345 lines
45 KiB
XML
345 lines
45 KiB
XML
<document id="C387AE1098F2E530F950471F7EE52EC5" ID-CLB-Dataset="21670" ID-DOI="10.5852/ejt.2020.664" ID-GBIF-Dataset="70213048-4950-4064-b085-e13bb570ae0a" ID-ISSN="2118-9773" ID-Zenodo-Dep="3898857" ID-ZooBank="C3B7A472-8052-46CF-BF62-755FEDDC33AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1592407223497" checkinUser="valdenar" docAuthor="Bogri, Amalia, Solodovnikov, Alexey & Hansen, Aslak Kappel" docDate="2020" docId="BE518788FF95FFF41981FDF1FE95FB5B" docLanguage="en" docName="ejt-664_hu_borgi_solodovnikov_hansen.pdf.imf" docOrigin="European Journal of Taxonomy 664" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Quedius adilus Bogri & Solodovnikov & Hansen 2020, sp. nov." docType="treatment" docUuid="2C75AA2D-F524-4308-9823-186131633E08" docUuidSource="ZooBank" docVersion="6" lastPageNumber="20" masterDocId="4268FFF0FF84FFE71B0FFF97FFDDFFAB" masterDocTitle="Hypogean Quedius of Taiwan and their biogeographic significance (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="18" updateTime="1698842574081" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="369624EE6114EBF627579A63C4CC5F51" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="966F636C4CD48B318392A4AFDA099224">
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<mods:title id="EC12A1DE23C3071978B1C7F8BFE5993A">Hypogean Quedius of Taiwan and their biogeographic significance (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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</mods:titleInfo>
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<mods:name id="38FFBBEAB15ADA6758BA7EFE19A4727C" type="personal">
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<mods:roleTerm id="41BD8AEFF3FB7D057B0B9A5BA0D8BAD1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B1EFC7CC069BF6ADDEB6795B2E979441">Bogri, Amalia</mods:namePart>
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<mods:nameIdentifier id="4FF60B6576132CA89C2D59D5F6C1D1D0" type="LSID">318FE942-705C-4540-92E1-0D51F63686B8</mods:nameIdentifier>
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<mods:affiliation id="2C53D3C713EB8EB76CB3DCE539D73946">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, Copenhagen 2100, Denmark. & Email: amalia. bogri. 93 @ gmail. com & urn: lsid: zoobank. org: author: 318 FE 942 - 705 C- 4540 - 92 E 1 - 0 D 51 F 63686 B 8</mods:affiliation>
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<mods:nameIdentifier id="DAA2A36C4061C15F5B8215BB82F50A91" type="email">bogri.93@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="238E2D6C2899FB87BBE461D923277B6D" type="personal">
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<mods:role id="AEEB2FAA8F554AF0FB6650019C11FFD8">
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<mods:roleTerm id="90C56EEE51006885DDA907FB1E4F8347">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7378074255374D66326C0782955782D7">Solodovnikov, Alexey</mods:namePart>
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<mods:nameIdentifier id="900B8902091F0A2FC1BF730677C60CD9" type="LSID">E56B0FF6-38F5-48D7-A5C1-0EA071EB4A99</mods:nameIdentifier>
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<mods:affiliation id="C563296DCF1F79E3FD8AAB2EB866067F">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, Copenhagen 2100, Denmark. & Email: asolodovnikov @ snm. ku. dk & urn: lsid: zoobank. org: author: E 56 B 0 FF 6 - 38 F 5 - 48 D 7 - A 5 C 1 - 0 EA 071 EB 4 A 99</mods:affiliation>
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<mods:nameIdentifier id="E616A3B109E44A6D02C95223C154BC8C" type="email">asolodovnikov@snm.ku.dk</mods:nameIdentifier>
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</mods:name>
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<mods:name id="C3E445BA89C07CB398396E1C5807B046" type="personal">
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<mods:role id="97250A442CBFEEA1234AE0CAA642C615">
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<mods:roleTerm id="1FB48FEF730EB9A5FF8FFD766906777D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="441B335BA28D30884015A20C74276E71">Hansen, Aslak Kappel</mods:namePart>
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<mods:nameIdentifier id="031D908DD5AB0FA6A543D9584E24FC24" type="LSID">D1FBF5E1-6876-48EF-978C-FFDADFF7334C</mods:nameIdentifier>
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<mods:affiliation id="42CC3620FAD702D77D32B737A54131D7">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, Copenhagen 2100, Denmark. & Natural History Museum Aarhus, Wilhelm Meyers Allé 10, 8000 Aarhus, Denmark. & Department of Bioscience, University of Aarhus, Ny Munkegade 116, 8000 Aarhus, Denmark. & urn: lsid: zoobank. org: author: D 1 FBF 5 E 1 - 6876 - 48 EF- 978 C-FFDADFF 7334 C & Corresponding author: akhansen @ snm. ku. dk</mods:affiliation>
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<mods:nameIdentifier id="67CA62FE5B37FCA537ACD183C422FE44" type="email">akhansen@snm.ku.dk</mods:nameIdentifier>
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<mods:typeOfResource id="A67F3DABB866FA9AB23072E8F96296B4">text</mods:typeOfResource>
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<mods:title id="92F905FBB6022F78B5B913B32855D2F3">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="49C9DA1DAF6BDC2512057FCE89A8C6A2">
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<mods:date id="C17BAE4E318FC6C5FE70B897A343B31E">2020</mods:date>
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<mods:number id="BCBFE3D5A10491A2E0A5B436D11B8E49">2020-06-11</mods:number>
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<mods:number id="CF91A85F08284271ED0561C6B59CEAF4">664</mods:number>
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<mods:classification id="99E9B7164F1644665C8F5032B503648F">journal article</mods:classification>
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<mods:identifier id="14AB98D2C1FACCBDE7923FF48C979DAA" type="DOI">10.5852/ejt.2020.664</mods:identifier>
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<mods:identifier id="86F9FDEA3866BEB095A3AC07CA8F2110" type="ISSN">2118-9773</mods:identifier>
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<treatment id="BE518788FF95FFF41981FDF1FE95FB5B" ID-GBIF-Taxon="164612891" LSID="urn:lsid:zoobank.org:act:2C75AA2D-F524-4308-9823-186131633E08" httpUri="http://treatment.plazi.org/id/BE518788FF95FFF41981FDF1FE95FB5B" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
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<subSubSection id="7EE26515FF95FFF61981FDF1FC7BFD2A" box="[654,934,614,641]" pageId="17" pageNumber="18" type="nomenclature">
