbreviations: ep = embolic part of apes of palpal organ. Scale bars: 0.20 mm.
</paragraph>
</caption>
<paragraph pageId="4" pageNumber="19">Female (paratype). Coloration as in male.</paragraph>
<paragraph pageId="4" pageNumber="19">Measurements: total length 4.50; carapace 1.85 long, 1.45 wide, 1.20 high; abdomen 2.67 long, 1.80 wide, 2.21 high; clypeus 0.43 high; sternum 1.14 long, 0.97 wide. Length of legs: I 7.35 (2.25, 0.55, 2.10, 1.65, 0.80); II 6.23 (1.95, 0.53, 1.65, 1.35, 0.75); III 5.22 (1.55, 0.50, 1.30, 1.22, 0.65); IV 7.20 (2.15, 0.54, 2.00, 1.80, 0.71). Length of palp: 1.58 (0.45, 0.21, 0.34, 0.38).</paragraph>
<paragraph pageId="4" pageNumber="19">
Carapace, abdomen, and legs as in male (Fig. 4
<normalizedToken originalValue="CD">C-D</normalizedToken>
, F); clypeus slightly lower than in male.
</paragraph>
<paragraph pageId="4" pageNumber="19">
Genitalia (Figs 4G; 6
<normalizedToken originalValue="AB">A-B</normalizedToken>
): epigynal area strongly sclerotized (Fig. 4G); postgenital scutum rugose, wider than preanal scutum; median ventrolateral plate triangular; preanal scutum rugose, wider than long (Fig. 6A). Vulva with a large, nearly rectangular atrium, laterally rugose, and protruding anteromedially; three adjacent, equal sizes bridge fragments of MV below the atrium, and close to the dorso-posterior margin of postgenital scutum (Fig. 6B).
</paragraph>
<caption pageId="4" pageNumber="19">
<paragraph pageId="4" pageNumber="19">
Figure 6.
<taxonomicName class="Arachnida" family="Pacullidae" genus="Paculla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paculla globosa" order="Araneae" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="globosa">Paculla globosa</taxonomicName>
sp. n., female paratype. A genital area (lactic acid-treated), ventral Bditto, dorsal. Abbreviations: AT = atrium; AV = anterior ventrolateral plate; MV = median ventrolateral plate; MVB = bridge fragments of MV; P = pulmonary plate; PA = preanal plate; PG = postgenital plate; PV = posterior ventrolateral plate. Scale bars: 0.50 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="4" pageNumber="19" type="distribution">
<paragraph pageId="4" pageNumber="19">Distribution.</paragraph>
<paragraph pageId="4" pageNumber="19">Singapore.</paragraph>
</subSubSection>
</treatment>
</document>