1052 lines
200 KiB
XML
1052 lines
200 KiB
XML
<document id="6CAC445A885C2D32E7C90FDD68E7BE46" ID-CLB-Dataset="54295" ID-DOI="10.11646/zootaxa.5325.3.4" ID-GBIF-Dataset="df319cf7-2755-4529-bb7f-37ee52029320" ID-ISSN="1175-5326" ID-Zenodo-Dep="8243460" ID-ZooBank="7C6BF79E-BE56-4A46-8746-CDA856CA8BF9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1691942007322" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2023" docId="A02A8786FFCCDD64FF5178382D4BFAC6" docLanguage="en" docName="zootaxa.5325.3.4.pdf" docOrigin="Zootaxa 5325 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5325.3.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Arthromelodes tokushigei Jałoszyński 2023, sp. n." docType="treatment" docVersion="4" lastPageNumber="407" masterDocId="5C13FFFEFFCDDD6AFFC67F4B293CFF93" masterDocTitle="A new species of Arthromelodes Jeannel from Okinawa-jima, Japan (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="408" masterPageNumber="393" pageNumber="394" updateTime="1692029459539" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C4CB432129EEDE4DD1396C41020594E1">A new species of Arthromelodes Jeannel from Okinawa-jima, Japan (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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<mods:namePart id="92A9330909E23DAC578312AF52406EA5">Jałoszyński, Paweł</mods:namePart>
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<mods:date id="B04845449D6BF940F13951AB5484F8E5">2023</mods:date>
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<subSubSection id="6099651BFFCCDD6BFF5178382B25F81D" box="[151,537,1907,1934]" pageId="1" pageNumber="394" type="nomenclature">
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<paragraph id="283C3690FFCCDD6BFF5178382B25F81D" blockId="1.[151,537,1907,1970]" box="[151,537,1907,1934]" pageId="1" pageNumber="394">
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<heading id="737481FCFFCCDD6BFF5178382B25F81D" bold="true" box="[151,537,1907,1934]" fontSize="11" level="1" pageId="1" pageNumber="394" reason="1">
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<emphasis id="1AF7EA82FFCCDD6BFF5178382B25F81D" bold="true" box="[151,537,1907,1934]" pageId="1" pageNumber="394">
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<taxonomicName id="EF834D13FFCCDD6BFF51783828ECF81D" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[151,464,1907,1934]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="tokushigei" status="sp. n.">
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<emphasis id="1AF7EA82FFCCDD6BFF51783828ECF81D" bold="true" box="[151,464,1907,1934]" italics="true" pageId="1" pageNumber="394">Arthromelodes tokushigei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="01C457F9FFCCDD6BFE11783F2B25F81D" box="[471,537,1908,1934]" pageId="1" pageNumber="394" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="6099651BFFCCDD6BFF5178D32825F821" box="[151,281,1944,1970]" pageId="1" pageNumber="394" type="description">
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<paragraph id="283C3690FFCCDD6BFF5178D32825F821" blockId="1.[151,537,1907,1970]" box="[151,281,1944,1970]" pageId="1" pageNumber="394">
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(
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<figureCitation id="B0B82A15FFCCDD6BFF5978D3282DF821" box="[159,273,1944,1970]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–6" captionStart-10="FIGURES 43–48" captionStart-2="FIGURES 7–10" captionStart-3="FIGURES 11–14" captionStart-4="FIGURES 15–17" captionStart-5="FIGURES 18–20" captionStart-6="FIGURES 21–26" captionStart-7="FIGURES 27–30" captionStart-8="FIGURES 31–36" captionStart-9="FIGURES 37–42" captionStartId-0="2.[151,264,1936,1960]" captionStartId-1="3.[151,264,1828,1852]" captionStartId-10="13.[151,264,1561,1585]" captionStartId-2="5.[151,264,1573,1597]" captionStartId-3="6.[151,264,1604,1628]" captionStartId-4="7.[151,264,1506,1530]" captionStartId-5="8.[151,264,1680,1704]" captionStartId-6="9.[151,264,1280,1304]" captionStartId-7="10.[151,261,1754,1778]" captionStartId-8="11.[151,264,1693,1717]" captionStartId-9="12.[151,264,1376,1400]" captionTargetBox-0="[157,1430,600,1904]" captionTargetBox-1="[151,1436,181,1804]" captionTargetBox-10="[157,1430,187,1531]" captionTargetBox-2="[151,1435,323,1550]" captionTargetBox-3="[151,1435,345,1572]" captionTargetBox-4="[151,1435,345,1483]" captionTargetBox-5="[152,1429,525,1657]" captionTargetBox-6="[155,1435,381,1257]" captionTargetBox-7="[151,1417,489,1721]" captionTargetBox-8="[151,1435,237,1670]" captionTargetBox-9="[157,1429,187,1346]" captionTargetId-0="figure-205@2.[151,1436,594,1910]" captionTargetId-1="figure-15@3.[151,1436,181,1804]" captionTargetId-10="figure-15@13.[151,1436,181,1537]" captionTargetId-2="figure-74@5.[151,1435,309,1550]" captionTargetId-3="figure-101@6.[151,1435,345,1581]" captionTargetId-4="figure-81@7.[151,1435,345,1483]" captionTargetId-5="figure-164@8.[151,1436,525,1657]" captionTargetId-6="figure-112@9.[151,1435,381,1257]" captionTargetId-7="figure-172@10.[151,1435,489,1731]" captionTargetId-8="figure-36@11.[151,1435,237,1670]" captionTargetId-9="figure-21@12.[151,1436,181,1352]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-10="13" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="11" captionTargetPageId-9="12" captionText-0="FIGURES 1–2. Arthromelodes tokushigei sp. n., male. Dorsal habitus of male (1) and female (2)." captionText-1="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." captionText-10="FIGURES 43–48. Arthromelodes tokushigei sp. n., male. Aedeagus in ventral (43, 46), ventral and tilted dorsally to visualize shape of ventral projection (44, 47), and lateral (45, 48) views." captionText-2="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." captionText-3="FIGURES 11–14. Arthromelodes tokushigei sp. n., male. Prothorax in dorsal (11), ventral (12, 13) and left lateral (14) views. Arrowheads indicate ventral foveae. Abbreviations: abg, antebasal groove; abp, antebasal pit; bst, basisternal region of prosternum; fs, furcasternum; hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; mls, median longitudinal sulcus." captionText-4="FIGURES 15–17. Arthromelodes tokushigei sp. n., male. Mesonotum and elytral base in dorsal view (15); left elytron in dorsal view (16); right elytron in dorsal view (17). Abbreviations: as, adsutural sulcus; bef, basal elytral fovea; ds, discal sulcus; eal, elytral articulating lobe; ls, lateral sulcus; phf, posthumeral fovea; sc2+scl2, fused mesoscutum and mesoscutellum." captionText-5="FIGURES 18–20. Arthromelodes tokushigei sp. n., male. Pterothorax in ventral view (18, 19); anterior region of mesoventrite in ventral view (20). Arrowheads indicate ventral foveae. Abbreviations: amtvp, anterior metaventral process; msmtb, mesometathoracic border; msvp, mesoventral intermesocoxal process; mtvp, metaventral intermetacoxal process; ppc, prepectus; v2, mesoventrite; v3, metaventrite." captionText-6="FIGURES 21–26. Arthromelodes tokushigei sp. n., male. Fore (21), middle (22) and hind (23) leg in posterior view; procoxae in ventral view (24); procoxal glandular opening (25); ventral field of carinulae on profemur (26)." captionText-7="FIGURES 27–30. Arthromelodes tokushigei sp. n., male.Abdomen in dorsal view (27); abdominal segments III–IV in dorsal (with ventral structures visible) (28) and left lateral (29) views; tergite V in dorsal view (30). Arrowheads indicate foveae (d, dorsal; v, ventral).Abbreviations: imcp, intermetacoxal process; lsp, lateral setal patch; mt, median tubercle; tIII, tergite III; tIV, tergite IV." captionText-8="FIGURES 31–36. Arthromelodes tokushigei sp. n., male. Lateral fovea on abdominal tergite IV (31); median impression on tergite IV (32); posterior region of median impression on tergite IV (33); anterior region of impression on tergite IV (34); median process of median impression on tergite IV (35); spatulate setae on lateral setal patch of tergite IV (36). Abbreviations: lp, lateral process; mp, median process; mt, median tubercle; smp, submedian process." captionText-9="FIGURES 37–42. Arthromelodes tokushigei sp. n., male (37–39) and female (40–42). Tergite VIII in dorsal view (37, 40); sternite VIII in ventral view (38, 42); sternite IX in ventral view, with internal membrane (39); postabdominal sclerites (41)." figureDoi-0="http://doi.org/10.5281/zenodo.8243462" figureDoi-1="http://doi.org/10.5281/zenodo.8243464" figureDoi-10="http://doi.org/10.5281/zenodo.8243482" figureDoi-2="http://doi.org/10.5281/zenodo.8243466" figureDoi-3="http://doi.org/10.5281/zenodo.8243468" figureDoi-4="http://doi.org/10.5281/zenodo.8243470" figureDoi-5="http://doi.org/10.5281/zenodo.8243472" figureDoi-6="http://doi.org/10.5281/zenodo.8243474" figureDoi-7="http://doi.org/10.5281/zenodo.8243476" figureDoi-8="http://doi.org/10.5281/zenodo.8243478" figureDoi-9="http://doi.org/10.5281/zenodo.8243480" httpUri-0="https://zenodo.org/record/8243462/files/figure.png" httpUri-1="https://zenodo.org/record/8243464/files/figure.png" httpUri-10="https://zenodo.org/record/8243482/files/figure.png" httpUri-2="https://zenodo.org/record/8243466/files/figure.png" httpUri-3="https://zenodo.org/record/8243468/files/figure.png" httpUri-4="https://zenodo.org/record/8243470/files/figure.png" httpUri-5="https://zenodo.org/record/8243472/files/figure.png" httpUri-6="https://zenodo.org/record/8243474/files/figure.png" httpUri-7="https://zenodo.org/record/8243476/files/figure.png" httpUri-8="https://zenodo.org/record/8243478/files/figure.png" httpUri-9="https://zenodo.org/record/8243480/files/figure.png" pageId="1" pageNumber="394">Figs 1–48</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6099651BFFCCDD68FF5178AB2D20FF6A" lastPageId="2" lastPageNumber="395" pageId="1" pageNumber="394" type="materials_examined">
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<paragraph id="283C3690FFCCDD68FF5178AB2D20FF6A" blockId="1.[151,1436,2016,2044]" lastBlockId="2.[151,1436,150,573]" lastPageId="2" lastPageNumber="395" pageId="1" pageNumber="394">
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<emphasis id="1AF7EA82FFCCDD6BFF5178AB2874F869" bold="true" box="[151,328,2016,2042]" pageId="1" pageNumber="394">Type material.</emphasis>
|
||
<materialsCitation id="98EB3CCDFFCCDD68FE9478AB2DCCFF46" collectingDate="2023-02-26" collectionCode="NSMT" collectorName="P. Jaloszynski" country="Japan" lastPageId="2" lastPageNumber="395" location="Okinawa Pref." pageId="1" pageNumber="394" specimenCount="1" stateProvince="Okinawa Pref." typeStatus="holotype">
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<emphasis id="1AF7EA82FFCCDD6BFE9478AB2A3FF869" bold="true" box="[338,771,2016,2042]" pageId="1" pageNumber="394">
|
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<typeStatus id="F7388832FFCCDD6BFE9478AB28FFF869" box="[338,451,2016,2042]" pageId="1" pageNumber="394">Holotype</typeStatus>
|
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:
|
||
<collectingCountry id="50947600FFCCDD6BFE1478AB2B10F869" box="[466,556,2016,2042]" name="Japan" pageId="1" pageNumber="394">JAPAN</collectingCountry>
|
||
(
|
||
<collectingRegion id="EA47F872FFCCDD6BFDF878AB2BCAF869" box="[574,758,2016,2042]" country="Japan" name="Okinawa" pageId="1" pageNumber="394">Okinawa Pref.</collectingRegion>
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):
|
||
</emphasis>
|
||
♁, two labels: “
|
||
<emphasis id="1AF7EA82FFCCDD6BFC0178AB2D1DF869" bold="true" box="[967,1057,2016,2042]" pageId="1" pageNumber="394">JAPAN</emphasis>
|
||
, OKINAWA Pref. / OKINAWAjima,
|
||
<date id="5C3D1050FFCFDD68FF1E7FDC2870FF22" box="[216,332,151,177]" pageId="2" pageNumber="395" value="2023-02-26">
|
||
<collectingDate id="4C79E9B8FFCFDD68FF1E7FDC2870FF22" box="[216,332,151,177]" pageId="2" pageNumber="395" value="2023-02-26">26 ii 2023</collectingDate>
|
||
</date>
|
||
/ Ohkuni-rindo, Jashiki / 2645’15.7’’N/12814’44.8’’E / leg. P. JAŁOSZYŃSKI” [white, printed], “
|
||
<taxonomicName id="EF834D13FFCFDD68FF627FF028B3FF46" authorityName="Jeannel" authorityYear="1954" box="[164,399,187,213]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="395" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1AF7EA82FFCFDD68FF627FF028B3FF46" box="[164,399,187,213]" italics="true" pageId="2" pageNumber="395">ARTHROMELODES</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="EF834D13FFCFDD68FE647FF02B29FF46" authorityName="Jałoszyński" authorityYear="2023" box="[418,533,187,213]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="395" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFCFDD68FE647FF02B29FF46" box="[418,533,187,213]" italics="true" pageId="2" pageNumber="395">tokushigei</emphasis>
|
||
</taxonomicName>
|
||
m. /
|
||
<collectorName id="85765346FFCFDD68FD8A7FF02BD6FF46" box="[588,746,187,213]" pageId="2" pageNumber="395">P. Jałoszyński</collectorName>
|
||
, 2023/ HOLOTYPUS” [red, printed] (
|
||
<collectionCode id="4E92AE55FFCFDD68FB557FF02DDBFF46" box="[1171,1255,187,213]" collectionName="Japan, Tokyo, National Science Museum (Natural History)" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="2" pageNumber="395">NSMT</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="98EB3CCDFFCFDD68FB3F7FF02D24FF6A" collectingDate="2023-02-26" collectionCode="NSMT" collectorName="P. Jaloszynski" country="Japan" location="Okinawa Pref." pageId="2" pageNumber="395" specimenCount="10" specimenCount-female="3" stateProvince="Okinawa" typeStatus="paratype">
|
||
<emphasis id="1AF7EA82FFCFDD68FB3F7FF029E6FF6B" bold="true" pageId="2" pageNumber="395">
|
||
<typeStatus id="F7388832FFCFDD68FB3F7FF02C4DFF46" box="[1273,1393,187,213]" pageId="2" pageNumber="395">Paratypes</typeStatus>
|
||
(10 exx.):
|
||
</emphasis>
|
||
<specimenCount id="3E85FD19FFCFDD68FF267F942823FF68" box="[224,287,223,251]" pageId="2" pageNumber="395">7 ♁♁</specimenCount>
|
||
(one disarticulated),
|
||
<specimenCount id="3E85FD19FFCFDD68FDCB7F942B71FF6A" box="[525,589,223,249]" pageId="2" pageNumber="395" type="female">3 ♀♀</specimenCount>
|
||
, same data as for holotype (cPJ,
|
||
<collectionCode id="4E92AE55FFCFDD68FC787F942D2DFF6A" box="[958,1041,223,249]" collectionName="Japan, Tokyo, National Science Museum (Natural History)" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="2" pageNumber="395">NSMT</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6099651BFFCFDD68FF017E492BBBFDAE" pageId="2" pageNumber="395" type="diagnosis">
|
||
<paragraph id="283C3690FFCFDD68FF017E492BBBFDAE" blockId="2.[151,1436,150,573]" pageId="2" pageNumber="395">
|
||
<emphasis id="1AF7EA82FFCFDD68FF017E49287EFE8F" bold="true" box="[199,322,258,284]" pageId="2" pageNumber="395">Diagnosis.</emphasis>
|
||
Punctures on dorsum of head dense and distinct in anterior half, posteriorly becoming shallower and sparser; punctures on pronotal disc distinct and dense but shallow; in males each profemur with elongate transversely carinulate field in basal half of ventral surface and mesotibia with elongate ventral apical tooth or mucro, but all trochanters unmodified; abdominal tergite IV in male with large posterior transverse impression bearing large oval median tubercle near posterior margin of tergite and three pairs of minute elongate processes in front of tubercle, lateral setal patches of male tergite IV situated entirely laterally to posteromedian impression; aedeagus in ventral view with ventral projection of median lobe (inserted just above orifice) recurved and with subtriangular apical region with strongly oblique distal margin, dorsal projection in lateral view bent ventrad nearly at right angle and with minute subtriangular proximal process.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="7CFC6618FFCFDD68FF5178DB2D4AF83B" ID-DOI="http://doi.org/10.5281/zenodo.8243462" ID-Zenodo-Dep="8243462" box="[151,1142,1936,1960]" httpUri="https://zenodo.org/record/8243462/files/figure.png" pageId="2" pageNumber="395" startId="2.[151,264,1936,1960]" targetBox="[157,1430,600,1904]" targetPageId="2" targetType="figure">
|
||
<paragraph id="283C3690FFCFDD68FF5178DB2D4AF83B" blockId="2.[151,1142,1936,1960]" box="[151,1142,1936,1960]" pageId="2" pageNumber="395">
|
||
<emphasis id="1AF7EA82FFCFDD68FF5178DB2806F83B" bold="true" box="[151,314,1936,1960]" pageId="2" pageNumber="395">FIGURES 1–2.</emphasis>
|
||
<taxonomicName id="EF834D13FFCFDD68FE8778DB2B79F83B" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[321,581,1936,1960]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="395" phylum="Arthropoda" rank="species" species="tokushigei" status="sp. n.">
|
||
<emphasis id="1AF7EA82FFCFDD68FE8778DB2B79F83B" box="[321,581,1936,1960]" italics="true" pageId="2" pageNumber="395">Arthromelodes tokushigei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="01C457F9FFCFDD68FD8D78DB2BBFF834" box="[587,643,1936,1959]" pageId="2" pageNumber="395" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male. Dorsal habitus of male (
|
||
<emphasis id="1AF7EA82FFCFDD68FC0778DB2AF2F83B" bold="true" box="[961,974,1936,1960]" pageId="2" pageNumber="395">1</emphasis>
|
||
) and female (
|
||
<emphasis id="1AF7EA82FFCFDD68FB9D78DB2D54F83B" bold="true" box="[1115,1128,1936,1960]" pageId="2" pageNumber="395">2</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7CFC6618FFCEDD69FF51786F2DE5F83B" ID-DOI="http://doi.org/10.5281/zenodo.8243464" ID-Zenodo-Dep="8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="3" pageNumber="396" startId="3.[151,264,1828,1852]" targetBox="[151,1436,181,1804]" targetPageId="3" targetType="figure">
|
||
<paragraph id="283C3690FFCEDD69FF51786F2DE5F83B" blockId="3.[151,1437,1828,1960]" pageId="3" pageNumber="396">
|
||
<emphasis id="1AF7EA82FFCEDD69FF51786F2800F8AF" bold="true" box="[151,316,1828,1852]" pageId="3" pageNumber="396">FIGURES 3–6.</emphasis>
|
||
<taxonomicName id="EF834D13FFCEDD69FE82786F2B76F8AF" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[324,586,1828,1852]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="tokushigei" status="sp. n.">
|
||
<emphasis id="1AF7EA82FFCEDD69FE82786F2B76F8AF" box="[324,586,1828,1852]" italics="true" pageId="3" pageNumber="396">Arthromelodes tokushigei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="01C457F9FFCEDD69FD94786E2BB7F8AF" box="[594,651,1829,1852]" pageId="3" pageNumber="396" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male. Head in dorsal (
|
||
<emphasis id="1AF7EA82FFCEDD69FCB9786F2AB0F8AF" bold="true" box="[895,908,1828,1852]" pageId="3" pageNumber="396">3</emphasis>
|
||
), ventral (
|
||
<emphasis id="1AF7EA82FFCEDD69FC3F786F2D3AF8AF" bold="true" box="[1017,1030,1828,1852]" pageId="3" pageNumber="396">4</emphasis>
|
||
) and lateral (
|
||
<emphasis id="1AF7EA82FFCEDD69FB55786F2D9CF8AF" bold="true" box="[1171,1184,1828,1852]" pageId="3" pageNumber="396">5</emphasis>
|
||
) views; right antenna in dorsal view (
|
||
<emphasis id="1AF7EA82FFCEDD69FEDD78032814F8F3" bold="true" box="[283,296,1864,1888]" pageId="3" pageNumber="396">6</emphasis>
|
||
). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="6099651BFFC9DD66FF017FDD2DC5F92F" lastPageId="12" lastPageNumber="405" pageId="4" pageNumber="397" type="description">
|
||
<paragraph id="283C3690FFC9DD6EFF017FDD2A00FF46" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="397">
|
||
<emphasis id="1AF7EA82FFC9DD6EFF017FDD2865FF23" bold="true" box="[199,345,150,176]" pageId="4" pageNumber="397">Description.</emphasis>
|
||
Body of male (
|
||
<figureCitation id="B0B82A15FFC9DD6EFDCC7FDC2B77FF22" box="[522,587,151,177]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1936,1960]" captionTargetBox="[157,1430,600,1904]" captionTargetId="figure-205@2.[151,1436,594,1910]" captionTargetPageId="2" captionText="FIGURES 1–2. Arthromelodes tokushigei sp. n., male. Dorsal habitus of male (1) and female (2)." figureDoi="http://doi.org/10.5281/zenodo.8243462" httpUri="https://zenodo.org/record/8243462/files/figure.png" pageId="4" pageNumber="397">Fig. 1</figureCitation>