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<paragraph id="3647369EFF95FFF61981FDF1FC7BFD2A" blockId="17.[378,1209,614,711]" box="[654,934,614,641]" pageId="17" pageNumber="18">
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<heading id="6D0F81F2FF95FFF61981FDF1FC7BFD2A" box="[654,934,614,641]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
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<taxonomicName id="F1F84D1DFF95FFF61981FDF1FC9EFD2A" ID-CoL="8Y93B" authority="Bogri & Solodovnikov & Hansen, 2020" authorityName="Bogri & Solodovnikov & Hansen" authorityYear="2020" box="[654,835,614,641]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="adilus" status="sp. nov.">
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<emphasis id="048CEA8CFF95FFF61981FDF1FC9EFD2A" bold="true" box="[654,835,614,641]" italics="true" pageId="17" pageNumber="18">Quedius adilus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1FBF57F7FF95FFF61844FDF0FC7BFD2A" box="[843,934,615,641]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="7EE26515FF95FFF61A75FD1DFCA7FD6C" pageId="17" pageNumber="18" type="description">
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<paragraph id="3647369EFF95FFF61A75FD1DFB64FD0E" blockId="17.[378,1209,614,711]" box="[378,1209,650,677]" pageId="17" pageNumber="18">
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<uri id="42693A9CFF95FFF61A75FD1DFB64FD0E" box="[378,1209,650,677]" pageId="17" pageNumber="18">
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urn:lsid:zoobank.org:act:
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<uuid id="425E0C4BFF95FFF61999FD1DFB64FD0E" box="[662,1209,650,677]" pageId="17" pageNumber="18">2C75AA2D-F524-4308-9823-186131633E08</uuid>
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</uri>
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</paragraph>
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<paragraph id="3647369EFF95FFF619B6FD3AFCA7FD6C" blockId="17.[378,1209,614,711]" box="[697,890,685,711]" pageId="17" pageNumber="18">
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<figureCitation id="AEC32A1BFF95FFF619B6FD3AFCD9FD6C" box="[697,772,685,711]" captionStart="Fig" captionStartId="5.[189,232,1824,1850]" captionTargetBox="[221,1368,264,1771]" captionTargetId="figure@5.[220,1368,264,1771]" captionTargetPageId="5" captionText="Fig. 2. Distribution of species in the Quedius abnormalis group in Taiwan. Filled square - refers to the new specimens of adults and larvae of Quedius masuzoi Watanabe, 1989 reported in this paper. Square with cross Oi is the type locality of Quedius masuzoi. Unfilled square D is the type locality of Quedius chiangi Smetana, 1995, junior synonym of Q. masuzoi. Question mark (?) refers to an undescribed female specimen reported in Watanabe (1989). Unfilled circle O is the type locality of Quedius nishikawai Watanabe, 1991. Filled star * is the type locality of Quedius adilus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3898861" httpUri="https://zenodo.org/record/3898861/files/figure.png" pageId="17" pageNumber="18">Figs 2</figureCitation>
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,
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<figureCitation id="AEC32A1BFF95FFF6181FFD3AFC8FFD6C" box="[784,850,685,711]" captionStart="Fig" captionStartId="7.[189,232,1755,1781]" captionTargetBox="[249,1339,264,1705]" captionTargetId="figure@7.[249,1339,264,1705]" captionTargetPageId="7" captionText="Fig. 3. Aedeagi and female terminalia of species of the Quedius abnormalis group from Taiwan. The graphic symbol on each illustration refers to the collection locality on Fig. 2 as follows: filled square - to the new specimens of Quedius masuzoi Watanabe, 1989 reported in this paper; square with cross Oi to the male holotype of Quedius masuzoi; unfilled square D to the female holotype of Quedius chiangi Smetana, 1995, synonym of Q. masuzoi; unfilled circle O to the female holotype of Quedius nishikawai Watanabe, 1991; filled star * to the male holotype of Quedius adilus sp. nov. Top row (A–C): parameral view of aedeagus. Middle row (D–F): lateral view of aedeagus. Bottom row (G–I): female tergite X. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3898865" httpUri="https://zenodo.org/record/3898865/files/figure.png" pageId="17" pageNumber="18">3C, F</figureCitation>
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,
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||
<figureCitation id="AEC32A1BFF95FFF61851FD3AFCA7FD6C" box="[862,890,685,711]" captionStart="Fig" captionStartId="18.[189,232,1965,1991]" captionTargetBox="[193,1389,693,1930]" captionTargetId="figure@18.[189,1399,675,1944]" captionTargetPageId="18" captionText="Fig. 11. Quedius adilus sp. nov. A. Habitus, legs and antennae of left side cropped out of image to preserve space. B. Aedeagus in lateral view. C. Aedeagus in parameral view. Scale bar: A = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3898883" httpUri="https://zenodo.org/record/3898883/files/figure.png" pageId="17" pageNumber="18">11</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="7EE26515FF95FFF61BB2FD78FE43FCD6" pageId="17" pageNumber="18" type="etymology">
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||
<paragraph id="3647369EFF95FFF61BB2FD78FAABFC9C" blockId="17.[189,324,751,776]" box="[189,1398,751,823]" lastBlockId="17.[189,1399,796,893]" pageId="17" pageNumber="18">
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||
<emphasis id="048CEA8CFF95FFF61BB2FD78FE99FCA2" bold="true" box="[189,324,751,777]" pageId="17" pageNumber="18">Etymology</emphasis>
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The species name
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<taxonomicName id="F1F84D1DFF95FFF61A95FC8AFE3FFC9C" authorityName="Bogri & Solodovnikov & Hansen" authorityYear="2020" box="[410,482,797,823]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="adilus">
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<emphasis id="048CEA8CFF95FFF61A95FC8AFE3FFC9C" box="[410,482,797,823]" italics="true" pageId="17" pageNumber="18">adilus</emphasis>
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</taxonomicName>
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is derived from the Greek word ‘άδηλος’, which means ‘latent’, ‘existing or
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</paragraph>
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<paragraph id="3647369EFF95FFF61BB2FCD6FE43FCD6" blockId="17.[189,1399,796,893]" pageId="17" pageNumber="18">present but concealed’. This name hints at the difficulty of collecting this species, which is hidden in hypogean crevices.</paragraph>
