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<document ID-CLB-Dataset="122795" ID-DOI="http://dx.doi.org/10.3897/asp.80.e73829" ID-GBIF-Dataset="5f938945-980c-433c-942a-f05d39529a25" ID-Pensoft-Pub="1864-8312-80-309" ID-Pensoft-UUID="11907D1B6F665D96B5FD2F9751431366" ID-ZooBank="D5C0468B99A14EF39237D9BC51A8BDA3" ModsDocID="1864-8312-80-309" checkinTime="1660711556436" checkinUser="pensoft" docAuthor="Benedetti, Alipio Rezende &amp; Pinto-da-Rocha, Ricardo" docDate="2022" docId="AB4B8BE579B25EDD928A54C11536A1B0" docLanguage="en" docName="ArthropodSystPhyl 80: 309-388" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-08-16" docSource="http://dx.doi.org/10.3897/asp.80.e73829" docTitle="Huancabamba kubricki gen. et. 2022, sp. nov." docType="treatment" docUuid="1789853E-3528-489A-8F3E-1E4309E56C74" docUuidSource="ZooBank" docVersion="5" id="11907D1B6F665D96B5FD2F9751431366" lastPageNumber="309" masterDocId="11907D1B6F665D96B5FD2F9751431366" masterDocTitle="Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species" masterLastPageNumber="388" masterPageNumber="309" pageNumber="309" updateTime="1732651490011" updateUser="ExternalLinkService">
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<mods:title id="F9575033A847950F42F5ABF3E14F3A59">Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
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<mods:namePart id="780AC6FBB23A065E2B153CC6676307D5">Benedetti, Alipio Rezende</mods:namePart>
<mods:affiliation id="6574BB2C8FE87648AC37C75A9A190EA2">Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil &amp; Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
<mods:nameIdentifier id="7469FF2FED28AAA3F8539B4D20EA3287" type="email">alipiobenedetti@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FA5C30537708D38C260386A7A4A9296F">Pinto-da-Rocha, Ricardo</mods:namePart>
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<mods:affiliation id="CCD119B36E162936848D9786771B0E4C">Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
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<mods:date id="567D7E561D4907F2C8C320FD12C56814">2022</mods:date>
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<treatment id="AB4B8BE579B25EDD928A54C11536A1B0" ID-GBIF-Taxon="199468650" LSID="urn:lsid:zoobank.org:act:1789853E-3528-489A-8F3E-1E4309E56C74" httpUri="http://treatment.plazi.org/id/AB4B8BE579B25EDD928A54C11536A1B0" lastPageNumber="309" pageId="0" pageNumber="309" scope_family="Metasarcidae" scope_order="Opiliones" scope_subOrder="Laniatores">
<subSubSection id="95A3556742DDF91F3F94350D728E9E9D" pageId="0" pageNumber="309" type="nomenclature">
<paragraph id="BEC7C9EB274340225C5FDBD42813FB70" pageId="0" pageNumber="309">
3.29.
<taxonomicName id="6214B5D0682E7625726E1653D0844C74" ID-CoL="9XB4S" LSID="http://zoobank.org/1789853E-3528-489A-8F3E-1E4309E56C74" authority="gen. et." authorityName="gen. et." authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Huancabamba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Huancabamba kubricki" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="kubricki" status="sp. nov.">Huancabamba kubricki gen. et.</taxonomicName>
<taxonomicNameLabel id="CFA3D31BFB342045E7BE854AD531E2F8" pageId="0" pageNumber="309">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="DE6C2002826001C58AC5804E8E98543A" pageId="0" pageNumber="309" type="description">
<paragraph id="33E32E981F2842D8077B2C02AAD8B25E" pageId="0" pageNumber="309">
<figureCitation id="BC4ACA1F5CECC55930C7879D01CD3EFA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus, dorsal of males of Huancabamba gen. nov. and Incasarcus. A H. kubricki gen. et sp. nov.; B I. argenteus Kury &amp; Maury, 1998; C I. dianae Kury &amp; Maury, 1998; D I. ochoai Kury &amp; Maury, 1998; E I. pictus Kury &amp; Maury, 1998; F I. viracocha Kury &amp; Maury, 1998; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure4" httpUri="https://binary.pensoft.net/fig/729878" pageId="0" pageNumber="309">Figs 4A</figureCitation>