|
||
) elongate, strongly convex, moderately to darkly brown, with tarsi slightly lighter, setae slightly lighter than cuticle; BL
|
||
<quantity id="EF7B9B75FFC9DD6EFD547FF02A04FF46" box="[658,824,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.115" metricValueMax="2.19" metricValueMin="2.04" pageId="4" pageNumber="397" unit="mm" value="2.115" valueMax="2.19" valueMin="2.04">2.04–2.19 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="283C3690FFC9DD6EFF017F942DE3FC36" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="397">
|
||
Head (
|
||
<figureCitation id="B0B82A15FFC9DD6EFED67F94284FFF6A" box="[272,371,222,249]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Figs 3–5</figureCitation>
|
||
) subhexagonal, flattened, broadest at eyes, HL
|
||
<quantity id="EF7B9B75FFC9DD6EFC477F942D19FF6A" box="[897,1061,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="4.449999999999999" metricValueMax="4.8" metricValueMin="4.1" pageId="4" pageNumber="397" unit="mm" value="0.44499999999999995" valueMax="0.48" valueMin="0.41">0.41–0.48 mm</quantity>
|
||
, HW
|
||
<quantity id="EF7B9B75FFC9DD6EFBA27F952C36FF6A" box="[1124,1290,222,249]" metricMagnitude="-4" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" pageId="4" pageNumber="397" unit="mm" value="0.445" valueMax="0.45" valueMin="0.44">0.44–0.45 mm</quantity>
|
||
. Neck region (
|
||
<figureCitation id="B0B82A15FFC9DD6EFF597E4829D4FE8E" box="[159,232,259,285]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Fig. 3</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFF3E7E48282DFE8F" box="[248,273,259,284]" italics="true" pageId="4" pageNumber="397">nr</emphasis>
|
||
) dorsally smooth and with short posteromedian occipital carina (
|
||
<figureCitation id="B0B82A15FFC9DD6EFBCA7E482D68FE8E" box="[1036,1108,259,285]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Fig. 3</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFBA27E482DB7FE8F" box="[1124,1163,259,284]" italics="true" pageId="4" pageNumber="397">occ</emphasis>
|
||
); ventrally neck region with distinct gular sutures (
|
||
<figureCitation id="B0B82A15FFC9DD6EFE1E7E6C2B1EFED2" box="[472,546,295,321]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Fig. 4</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFDF77E6C2B76FED3" box="[561,586,295,320]" italics="true" pageId="4" pageNumber="397">gs</emphasis>
|
||
) obliterated anteriorly and demarcating subtriangular gular plate (
|
||
<figureCitation id="B0B82A15FFC9DD6EFA887E6C2CABFED2" box="[1358,1431,295,321]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Fig. 4</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFF517E00298FFEF7" box="[151,179,331,356]" italics="true" pageId="4" pageNumber="397">gp</emphasis>
|
||
) covered with strongly transverse microreticulation; vertex (
|
||
<figureCitation id="B0B82A15FFC9DD6EFCAF7E002A93FEF6" box="[873,943,331,357]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Fig. 3</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFC787E002AEEFEF7" box="[958,978,331,356]" italics="true" pageId="4" pageNumber="397">vt</emphasis>
|
||
) weakly convex medially and distinctly impressed laterally, with short median longitudinal vertexal carina (
|
||
<figureCitation id="B0B82A15FFC9DD6EFC427E242AF4FE1A" box="[900,968,367,393]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Fig. 3</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFC157E242AD0FE1B" box="[979,1004,367,392]" italics="true" pageId="4" pageNumber="397">vc</emphasis>
|
||
) adjacent to posterior margin of vertex and not connected with occipital carina; dorsal tentorial pits (
|
||
<figureCitation id="B0B82A15FFC9DD6EFC807ED82AB7FE3E" box="[838,907,403,429]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Fig. 3</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFC5E7ED82A80FE3E" box="[920,956,403,429]" italics="true" pageId="4" pageNumber="397">dtp</emphasis>
|
||
) large and deep, circular and asetose, each situated in lateral impressed region of vertex; vertex in front of dorsal tentorial pits and posterior margin of frons (
|
||
<figureCitation id="B0B82A15FFC9DD6EFF597E9029DFFE66" box="[159,227,475,501]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Fig. 3</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFF287E90283DFE66" box="[238,257,475,501]" italics="true" pageId="4" pageNumber="397">fr</emphasis>
|
||
) in front of eyes with shallow vertexal sulcus (
|
||
<figureCitation id="B0B82A15FFC9DD6EFCC27E902A7BFE66" box="[772,839,475,501]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Fig. 3</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFC947E902A55FE67" box="[850,873,475,500]" italics="true" pageId="4" pageNumber="397">vs</emphasis>
|
||
) forming M-shaped figure with shallow transverse portion; frons at middle slightly impressed, impressed region flanked by prominent supraantennal tubercles. Eyes (
|
||
<figureCitation id="B0B82A15FFC9DD6EFF597D682839FDAE" box="[159,261,546,573]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Figs 3–5</figureCitation>
|
||
) large, strongly convex, bean-shaped, each with 18 convex corneal lenses. Occipital constriction (
|
||
<figureCitation id="B0B82A15FFC9DD6EFAA97D68299BFDF2" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Fig. 3</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFF737D0C29E0FDF3" box="[181,220,583,608]" italics="true" pageId="4" pageNumber="397">occ</emphasis>
|
||
) about as wide as 1/3 of HW including eyes; tempora (
|
||
<figureCitation id="B0B82A15FFC9DD6EFC8A7D0C2AADFDF2" box="[844,913,583,609]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Fig. 3</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFC587D0C2A86FDF3" box="[926,954,583,608]" italics="true" pageId="4" pageNumber="397">tm</emphasis>
|
||
) in dorsal view nearly as long as length of eye, nearly straight and strongly converging posterad. Clypeus (
|
||
<figureCitation id="B0B82A15FFC9DD6EFCA47D202A9BFD16" box="[866,935,619,645]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Fig. 3</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFC757D202AFBFD16" box="[947,967,619,645]" italics="true" pageId="4" pageNumber="397">cl</emphasis>
|
||
) semicircular with rounded and distinctly carinate and upturned anterior margin. Punctures on dorsum of head dense and composed of large deep punctures on clypeus, posteriorly on frons and vertex gradually reducing in depth and becoming sparser, so that posterior margin of vertex remains virtually impunctate; setae on clypeus, frons and vertex sparse, short and suberect. Genae and postgenae (
|
||
<figureCitation id="B0B82A15FFC9DD6EFED07DB02867FC86" box="[278,347,763,789]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Fig. 4</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFEA17DB028B3FC87" box="[359,399,763,788]" italics="true" pageId="4" pageNumber="397">gen</emphasis>
|
||
,
|
||
<emphasis id="1AF7EA82FFC9DD6EFE5B7DB028EFFC87" box="[413,467,763,788]" italics="true" pageId="4" pageNumber="397">pgen</emphasis>
|
||
) weakly convex and sparsely, distinctly punctate, sparsely covered with long and erect setae directed mainly laterally and anterolaterally. Antennal fossa conspicuously large and ventrally accentuated by distinct ocular-mandibular carina (
|
||
<figureCitation id="B0B82A15FFC9DD6EFDD37C082B64FCCE" box="[533,600,834,861]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Fig. 5</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFDA57C082BAEFCCF" box="[611,658,835,860]" italics="true" pageId="4" pageNumber="397">omc</emphasis>
|
||
) running from ventral margin of eye to mandibular base and anteriorly confluent with marginal carina of clypeus. Posterior tentorial pits (marking border between gula and submentum) fused at middle to form single slightly transverse pit situated at level of posterior margins of eyes.
|
||
</paragraph>
|
||
<paragraph id="283C3690FFC9DD6EFF017CE42C67FB32" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="397">
|
||
Antennae (
|
||
<figureCitation id="B0B82A15FFC9DD6EFE867CE428BAFC5A" box="[320,390,943,969]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1936,1960]" captionTargetBox="[157,1430,600,1904]" captionTargetId="figure-205@2.[151,1436,594,1910]" captionTargetPageId="2" captionText="FIGURES 1–2. Arthromelodes tokushigei sp. n., male. Dorsal habitus of male (1) and female (2)." figureDoi="http://doi.org/10.5281/zenodo.8243462" httpUri="https://zenodo.org/record/8243462/files/figure.png" pageId="4" pageNumber="397">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="B0B82A15FFC9DD6EFE527CE4289FFC5A" box="[404,419,943,969]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">6</figureCitation>
|
||
) slightly shorter than head, pronotum and elytra together, weakly thickening toward apices; AnL
|
||
<quantity id="EF7B9B75FFC9DD6EFF147C982845FC7E" box="[210,377,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="1.2149999999999999" metricValueMax="1.23" metricValueMin="1.2" pageId="4" pageNumber="397" unit="mm" value="1.2149999999999999" valueMax="1.23" valueMin="1.2">1.20–1.23 mm</quantity>
|
||
; scape distinctly elongate, subcylindrical, with deep dorsolateral emargination and flattening, pedicel slightly broadening distad, distinctly elongate but clearly shorter and narrower than scape; antennomeres 3–4 each about as long as broad, antennomeres 5–7 each slightly elongate (7 longest), antennomere 8 distinctly shorter than 7, about as long as broad, antennomeres 9–10 each about as long as broad, antennomere 11 nearly as long as 9–10 combined, distinctly broader than 10, fusiform, unmodified. All antennomeres covered with coarse microreticulation and dense setae, those on antennomeres 9–11 longer than those on proximal antennomeres.
|
||
</paragraph>
|
||
<paragraph id="283C3690FFC9DD6EFF017BE02D7BFA9E" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="397">
|
||
Labrum (
|
||
<figureCitation id="B0B82A15FFC9DD6EFEE87BE0284CFB56" box="[302,368,1195,1221]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">Fig. 7</figureCitation>
|
||
) steeply declining anterad, broadening anteriorly, strongly transverse, with distinct subtriangular anterolateral lobes and nearly straight anterior margin; dorsal surface with sparse paired setae, four peg-like sensilla (
|
||
<figureCitation id="B0B82A15FFC9DD6EFF597BB829D8FA9E" box="[159,228,1267,1293]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">Fig. 7</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFF377BB8282EFA9E" box="[241,274,1267,1293]" italics="true" pageId="4" pageNumber="397">pls</emphasis>
|
||
) of anterior epipharynx long, apically pointed and adjacent to each other.
|
||
</paragraph>
|
||
<paragraph id="283C3690FFC9DD6EFF017A5C28D4FA0E" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="397">
|
||
Mandibles (
|
||
<figureCitation id="B0B82A15FFC9DD6EFE8B7A5C288EFAA2" box="[333,434,1303,1329]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">Figs 8, 9</figureCitation>
|
||
) subtriangular, flattened dorsoventrally, each with sharp apical incisor and preapical row of four teeth reducing in size toward mandibular base; outer mandibular margin strongly convex and indistinctly undulate, with long seta inserted dorsolaterally in distal 1/3 of mandible, and a few short setae distributed along outer margin in posterior half.
|
||
</paragraph>
|
||
<paragraph id="283C3690FFC9DD6EFF017AEC2B24F8BA" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="397">
|
||
Each maxilla(
|
||
<figureCitation id="B0B82A15FFC9DD6EFEA47AEC2893FA52" box="[354,431,1447,1473]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">Fig.10</figureCitation>
|
||
)with transverse cardo(
|
||
<figureCitation id="B0B82A15FFC9DD6EFD6B7AEC2BC1FA52" box="[685,765,1447,1473]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">Fig.10</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFCC37AEC2A23FA52" box="[773,799,1447,1473]" italics="true" pageId="4" pageNumber="397">cd</emphasis>
|
||
),subtriangular basistipes(
|
||
<figureCitation id="B0B82A15FFC9DD6EFB847AEC2DADFA52" box="[1090,1169,1447,1473]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">Fig.10</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFB5F7AEC2D86FA52" box="[1177,1210,1447,1473]" italics="true" pageId="4" pageNumber="397">bst</emphasis>
|
||
)and subrectangular, elongate mediostipes (Fog. 10;
|
||
<emphasis id="1AF7EA82FFC9DD6EFE337A802B20FA77" box="[501,540,1483,1508]" italics="true" pageId="4" pageNumber="397">mst</emphasis>
|
||
); palpifer (
|
||
<figureCitation id="B0B82A15FFC9DD6EFD5F7A802BD0FA76" box="[665,748,1483,1509]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFD317A802A27FA76" box="[759,795,1483,1509]" italics="true" pageId="4" pageNumber="397">ppf</emphasis>
|
||
) elongate, clearly demarcated from stipes; galea (
|
||
<figureCitation id="B0B82A15FFC9DD6EFA837A802CABFA76" box="[1349,1431,1483,1509]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFF517AA42987F99A" box="[151,187,1519,1545]" italics="true" pageId="4" pageNumber="397">gal</emphasis>
|
||
) weakly elongate, with dense fringe of long setae along distal and mesal margins; lacinia (
|
||
<figureCitation id="B0B82A15FFC9DD6EFB0E7AA42C21F99A" box="[1224,1309,1519,1545]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFAEC7AA42C70F99A" box="[1322,1356,1519,1545]" italics="true" pageId="4" pageNumber="397">lac</emphasis>
|
||
) much shorter than galea and with sparser and thicker setae along mesal margin. Maxillary palp (
|
||
<figureCitation id="B0B82A15FFC9DD6EFBBA79582DF8F9BE" box="[1148,1220,1555,1581]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="B0B82A15FFC9DD6EFB0979582DD0F9BE" box="[1231,1260,1555,1581]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">10</figureCitation>
|
||
) slightly longer than exposed anterior region of head capsule; palpomere 1 (
|
||
<figureCitation id="B0B82A15FFC9DD6EFCE8797C2ABDF9C2" box="[814,897,1591,1617]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFC4A797C2AF5F9C2" box="[908,969,1591,1617]" italics="true" pageId="4" pageNumber="397">mxp1</emphasis>
|
||
) minute, elongate and asetose; palpomere 2 (
|
||
<figureCitation id="B0B82A15FFC9DD6EFF7279102834F9E6" box="[180,264,1627,1653]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFED27910286CF9E6" box="[276,336,1627,1653]" italics="true" pageId="4" pageNumber="397">mxp2</emphasis>
|
||
) pipe-shaped, strongly elongate and weakly curved in distal, broadened region, with distal margin oblique and carinate, surface covered with several sparsely distributed setae; palpomere 3 (
|
||
<figureCitation id="B0B82A15FFC9DD6EFB6179342DC0F90A" box="[1191,1276,1663,1689]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFACC79342C7AF90A" box="[1290,1350,1663,1689]" italics="true" pageId="4" pageNumber="397">mxp3</emphasis>
|
||
) small, subtriangular in lateral view, with sparse long setae distributed mainly on posterior margin; palpomere 4 (
|
||
<figureCitation id="B0B82A15FFC9DD6EFA8579E82CABF92E" box="[1347,1431,1699,1725]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFF51798C29E8F972" box="[151,212,1735,1761]" italics="true" pageId="4" pageNumber="397">mxp4</emphasis>
|
||
) fusiform, strongly enlarged, covered with sparse and almost evenly distributed short setae and with 2–3 longer and more erect setae (in
|
||
<figureCitation id="B0B82A15FFC9DD6EFE3A79A02B70F896" box="[508,588,1771,1797]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">Fig. 10</figureCitation>
|
||
one long seta is visible near base); apical sensory appendage of palpomere 4 (
|
||
<figureCitation id="B0B82A15FFC9DD6EFF7278442834F8BA" box="[180,264,1807,1833]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFED278442811F8BB" box="[276,301,1807,1832]" italics="true" pageId="4" pageNumber="397">sa</emphasis>
|
||
) minute, subconical.
|
||
</paragraph>
|
||
<paragraph id="283C3690FFC9DD6FFF0178782B9CFE8E" blockId="4.[151,1437,150,2013]" lastBlockId="5.[151,1436,151,285]" lastPageId="5" lastPageNumber="398" pageId="4" pageNumber="397">
|
||
Labium (
|
||
<figureCitation id="B0B82A15FFC9DD6EFEE978782845F8DE" box="[303,377,1843,1869]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="B0B82A15FFC9DD6EFE4178782898F8DE" box="[391,420,1843,1869]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">10</figureCitation>
|
||
) with submentum (
|
||
<figureCitation id="B0B82A15FFC9DD6EFD4278782BF7F8DE" box="[644,715,1843,1869]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Fig. 4</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFD1E78782A39F8DF" box="[728,773,1843,1868]" italics="true" pageId="4" pageNumber="397">smn</emphasis>
|
||
) discernible as short and slightly convex transverse stripe just behind mentum and prolonged posteriorly as narrow median submental carina (
|
||
<figureCitation id="B0B82A15FFC9DD6EFB8D781C2DAEF8E2" box="[1099,1170,1879,1905]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="4" pageNumber="397">Fig. 4</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFB58781C2DF6F8E3" box="[1182,1226,1879,1904]" italics="true" pageId="4" pageNumber="397">smc</emphasis>
|
||
) between strongly expanded mesally genae and postgenae; mentum (
|
||
<figureCitation id="B0B82A15FFC9DD6EFD2978302A74F806" box="[751,840,1915,1941]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFC9C78302A40F807" box="[858,892,1915,1940]" italics="true" pageId="4" pageNumber="397">mn</emphasis>
|
||
) inversely subtrapezoidal with deep posterior constriction, so that posterior and anterior corners are projecting laterally, anterior margin convex at middle and emarginate at each side. Prelabium (
|
||
<figureCitation id="B0B82A15FFC9DD6EFDE878882BBEF84E" box="[558,642,1987,2013]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="4" pageNumber="397">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC9DD6EFD4878882B8EF84E" box="[654,690,1987,2013]" italics="true" pageId="4" pageNumber="397">plb</emphasis>
|
||
) largely membranous, with poorly discernible premental sclerites; insertions of labial palps narrowly separated; labial palps trimerous, palpomere 1 (
|
||
<figureCitation id="B0B82A15FFC8DD6FFB807FDC2DA0FF22" box="[1094,1180,151,177]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="5" pageNumber="398">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC8DD6FFB6C7FDC2DF1FF22" box="[1194,1229,151,177]" italics="true" pageId="5" pageNumber="398">lp1</emphasis>
|
||
) minute, annulate and asetose; palpomere 2 (
|
||
<figureCitation id="B0B82A15FFC8DD6FFE0C7FF02B1CFF46" box="[458,544,187,213]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="5" pageNumber="398">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC8DD6FFDE87FF02B6EFF46" box="[558,594,187,213]" italics="true" pageId="5" pageNumber="398">lp2</emphasis>
|
||
) strongly elongate, weakly curved and slightly broadening towards apex, with several short and sparse setae mainly distributed in mesal apical region; palpomere 3 (
|
||
<figureCitation id="B0B82A15FFC8DD6FFB647F942DCAFF6A" box="[1186,1270,223,249]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1573,1597]" captionTargetBox="[151,1435,323,1550]" captionTargetId="figure-74@5.[151,1435,309,1550]" captionTargetPageId="5" captionText="FIGURES 7–10. Arthromelodes tokushigei sp. n., male. Labrum in dorsal view (7); left (8) and right (9) mandible in dorsal view; maxillary-labial complex in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage." figureDoi="http://doi.org/10.5281/zenodo.8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="5" pageNumber="398">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC8DD6FFAC57F942C1AFF6A" box="[1283,1318,223,249]" italics="true" pageId="5" pageNumber="398">lp3</emphasis>
|
||
) setiform, strongly elongate, slender, curved and asetose.