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</subSubSection>
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<subSubSection id="7EE26515FF95FFF61BB2FC32FDE1FB4F" pageId="17" pageNumber="18" type="materials_examined">
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<paragraph id="3647369EFF95FFF61BB2FC32FE77FC14" blockId="17.[189,426,933,959]" box="[189,426,933,959]" pageId="17" pageNumber="18">
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||
<heading id="6D0F81F2FF95FFF61BB2FC32FE77FC14" bold="true" box="[189,426,933,959]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
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<emphasis id="048CEA8CFF95FFF61BB2FC32FE77FC14" bold="true" box="[189,426,933,959]" pageId="17" pageNumber="18">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="3647369EFF95FFF61BE5FC43FE81FC45" blockId="17.[189,1399,980,1252]" box="[234,348,980,1006]" pageId="17" pageNumber="18">
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<emphasis id="048CEA8CFF95FFF61BE5FC43FE81FC45" bold="true" box="[234,348,980,1006]" pageId="17" pageNumber="18">
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<typeStatus id="E943883CFF95FFF61BE5FC43FE81FC45" box="[234,348,980,1006]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="3647369EFF95FFF61BB2FC60FEBFFBFC" blockId="17.[189,1399,980,1252]" pageId="17" pageNumber="18">
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<materialsCitation id="86903CC3FF95FFF61BB2FC60FEBFFBFC" ID-GBIF-Occurrence="2636385302" collectingDate="2004-04-06" collectingDateMax="2004-05-11" collectingDateMin="2004-04-06" collectingMethod="Malaise trap" collectionCode="NMNS" collectorName="C. S. Lin & W. T. Yang" country="Taiwan" determinerName="Det. A. Brunke" location="Yuanfeng" pageId="17" pageNumber="18" specimenCode="NMNS ENT, 6660- 241" specimenCount="1" specimenCount-male="1" stateProvince="Nantou County" typeStatus="holotype">
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<collectingCountry id="4EEF760EFF95FFF61BB2FC60FEF3FBBA" box="[189,302,1015,1041]" name="Taiwan" pageId="17" pageNumber="18">TAIWAN</collectingCountry>
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•
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<specimenCount id="20FEFD17FF95FFF61A44FC62FE82FBBA" box="[331,351,1013,1041]" pageId="17" pageNumber="18" type="male">♂</specimenCount>
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;
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<collectingRegion id="F43CF87CFF95FFF61A7CFC60FDF6FBBA" box="[371,555,1015,1041]" country="Taiwan" name="Nantou" pageId="17" pageNumber="18">Nantou County</collectingRegion>
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,
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<location id="33276045FF95FFF61938FC61FD7AFBBB" LSID="urn:lsid:plazi:treatment:BE518788FF95FFF41981FDF1FE95FB5B:33276045FF95FFF61938FC61FD7AFBBB" box="[567,679,1014,1040]" country="Taiwan" name="Yuanfeng" pageId="17" pageNumber="18" stateProvince="Nantou County">Yuanfeng</location>
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[
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<emphasis id="048CEA8CFF95FFF619B4FC62FD28FBB9" bold="true" box="[699,757,1013,1042]" pageId="17" pageNumber="18">鳶 峰</emphasis>
|
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];
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<date id="4246105EFF95FFF6181EFC61FC46FBBA" box="[785,923,1014,1041]" pageId="17" pageNumber="18" value="2004-04-06">
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<collectingDate id="5202E9B6FF95FFF6181EFC61FC46FBBA" box="[785,923,1014,1041]" pageId="17" pageNumber="18" value="2004-04-06">6 Apr 2004</collectingDate>
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</date>
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–
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<date id="4246105EFF95FFF618A4FC60FB97FBBA" box="[939,1098,1015,1041]" pageId="17" pageNumber="18" value="2004-05-11">
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<collectingDate id="5202E9B6FF95FFF618A4FC60FB97FBBA" box="[939,1098,1015,1041]" pageId="17" pageNumber="18" value="2004-05-11">11 May 2004</collectingDate>
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</date>
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;
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<collectorName id="9B0D5348FF95FFF61F56FC60FB63FBBB" box="[1113,1214,1014,1041]" pageId="17" pageNumber="18">C.S. Lin</collectorName>
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and
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<collectorName id="9B0D5348FF95FFF61FF5FC60FAAAFBBA" box="[1274,1399,1015,1041]" pageId="17" pageNumber="18">W.T. Yang</collectorName>
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leg.; “
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<collectingMethod id="EFB94E89FF95FFF61A0AFB8DFE40FB9F" box="[261,413,1050,1076]" pageId="17" pageNumber="18">Malaise trap</collectingMethod>
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(KCN) /
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<taxonomicName id="F1F84D1DFF95FFF61919FB8DFCF7FB9F" authority="" authorityName="Dejean" authorityYear="1833" box="[534,810,1050,1076]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subGenus" subGenus="Microsaurus">
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<emphasis id="048CEA8CFF95FFF61919FB8DFDA5FB9F" box="[534,632,1050,1076]" italics="true" pageId="17" pageNumber="18">Quedius</emphasis>
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(
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<emphasis id="048CEA8CFF95FFF61983FB8DFCFCFB98" box="[652,801,1050,1075]" italics="true" pageId="17" pageNumber="18">Microsaurus</emphasis>
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||
)
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</taxonomicName>
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sp. Det. A. Brunke 2018”; “
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<specimenCode id="665E9EE5FF95FFF61F88FB8DFF33FBFC" pageId="17" pageNumber="18">NMNS ENT, 6660- 241</specimenCode>
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||
”;
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||
<collectionCode id="50E9AE5BFF95FFF61A0AFBAAFE83FBFC" box="[261,350,1085,1111]" collectionName="Taiwan [Republic of China], Taichung, National Museum of Natural Science" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="17" pageNumber="18">NMNS</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="3647369EFF95FFF61BB2FBF7FDE1FB4F" blockId="17.[189,1399,980,1252]" pageId="17" pageNumber="18">