<figureCitation id="853930D9FEF05786A362238DE71C0335" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Male trochanter-patella / tibia IV of Huancabamba gen. nov., Lumieria gen. nov. and Tschaidicancha, dorsal and ventral view, respectively. A - B H. kubricki gen. et sp. nov.; C - D L. antonionii gen. et sp. nov.; E - F L. woodyalleni gen. et sp. nov.; G - H T. chaplini sp. nov.; I - J T. joseochoai sp. nov.; K - L T. scorsesei sp. nov.; M - N T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure11" httpUri="https://binary.pensoft.net/fig/729885" pageId="0" pageNumber="309">, 11A, B</figureCitation>
<figureCitation id="56F90D5AD4458BB8FB48BAA4CDD61FB1" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Penis of Huancabamba gen. nov. and Incasarcus, dorsal, lateral and ventral views, respectively. A - C H. kubricki gen. et sp. nov.; D - F I. ochoai Kury &amp; Maury, 1998; G - I I. viracocha Kury &amp; Maury, 1998; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure18" httpUri="https://binary.pensoft.net/fig/729892" pageId="0" pageNumber="309">, 18A-C</figureCitation>
<figureCitation id="971928A0C380482A155DEF1D91184427" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Live specimens of Huancabamba gen. nov. and Lumieria gen. nov. A H. kubricki gen. nov. et sp. nov., male; B female; C L. antonionii gen. nov. sp. nov., male; D female; E L. woodyalleni gen. nov. sp. nov., male; F female." figureDoi="10.3897/asp.80.e73829.figure24" httpUri="https://binary.pensoft.net/fig/729898" pageId="0" pageNumber="309">, 24A, B</figureCitation>
<figureCitation id="7EB965EDC5869B5D682EE6F640D4778E" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Distribution of Huancabamba gen. nov., Lumieria gen. nov. and Tschaidicancha in Peru. A H. kubricki gen. et sp. nov., L. antonionii gen. et sp. nov., T. chaplini sp. nov., T. scorsesei sp. nov., T. weyrauchi; B L. woodyalleni gen. et sp. nov., T. joseochoai sp. nov." figureDoi="10.3897/asp.80.e73829.figure29" httpUri="https://binary.pensoft.net/fig/729903" pageId="0" pageNumber="309">, 29A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="06B3D1117EDE71E124155559D4ED55A1" pageId="0" pageNumber="309" type="description">
<paragraph id="47EC00788333DDB9D564802C2F6504C8" pageId="0" pageNumber="309">Description.</paragraph>
<paragraph id="44F482FDCEA9229B7574291DC9F2ABFE" pageId="0" pageNumber="309">
<emphasis id="DB5541DD0BF504C1F4723580EA84DE69" bold="true" pageId="0" pageNumber="309">
MALE:
<emphasis id="4C1BB75671582A29CB3C2CC66179A8A5" italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis id="64789ACC59FAB86011B82970A395AA0A" italics="true" pageId="0" pageNumber="309">n</emphasis>
=3) DSW: 4.0-4.2 (4.2); DSL: 3.6-4.3 (4.1); CL: 1.5-2.0 (2.0). FIVL: 6.6-7.3 (7.3). ChL: 1.2-3.6 (3.2).
<emphasis id="D0A42B794433BB44EB6AA4BA890EEA6F" bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
: (Fig.
<figureCitation id="B4FBF1C1F240DC7CA4ACD67198ECC838" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Live specimens of Huancabamba gen. nov. and Lumieria gen. nov. A H. kubricki gen. nov. et sp. nov., male; B female; C L. antonionii gen. nov. sp. nov., male; D female; E L. woodyalleni gen. nov. sp. nov., male; F female." figureDoi="10.3897/asp.80.e73829.figure24" httpUri="https://binary.pensoft.net/fig/729898" pageId="0" pageNumber="309">24A</figureCitation>
) Reddish carapace with blackish sides. Dorsal scutum and free tergites brownish orange. Lateral margins of DS, small tubercles on scutal areas, spines on area III and free tergites yellow. A dry-mark on depression of ocularium, carapace, and lateral region to ocularium.
<taxonomicName id="3D1085A1568F986D25FDFE6C681A1D1F" genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">Pedipalpus</taxonomicName>
and chelicerae yellow with small black spots. Legs I-II with coloration similar to dorsal scutum. Legs III-IV reddish brown.
<emphasis id="AD03C682C5CCE926A8FDEFC4024E88DC" bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
: (Fig.
