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<document id="29B31B9E8FDFBE83DF6EAAC8E6AC86B5" ID-CLB-Dataset="26196" ID-DOI="10.11646/zootaxa.4638.2.8" ID-GBIF-Dataset="3029fd16-4e07-46e3-89dc-9701f80d7533" ID-ISSN="1175-5326" ID-Zenodo-Dep="3337751" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1563263341028" checkinUser="plazi" docAuthor="Lin, Yucheng" docDate="2019" docId="3C4787C1FFEF471416ADF96D03EB6852" docLanguage="en" docName="zootaxa.4638.2.8.pdf" docOrigin="Zootaxa 4638 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Symphytognatha milleri Lin 2019, new species" docType="treatment" docVersion="9" lastPageNumber="294" masterDocId="C07EFFB9FFEF4717163AFFD5013C6C40" masterDocTitle="First report of the spider genus Symphytognatha from Asia (Araneae, Symphytognathidae)" masterLastPageNumber="295" masterPageNumber="291" pageNumber="291" updateTime="1698745592840" updateUser="ExternalLinkService">
<mods:mods id="DA041E83AFFA4EBDD33464BFCF97CA95" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1F4F17A2ACB8986518CA972C2B7A12A8">
<mods:title id="3C531FBA4271992EE89ED1B819818633">First report of the spider genus Symphytognatha from Asia (Araneae, Symphytognathidae)</mods:title>
</mods:titleInfo>
<mods:name id="851FB4BA023619C130187AB23C84F6DA" type="personal">
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<mods:roleTerm id="53448144B1E90EA93F28A1B8750A710C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0DC07E38C3150E639C2AABE54B1B62C9">Lin, Yucheng</mods:namePart>
</mods:name>
<mods:typeOfResource id="5DBEA0925EFC1E10C4B9F1C707ADE862">text</mods:typeOfResource>
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<mods:title id="A207FEA5AC221D63555D6C2840BCCE21">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9B24F58C3D637FFBD60CA04C25C579A8">
<mods:date id="634C73C50CB4E86A3ACDCB4270584FDF">2019</mods:date>
<mods:detail id="C0217343FA591034DD7140250173F77A" type="pubDate">
<mods:number id="A75C4CABFFE68FE7E37145983AEE0424">2019-07-16</mods:number>
</mods:detail>
<mods:detail id="D3C7B4B64E8C3DEDEE4C85A26555F111" type="volume">
<mods:number id="870B69A97959D71EA746FA3BE81E9D52">4638</mods:number>
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<mods:detail id="18BCB340F44138A832830C1501F129B1" type="issue">
<mods:number id="E4350F95A421053B5AB8153815A58E11">2</mods:number>
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<mods:classification id="3E1DB118B572B2596F0F83E22847A2BB">journal article</mods:classification>
<mods:identifier id="C2AC307F462FCA2A1962ADFD22524E2C" type="CLB-Dataset">26196</mods:identifier>
<mods:identifier id="1C78904B7EFE63C5E25C5A2BFEB3188A" type="DOI">10.11646/zootaxa.4638.2.8</mods:identifier>
<mods:identifier id="13C42769658253E076866A961E9C3F29" type="GBIF-Dataset">3029fd16-4e07-46e3-89dc-9701f80d7533</mods:identifier>
<mods:identifier id="3594F52FD7BDDB0F935C0941F2BACFD2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C535C820687B180CB1BBB7118EFFFF74" type="Zenodo-Dep">3337751</mods:identifier>
</mods:mods>
<treatment id="3C4787C1FFEF471416ADF96D03EB6852" ID-DOI="http://doi.org/10.5281/zenodo.5923797" ID-GBIF-Taxon="158691035" ID-Zenodo-Dep="5923797" LSID="urn:lsid:plazi:treatment:3C4787C1FFEF471416ADF96D03EB6852" httpUri="http://treatment.plazi.org/id/3C4787C1FFEF471416ADF96D03EB6852" lastPageId="3" lastPageNumber="294" pageId="0" pageNumber="291">
<subSubSection id="FCF4655CFFEF471716ADF96D03E86A92" box="[151,724,1720,1747]" pageId="0" pageNumber="291" type="nomenclature">
<paragraph id="B45136D7FFEF471716ADF96D03E86A92" blockId="0.[151,1436,1720,2034]" box="[151,724,1720,1747]" pageId="0" pageNumber="291">
<heading id="EF1981BBFFEF471716ADF96D03E86A92" bold="true" box="[151,724,1720,1747]" fontSize="11" level="1" pageId="0" pageNumber="291" reason="1">
<emphasis id="869AEAC5FFEF471716ADF96D03E86A92" bold="true" box="[151,724,1720,1747]" pageId="0" pageNumber="291">
<taxonomicName id="73EE4D54FFEF471716ADF96D00866A93" ID-CoL="98ZYX" authority="Lin, 2019" authorityName="Lin" authorityYear="2019" box="[151,442,1720,1747]" class="Arachnida" family="Symphytognathidae" genus="Symphytognatha" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="291" phylum="Arthropoda" rank="species" species="milleri" status="sp. nov.">
<emphasis id="869AEAC5FFEF471716ADF96D00866A93" bold="true" box="[151,442,1720,1747]" italics="true" pageId="0" pageNumber="291">Symphytognatha milleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9DA957BEFFEF471717FBF96C036D6A92" box="[449,593,1720,1747]" pageId="0" pageNumber="291" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="2CD52A52FFEF4717145BF96D03F76A92" box="[609,715,1720,1746]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,244,1984,2006]" captionStartId-1="2.[151,244,1930,1952]" captionStartId-2="3.[151,244,1634,1656]" captionTargetBox-0="[189,1398,185,1951]" captionTargetBox-1="[201,1386,167,1904]" captionTargetBox-2="[151,1435,1065,1614]" captionTargetId-0="figure@1.[189,1398,185,1964]" captionTargetId-1="figure@2.[201,1386,167,1910]" captionTargetId-2="figure@3.