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Since the originally pinned
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<typeStatus id="E943883CFF95FFF61902FBF7FDA9FBD1" box="[525,628,1120,1146]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
|
||
was remounted on a card in this study, its pronotal disc on the left side bears a hole from the pin. The genital segment and aedeagus were dissected and preserved in glycerin in a microvial pinned with the card. The last segment of the right antenna, as well as the middle and posterior tibiae, are missing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7EE26515FF95FFF51BB2FA9CFEE4FE5F" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
|
||
<paragraph id="3647369EFF95FFF61BB2FA9CFE93FA8E" blockId="17.[189,334,1291,1317]" box="[189,334,1291,1317]" pageId="17" pageNumber="18">
|
||
<heading id="6D0F81F2FF95FFF61BB2FA9CFE93FA8E" bold="true" box="[189,334,1291,1317]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
|
||
<emphasis id="048CEA8CFF95FFF61BB2FA9CFE93FA8E" bold="true" box="[189,334,1291,1317]" pageId="17" pageNumber="18">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="3647369EFF95FFF61BE5FAADFEF7FAFF" blockId="17.[189,1399,1338,1435]" box="[234,298,1338,1364]" pageId="17" pageNumber="18">
|
||
<emphasis id="048CEA8CFF95FFF61BE5FAADFEF7FAFF" bold="true" box="[234,298,1338,1364]" pageId="17" pageNumber="18">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="3647369EFF95FFF61BB2FACAFAE5FA30" blockId="17.[189,1399,1338,1435]" pageId="17" pageNumber="18">MEASUREMENTS. HW: 2.47, HL: 2.12, HW/HL ratio: 1.17, TL/EL ratio: 7, PL: 2.54, PW: 2.83, PL/PW ratio: 0.90, A4L/A4W ratio: 1.25, total body length: 14.46, body length without terminalia: 11.05.</paragraph>
|
||
<paragraph id="3647369EFF95FFF61BB2FA29FC54FA50" blockId="17.[189,1398,1470,1531]" pageId="17" pageNumber="18">
|
||
HABITUS. Dark brown-reddish, with paler apical part of antenna, tibia and tarsi (
|
||
<figureCitation id="AEC32A1BFF95FFF61F5CFA29FB63FA73" box="[1107,1214,1470,1496]" captionStart="Fig" captionStartId="18.[189,232,1965,1991]" captionTargetBox="[193,1389,693,1930]" captionTargetId="figure@18.[189,1399,675,1944]" captionTargetPageId="18" captionText="Fig. 11. Quedius adilus sp. nov. A. Habitus, legs and antennae of left side cropped out of image to preserve space. B. Aedeagus in lateral view. C. Aedeagus in parameral view. Scale bar: A = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3898883" httpUri="https://zenodo.org/record/3898883/files/figure.png" pageId="17" pageNumber="18">Fig. 11A</figureCitation>
|
||
). Large species, dorso-ventrally flattened, with very small eyes and long legs.
|
||
</paragraph>
|
||
<paragraph id="3647369EFF95FFF61BB2F988FC26F9D4" blockId="17.[189,1399,1566,1664]" pageId="17" pageNumber="18">HEAD. Large, slightly wider than long, flattened and with oblique longitudinal ridge behind eyes; two basal punctures on each side (apical one is smaller) and five additional punctures near basal punctures. Antennae long and slender; antennomeres 4–10 more or less spherical.</paragraph>
|
||
<paragraph id="3647369EFF95FFF61BB2F934FA9BF88D" blockId="17.[189,1398,1699,1831]" pageId="17" pageNumber="18">THORAX. Pronotum distinctly narrowing anteriad; dorsal rows each with three punctures; left sublateral row with two punctures; right sublateral row with three punctures; one additional puncture between left dorsal and sublateral rows of punctures; two additional punctures between right dorsal and sublateral rows of punctures. Scutellum sparsely punctured. Elytra shorter than pronotum; hind wings absent.</paragraph>
|
||
<paragraph id="3647369EFF95FFF61BB2F8DDFCC6F8CF" blockId="17.[189,795,1866,1892]" box="[189,795,1866,1892]" pageId="17" pageNumber="18">LEGS. First four segments of tarsus strongly dilated.</paragraph>
|
||
<paragraph id="3647369EFF95FFF61BB2F81FFBD5F843" blockId="17.[189,1399,1927,2024]" pageId="17" pageNumber="18">ABDOMEN. Sternite VIII broad and sub-parallel, with distinct medial emargination apically and five setiferous punctures present at each lateral side. Sternite IX basally slightly oval, apically emarginate. Tergite X with distinct emargination basally, with several setae apically.</paragraph>
|
||
<paragraph id="3647369EFF96FFF51BB1FE9BFC7CFE3B" blockId="18.[190,1399,267,400]" pageId="18" pageNumber="19">
|
||
AEDEAGUS (
|
||
<figureCitation id="AEC32A1BFF96FFF51A43FE9CFE1AFE8D" box="[332,455,267,294]" captionStart="Fig" captionStartId="7.[189,232,1755,1781]" captionTargetBox="[249,1339,264,1705]" captionTargetId="figure@7.[249,1339,264,1705]" captionTargetPageId="7" captionText="Fig. 3. Aedeagi and female terminalia of species of the Quedius abnormalis group from Taiwan. The graphic symbol on each illustration refers to the collection locality on Fig. 2 as follows: filled square - to the new specimens of Quedius masuzoi Watanabe, 1989 reported in this paper; square with cross Oi to the male holotype of Quedius masuzoi; unfilled square D to the female holotype of Quedius chiangi Smetana, 1995, synonym of Q. masuzoi; unfilled circle O to the female holotype of Quedius nishikawai Watanabe, 1991; filled star * to the male holotype of Quedius adilus sp. nov. Top row (A–C): parameral view of aedeagus. Middle row (D–F): lateral view of aedeagus. Bottom row (G–I): female tergite X. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3898865" httpUri="https://zenodo.org/record/3898865/files/figure.png" pageId="18" pageNumber="19">Fig. 3C, F</figureCitation>
|
||
,
|
||
<figureCitation id="AEC32A1BFF96FFF51ADBFE9BFE32FE8D" box="[468,495,268,294]" captionStart="Fig" captionStartId="18.[189,232,1965,1991]" captionTargetBox="[193,1389,693,1930]" captionTargetId="figure@18.[189,1399,675,1944]" captionTargetPageId="18" captionText="Fig. 11. Quedius adilus sp. nov. A. Habitus, legs and antennae of left side cropped out of image to preserve space. B. Aedeagus in lateral view. C. Aedeagus in parameral view. Scale bar: A = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3898883" httpUri="https://zenodo.org/record/3898883/files/figure.png" pageId="18" pageNumber="19">11</figureCitation>
|
||
B–C). Median lobe slightly broader than paramere, apically protruding over paramere (in parameral view), slightly sharpened apically, its subapical protrusion axe-like in lateral view; paramere large, apically bilobed, each lobe with four long apical setae and sparse sensory peg setae, apical part of paramere slightly narrower than basal part.