<figureCitation id="EBB02B4FE48F06B374B550B740E9070F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus, dorsal of males of Huancabamba gen. nov. and Incasarcus. A H. kubricki gen. et sp. nov.; B I. argenteus Kury &amp; Maury, 1998; C I. dianae Kury &amp; Maury, 1998; D I. ochoai Kury &amp; Maury, 1998; E I. pictus Kury &amp; Maury, 1998; F I. viracocha Kury &amp; Maury, 1998; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure4" httpUri="https://binary.pensoft.net/fig/729878" pageId="0" pageNumber="309">4A</figureCitation>
) Row of granules on the anterior margin of dorsal scutum and the median elevation. Granules more concentrated on carapace sides and on posterior region to ocularium. Ocularium with a relative strong depr ession; with pair of small tubercles and few granules near the eyes. Dorsal scutum with scutal grooves almost inconspicuous; grooves I and II more visible than others; granules sparsely distributed throughout scutal areas. Areas I-IV with a lateral pair of tubercles; I with a pair of median tubercles, larger than those on the lateral portion of scutal areas; III with a pair of large median blunt spines, base with small tubercles. Lateral margins of DS sparsely covered with granules throughout extension. Posterior margin of DS and free tergites I-III each one with a median pair of tubercles and a row of few granules.
<emphasis id="4BBADB120417AFFCFFD4DB0E05CF8389" bold="true" italics="true" pageId="0" pageNumber="309">Chelicerae</emphasis>
: (Fig.
<figureCitation id="13366BD5A3D31DC262644F07A7B422E5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus, dorsal of males of Huancabamba gen. nov. and Incasarcus. A H. kubricki gen. et sp. nov.; B I. argenteus Kury &amp; Maury, 1998; C I. dianae Kury &amp; Maury, 1998; D I. ochoai Kury &amp; Maury, 1998; E I. pictus Kury &amp; Maury, 1998; F I. viracocha Kury &amp; Maury, 1998; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure4" httpUri="https://binary.pensoft.net/fig/729878" pageId="0" pageNumber="309">4A</figureCitation>
) Swollen in large males, similar to females in the small males. Segment I with three small prolateral tubercles and one large tubercle located medially or retrolaterally. Segment II with small setiferous granules, concentrated more distally; one tooth. Segment III with two teeth.
<taxonomicName id="A99167F6FB4A703D81A9DDC5FEBDCA0A" genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
<emphasis id="63FE2E48BBE3227544E511BA57873ABE" bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
</taxonomicName>
: Trochanter with a ventroapical setiferous tubercle. Femur with a ventral row of 6-7 setiferous tubercles of irregular size, the largest being the most apical; one proapical spine. Patella with a promedian spine. Tibia: retrolateral iiIIIi, prolateral IIII. Tarsus: retrolateral iIiIi, prolateral iIIi.
<emphasis id="D860D12B5C6F04C71ED469E08753AC74" bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
: Coxa I with a median row of 5-6 setiferous tubercles, the most median of which is larger; III-IV with small granules sparsely distributed. Rows of tubercles between the coxae II-III and III-IV. Genital area, anal operculum and free sternites with few granules.
<emphasis id="AEB0DF7CCFB978C7918AFA4B2E052C31" bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
: (Figs
<figureCitation id="B418CFD6A6BA7CF70DBED0C796CC525A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus, dorsal of males of Huancabamba gen. nov. and Incasarcus. A H. kubricki gen. et sp. nov.; B I. argenteus Kury &amp; Maury, 1998; C I. dianae Kury &amp; Maury, 1998; D I. ochoai Kury &amp; Maury, 1998; E I. pictus Kury &amp; Maury, 1998; F I. viracocha Kury &amp; Maury, 1998; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure4" httpUri="https://binary.pensoft.net/fig/729878" pageId="0" pageNumber="309">4A</figureCitation>
,
<figureCitation id="86D7F70F9E3FE502C48A887706298124" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Male trochanter-patella / tibia IV of Huancabamba gen. nov., Lumieria gen. nov. and Tschaidicancha, dorsal and ventral view, respectively. A - B H. kubricki gen. et sp. nov.; C - D L. antonionii gen. et sp. nov.; E - F L. woodyalleni gen. et sp. nov.; G - H T. chaplini sp. nov.; I - J T. joseochoai sp. nov.; K - L T. scorsesei sp. nov.; M - N T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure11" httpUri="https://binary.pensoft.net/fig/729885" pageId="0" pageNumber="309">11A, B</figureCitation>
) Coxa I-II each one with a prolateral and a retrolateral apophysis. Coxa III unarmed. Coxa IV with setiferous granules distributed throughout its surface, with the largest clustered apically. Trochanters I-IV unarmed and with a few granules. Femora I-III unarmed and with few granules. Femur IV unarmed and with more abundant and larger granulation than those of the other femora. Patellae-tibiae I-IV unarmed with few granules. Tarsal segmentation: (
<emphasis id="0A7D1E6DED8BAFDBBDC9636A5E9C7DB1" italics="true" pageId="0" pageNumber="309">n</emphasis>
=3) 7-8 (7), 16-17 (17), 15-17 (17), 16-17 (17).
