treatments-xml/data/A2/37/5B/A2375B97C35B56828B5963F249F49EB7.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1081.75927" ID-GBIF-Dataset="6ebea377-f492-43a0-b87f-eea36d326201" ID-PMC="PMC8782822" ID-Pensoft-Pub="1313-2970-1081-127" ID-Pensoft-UUID="A101072FE2F95283985CF69DC922CED0" ID-PubMed="35087299" ID-ZooBank="AD2D54F556D14DB2B7F8F3B3E0528394" ModsDocID="1313-2970-1081-127" checkinTime="1642207361831" checkinUser="pensoft" docAuthor="Yoshitomi, Hiroyuki &amp; Pham, Thai Hong" docDate="2022" docId="A2375B97C35B56828B5963F249F49EB7" docLanguage="en" docName="ZooKeys 1081: 127-136" docOrigin="ZooKeys 1081" docPubDate="2022-01-14" docSource="http://dx.doi.org/10.3897/zookeys.1081.75927" docTitle="Cacodaemon laotinus subsp. laotinus laotinus (Arrow 1920" docType="treatment" docVersion="4" id="A101072FE2F95283985CF69DC922CED0" lastPageNumber="127" masterDocId="A101072FE2F95283985CF69DC922CED0" masterDocTitle="The genus Cacodaemon (Coleoptera, Endomychidae) of Vietnam" masterLastPageNumber="136" masterPageNumber="127" pageNumber="127" updateTime="1668151305564" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The genus Cacodaemon (Coleoptera, Endomychidae) of Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Yoshitomi, Hiroyuki</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8416-9249</mods:nameIdentifier>
<mods:affiliation>Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, 790 - 8566 Japan</mods:affiliation>
<mods:nameIdentifier type="email">hymushi@agr.ehime-u.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pham, Thai Hong</mods:namePart>
<mods:affiliation>Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, VAST, 321 Huynh Thuc Khang, Hue, Vietnam &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2022</mods:date>
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<mods:number>2022-01-14</mods:number>
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<mods:number>1081</mods:number>
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<mods:start>127</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1081.75927</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1081.75927</mods:identifier>
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<treatment ID-GBIF-Taxon="192143025" LSID="urn:lsid:plazi:treatment:A2375B97C35B56828B5963F249F49EB7" httpUri="http://treatment.plazi.org/id/A2375B97C35B56828B5963F249F49EB7" lastPageNumber="127" pageId="0" pageNumber="127">
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<paragraph pageId="0" pageNumber="127">
<taxonomicName LSID="A2375B97-C35B-5682-8B59-63F249F49EB7" authority="laotinus (Arrow, 1920)" authorityName="laotinus (Arrow" authorityYear="1920" baseAuthorityName="Arrow" baseAuthorityYear="1920" class="Insecta" family="Endomychidae" genus="Cacodaemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacodaemon laotinus subsp. laotinus" order="Coleoptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="laotinus" subSpecies="laotinus">Cacodaemon laotinus laotinus (Arrow, 1920)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="description">
<paragraph pageId="0" pageNumber="127">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, B male C, D-F female. Scales: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure1" httpUri="https://binary.pensoft.net/fig/634063" pageId="0" pageNumber="127">Figs 1A-1D</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Fore tibiae of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure2" httpUri="https://binary.pensoft.net/fig/634064" pageId="0" pageNumber="127">, 2A-2D</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Elytral appendages of Cacodaemon spp. in lateral view A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure3" httpUri="https://binary.pensoft.net/fig/634065" pageId="0" pageNumber="127">, 3A-3D</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Aedeagus of Cacodaemon laotinus laotinus A-D vietnamese specimen E-H laotian specimen. A, E dorsal B, F lateral C, G ventral D, H apical. Abbreviations ab: apical branch; sb: subapical branch. Scales: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure4" httpUri="https://binary.pensoft.net/fig/634066" pageId="0" pageNumber="127">, 4A-4H</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Female genitalia of Cacodaemon spp. A, B C. laotinus laotinus C, D C. vietnamensis sp. nov. E, F C. proavus A, C, E sternite VIII B, D, F ovipositor. Scale: 0.5 mm." figureDoi="10.3897/zookeys.1081.75927.figure5" httpUri="https://binary.pensoft.net/fig/634067" pageId="0" pageNumber="127">, 5A</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Female genitalia of Cacodaemon spp. A, B C. laotinus laotinus C, D C. vietnamensis sp. nov. E, F C. proavus A, C, E sternite VIII B, D, F ovipositor. Scale: 0.5 mm." figureDoi="10.3897/zookeys.1081.75927.figure5" httpUri="https://binary.pensoft.net/fig/634067" pageId="0" pageNumber="127">, 5B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="reference_group">
<paragraph pageId="0" pageNumber="127">
<taxonomicName class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Amphisternus bellicosus subsp. var. var. laotinus" order="Coleoptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="variety" species="bellicosus" subSpecies="var." variety="laotinus">Amphisternus bellicosus var. laotinus</taxonomicName>
Arrow, 1920: 322.