|
||
</paragraph>
|
||
<paragraph id="3647369EFF96FFF51BE4FE20FE9BFE7A" blockId="18.[190,326,439,500]" box="[235,326,439,465]" pageId="18" pageNumber="19">
|
||
<emphasis id="048CEA8CFF96FFF51BE4FE20FE9BFE7A" bold="true" box="[235,326,439,465]" pageId="18" pageNumber="19">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="3647369EFF96FFF51BB1FE4DFEE4FE5F" blockId="18.[190,326,439,500]" box="[190,313,474,500]" pageId="18" pageNumber="19">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7EE26515FF96FFF41BB1FD8CFE29FD2D" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="diagnosis">
|
||
<paragraph id="3647369EFF96FFF51BB1FD8CFE1AFD9E" blockId="18.[190,455,539,565]" box="[190,455,539,565]" pageId="18" pageNumber="19">
|
||
<heading id="6D0F81F2FF96FFF51BB1FD8CFE1AFD9E" bold="true" box="[190,455,539,565]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
|
||
<emphasis id="048CEA8CFF96FFF51BB1FD8CFE1AFD9E" bold="true" box="[190,455,539,565]" pageId="18" pageNumber="19">Differential diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="3647369EFF96FFF41BB1FDDDFE29FD2D" blockId="18.[190,1399,586,647]" lastBlockId="19.[189,1399,267,646]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
|
||
This new species is similar to
|
||
<taxonomicName id="F1F84D1DFF96FFF51928FDDDFD70FDC8" authorityName="in Watanabe" authorityYear="1989" box="[551,685,586,612]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="masuzoi">
|
||
<emphasis id="048CEA8CFF96FFF51928FDDDFDE1FDCF" box="[551,572,586,612]" italics="true" pageId="18" pageNumber="19">Q</emphasis>
|
||
.
|
||
<emphasis id="048CEA8CFF96FFF51942FDDDFD70FDC8" box="[589,685,586,611]" italics="true" pageId="18" pageNumber="19">masuzoi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F1F84D1DFF96FFF519E6FDDDFC50FDCF" authorityName="Watanabe" authorityYear="1991" box="[745,909,586,612]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="nishikawai">
|
||
<emphasis id="048CEA8CFF96FFF519E6FDDDFD23FDCF" box="[745,766,586,612]" italics="true" pageId="18" pageNumber="19">Q</emphasis>
|
||
.
|
||
<emphasis id="048CEA8CFF96FFF51801FDDDFC50FDCF" box="[782,909,586,612]" italics="true" pageId="18" pageNumber="19">nishikawai</emphasis>
|
||
</taxonomicName>
|
||
, from which it differs in the more dorsoventrally flattened head, with an oblique longitudinal ridge behind the eyes. Also, unlike both these species,
|
||
<taxonomicName id="F1F84D1DFF97FFF41A2CFE9CFE0AFE8E" authority="Bogri & Solodovnikov & Hansen, 2020" authorityName="Bogri & Solodovnikov & Hansen" authorityYear="2020" box="[291,471,267,293]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="adilus" status="sp. nov.">
|
||
<emphasis id="048CEA8CFF97FFF41A2CFE9CFE0AFE8E" box="[291,471,267,293]" italics="true" pageId="19" pageNumber="20">Quedius adilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1FBF57F7FF97FFF41AEEFE9BFDE3FE8D" box="[481,574,268,294]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has two basal punctures on each side, instead of the usual one, and numerous (five) additional punctures near each basal puncture. Its antennomeres 4–10 are rather spherical and antennomeres 8–10 are relatively shorter than in
|
||
<taxonomicName id="F1F84D1DFF97FFF41890FEC5FB98FEC7" authorityName="Watanabe" authorityYear="1991" box="[927,1093,338,364]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nishikawai">
|
||
<emphasis id="048CEA8CFF97FFF41890FEC5FC69FEC7" box="[927,948,338,364]" italics="true" pageId="19" pageNumber="20">Q</emphasis>
|
||
.
|
||
<emphasis id="048CEA8CFF97FFF418CDFEC5FB98FEC7" box="[962,1093,338,364]" italics="true" pageId="19" pageNumber="20">nishikawai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="048CEA8CFF97FFF41F4AFEC5FB94FEC7" box="[1093,1097,338,364]" italics="true" pageId="19" pageNumber="20">.</emphasis>
|
||
As these characters seem to be highly variable within this species group, reliable identification of the new species should be made through an examination of the male genitalia. Unlike in
|
||
<taxonomicName id="F1F84D1DFF97FFF4185EFE0EFC08FE19" authorityName="in Watanabe" authorityYear="1989" box="[849,981,409,435]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="masuzoi">
|
||
<emphasis id="048CEA8CFF97FFF4185EFE0EFCBBFE18" box="[849,870,409,435]" italics="true" pageId="19" pageNumber="20">Q</emphasis>
|
||
.
|
||
<emphasis id="048CEA8CFF97FFF4187AFE0EFC08FE19" box="[885,981,409,434]" italics="true" pageId="19" pageNumber="20">masuzoi</emphasis>
|
||
</taxonomicName>
|
||
, the median lobe in
|
||
<taxonomicName id="F1F84D1DFF97FFF41FCAFE0FFAAAFE19" authority="Bogri & Solodovnikov & Hansen, 2020" authorityName="Bogri & Solodovnikov & Hansen" authorityYear="2020" box="[1221,1399,408,434]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="adilus" status="sp. nov.">
|
||
<emphasis id="048CEA8CFF97FFF41FCAFE0FFAAAFE19" box="[1221,1399,408,434]" italics="true" pageId="19" pageNumber="20">Quedius adilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1FBF57F7FF97FFF41BB2FE2BFEC5FE7D" box="[189,280,444,470]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is more narrowly elongate and forms a sharper point at the apex; it strongly protrudes over the apical margin of the bilobed paramere (in parameral view) (compare
|
||
<figureCitation id="AEC32A1BFF97FFF418FCFE48FB48FE52" box="[1011,1173,479,505]" captionStart="Fig" captionStartId="7.[189,232,1755,1781]" captionTargetBox="[249,1339,264,1705]" captionTargetId="figure@7.[249,1339,264,1705]" captionTargetPageId="7" captionText="Fig. 3. Aedeagi and female terminalia of species of the Quedius abnormalis group from Taiwan. The graphic symbol on each illustration refers to the collection locality on Fig. 2 as follows: filled square - to the new specimens of Quedius masuzoi Watanabe, 1989 reported in this paper; square with cross Oi to the male holotype of Quedius masuzoi; unfilled square D to the female holotype of Quedius chiangi Smetana, 1995, synonym of Q. masuzoi; unfilled circle O to the female holotype of Quedius nishikawai Watanabe, 1991; filled star * to the male holotype of Quedius adilus sp. nov. Top row (A–C): parameral view of aedeagus. Middle row (D–F): lateral view of aedeagus. Bottom row (G–I): female tergite X. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3898865" httpUri="https://zenodo.org/record/3898865/files/figure.png" pageId="19" pageNumber="20">Fig. 3C and 3</figureCitation>
|
||
A–B); the elevated dents of its subapical carina are positioned more remotely from its apex (in lateral view) (compare
|
||
<figureCitation id="AEC32A1BFF97FFF41BB2FDB2FE81FD94" box="[189,348,549,575]" captionStart="Fig" captionStartId="7.[189,232,1755,1781]" captionTargetBox="[249,1339,264,1705]" captionTargetId="figure@7.[249,1339,264,1705]" captionTargetPageId="7" captionText="Fig. 3. Aedeagi and female terminalia of species of the Quedius abnormalis group from Taiwan. The graphic symbol on each illustration refers to the collection locality on Fig. 2 as follows: filled square - to the new specimens of Quedius masuzoi Watanabe, 1989 reported in this paper; square with cross Oi to the male holotype of Quedius masuzoi; unfilled square D to the female holotype of Quedius chiangi Smetana, 1995, synonym of Q. masuzoi; unfilled circle O to the female holotype of Quedius nishikawai Watanabe, 1991; filled star * to the male holotype of Quedius adilus sp. nov. Top row (A–C): parameral view of aedeagus. Middle row (D–F): lateral view of aedeagus. Bottom row (G–I): female tergite X. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3898865" httpUri="https://zenodo.org/record/3898865/files/figure.png" pageId="19" pageNumber="20">Fig. 3F and 3</figureCitation>
|
||
D–E). Also, compared to conditions in
|
||
<taxonomicName id="F1F84D1DFF97FFF4183CFDB2FC65FD94" authorityName="in Watanabe" authorityYear="1989" box="[819,952,549,576]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="masuzoi">
|
||
<emphasis id="048CEA8CFF97FFF4183CFDB2FC95FD94" box="[819,840,549,575]" italics="true" pageId="19" pageNumber="20">Q</emphasis>
|
||
.
|
||
<emphasis id="048CEA8CFF97FFF41857FDB1FC65FD94" box="[856,952,550,575]" italics="true" pageId="19" pageNumber="20">masuzoi</emphasis>
|
||
</taxonomicName>
|
||
, the split between the apical lobes of the paramere in
|
||
<taxonomicName id="F1F84D1DFF97FFF41A74FDDFFDF1FDC9" authority="Bogri & Solodovnikov & Hansen, 2020" authorityName="Bogri & Solodovnikov & Hansen" authorityYear="2020" box="[379,556,584,610]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="adilus" status="sp. nov.">
|
||
<emphasis id="048CEA8CFF97FFF41A74FDDFFDF1FDC9" box="[379,556,584,610]" italics="true" pageId="19" pageNumber="20">Quedius adilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1FBF57F7FF97FFF4193CFDDEFD50FDC8" box="[563,653,585,611]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is more narrow and the arrangement of the sensory peg setae is more sparse (
|
||
<figureCitation id="AEC32A1BFF97FFF41A55FDFCFE71FD2D" box="[346,428,619,646]" captionStart="Fig" captionStartId="18.[189,232,1965,1991]" captionTargetBox="[193,1389,693,1930]" captionTargetId="figure@18.[189,1399,675,1944]" captionTargetPageId="18" captionText="Fig. 11. Quedius adilus sp. nov. A. Habitus, legs and antennae of left side cropped out of image to preserve space. B. Aedeagus in lateral view. C. Aedeagus in parameral view. Scale bar: A = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3898883" httpUri="https://zenodo.org/record/3898883/files/figure.png" pageId="19" pageNumber="20">Fig. 11</figureCitation>
|
||
B–C).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="62876616FF96FFF51BB2F83AFAE3F840" ID-DOI="http://doi.org/10.5281/zenodo.3898883" ID-Zenodo-Dep="3898883" httpUri="https://zenodo.org/record/3898883/files/figure.png" pageId="18" pageNumber="19" startId="18.[189,232,1965,1991]" targetBox="[193,1389,693,1930]" targetPageId="18">
|
||
<paragraph id="3647369EFF96FFF51BB2F83AFAE3F840" blockId="18.[189,1399,1965,2027]" pageId="18" pageNumber="19">
|
||
<emphasis id="048CEA8CFF96FFF51BB2F83AFEC6F86C" bold="true" box="[189,283,1965,1991]" pageId="18" pageNumber="19">Fig. 11.</emphasis>
|
||
<taxonomicName id="F1F84D1DFF96FFF51A2AF83AFE07F86C" authority="Bogri & Solodovnikov & Hansen, 2020" authorityName="Bogri & Solodovnikov & Hansen" authorityYear="2020" box="[293,474,1965,1991]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="adilus" status="sp. nov.">
|
||
<emphasis id="048CEA8CFF96FFF51A2AF83AFE07F86C" box="[293,474,1965,1991]" italics="true" pageId="18" pageNumber="19">Quedius adilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1FBF57F7FF96FFF51AEBF83AFD9FF86C" box="[484,578,1965,1991]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="048CEA8CFF96FFF51943F83AFDBCF86C" bold="true" box="[588,609,1965,1991]" pageId="18" pageNumber="19">A</emphasis>
|
||
. Habitus, legs and antennae of left side cropped out of image to preserve space.