|
||
</paragraph>
|
||
<caption id="7CFC6618FFC8DD6FFF51796E2B37F93A" ID-DOI="http://doi.org/10.5281/zenodo.8243466" ID-Zenodo-Dep="8243466" httpUri="https://zenodo.org/record/8243466/files/figure.png" pageId="5" pageNumber="398" startId="5.[151,264,1573,1597]" targetBox="[151,1435,323,1550]" targetPageId="5" targetType="figure">
|
||
<paragraph id="283C3690FFC8DD6FFF51796E2B37F93A" blockId="5.[151,1436,1573,1706]" pageId="5" pageNumber="398">
|
||
<emphasis id="1AF7EA82FFC8DD6FFF51796E2875F9AE" bold="true" box="[151,329,1573,1597]" pageId="5" pageNumber="398">FIGURES 7–10.</emphasis>
|
||
<taxonomicName id="EF834D13FFC8DD6FFE97796E2B6AF9AE" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[337,598,1573,1597]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="398" phylum="Arthropoda" rank="species" species="tokushigei" status="sp. n.">
|
||
<emphasis id="1AF7EA82FFC8DD6FFE97796E2B6AF9AE" box="[337,598,1573,1597]" italics="true" pageId="5" pageNumber="398">Arthromelodes tokushigei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="01C457F9FFC8DD6FFD98796D2BA4F9AE" box="[606,664,1574,1597]" pageId="5" pageNumber="398" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male. Labrum in dorsal view (
|
||
<emphasis id="1AF7EA82FFC8DD6FFC19796E2AD0F9AE" bold="true" box="[991,1004,1573,1597]" pageId="5" pageNumber="398">7</emphasis>
|
||
); left (
|
||
<emphasis id="1AF7EA82FFC8DD6FFBF0796E2D7FF9AE" bold="true" box="[1078,1091,1573,1597]" pageId="5" pageNumber="398">8</emphasis>
|
||
) and right (
|
||
<emphasis id="1AF7EA82FFC8DD6FFB79796E2DF0F9AE" bold="true" box="[1215,1228,1573,1597]" pageId="5" pageNumber="398">9</emphasis>
|
||
) mandible in dorsal view; maxillary-labial complex in ventral view (
|
||
<emphasis id="1AF7EA82FFC8DD6FFD4979022B94F9F2" bold="true" box="[655,680,1609,1633]" pageId="5" pageNumber="398">10</emphasis>
|
||
). Abbreviations: bst, basistipes; cd, cardo; gal, galea; lac, lacinia; lp1–3, labial palpomere 1–3; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere 1–4; plb, prelabium; pls, peg-like sensilla; ppf, palpifer; sa, sensory appendage.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="283C3690FFC8DD6CFF0179B02A7AFED2" blockId="5.[151,1437,1787,2030]" lastBlockId="6.[151,1437,151,321]" lastPageId="6" lastPageNumber="399" pageId="5" pageNumber="398">
|
||
Prothorax (
|
||
<figureCitation id="B0B82A15FFC8DD6FFE8579B02883F886" box="[323,447,1787,1813]" captionStart="FIGURES 11–14" captionStartId="6.[151,264,1604,1628]" captionTargetBox="[151,1435,345,1572]" captionTargetId="figure-101@6.[151,1435,345,1581]" captionTargetPageId="6" captionText="FIGURES 11–14. Arthromelodes tokushigei sp. n., male. Prothorax in dorsal (11), ventral (12, 13) and left lateral (14) views. Arrowheads indicate ventral foveae. Abbreviations: abg, antebasal groove; abp, antebasal pit; bst, basisternal region of prosternum; fs, furcasternum; hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; mls, median longitudinal sulcus." figureDoi="http://doi.org/10.5281/zenodo.8243468" httpUri="https://zenodo.org/record/8243468/files/figure.png" pageId="5" pageNumber="398">Figs 11–14</figureCitation>
|
||
) convex dorsally and flattened ventrally, slightly transverse, in dorsal view subhexagonal. Pronotum (
|
||
<figureCitation id="B0B82A15FFC8DD6FFEDE785428A2F8AA" box="[280,414,1823,1849]" captionStart="FIGURES 11–14" captionStartId="6.[151,264,1604,1628]" captionTargetBox="[151,1435,345,1572]" captionTargetId="figure-101@6.[151,1435,345,1581]" captionTargetPageId="6" captionText="FIGURES 11–14. Arthromelodes tokushigei sp. n., male. Prothorax in dorsal (11), ventral (12, 13) and left lateral (14) views. Arrowheads indicate ventral foveae. Abbreviations: abg, antebasal groove; abp, antebasal pit; bst, basisternal region of prosternum; fs, furcasternum; hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; mls, median longitudinal sulcus." figureDoi="http://doi.org/10.5281/zenodo.8243468" httpUri="https://zenodo.org/record/8243468/files/figure.png" pageId="5" pageNumber="398">Figs 11, 14</figureCitation>
|
||
) broadest distinctly in front of middle; PL
|
||
<quantity id="EF7B9B75FFC8DD6FFC5D78542D78F8A9" box="[923,1092,1823,1850]" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="5" pageNumber="398" unit="mm" value="0.475" valueMax="0.5" valueMin="0.45">0.45–0.50 mm</quantity>
|
||
, PW
|
||
<quantity id="EF7B9B75FFC8DD6FFB4178542C0DF8A9" box="[1159,1329,1823,1850]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.8" metricValueMin="4.5" pageId="5" pageNumber="398" unit="mm" value="0.46499999999999997" valueMax="0.48" valueMin="0.45">0.45–0.48 mm</quantity>
|
||
. Anterior margin nearly straight; lateral margins strongly convex in submedian region, slightly sinuate anteriorly and posteriorly; posterior corners nearly right-angled and blunt; posterior margin nearly straight, with marginal carina on entire length. Median longitudinal sulcus (
|
||
<figureCitation id="B0B82A15FFC8DD6FFD5878C02BCEF836" box="[670,754,1931,1957]" captionStart="FIGURES 11–14" captionStartId="6.[151,264,1604,1628]" captionTargetBox="[151,1435,345,1572]" captionTargetId="figure-101@6.[151,1435,345,1581]" captionTargetPageId="6" captionText="FIGURES 11–14. Arthromelodes tokushigei sp. n., male. Prothorax in dorsal (11), ventral (12, 13) and left lateral (14) views. Arrowheads indicate ventral foveae. Abbreviations: abg, antebasal groove; abp, antebasal pit; bst, basisternal region of prosternum; fs, furcasternum; hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; mls, median longitudinal sulcus." figureDoi="http://doi.org/10.5281/zenodo.8243468" httpUri="https://zenodo.org/record/8243468/files/figure.png" pageId="5" pageNumber="398">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC8DD6FFD3978C02A1AF836" box="[767,806,1931,1957]" italics="true" pageId="5" pageNumber="398">mls</emphasis>
|
||
) narrow and sharply marked, anteriorly obliterated just before anterior pronotal margin and posteriorly deepened to form indistinct, vestigial and asetose median antebasal fovea; lateral longitudinal sulci (
|
||
<figureCitation id="B0B82A15FFC8DD6FFDC278982B6AF87E" box="[516,598,2003,2029]" captionStart="FIGURES 11–14" captionStartId="6.[151,264,1604,1628]" captionTargetBox="[151,1435,345,1572]" captionTargetId="figure-101@6.[151,1435,345,1581]" captionTargetPageId="6" captionText="FIGURES 11–14. Arthromelodes tokushigei sp. n., male. Prothorax in dorsal (11), ventral (12, 13) and left lateral (14) views. Arrowheads indicate ventral foveae. Abbreviations: abg, antebasal groove; abp, antebasal pit; bst, basisternal region of prosternum; fs, furcasternum; hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; mls, median longitudinal sulcus." figureDoi="http://doi.org/10.5281/zenodo.8243468" httpUri="https://zenodo.org/record/8243468/files/figure.png" pageId="5" pageNumber="398">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC8DD6FFDA478982B40F87E" box="[610,636,2003,2029]" italics="true" pageId="5" pageNumber="398">lls</emphasis>
|
||
) sharply marked and each posteriorly connected with large and asetose lateral antebasal fovea (
|
||
<figureCitation id="B0B82A15FFCBDD6CFE667FDC2B1CFF22" box="[416,544,151,177]" captionStart="FIGURES 11–14" captionStartId="6.[151,264,1604,1628]" captionTargetBox="[151,1435,345,1572]" captionTargetId="figure-101@6.[151,1435,345,1581]" captionTargetPageId="6" captionText="FIGURES 11–14. Arthromelodes tokushigei sp. n., male. Prothorax in dorsal (11), ventral (12, 13) and left lateral (14) views. Arrowheads indicate ventral foveae. Abbreviations: abg, antebasal groove; abp, antebasal pit; bst, basisternal region of prosternum; fs, furcasternum; hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; mls, median longitudinal sulcus." figureDoi="http://doi.org/10.5281/zenodo.8243468" httpUri="https://zenodo.org/record/8243468/files/figure.png" pageId="6" pageNumber="399">Figs 11, 14</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFCBDD6CFDEA7FDC2B76FF22" box="[556,586,151,177]" italics="true" pageId="6" pageNumber="399">laf</emphasis>
|
||
); lateral and median antebasal foveae connected by biarcuate and anteriorly convex antebasal transverse groove (
|
||
<figureCitation id="B0B82A15FFCBDD6CFDF27FF02BBBFF46" box="[564,647,187,213]" captionStart="FIGURES 11–14" captionStartId="6.[151,264,1604,1628]" captionTargetBox="[151,1435,345,1572]" captionTargetId="figure-101@6.[151,1435,345,1581]" captionTargetPageId="6" captionText="FIGURES 11–14. Arthromelodes tokushigei sp. n., male. Prothorax in dorsal (11), ventral (12, 13) and left lateral (14) views. Arrowheads indicate ventral foveae. Abbreviations: abg, antebasal groove; abp, antebasal pit; bst, basisternal region of prosternum; fs, furcasternum; hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; mls, median longitudinal sulcus." figureDoi="http://doi.org/10.5281/zenodo.8243468" httpUri="https://zenodo.org/record/8243468/files/figure.png" pageId="6" pageNumber="399">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFCBDD6CFD557FF02B81FF46" box="[659,701,187,213]" italics="true" pageId="6" pageNumber="399">abg</emphasis>
|
||
); pronotal base with inner and outer antebasal pits (
|
||
<figureCitation id="B0B82A15FFCBDD6CFAC57FF02C6AFF46" box="[1283,1366,187,213]" captionStart="FIGURES 11–14" captionStartId="6.[151,264,1604,1628]" captionTargetBox="[151,1435,345,1572]" captionTargetId="figure-101@6.[151,1435,345,1581]" captionTargetPageId="6" captionText="FIGURES 11–14. Arthromelodes tokushigei sp. n., male. Prothorax in dorsal (11), ventral (12, 13) and left lateral (14) views. Arrowheads indicate ventral foveae. Abbreviations: abg, antebasal groove; abp, antebasal pit; bst, basisternal region of prosternum; fs, furcasternum; hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; mls, median longitudinal sulcus." figureDoi="http://doi.org/10.5281/zenodo.8243468" httpUri="https://zenodo.org/record/8243468/files/figure.png" pageId="6" pageNumber="399">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFCBDD6CFAA47FF02CB0FF46" box="[1378,1420,187,213]" italics="true" pageId="6" pageNumber="399">abp</emphasis>
|
||
), each distinct and large but shallow and with diffuse margins. Punctures on disc distinct and large but shallow and somewhat diffuse, distances between punctures subequal to their diameters; setae sparse, short and suberect. Hypomera (
|
||
<figureCitation id="B0B82A15FFCBDD6CFEDB7E6C284CFED2" box="[285,368,295,321]" captionStart="FIGURES 11–14" captionStartId="6.[151,264,1604,1628]" captionTargetBox="[151,1435,345,1572]" captionTargetId="figure-101@6.[151,1435,345,1581]" captionTargetPageId="6" captionText="FIGURES 11–14. Arthromelodes tokushigei sp. n., male. Prothorax in dorsal (11), ventral (12, 13) and left lateral (14) views. Arrowheads indicate ventral foveae. Abbreviations: abg, antebasal groove; abp, antebasal pit; bst, basisternal region of prosternum; fs, furcasternum; hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; mls, median longitudinal sulcus." figureDoi="http://doi.org/10.5281/zenodo.8243468" httpUri="https://zenodo.org/record/8243468/files/figure.png" pageId="6" pageNumber="399">Fig. 12</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFCBDD6CFEBA7E6C28AAFED2" box="[380,406,295,321]" italics="true" pageId="6" pageNumber="399">hy</emphasis>
|
||
) anteriorly setose, posteriorly asetose.
|
||
</paragraph>
|
||
<caption id="7CFC6618FFCBDD6CFF51790F2B92F95B" ID-DOI="http://doi.org/10.5281/zenodo.8243468" ID-Zenodo-Dep="8243468" httpUri="https://zenodo.org/record/8243468/files/figure.png" pageId="6" pageNumber="399" startId="6.[151,264,1604,1628]" targetBox="[151,1435,345,1572]" targetPageId="6" targetType="figure">
|
||
<paragraph id="283C3690FFCBDD6CFF51790F2B92F95B" blockId="6.[151,1437,1604,1736]" pageId="6" pageNumber="399">
|
||
<emphasis id="1AF7EA82FFCBDD6CFF51790F2864F9CF" bold="true" box="[151,344,1604,1628]" pageId="6" pageNumber="399">FIGURES 11–14.</emphasis>
|
||
<taxonomicName id="EF834D13FFCBDD6CFEA5790F2B50F9CF" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[355,620,1604,1628]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="399" phylum="Arthropoda" rank="species" species="tokushigei" status="sp. n.">
|
||
<emphasis id="1AF7EA82FFCBDD6CFEA5790F2B50F9CF" box="[355,620,1604,1628]" italics="true" pageId="6" pageNumber="399">Arthromelodes tokushigei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="01C457F9FFCBDD6CFDB1790F2B88F9C8" box="[631,692,1604,1627]" pageId="6" pageNumber="399" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male. Prothorax in dorsal (
|
||
<emphasis id="1AF7EA82FFCBDD6CFC20790F2AC2F9CF" bold="true" box="[998,1022,1604,1628]" pageId="6" pageNumber="399">11</emphasis>
|
||
), ventral (
|
||
<emphasis id="1AF7EA82FFCBDD6CFBB5790F2D8AF9CF" bold="true" box="[1139,1206,1604,1628]" pageId="6" pageNumber="399">12, 13</emphasis>
|
||
) and left lateral (
|
||
<emphasis id="1AF7EA82FFCBDD6CFABC790F2CAFF9CF" bold="true" box="[1402,1427,1604,1628]" pageId="6" pageNumber="399">14</emphasis>
|
||
) views. Arrowheads indicate ventral foveae. Abbreviations: abg, antebasal groove; abp, antebasal pit; bst, basisternal region of prosternum; fs, furcasternum; hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; mls, median longitudinal sulcus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="283C3690FFCBDD6DFF0178502C23FF6A" blockId="6.[151,1437,1818,2025]" lastBlockId="7.[151,1436,151,321]" lastPageId="7" lastPageNumber="400" pageId="6" pageNumber="399">
|
||
Prosternum (
|
||
<figureCitation id="B0B82A15FFCBDD6CFE92785128F2F8A7" box="[340,462,1818,1844]" captionStart="FIGURES 11–14" captionStartId="6.[151,264,1604,1628]" captionTargetBox="[151,1435,345,1572]" captionTargetId="figure-101@6.[151,1435,345,1581]" captionTargetPageId="6" captionText="FIGURES 11–14. Arthromelodes tokushigei sp. n., male. Prothorax in dorsal (11), ventral (12, 13) and left lateral (14) views. Arrowheads indicate ventral foveae. Abbreviations: abg, antebasal groove; abp, antebasal pit; bst, basisternal region of prosternum; fs, furcasternum; hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; mls, median longitudinal sulcus." figureDoi="http://doi.org/10.5281/zenodo.8243468" httpUri="https://zenodo.org/record/8243468/files/figure.png" pageId="6" pageNumber="399">Figs 12, 13</figureCitation>
|
||
) with basisternal (precoxal)region much longer than procoxal rests (which are composite structures combining posterior basisternal region and furcasternum situated behind profurcal invagination sites); notosternal sutures lacking; prosternum with one pair of foveae (
|
||
<figureCitation id="B0B82A15FFCBDD6CFCAF78292AD5F8EF" box="[873,1001,1890,1916]" captionStart="FIGURES 11–14" captionStartId="6.[151,264,1604,1628]" captionTargetBox="[151,1435,345,1572]" captionTargetId="figure-101@6.[151,1435,345,1581]" captionTargetPageId="6" captionText="FIGURES 11–14. Arthromelodes tokushigei sp. n., male. Prothorax in dorsal (11), ventral (12, 13) and left lateral (14) views. Arrowheads indicate ventral foveae. Abbreviations: abg, antebasal groove; abp, antebasal pit; bst, basisternal region of prosternum; fs, furcasternum; hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; mls, median longitudinal sulcus." figureDoi="http://doi.org/10.5281/zenodo.8243468" httpUri="https://zenodo.org/record/8243468/files/figure.png" pageId="6" pageNumber="399">Figs 12, 13</figureCitation>
|
||
, indicated by arrowheads) situated just in front of and slightly mesad procoxal cavity, each directed posteromesad and with asetose opening. Basisternal region posteriorly forming short and broadly subtriangular prosternal process not separating procoxae; surface of basisternal region densely covered with long and recumbent to suberect setae, especially dense in anterior half of prosternum. Hypomeral ridge (
|
||
<figureCitation id="B0B82A15FFCADD6DFE2D7FDC2B5BFF22" box="[491,615,151,177]" captionStart="FIGURES 11–14" captionStartId="6.[151,264,1604,1628]" captionTargetBox="[151,1435,345,1572]" captionTargetId="figure-101@6.[151,1435,345,1581]" captionTargetPageId="6" captionText="FIGURES 11–14. Arthromelodes tokushigei sp. n., male. Prothorax in dorsal (11), ventral (12, 13) and left lateral (14) views. Arrowheads indicate ventral foveae. Abbreviations: abg, antebasal groove; abp, antebasal pit; bst, basisternal region of prosternum; fs, furcasternum; hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; mls, median longitudinal sulcus." figureDoi="http://doi.org/10.5281/zenodo.8243468" httpUri="https://zenodo.org/record/8243468/files/figure.png" pageId="7" pageNumber="400">Figs 12, 13</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFCADD6DFDB77FDC2BAAFF22" box="[625,662,151,177]" italics="true" pageId="7" pageNumber="400">hyr</emphasis>
|
||
) developed along adcoxal margin of each hypomeron and demarcating narrow inner region of hypomeron; hypomeral groove (
|
||
<figureCitation id="B0B82A15FFCADD6DFCD37FF02A57FF46" box="[789,875,187,213]" captionStart="FIGURES 11–14" captionStartId="6.[151,264,1604,1628]" captionTargetBox="[151,1435,345,1572]" captionTargetId="figure-101@6.[151,1435,345,1581]" captionTargetPageId="6" captionText="FIGURES 11–14. Arthromelodes tokushigei sp. n., male. Prothorax in dorsal (11), ventral (12, 13) and left lateral (14) views. Arrowheads indicate ventral foveae. Abbreviations: abg, antebasal groove; abp, antebasal pit; bst, basisternal region of prosternum; fs, furcasternum; hg, hypomeral groove; hy, hypomeron; hyr, hypomeral ridge; laf, lateral fovea; lls, lateral longitudinal sulcus; mls, median longitudinal sulcus." figureDoi="http://doi.org/10.5281/zenodo.8243468" httpUri="https://zenodo.org/record/8243468/files/figure.png" pageId="7" pageNumber="400">Fig. 14</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFCADD6DFCBF7FF02AA9FF46" box="[889,917,187,213]" italics="true" pageId="7" pageNumber="400">hg</emphasis>
|
||
) present, but externally indistinct and poorly visible, in cleared specimens accentuated by line of darker cuticle marking internal strenghtening ridge.
|
||
</paragraph>
|
||
<paragraph id="283C3690FFCADD6DFF017E482B6AFED2" blockId="7.[151,1436,151,321]" pageId="7" pageNumber="400">
|
||
Mesonotum (
|
||
<figureCitation id="B0B82A15FFCADD6DFE9C7E482895FE8E" box="[346,425,258,285]" captionStart="FIGURES 15–17" captionStartId="7.[151,264,1506,1530]" captionTargetBox="[151,1435,345,1483]" captionTargetId="figure-81@7.[151,1435,345,1483]" captionTargetPageId="7" captionText="FIGURES 15–17. Arthromelodes tokushigei sp. n., male. Mesonotum and elytral base in dorsal view (15); left elytron in dorsal view (16); right elytron in dorsal view (17). Abbreviations: as, adsutural sulcus; bef, basal elytral fovea; ds, discal sulcus; eal, elytral articulating lobe; ls, lateral sulcus; phf, posthumeral fovea; sc2+scl2, fused mesoscutum and mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.8243470" httpUri="https://zenodo.org/record/8243470/files/figure.png" pageId="7" pageNumber="400">Fig. 15</figureCitation>
|
||
) strongly elongate, subtriangular, lacking scutoscutellar suture; scutellar shield not exposed, hidden under posterior pronotal margin.