<emphasis id="17ADE1F0EFE7CC85E9A1988363BAEFE2" bold="true" italics="true" pageId="0" pageNumber="309">Penis</emphasis>
: (18A-C) VP rectangular, with distal margin straight; VP wide and robust in lateral view. MS C1-C13(C14) subapical long and apically curved; MS A1 median short and straight; MS B1-B2 median-basal long and straight (longer than MS A, shorter than MS C); MS D1 very short, placed dorsally, near MS C. Lateral sacs very short (twice as long as wide), apically blunt, with short T3-like microsetae. Stylus with apex dorsoventrally expanded and with apical projections. Dorsal process absent. Promontory convex. -
<emphasis id="62162C691F3C2D01AB7C94CE06EBC058" bold="true" pageId="0" pageNumber="309">
FEMALE:
<emphasis id="876237BB1AEA84FCE546B21AB091352F" italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis id="58A16E686C6DCFC23A622D3BE1DE5041" italics="true" pageId="0" pageNumber="309">n</emphasis>
=3) DSW: 4.1-4.3; DSL: 4.1-4.3; CL: 1.6-1.8 FIVL: 6.9-7.0. ChL: 1.6-1.7. (Fig.
<figureCitation id="5A6E5BADFCD5D46DB3AD4D7BD92CBEE0" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Live specimens of Huancabamba gen. nov. and Lumieria gen. nov. A H. kubricki gen. nov. et sp. nov., male; B female; C L. antonionii gen. nov. sp. nov., male; D female; E L. woodyalleni gen. nov. sp. nov., male; F female." figureDoi="10.3897/asp.80.e73829.figure24" httpUri="https://binary.pensoft.net/fig/729898" pageId="0" pageNumber="309">24B</figureCitation>
) Chelicerae small, not swollen. Femur IV with less dense granulation, with granules smaller than those of males. Tarsal segmentation: (
<emphasis id="4890975FB12F7AE43F5FB82A36AFA043" italics="true" pageId="0" pageNumber="309">n</emphasis>
=3) 7, 13-16, 15-17, 15-17.
</paragraph>
</subSubSection>
<subSubSection id="0DC7ECA0C32D6237E0082766932DDFD5" pageId="0" pageNumber="309" type="diagnosis">
<paragraph id="DA276B85AA0529B59ED531B2EDB89570" pageId="0" pageNumber="309">Diagnosis.</paragraph>
<paragraph id="08C18E77A0DD5EC6E590C64A2A5B1DD7" pageId="0" pageNumber="309">As for the genus.</paragraph>
</subSubSection>
<subSubSection id="06D024695B7663FE3A1FCE6F5318E3ED" pageId="0" pageNumber="309" type="etymology">
<paragraph id="D7A8013925011005127A781B6BC0A15A" pageId="0" pageNumber="309">Derivatio nominis.</paragraph>
<paragraph id="22178DBF783F352C9D450AC6705E33C3" pageId="0" pageNumber="309">The specific epithet of masculine gender, in the genitive form, dedicated to the American director, producer and screenwriter Stanley Kubrick (1928-1999).</paragraph>
</subSubSection>
<subSubSection id="ED0531681B67BCE2E89ECB3E855701B3" pageId="0" pageNumber="309" type="distribution">
<paragraph id="52733773ED6D8460C5C45FB4777350CC" pageId="0" pageNumber="309">Distribution.</paragraph>
<paragraph id="79314211DD92B22C25A4597A6969DD00" pageId="0" pageNumber="309">
(Fig.
<figureCitation id="16E89609F97BCE626F710621F3B176C8" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Distribution of Huancabamba gen. nov., Lumieria gen. nov. and Tschaidicancha in Peru. A H. kubricki gen. et sp. nov., L. antonionii gen. et sp. nov., T. chaplini sp. nov., T. scorsesei sp. nov., T. weyrauchi; B L. woodyalleni gen. et sp. nov., T. joseochoai sp. nov." figureDoi="10.3897/asp.80.e73829.figure29" httpUri="https://binary.pensoft.net/fig/729903" pageId="0" pageNumber="309">29A</figureCitation>
) PERU.
<taxonomicName id="954789AC192181D67FD36DE70AA36CAC" authorityName="Roewer" authorityYear="1952" class="Arachnida" family="Metasarcidae" genus="Cajamarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cajamarca" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Cajamarca</taxonomicName>
. Near Cutervo.
</paragraph>
</subSubSection>
<subSubSection id="3E5BBAB650B9DDA464674EEC05DF8798" pageId="0" pageNumber="309" type="materials_examined">
<paragraph id="B68B08A8265983240D056C25B2635181" pageId="0" pageNumber="309">Material examined.</paragraph>
<paragraph id="DE0FB1199C57AE35448235348E9FC2EB" pageId="0" pageNumber="309">
<emphasis id="B4C45B6768448BBC7D20FA323F94F7E4" bold="true" pageId="0" pageNumber="309">
<emphasis id="5D7E9D4B6E2047763DB238B2B33FA283" italics="true" pageId="0" pageNumber="309">Type material</emphasis>
:
<materialsCitation id="65906095B377CABB7955A23665C1FC08" ID-GBIF-Occurrence="3889641315" collectingDate="2011-05-20" collectorName="R. Pinto-da-Rocha, D. Silva" country="PERU" latitude="-6.3450003" location="Cutervo" longLatPrecision="21" longitude="-78.821945" specimenCount="♂" typeStatus="Holotype">
<typeStatus id="E239C75380D3A03E88BE3F4D8323C364">Holotype</typeStatus>
<specimenCount id="D22E3D2D160D55CCD214F563CF914638"></specimenCount>
, '
<collectingCountry id="7E2C8A0BCAAD2A132987E59C0A76EEA7" name="Peru">PERU</collectingCountry>
,
<taxonomicName id="4240B4FF4E1D7BF2FB1672E5750877A9" authorityName="Roewer" authorityYear="1952" class="Arachnida" family="Metasarcidae" genus="Cajamarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cajamarca" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Cajamarca</taxonomicName>
, near
<location id="47BE1C749C23AF05E95D8A09DC48691A" LSID="urn:lsid:plazi:treatment:AB4B8BE579B25EDD928A54C11536A1B0:47BE1C749C23AF05E95D8A09DC48691A" country="PERU" latitude="-6.3450003" longLatPrecision="21" longitude="-78.821945" name="Cutervo">Cutervo</location>
,
<geoCoordinate id="3439D8E6D38A59D18E7FB86E16155138" degrees="06" direction="south" minutes="20" orientation="latitude" precision="15" seconds="42" value="-6.3450003">06°2042″S</geoCoordinate>
<geoCoordinate id="5CB6851882F82C744F2ACEEDBE9E187F" degrees="78" direction="west" minutes="49" orientation="longitude" precision="15" seconds="19" value="-78.821945">78°4919″W</geoCoordinate>
,
<collectingDate id="AF4352208621FC63E812A3B768BDF81E" value="2011-05-20">20/V/2011</collectingDate>
,
<collectorName id="FC7822BC8C9953C82B1EEAA0B5932D5C">R. Pinto-da-Rocha</collectorName>
&amp;
<collectorName id="829FD62F9E0E798ED530C6F49B19E8A9">D. Silva</collectorName>
leg. (MUSM)
</materialsCitation>
</emphasis>
-
<emphasis id="7EF55B1953718C9F6BC3239736D15FB5" bold="true" pageId="0" pageNumber="309">
<typeStatus id="986DE73700B18E30DC28F29816BA6006">Paratypes</typeStatus>
</emphasis>
<specimenCount id="14978E9EB78B817CD14E933D096B8954" type="female">7 ♀</specimenCount>
,
<normalizedToken id="E2F0AEB7B5DD17D8DE90A852344FF744" originalValue="ditto">'ditto'</normalizedToken>
(MUSM);
<emphasis id="5F768E68141D9E1D609A9210BC1A3FF7" bold="true" pageId="0" pageNumber="309">
<typeStatus id="A17D603C8AEF55CE27221335F7001F1B">Paratypes</typeStatus>
</emphasis>
<specimenCount id="221DED007219EACF26A2EEFCD71A039A" type="male">2 ♂</specimenCount>
,
<specimenCount id="805A2B32CC13DDD1D463790EAA91304E" type="female">7 ♀</specimenCount>
,
<normalizedToken id="D1B0C712A50BE8A1EAA45882908C5B57" originalValue="ditto">'ditto'</normalizedToken>
(MZSP 36989).
</paragraph>
</subSubSection>
</treatment>
</document>