</paragraph>
<paragraph pageId="0" pageNumber="127">
<taxonomicName class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Amphisternus laotinus" order="Coleoptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="laotinus">Amphisternus laotinus</taxonomicName>
: 
<bibRefCitation author="Arrow, GJ" journalOrPublisher="Faune des Colonies Francaises" pageId="0" pageNumber="127" pagination="329 - 357" refId="B2" refString="Arrow, GJ, 1928. Coleopteres Erotylides et Endomychides de L'Indochine Francaise. Faune des Colonies Francaises 2: 329 - 357" title="Coleopteres Erotylides et Endomychides de L'Indochine Francaise." volume="2" year="1928">Arrow 1928</bibRefCitation>
: 343; 
<bibRefCitation author="Strohecker, HF" journalOrPublisher="Zootaxa" pageId="0" pageNumber="127" refId="B6" refString="Strohecker, HF, 1953. Coleoptera, Endomychidae. In: Wytsman P (Ed.) Genera Insectorum. Louis Desmet-Verteneuil, Bruxelles, 1-145." title="Coleoptera, Endomychidae. In: Wytsman P (Ed.) Genera Insectorum. Louis Desmet-Verteneuil, Bruxelles, 1 - 145." year="1953">Strohecker 1953</bibRefCitation>
: 110.
</paragraph>
<paragraph pageId="0" pageNumber="127">
<taxonomicName class="Insecta" family="Endomychidae" genus="Cacodaemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacodaemon laotinus" order="Coleoptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="laotinus">Cacodaemon laotinus</taxonomicName>
: 
<bibRefCitation author="Strohecker, HF" journalOrPublisher="Pacific Insects" pageId="0" pageNumber="127" pagination="319 - 357" refId="B7" refString="Strohecker, HF, 1964. A synopsis of the Amphisternini (Coleoptera: Endomychidae). Pacific Insects 6: 319 - 357" title="A synopsis of the Amphisternini (Coleoptera: Endomychidae)." volume="6" year="1964">Strohecker 1964</bibRefCitation>
: 350; 
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1999.1.1" author="Shockley, FW" journalOrPublisher="Zootaxa" pageId="0" pageNumber="127" pagination="1 - 113" refId="B5" refString="Shockley, FW, Tomaszewska, KW, McHugh, JV, 2009. An annotated checklist of the handsome fungus beetles of the world (Coleoptera: Cucujoidea: Endomychidae). Zootaxa 1999: 1 - 113, DOI: https://doi.org/10.11646/zootaxa.1999.1.1" title="An annotated checklist of the handsome fungus beetles of the world (Coleoptera: Cucujoidea: Endomychidae)." url="https://doi.org/10.11646/zootaxa.1999.1.1" volume="1999" year="2009">Shockley et al. 2009</bibRefCitation>
: 36.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="materials_examined">
<paragraph pageId="0" pageNumber="127">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="127">
<materialsCitation collectingDate="2006-06-25" collectorName="J. Yamasako" country="Laos" location="East Nong Het" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Xieng Khouang Prov.">
<specimenCount type="male">1 male</specimenCount>
&amp; 
<specimenCount type="female">1 female</specimenCount>
(EUMJ), &quot;[
<collectingCountry name="Laos">Laos</collectingCountry>
] 
<location LSID="urn:lsid:plazi:treatment:A2375B97C35B56828B5963F249F49EB7:9867AB942D645864BDAB0242CAFB42F9" country="Laos" name="East Nong Het" stateProvince="Xieng Khouang Prov.">East Nong Het</location>
<collectingRegion country="Laos" name="Xiangkhouang">Xieng Khouang Prov.</collectingRegion>
<collectingDate value="2006-06-25">25. VI. 2006</collectingDate>
<collectorName>J. Yamasako</collectorName>
leg.&quot;
</materialsCitation>
; 
<materialsCitation collectingDate="1996-06" collectorName="T. Kumasawa Collection" country="Vietnam" location="Tam Dao" specimenCount="4" specimenCount-female="1" specimenCount-male="3">
<specimenCount type="male">3 males</specimenCount>
&amp; 
<specimenCount type="female">1 female</specimenCount>
(NIAS, VNMN), &quot; 
<location LSID="urn:lsid:plazi:treatment:A2375B97C35B56828B5963F249F49EB7:6975DAC13955EDFA5892C97B779D1774" country="Vietnam" name="Tam Dao">Tam Dao</location>
N. 
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
<collectingDate value="1996-06">June 1996</collectingDate>
&quot;, &quot; 
<collectorName>T. Kumasawa Collection</collectorName>
&quot;
</materialsCitation>
; 
<materialsCitation collectingDate="1997-05-01" collectorName="T. Kumasawa Collection" country="Vietnam" location="Tam Dao" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1 female</specimenCount>
(NIAS), &quot; 
<location LSID="urn:lsid:plazi:treatment:A2375B97C35B56828B5963F249F49EB7:F0842ED36FA9649B06AD23B8B80F8595" country="Vietnam" name="Tam Dao">Tam Dao</location>
N. 
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
<collectingDate value="1997-05-01">1 May 1997</collectingDate>
&quot;, &quot; 
<collectorName>T. Kumasawa Collection</collectorName>
&quot;
</materialsCitation>
; 
<materialsCitation collectingDate="1994-06" country="Japan" location="Tam Dao" specimenCount="5" specimenCount-female="1" specimenCount-male="4">
<specimenCount type="male">4 males</specimenCount>
&amp; 
<specimenCount type="female">1 female</specimenCount>
(SEHU), &quot; 
<location LSID="urn:lsid:plazi:treatment:A2375B97C35B56828B5963F249F49EB7:760673917999D46E646242EE4E33AF1E" country="Japan" name="Tam Dao">Tam Dao</location>
<collectingDate value="1994-06">VI 1994</collectingDate>
&quot;, &quot;A. TANAKA Coll. (田中 
<normalizedToken originalValue="明">明</normalizedToken>
) Sehu 
<collectingCountry name="Japan">Japan</collectingCountry>
2005&quot;
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="diagnosis">
<paragraph pageId="0" pageNumber="127">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="127">This is a distinct species in the genus by having the following characteristics: elytral appendages, apical parts of femur, and antennomere 1 tinged with orange/dull orange; DA conoidal, not simply pointed.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="redescription">
<paragraph pageId="0" pageNumber="127">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Male.</emphasis>
Body (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, B male C, D-F female. Scales: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure1" httpUri="https://binary.pensoft.net/fig/634063" pageId="0" pageNumber="127">1A</figureCitation>
, 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, B male C, D-F female. Scales: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure1" httpUri="https://binary.pensoft.net/fig/634063" pageId="0" pageNumber="127">1C</figureCitation>
) oval, convex dorsally, weakly shiny. Coloration of body black; BA, DA, PA, humeral parts, and apical parts of femur orange; antennomere 1 I and anterolateral corners of pronotum dull orange.
</paragraph>
<caption doi="10.3897/zookeys.1081.75927.figure1" httpUri="https://binary.pensoft.net/fig/634063" pageId="0" pageNumber="127" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Figure 1.</emphasis>
Habitus of 
<taxonomicName authorityName="Thomson" authorityYear="1857" class="Insecta" family="Endomychidae" genus="Cacodaemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacodaemon" order="Coleoptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="127">Cacodaemon</emphasis>
</taxonomicName>
spp. 
<emphasis bold="true" pageId="0" pageNumber="127">A-D</emphasis>
<taxonomicName authority="laotinus" lsidName="C. laotinus subsp. laotinus" pageId="0" pageNumber="127" rank="subSpecies" species="laotinus" subSpecies="laotinus">
<emphasis italics="true" pageId="0" pageNumber="127">C. laotinus laotinus</emphasis>
</taxonomicName>
, Vietnamese specimens (
<emphasis bold="true" pageId="0" pageNumber="127">A, B</emphasis>
) and Laotian specimens (
<emphasis bold="true" pageId="0" pageNumber="127">C, D</emphasis>
) 
<emphasis bold="true" pageId="0" pageNumber="127">E</emphasis>
<taxonomicName lsidName="C. vietnamensis" pageId="0" pageNumber="127" rank="species" species="vietnamensis">
<emphasis italics="true" pageId="0" pageNumber="127">C. vietnamensis</emphasis>
</taxonomicName>
sp. nov., holotype 
<emphasis bold="true" pageId="0" pageNumber="127">F</emphasis>
<taxonomicName lsidName="C. proavus" pageId="0" pageNumber="127" rank="species" species="proavus">
<emphasis italics="true" pageId="0" pageNumber="127">C. proavus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="127">A, B</emphasis>
male 
<emphasis bold="true" pageId="0" pageNumber="127">C, D-F</emphasis>
female. Scales: 1.0 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="127">
Head finely punctate. Antennae long and slender; antennomere 3 longest, a little shorter than 4 and 5 combined; club (antennomeres 9-11) distinctly wide. Pronotum impunctate, microreticulate, widest at middle; basal and lateral grooves shallow; front corners projecting and pointed; lateral margins slightly tapered anteriorly and posteriorly; posterior corners slightly projecting posterolaterally; PLM/PLS 0.74-0.77 (0.75); PWM/PWA 1.14-1.23 (1.20); PWM/PLM 1.43-1.52 (1.49); PWM/PLS 1.06-1.17 (1.12). Fore tibia (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Fore tibiae of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure2" httpUri="https://binary.pensoft.net/fig/634064" pageId="0" pageNumber="127">2A</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Fore tibiae of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure2" httpUri="https://binary.pensoft.net/fig/634064" pageId="0" pageNumber="127">2C</figureCitation>
) straight and long, slender, with a sharp denticle at middle of inner margin. Elytra (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Elytral appendages of Cacodaemon spp. in lateral view A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure3" httpUri="https://binary.pensoft.net/fig/634065" pageId="0" pageNumber="127">3A</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Elytral appendages of Cacodaemon spp. in lateral view A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure3" httpUri="https://binary.pensoft.net/fig/634065" pageId="0" pageNumber="127">3C</figureCitation>
) finely and sparsely punctate, microreticulate; BA large tubercle; HA long spine, projecting laterally, stout in basal part; DA projecting dorsally, conoidal, with small spine or dull apex, stout in basal part; PA a pair of tubercles; apex of elytra with long spines; EL/EWH 0.90-0.92 (0.91); EL/PLM 3.43-3.51 (3.46); EWH/PWM 2.46-2.67 (2.58); EWH/PWM 1.73-1.67 (1.71); TL/EWH 1.15-1.19 (1.17). Aedeagus (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Aedeagus of Cacodaemon laotinus laotinus A-D vietnamese specimen E-H laotian specimen. A, E dorsal B, F lateral C, G ventral D, H apical. Abbreviations ab: apical branch; sb: subapical branch. Scales: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure4" httpUri="https://binary.pensoft.net/fig/634066" pageId="0" pageNumber="127">4</figureCitation>
) stout; apical branch (ab) short, with acute apex; subapical branch (sb) long, slightly curved, with pointed apex.
</paragraph>
<caption doi="10.3897/zookeys.1081.75927.figure2" httpUri="https://binary.pensoft.net/fig/634064" pageId="0" pageNumber="127" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Figure 2.</emphasis>
Fore tibiae of 
<taxonomicName authorityName="Thomson" authorityYear="1857" class="Insecta" family="Endomychidae" genus="Cacodaemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacodaemon" order="Coleoptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="127">Cacodaemon</emphasis>
</taxonomicName>
spp. 
<emphasis bold="true" pageId="0" pageNumber="127">A-D</emphasis>
<taxonomicName authority="laotinus" lsidName="C. laotinus subsp. laotinus" pageId="0" pageNumber="127" rank="subSpecies" species="laotinus" subSpecies="laotinus">
<emphasis italics="true" pageId="0" pageNumber="127">C. laotinus laotinus</emphasis>
</taxonomicName>
, Vietnamese specimens (
<emphasis bold="true" pageId="0" pageNumber="127">A, B</emphasis>
) and Laotian specimens (
<emphasis bold="true" pageId="0" pageNumber="127">C, D</emphasis>
) 
<emphasis bold="true" pageId="0" pageNumber="127">E</emphasis>
<taxonomicName lsidName="C. vietnamensis" pageId="0" pageNumber="127" rank="species" species="vietnamensis">
<emphasis italics="true" pageId="0" pageNumber="127">C. vietnamensis</emphasis>
</taxonomicName>
sp. nov., holotype 
<emphasis bold="true" pageId="0" pageNumber="127">F</emphasis>
<taxonomicName lsidName="C. proavus" pageId="0" pageNumber="127" rank="species" species="proavus">
<emphasis italics="true" pageId="0" pageNumber="127">C. proavus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="127">A, C</emphasis>
male 
<emphasis bold="true" pageId="0" pageNumber="127">B, D-F</emphasis>
female. Scale: 1.0 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Female</emphasis>
(Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, B male C, D-F female. Scales: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure1" httpUri="https://binary.pensoft.net/fig/634063" pageId="0" pageNumber="127">1B</figureCitation>
, 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, B male C, D-F female. Scales: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure1" httpUri="https://binary.pensoft.net/fig/634063" pageId="0" pageNumber="127">1I</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Elytral appendages of Cacodaemon spp. in lateral view A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure3" httpUri="https://binary.pensoft.net/fig/634065" pageId="0" pageNumber="127">3B</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Elytral appendages of Cacodaemon spp. in lateral view A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure3" httpUri="https://binary.pensoft.net/fig/634065" pageId="0" pageNumber="127">3D</figureCitation>
). Sexual dimorphism distinct in the following characteristics: fore tibia (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Fore tibiae of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure2" httpUri="https://binary.pensoft.net/fig/634064" pageId="0" pageNumber="127">2B</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Fore tibiae of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure2" httpUri="https://binary.pensoft.net/fig/634064" pageId="0" pageNumber="127">2D</figureCitation>
) straight and long, lacking denticle; posterior corners of pronotum right-angled. PLM/PLS 0.73-0.74 (0.74); PWM/PWA 1.26-1.27 (1.27); PWM/PLM 1.52-1.57 (1.54); PWM/PLS 1.13-1.14 (1.14); EL/EWH 0.92; EL/PLM 3.43-3.74 (3.59); EWH/PWM 2.46-2.58 (2.52); EWH/EWM 1.76-1.72 (1.74); TL/EWH 1.17-1.19 (1.18). Sternite VIII (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Female genitalia of Cacodaemon spp. A, B C. laotinus laotinus C, D C. vietnamensis sp. nov. E, F C. proavus A, C, E sternite VIII B, D, F ovipositor. Scale: 0.5 mm." figureDoi="10.3897/zookeys.1081.75927.figure5" httpUri="https://binary.pensoft.net/fig/634067" pageId="0" pageNumber="127">5A</figureCitation>
) deeply emarginate at posterior margin. Ovipositor (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Female genitalia of Cacodaemon spp. A, B C. laotinus laotinus C, D C. vietnamensis sp. nov. E, F C. proavus A, C, E sternite VIII B, D, F ovipositor. Scale: 0.5 mm." figureDoi="10.3897/zookeys.1081.75927.figure5" httpUri="https://binary.pensoft.net/fig/634067" pageId="0" pageNumber="127">5B</figureCitation>
) bearing long setae along lateral margins of fused coxites; posterior margin of fused coxites arcuate; styli bearing long setae.
</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Measurement.</emphasis>
Male from Vietnam (
<emphasis italics="true" pageId="0" pageNumber="127">N</emphasis>
= 3). TL 10.20-10.45 (10.34) mm; PWM 3.30-3.58 (3.44) mm; PWA 2.80-2.90 (2.87) mm; PLM 2.30-2.35 (2.32) mm; PLS 3.05-3.10 (3.08) mm; EL 7.90-8.10 (8.03) mm; EWH 8.80-9.00 (8.87) mm; EWM 5.10-5.40 (5.20) mm. Male from Laos (
<emphasis italics="true" pageId="0" pageNumber="127">N</emphasis>
= 1). TL 10.10 mm; PWM 3.45 mm; PWA 2.80 mm; PLM 2.30 mm; PLS 3.10 mm; EL 7.80 mm; EWH 8.28 mm; EWM 5.00 mm. Female from Vietnam (
<emphasis italics="true" pageId="0" pageNumber="127">N</emphasis>
= 2). TL 10.20-10.90 (10.55) mm; PWM 3.50-3.60 (3.55) mm; PWA 2.75-2.85 (2.80) mm; PLM 2.30 mm; PLS 3.10-3.15 (3.13) mm; EL 7.90-8.60 (8.25) mm; EWH 8.60-9.30 (8.95) mm; EWM 4.90-5.40 (5.15) mm. Female from Laos (
<emphasis italics="true" pageId="0" pageNumber="127">N</emphasis>
= 1). TL 9.00 mm; PWM 3.25 mm; PWA 2.70 mm; PLM 2.00 mm; PLS 3.00 mm; EL 7.00 mm; EWH 7.60 mm; EWM 4.70 mm.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="distribution">
<paragraph pageId="0" pageNumber="127">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="127">Laos, Vietnam (new record).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="remarks">
<paragraph pageId="0" pageNumber="127">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="127">
As already mentioned by 
<bibRefCitation author="Strohecker, HF" journalOrPublisher="Pacific Insects" pageId="0" pageNumber="127" pagination="319 - 357" refId="B7" refString="Strohecker, HF, 1964. A synopsis of the Amphisternini (Coleoptera: Endomychidae). Pacific Insects 6: 319 - 357" title="A synopsis of the Amphisternini (Coleoptera: Endomychidae)." volume="6" year="1964">Strohecker (1964)</bibRefCitation>
, 
<taxonomicName class="Insecta" family="Endomychidae" genus="Cacodaemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacodaemon laotinus subsp. yunnanensis" order="Coleoptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="laotinus" subSpecies="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="127">Cacodaemon laotinus yunnanensis</emphasis>
</taxonomicName>
(Kryzhanovskij, 1960) described from Yunnan (China), is thought to be an infraspecific variation. We do not treat this subspecies in this paper because the holotype could not be examined.
</paragraph>
</subSubSection>
</treatment>
</document>