|
||
</paragraph>
|
||
<caption id="7CFC6618FFCADD6DFF517AA92C17F9D0" ID-DOI="http://doi.org/10.5281/zenodo.8243470" ID-Zenodo-Dep="8243470" httpUri="https://zenodo.org/record/8243470/files/figure.png" pageId="7" pageNumber="400" startId="7.[151,264,1506,1530]" targetBox="[151,1435,345,1483]" targetPageId="7" targetType="figure">
|
||
<paragraph id="283C3690FFCADD6DFF517AA92C17F9D0" blockId="7.[151,1436,1506,1603]" pageId="7" pageNumber="400">
|
||
<emphasis id="1AF7EA82FFCADD6DFF517AA92868FA68" bold="true" box="[151,340,1506,1531]" pageId="7" pageNumber="400">FIGURES 15–17.</emphasis>
|
||
<taxonomicName id="EF834D13FFCADD6DFE9F7AA82B61FA68" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[345,605,1507,1531]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="400" phylum="Arthropoda" rank="species" species="tokushigei" status="sp. n.">
|
||
<emphasis id="1AF7EA82FFCADD6DFE9F7AA82B61FA68" box="[345,605,1507,1531]" italics="true" pageId="7" pageNumber="400">Arthromelodes tokushigei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="01C457F9FFCADD6DFDA47AA82BA6FA69" box="[610,666,1507,1530]" pageId="7" pageNumber="400" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male. Mesonotum and elytral base in dorsal view (
|
||
<emphasis id="1AF7EA82FFCADD6DFB677AA82D86FA68" bold="true" box="[1185,1210,1507,1531]" pageId="7" pageNumber="400">15</emphasis>
|
||
); left elytron in dorsal view (
|
||
<emphasis id="1AF7EA82FFCADD6DFF1E794C29CDF98C" bold="true" box="[216,241,1543,1567]" pageId="7" pageNumber="400">16</emphasis>
|
||
); right elytron in dorsal view (
|
||
<emphasis id="1AF7EA82FFCADD6DFDE8794C2B7BF98C" bold="true" box="[558,583,1543,1567]" pageId="7" pageNumber="400">17</emphasis>
|
||
). Abbreviations: as, adsutural sulcus; bef, basal elytral fovea; ds, discal sulcus; eal, elytral articulating lobe; ls, lateral sulcus; phf, posthumeral fovea; sc2+scl2, fused mesoscutum and mesoscutellum.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="283C3690FFCADD6DFF0179DE2B91F85C" blockId="7.[151,1437,1685,2035]" pageId="7" pageNumber="400">
|
||
Elytra (
|
||
<figureCitation id="B0B82A15FFCADD6DFED879DE289FF93C" box="[286,419,1685,1711]" captionStart="FIGURES 15–17" captionStartId="7.[151,264,1506,1530]" captionTargetBox="[151,1435,345,1483]" captionTargetId="figure-81@7.[151,1435,345,1483]" captionTargetPageId="7" captionText="FIGURES 15–17. Arthromelodes tokushigei sp. n., male. Mesonotum and elytral base in dorsal view (15); left elytron in dorsal view (16); right elytron in dorsal view (17). Abbreviations: as, adsutural sulcus; bef, basal elytral fovea; ds, discal sulcus; eal, elytral articulating lobe; ls, lateral sulcus; phf, posthumeral fovea; sc2+scl2, fused mesoscutum and mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.8243470" httpUri="https://zenodo.org/record/8243470/files/figure.png" pageId="7" pageNumber="400">Figs 16, 17</figureCitation>
|
||
) together transverse subtrapezoidal with strongly rounded sides, broadest near posterior third; EL
|
||
<quantity id="EF7B9B75FFCADD6DFEC279F22896F940" box="[260,426,1721,1748]" metricMagnitude="-4" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.1" metricValueMin="5.8" pageId="7" pageNumber="400" unit="mm" value="0.595" valueMax="0.61" valueMin="0.58">0.58–0.61 mm</quantity>
|
||
, EW
|
||
<quantity id="EF7B9B75FFCADD6DFE2E79F22BB3F940" box="[488,655,1721,1748]" metricMagnitude="-4" metricUnit="m" metricValue="7.15" metricValueMax="7.5" metricValueMin="6.8" pageId="7" pageNumber="400" unit="mm" value="0.715" valueMax="0.75" valueMin="0.68">0.68–0.75 mm</quantity>
|
||
. Each elytron with complete adsutural sulcus (
|
||
<figureCitation id="B0B82A15FFCADD6DFB5D79F22C22F940" box="[1179,1310,1721,1747]" captionStart="FIGURES 15–17" captionStartId="7.[151,264,1506,1530]" captionTargetBox="[151,1435,345,1483]" captionTargetId="figure-81@7.[151,1435,345,1483]" captionTargetPageId="7" captionText="FIGURES 15–17. Arthromelodes tokushigei sp. n., male. Mesonotum and elytral base in dorsal view (15); left elytron in dorsal view (16); right elytron in dorsal view (17). Abbreviations: as, adsutural sulcus; bef, basal elytral fovea; ds, discal sulcus; eal, elytral articulating lobe; ls, lateral sulcus; phf, posthumeral fovea; sc2+scl2, fused mesoscutum and mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.8243470" httpUri="https://zenodo.org/record/8243470/files/figure.png" pageId="7" pageNumber="400">Figs 16, 17</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFCADD6DFAEC79F12C7FF940" box="[1322,1347,1722,1747]" italics="true" pageId="7" pageNumber="400">as</emphasis>
|
||
), discal sulcus ((
|
||
<figureCitation id="B0B82A15FFCADD6DFF337996284FF964" box="[245,371,1757,1783]" captionStart="FIGURES 15–17" captionStartId="7.[151,264,1506,1530]" captionTargetBox="[151,1435,345,1483]" captionTargetId="figure-81@7.[151,1435,345,1483]" captionTargetPageId="7" captionText="FIGURES 15–17. Arthromelodes tokushigei sp. n., male. Mesonotum and elytral base in dorsal view (15); left elytron in dorsal view (16); right elytron in dorsal view (17). Abbreviations: as, adsutural sulcus; bef, basal elytral fovea; ds, discal sulcus; eal, elytral articulating lobe; ls, lateral sulcus; phf, posthumeral fovea; sc2+scl2, fused mesoscutum and mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.8243470" httpUri="https://zenodo.org/record/8243470/files/figure.png" pageId="7" pageNumber="400">Figs 16, 17</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFCADD6DFEBB799628AAF964" box="[381,406,1757,1783]" italics="true" pageId="7" pageNumber="400">ds</emphasis>
|
||
) developed in anterior 2/3, lateral sulcus (
|
||
<figureCitation id="B0B82A15FFCADD6DFCA079962A84F964" box="[870,952,1757,1783]" captionStart="FIGURES 15–17" captionStartId="7.[151,264,1506,1530]" captionTargetBox="[151,1435,345,1483]" captionTargetId="figure-81@7.[151,1435,345,1483]" captionTargetPageId="7" captionText="FIGURES 15–17. Arthromelodes tokushigei sp. n., male. Mesonotum and elytral base in dorsal view (15); left elytron in dorsal view (16); right elytron in dorsal view (17). Abbreviations: as, adsutural sulcus; bef, basal elytral fovea; ds, discal sulcus; eal, elytral articulating lobe; ls, lateral sulcus; phf, posthumeral fovea; sc2+scl2, fused mesoscutum and mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.8243470" httpUri="https://zenodo.org/record/8243470/files/figure.png" pageId="7" pageNumber="400">Fig. 16</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFCADD6DFC0579962AEAF964" box="[963,982,1757,1783]" italics="true" pageId="7" pageNumber="400">ls</emphasis>
|
||
) not visible in dorsal view and extending from posthumeral fovea to posterolateral elytral corner. Each elytron with three distinct asetose circular foveae: pair of basal elytral foveae (
|
||
<figureCitation id="B0B82A15FFCADD6DFE5B786E2B22F8AC" box="[413,542,1829,1855]" captionStart="FIGURES 15–17" captionStartId="7.[151,264,1506,1530]" captionTargetBox="[151,1435,345,1483]" captionTargetId="figure-81@7.[151,1435,345,1483]" captionTargetPageId="7" captionText="FIGURES 15–17. Arthromelodes tokushigei sp. n., male. Mesonotum and elytral base in dorsal view (15); left elytron in dorsal view (16); right elytron in dorsal view (17). Abbreviations: as, adsutural sulcus; bef, basal elytral fovea; ds, discal sulcus; eal, elytral articulating lobe; ls, lateral sulcus; phf, posthumeral fovea; sc2+scl2, fused mesoscutum and mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.8243470" httpUri="https://zenodo.org/record/8243470/files/figure.png" pageId="7" pageNumber="400">Figs 16, 17</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFCADD6DFDEC786E2B70F8AC" box="[554,588,1829,1855]" italics="true" pageId="7" pageNumber="400">bef</emphasis>
|
||
) situated in distinct transverse impression just behind anterior ridge of disc, and posthumeral fovea (
|
||
<figureCitation id="B0B82A15FFCADD6DFE6A78022B3DF8F0" box="[428,513,1865,1891]" captionStart="FIGURES 15–17" captionStartId="7.[151,264,1506,1530]" captionTargetBox="[151,1435,345,1483]" captionTargetId="figure-81@7.[151,1435,345,1483]" captionTargetPageId="7" captionText="FIGURES 15–17. Arthromelodes tokushigei sp. n., male. Mesonotum and elytral base in dorsal view (15); left elytron in dorsal view (16); right elytron in dorsal view (17). Abbreviations: as, adsutural sulcus; bef, basal elytral fovea; ds, discal sulcus; eal, elytral articulating lobe; ls, lateral sulcus; phf, posthumeral fovea; sc2+scl2, fused mesoscutum and mesoscutellum." figureDoi="http://doi.org/10.5281/zenodo.8243470" httpUri="https://zenodo.org/record/8243470/files/figure.png" pageId="7" pageNumber="400">Fig. 16</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFCADD6DFDC978022B0FF8F0" box="[527,563,1865,1891]" italics="true" pageId="7" pageNumber="400">phf</emphasis>
|
||
) situated on lateral elytral surface far behind humerus. Posterolateral elytral corner obtuse-angled and broadly rounded; posterior elytral margin weakly arcuate, with shallow emargination at adsutural and lateral corners. Punctures on elytra much less distinct than those on pronotal disc, superficial and inconspicuous; setae sparse, short and suberect.
|
||
</paragraph>
|
||
<paragraph id="283C3690FFCADD62FF0178922A0EFE66" blockId="7.[151,1437,1685,2035]" lastBlockId="8.[151,1437,151,501]" lastPageId="8" lastPageNumber="401" pageId="7" pageNumber="400">
|
||
Mesoventrite (
|
||
<figureCitation id="B0B82A15FFCADD6DFEA9789228FBF860" box="[367,455,2009,2035]" captionStart="FIGURES 18–20" captionStartId="8.[151,264,1680,1704]" captionTargetBox="[152,1429,525,1657]" captionTargetId="figure-164@8.[151,1436,525,1657]" captionTargetPageId="8" captionText="FIGURES 18–20. Arthromelodes tokushigei sp. n., male. Pterothorax in ventral view (18, 19); anterior region of mesoventrite in ventral view (20). Arrowheads indicate ventral foveae. Abbreviations: amtvp, anterior metaventral process; msmtb, mesometathoracic border; msvp, mesoventral intermesocoxal process; mtvp, metaventral intermetacoxal process; ppc, prepectus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.8243472" httpUri="https://zenodo.org/record/8243472/files/figure.png" pageId="7" pageNumber="400">Fig. 18</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFCADD6DFE1E789228CEF860" box="[472,498,2009,2035]" italics="true" pageId="7" pageNumber="400">
|
||
v
|
||
<figureCitation id="B0B82A15FFCADD6DFE22789228CEF860" box="[484,498,2009,2035]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1936,1960]" captionTargetBox="[157,1430,600,1904]" captionTargetId="figure-205@2.[151,1436,594,1910]" captionTargetPageId="2" captionText="FIGURES 1–2. Arthromelodes tokushigei sp. n., male. Dorsal habitus of male (1) and female (2)." figureDoi="http://doi.org/10.5281/zenodo.8243462" httpUri="https://zenodo.org/record/8243462/files/figure.png" pageId="7" pageNumber="400">2</figureCitation>
|
||
</emphasis>
|
||
) transverse, with lateral regions fully demarcated from metaventrite (i.e, mesometaventral border clearly marked;
|
||
<figureCitation id="B0B82A15FFC5DD62FDE87FDC2BBEFF22" box="[558,642,151,177]" captionStart="FIGURES 18–20" captionStartId="8.[151,264,1680,1704]" captionTargetBox="[152,1429,525,1657]" captionTargetId="figure-164@8.[151,1436,525,1657]" captionTargetPageId="8" captionText="FIGURES 18–20. Arthromelodes tokushigei sp. n., male. Pterothorax in ventral view (18, 19); anterior region of mesoventrite in ventral view (20). Arrowheads indicate ventral foveae. Abbreviations: amtvp, anterior metaventral process; msmtb, mesometathoracic border; msvp, mesoventral intermesocoxal process; mtvp, metaventral intermetacoxal process; ppc, prepectus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.8243472" httpUri="https://zenodo.org/record/8243472/files/figure.png" pageId="8" pageNumber="401">Fig. 19</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC5DD62FD497FDC2BE4FF22" box="[655,728,151,177]" italics="true" pageId="8" pageNumber="401">msmtb</emphasis>
|
||
). Mesanepisterna and anterior region of mesoventrite forming massive prepectus (
|
||
<figureCitation id="B0B82A15FFC5DD62FEB37FF028F5FF46" box="[373,457,187,213]" captionStart="FIGURES 18–20" captionStartId="8.[151,264,1680,1704]" captionTargetBox="[152,1429,525,1657]" captionTargetId="figure-164@8.[151,1436,525,1657]" captionTargetPageId="8" captionText="FIGURES 18–20. Arthromelodes tokushigei sp. n., male. Pterothorax in ventral view (18, 19); anterior region of mesoventrite in ventral view (20). Arrowheads indicate ventral foveae. Abbreviations: amtvp, anterior metaventral process; msmtb, mesometathoracic border; msvp, mesoventral intermesocoxal process; mtvp, metaventral intermetacoxal process; ppc, prepectus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.8243472" httpUri="https://zenodo.org/record/8243472/files/figure.png" pageId="8" pageNumber="401">Fig. 19</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC5DD62FE107FF028C2FF47" box="[470,510,187,212]" italics="true" pageId="8" pageNumber="401">ppc</emphasis>
|
||
), posteriorly mesoventrite strongly and abruply broadeneing, with lateral margins strongly diverging posterad. Mesoventral process (
|
||
<figureCitation id="B0B82A15FFC5DD62FD097F942A1FFF6A" box="[719,803,223,249]" captionStart="FIGURES 18–20" captionStartId="8.[151,264,1680,1704]" captionTargetBox="[152,1429,525,1657]" captionTargetId="figure-164@8.[151,1436,525,1657]" captionTargetPageId="8" captionText="FIGURES 18–20. Arthromelodes tokushigei sp. n., male. Pterothorax in ventral view (18, 19); anterior region of mesoventrite in ventral view (20). Arrowheads indicate ventral foveae. Abbreviations: amtvp, anterior metaventral process; msmtb, mesometathoracic border; msvp, mesoventral intermesocoxal process; mtvp, metaventral intermetacoxal process; ppc, prepectus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.8243472" httpUri="https://zenodo.org/record/8243472/files/figure.png" pageId="8" pageNumber="401">Fig. 18</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC5DD62FCE97F942A55FF6B" box="[815,873,223,248]" italics="true" pageId="8" pageNumber="401">msvp</emphasis>
|
||
) situated between anterior margins of mesocoxae, broadly subtriangular and posteriorly widely separated from anterior metaventral process. Prepectus covered with transverse microreticulation; lateral and posteromedian regions of mesoventrite smooth, sides asetose, median region covered with sparse suberect setae. Three pairs of foveae are situated on mesoventrite (indicated by arrowheads in
|
||
<figureCitation id="B0B82A15FFC5DD62FF7E7E242802FE1A" box="[184,318,367,393]" captionStart="FIGURES 18–20" captionStartId="8.[151,264,1680,1704]" captionTargetBox="[152,1429,525,1657]" captionTargetId="figure-164@8.[151,1436,525,1657]" captionTargetPageId="8" captionText="FIGURES 18–20. Arthromelodes tokushigei sp. n., male. Pterothorax in ventral view (18, 19); anterior region of mesoventrite in ventral view (20). Arrowheads indicate ventral foveae. Abbreviations: amtvp, anterior metaventral process; msmtb, mesometathoracic border; msvp, mesoventral intermesocoxal process; mtvp, metaventral intermetacoxal process; ppc, prepectus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.8243472" httpUri="https://zenodo.org/record/8243472/files/figure.png" pageId="8" pageNumber="401">Figs 18–20</figureCitation>
|
||
): one pair ventrally and sublaterally just behind median transverse region of prepectus, directed anterolaterad; one pair ventrally and laterally between lateral region of prepectus and lateral asetose region of mesoventrite, directed anteromesad; and one pair sharing opening with the latter, but much broader and directed mesad. Opeinings of all mesoventral foveae densely setose.
|
||
</paragraph>
|
||
<caption id="7CFC6618FFC5DD62FF5179DB28EFF887" ID-DOI="http://doi.org/10.5281/zenodo.8243472" ID-Zenodo-Dep="8243472" httpUri="https://zenodo.org/record/8243472/files/figure.png" pageId="8" pageNumber="401" startId="8.[151,264,1680,1704]" targetBox="[152,1429,525,1657]" targetPageId="8" targetType="figure">
|
||
<paragraph id="283C3690FFC5DD62FF5179DB28EFF887" blockId="8.[151,1436,1680,1812]" pageId="8" pageNumber="401">
|
||
<emphasis id="1AF7EA82FFC5DD62FF5179DB2868F93B" bold="true" box="[151,340,1680,1704]" pageId="8" pageNumber="401">FIGURES 18–20.</emphasis>
|
||
<taxonomicName id="EF834D13FFC5DD62FE9D79DB2B63F93B" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[347,607,1680,1704]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="401" phylum="Arthropoda" rank="species" species="tokushigei" status="sp. n.">
|
||
<emphasis id="1AF7EA82FFC5DD62FE9D79DB2B63F93B" box="[347,607,1680,1704]" italics="true" pageId="8" pageNumber="401">Arthromelodes tokushigei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="01C457F9FFC5DD62FDA079DB2BA2F934" box="[614,670,1680,1703]" pageId="8" pageNumber="401" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male. Pterothorax in ventral view (
|
||
<emphasis id="1AF7EA82FFC5DD62FBCD79DB2D77F93B" bold="true" box="[1035,1099,1680,1704]" pageId="8" pageNumber="401">18, 19</emphasis>
|
||
); anterior region of mesoventrite in ventral view (
|
||
<emphasis id="1AF7EA82FFC5DD62FE8579FF2860F95F" bold="true" box="[323,348,1716,1740]" pageId="8" pageNumber="401">20</emphasis>
|
||
). Arrowheads indicate ventral foveae. Abbreviations: amtvp, anterior metaventral process; msmtb, mesometathoracic border; msvp, mesoventral intermesocoxal process; mtvp, metaventral intermetacoxal process; ppc, prepectus; v2, mesoventrite; v3, metaventrite.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="283C3690FFC5DD62FF01782C28A4F837" blockId="8.[151,1437,1894,2029]" pageId="8" pageNumber="401">Metanotum (not shown) strongly shortened and largely membranous, lacking alacristae and apodemes; hind wings entirely lacking.</paragraph>
|
||
<paragraph id="283C3690FFC5DD63FF0178E52C65FEF6" blockId="8.[151,1437,1894,2029]" lastBlockId="9.[151,1437,151,357]" lastPageId="9" lastPageNumber="402" pageId="8" pageNumber="401">
|
||
Metaventrite (
|
||
<figureCitation id="B0B82A15FFC5DD62FEA378E52885F85B" box="[357,441,1966,1992]" captionStart="FIGURES 18–20" captionStartId="8.[151,264,1680,1704]" captionTargetBox="[152,1429,525,1657]" captionTargetId="figure-164@8.[151,1436,525,1657]" captionTargetPageId="8" captionText="FIGURES 18–20. Arthromelodes tokushigei sp. n., male. Pterothorax in ventral view (18, 19); anterior region of mesoventrite in ventral view (20). Arrowheads indicate ventral foveae. Abbreviations: amtvp, anterior metaventral process; msmtb, mesometathoracic border; msvp, mesoventral intermesocoxal process; mtvp, metaventral intermetacoxal process; ppc, prepectus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.8243472" httpUri="https://zenodo.org/record/8243472/files/figure.png" pageId="8" pageNumber="401">Fig. 18</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC5DD62FE0178E528DDF85B" box="[455,481,1966,1992]" italics="true" pageId="8" pageNumber="401">
|
||
v
|
||
<figureCitation id="B0B82A15FFC5DD62FE1578E528DDF85B" box="[467,481,1966,1992]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[151,1436,181,1804]" captionTargetId="figure-15@3.[151,1436,181,1804]" captionTargetPageId="3" captionText="FIGURES 3–6. Arthromelodes tokushigei sp. n., male. Head in dorsal (3), ventral (4) and lateral (5) views; right antenna in dorsal view (6). Abbreviations: cl, clypeus; dtt, dorsal tentorial pit; fr, frons; gen, gena; gp, gular plate; gs, gular suture; nr, neck region; oc, occipital carina; occ, occipital constriction; omc, ocular-mandibular carina; pgen, postgena; ptp, posterior tentorial pit; smc, submental carina; smn, submentum; tm, temple; vc, vertexal carina; vs, vertexal sulcus; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.8243464" httpUri="https://zenodo.org/record/8243464/files/figure.png" pageId="8" pageNumber="401">3</figureCitation>
|
||
</emphasis>
|
||
) about twice as wide as long, weakly impressed at middle and convex at sides, with broadly subtriangular anterior metaventral (intermesocoxal) process (
|
||
<figureCitation id="B0B82A15FFC5DD62FC6D78992D0DF87F" box="[939,1073,2002,2028]" captionStart="FIGURES 18–20" captionStartId="8.[151,264,1680,1704]" captionTargetBox="[152,1429,525,1657]" captionTargetId="figure-164@8.[151,1436,525,1657]" captionTargetPageId="8" captionText="FIGURES 18–20. Arthromelodes tokushigei sp. n., male. Pterothorax in ventral view (18, 19); anterior region of mesoventrite in ventral view (20). Arrowheads indicate ventral foveae. Abbreviations: amtvp, anterior metaventral process; msmtb, mesometathoracic border; msvp, mesoventral intermesocoxal process; mtvp, metaventral intermetacoxal process; ppc, prepectus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.8243472" httpUri="https://zenodo.org/record/8243472/files/figure.png" pageId="8" pageNumber="401">Figs 18, 19</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC5DD62FB8778982DB9F87F" box="[1089,1157,2003,2028]" italics="true" pageId="8" pageNumber="401">amtvp</emphasis>
|
||
). Metaventrite with two pairs of foveae (indicated by arrowheads in
|
||
<figureCitation id="B0B82A15FFC4DD63FDB97FDC2BC2FF22" box="[639,766,151,177]" captionStart="FIGURES 18–20" captionStartId="8.[151,264,1680,1704]" captionTargetBox="[152,1429,525,1657]" captionTargetId="figure-164@8.[151,1436,525,1657]" captionTargetPageId="8" captionText="FIGURES 18–20. Arthromelodes tokushigei sp. n., male. Pterothorax in ventral view (18, 19); anterior region of mesoventrite in ventral view (20). Arrowheads indicate ventral foveae. Abbreviations: amtvp, anterior metaventral process; msmtb, mesometathoracic border; msvp, mesoventral intermesocoxal process; mtvp, metaventral intermetacoxal process; ppc, prepectus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.8243472" httpUri="https://zenodo.org/record/8243472/files/figure.png" pageId="9" pageNumber="402">Figs 18, 19</figureCitation>
|
||
): just behind and laterad mesocoxal cavities, directed masad and shallow, developed rather as deep impressions than typical foveal cuticular invaginations; and just behind middle of each mesocoxal rest, directed anterodorsad; openings of all metaventral foveae densely setose, those of posterior foveae situated in circular impressions much larger than diameter of fovea. Metaventral intermetacoxal process (
|
||
<figureCitation id="B0B82A15FFC4DD63FF597E6C29CDFED2" box="[159,241,295,321]" captionStart="FIGURES 18–20" captionStartId="8.[151,264,1680,1704]" captionTargetBox="[152,1429,525,1657]" captionTargetId="figure-164@8.[151,1436,525,1657]" captionTargetPageId="8" captionText="FIGURES 18–20. Arthromelodes tokushigei sp. n., male. Pterothorax in ventral view (18, 19); anterior region of mesoventrite in ventral view (20). Arrowheads indicate ventral foveae. Abbreviations: amtvp, anterior metaventral process; msmtb, mesometathoracic border; msvp, mesoventral intermesocoxal process; mtvp, metaventral intermetacoxal process; ppc, prepectus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.8243472" httpUri="https://zenodo.org/record/8243472/files/figure.png" pageId="9" pageNumber="402">Fig. 18</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC4DD63FF387E6C2808FED3" box="[254,308,295,320]" italics="true" pageId="9" pageNumber="402">mtvp</emphasis>
|
||
) broadly subtriangular, divided medially by narrow and long emargination into rounded lateral lobes. Metaventrite virtually impunctate, covered with sparse and short setae denser on median region than on sides.
|
||
</paragraph>
|
||
<caption id="7CFC6618FFC4DD63FF517A4B2D4AFAAE" ID-DOI="http://doi.org/10.5281/zenodo.8243474" ID-Zenodo-Dep="8243474" httpUri="https://zenodo.org/record/8243474/files/figure.png" pageId="9" pageNumber="402" startId="9.[151,264,1280,1304]" targetBox="[155,1435,381,1257]" targetPageId="9" targetType="figure">
|
||
<paragraph id="283C3690FFC4DD63FF517A4B2D4AFAAE" blockId="9.[151,1436,1280,1341]" pageId="9" pageNumber="402">
|
||
<emphasis id="1AF7EA82FFC4DD63FF517A4B2868FA8B" bold="true" box="[151,340,1280,1304]" pageId="9" pageNumber="402">FIGURES 21–26.</emphasis>
|
||
<taxonomicName id="EF834D13FFC4DD63FE9D7A4A2B63FA8A" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[347,607,1281,1305]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="402" phylum="Arthropoda" rank="species" species="tokushigei" status="sp. n.">
|
||
<emphasis id="1AF7EA82FFC4DD63FE9D7A4A2B63FA8A" box="[347,607,1281,1305]" italics="true" pageId="9" pageNumber="402">Arthromelodes tokushigei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="01C457F9FFC4DD63FDA37A4A2BA1FA8B" box="[613,669,1281,1304]" pageId="9" pageNumber="402" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male. Fore (
|
||
<emphasis id="1AF7EA82FFC4DD63FCE27A4B2A01FA8B" bold="true" box="[804,829,1280,1304]" pageId="9" pageNumber="402">21</emphasis>
|
||
), middle (
|
||
<emphasis id="1AF7EA82FFC4DD63FC6E7A4B2AFDFA8B" bold="true" box="[936,961,1280,1304]" pageId="9" pageNumber="402">22</emphasis>
|
||
) and hind (
|
||
<emphasis id="1AF7EA82FFC4DD63FBFE7A4B2D6DFA8B" bold="true" box="[1080,1105,1280,1304]" pageId="9" pageNumber="402">23</emphasis>
|
||
) leg in posterior view; procoxae in ventral view (
|
||
<emphasis id="1AF7EA82FFC4DD63FEF87A6F286BFAAF" bold="true" box="[318,343,1316,1340]" pageId="9" pageNumber="402">24</emphasis>
|
||
); procoxal glandular opening (
|
||
<emphasis id="1AF7EA82FFC4DD63FD547A6F2B97FAAF" bold="true" box="[658,683,1316,1340]" pageId="9" pageNumber="402">25</emphasis>
|
||
); ventral field of carinulae on profemur (
|
||
<emphasis id="1AF7EA82FFC4DD63FB887A6F2D5BFAAF" bold="true" box="[1102,1127,1316,1340]" pageId="9" pageNumber="402">26</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="283C3690FFC4DD63FF017AC42DCEF936" blockId="9.[151,1437,1423,2025]" pageId="9" pageNumber="402">
|
||
Legs (
|
||
<figureCitation id="B0B82A15FFC4DD63FECD7AC428B5FA3A" box="[267,393,1423,1449]" captionStart="FIGURES 21–26" captionStartId="9.[151,264,1280,1304]" captionTargetBox="[155,1435,381,1257]" captionTargetId="figure-112@9.[151,1435,381,1257]" captionTargetPageId="9" captionText="FIGURES 21–26. Arthromelodes tokushigei sp. n., male. Fore (21), middle (22) and hind (23) leg in posterior view; procoxae in ventral view (24); procoxal glandular opening (25); ventral field of carinulae on profemur (26)." figureDoi="http://doi.org/10.5281/zenodo.8243474" httpUri="https://zenodo.org/record/8243474/files/figure.png" pageId="9" pageNumber="402">Figs 21–26</figureCitation>
|
||
) long and slender. Fore legs (
|
||
<figureCitation id="B0B82A15FFC4DD63FD0F7AC42A24FA3A" box="[713,792,1423,1449]" captionStart="FIGURES 21–26" captionStartId="9.[151,264,1280,1304]" captionTargetBox="[155,1435,381,1257]" captionTargetId="figure-112@9.[151,1435,381,1257]" captionTargetPageId="9" captionText="FIGURES 21–26. Arthromelodes tokushigei sp. n., male. Fore (21), middle (22) and hind (23) leg in posterior view; procoxae in ventral view (24); procoxal glandular opening (25); ventral field of carinulae on profemur (26)." figureDoi="http://doi.org/10.5281/zenodo.8243474" httpUri="https://zenodo.org/record/8243474/files/figure.png" pageId="9" pageNumber="402">Fig. 21</figureCitation>
|
||
) with subconical coxa proximally prolonged by trochantin fused with cryptopleuron (not visible in intact specimens), each procoxa with glandular or sensory asetose opening in distal region of anterior surface (
|
||
<figureCitation id="B0B82A15FFC4DD63FDFE7A9C2BFDFA61" box="[568,705,1495,1522]" captionStart="FIGURES 21–26" captionStartId="9.[151,264,1280,1304]" captionTargetBox="[155,1435,381,1257]" captionTargetId="figure-112@9.[151,1435,381,1257]" captionTargetPageId="9" captionText="FIGURES 21–26. Arthromelodes tokushigei sp. n., male. Fore (21), middle (22) and hind (23) leg in posterior view; procoxae in ventral view (24); procoxal glandular opening (25); ventral field of carinulae on profemur (26)." figureDoi="http://doi.org/10.5281/zenodo.8243474" httpUri="https://zenodo.org/record/8243474/files/figure.png" pageId="9" pageNumber="402">Figs 24, 25</figureCitation>
|
||
); trochanter small, elongate subtriangular with rounded ventral margin; femur clavate, in proximal region of ventral surface with elongate field of transverse irregular carinulae (
|
||
<figureCitation id="B0B82A15FFC4DD63FF59795429C9F9AA" box="[159,245,1567,1593]" captionStart="FIGURES 21–26" captionStartId="9.[151,264,1280,1304]" captionTargetBox="[155,1435,381,1257]" captionTargetId="figure-112@9.[151,1435,381,1257]" captionTargetPageId="9" captionText="FIGURES 21–26. Arthromelodes tokushigei sp. n., male. Fore (21), middle (22) and hind (23) leg in posterior view; procoxae in ventral view (24); procoxal glandular opening (25); ventral field of carinulae on profemur (26)." figureDoi="http://doi.org/10.5281/zenodo.8243474" httpUri="https://zenodo.org/record/8243474/files/figure.png" pageId="9" pageNumber="402">Fig. 26</figureCitation>
|
||
); tibia straight, broadening from base to around distal 1/3 and then tapering towards apex; tarsus slightly longer than half length of tibia, with tarsomere 3 indistinctly longer than 2. Middle leg (
|
||
<figureCitation id="B0B82A15FFC4DD63FBB479082DF8F9CE" box="[1138,1220,1603,1629]" captionStart="FIGURES 21–26" captionStartId="9.[151,264,1280,1304]" captionTargetBox="[155,1435,381,1257]" captionTargetId="figure-112@9.[151,1435,381,1257]" captionTargetPageId="9" captionText="FIGURES 21–26. Arthromelodes tokushigei sp. n., male. Fore (21), middle (22) and hind (23) leg in posterior view; procoxae in ventral view (24); procoxal glandular opening (25); ventral field of carinulae on profemur (26)." figureDoi="http://doi.org/10.5281/zenodo.8243474" httpUri="https://zenodo.org/record/8243474/files/figure.png" pageId="9" pageNumber="402">Fig. 22</figureCitation>
|
||
) similar to fore leg but femur ventrally smooth and tibia with slender ventral apical mucro. Hind leg (
|
||
<figureCitation id="B0B82A15FFC4DD63FBDB792C2D51F912" box="[1053,1133,1639,1665]" captionStart="FIGURES 21–26" captionStartId="9.[151,264,1280,1304]" captionTargetBox="[155,1435,381,1257]" captionTargetId="figure-112@9.[151,1435,381,1257]" captionTargetPageId="9" captionText="FIGURES 21–26. Arthromelodes tokushigei sp. n., male. Fore (21), middle (22) and hind (23) leg in posterior view; procoxae in ventral view (24); procoxal glandular opening (25); ventral field of carinulae on profemur (26)." figureDoi="http://doi.org/10.5281/zenodo.8243474" httpUri="https://zenodo.org/record/8243474/files/figure.png" pageId="9" pageNumber="402">Fig. 23</figureCitation>
|
||
) longest and similar to fore and middle leg except for strongly transverse and subtriangular coxa and lack of any modifications.
|
||
</paragraph>
|
||
<paragraph id="283C3690FFC4DD60FF0179E42D17FE42" blockId="9.[151,1437,1423,2025]" lastBlockId="10.[151,1437,151,465]" lastPageId="10" lastPageNumber="403" pageId="9" pageNumber="402">
|
||
Abdomen (
|
||
<figureCitation id="B0B82A15FFC4DD63FE8F79E428F2F95A" box="[329,462,1711,1737]" captionStart-0="FIGURES 27–30" captionStart-1="FIGURES 31–36" captionStart-2="FIGURES 37–42" captionStartId-0="10.[151,261,1754,1778]" captionStartId-1="11.[151,264,1693,1717]" captionStartId-2="12.[151,264,1376,1400]" captionTargetBox-0="[151,1417,489,1721]" captionTargetBox-1="[151,1435,237,1670]" captionTargetBox-2="[157,1429,187,1346]" captionTargetId-0="figure-172@10.[151,1435,489,1731]" captionTargetId-1="figure-36@11.[151,1435,237,1670]" captionTargetId-2="figure-21@12.[151,1436,181,1352]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURES 27–30. Arthromelodes tokushigei sp. n., male.Abdomen in dorsal view (27); abdominal segments III–IV in dorsal (with ventral structures visible) (28) and left lateral (29) views; tergite V in dorsal view (30). Arrowheads indicate foveae (d, dorsal; v, ventral).Abbreviations: imcp, intermetacoxal process; lsp, lateral setal patch; mt, median tubercle; tIII, tergite III; tIV, tergite IV." captionText-1="FIGURES 31–36. Arthromelodes tokushigei sp. n., male. Lateral fovea on abdominal tergite IV (31); median impression on tergite IV (32); posterior region of median impression on tergite IV (33); anterior region of impression on tergite IV (34); median process of median impression on tergite IV (35); spatulate setae on lateral setal patch of tergite IV (36). Abbreviations: lp, lateral process; mp, median process; mt, median tubercle; smp, submedian process." captionText-2="FIGURES 37–42. Arthromelodes tokushigei sp. n., male (37–39) and female (40–42). Tergite VIII in dorsal view (37, 40); sternite VIII in ventral view (38, 42); sternite IX in ventral view, with internal membrane (39); postabdominal sclerites (41)." figureDoi-0="http://doi.org/10.5281/zenodo.8243476" figureDoi-1="http://doi.org/10.5281/zenodo.8243478" figureDoi-2="http://doi.org/10.5281/zenodo.8243480" httpUri-0="https://zenodo.org/record/8243476/files/figure.png" httpUri-1="https://zenodo.org/record/8243478/files/figure.png" httpUri-2="https://zenodo.org/record/8243480/files/figure.png" pageId="9" pageNumber="402">Figs 27–39</figureCitation>
|
||
) equal to or slightly longer than elytra, AbL
|
||
<quantity id="EF7B9B75FFC4DD63FC2B79E42DABF95A" box="[1005,1175,1711,1738]" metricMagnitude="-4" metricUnit="m" metricValue="6.05" metricValueMax="6.8" metricValueMin="5.3" pageId="9" pageNumber="402" unit="mm" value="0.605" valueMax="0.68" valueMin="0.53">0.53–0.68 mm</quantity>
|
||
, AbW
|
||
<quantity id="EF7B9B75FFC4DD63FB2879E42CA5F95A" box="[1262,1433,1711,1738]" metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="7.5" metricValueMin="6.0" pageId="9" pageNumber="402" unit="mm" value="0.675" valueMax="0.75" valueMin="0.6">0.60–0.75 mm</quantity>
|
||
. Tergite IV (
|
||
<figureCitation id="B0B82A15FFC4DD63FEDA799828A2F97E" box="[284,414,1747,1773]" captionStart="FIGURES 27–30" captionStartId="10.[151,261,1754,1778]" captionTargetBox="[151,1417,489,1721]" captionTargetId="figure-172@10.[151,1435,489,1731]" captionTargetPageId="10" captionText="FIGURES 27–30. Arthromelodes tokushigei sp. n., male.Abdomen in dorsal view (27); abdominal segments III–IV in dorsal (with ventral structures visible) (28) and left lateral (29) views; tergite V in dorsal view (30). Arrowheads indicate foveae (d, dorsal; v, ventral).Abbreviations: imcp, intermetacoxal process; lsp, lateral setal patch; mt, median tubercle; tIII, tergite III; tIV, tergite IV." figureDoi="http://doi.org/10.5281/zenodo.8243476" httpUri="https://zenodo.org/record/8243476/files/figure.png" pageId="9" pageNumber="402">Figs 27, 28</figureCitation>
|
||
) broadly subtrapezoidal, weakly narrowing posterad, convex in anterior half and strongly modified posteriorly; base with two pairs of foveae (
|
||
<figureCitation id="B0B82A15FFC4DD63FD2979BC2A7FF882" box="[751,835,1783,1809]" captionStart="FIGURES 27–30" captionStartId="10.[151,261,1754,1778]" captionTargetBox="[151,1417,489,1721]" captionTargetId="figure-172@10.[151,1435,489,1731]" captionTargetPageId="10" captionText="FIGURES 27–30. Arthromelodes tokushigei sp. n., male.Abdomen in dorsal view (27); abdominal segments III–IV in dorsal (with ventral structures visible) (28) and left lateral (29) views; tergite V in dorsal view (30). Arrowheads indicate foveae (d, dorsal; v, ventral).Abbreviations: imcp, intermetacoxal process; lsp, lateral setal patch; mt, median tubercle; tIII, tergite III; tIV, tergite IV." figureDoi="http://doi.org/10.5281/zenodo.8243476" httpUri="https://zenodo.org/record/8243476/files/figure.png" pageId="9" pageNumber="402">Fig. 28</figureCitation>
|
||
, indicated with arrowheads marked as “d”): one pair of shallow sublateral foveae directed mesad, with large setose openings (
|
||
<figureCitation id="B0B82A15FFC4DD63FC2878502D78F8A6" box="[1006,1092,1819,1845]" captionStart="FIGURES 31–36" captionStartId="11.[151,264,1693,1717]" captionTargetBox="[151,1435,237,1670]" captionTargetId="figure-36@11.[151,1435,237,1670]" captionTargetPageId="11" captionText="FIGURES 31–36. Arthromelodes tokushigei sp. n., male. Lateral fovea on abdominal tergite IV (31); median impression on tergite IV (32); posterior region of median impression on tergite IV (33); anterior region of impression on tergite IV (34); median process of median impression on tergite IV (35); spatulate setae on lateral setal patch of tergite IV (36). Abbreviations: lp, lateral process; mp, median process; mt, median tubercle; smp, submedian process." figureDoi="http://doi.org/10.5281/zenodo.8243478" httpUri="https://zenodo.org/record/8243478/files/figure.png" pageId="9" pageNumber="402">Fig. 31</figureCitation>
|
||
), and with median transverse impression filled with setae (
|
||
<figureCitation id="B0B82A15FFC4DD63FE1878742B0CF8CA" box="[478,560,1855,1881]" captionStart="FIGURES 31–36" captionStartId="11.[151,264,1693,1717]" captionTargetBox="[151,1435,237,1670]" captionTargetId="figure-36@11.[151,1435,237,1670]" captionTargetPageId="11" captionText="FIGURES 31–36. Arthromelodes tokushigei sp. n., male. Lateral fovea on abdominal tergite IV (31); median impression on tergite IV (32); posterior region of median impression on tergite IV (33); anterior region of impression on tergite IV (34); median process of median impression on tergite IV (35); spatulate setae on lateral setal patch of tergite IV (36). Abbreviations: lp, lateral process; mp, median process; mt, median tubercle; smp, submedian process." figureDoi="http://doi.org/10.5281/zenodo.8243478" httpUri="https://zenodo.org/record/8243478/files/figure.png" pageId="9" pageNumber="402">Fig. 32</figureCitation>
|
||
) and flanked by pair of submedian foveae directed laterad, but lacking lateral discal carinae. Posterior half of tergite IV with large transverse impression subdivided into small anterior and much larger posterior cavities, the latter with large oval posteromedian tubercle (
|
||
<figureCitation id="B0B82A15FFC4DD63FC1C78CC2D17F832" box="[986,1067,1927,1953]" captionStart="FIGURES 31–36" captionStartId="11.[151,264,1693,1717]" captionTargetBox="[151,1435,237,1670]" captionTargetId="figure-36@11.[151,1435,237,1670]" captionTargetPageId="11" captionText="FIGURES 31–36. Arthromelodes tokushigei sp. n., male. Lateral fovea on abdominal tergite IV (31); median impression on tergite IV (32); posterior region of median impression on tergite IV (33); anterior region of impression on tergite IV (34); median process of median impression on tergite IV (35); spatulate setae on lateral setal patch of tergite IV (36). Abbreviations: lp, lateral process; mp, median process; mt, median tubercle; smp, submedian process." figureDoi="http://doi.org/10.5281/zenodo.8243478" httpUri="https://zenodo.org/record/8243478/files/figure.png" pageId="9" pageNumber="402">Fig. 33</figureCitation>
|
||
) sparsely covered with setae and surrounded laterally and anteriorly by asetose and smooth concave integument. In front of posteromedian tubercle, in small anterior part of impression, there are three pairs of slender cuticular processes: lateral process (
|
||
<figureCitation id="B0B82A15FFC4DD63FA8478842CABF87A" box="[1346,1431,1999,2025]" captionStart="FIGURES 31–36" captionStartId="11.[151,264,1693,1717]" captionTargetBox="[151,1435,237,1670]" captionTargetId="figure-36@11.[151,1435,237,1670]" captionTargetPageId="11" captionText="FIGURES 31–36. Arthromelodes tokushigei sp. n., male. Lateral fovea on abdominal tergite IV (31); median impression on tergite IV (32); posterior region of median impression on tergite IV (33); anterior region of impression on tergite IV (34); median process of median impression on tergite IV (35); spatulate setae on lateral setal patch of tergite IV (36). Abbreviations: lp, lateral process; mp, median process; mt, median tubercle; smp, submedian process." figureDoi="http://doi.org/10.5281/zenodo.8243478" httpUri="https://zenodo.org/record/8243478/files/figure.png" pageId="9" pageNumber="402">Fig. 34</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC7DD60FF517FDC2991FF22" box="[151,173,151,177]" italics="true" pageId="10" pageNumber="403">lp</emphasis>
|
||
), submedian process (
|
||
<figureCitation id="B0B82A15FFC7DD60FE6E7FDC28C7FF22" box="[424,507,151,177]" captionStart="FIGURES 31–36" captionStartId="11.[151,264,1693,1717]" captionTargetBox="[151,1435,237,1670]" captionTargetId="figure-36@11.[151,1435,237,1670]" captionTargetPageId="11" captionText="FIGURES 31–36. Arthromelodes tokushigei sp. n., male. Lateral fovea on abdominal tergite IV (31); median impression on tergite IV (32); posterior region of median impression on tergite IV (33); anterior region of impression on tergite IV (34); median process of median impression on tergite IV (35); spatulate setae on lateral setal patch of tergite IV (36). Abbreviations: lp, lateral process; mp, median process; mt, median tubercle; smp, submedian process." figureDoi="http://doi.org/10.5281/zenodo.8243478" httpUri="https://zenodo.org/record/8243478/files/figure.png" pageId="10" pageNumber="403">Fig. 34</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC7DD60FDC17FDC2B08FF23" box="[519,564,151,176]" italics="true" pageId="10" pageNumber="403">smp</emphasis>
|
||
), and median process (
|
||
<figureCitation id="B0B82A15FFC7DD60FCFE7FDC2AB7FF22" box="[824,907,151,177]" captionStart="FIGURES 31–36" captionStartId="11.[151,264,1693,1717]" captionTargetBox="[151,1435,237,1670]" captionTargetId="figure-36@11.[151,1435,237,1670]" captionTargetPageId="11" captionText="FIGURES 31–36. Arthromelodes tokushigei sp. n., male. Lateral fovea on abdominal tergite IV (31); median impression on tergite IV (32); posterior region of median impression on tergite IV (33); anterior region of impression on tergite IV (34); median process of median impression on tergite IV (35); spatulate setae on lateral setal patch of tergite IV (36). Abbreviations: lp, lateral process; mp, median process; mt, median tubercle; smp, submedian process." figureDoi="http://doi.org/10.5281/zenodo.8243478" httpUri="https://zenodo.org/record/8243478/files/figure.png" pageId="10" pageNumber="403">Fig. 34</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC7DD60FC517FDC2A85FF23" box="[919,953,151,176]" italics="true" pageId="10" pageNumber="403">mp</emphasis>
|
||
). Lateral and median processes bear apical setae, each median process bears subapical and directed posteriorly penicillus composed of a few short setae (
|
||
<figureCitation id="B0B82A15FFC7DD60FAA97FF0298AFF6A" captionStart="FIGURES 31–36" captionStartId="11.[151,264,1693,1717]" captionTargetBox="[151,1435,237,1670]" captionTargetId="figure-36@11.[151,1435,237,1670]" captionTargetPageId="11" captionText="FIGURES 31–36. Arthromelodes tokushigei sp. n., male. Lateral fovea on abdominal tergite IV (31); median impression on tergite IV (32); posterior region of median impression on tergite IV (33); anterior region of impression on tergite IV (34); median process of median impression on tergite IV (35); spatulate setae on lateral setal patch of tergite IV (36). Abbreviations: lp, lateral process; mp, median process; mt, median tubercle; smp, submedian process." figureDoi="http://doi.org/10.5281/zenodo.8243478" httpUri="https://zenodo.org/record/8243478/files/figure.png" pageId="10" pageNumber="403">Fig. 35</figureCitation>
|
||
). Anterior margin of abdominal impression with strongly thickened cuticular walls (visible in cleared specimen:
|
||
<figureCitation id="B0B82A15FFC7DD60FF517E4829D7FE8E" box="[151,235,259,285]" captionStart="FIGURES 27–30" captionStartId="10.[151,261,1754,1778]" captionTargetBox="[151,1417,489,1721]" captionTargetId="figure-172@10.[151,1435,489,1731]" captionTargetPageId="10" captionText="FIGURES 27–30. Arthromelodes tokushigei sp. n., male.Abdomen in dorsal view (27); abdominal segments III–IV in dorsal (with ventral structures visible) (28) and left lateral (29) views; tergite V in dorsal view (30). Arrowheads indicate foveae (d, dorsal; v, ventral).Abbreviations: imcp, intermetacoxal process; lsp, lateral setal patch; mt, median tubercle; tIII, tergite III; tIV, tergite IV." figureDoi="http://doi.org/10.5281/zenodo.8243476" httpUri="https://zenodo.org/record/8243476/files/figure.png" pageId="10" pageNumber="403">Fig. 28</figureCitation>
|
||
). Median impression flanked by large and slightly impressed lateral setose patches (
|
||
<figureCitation id="B0B82A15FFC7DD60FB5C7E482DD1FE8E" box="[1178,1261,259,285]" captionStart="FIGURES 27–30" captionStartId="10.[151,261,1754,1778]" captionTargetBox="[151,1417,489,1721]" captionTargetId="figure-172@10.[151,1435,489,1731]" captionTargetPageId="10" captionText="FIGURES 27–30. Arthromelodes tokushigei sp. n., male.Abdomen in dorsal view (27); abdominal segments III–IV in dorsal (with ventral structures visible) (28) and left lateral (29) views; tergite V in dorsal view (30). Arrowheads indicate foveae (d, dorsal; v, ventral).Abbreviations: imcp, intermetacoxal process; lsp, lateral setal patch; mt, median tubercle; tIII, tergite III; tIV, tergite IV." figureDoi="http://doi.org/10.5281/zenodo.8243476" httpUri="https://zenodo.org/record/8243476/files/figure.png" pageId="10" pageNumber="403">Fig. 27</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC7DD60FB3F7E482C26FE8E" box="[1273,1306,259,285]" italics="true" pageId="10" pageNumber="403">lsp</emphasis>
|
||
) composed of dense and short spatulate setae (
|
||
<figureCitation id="B0B82A15FFC7DD60FDE57E6C2B44FED2" box="[547,632,295,321]" captionStart="FIGURES 31–36" captionStartId="11.[151,264,1693,1717]" captionTargetBox="[151,1435,237,1670]" captionTargetId="figure-36@11.[151,1435,237,1670]" captionTargetPageId="11" captionText="FIGURES 31–36. Arthromelodes tokushigei sp. n., male. Lateral fovea on abdominal tergite IV (31); median impression on tergite IV (32); posterior region of median impression on tergite IV (33); anterior region of impression on tergite IV (34); median process of median impression on tergite IV (35); spatulate setae on lateral setal patch of tergite IV (36). Abbreviations: lp, lateral process; mp, median process; mt, median tubercle; smp, submedian process." figureDoi="http://doi.org/10.5281/zenodo.8243478" httpUri="https://zenodo.org/record/8243478/files/figure.png" pageId="10" pageNumber="403">Fig. 36</figureCitation>
|
||
), each seta with convex ventral/posterior and concave, grooved dorsal/ anterior surface, presumably forming glandular evaporation apparatus, with glandular opening situated anteriorly on setal papilla at base of each modified seta. Tergites V (
|
||
<figureCitation id="B0B82A15FFC7DD60FCC67E242A6EFE1A" box="[768,850,367,393]" captionStart="FIGURES 27–30" captionStartId="10.[151,261,1754,1778]" captionTargetBox="[151,1417,489,1721]" captionTargetId="figure-172@10.[151,1435,489,1731]" captionTargetPageId="10" captionText="FIGURES 27–30. Arthromelodes tokushigei sp. n., male.Abdomen in dorsal view (27); abdominal segments III–IV in dorsal (with ventral structures visible) (28) and left lateral (29) views; tergite V in dorsal view (30). Arrowheads indicate foveae (d, dorsal; v, ventral).Abbreviations: imcp, intermetacoxal process; lsp, lateral setal patch; mt, median tubercle; tIII, tergite III; tIV, tergite IV." figureDoi="http://doi.org/10.5281/zenodo.8243476" httpUri="https://zenodo.org/record/8243476/files/figure.png" pageId="10" pageNumber="403">Fig. 30</figureCitation>
|
||
) to VII short and each with pair of dorsal sublateral foveae (
|
||
<figureCitation id="B0B82A15FFC7DD60FF367ED8287DFE3E" box="[240,321,403,429]" captionStart="FIGURES 27–30" captionStartId="10.[151,261,1754,1778]" captionTargetBox="[151,1417,489,1721]" captionTargetId="figure-172@10.[151,1435,489,1731]" captionTargetPageId="10" captionText="FIGURES 27–30. Arthromelodes tokushigei sp. n., male.Abdomen in dorsal view (27); abdominal segments III–IV in dorsal (with ventral structures visible) (28) and left lateral (29) views; tergite V in dorsal view (30). Arrowheads indicate foveae (d, dorsal; v, ventral).Abbreviations: imcp, intermetacoxal process; lsp, lateral setal patch; mt, median tubercle; tIII, tergite III; tIV, tergite IV." figureDoi="http://doi.org/10.5281/zenodo.8243476" httpUri="https://zenodo.org/record/8243476/files/figure.png" pageId="10" pageNumber="403">Fig. 30</figureCitation>
|
||
; indicated by arrowhead) directed posteromesad and slightly ventrad. Tergite VIII (
|
||
<figureCitation id="B0B82A15FFC7DD60FB107ED82C1AFE3E" box="[1238,1318,403,429]" captionStart="FIGURES 37–42" captionStartId="12.[151,264,1376,1400]" captionTargetBox="[157,1429,187,1346]" captionTargetId="figure-21@12.[151,1436,181,1352]" captionTargetPageId="12" captionText="FIGURES 37–42. Arthromelodes tokushigei sp. n., male (37–39) and female (40–42). Tergite VIII in dorsal view (37, 40); sternite VIII in ventral view (38, 42); sternite IX in ventral view, with internal membrane (39); postabdominal sclerites (41)." figureDoi="http://doi.org/10.5281/zenodo.8243480" httpUri="https://zenodo.org/record/8243480/files/figure.png" pageId="10" pageNumber="403">Fig. 37</figureCitation>
|
||
) lentiform with deep arcuate posteromedian emargination and two pairs of long lateral setae.
|
||
</paragraph>
|
||
<caption id="7CFC6618FFC7DD60FF5179912CA9F8A9" ID-DOI="http://doi.org/10.5281/zenodo.8243476" ID-Zenodo-Dep="8243476" httpUri="https://zenodo.org/record/8243476/files/figure.png" pageId="10" pageNumber="403" startId="10.[151,261,1754,1778]" targetBox="[151,1417,489,1721]" targetPageId="10" targetType="figure">
|
||
<paragraph id="283C3690FFC7DD60FF5179912CA9F8A9" blockId="10.[151,1437,1754,1850]" pageId="10" pageNumber="403">
|
||
<emphasis id="1AF7EA82FFC7DD60FF5179912872F961" bold="true" box="[151,334,1754,1778]" pageId="10" pageNumber="403">FIGURES 27–30.</emphasis>
|
||
<taxonomicName id="EF834D13FFC7DD60FE9279912B6DF961" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[340,593,1754,1778]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="403" phylum="Arthropoda" rank="species" species="tokushigei" status="sp. n.">
|
||
<emphasis id="1AF7EA82FFC7DD60FE9279912B6DF961" box="[340,593,1754,1778]" italics="true" pageId="10" pageNumber="403">Arthromelodes tokushigei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="01C457F9FFC7DD60FD9079902BB0F961" box="[598,652,1755,1778]" pageId="10" pageNumber="403" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male.Abdomen in dorsal view (
|
||
<emphasis id="1AF7EA82FFC7DD60FC0879912ADBF961" bold="true" box="[974,999,1754,1778]" pageId="10" pageNumber="403">27</emphasis>
|
||
); abdominal segments III–IV in dorsal (with ventral structures visible) (
|
||
<emphasis id="1AF7EA82FFC7DD60FE6279B52881F885" bold="true" box="[420,445,1790,1814]" pageId="10" pageNumber="403">28</emphasis>
|
||
) and left lateral (
|
||
<emphasis id="1AF7EA82FFC7DD60FDAD79B52BB8F885" bold="true" box="[619,644,1790,1814]" pageId="10" pageNumber="403">29</emphasis>
|
||
) views; tergite V in dorsal view (
|
||
<emphasis id="1AF7EA82FFC7DD60FC1279B52AD1F885" bold="true" box="[980,1005,1790,1814]" pageId="10" pageNumber="403">30</emphasis>
|
||
). Arrowheads indicate foveae (d, dorsal; v, ventral). Abbreviations: imcp, intermetacoxal process; lsp, lateral setal patch; mt, median tubercle; tIII, tergite III; tIV, tergite IV.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="283C3690FFC7DD61FF0178C628CDFF46" blockId="10.[151,1437,1933,2031]" lastBlockId="11.[151,1436,151,213]" lastPageId="11" lastPageNumber="404" pageId="10" pageNumber="403">
|
||
Sternite IV (
|
||
<figureCitation id="B0B82A15FFC7DD60FE9278C62894F834" box="[340,424,1933,1959]" captionStart="FIGURES 27–30" captionStartId="10.[151,261,1754,1778]" captionTargetBox="[151,1417,489,1721]" captionTargetId="figure-172@10.[151,1435,489,1731]" captionTargetPageId="10" captionText="FIGURES 27–30. Arthromelodes tokushigei sp. n., male.Abdomen in dorsal view (27); abdominal segments III–IV in dorsal (with ventral structures visible) (28) and left lateral (29) views; tergite V in dorsal view (30). Arrowheads indicate foveae (d, dorsal; v, ventral).Abbreviations: imcp, intermetacoxal process; lsp, lateral setal patch; mt, median tubercle; tIII, tergite III; tIV, tergite IV." figureDoi="http://doi.org/10.5281/zenodo.8243476" httpUri="https://zenodo.org/record/8243476/files/figure.png" pageId="10" pageNumber="403">Fig. 28</figureCitation>
|
||
, structures indicated with “v“, and
|
||
<figureCitation id="B0B82A15FFC7DD60FCFD78C62AB1F834" box="[827,909,1933,1959]" captionStart="FIGURES 27–30" captionStartId="10.[151,261,1754,1778]" captionTargetBox="[151,1417,489,1721]" captionTargetId="figure-172@10.[151,1435,489,1731]" captionTargetPageId="10" captionText="FIGURES 27–30. Arthromelodes tokushigei sp. n., male.Abdomen in dorsal view (27); abdominal segments III–IV in dorsal (with ventral structures visible) (28) and left lateral (29) views; tergite V in dorsal view (30). Arrowheads indicate foveae (d, dorsal; v, ventral).Abbreviations: imcp, intermetacoxal process; lsp, lateral setal patch; mt, median tubercle; tIII, tergite III; tIV, tergite IV." figureDoi="http://doi.org/10.5281/zenodo.8243476" httpUri="https://zenodo.org/record/8243476/files/figure.png" pageId="10" pageNumber="403">Fig. 29</figureCitation>
|
||
) with narrow keel-like intermetacoxal process (
|
||
<figureCitation id="B0B82A15FFC7DD60FF5978FA29CFF858" box="[159,243,1969,1995]" captionStart="FIGURES 27–30" captionStartId="10.[151,261,1754,1778]" captionTargetBox="[151,1417,489,1721]" captionTargetId="figure-172@10.[151,1435,489,1731]" captionTargetPageId="10" captionText="FIGURES 27–30. Arthromelodes tokushigei sp. n., male.Abdomen in dorsal view (27); abdominal segments III–IV in dorsal (with ventral structures visible) (28) and left lateral (29) views; tergite V in dorsal view (30). Arrowheads indicate foveae (d, dorsal; v, ventral).Abbreviations: imcp, intermetacoxal process; lsp, lateral setal patch; mt, median tubercle; tIII, tergite III; tIV, tergite IV." figureDoi="http://doi.org/10.5281/zenodo.8243476" httpUri="https://zenodo.org/record/8243476/files/figure.png" pageId="10" pageNumber="403">Fig. 29</figureCitation>
|
||
;
|
||
<emphasis id="1AF7EA82FFC7DD60FF3878FA2808F859" box="[254,308,1969,1994]" italics="true" pageId="10" pageNumber="403">imcp</emphasis>
|
||
) and two pairs of foveae, all directed mesad: ventral sublateral foveae and ventral submedian foveae (
|
||
<figureCitation id="B0B82A15FFC7DD60FF59789E29C9F87C" box="[159,245,2005,2031]" captionStart="FIGURES 27–30" captionStartId="10.[151,261,1754,1778]" captionTargetBox="[151,1417,489,1721]" captionTargetId="figure-172@10.[151,1435,489,1731]" captionTargetPageId="10" captionText="FIGURES 27–30. Arthromelodes tokushigei sp. n., male.Abdomen in dorsal view (27); abdominal segments III–IV in dorsal (with ventral structures visible) (28) and left lateral (29) views; tergite V in dorsal view (30). Arrowheads indicate foveae (d, dorsal; v, ventral).Abbreviations: imcp, intermetacoxal process; lsp, lateral setal patch; mt, median tubercle; tIII, tergite III; tIV, tergite IV." figureDoi="http://doi.org/10.5281/zenodo.8243476" httpUri="https://zenodo.org/record/8243476/files/figure.png" pageId="10" pageNumber="403">Fig. 28</figureCitation>
|
||
; indicated with arrowheads marked with “v“). Sternites V–VII (not shown) narrow and lacking foveae; sternite VIII (
|
||
<figureCitation id="B0B82A15FFC6DD61FEE87FDC2842FF22" box="[302,382,151,177]" captionStart="FIGURES 37–42" captionStartId="12.[151,264,1376,1400]" captionTargetBox="[157,1429,187,1346]" captionTargetId="figure-21@12.[151,1436,181,1352]" captionTargetPageId="12" captionText="FIGURES 37–42. Arthromelodes tokushigei sp. n., male (37–39) and female (40–42). Tergite VIII in dorsal view (37, 40); sternite VIII in ventral view (38, 42); sternite IX in ventral view, with internal membrane (39); postabdominal sclerites (41)." figureDoi="http://doi.org/10.5281/zenodo.8243480" httpUri="https://zenodo.org/record/8243480/files/figure.png" pageId="11" pageNumber="404">Fig. 38</figureCitation>
|
||
) crescent-shaped with shallow posteromedian emargination and one pair of long setae; sternite IX (
|
||
<figureCitation id="B0B82A15FFC6DD61FF027FF02829FF46" box="[196,277,187,213]" captionStart="FIGURES 37–42" captionStartId="12.[151,264,1376,1400]" captionTargetBox="[157,1429,187,1346]" captionTargetId="figure-21@12.[151,1436,181,1352]" captionTargetPageId="12" captionText="FIGURES 37–42. Arthromelodes tokushigei sp. n., male (37–39) and female (40–42). Tergite VIII in dorsal view (37, 40); sternite VIII in ventral view (38, 42); sternite IX in ventral view, with internal membrane (39); postabdominal sclerites (41)." figureDoi="http://doi.org/10.5281/zenodo.8243480" httpUri="https://zenodo.org/record/8243480/files/figure.png" pageId="11" pageNumber="404">Fig. 39</figureCitation>
|
||
) minute, lentiform.
|
||
</paragraph>
|
||
<caption id="7CFC6618FFC6DD61FF5179D62AA1F8B2" ID-DOI="http://doi.org/10.5281/zenodo.8243478" ID-Zenodo-Dep="8243478" httpUri="https://zenodo.org/record/8243478/files/figure.png" pageId="11" pageNumber="404" startId="11.[151,264,1693,1717]" targetBox="[151,1435,237,1670]" targetPageId="11" targetType="figure">
|
||
<paragraph id="283C3690FFC6DD61FF5179D62AA1F8B2" blockId="11.[151,1437,1693,1825]" pageId="11" pageNumber="404">
|
||
<emphasis id="1AF7EA82FFC6DD61FF5179D6286AF926" bold="true" box="[151,342,1693,1717]" pageId="11" pageNumber="404">FIGURES 31–36.</emphasis>
|
||
<taxonomicName id="EF834D13FFC6DD61FE9B79D62B5FF926" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[349,611,1693,1717]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="404" phylum="Arthropoda" rank="species" species="tokushigei" status="sp. n.">
|
||
<emphasis id="1AF7EA82FFC6DD61FE9B79D62B5FF926" box="[349,611,1693,1717]" italics="true" pageId="11" pageNumber="404">Arthromelodes tokushigei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="01C457F9FFC6DD61FDAD79D52B99F926" box="[619,677,1694,1717]" pageId="11" pageNumber="404" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male. Lateral fovea on abdominal tergite IV (
|
||
<emphasis id="1AF7EA82FFC6DD61FB4E79D62D9DF926" bold="true" box="[1160,1185,1693,1717]" pageId="11" pageNumber="404">31</emphasis>
|
||
); median impression on tergite IV (
|
||
<emphasis id="1AF7EA82FFC6DD61FEC2798A2821F94A" bold="true" box="[260,285,1729,1753]" pageId="11" pageNumber="404">32</emphasis>
|
||
); posterior region of median impression on tergite IV (
|
||
<emphasis id="1AF7EA82FFC6DD61FC84798A2A67F94A" bold="true" box="[834,859,1729,1753]" pageId="11" pageNumber="404">33</emphasis>
|
||
); anterior region of impression on tergite IV (
|
||
<emphasis id="1AF7EA82FFC6DD61FAE2798A2C01F94A" bold="true" box="[1316,1341,1729,1753]" pageId="11" pageNumber="404">34</emphasis>
|
||
); median process of median impression on tergite IV (
|
||
<emphasis id="1AF7EA82FFC6DD61FD9279AE2B51F96E" bold="true" box="[596,621,1765,1789]" pageId="11" pageNumber="404">35</emphasis>
|
||
); spatulate setae on lateral setal patch of tergite IV (
|
||
<emphasis id="1AF7EA82FFC6DD61FBB379AE2DB2F96E" bold="true" box="[1141,1166,1765,1789]" pageId="11" pageNumber="404">36</emphasis>
|
||
). Abbreviations: lp, lateral process; mp, median process; mt, median tubercle; smp, submedian process.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="283C3690FFC6DD61FF0178382DB8F869" blockId="11.[151,1437,1907,2042]" pageId="11" pageNumber="404">
|
||
Aedeagus (
|
||
<figureCitation id="B0B82A15FFC6DD61FE83783828FAF81E" box="[325,454,1907,1933]" captionStart="FIGURES 43–48" captionStartId="13.[151,264,1561,1585]" captionTargetBox="[157,1430,187,1531]" captionTargetId="figure-15@13.[151,1436,181,1537]" captionTargetPageId="13" captionText="FIGURES 43–48. Arthromelodes tokushigei sp. n., male. Aedeagus in ventral (43, 46), ventral and tilted dorsally to visualize shape of ventral projection (44, 47), and lateral (45, 48) views." figureDoi="http://doi.org/10.5281/zenodo.8243482" httpUri="https://zenodo.org/record/8243482/files/figure.png" pageId="11" pageNumber="404">Figs 43–48</figureCitation>
|
||
) with basal capsular region of median lobe stout, broader than long, ventral projection above ventral orifice slightly recurved, with subtriangular apex and strongly oblique apical margin; dorsal projection strongly bent ventrad at nearly right angle, with slander gradually tapering and pointed apex, proximal region of dorsal projection with small elongate subtriangular and pointed tooth directed proximally.
|
||
</paragraph>
|
||
<caption id="7CFC6618FFC1DD66FF517A2B2C43FA0F" ID-DOI="http://doi.org/10.5281/zenodo.8243480" ID-Zenodo-Dep="8243480" httpUri="https://zenodo.org/record/8243480/files/figure.png" pageId="12" pageNumber="405" startId="12.[151,264,1376,1400]" targetBox="[157,1429,187,1346]" targetPageId="12" targetType="figure">
|
||
<paragraph id="283C3690FFC1DD66FF517A2B2C43FA0F" blockId="12.[151,1436,1376,1436]" pageId="12" pageNumber="405">
|
||
<emphasis id="1AF7EA82FFC1DD66FF517A2B286BFAEB" bold="true" box="[151,343,1376,1400]" pageId="12" pageNumber="405">FIGURES 37–42.</emphasis>
|
||
<taxonomicName id="EF834D13FFC1DD66FE997A2B2B59FAEB" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[351,613,1376,1400]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="405" phylum="Arthropoda" rank="species" species="tokushigei" status="sp. n.">
|
||
<emphasis id="1AF7EA82FFC1DD66FE997A2B2B59FAEB" box="[351,613,1376,1400]" italics="true" pageId="12" pageNumber="405">Arthromelodes tokushigei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="01C457F9FFC1DD66FDA87A2A2B94FAEB" box="[622,680,1377,1400]" pageId="12" pageNumber="405" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male (
|
||
<emphasis id="1AF7EA82FFC1DD66FD3E7A2B2A0BFAEB" bold="true" box="[760,823,1376,1400]" pageId="12" pageNumber="405">37–39</emphasis>
|
||
) and female (
|
||
<emphasis id="1AF7EA82FFC1DD66FC0A7A2B2D37FAEB" bold="true" box="[972,1035,1376,1400]" pageId="12" pageNumber="405">40–42</emphasis>
|
||
). Tergite VIII in dorsal view (
|
||
<emphasis id="1AF7EA82FFC1DD66FA8D7A2B2CB0FAEB" bold="true" box="[1355,1420,1376,1400]" pageId="12" pageNumber="405">37, 40</emphasis>
|
||
); sternite VIII in ventral view (
|
||
<emphasis id="1AF7EA82FFC1DD66FE067ACF28C3FA0F" bold="true" box="[448,511,1412,1436]" pageId="12" pageNumber="405">38, 42</emphasis>
|
||
); sternite IX in ventral view, with internal membrane (
|
||
<emphasis id="1AF7EA82FFC1DD66FBED7ACF2D78FA0F" bold="true" box="[1067,1092,1412,1436]" pageId="12" pageNumber="405">39</emphasis>
|
||
); postabdominal sclerites (
|
||
<emphasis id="1AF7EA82FFC1DD66FA9E7ACF2C4DFA0F" bold="true" box="[1368,1393,1412,1436]" pageId="12" pageNumber="405">41</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="283C3690FFC1DD66FF017AA52DC5F92F" blockId="12.[151,1436,1518,2013]" pageId="12" pageNumber="405">
|
||
Female. Externally differs from male in unmodified abdomen with sternites IV–VIII exposed, unmodified porfemora, mesotibiae lacking apical mucro, and slightly shorter antennae in relation to body length. BL
|
||
<quantity id="EF7B9B75FFC1DD66FAEA795929FAF9C2" metricMagnitude="-3" metricUnit="m" metricValue="2.105" metricValueMax="2.13" metricValueMin="2.08" pageId="12" pageNumber="405" unit="mm" value="2.105" valueMax="2.13" valueMin="2.08">2.08–2.13 mm</quantity>
|
||
; HL
|
||
<quantity id="EF7B9B75FFC1DD66FEC6797D289BF9C2" box="[256,423,1590,1617]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="12" pageNumber="405" unit="mm" value="0.44" valueMax="0.45" valueMin="0.43">0.43–0.45 mm</quantity>
|
||
, HW
|
||
<quantity id="EF7B9B75FFC1DD66FE2D797D2BAEF9C2" box="[491,658,1590,1617]" metricMagnitude="-4" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" pageId="12" pageNumber="405" unit="mm" value="0.445" valueMax="0.45" valueMin="0.44">0.44–0.45 mm</quantity>
|
||
, AnL
|
||
<quantity id="EF7B9B75FFC1DD66FD1E797D2A7DF9C2" box="[728,833,1590,1617]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="12" pageNumber="405" unit="mm" value="1.15">1.15 mm</quantity>
|
||
; PL
|
||
<quantity id="EF7B9B75FFC1DD66FCB1797D2AE2F9C2" box="[887,990,1590,1617]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="12" pageNumber="405" unit="mm" value="0.48">0.48 mm</quantity>
|
||
, PW
|
||
<quantity id="EF7B9B75FFC1DD66FBD8797D2DF9F9C2" box="[1054,1221,1590,1617]" metricMagnitude="-4" metricUnit="m" metricValue="4.55" metricValueMax="4.6" metricValueMin="4.5" pageId="12" pageNumber="405" unit="mm" value="0.455" valueMax="0.46" valueMin="0.45">0.45–0.46 mm</quantity>
|
||
; EL
|
||
<quantity id="EF7B9B75FFC1DD66FB3A797D2C58F9C2" box="[1276,1380,1590,1617]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="12" pageNumber="405" unit="mm" value="0.58">0.58 mm</quantity>
|
||
, EW
|
||
<quantity id="EF7B9B75FFC1DD66FF5179112801F9E6" box="[151,317,1626,1653]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" metricValueMax="7.5" metricValueMin="7.3" pageId="12" pageNumber="405" unit="mm" value="0.74" valueMax="0.75" valueMin="0.73">0.73–0.75 mm</quantity>
|
||
; AbdL
|
||
<quantity id="EF7B9B75FFC1DD66FE4979112B08F9E6" box="[399,564,1626,1653]" metricMagnitude="-4" metricUnit="m" metricValue="6.15" metricValueMax="6.3" metricValueMin="6.0" pageId="12" pageNumber="405" unit="mm" value="0.615" valueMax="0.63" valueMin="0.6">0.60–0.63 mm</quantity>
|
||
, AbdW
|
||
<quantity id="EF7B9B75FFC1DD66FD5679112BCBF9E6" box="[656,759,1626,1653]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="12" pageNumber="405" unit="mm" value="0.75">0.75 mm</quantity>
|
||
. Tergite VIII (
|
||
<figureCitation id="B0B82A15FFC1DD66FC5079112AD4F9E7" box="[918,1000,1626,1652]" captionStart="FIGURES 37–42" captionStartId="12.[151,264,1376,1400]" captionTargetBox="[157,1429,187,1346]" captionTargetId="figure-21@12.[151,1436,181,1352]" captionTargetPageId="12" captionText="FIGURES 37–42. Arthromelodes tokushigei sp. n., male (37–39) and female (40–42). Tergite VIII in dorsal view (37, 40); sternite VIII in ventral view (38, 42); sternite IX in ventral view, with internal membrane (39); postabdominal sclerites (41)." figureDoi="http://doi.org/10.5281/zenodo.8243480" httpUri="https://zenodo.org/record/8243480/files/figure.png" pageId="12" pageNumber="405">Fig. 40</figureCitation>
|
||
) lentiform, posteriorly not emarginate; sternite VIII (
|
||
<figureCitation id="B0B82A15FFC1DD66FEFD793528AEF90B" box="[315,402,1662,1688]" captionStart="FIGURES 37–42" captionStartId="12.[151,264,1376,1400]" captionTargetBox="[157,1429,187,1346]" captionTargetId="figure-21@12.[151,1436,181,1352]" captionTargetPageId="12" captionText="FIGURES 37–42. Arthromelodes tokushigei sp. n., male (37–39) and female (40–42). Tergite VIII in dorsal view (37, 40); sternite VIII in ventral view (38, 42); sternite IX in ventral view, with internal membrane (39); postabdominal sclerites (41)." figureDoi="http://doi.org/10.5281/zenodo.8243480" httpUri="https://zenodo.org/record/8243480/files/figure.png" pageId="12" pageNumber="405">Fig. 42</figureCitation>
|
||
) crescent-shaped with one pair of long setae and lacking posteromedian emargination; postabdominal sclerites (
|
||
<figureCitation id="B0B82A15FFC1DD66FE6879E92B3CF92F" box="[430,512,1698,1724]" captionStart="FIGURES 37–42" captionStartId="12.[151,264,1376,1400]" captionTargetBox="[157,1429,187,1346]" captionTargetId="figure-21@12.[151,1436,181,1352]" captionTargetPageId="12" captionText="FIGURES 37–42. Arthromelodes tokushigei sp. n., male (37–39) and female (40–42). Tergite VIII in dorsal view (37, 40); sternite VIII in ventral view (38, 42); sternite IX in ventral view, with internal membrane (39); postabdominal sclerites (41)." figureDoi="http://doi.org/10.5281/zenodo.8243480" httpUri="https://zenodo.org/record/8243480/files/figure.png" pageId="12" pageNumber="405">Fig. 41</figureCitation>
|
||
) complex, symmetrical (preparation in
|
||
<figureCitation id="B0B82A15FFC1DD66FC7D79E92D37F92F" box="[955,1035,1698,1724]" captionStart="FIGURES 37–42" captionStartId="12.[151,264,1376,1400]" captionTargetBox="[157,1429,187,1346]" captionTargetId="figure-21@12.[151,1436,181,1352]" captionTargetPageId="12" captionText="FIGURES 37–42. Arthromelodes tokushigei sp. n., male (37–39) and female (40–42). Tergite VIII in dorsal view (37, 40); sternite VIII in ventral view (38, 42); sternite IX in ventral view, with internal membrane (39); postabdominal sclerites (41)." figureDoi="http://doi.org/10.5281/zenodo.8243480" httpUri="https://zenodo.org/record/8243480/files/figure.png" pageId="12" pageNumber="405">Fig. 41</figureCitation>
|
||
is slightly distorted).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6099651BFFC1DD66FF01798D2A5FF972" box="[199,867,1734,1761]" pageId="12" pageNumber="405" type="distribution">
|
||
<paragraph id="283C3690FFC1DD66FF01798D2A5FF972" blockId="12.[151,1436,1518,2013]" box="[199,867,1734,1761]" pageId="12" pageNumber="405">
|
||
<emphasis id="1AF7EA82FFC1DD66FF01798D285DF973" bold="true" box="[199,353,1734,1760]" pageId="12" pageNumber="405">Distribution.</emphasis>
|
||
<collectingRegion id="EA47F872FFC1DD66FEA1798D28F0F973" box="[359,460,1734,1760]" country="Japan" name="Okinawa" pageId="12" pageNumber="405">Okinawa</collectingRegion>
|
||
Island, Ryȗkyȗ Archipelago,
|
||
<collectingCountry id="50947600FFC1DD66FCDD798C2A62F972" box="[795,862,1735,1761]" name="Japan" pageId="12" pageNumber="405">Japan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6099651BFFC1DD66FF0179A128A1F8E3" pageId="12" pageNumber="405" type="etymology">
|
||
<paragraph id="283C3690FFC1DD66FF0179A128A1F8E3" blockId="12.[151,1436,1518,2013]" pageId="12" pageNumber="405">
|
||
<emphasis id="1AF7EA82FFC1DD66FF0179A12872F897" bold="true" box="[199,334,1770,1796]" pageId="12" pageNumber="405">Etymology.</emphasis>
|
||
The new species is named in honor of Mr. Norihide Tokushige, who organized the field trip on which specimens of the first
|
||
<taxonomicName id="EF834D13FFC1DD66FE3678452BA9F8BB" authorityName="Jeannel" authorityYear="1954" box="[496,661,1806,1832]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="405" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1AF7EA82FFC1DD66FE3678452BA9F8BB" box="[496,661,1806,1832]" italics="true" pageId="12" pageNumber="405">Arthromelodes</emphasis>
|
||
</taxonomicName>
|
||
known to inhabit Okinawa-jima were collected, and who himself discovered many interesting
|
||
<taxonomicName id="EF834D13FFC1DD66FE1A78792B5EF8DF" box="[476,610,1842,1868]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="405" phylum="Arthropoda" rank="subFamily" subFamily="Pselaphinae">Pselaphinae</taxonomicName>
|
||
and
|
||
<taxonomicName id="EF834D13FFC1DD66FD5C78792A01F8DF" box="[666,829,1842,1868]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="405" phylum="Arthropoda" rank="subFamily" subFamily="Scydmaeninae">Scydmaeninae</taxonomicName>
|
||
and significantly contributed to the knowledge of the Okinawan biodiversity.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6099651BFFC1DD64FF0178312D4BFAC6" lastPageId="14" lastPageNumber="407" pageId="12" pageNumber="405" type="discussion">
|
||
<paragraph id="283C3690FFC1DD64FF0178312B9FFE66" blockId="12.[151,1436,1518,2013]" lastBlockId="14.[151,1437,151,1365]" lastPageId="14" lastPageNumber="407" pageId="12" pageNumber="405">
|
||
<emphasis id="1AF7EA82FFC1DD66FF0178312878F807" bold="true" box="[199,324,1914,1940]" pageId="12" pageNumber="405">Remarks.</emphasis>
|
||
To date, eighteen species and two subspecies of
|
||
<taxonomicName id="EF834D13FFC1DD66FC4578312D0AF807" authorityName="Jeannel" authorityYear="1954" box="[899,1078,1914,1940]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="405" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1AF7EA82FFC1DD66FC4578312D0AF807" box="[899,1078,1914,1940]" italics="true" pageId="12" pageNumber="405">Arthromelodes</emphasis>
|
||
</taxonomicName>
|
||
were known to occur in
|
||
<collectingCountry id="50947600FFC1DD66FA9178302CA0F806" box="[1367,1436,1915,1941]" name="Japan" pageId="12" pageNumber="405">Japan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="4C124B61FFC1DD66FF5978D528B6F82B" author="Shibata, Y. & Maruyama, M. & Hoshina, H. & Kishimoto, T. & Naomi, S. - I. & Nomura, S. & Puthz, V. & Shimada, T. & Watanabe, Y. & Yamamoto, S." box="[159,394,1950,1976]" pageId="12" pageNumber="405" pagination="69 - 218" refId="ref6900" refString="Shibata, Y., Maruyama, M., Hoshina, H., Kishimoto, T., Naomi, S. - I., Nomura, S., Puthz, V., Shimada, T., Watanabe, Y. & Yamamoto, S. (2013) Catalogue of Japanese Staphylinidae (Insecta: Coleoptera). Bulletin of the Kyushu University Museum, 11, 69 - 218." type="journal article" year="2013">
|
||
Shibata
|
||
<emphasis id="1AF7EA82FFC1DD66FEC278D4287DF82B" box="[260,321,1950,1976]" italics="true" pageId="12" pageNumber="405">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
). Fifteen of them inhabit the large ‘mainland’ islands (Honshȗ, Kyȗshȗ and Shikoku), one was discovered in Yakushima (~
|
||
<quantity id="EF7B9B75FFC1DD66FDF278892B41F84F" box="[564,637,1986,2012]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="405" unit="km" value="60.0">60 km</quantity>
|
||
S of Kyȗshȗ), and two on the northeastern Ryȗkyȗan Amami Islands. No species have been recorded from central or southwestern Ryȗkyȗs. When the identification key of
|
||
<bibRefCitation id="4C124B61FFC0DD67FA8679E329DEF976" author="Nomura, S." pageId="13" pageNumber="406" pagination="1 - 462" refId="ref6867" refString="Nomura, S. (1991) Systematic study on the genus Batrisoplisus and its allied genera from Japan (Coleoptera, Pselaphidae). Esakia, 30, 1 - 462." type="journal article" year="1991">Nomura (1991)</bibRefCitation>
|
||
is used,
|
||
<taxonomicName id="EF834D13FFC0DD67FE83798028EBF976" authorityName="Jałoszyński" authorityYear="2023" box="[325,471,1739,1765]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFC0DD67FE83798028EBF976" box="[325,471,1739,1765]" italics="true" pageId="13" pageNumber="406">A. tokushigei</emphasis>
|
||
</taxonomicName>
|
||
may key out as
|
||
<taxonomicName id="EF834D13FFC0DD67FD5679802A50F975" authority="Nomura" authorityName="Nomura" authorityYear="1991" box="[656,876,1739,1766]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="crucifer">
|
||
<emphasis id="1AF7EA82FFC0DD67FD5679802A35F976" box="[656,777,1739,1765]" italics="true" pageId="13" pageNumber="406">A. crucifer</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
or the pair
|
||
<taxonomicName id="EF834D13FFC0DD67FC2879802DDEF975" authority="Nomura" authorityName="Nomura" authorityYear="1991" box="[1006,1250,1739,1766]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="sinuatipes">
|
||
<emphasis id="1AF7EA82FFC0DD67FC2879802D43F976" box="[1006,1151,1739,1765]" italics="true" pageId="13" pageNumber="406">A. sinuatipes</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EF834D13FFC0DD67FADF798029CAF899" authority="Nomura" authorityName="Nomura" authorityYear="1991" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="aizuanus">
|
||
<emphasis id="1AF7EA82FFC0DD67FADF79802CA1F976" box="[1305,1437,1739,1765]" italics="true" pageId="13" pageNumber="406">A. aizuanus</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
, depending on interpretation of the punctures on head and pronotum (densely and coarsely punctate vs. sparsely punctate—it is difficult to decide which
|
||
<typeStatus id="F7388832FFC0DD67FD7A785F2BD0F8BD" box="[700,748,1812,1838]" pageId="13" pageNumber="406">type</typeStatus>
|
||
of punctation better describes the condition in
|
||
<taxonomicName id="EF834D13FFC0DD67FB3C78582CB0F8BE" authorityName="Jałoszyński" authorityYear="2023" box="[1274,1420,1811,1837]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFC0DD67FB3C78582CB0F8BE" box="[1274,1420,1811,1837]" italics="true" pageId="13" pageNumber="406">A. tokushigei</emphasis>
|
||
</taxonomicName>
|
||
). However, none of the Japanese species revised by
|
||
<bibRefCitation id="4C124B61FFC0DD67FD1378732ABFF8C2" author="Nomura, S." box="[725,899,1847,1874]" pageId="13" pageNumber="406" pagination="1 - 462" refId="ref6867" refString="Nomura, S. (1991) Systematic study on the genus Batrisoplisus and its allied genera from Japan (Coleoptera, Pselaphidae). Esakia, 30, 1 - 462." type="journal article" year="1991">Nomura (1991)</bibRefCitation>
|
||
and later described by
|
||
<bibRefCitation id="4C124B61FFC0DD67FB4D787C2C2CF8C2" author="Arai, S." box="[1163,1296,1847,1873]" pageId="13" pageNumber="406" pagination="275 - 279" refId="ref6567" refString="Arai, S. (2002) A new species of the genus Arthromelodes (Coleoptera, Staphylinidae, Pselaphinae) from Kanto District, central Japan. Special Bulletin of the Japanese Society of Coleopterology, 5, 275 - 279." type="journal article" year="2002">Arai (2002)</bibRefCitation>
|
||
has an even remotely similar aedeagus. Among the previously known Japanese species, only seven have the abdominal tergite IV modified:
|
||
<taxonomicName id="EF834D13FFC0DD67FEF578342B1CF809" authority="Nomura" authorityName="Nomura" authorityYear="1991" box="[307,544,1919,1946]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="aizuanus">
|
||
<emphasis id="1AF7EA82FFC0DD67FEF578342884F80A" box="[307,440,1919,1945]" italics="true" pageId="13" pageNumber="406">A. aizuanus</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EF834D13FFC0DD67FDEB78342A33F809" authority="Nomura" authorityName="Nomura" authorityYear="1991" box="[557,783,1919,1946]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="crucifer">
|
||
<emphasis id="1AF7EA82FFC0DD67FDEB78342B9BF80A" box="[557,679,1919,1945]" italics="true" pageId="13" pageNumber="406">A. crucifer</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EF834D13FFC0DD67FCDA78342D2FF80A" authority="(Raffray)" baseAuthorityName="Raffray" baseAuthorityYear="1909" box="[796,1043,1919,1945]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="dilatatus">
|
||
<emphasis id="1AF7EA82FFC0DD67FCDA78342AA3F80A" box="[796,927,1919,1945]" italics="true" pageId="13" pageNumber="406">A. dilatatus</emphasis>
|
||
(Raffray)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EF834D13FFC0DD67FBD978342DCFF80A" authority="(Sharp)" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[1055,1267,1919,1945]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="optatus">
|
||
<emphasis id="1AF7EA82FFC0DD67FBD978342DAFF80A" box="[1055,1171,1919,1945]" italics="true" pageId="13" pageNumber="406">A. optatus</emphasis>
|
||
(Sharp)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EF834D13FFC0DD67FAC6783429CAF82D" authority="Nomura" authorityName="Nomura" authorityYear="1991" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="saikaiensis">
|
||
<emphasis id="1AF7EA82FFC0DD67FAC678342CA0F80A" box="[1280,1436,1919,1945]" italics="true" pageId="13" pageNumber="406">A. saikaiensis</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EF834D13FFC0DD67FEC278E828C3F82D" authority="Nomura" authorityName="Nomura" authorityYear="1991" box="[260,511,1955,1982]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="sinuatipes">
|
||
<emphasis id="1AF7EA82FFC0DD67FEC278E828ABF82E" box="[260,407,1955,1981]" italics="true" pageId="13" pageNumber="406">A. sinuatipes</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EF834D13FFC0DD67FDF978E82A2BF82E" authority="Arai." authorityName="Arai." box="[575,791,1955,1981]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="watanabei">
|
||
<emphasis id="1AF7EA82FFC0DD67FDF978E82BEAF82E" box="[575,726,1955,1981]" italics="true" pageId="13" pageNumber="406">A. watanabei</emphasis>
|
||
Arai.