|
||
<emphasis id="048CEA8CFF96FFF51A78F847FE56F841" bold="true" box="[375,395,2000,2026]" pageId="18" pageNumber="19">B</emphasis>
|
||
. Aedeagus in lateral view.
|
||
<emphasis id="048CEA8CFF96FFF519C7F847FD00F841" bold="true" box="[712,733,2000,2026]" pageId="18" pageNumber="19">C</emphasis>
|
||
. Aedeagus in parameral view. Scale bar: A = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="7EE26515FF97FFF41BB2FD3AFBB8FD5D" box="[189,1125,685,758]" pageId="19" pageNumber="20" type="distribution">
|
||
<paragraph id="3647369EFF97FFF41BB2FD3AFE8BFD6C" blockId="19.[189,342,685,711]" box="[189,342,685,711]" pageId="19" pageNumber="20">
|
||
<heading id="6D0F81F2FF97FFF41BB2FD3AFE8BFD6C" bold="true" box="[189,342,685,711]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
|
||
<emphasis id="048CEA8CFF97FFF41BB2FD3AFE8BFD6C" bold="true" box="[189,342,685,711]" pageId="19" pageNumber="20">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="3647369EFF97FFF41BB2FD4BFBB8FD5D" blockId="19.[189,1125,731,758]" box="[189,1125,731,758]" pageId="19" pageNumber="20">
|
||
Known only from the
|
||
<typeStatus id="E943883CFF97FFF41ACDFD4BFE28FD5D" box="[450,501,732,758]" pageId="19" pageNumber="20">type</typeStatus>
|
||
locality in
|
||
<collectingRegion id="F43CF87CFF97FFF41976FD4BFCF0FD5D" box="[633,813,732,758]" country="Taiwan" name="Nantou" pageId="19" pageNumber="20">Nantou County</collectingRegion>
|
||
in central
|
||
<collectingCountry id="4EEF760EFF97FFF418A6FD4CFC22FD5E" box="[937,1023,731,757]" name="Taiwan" pageId="19" pageNumber="20">Taiwan</collectingCountry>
|
||
(
|
||
<figureCitation id="AEC32A1BFF97FFF41F00FD4CFB8AFD5D" box="[1039,1111,731,758]" captionStart="Fig" captionStartId="5.[189,232,1824,1850]" captionTargetBox="[221,1368,264,1771]" captionTargetId="figure@5.[220,1368,264,1771]" captionTargetPageId="5" captionText="Fig. 2. Distribution of species in the Quedius abnormalis group in Taiwan. Filled square - refers to the new specimens of adults and larvae of Quedius masuzoi Watanabe, 1989 reported in this paper. Square with cross Oi is the type locality of Quedius masuzoi. Unfilled square D is the type locality of Quedius chiangi Smetana, 1995, junior synonym of Q. masuzoi. Question mark (?) refers to an undescribed female specimen reported in Watanabe (1989). Unfilled circle O is the type locality of Quedius nishikawai Watanabe, 1991. Filled star * is the type locality of Quedius adilus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3898861" httpUri="https://zenodo.org/record/3898861/files/figure.png" pageId="19" pageNumber="20">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7EE26515FF97FFF41BB2FC8AFE95FB5B" pageId="19" pageNumber="20" type="discussion">
|
||
<paragraph id="3647369EFF97FFF41BB2FC8AFEE2FC9C" blockId="19.[189,319,797,823]" box="[189,319,797,823]" pageId="19" pageNumber="20">
|
||
<heading id="6D0F81F2FF97FFF41BB2FC8AFEE2FC9C" bold="true" box="[189,319,797,823]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
|
||
<emphasis id="048CEA8CFF97FFF41BB2FC8AFEE2FC9C" bold="true" box="[189,319,797,823]" pageId="19" pageNumber="20">Bionomics</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="3647369EFF97FFF41BB2FCDCFDC0FC22" blockId="19.[189,1399,843,905]" pageId="19" pageNumber="20">
|
||
The
|
||
<typeStatus id="E943883CFF97FFF41BFDFCDCFE84FCCE" box="[242,345,843,869]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
was collected by Malaise trap, probably accidentally. No information about the habitat is provided on the locality label.
|
||
</paragraph>
|
||
<paragraph id="3647369EFF97FFF41BB2FC27FEF2FC61" blockId="19.[189,303,944,970]" box="[189,303,944,970]" pageId="19" pageNumber="20">
|
||
<heading id="6D0F81F2FF97FFF41BB2FC27FEF2FC61" bold="true" box="[189,303,944,970]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
|
||
<emphasis id="048CEA8CFF97FFF41BB2FC27FEF2FC61" bold="true" box="[189,303,944,970]" pageId="19" pageNumber="20">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="3647369EFF97FFF41BB2FC48FE95FB5B" blockId="19.[189,1399,991,1264]" pageId="19" pageNumber="20">
|
||
As established in this paper, the aedeagus shape in
|
||
<taxonomicName id="F1F84D1DFF97FFF41825FC48FC6DFC53" authorityName="in Watanabe" authorityYear="1989" box="[810,944,991,1017]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="masuzoi">
|
||
<emphasis id="048CEA8CFF97FFF41825FC48FCE2FC52" box="[810,831,991,1017]" italics="true" pageId="19" pageNumber="20">Q</emphasis>
|
||
.
|
||
<emphasis id="048CEA8CFF97FFF4185FFC48FC6DFC53" box="[848,944,991,1016]" italics="true" pageId="19" pageNumber="20">masuzoi</emphasis>
|
||
</taxonomicName>
|
||
is stable, contrary to other characters earlier chosen to delimit that species. Therefore, the clearly visible difference in the shape of the aedeagus between
|
||
<taxonomicName id="F1F84D1DFF97FFF41A29FBB2FE4CFB94" authority="Bogri & Solodovnikov & Hansen, 2020" authorityName="Bogri & Solodovnikov & Hansen" authorityYear="2020" box="[294,401,1061,1088]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="adilus" status="sp. nov.">
|
||
<emphasis id="048CEA8CFF97FFF41A29FBB2FEE6FB94" box="[294,315,1061,1087]" italics="true" pageId="19" pageNumber="20">Q</emphasis>
|
||
.
|
||
<emphasis id="048CEA8CFF97FFF41A46FBB2FE4CFB94" box="[329,401,1061,1087]" italics="true" pageId="19" pageNumber="20">adilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1FBF57F7FF97FFF41A97FBB1FE2FFBEB" box="[408,498,1062,1088]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="F1F84D1DFF97FFF41925FBB2FD70FB94" authorityName="in Watanabe" authorityYear="1989" box="[554,685,1061,1088]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="masuzoi">
|
||
<emphasis id="048CEA8CFF97FFF41925FBB2FDE2FB94" box="[554,575,1061,1087]" italics="true" pageId="19" pageNumber="20">Q</emphasis>
|
||
.
|
||
<emphasis id="048CEA8CFF97FFF41942FBB1FD70FB94" box="[589,685,1062,1087]" italics="true" pageId="19" pageNumber="20">masuzoi</emphasis>
|
||
</taxonomicName>
|
||
, in addition to several external characters separating the new species from
|
||
<taxonomicName id="F1F84D1DFF97FFF41A53FBDEFDDDFBC9" authorityName="Watanabe" authorityYear="1991" box="[348,512,1096,1123]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nishikawai">
|
||
<emphasis id="048CEA8CFF97FFF41A53FBDEFEACFBC8" box="[348,369,1097,1123]" italics="true" pageId="19" pageNumber="20">Q</emphasis>
|
||
.
|
||
<emphasis id="048CEA8CFF97FFF41A8EFBDFFDDDFBC9" box="[385,512,1096,1122]" italics="true" pageId="19" pageNumber="20">nishikawai</emphasis>
|
||
</taxonomicName>
|
||
, advocate for its formal description even based on the available singleton. The
|
||
<typeStatus id="E943883CFF97FFF41BFEFBFBFEF9FB2D" box="[241,292,1132,1158]" pageId="19" pageNumber="20">type</typeStatus>
|
||
locality (Yuangfeng) of the new species is only
|
||
<quantity id="F1009B7BFF97FFF4185BFBFBFC40FB2E" box="[852,925,1131,1158]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="20" unit="km" value="10.0">10 km</quantity>
|
||
away from Nanhua Mt, the
|
||
<typeStatus id="E943883CFF97FFF41FEAFBFBFAC5FB2D" box="[1253,1304,1132,1158]" pageId="19" pageNumber="20">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="F1F84D1DFF97FFF41BEEFB18FEB4FB03" authorityName="in Watanabe" authorityYear="1989" box="[225,361,1167,1193]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="masuzoi">
|
||
<emphasis id="048CEA8CFF97FFF41BEEFB18FF2BFB02" box="[225,246,1167,1193]" italics="true" pageId="19" pageNumber="20">Q</emphasis>
|
||
.
|
||
<emphasis id="048CEA8CFF97FFF41A06FB18FEB4FB03" box="[265,361,1167,1192]" italics="true" pageId="19" pageNumber="20">masuzoi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="AEC32A1BFF97FFF41A72FB18FE15FB02" box="[381,456,1167,1193]" captionStart="Fig" captionStartId="5.[189,232,1824,1850]" captionTargetBox="[221,1368,264,1771]" captionTargetId="figure@5.[220,1368,264,1771]" captionTargetPageId="5" captionText="Fig. 2. Distribution of species in the Quedius abnormalis group in Taiwan. Filled square - refers to the new specimens of adults and larvae of Quedius masuzoi Watanabe, 1989 reported in this paper. Square with cross Oi is the type locality of Quedius masuzoi. Unfilled square D is the type locality of Quedius chiangi Smetana, 1995, junior synonym of Q. masuzoi. Question mark (?) refers to an undescribed female specimen reported in Watanabe (1989). Unfilled circle O is the type locality of Quedius nishikawai Watanabe, 1991. Filled star * is the type locality of Quedius adilus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3898861" httpUri="https://zenodo.org/record/3898861/files/figure.png" pageId="19" pageNumber="20">Fig. 2</figureCitation>
|
||
), which signals that the two species may be sympatric, further corroborating our hypothesis that the aedeagal difference we observe is indicative of a new species different from
|
||
<taxonomicName id="F1F84D1DFF97FFF41BB2FB42FE9CFB44" authorityName="in Watanabe" authorityYear="1989" box="[189,321,1237,1264]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="masuzoi">
|
||
<emphasis id="048CEA8CFF97FFF41BB2FB42FF0FFB44" box="[189,210,1237,1263]" italics="true" pageId="19" pageNumber="20">Q</emphasis>
|
||
.
|
||
<emphasis id="048CEA8CFF97FFF41BEEFB41FE9CFB44" box="[225,321,1238,1263]" italics="true" pageId="19" pageNumber="20">masuzoi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |