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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1077.67081" ID-GBIF-Dataset="66b827c0-65ac-44a6-90e0-b78cd9ef81c2" ID-PMC="PMC8692307" ID-Pensoft-Pub="1313-2970-1077-1" ID-Pensoft-UUID="734088641608531C8E2CC69397B000ED" ID-PubMed="35027858" ID-ZooBank="C28AD65A76F242CFBED7DFB3702CABCE" ModsDocID="1313-2970-1077-1" checkinTime="1639570129388" checkinUser="pensoft" docAuthor="De Mattia, Willy, Reier, Susanne &amp; Haring, Elisabeth" docDate="2021" docId="389B22079F0A5AE4A23B9CE1D26A3029" docLanguage="en" docName="ZooKeys 1077: 1-175" docOrigin="ZooKeys 1077" docPubDate="2021-12-14" docSource="http://dx.doi.org/10.3897/zookeys.1077.67081" docTitle="Siciliaria grohmanniana subsp. grohmanniana grohmanniana (Rossmaessler 1836" docType="treatment" docVersion="4" id="734088641608531C8E2CC69397B000ED" lastPageNumber="1" masterDocId="734088641608531C8E2CC69397B000ED" masterDocTitle="Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)" masterLastPageNumber="175" masterPageNumber="1" pageNumber="1" updateTime="1668151235153" updateUser="ExternalLinkService">
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<mods:title>Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>De Mattia, Willy</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0056-467X</mods:nameIdentifier>
<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
<mods:nameIdentifier type="email">willy.demattia@icgeb.org</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reier, Susanne</mods:namePart>
<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Haring, Elisabeth</mods:namePart>
<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-12-14</mods:number>
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<mods:detail type="volume">
<mods:number>1077</mods:number>
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<mods:start>1</mods:start>
<mods:end>175</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1077.67081</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1077.67081</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1077-1</mods:identifier>
<mods:identifier type="ZooBank">C28AD65A76F242CFBED7DFB3702CABCE</mods:identifier>
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<treatment ID-GBIF-Taxon="191645973" LSID="urn:lsid:plazi:treatment:389B22079F0A5AE4A23B9CE1D26A3029" httpUri="http://treatment.plazi.org/id/389B22079F0A5AE4A23B9CE1D26A3029" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="389B2207-9F0A-5AE4-A23B-9CE1D26A3029" authority="grohmanniana (Rossmaessler, 1836)" authorityName="grohmanniana (Rossmaessler" authorityYear="1836" baseAuthorityName="Rossmässler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria grohmanniana subsp. grohmanniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="grohmanniana" subSpecies="grohmanniana">
Siciliaria grohmanniana grohmanniana (
<normalizedToken originalValue="Rossmässler">Rossmaessler</normalizedToken>
, 1836)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Wagner's (1913, 1925) and Brandt's (1961) Siciliaria genital drawings A Siciliaria septemplicata, Wagner, 1913: plate 572, fig. 14 B Siciliaria grohmanniana grohmanniana, Wagner, 1925: plate 1, fig. 8 C Siciliaria calcarae calcarae, 1925: plate 3, fig. 25 D Siciliaria ferrox, Brandt, 1961: fig. 1 E overall Siciliaria / Sicania and Sicilian Mauritanica distribution F Siciliaria / Sicania and Sicilian Mauritanica distribution of taxa." figureDoi="10.3897/zookeys.1077.67081.figure1" httpUri="https://binary.pensoft.net/fig/622793" pageId="0" pageNumber="1">Figs 1.F, 7.1-7.3, 8.1-8.4, 13.1-13.4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Rossmassler" authorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Clausilia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Clausilia grohmanniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="grohmanniana">Clausilia grohmanniana</taxonomicName>
<normalizedToken originalValue="Rossmässler">Rossmaessler</normalizedToken>
1836: 7.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Schmidt" authorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Clausilia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Clausilia grohmanniana subsp. var. var. minor" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="grohmanniana" subSpecies="var." variety="minor">Clausilia grohmanniana var. minor</taxonomicName>
A.
<bibRefCitation author="Schmidt, A" journalOrPublisher="Ruthenica, Supplement" pageId="0" pageNumber="1" refId="B89" refString="Schmidt, A, 1868. System der europaeischen Clausilien und ihrer naechsten Verwandten. Cassel, 175 pp." title="System der europaeischen Clausilien und ihrer naechsten Verwandten. Cassel, 175 pp." year="1868">Schmidt 1868</bibRefCitation>
: 40 [non
<normalizedToken originalValue="Rossmässler">Rossmaessler</normalizedToken>
].
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Clausilia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Clausilia grohmanuiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="grohmanuiana">Clausilia grohmanuiana</taxonomicName>
[sic!] -
<bibRefCitation author="Benoit, L" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B6" refString="Benoit, L, 1876. Illustrazione sistematica critica iconografica de'testacei estramarini della Sicilia ulteriore e delle isole circostanti. Nobile, Napoli, [I-XVI,] 248 pp. [Tav. I-IX, XI-XII]" title="Illustrazione sistematica critica iconografica de'testacei estramarini della Sicilia ulteriore e delle isole circostanti. Nobile, Napoli, [I-XVI,] 248 pp. [Tav. I-IX, XI-XII]" year="1876">Benoit 1876</bibRefCitation>
: 151.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Clausilia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Clausilia grohmaniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="grohmaniana">Clausilia grohmaniana</taxonomicName>
[sic!] -
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.51778" author="Benoit, L" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B7" refString="Benoit, L, 1881. Nuovo catalogo delle conchiglie terrestri e fluviatili della Sicilia. Atti della Reale Accademia Peloritana 3: 43-216. [Sep.: I-VI, 7-176]. https://doi.org/10.5962/bhl.title.51778" title="Nuovo catalogo delle conchiglie terrestri e fluviatili della Sicilia. Atti della Reale Accademia Peloritana 3: 43 - 216. [Sep.: I-VI, 7 - 176]." url="https://doi.org/10.5962/bhl.title.51778" year="1881">Benoit 1881</bibRefCitation>
: 108.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Rossmassler" authorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Clausilia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Clausilia (Siciliaria) grohmanniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="grohmanniana" subGenus="Siciliaria">Clausilia (Siciliaria) grohmanniana</taxonomicName>
-
<bibRefCitation author="Monterosato, TA" journalOrPublisher="Il Naturalista Siciliano" pageId="0" pageNumber="1" pagination="165 - 173" refId="B54" refString="Monterosato, TA, 1894. Conchiglie terrestri viventi e fossili di Monte Pellegrino. Il Naturalista Siciliano 13 (9): 165 - 173" title="Conchiglie terrestri viventi e fossili di Monte Pellegrino." volume="13" year="1894">Monterosato 1894</bibRefCitation>
: 170.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Delima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Delima (Siciliaria) grohmanniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="grohmanniana" subGenus="Siciliaria">Delima (Siciliaria) grohmanniana</taxonomicName>
-
<bibRefCitation author="Wagner, AJ" journalOrPublisher="Annales Zoologici Musei Polonici Historiae Naturalis" pageId="0" pageNumber="1" pagination="1 - 73" refId="B100" refString="Wagner, AJ, 1925. Studien ueber die Systematik, Stammesgeschichte und geographische Verbreitung des Genus Delima (Hartmann) A. J. Wagner. Annales Zoologici Musei Polonici Historiae Naturalis 4 (1): 1 - 73" title="Studien ueber die Systematik, Stammesgeschichte und geographische Verbreitung des Genus Delima (Hartmann) A. J. Wagner." volume="4" year="1925">Wagner 1925</bibRefCitation>
: 67.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="s. l." baseAuthorityName="Rossmaessler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria grohmanniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="grohmanniana">Siciliaria grohmanniana</taxonomicName>
-
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Ruthenica, Supplement" pageId="0" pageNumber="1" pagination="563 - 729" refId="B88" refString="Schileyko, AA, 2000. Treatise on recent terrestrial pulmonate molluscs. 5. Clausiliidae. Ruthenica, Supplement 2: 563 - 729" title="Treatise on recent terrestrial pulmonate molluscs. 5. Clausiliidae." volume="2" year="2000">Schileyko 2000</bibRefCitation>
: 665.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Rossmassler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria grohmanniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="grohmanniana">Charpentieria grohmanniana</taxonomicName>
-
<bibRefCitation DOI="https://doi.org/10.1127/arch.moll/133/2004/185" author="Beckmann, KH" journalOrPublisher="Basteria" pageId="0" pageNumber="1" refId="B4" refString="Beckmann, KH, 2004. Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133(1/2): 185-191. https://doi.org/10.1127/arch.moll/133/2004/185" title="Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133 (1 / 2): 185 - 191." url="https://doi.org/10.1127/arch.moll/133/2004/185" year="2004">Beckmann 2004</bibRefCitation>
: 188.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="s. l." baseAuthorityName="Rossmaessler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria grohmanniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="grohmanniana">Siciliaria grohmanniana</taxonomicName>
-
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" refId="B65" refString="Nordsieck, H, 2007. Worldwide Door Snails (Clausiliidae), recent and fossil. Conchbooks Hackenheim, 214 pp." title="Worldwide Door Snails (Clausiliidae), recent and fossil. Conchbooks Hackenheim, 214 pp." year="2007">Nordsieck 2007</bibRefCitation>
: 53.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="s. l." baseAuthorityName="Rossmaessler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria grohmanniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="grohmanniana">Siciliaria grohmanniana</taxonomicName>
-
<bibRefCitation author="Welter-Schultes, F" journalOrPublisher="Planet Poster Editions, Goettingen" pageId="0" pageNumber="1" refId="B101" refString="Welter-Schultes, F, 2012. European non-marine molluscs, a quide for species identification. Planet Poster Editions, Goettingen" title="European non-marine molluscs, a quide for species identification." year="2012">Welter-Schultes 2012</bibRefCitation>
: 340.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="s. l." baseAuthorityName="Rossmaessler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria grohmanniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="grohmanniana">Siciliaria grohmanniana</taxonomicName>
-
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck 2013b</bibRefCitation>
: 6.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Rossmassler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria grohmanniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="grohmanniana">Charpentieria grohmanniana</taxonomicName>
-
<bibRefCitation author="De Mattia, W" journalOrPublisher="Molecular Ecology Resources" pageId="0" pageNumber="1" publicationUrl="https://www.iucnredlist.org/species/171033/1320304" refId="B14" refString="De Mattia, W, 2017a. Charpentieria grohmanniana. The IUCN Red List of Threatened Species. https://www.iucnredlist.org/species/171033/1320304" title="Charpentieria grohmanniana. The IUCN Red List of Threatened Species." url="https://www.iucnredlist.org/species/171033/1320304" year="2017 a">De Mattia 2017a</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2003-12-20" collectorName="Monte Pellegrino, N, W. De Mattia, J. Macor" country="Italy" elevation="380" latitude="38.18202" location="Palermo" longLatPrecision="1" longitude="13.350462" specimenCount="1" stateProvince="Sicily">
<collectingCountry name="Italy">Italy</collectingCountry>
,
<collectingRegion country="Italy" name="Sicilia">Sicily</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:389B22079F0A5AE4A23B9CE1D26A3029:9BCEF84B8D7A08AAB7229B874F31EB66" country="Italy" latitude="38.18202" longLatPrecision="1" longitude="13.350462" name="Palermo" stateProvince="Sicily">Palermo</location>
,
<collectorName>Monte Pellegrino, N</collectorName>
side of the top plateau,
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.8" unit="m" value="380.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.8" unit="m" value="380.0">380 m</quantity>
asl
</elevation>
,
<geoCoordinate degrees="38" direction="north" minutes="10" orientation="latitude" precision="1" seconds="55.26" value="38.18202">38°10'55.26&quot;N</geoCoordinate>
,
<geoCoordinate degrees="13" direction="east" minutes="21" orientation="longitude" precision="1" seconds="1.66" value="13.350462">13°21'1.66&quot;E</geoCoordinate>
,
<collectorName>W. De Mattia</collectorName>
and
<collectorName>J. Macor</collectorName>
leg.,
<collectingDate value="2003-12-20">20.xii.2003</collectingDate>
. 15 live spm, 3 dissected spm
</materialsCitation>
.
<materialsCitation accessionNumber="MW758886, MW758887, MW757091MW757092, MW758885" collectingDate="2017-04-15" collectorName="Lab ID, W. De Mattia, J. Macor" country="Italy" county="Palermo" elevation="420" latitude="38.167892" location="Santuario Santa Rosalia" longLatPrecision="1" longitude="13.350722" municipality="Monte Pellegrino" specimenCount="1" stateProvince="Sicily">
<collectingCountry name="Italy">Italy</collectingCountry>
,
<collectingRegion country="Italy" name="Sicilia">Sicily</collectingRegion>
,
<collectingCounty>Palermo</collectingCounty>
,
<collectingMunicipality>Monte Pellegrino</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:389B22079F0A5AE4A23B9CE1D26A3029:789A6A560896B0DF02C60D0F1F287A4A" country="Italy" county="Palermo" latitude="38.167892" longLatPrecision="1" longitude="13.350722" municipality="Monte Pellegrino" name="Santuario Santa Rosalia" stateProvince="Sicily">Santuario Santa Rosalia</location>
,
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.2" unit="m" value="420.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.2" unit="m" value="420.0">420 m</quantity>
asl
</elevation>
,
<geoCoordinate degrees="38" direction="north" minutes="10" orientation="latitude" precision="1" seconds="4.41" value="38.167892">38°10'4.41&quot;N</geoCoordinate>
,
<geoCoordinate degrees="13" direction="east" minutes="21" orientation="longitude" precision="1" seconds="2.60" value="13.350722">13°21'2.60&quot;E</geoCoordinate>
, [
<collectorName>Lab ID</collectorName>
42_1, COI:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW758886">MW758886</accessionNumber>
;
<collectorName>Lab ID</collectorName>
42_2, COI:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW758887">MW758887</accessionNumber>
, ITS2:
<accessionNumber isEnumeration="true">MW757091MW757092</accessionNumber>
;
<collectorName>Lab ID</collectorName>
42_3, COI:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW758885">MW758885</accessionNumber>
],
<collectorName>W. De Mattia</collectorName>
and
<collectorName>J. Macor</collectorName>
leg.,
<collectingDate value="2017-04-15">15.iv.2017</collectingDate>
. 12 live spm, 3 dissected spm
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="shell (figs 8.18.4, 13.113.4)">
<paragraph pageId="0" pageNumber="1">
Shell (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Siciliaria grohmanniana grohmanniana (Rossmaessler, 1836), Monte Pellegrino, Palermo 8.1 shell 8.2 shell 8.3 detail of the aperture 8.4 clausiliar plate double side. Siciliaria grohmanniana addaurae ssp. nov. Grotta dell'Addaura, Punta Priola 8.5 shell 8.6 shell 8.7 detail of the aperture 8.8 detail of the columellar side of last whorl 8.9 clausiliar plate double side." figureDoi="10.3897/zookeys.1077.67081.figure8" httpUri="https://binary.pensoft.net/fig/622800" pageId="0" pageNumber="1">8</figureCitation>
.1-8.4, 13.1-13.4).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Shell mostly decollate; whorls ribbed; dorsal keel indistinct or missing; inferior lamella very high; two anterior upper palatal plicae present, upper one mostly separated from upper palatal plica; palatal edge of clausilium plate distally receding, plate gutter-like narrowed, palatal edge against distal end bent upwards and more or less pointed (as in
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck 2013b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements</paragraph>
<paragraph pageId="0" pageNumber="1">
(n = 25, decollate): shell height 19.4
<normalizedToken originalValue="±">+/-</normalizedToken>
0.8, whorl width 5.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2, aperture height 3.9
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2, aperture width 2.7
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="external morphology of the genital organs (fig. 7.1)">
<paragraph pageId="0" pageNumber="1">
External morphology of the genital organs (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Siciliaria grohmanniana grohmanniana (Rossmaessler, 1836), Monte Pellegrino, Palermo 7.1 whole distal genital organs 7.2 internal distal part of genital organs 7.3 penial pseudopapilla detail. Siciliaria grohmanniana addaurae ssp. nov. Grotta dell'Addaura, Punta Priola 7.4 whole distal genital organs 7.5 internal distal part of genital organs 7.6 penial pseudopapilla detail." figureDoi="10.3897/zookeys.1077.67081.figure7" httpUri="https://binary.pensoft.net/fig/622799" pageId="0" pageNumber="1">7</figureCitation>
.1).
</paragraph>
<paragraph pageId="0" pageNumber="1">The FO is longer than the V (FO/V range from 1.6 to 1.8). The VD is thin along its whole course. The FDBC is shorter than the BC+SDBC (FDBC/BC+SDBC range from 0.7 to 0.8). The BC+SDBC is cylindrical or club-like and slightly longer than the V (BC+SDBC/V range 1.3-1.4), with no clear distinction between the SDBC and the BC. The apex is wide and round or pointed. The D is longer than the V (D/V range 2.0-2.1) and longer that the BC+SDBC (D/BC+SDBC range 1.4-1.7), thinner than the BC+SDBC and with a pointed apex. The V is short and cylindrical. The A is large. The PC is longer than the V (P+E/V range 2.4-2.6). The PR is long and robust. There is a clear distinction between P and E as there is a visible ER and a proximal narrowing. The E is thinner but longer than the P (E/P range 1.1-1.3), almost gradually shrinking and turning into the VD.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="internal morphology of the genital organs (figs 7.2, 7.3)">
<paragraph pageId="0" pageNumber="1">
Internal morphology of the genital organs (Figs
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Siciliaria grohmanniana grohmanniana (Rossmaessler, 1836), Monte Pellegrino, Palermo 7.1 whole distal genital organs 7.2 internal distal part of genital organs 7.3 penial pseudopapilla detail. Siciliaria grohmanniana addaurae ssp. nov. Grotta dell'Addaura, Punta Priola 7.4 whole distal genital organs 7.5 internal distal part of genital organs 7.6 penial pseudopapilla detail." figureDoi="10.3897/zookeys.1077.67081.figure7" httpUri="https://binary.pensoft.net/fig/622799" pageId="0" pageNumber="1">7</figureCitation>
.2, 7.3).
</paragraph>
<paragraph pageId="0" pageNumber="1">The A is smooth or with weak traces of the distal penial pleats. The P presents 4-6 longitudinal and very irregular pleats. These pleats are very variable in thickness and sculpture, being both smooth or segmented along the same pleat. These pleats often split (proximally, distally or both) into smaller pleats. The fine structure of the wall is smooth. The PP is big, irregular, wrinkly and pointed. It can be smooth or with small tubercles. The P-E transition presents one ER with the PP and ELP originating from the ER. The epiphallar formula is: 1ER(PP+ELP). The E shows 4 to 6 main longitudinal finely fringed pleats. The V is almost completely smooth. The wall is finely granulated.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authority="grohmanniana" authorityName="grohmanniana (Rossmaessler" authorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria grohmanniana subsp. grohmanniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="grohmanniana" subSpecies="grohmanniana">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria grohmanniana grohmanniana</emphasis>
</taxonomicName>
is widespread and common throughout its range. It is found to live on exposed limestone walls hiding in cracks and under crags. It is also found in
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" lsidName="Pinus" order="Pinales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pinus</emphasis>
</taxonomicName>
sp. forest on decaying woods and tree trunks, including shady and humid spots on mosses and ferns. According to
<bibRefCitation author="De Mattia, W" journalOrPublisher="Molecular Ecology Resources" pageId="0" pageNumber="1" publicationUrl="https://www.iucnredlist.org/species/171033/1320304" refId="B14" refString="De Mattia, W, 2017a. Charpentieria grohmanniana. The IUCN Red List of Threatened Species. https://www.iucnredlist.org/species/171033/1320304" title="Charpentieria grohmanniana. The IUCN Red List of Threatened Species." url="https://www.iucnredlist.org/species/171033/1320304" year="2017 a">De Mattia (2017a)</bibRefCitation>
<taxonomicName authority="grohmanniana" authorityName="grohmanniana (Rossmaessler" authorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria grohmanniana subsp. grohmanniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="grohmanniana" subSpecies="grohmanniana">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria grohmanniana grohmanniana</emphasis>
</taxonomicName>
is considered as Endangered following the IUCN criteria B1ab(i,ii,iii,v)+2ab(i,ii,iii,v). The whole area of distribution is currently included in protected areas.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">The taxon is limited to the calcareous mountains northern of Palermo: Monte Pellegrino and the southern slopes of Monte Gallo.</paragraph>
</subSubSection>
</treatment>
</document>