180 lines
13 KiB
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180 lines
13 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.661.10677" ID-GBIF-Dataset="9bee49e3-7184-47ae-8c81-8087c21739a6" ID-PMC="PMC5539366" ID-Pensoft-Pub="1313-2970-661-15" ID-PubMed="28769602" ID-ZBK="7ADEBBF244A04276AB7D1EB2BBFD8953" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-661-15" ModsDocOrigin="ZooKeys 661" ModsDocTitle="Taxonomic notes on the armored spiders of the families Pacullidae and Tetrablemmidae (Arachnida, Araneae) from Singapore" checkinTime="1489572793026" checkinUser="pensoft" docAuthor="Lin, Yucheng, Koh, Joseph K. H., Koponen, Seppo & Li, Shuqiang" docDate="2017" docId="A4265D80A53F3EEADF848C5861F6E866" docLanguage="en" docName="ZooKeys 661: 15-60" docOrigin="ZooKeys 661" docSource="http://dx.doi.org/10.3897/zookeys.661.10677" docTitle="Paculla globosa Lin & Li, sp. n." docType="treatment" docUuid="CAD62F40-4F41-4D91-8E0E-7E9AAAA1632C" docUuidSource="ZooBank" docVersion="5" lastPageNumber="19" masterDocId="D064C572FFEFFFD88F28FFD23F2FFFDE" masterDocTitle="Taxonomic notes on the armored spiders of the families Pacullidae and Tetrablemmidae (Arachnida, Araneae) from Singapore" masterLastPageNumber="60" masterPageNumber="15" pageNumber="17" updateTime="1668164295527" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic notes on the armored spiders of the families Pacullidae and Tetrablemmidae (Arachnida, Araneae) from Singapore</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lin, Yucheng</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Koh, Joseph K. H.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Koponen, Seppo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Li, Shuqiang</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>661</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>15</mods:start>
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<mods:end>60</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.661.10677</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.661.10677</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-661-15</mods:identifier>
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<mods:identifier type="ZBK">7ADEBBF244A04276AB7D1EB2BBFD8953</mods:identifier>
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<mods:identifier type="ZooBank">7ADEBBF244A04276AB7D1EB2BBFD8953</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127875378" LSID="urn:lsid:zoobank.org:act:CAD62F40-4F41-4D91-8E0E-7E9AAAA1632C" httpUri="http://treatment.plazi.org/id/A4265D80A53F3EEADF848C5861F6E866" lastPageId="4" lastPageNumber="19" pageId="2" pageNumber="17">
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<subSubSection pageId="2" pageNumber="17" type="nomenclature">
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<paragraph pageId="2" pageNumber="17">
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<taxonomicName LSID="http://zoobank.org/CAD62F40-4F41-4D91-8E0E-7E9AAAA1632C" authority="Lin & Li" class="Arachnida" family="Pacullidae" genus="Paculla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paculla globosa" order="Araneae" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="globosa">Paculla globosa Lin & Li</taxonomicName>
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<taxonomicNameLabel pageId="2" pageNumber="17">sp. n.</taxonomicNameLabel>
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Figs 4, 5, 6
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="18" type="type material">
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<paragraph pageId="3" pageNumber="18">
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<pageBreakToken pageId="3" pageNumber="18" start="start">Type</pageBreakToken>
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material.
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</paragraph>
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<paragraph pageId="3" pageNumber="18">
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Holotype ♂ (LKCNHM), SINGAPORE: Bukit Timah Nature Reserve, Jungle Fall Stream, altitude 118 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="1.3570555">1°21'25.4"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="103.7737">103°46'25.3"E</geoCoordinate>
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, 21 August 2015, S. Li and Y. Tong leg. Paratype 1♀ (LKCNHM), SINGAPORE: Bukit Timah Nature Reserve, Bukit Timah Summit, altitude 163 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="1.3546389">1°21'16.7"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="103.77639">103°46'35.0"E</geoCoordinate>
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, 19 August 2015, S. Li and Y. Tong leg.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="18" type="etymology">
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<paragraph pageId="3" pageNumber="18">Etymology.</paragraph>
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<paragraph pageId="3" pageNumber="18">
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The specific epithet derives from the Latin word
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<normalizedToken originalValue="“globosus”">"globosus"</normalizedToken>
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= globular, drawing attention to the shape of palpal bulb; adjective.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="18" type="diagnosis">
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<paragraph pageId="3" pageNumber="18">Diagnosis.</paragraph>
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<paragraph pageId="3" pageNumber="18">
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This new species can be distinguished from all congeners with the exception of
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<taxonomicName class="Arachnida" family="Pacullidae" genus="Paculla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paculla sulaimani" order="Araneae" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="sulaimani">Paculla sulaimani</taxonomicName>
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Lehtinen, 1981 by the globular bulb (Fig. 5
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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), the long, finger-like embolus (Fig. 5
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<normalizedToken originalValue="A–D">A-D</normalizedToken>
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) and the presence of three bridge fragments of MV (Fig. 6B). Its male differs from
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<taxonomicName class="Arachnida" family="Pacullidae" genus="Paculla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paculla sulaimani" order="Araneae" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="sulaimani">Paculla sulaimani</taxonomicName>
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(see Lehtinen, 1981: 18, fig. 13) by the narrower embolus with weakly lobed end (Fig. 5A,
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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). The female differs from
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<taxonomicName class="Arachnida" family="Pacullidae" genus="Paculla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paculla bukittimahensis" order="Araneae" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="bukittimahensis">Paculla bukittimahensis</taxonomicName>
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sp. n. (Fig. 3B) by the anterior median part of the atrium protruding broadly and the larger, bridge fragments of MV almost touching each other (Fig. 6B).
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</paragraph>
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<caption pageId="3" pageNumber="18">
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<paragraph pageId="3" pageNumber="18">
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Figure 4.
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<taxonomicName class="Arachnida" family="Pacullidae" genus="Paculla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paculla globosa" order="Araneae" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="globosa">Paculla globosa</taxonomicName>
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sp. n., male holotype (
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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, E) and female paratype (
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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,
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<normalizedToken originalValue="F–G">F-G</normalizedToken>
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).
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<normalizedToken originalValue="A–F">A-F</normalizedToken>
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habitus G genital area (untreated). A, C dorsal B, D, G ventral
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<normalizedToken originalValue="E–F">E-F</normalizedToken>
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lateral. Abbreviations: A = anal plate; AV = anterior ventrolateral plate; L = lateral plate; MV = median ventrolateral plate; P = pulmonary plate; PA = preanal plate; PG = postgenital plate. Scale bars: 0.50 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="19" pageId="3" pageNumber="18" type="description">
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<paragraph pageId="3" pageNumber="18">Description.</paragraph>
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<paragraph pageId="3" pageNumber="18">Male (holotype). Coloration: body dark reddish brown; legs reddish-brown.</paragraph>
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<paragraph pageId="3" pageNumber="18">Measurements: total length 4.40; carapace 1.95 long, 1.40 wide, 1.20 high; abdomen 2.50 long, 1.90 wide, 2.30 high; clypeus 0.49 high; sternum 1.30 long, 1.00 wide. Length of legs: I 8.28 (2.53, 0.65, 2.35, 1.75, 1.00); II 6.95 (2.10, 0.60, 1.85, 1.50, 0.90); III 5.75 (1.70, 0.54, 1.41, 1.30, 0.80); IV 8.16 (2.50, 0.62, 2.30, 1.91, 0.83).</paragraph>
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<paragraph lastPageId="4" lastPageNumber="19" pageId="3" pageNumber="18">
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Prosoma (Fig. 4
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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, E): carapace low, sparsely granulated over whole surface, but surface between the granules smooth; eyes white, ALE=AME=PLE; cephalic part moderately raised; cervical groove distinct; thoracic fovea shallow; clypeus vertical an
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<pageBreakToken pageId="4" pageNumber="19" start="start">teriorly</pageBreakToken>
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; labium triangular, distally obtuse; sternum rough, covered with thin setae, posterior corner protruded. Legs: cuticle striated, weakly granular.
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</paragraph>
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<paragraph pageId="4" pageNumber="19">
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Opisthosoma (Fig. 4
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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, E): dorsal scutum long, oval, covered with thin setae, margin rugose, center modified by sparse pits; rows of small sclerites between lateral scuta, lateral scutum I short; ventral scutum smooth, margin rugose; perigenital scutum broad, triangular.
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</paragraph>
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<paragraph pageId="4" pageNumber="19">
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Palp (Fig. 5
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<normalizedToken originalValue="A–D">A-D</normalizedToken>
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): femoral cuticle smooth, approximately two times as long as patella; patella proximally narrow, distally wide; tibia broad, swollen, 1.3 times as wide as femur; cymbium compressed, slightly shorter than tibia, distally obtuse bifurcate; bulb globular, surface smooth (Fig. 5
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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); embolus long, clavate, starting from subdistal-ventral 1/3 position of bulbous surface (Fig. 5B), proximally sclerotized and bent, distally shallow furcated; embolic tip faint, with split ends (Fig. 5A).
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</paragraph>
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<caption pageId="4" pageNumber="19">
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<paragraph pageId="4" pageNumber="19">
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Figure 5.
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<taxonomicName class="Arachnida" family="Pacullidae" genus="Paculla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paculla globosa" order="Araneae" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="globosa">Paculla globosa</taxonomicName>
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sp. n., male holotype. A,
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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left palp B palpal bulb. A anterior B ventral C prolateral D retrolateral. Abbreviations: ep = embolic part of apes of palpal organ. Scale bars: 0.20 mm.
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</paragraph>
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</caption>
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<paragraph pageId="4" pageNumber="19">Female (paratype). Coloration as in male.</paragraph>
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<paragraph pageId="4" pageNumber="19">Measurements: total length 4.50; carapace 1.85 long, 1.45 wide, 1.20 high; abdomen 2.67 long, 1.80 wide, 2.21 high; clypeus 0.43 high; sternum 1.14 long, 0.97 wide. Length of legs: I 7.35 (2.25, 0.55, 2.10, 1.65, 0.80); II 6.23 (1.95, 0.53, 1.65, 1.35, 0.75); III 5.22 (1.55, 0.50, 1.30, 1.22, 0.65); IV 7.20 (2.15, 0.54, 2.00, 1.80, 0.71). Length of palp: 1.58 (0.45, 0.21, 0.34, 0.38).</paragraph>
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<paragraph pageId="4" pageNumber="19">
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Carapace, abdomen, and legs as in male (Fig. 4
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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, F); clypeus slightly lower than in male.
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</paragraph>
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<paragraph pageId="4" pageNumber="19">
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Genitalia (Figs 4G; 6
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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): epigynal area strongly sclerotized (Fig. 4G); postgenital scutum rugose, wider than preanal scutum; median ventrolateral plate triangular; preanal scutum rugose, wider than long (Fig. 6A). Vulva with a large, nearly rectangular atrium, laterally rugose, and protruding anteromedially; three adjacent, equal sizes bridge fragments of MV below the atrium, and close to the dorso-posterior margin of postgenital scutum (Fig. 6B).
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</paragraph>
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<caption pageId="4" pageNumber="19">
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<paragraph pageId="4" pageNumber="19">
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Figure 6.
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<taxonomicName class="Arachnida" family="Pacullidae" genus="Paculla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paculla globosa" order="Araneae" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="globosa">Paculla globosa</taxonomicName>
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sp. n., female paratype. A genital area (lactic acid-treated), ventral Bditto, dorsal. Abbreviations: AT = atrium; AV = anterior ventrolateral plate; MV = median ventrolateral plate; MVB = bridge fragments of MV; P = pulmonary plate; PA = preanal plate; PG = postgenital plate; PV = posterior ventrolateral plate. Scale bars: 0.50 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="19" type="distribution">
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<paragraph pageId="4" pageNumber="19">Distribution.</paragraph>
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<paragraph pageId="4" pageNumber="19">Singapore.</paragraph>
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</subSubSection>
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</treatment>
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</document>
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