[151,1435,1065,1614]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Symphytognatha milleri n. sp., female holotype (A–C, G) and subadult male paratype (D–F, H). A–F, Habitus. G–H, Prosoma. A, D, Dorsal. B, E, Ventral. C, F, Lateral. G–H, Frontal. Scale bars: 0.50 (A–F); 0.20 (G–H)." captionText-1="FIGURE 2. Symphytognatha milleri n. sp., female holotype. A, Habitus (omitted left legs). B–C, Chelicerae. D–E, Epigynum. F–G, Vulva (lactic acid treated). A, Lateral. B, D, Posterior. C, Anterior. E, F, Ventral. G, Dorsal. Abbreviations: cd, copulatory duct; fd, fertilization duct; sp, spermatheca; vs, vulval septum. Scale bars: 0.20 (A, D–E); 0.10 (B–C, F–G)." captionText-2="FIGURE 3. Symphytognatha milleri n. sp., legs’ claws of female holotype.A, Multidentate claw on leg I. B, Multidentate claw on leg II. C, edentate claw on leg III. D, edentate claw on leg IV. A, B, Retrolateral. C, D, Prolateral. Scale bar: 0.05." httpUri-0="https://zenodo.org/record/3337753/files/figure.png" httpUri-1="https://zenodo.org/record/3337755/files/figure.png" httpUri-2="https://zenodo.org/record/3337757/files/figure.png" pageId="0" pageNumber="291">Figs 1–3</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FCF4655CFFEF471716ADF90B030A6B22" pageId="0" pageNumber="291" type="materials_examined">
<paragraph id="B45136D7FFEF471716ADF90B030A6B22" blockId="0.[151,1436,1720,2034]" pageId="0" pageNumber="291">
<emphasis id="869AEAC5FFEF471716ADF90B007D6AB6" bold="true" box="[151,321,1758,1782]" pageId="0" pageNumber="291">Type material.</emphasis>
<materialsCitation id="04863C8AFFEF47171771F90B030A6B22" ID-GBIF-Occurrence="2283872132" pageId="0" pageNumber="291" typeStatus="holotype">
<typeStatus id="6B558875FFEF47171771F90B00916AB6" box="[331,429,1758,1782]" pageId="0" pageNumber="291" type="holotype">
<emphasis id="869AEAC5FFEF47171771F90B00916AB6" box="[331,429,1758,1782]" italics="true" pageId="0" pageNumber="291">Holotype</emphasis>
</typeStatus>
<specimenCount id="A2E8FD5EFFEF4717178DF90B00F76AB5" box="[439,459,1758,1781]" pageId="0" pageNumber="291" type="female">♀</specimenCount>
and
<typeStatus id="6B558875FFEF4717143DF90A035A6AB6" box="[519,614,1759,1782]" pageId="0" pageNumber="291" type="paratype">
<emphasis id="869AEAC5FFEF4717143DF90A035A6AB6" box="[519,614,1759,1782]" italics="true" pageId="0" pageNumber="291">paratype</emphasis>
</typeStatus>
<specimenCount id="A2E8FD5EFFEF4717144AF90B03AE6AB6" box="[624,658,1758,1782]" pageId="0" pageNumber="291" type="male">1♂</specimenCount>
(subadult) (
<collectionCode id="D2FFAE12FFEF47171520F90B02BC6AB6" box="[794,896,1758,1782]" pageId="0" pageNumber="291">NHMSU</collectionCode>
),
<collectingCountry id="CCF97647FFEF471715ACF90B02D36AB6" box="[918,1007,1758,1782]" name="China" pageId="0" pageNumber="291">CHINA</collectingCountry>
:
<collectingRegion id="762AF835FFEF471715C6F90B05866AB6" box="[1020,1210,1758,1782]" country="China" name="Yunnan" pageId="0" pageNumber="291">Yunnan Province</collectingRegion>
,
<collectingMunicipality id="5435ACADFFEF471712F2F90B04A46AB6" box="[1224,1432,1758,1782]" pageId="0" pageNumber="291">Nujiang Prefecture</collectingMunicipality>
,
<collectingCounty id="5D304E5BFFEF471716ADF8D700606B5A" box="[151,348,1794,1818]" pageId="0" pageNumber="291">Gongshan County</collectingCounty>
,
<location id="B131600CFFEF47171752F8D7021D6B59" LSID="urn:lsid:plazi:treatment:3C4787C1FFEF471416ADF96D03EB6852:B131600CFFEF47171752F8D7021D6B59" box="[360,801,1794,1818]" country="China" county="Gongshan County" latitude="27.67934" longLatPrecision="1" longitude="98.27291" municipality="Nujiang Prefecture" name="Gaoligongshan National Nature Reserve" pageId="0" pageNumber="291" stateProvince="Yunnan">Gaoligongshan National Nature Reserve</location>
,
<location id="B131600CFFEF47171514F8D702C66B59" LSID="urn:lsid:plazi:treatment:3C4787C1FFEF471416ADF96D03EB6852:B131600CFFEF47171514F8D702C66B59" box="[814,1018,1794,1818]" country="China" county="Gongshan County" latitude="27.67934" longLatPrecision="1" longitude="98.27291" municipality="Nujiang Prefecture" name="Dulongjiang Town" pageId="0" pageNumber="291" stateProvince="Yunnan">Dulongjiang Town</location>
,
<location id="B131600CFFEF4717123DF8D705D56B5A" LSID="urn:lsid:plazi:treatment:3C4787C1FFEF471416ADF96D03EB6852:B131600CFFEF4717123DF8D705D56B5A" box="[1031,1257,1794,1818]" country="China" county="Gongshan County" latitude="27.67934" longLatPrecision="1" longitude="98.27291" municipality="Nujiang Prefecture" name="Qinlangdang Village" pageId="0" pageNumber="291" stateProvince="Yunnan">Qinlangdang Village</location>
, near to
<location id="B131600CFFEF47171373F8D701C06B7E" LSID="urn:lsid:plazi:treatment:3C4787C1FFEF471416ADF96D03EB6852:B131600CFFEF47171373F8D701C06B7E" country="China" county="Gongshan County" latitude="27.67934" longLatPrecision="1" longitude="98.27291" municipality="Nujiang Prefecture" name="Habang Waterfall" pageId="0" pageNumber="291" stateProvince="Yunnan">Habang Waterfall</location>
, primary subtropical broadleaf deciduous forest, in leaf litter,
<geoCoordinate id="D1DA5010FFEF471715BDF8F305306B7E" box="[903,1036,1830,1854]" degrees="27.67934" direction="north" orientation="latitude" pageId="0" pageNumber="291" precision="1" value="27.67934">27.67934°N</geoCoordinate>
,
<geoCoordinate id="D1DA5010FFEF4717122FF8F305996B7E" box="[1045,1189,1830,1854]" degrees="098.27291" direction="east" orientation="longitude" pageId="0" pageNumber="291" precision="1" value="98.27291">098.27291°E</geoCoordinate>
;
<elevation id="3FC3D1E4FFEF47171294F8F3043C6B7D" box="[1198,1280,1830,1854]" metricMagnitude="3" metricUnit="m" metricValue="1.205" pageId="0" pageNumber="291" unit="m" value="1205.0">
<quantity id="73169B32FFEF47171294F8F3043C6B7D" box="[1198,1280,1830,1854]" metricMagnitude="3" metricUnit="m" metricValue="1.205" pageId="0" pageNumber="291" unit="m" value="1205.0">1205 m</quantity>
</elevation>
,
<collectingDate id="D014E9FFFFEF47171330F8F304A46B7E" box="[1290,1432,1830,1854]" pageId="0" pageNumber="291" value="2018-08-14">
<date id="C0501017FFEF47171330F8F304A46B7E" box="[1290,1432,1830,1854]" pageId="0" pageNumber="291" value="2018-08-14">14.VIII.2018</date>
</collectingDate>
,
<collectorName id="191B5301FFEF471716ADF89F01DB6B22" box="[151,231,1866,1890]" pageId="0" pageNumber="291">Y.C. Li</collectorName>
,
<collectorName id="191B5301FFEF471716CBF89F00146B22" box="[241,296,1866,1890]" pageId="0" pageNumber="291">Y. Li</collectorName>
,
<collectorName id="191B5301FFEF47171708F89F00B76B22" box="[306,395,1866,1890]" pageId="0" pageNumber="291">Y.F. Shu</collectorName>
&amp;
<collectorName id="191B5301FFEF47171797F89F03346B22" box="[429,520,1866,1890]" pageId="0" pageNumber="291">Y.C. Lin</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="FCF4655CFFEF471716FDF8BB02FA6BEA" pageId="0" pageNumber="291" type="etymology">
<paragraph id="B45136D7FFEF471716FDF8BB02FA6BEA" blockId="0.[151,1436,1720,2034]" pageId="0" pageNumber="291">
<emphasis id="869AEAC5FFEF471716FDF8BB00746BC6" bold="true" box="[199,328,1902,1926]" pageId="0" pageNumber="291">Etymology.</emphasis>
Patronymic in honor of Dr. Jeremy A. Miller (Naturalis Biodiversity Center,
<collectingCountry id="CCF97647FFEF47171249F8BB05C46BC6" box="[1139,1272,1902,1926]" name="Netherlands" pageId="0" pageNumber="291">Netherlands</collectingCountry>
), in recognition of his outstanding efforts in the study of Chinese symphytognathoid spiders.
</paragraph>
</subSubSection>
<subSubSection id="FCF4655CFFEF471416FDF86302806D26" lastPageId="3" lastPageNumber="294" pageId="0" pageNumber="291" type="diagnosis">
<paragraph id="B45136D7FFEF471416FDF86302806D26" blockId="0.[151,1436,1720,2034]" lastBlockId="3.[151,1437,154,1042]" lastPageId="3" lastPageNumber="294" pageId="0" pageNumber="291">
<emphasis id="869AEAC5FFEF471716FDF86300006B8E" bold="true" box="[199,316,1974,1998]" pageId="0" pageNumber="291">Diagnosis.</emphasis>
With the exception of
<taxonomicName id="73EE4D54FFEF47171401F86302626B8E" authority="Hickman, 1931" authorityName="Hickman" authorityYear="1931" box="[571,862,1974,1998]" class="Arachnida" family="Symphytognathidae" genus="Symphytognatha" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="291" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="869AEAC5FFEF47171401F86303976B8E" box="[571,683,1974,1998]" italics="true" pageId="0" pageNumber="291">S. globosa</emphasis>
<bibRefCitation id="D07F4B26FFEF4717148FF86302626B8E" author="Hickman, V. V." box="[693,862,1974,1998]" pageId="0" pageNumber="291" pagination="1321 - 1328" refId="ref2000" refString="Hickman, V. V. (1931) A new family of spiders. Proceedings of the Zoological Society of London (B), 1931, 1321 - 1328. https: // doi. org / 10.1111 / j. 1096 - 3642.1931. tb 01063. x" type="journal article" year="1931">Hickman, 1931</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="73EE4D54FFEF47171551F86305E46B8E" authority="Forster &amp; Platnick, 1977" authorityName="Forster &amp; Platnick" authorityYear="1977" box="[875,1240,1974,1998]" class="Arachnida" family="Symphytognathidae" genus="Symphytognatha" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="291" phylum="Arthropoda" rank="species" species="blesti">
<emphasis id="869AEAC5FFEF47171551F86302FE6B8E" box="[875,962,1974,1998]" italics="true" pageId="0" pageNumber="291">S. blesti</emphasis>
<bibRefCitation id="D07F4B26FFEF471715F0F86305E46B8E" author="Forster, R. R. &amp; Platnick, N. I." box="[970,1240,1974,1998]" pageId="0" pageNumber="291" pagination="1 - 29" refId="ref1809" refString="Forster, R. R. &amp; Platnick, N. I. (1977) A review of the spider family Symphytognathidae (Arachnida, Araneae). American Museum Novitates, 2619, 1 - 29." type="journal article" year="1977">Forster &amp; Platnick, 1977</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="73EE4D54FFEF4717132BF86300386BB2" authority="Harvey, 1992" authorityName="Harvey" authorityYear="1992" class="Arachnida" family="Symphytognathidae" genus="Symphytognatha" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="291" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="869AEAC5FFEF4717132BF863045F6B8E" box="[1297,1379,1974,1998]" italics="true" pageId="0" pageNumber="291">S. picta</emphasis>
<bibRefCitation id="D07F4B26FFEF47171356F86300386BB2" author="Harvey, M. S." pageId="0" pageNumber="291" pagination="685 - 689" refId="ref1926" refString="Harvey, M. S. (1992) A new species of Symphytognatha Hickman (Araneae: Symphytognathidae) from Western Australia. Records of the Western Australian Museum, 15, 685 - 689." type="journal article" year="1992">Harvey, 1992</bibRefCitation>
</taxonomicName>
, the new species can be distinguished from other congeners by the presence of a vulval septum and coiled spermathecae (
<figureCitation id="2CD52A52FFEC47141702FF4F00B76CF2" box="[312,395,154,178]" captionStart="FIGURE 2" captionStartId="2.[151,244,1930,1952]" captionTargetBox="[201,1386,167,1904]" captionTargetId="figure@2.[201,1386,167,1910]" captionTargetPageId="2" captionText="FIGURE 2. Symphytognatha milleri n. sp., female holotype. A, Habitus (omitted left legs). B–C, Chelicerae. D–E, Epigynum. F–G, Vulva (lactic acid treated). A, Lateral. B, D, Posterior. C, Anterior. E, F, Ventral. G, Dorsal. Abbreviations: cd, copulatory duct; fd, fertilization duct; sp, spermatheca; vs, vulval septum. Scale bars: 0.20 (A, D–E); 0.10 (B–C, F–G)." httpUri="https://zenodo.org/record/3337755/files/figure.png" pageId="3" pageNumber="294">Fig. 2G</figureCitation>
). It differs from
<taxonomicName id="73EE4D54FFEC47141479FF4F038F6CF2" authorityName="Hickman" authorityYear="1931" box="[579,691,154,178]" class="Arachnida" family="Symphytognathidae" genus="Symphytognatha" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="294" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="869AEAC5FFEC47141479FF4F038F6CF2" box="[579,691,154,178]" italics="true" pageId="3" pageNumber="294">S. globosa</emphasis>
</taxonomicName>
and
<taxonomicName id="73EE4D54FFEC471414D3FF4F02036CF2" authorityName="Forster &amp; Platnick" authorityYear="1977" box="[745,831,154,178]" class="Arachnida" family="Symphytognathidae" genus="Symphytognatha" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="294" phylum="Arthropoda" rank="species" species="blesti">
<emphasis id="869AEAC5FFEC471414D3FF4F02036CF2" box="[745,831,154,178]" italics="true" pageId="3" pageNumber="294">S. blesti</emphasis>
</taxonomicName>
by a fingerlike vulval septum (needlelike in
<taxonomicName id="73EE4D54FFEC4714131FFF4F04A96CF2" authorityName="Hickman" authorityYear="1931" box="[1317,1429,154,178]" class="Arachnida" family="Symphytognathidae" genus="Symphytognatha" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="294" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="869AEAC5FFEC4714131FFF4F04A96CF2" box="[1317,1429,154,178]" italics="true" pageId="3" pageNumber="294">S. globosa</emphasis>
</taxonomicName>
, absent in
<taxonomicName id="73EE4D54FFEC47141738FF6B00656C96" authorityName="Forster &amp; Platnick" authorityYear="1977" box="[258,345,190,214]" class="Arachnida" family="Symphytognathidae" genus="Symphytognatha" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="294" phylum="Arthropoda" rank="species" species="blesti">
<emphasis id="869AEAC5FFEC47141738FF6B00656C96" box="[258,345,190,214]" italics="true" pageId="3" pageNumber="294">S. blesti</emphasis>
</taxonomicName>
), the wider spermatheca and the copulatory duct starts far from the base of vulval septum (starts at the base of the vulval septum in
<taxonomicName id="73EE4D54FFEC471417C7FF3703526CBA" authorityName="Hickman" authorityYear="1931" box="[509,622,226,250]" class="Arachnida" family="Symphytognathidae" genus="Symphytognatha" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="294" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="869AEAC5FFEC471417C7FF3703526CBA" box="[509,622,226,250]" italics="true" pageId="3" pageNumber="294">S. globosa</emphasis>
</taxonomicName>
, starts in the posteromarginal center of epigynum in
<taxonomicName id="73EE4D54FFEC47141288FF3704346CBA" authorityName="Forster &amp; Platnick" authorityYear="1977" box="[1202,1288,226,250]" class="Arachnida" family="Symphytognathidae" genus="Symphytognatha" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="294" phylum="Arthropoda" rank="species" species="blesti">
<emphasis id="869AEAC5FFEC47141288FF3704346CBA" box="[1202,1288,226,250]" italics="true" pageId="3" pageNumber="294">S. blesti</emphasis>
</taxonomicName>
) (
<figureCitation id="2CD52A52FFEC4714131BFF3704496CBA" box="[1313,1397,226,250]" captionStart="FIGURE 2" captionStartId="2.[151,244,1930,1952]" captionTargetBox="[201,1386,167,1904]" captionTargetId="figure@2.[201,1386,167,1910]" captionTargetPageId="2" captionText="FIGURE 2. Symphytognatha milleri n. sp., female holotype. A, Habitus (omitted left legs). B–C, Chelicerae. D–E, Epigynum. F–G, Vulva (lactic acid treated). A, Lateral. B, D, Posterior. C, Anterior. E, F, Ventral. G, Dorsal. Abbreviations: cd, copulatory duct; fd, fertilization duct; sp, spermatheca; vs, vulval septum. Scale bars: 0.20 (A, D–E); 0.10 (B–C, F–G)." httpUri="https://zenodo.org/record/3337755/files/figure.png" pageId="3" pageNumber="294">Fig. 2G</figureCitation>
vs.
<bibRefCitation id="D07F4B26FFEC471416ADFED300036D5E" author="Hickman, V. V." box="[151,319,262,286]" pageId="3" pageNumber="294" pagination="1321 - 1328" refId="ref2000" refString="Hickman, V. V. (1931) A new family of spiders. Proceedings of the Zoological Society of London (B), 1931, 1321 - 1328. https: // doi. org / 10.1111 / j. 1096 - 3642.1931. tb 01063. x" type="journal article" year="1931">Hickman, 1931</bibRefCitation>
: fig. 6, plate 2;
<bibRefCitation id="D07F4B26FFEC471417DFFED303CD6D5E" author="Forster, R. R. &amp; Platnick, N. I." box="[485,753,262,286]" pageId="3" pageNumber="294" pagination="1 - 29" refId="ref1809" refString="Forster, R. R. &amp; Platnick, N. I. (1977) A review of the spider family Symphytognathidae (Arachnida, Araneae). American Museum Novitates, 2619, 1 - 29." type="journal article" year="1977">Forster &amp; Platnick, 1977</bibRefCitation>
: figs 12, 13). Differs from
<taxonomicName id="73EE4D54FFEC47141235FED305636D5D" authorityName="Harvey" authorityYear="1992" box="[1039,1119,262,285]" class="Arachnida" family="Symphytognathidae" genus="Symphytognatha" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="294" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="869AEAC5FFEC47141235FED305636D5D" box="[1039,1119,262,285]" italics="true" pageId="3" pageNumber="294">S. picta</emphasis>
</taxonomicName>
by the long, fingerlike vulval septum (short, nearly triangular in
<taxonomicName id="73EE4D54FFEC47141434FEFF03626D01" authorityName="Harvey" authorityYear="1992" box="[526,606,298,321]" class="Arachnida" family="Symphytognathidae" genus="Symphytognatha" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="294" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="869AEAC5FFEC47141434FEFF03626D01" box="[526,606,298,321]" italics="true" pageId="3" pageNumber="294">S. picta</emphasis>
</taxonomicName>
), and the copulatory ducts starts far from the base of vulval septum (starts at the base of the vulval septum in
<taxonomicName id="73EE4D54FFEC471417CBFE9B037D6D25" authorityName="Harvey" authorityYear="1992" box="[497,577,334,357]" class="Arachnida" family="Symphytognathidae" genus="Symphytognatha" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="294" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="869AEAC5FFEC471417CBFE9B037D6D25" box="[497,577,334,357]" italics="true" pageId="3" pageNumber="294">S. picta</emphasis>
</taxonomicName>
) (
<figureCitation id="2CD52A52FFEC47141463FE9B03976D26" box="[601,683,334,358]" captionStart="FIGURE 2" captionStartId="2.[151,244,1930,1952]" captionTargetBox="[201,1386,167,1904]" captionTargetId="figure@2.[201,1386,167,1910]" captionTargetPageId="2" captionText="FIGURE 2. Symphytognatha milleri n. sp., female holotype. A, Habitus (omitted left legs). B–C, Chelicerae. D–E, Epigynum. F–G, Vulva (lactic acid treated). A, Lateral. B, D, Posterior. C, Anterior. E, F, Ventral. G, Dorsal. Abbreviations: cd, copulatory duct; fd, fertilization duct; sp, spermatheca; vs, vulval septum. Scale bars: 0.20 (A, D–E); 0.10 (B–C, F–G)." httpUri="https://zenodo.org/record/3337755/files/figure.png" pageId="3" pageNumber="294">Fig. 2G</figureCitation>
vs.
<bibRefCitation id="D07F4B26FFEC471414EDFE9B02566D26" author="Harvey, M. S." box="[727,874,334,358]" pageId="3" pageNumber="294" pagination="685 - 689" refId="ref1926" refString="Harvey, M. S. (1992) A new species of Symphytognatha Hickman (Araneae: Symphytognathidae) from Western Australia. Records of the Western Australian Museum, 15, 685 - 689." type="journal article" year="1992">Harvey, 1992</bibRefCitation>
: fig. 3).
</paragraph>
</subSubSection>
<caption id="E091665FFFEE471616ADF81505706BBA" httpUri="https://zenodo.org/record/3337753/files/figure.png" pageId="1" pageNumber="292" startId="1.[151,244,1984,2006]" targetBox="[189,1398,185,1951]" targetPageId="1">
<paragraph id="B45136D7FFEE471616ADF81505706BBA" blockId="1.[151,1436,1984,2043]" pageId="1" pageNumber="292">
<emphasis id="869AEAC5FFEE471616ADF81500316B96" bold="true" box="[151,269,1984,2006]" pageId="1" pageNumber="292">FIGURE 1.</emphasis>
<taxonomicName id="73EE4D54FFEE47161728F81400C46B96" authority="Lin, 2019" authorityName="Lin" authorityYear="2019" box="[274,504,1985,2006]" class="Arachnida" family="Symphytognathidae" genus="Symphytognatha" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="292" phylum="Arthropoda" rank="species" species="milleri" status="sp. nov.">
<emphasis id="869AEAC5FFEE47161728F81400C46B96" box="[274,504,1985,2006]" italics="true" pageId="1" pageNumber="292">Symphytognatha milleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9DA957BEFFEE471617C4F81403086B96" box="[510,564,1985,2006]" pageId="1" pageNumber="292" rank="species">
<emphasis id="869AEAC5FFEE471617C4F81403086B96" bold="true" box="[510,564,1985,2006]" pageId="1" pageNumber="292">n. sp.</emphasis>
</taxonomicNameLabel>
, female holotype (A–C, G) and subadult male paratype (D–F, H). A–F, Habitus. G–H, Prosoma. A, D, Dorsal. B, E, Ventral. C, F, Lateral. G–H, Frontal. Scale bars: 0.50 (A–F); 0.20 (G–H).
</paragraph>
</caption>
<caption id="E091665FFFED471516ADF85F05776BA8" httpUri="https://zenodo.org/record/3337755/files/figure.png" pageId="2" pageNumber="293" startId="2.[151,244,1930,1952]" targetBox="[201,1386,167,1904]" targetPageId="2">
<paragraph id="B45136D7FFED471516ADF85F05776BA8" blockId="2.[151,1437,1930,2024]" pageId="2" pageNumber="293">
<emphasis id="869AEAC5FFED471516ADF85F00316BDF" bold="true" box="[151,269,1930,1952]" pageId="2" pageNumber="293">FIGURE 2.</emphasis>
<taxonomicName id="73EE4D54FFED4715172EF85F00C76BDF" authority="Lin, 2019" authorityName="Lin" authorityYear="2019" box="[276,507,1930,1951]" class="Arachnida" family="Symphytognathidae" genus="Symphytognatha" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="293" phylum="Arthropoda" rank="species" species="milleri" status="sp. nov.">
<emphasis id="869AEAC5FFED4715172EF85F00C76BDF" box="[276,507,1930,1951]" italics="true" pageId="2" pageNumber="293">Symphytognatha milleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9DA957BEFFED4715143BF85E03056BE0" box="[513,569,1931,1952]" pageId="2" pageNumber="293" rank="species">
<emphasis id="869AEAC5FFED4715143BF85E03056BE0" bold="true" box="[513,569,1931,1952]" pageId="2" pageNumber="293">n. sp.</emphasis>
</taxonomicNameLabel>
, female holotype. A, Habitus (omitted left legs). B–C, Chelicerae. D–E, Epigynum. F–G, Vulva (lactic acid treated). A, Lateral. B, D, Posterior. C, Anterior. E, F, Ventral. G, Dorsal. Abbreviations: cd, copulatory duct; fd, fertilization duct; sp, spermatheca; vs, vulval septum. Scale bars: 0.20 (A, D–E); 0.10 (B–C, F–G).
</paragraph>
</caption>
<subSubSection id="FCF4655CFFEC471416FDFEA7045C6FAE" pageId="3" pageNumber="294" type="description">
<paragraph id="B45136D7FFEC471416FDFEA701F86F7A" blockId="3.[151,1437,154,1042]" pageId="3" pageNumber="294">
<emphasis id="869AEAC5FFEC471416FDFEA700926DCA" bold="true" box="[199,430,370,394]" pageId="3" pageNumber="294">Description. Female</emphasis>
(
<typeStatus id="6B558875FFEC47141785FEA7031B6DCA" box="[447,551,370,394]" pageId="3" pageNumber="294" type="holotype">Holotype</typeStatus>
).
<emphasis id="869AEAC5FFEC47141401FEA603EA6DCA" box="[571,726,371,394]" italics="true" pageId="3" pageNumber="294">Measurements</emphasis>
: Total length, not including chelicerae 0.84. Carapace 0.32 long, 0.30 wide, 0.28 high. Clypeus 0.08 high. Sternum 0.25 long, 0.21 wide. Abdomen 0.67 long, 0.62 wide, 0.68 high. Length of legs: I 0.74 (0.23, 0.09, 0.16, 0.11, 0.15); II 0.63 (0.20, 0.07, 0.13, 0.11, 0.12); III 0.53 (0.15, 0.07, 0.10, 0.08, 0.13); IV 0.91 (0.29, 0.10, 0.20, 0.15, 0.17). Leg formula IV-I-II-III. Somatic characters see
<figureCitation id="2CD52A52FFEC4714127FFE0B05BC6DB6" box="[1093,1152,478,502]" captionStart="FIGURE 1" captionStartId="1.[151,244,1984,2006]" captionTargetBox="[189,1398,185,1951]" captionTargetId="figure@1.[189,1398,185,1964]" captionTargetPageId="1" captionText="FIGURE 1. Symphytognatha milleri n. sp., female holotype (A–C, G) and subadult male paratype (D–F, H). A–F, Habitus. G–H, Prosoma. A, D, Dorsal. B, E, Ventral. C, F, Lateral. G–H, Frontal. Scale bars: 0.50 (A–F); 0.20 (G–H)." httpUri="https://zenodo.org/record/3337753/files/figure.png" pageId="3" pageNumber="294">Fig. 1</figureCitation>
D–F, H. Coloration: body chestnut red, but light color spots on sternum and abdomen, legs pale yellow. Cephalic pars strongly elevated. Thoracic pars sloping abruptly. Six eyes subequal in size, and in three diads, PME separated, lateral eyes contiguous, PLE-PME separated by ca. 1.3x PME diameter. Clypeus ca. 1.2x PME, concave. Chelicerae bright brown, projected forward, suture visible (
<figureCitation id="2CD52A52FFEC471416D4FDBB007A6EC5" box="[238,326,622,646]" captionStart="FIGURE 1" captionStartId="1.[151,244,1984,2006]" captionTargetBox="[189,1398,185,1951]" captionTargetId="figure@1.[189,1398,185,1964]" captionTargetPageId="1" captionText="FIGURE 1. Symphytognatha milleri n. sp., female holotype (A–C, G) and subadult male paratype (D–F, H). A–F, Habitus. G–H, Prosoma. A, D, Dorsal. B, E, Ventral. C, F, Lateral. G–H, Frontal. Scale bars: 0.50 (A–F); 0.20 (G–H)." httpUri="https://zenodo.org/record/3337753/files/figure.png" pageId="3" pageNumber="294">Figs 1H</figureCitation>
,
<figureCitation id="2CD52A52FFEC4714176FFDBB00446EC6" box="[341,376,622,646]" captionStart="FIGURE 2" captionStartId="2.[151,244,1930,1952]" captionTargetBox="[201,1386,167,1904]" captionTargetId="figure@2.[201,1386,167,1910]" captionTargetPageId="2" captionText="FIGURE 2. Symphytognatha milleri n. sp., female holotype. A, Habitus (omitted left legs). B–C, Chelicerae. D–E, Epigynum. F–G, Vulva (lactic acid treated). A, Lateral. B, D, Posterior. C, Anterior. E, F, Ventral. G, Dorsal. Abbreviations: cd, copulatory duct; fd, fertilization duct; sp, spermatheca; vs, vulval septum. Scale bars: 0.20 (A, D–E); 0.10 (B–C, F–G)." httpUri="https://zenodo.org/record/3337755/files/figure.png" pageId="3" pageNumber="294">2C</figureCitation>
), with two paired long, needle teeth on promargin (
<figureCitation id="2CD52A52FFEC47141593FDBB051C6EC6" box="[937,1056,622,646]" captionStart="FIGURE 2" captionStartId="2.[151,244,1930,1952]" captionTargetBox="[201,1386,167,1904]" captionTargetId="figure@2.[201,1386,167,1910]" captionTargetPageId="2" captionText="FIGURE 2. Symphytognatha milleri n. sp., female holotype. A, Habitus (omitted left legs). B–C, Chelicerae. D–E, Epigynum. F–G, Vulva (lactic acid treated). A, Lateral. B, D, Posterior. C, Anterior. E, F, Ventral. G, Dorsal. Abbreviations: cd, copulatory duct; fd, fertilization duct; sp, spermatheca; vs, vulval septum. Scale bars: 0.20 (A, D–E); 0.10 (B–C, F–G)." httpUri="https://zenodo.org/record/3337755/files/figure.png" pageId="3" pageNumber="294">Fig. 2B, C</figureCitation>
). Endites lack palp. Labium nearly rectangular, fused to sternum (
<figureCitation id="2CD52A52FFEC471417DAFD47030E6EEA" box="[480,562,658,682]" captionStart="FIGURE 1" captionStartId="1.[151,244,1984,2006]" captionTargetBox="[189,1398,185,1951]" captionTargetId="figure@1.[189,1398,185,1964]" captionTargetPageId="1" captionText="FIGURE 1. Symphytognatha milleri n. sp., female holotype (A–C, G) and subadult male paratype (D–F, H). A–F, Habitus. G–H, Prosoma. A, D, Dorsal. B, E, Ventral. C, F, Lateral. G–H, Frontal. Scale bars: 0.50 (A–F); 0.20 (G–H)." httpUri="https://zenodo.org/record/3337753/files/figure.png" pageId="3" pageNumber="294">Fig. 1E</figureCitation>
). Sternum slightly plump, anterior narrow and posterior wide, surface bears long setae, posteriorly truncated (
<figureCitation id="2CD52A52FFEC471417FEFD63030F6E8D" box="[452,563,694,718]" captionStart="FIGURE 1" captionStartId="1.[151,244,1984,2006]" captionTargetBox="[189,1398,185,1951]" captionTargetId="figure@1.[189,1398,185,1964]" captionTargetPageId="1" captionText="FIGURE 1. Symphytognatha milleri n. sp., female holotype (A–C, G) and subadult male paratype (D–F, H). A–F, Habitus. G–H, Prosoma. A, D, Dorsal. B, E, Ventral. C, F, Lateral. G–H, Frontal. Scale bars: 0.50 (A–F); 0.20 (G–H)." httpUri="https://zenodo.org/record/3337753/files/figure.png" pageId="3" pageNumber="294">Figs 1E, F</figureCitation>
,
<figureCitation id="2CD52A52FFEC47141407FD63035E6E8E" box="[573,610,694,718]" captionStart="FIGURE 2" captionStartId="2.[151,244,1930,1952]" captionTargetBox="[201,1386,167,1904]" captionTargetId="figure@2.[201,1386,167,1910]" captionTargetPageId="2" captionText="FIGURE 2. Symphytognatha milleri n. sp., female holotype. A, Habitus (omitted left legs). B–C, Chelicerae. D–E, Epigynum. F–G, Vulva (lactic acid treated). A, Lateral. B, D, Posterior. C, Anterior. E, F, Ventral. G, Dorsal. Abbreviations: cd, copulatory duct; fd, fertilization duct; sp, spermatheca; vs, vulval septum. Scale bars: 0.20 (A, D–E); 0.10 (B–C, F–G)." httpUri="https://zenodo.org/record/3337755/files/figure.png" pageId="3" pageNumber="294">2D</figureCitation>
). Multidentate superior claws on legs I and II, but edental claws on leg III and IV (
<figureCitation id="2CD52A52FFEC471416FBFD0F01C36EB2" box="[193,255,730,754]" captionStart="FIGURE 3" captionStartId="3.[151,244,1634,1656]" captionTargetBox="[151,1435,1065,1614]" captionTargetId="figure@3.[151,1435,1065,1614]" captionTargetPageId="3" captionText="FIGURE 3. Symphytognatha milleri n. sp., legs’ claws of female holotype.A, Multidentate claw on leg I. B, Multidentate claw on leg II. C, edentate claw on leg III. D, edentate claw on leg IV. A, B, Retrolateral. C, D, Prolateral. Scale bar: 0.05." httpUri="https://zenodo.org/record/3337757/files/figure.png" pageId="3" pageNumber="294">Fig. 3</figureCitation>
A–D). Abdomen nearly globular dorsally, covers with long setae, dorsally and ventrally with indistinct fuscous pattern (
<figureCitation id="2CD52A52FFEC471416CAFD2B00166F56" box="[240,298,766,790]" captionStart="FIGURE 1" captionStartId="1.[151,244,1984,2006]" captionTargetBox="[189,1398,185,1951]" captionTargetId="figure@1.[189,1398,185,1964]" captionTargetPageId="1" captionText="FIGURE 1. Symphytognatha milleri n. sp., female holotype (A–C, G) and subadult male paratype (D–F, H). A–F, Habitus. G–H, Prosoma. A, D, Dorsal. B, E, Ventral. C, F, Lateral. G–H, Frontal. Scale bars: 0.50 (A–F); 0.20 (G–H)." httpUri="https://zenodo.org/record/3337753/files/figure.png" pageId="3" pageNumber="294">Fig. 1</figureCitation>
D–F). Spinnerets pale brown, anteriors larger than posteriors. Colulus absent. Anal tubercle tiny, pale (
<figureCitation id="2CD52A52FFEC4714134BFD2B018B6F7A" captionStart="FIGURE 1" captionStartId="1.[151,244,1984,2006]" captionTargetBox="[189,1398,185,1951]" captionTargetId="figure@1.[189,1398,185,1964]" captionTargetPageId="1" captionText="FIGURE 1. Symphytognatha milleri n. sp., female holotype (A–C, G) and subadult male paratype (D–F, H). A–F, Habitus. G–H, Prosoma. A, D, Dorsal. B, E, Ventral. C, F, Lateral. G–H, Frontal. Scale bars: 0.50 (A–F); 0.20 (G–H)." httpUri="https://zenodo.org/record/3337753/files/figure.png" pageId="3" pageNumber="294">Fig. 1E</figureCitation>
).
</paragraph>
<paragraph id="B45136D7FFEC471416FDFC93053A6F8A" blockId="3.[151,1437,154,1042]" pageId="3" pageNumber="294">
<emphasis id="869AEAC5FFEC471416FDFC93000F6F1D" box="[199,307,838,861]" italics="true" pageId="3" pageNumber="294">Epigynum</emphasis>
(
<figureCitation id="2CD52A52FFEC4714177BFC9300416F1E" box="[321,381,838,862]" captionStart="FIGURE 2" captionStartId="2.[151,244,1930,1952]" captionTargetBox="[201,1386,167,1904]" captionTargetId="figure@2.[201,1386,167,1910]" captionTargetPageId="2" captionText="FIGURE 2. Symphytognatha milleri n. sp., female holotype. A, Habitus (omitted left legs). B–C, Chelicerae. D–E, Epigynum. F–G, Vulva (lactic acid treated). A, Lateral. B, D, Posterior. C, Anterior. E, F, Ventral. G, Dorsal. Abbreviations: cd, copulatory duct; fd, fertilization duct; sp, spermatheca; vs, vulval septum. Scale bars: 0.20 (A, D–E); 0.10 (B–C, F–G)." httpUri="https://zenodo.org/record/3337755/files/figure.png" pageId="3" pageNumber="294">Fig. 2</figureCitation>
D–G). Weakly sclerotized in posterior epigynal margin. Spermathecae twisted as Chinese doughnut. Copulatory ducts start at posterior margin of epigynum, almost run parallel to the fingerlike vulval septum. Short narrow fertilization ducts located at the bottom of spermathecae, on the outside of proximal copulatory ducts. Distal copulatory ducts membranous, translucent, connected with the outer laterals of spermathecae.
</paragraph>
<paragraph id="B45136D7FFEC471416FDFC03045C6FAE" blockId="3.[151,1437,154,1042]" box="[199,1376,982,1006]" pageId="3" pageNumber="294">
<emphasis id="869AEAC5FFEC471416FDFC03003D6FAE" bold="true" box="[199,257,982,1006]" pageId="3" pageNumber="294">Male</emphasis>
(subadult). Somatic characters as in
<figureCitation id="2CD52A52FFEC471414B0FC0303F86FAE" box="[650,708,982,1006]" captionStart="FIGURE 1" captionStartId="1.[151,244,1984,2006]" captionTargetBox="[189,1398,185,1951]" captionTargetId="figure@1.[189,1398,185,1964]" captionTargetPageId="1" captionText="FIGURE 1. Symphytognatha milleri n. sp., female holotype (A–C, G) and subadult male paratype (D–F, H). A–F, Habitus. G–H, Prosoma. A, D, Dorsal. B, E, Ventral. C, F, Lateral. G–H, Frontal. Scale bars: 0.50 (A–F); 0.20 (G–H)." httpUri="https://zenodo.org/record/3337753/files/figure.png" pageId="3" pageNumber="294">Fig. 1</figureCitation>
A–C, G. Deeper color than in female. Palpal details unknown.
</paragraph>
</subSubSection>
<subSubSection id="FCF4655CFFEC471416FDFC2F03EB6852" box="[199,727,1018,1042]" pageId="3" pageNumber="294" type="distribution">
<paragraph id="B45136D7FFEC471416FDFC2F03EB6852" blockId="3.[151,1437,154,1042]" box="[199,727,1018,1042]" pageId="3" pageNumber="294">
<emphasis id="869AEAC5FFEC471416FDFC2F00666852" bold="true" box="[199,346,1018,1042]" pageId="3" pageNumber="294">Distribution.</emphasis>
Known only from the
<typeStatus id="6B558875FFEC47141477FC2F03476852" box="[589,635,1018,1042]" pageId="3" pageNumber="294">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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