|
||
</taxonomicName>
|
||
<taxonomicName id="EF834D13FFC0DD67FCD978E82D0EF82E" authorityName="Nomura" authorityYear="1991" box="[799,1074,1955,1981]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="aizuanus">
|
||
<emphasis id="1AF7EA82FFC0DD67FCD978E82D0EF82E" box="[799,1074,1955,1981]" italics="true" pageId="13" pageNumber="406">Arthromelodes aizuanus</emphasis>
|
||
</taxonomicName>
|
||
has the lateral setal patches on tergite IV strongly elongate, anteriorly extending far beyond the anterior margin of the posteromedian impression (in
|
||
<taxonomicName id="EF834D13FFC3DD64FF7A7FDC2871FF22" authorityName="Jałoszyński" authorityYear="2023" box="[188,333,151,177]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFC3DD64FF7A7FDC2871FF22" box="[188,333,151,177]" italics="true" pageId="14" pageNumber="407">A. tokushigei</emphasis>
|
||
</taxonomicName>
|
||
not extending beyond the anterior margin of the impression);
|
||
<taxonomicName id="EF834D13FFC3DD64FC3C7FDC2D4CFF22" authorityName="Nomura" authorityYear="1991" box="[1018,1136,151,177]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="crucifer">
|
||
<emphasis id="1AF7EA82FFC3DD64FC3C7FDC2D4CFF22" box="[1018,1136,151,177]" italics="true" pageId="14" pageNumber="407">A. crucifer</emphasis>
|
||
</taxonomicName>
|
||
has the median impression on tergite IV elongate, with a “crossed carina“ (
|
||
<bibRefCitation id="4C124B61FFC3DD64FD0E7FF02A55FF46" author="Nomura, S." box="[712,873,187,213]" pageId="14" pageNumber="407" pagination="1 - 462" refId="ref6867" refString="Nomura, S. (1991) Systematic study on the genus Batrisoplisus and its allied genera from Japan (Coleoptera, Pselaphidae). Esakia, 30, 1 - 462." type="journal article" year="1991">Nomura 1991</bibRefCitation>
|
||
) and lacking oval posteromedian tubercle (in
|
||
<taxonomicName id="EF834D13FFC3DD64FA427FF02836FF6A" authorityName="Jałoszyński" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFC3DD64FA427FF02836FF6A" italics="true" pageId="14" pageNumber="407">A. tokushigei</emphasis>
|
||
</taxonomicName>
|
||
there is no carina in the impression, which is posteriorly occupied by a large oval tubercle); in
|
||
<taxonomicName id="EF834D13FFC3DD64FADA7F942CA0FF6A" baseAuthorityName="Raffray" baseAuthorityYear="1909" box="[1308,1436,223,249]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="dilatatus">
|
||
<emphasis id="1AF7EA82FFC3DD64FADA7F942CA0FF6A" box="[1308,1436,223,249]" italics="true" pageId="14" pageNumber="407">A. dilatatus</emphasis>
|
||
</taxonomicName>
|
||
the lateral setose patches on tergite IV are situated on broad and laterally protruding lobes (lobes lacking in
|
||
<taxonomicName id="EF834D13FFC3DD64FA427E482836FED2" authorityName="Jałoszyński" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFC3DD64FA427E482836FED2" italics="true" pageId="14" pageNumber="407">A. tokushigei</emphasis>
|
||
</taxonomicName>
|
||
); in
|
||
<taxonomicName id="EF834D13FFC3DD64FEF87E6C2893FED3" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[318,431,295,321]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="optatus">
|
||
<emphasis id="1AF7EA82FFC3DD64FEF87E6C2893FED3" box="[318,431,295,321]" italics="true" pageId="14" pageNumber="407">A. optatus</emphasis>
|
||
</taxonomicName>
|
||
the impression on tergite IV bears a median longitudinal carina (lacking in
|
||
<taxonomicName id="EF834D13FFC3DD64FB3F7E6C2CB7FED2" authorityName="Jałoszyński" authorityYear="2023" box="[1273,1419,295,321]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFC3DD64FB3F7E6C2CB7FED2" box="[1273,1419,295,321]" italics="true" pageId="14" pageNumber="407">A. tokushigei</emphasis>
|
||
</taxonomicName>
|
||
); in
|
||
<taxonomicName id="EF834D13FFC3DD64FF737E002873FEF6" authorityName="Nomura" authorityYear="1991" box="[181,335,331,357]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="saikaiensis">
|
||
<emphasis id="1AF7EA82FFC3DD64FF737E002873FEF6" box="[181,335,331,357]" italics="true" pageId="14" pageNumber="407">A. saikaiensis</emphasis>
|
||
</taxonomicName>
|
||
the abdomen is strongly constricted at base (not constricted in
|
||
<taxonomicName id="EF834D13FFC3DD64FBDE7E002D97FEF6" authorityName="Jałoszyński" authorityYear="2023" box="[1048,1195,331,357]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFC3DD64FBDE7E002D97FEF6" box="[1048,1195,331,357]" italics="true" pageId="14" pageNumber="407">A. tokushigei</emphasis>
|
||
</taxonomicName>
|
||
); in
|
||
<taxonomicName id="EF834D13FFC3DD64FB277E002C4FFEF7" authorityName="Nomura" authorityYear="1991" box="[1249,1395,331,357]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="sinuatipes">
|
||
<emphasis id="1AF7EA82FFC3DD64FB277E002C4FFEF7" box="[1249,1395,331,357]" italics="true" pageId="14" pageNumber="407">A. sinuatipes</emphasis>
|
||
</taxonomicName>
|
||
the abdominal tergite IV has a peculiar shape, it is rapidly narrowed behind the lateral setal patches and the median impression is distant from the posterior tergal margin (tergite not rapidly narrowed and the impression posteriorly adjacent to the posterior tergal margin); and
|
||
<taxonomicName id="EF834D13FFC3DD64FD4D7EFC2A23FE42" box="[651,799,439,465]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="watanabei">
|
||
<emphasis id="1AF7EA82FFC3DD64FD4D7EFC2A23FE42" box="[651,799,439,465]" italics="true" pageId="14" pageNumber="407">A. watanabei</emphasis>
|
||
</taxonomicName>
|
||
has strongly modified metatrochanters, each with a long curved ventral spine (lacking in
|
||
<taxonomicName id="EF834D13FFC3DD64FDC77E902BAFFE66" authorityName="Jałoszyński" authorityYear="2023" box="[513,659,475,501]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFC3DD64FDC77E902BAFFE66" box="[513,659,475,501]" italics="true" pageId="14" pageNumber="407">A. tokushigei</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption id="7CFC6618FFC0DD67FF5179522A2FF9C6" ID-DOI="http://doi.org/10.5281/zenodo.8243482" ID-Zenodo-Dep="8243482" httpUri="https://zenodo.org/record/8243482/files/figure.png" pageId="13" pageNumber="406" startId="13.[151,264,1561,1585]" targetBox="[157,1430,187,1531]" targetPageId="13" targetType="figure">
|
||
<paragraph id="283C3690FFC0DD67FF5179522A2FF9C6" blockId="13.[151,1436,1561,1621]" pageId="13" pageNumber="406">
|
||
<emphasis id="1AF7EA82FFC0DD67FF5179522869F9A2" bold="true" box="[151,341,1561,1585]" pageId="13" pageNumber="406">FIGURES 43–48.</emphasis>
|
||
<taxonomicName id="EF834D13FFC0DD67FE9A79522B5CF9A2" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[348,608,1561,1585]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="406" phylum="Arthropoda" rank="species" species="tokushigei" status="sp. n.">
|
||
<emphasis id="1AF7EA82FFC0DD67FE9A79522B5CF9A2" box="[348,608,1561,1585]" italics="true" pageId="13" pageNumber="406">Arthromelodes tokushigei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="01C457F9FFC0DD67FDAE79522B9CF9A3" box="[616,672,1561,1584]" pageId="13" pageNumber="406" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male. Aedeagus in ventral (
|
||
<emphasis id="1AF7EA82FFC0DD67FC0379522D38F9A2" bold="true" box="[965,1028,1561,1585]" pageId="13" pageNumber="406">43, 46</emphasis>
|
||
), ventral and tilted dorsally to visualize shape of ventral projection (
|
||
<emphasis id="1AF7EA82FFC0DD67FE73797628C8F9C6" bold="true" box="[437,500,1597,1621]" pageId="13" pageNumber="406">44, 47</emphasis>
|
||
), and lateral (
|
||
<emphasis id="1AF7EA82FFC0DD67FD4479762BFDF9C6" bold="true" box="[642,705,1597,1621]" pageId="13" pageNumber="406">45, 48</emphasis>
|
||
) views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="283C3690FFC3DD64FF017EB4286CFD62" blockId="14.[151,1437,151,1365]" pageId="14" pageNumber="407">
|
||
Among Eastern Palaearctic species (including those inhabiting the transition zone between Palaearctic and Oriental realms),
|
||
<taxonomicName id="EF834D13FFC3DD64FEA17D6828C3FDAE" authorityName="Jałoszyński" authorityYear="2023" box="[359,511,547,573]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFC3DD64FEA17D6828C3FDAE" box="[359,511,547,573]" italics="true" pageId="14" pageNumber="407">A. tokushigei</emphasis>
|
||
</taxonomicName>
|
||
somewhat resembles
|
||
<taxonomicName id="EF834D13FFC3DD64FCC17D682AD6FDAE" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[775,1002,547,573]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="choui">
|
||
<emphasis id="1AF7EA82FFC3DD64FCC17D682A56FDAE" box="[775,874,547,573]" italics="true" pageId="14" pageNumber="407">A. choui</emphasis>
|
||
<bibRefCitation id="4C124B61FFC3DD64FCB07D682AD6FDAE" author="Yin, Z. - W." box="[886,1002,547,573]" pageId="14" pageNumber="407" pagination="43 - 47" refId="ref6978" refString="Yin, Z. - W. (2018) First record of Arthromelodes Jeannel in China, with description of a new species (Coleoptera, Staphylinidae, Pselaphinae). ZooKeys, 744, 43 - 47. https: // doi. org / 10.3897 / zookeys. 744.23318" type="journal article" year="2018">Yin, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<collectingRegion id="EA47F872FFC3DD64FC397D682D60FDAE" box="[1023,1116,547,573]" country="China" name="Taiwan)" pageId="14" pageNumber="407">Taiwan)</collectingRegion>
|
||
in the modification of the abdominal tergite IV and in the aedeagal structure. However, in
|
||
<taxonomicName id="EF834D13FFC3DD64FCB67D0C2AF3FDF2" authorityName="Yin" authorityYear="2018" box="[880,975,583,609]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="choui">
|
||
<emphasis id="1AF7EA82FFC3DD64FCB67D0C2AF3FDF2" box="[880,975,583,609]" italics="true" pageId="14" pageNumber="407">A. choui</emphasis>
|
||
</taxonomicName>
|
||
the lateral setal patches on the abdomen extend anteriorly far beyond the posteromedian impression, and the latter is about as long as broad (in
|
||
<taxonomicName id="EF834D13FFC3DD64FACD7D202CA0FD16" authorityName="Jałoszyński" authorityYear="2023" box="[1291,1436,619,645]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFC3DD64FACD7D202CA0FD16" box="[1291,1436,619,645]" italics="true" pageId="14" pageNumber="407">A. tokushigei</emphasis>
|
||
</taxonomicName>
|
||
the setal patches anteriorly do not extend beyond the anterior margin of the impression, which is clearly transverse), the pronotal punctures are inconspicous (in
|
||
<taxonomicName id="EF834D13FFC3DD64FD4A7DF82A1CFD5E" authorityName="Jałoszyński" authorityYear="2023" box="[652,800,691,717]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFC3DD64FD4A7DF82A1CFD5E" box="[652,800,691,717]" italics="true" pageId="14" pageNumber="407">A. tokushigei</emphasis>
|
||
</taxonomicName>
|
||
distinct and dense), and the aedeagal projections are of different shapes.
|
||
</paragraph>
|
||
<paragraph id="283C3690FFC3DD64FF017DB0286DFBEE" blockId="14.[151,1437,151,1365]" pageId="14" pageNumber="407">
|
||
Interestingly, among
|
||
<taxonomicName id="EF834D13FFC3DD64FE707DB02B2AFC86" box="[438,534,763,789]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="tribe" tribe="Batrisini">Batrisini</taxonomicName>
|
||
of Okinawa-jima, females of
|
||
<taxonomicName id="EF834D13FFC3DD64FCAC7DB02AC1FC86" authorityName="Jałoszyński" authorityYear="2023" box="[874,1021,763,789]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFC3DD64FCAC7DB02AC1FC86" box="[874,1021,763,789]" italics="true" pageId="14" pageNumber="407">A. tokushigei</emphasis>
|
||
</taxonomicName>
|
||
can be easily confused with females of
|
||
<taxonomicName id="EF834D13FFC3DD64FF7F7C542B64FCAA" authority="Nomura, 1991" authorityName="Nomura" authorityYear="1991" box="[185,600,799,825]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="sakaii">
|
||
<emphasis id="1AF7EA82FFC3DD64FF7F7C54289AFCAA" box="[185,422,799,825]" italics="true" pageId="14" pageNumber="407">Batriscenellus sakaii</emphasis>
|
||
<bibRefCitation id="4C124B61FFC3DD64FE767C542B64FCAA" author="Nomura, S." box="[432,600,799,825]" pageId="14" pageNumber="407" pagination="1 - 462" refId="ref6867" refString="Nomura, S. (1991) Systematic study on the genus Batrisoplisus and its allied genera from Japan (Coleoptera, Pselaphidae). Esakia, 30, 1 - 462." type="journal article" year="1991">Nomura, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
. Both species were collected in the same spot, but
|
||
<taxonomicName id="EF834D13FFC3DD64FB687C542C2FFCAA" authorityName="Nomura" authorityYear="1991" box="[1198,1299,799,825]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="sakaii">
|
||
<emphasis id="1AF7EA82FFC3DD64FB687C542C2FFCAA" box="[1198,1299,799,825]" italics="true" pageId="14" pageNumber="407">B. sakaii</emphasis>
|
||
</taxonomicName>
|
||
is common also in other localities in the Yambaru area (Jałoszyński, personal obs.), while
|
||
<taxonomicName id="EF834D13FFC3DD64FBE47C082D8BFCCE" authorityName="Jałoszyński" authorityYear="2023" box="[1058,1207,835,861]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFC3DD64FBE47C082D8BFCCE" box="[1058,1207,835,861]" italics="true" pageId="14" pageNumber="407">A. tokushigei</emphasis>
|
||
</taxonomicName>
|
||
has not been found by Japanese collectors who explored this island previously (no specimens have been seen by S. Nomura; email dated
|
||
<date id="5C3D1050FFC3DD64FF1A7CC02863FC36" box="[220,351,907,933]" pageId="14" pageNumber="407" value="2023-04-26">26.04.2023</date>
|
||
). One of the reasons for overlooking
|
||
<taxonomicName id="EF834D13FFC3DD64FCC17CC02AA6FC36" authorityName="Jałoszyński" authorityYear="2023" box="[775,922,907,933]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFC3DD64FCC17CC02AA6FC36" box="[775,922,907,933]" italics="true" pageId="14" pageNumber="407">A. tokushigei</emphasis>
|
||
</taxonomicName>
|
||
may be that this is a wingless species, so the commonly used flight intercept traps will not detect its presence. The females of
|
||
<taxonomicName id="EF834D13FFC3DD64FBD57CE42D98FC5A" authorityName="Jałoszyński" authorityYear="2023" box="[1043,1188,943,969]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFC3DD64FBD57CE42D98FC5A" box="[1043,1188,943,969]" italics="true" pageId="14" pageNumber="407">A. tokushigei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EF834D13FFC3DD64FB1E7CE42C05FC5A" authorityName="Nomura" authorityYear="1991" box="[1240,1337,943,969]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="sakaii">
|
||
<emphasis id="1AF7EA82FFC3DD64FB1E7CE42C05FC5A" box="[1240,1337,943,969]" italics="true" pageId="14" pageNumber="407">B. sakaii</emphasis>
|
||
</taxonomicName>
|
||
have the same size and proportions of body parts; they clearly differ in distinct pronotal punctures in the former species vs. a smooth pronotal disc in
|
||
<taxonomicName id="EF834D13FFC3DD64FE607CBC2B35FB82" authorityName="Nomura" authorityYear="1991" box="[422,521,1015,1041]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="sakaii">
|
||
<emphasis id="1AF7EA82FFC3DD64FE607CBC2B35FB82" box="[422,521,1015,1041]" italics="true" pageId="14" pageNumber="407">B. sakaii</emphasis>
|
||
</taxonomicName>
|
||
, and in the structure of the postabdominal sclerites. As diagnoses of many genera of
|
||
<taxonomicName id="EF834D13FFC3DD64FF707B50282AFBA6" box="[182,278,1051,1077]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="tribe" tribe="Batrisini">Batrisini</taxonomicName>
|
||
seem extremely unclear (to the extent that they seem unusable) and this group will certainly require a profound reclassification, it is difficult to distinguish females of
|
||
<taxonomicName id="EF834D13FFC3DD64FCA47B742AC8FBCA" authorityName="Jałoszyński" authorityYear="2023" box="[866,1012,1087,1113]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFC3DD64FCA47B742AC8FBCA" box="[866,1012,1087,1113]" italics="true" pageId="14" pageNumber="407">A. tokushigei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EF834D13FFC3DD64FBEF7B742DB6FBCA" authorityName="Nomura" authorityYear="1991" box="[1065,1162,1087,1113]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="sakaii">
|
||
<emphasis id="1AF7EA82FFC3DD64FBEF7B742DB6FBCA" box="[1065,1162,1087,1113]" italics="true" pageId="14" pageNumber="407">B. sakaii</emphasis>
|
||
</taxonomicName>
|
||
based solely on external generic features.
|
||
</paragraph>
|
||
<paragraph id="283C3690FFC3DD64FF017BCC2D4BFAC6" blockId="14.[151,1437,151,1365]" pageId="14" pageNumber="407">
|
||
Specimens of
|
||
<taxonomicName id="EF834D13FFC3DD64FEA07BCC28C4FB32" authorityName="Jałoszyński" authorityYear="2023" box="[358,504,1159,1185]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="tokushigei">
|
||
<emphasis id="1AF7EA82FFC3DD64FEA07BCC28C4FB32" box="[358,504,1159,1185]" italics="true" pageId="14" pageNumber="407">A. tokushigei</emphasis>
|
||
</taxonomicName>
|
||
were collected by sifting leaf litter and rotten wood in a subtropical forest during a rather dry early spring, followed by extraction in a Winkler apparatus. Among many accompanying beetles found in the same sample of the substrate were
|
||
<taxonomicName id="EF834D13FFC3DD64FDBF7B842A25FB7A" box="[633,793,1231,1257]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="subFamily" subFamily="Euaesthetinae">Euaesthetinae</taxonomicName>
|
||
:
|
||
<taxonomicName id="EF834D13FFC3DD64FCE07B842D90FB7A" authority="Puthz" authorityName="Puthz" authorityYear="1986" box="[806,1196,1231,1257]" class="Insecta" family="Staphylinidae" genus="Stenaesthetus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="okinawaensis">
|
||
<emphasis id="1AF7EA82FFC3DD64FCE07B842D63FB7A" box="[806,1119,1231,1257]" italics="true" pageId="14" pageNumber="407">Stenaesthetus okinawaensis</emphasis>
|
||
Puthz
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="EF834D13FFC3DD64FB7C7B842C05FB7A" box="[1210,1337,1231,1257]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="subFamily" subFamily="Paederinae">Paederinae</taxonomicName>
|
||
:
|
||
<taxonomicName id="EF834D13FFC3DD64FA817B8428E1FA9E" authority="Ito" authorityName="Ito" class="Insecta" family="Staphylinidae" genus="Nazeris" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="subSpecies" species="okinawanus" subSpecies="okinawanus">
|
||
<emphasis id="1AF7EA82FFC3DD64FA817B842893FA9E" italics="true" pageId="14" pageNumber="407">Nazeris okinawanus okinawanus</emphasis>
|
||
Ito
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EF834D13FFC3DD64FE2B7BB82A99FA9E" authority="Watanabe" authorityName="Watanabe" box="[493,933,1267,1293]" class="Insecta" family="Staphylinidae" genus="Ochthephilum" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="okinawaense">
|
||
<emphasis id="1AF7EA82FFC3DD64FE2B7BB82A14FA9E" box="[493,808,1267,1293]" italics="true" pageId="14" pageNumber="407">Ochthephilum okinawaense</emphasis>
|
||
Watanabe
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="EF834D13FFC3DD64FC737BB82D03FA9E" box="[949,1087,1267,1293]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="subFamily" subFamily="Pselaphinae">Pselaphinae</taxonomicName>
|
||
:
|
||
<taxonomicName id="EF834D13FFC3DD64FB887BB82C0EFA9E" authority="Arai" authorityName="Arai" box="[1102,1330,1267,1293]" class="Insecta" family="Staphylinidae" genus="Morana" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="deigo">
|
||
<emphasis id="1AF7EA82FFC3DD64FB887BB82DCFFA9E" box="[1102,1267,1267,1293]" italics="true" pageId="14" pageNumber="407">Morana deigo</emphasis>
|
||
Arai
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EF834D13FFC3DD64FA847BB8287CFAA2" authority="Arai" authorityName="Arai" class="Insecta" family="Staphylinidae" genus="Morana" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="kazuyoae">
|
||
<emphasis id="1AF7EA82FFC3DD64FA847BB829C3FAA2" italics="true" pageId="14" pageNumber="407">Morana kazuyoae</emphasis>
|
||
Arai
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EF834D13FFC3DD64FE977A5C2A31FAA2" authority="Nomura" authorityName="Nomura" authorityYear="1991" box="[337,781,1303,1329]" class="Insecta" family="Staphylinidae" genus="Batriscenaulax" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="kunigamensis">
|
||
<emphasis id="1AF7EA82FFC3DD64FE977A5C2B9CFAA2" box="[337,672,1303,1329]" italics="true" pageId="14" pageNumber="407">Batriscenaulax kunigamensis</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EF834D13FFC3DD64FCD87A5C2D47FAA2" authority="Nomura" authorityName="Nomura" authorityYear="1991" box="[798,1147,1303,1329]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="sakaii">
|
||
<emphasis id="1AF7EA82FFC3DD64FCD87A5C2D33FAA2" box="[798,1039,1303,1329]" italics="true" pageId="14" pageNumber="407">Batriscenellus sakaii</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EF834D13FFC3DD64FB4B7A5C29CAFAC6" authority="Nomura" authorityName="Nomura" authorityYear="1986" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="picticornis">
|
||
<emphasis id="1AF7EA82FFC3DD64FB4B7A5C2CA0FAA3" box="[1165,1436,1303,1329]" italics="true" pageId="14" pageNumber="407">Tribasodites picticornis</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EF834D13FFC3DD64FEC77A702B74FAC6" authority="Nomura" authorityName="Nomura" box="[257,584,1339,1365]" class="Insecta" family="Staphylinidae" genus="Trisinus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="monostatos">
|
||
<emphasis id="1AF7EA82FFC3DD64FEC77A7028DEFAC7" box="[257,482,1339,1364]" italics="true" pageId="14" pageNumber="407">Trisinus monostatos</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="EF834D13FFC3DD64FD457A702A1AFAC6" authorityName="Erichson" authorityYear="1839" box="[643,806,1339,1365]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="subFamily" subFamily="Xantholininae">Xantholininae</taxonomicName>
|
||
:
|
||
<taxonomicName id="EF834D13FFC3DD64FCF67A702D4BFAC6" authority="Cameron." authorityName="Cameron." box="[816,1143,1339,1365]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="407" phylum="Arthropoda" rank="species" species="japonicus">
|
||
<emphasis id="1AF7EA82FFC3DD64FCF67A702D3DFAC7" box="[816,1025,1339,1365]" italics="true" pageId="14" pageNumber="407">Diochus japonicus</emphasis>
|
||
Cameron.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |