Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species Author Benedetti, Alipio Rezende Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil & Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil alipiobenedetti@gmail.com Author Pinto-da-Rocha, Ricardo https://orcid.org/0000-0002-3959-2205 Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil text Arthropod Systematics & amp; Phylogeny 2022 2022-08-16 80 309 388 http://dx.doi.org/10.3897/asp.80.e73829 journal article http://dx.doi.org/10.3897/asp.80.e73829 1864-8312-80-309 D5C0468B99A14EF39237D9BC51A8BDA3 11907D1B6F665D96B5FD2F9751431366 3.29. Huancabamba kubricki gen. et. sp. nov. Figs 4A , 11A, B , 18A-C , 24A, B , 29A Description. MALE: Measurements ( n =3) DSW: 4.0-4.2 (4.2); DSL: 3.6-4.3 (4.1); CL: 1.5-2.0 (2.0). FIVL: 6.6-7.3 (7.3). ChL: 1.2-3.6 (3.2). Coloration : (Fig. 24A ) Reddish carapace with blackish sides. Dorsal scutum and free tergites brownish orange. Lateral margins of DS, small tubercles on scutal areas, spines on area III and free tergites yellow. A dry-mark on depression of ocularium, carapace, and lateral region to ocularium. Pedipalpus and chelicerae yellow with small black spots. Legs I-II with coloration similar to dorsal scutum. Legs III-IV reddish brown. Dorsum : (Fig. 4A ) Row of granules on the anterior margin of dorsal scutum and the median elevation. Granules more concentrated on carapace sides and on posterior region to ocularium. Ocularium with a relative strong depr ession; with pair of small tubercles and few granules near the eyes. Dorsal scutum with scutal grooves almost inconspicuous; grooves I and II more visible than others; granules sparsely distributed throughout scutal areas. Areas I-IV with a lateral pair of tubercles; I with a pair of median tubercles, larger than those on the lateral portion of scutal areas; III with a pair of large median blunt spines, base with small tubercles. Lateral margins of DS sparsely covered with granules throughout extension. Posterior margin of DS and free tergites I-III each one with a median pair of tubercles and a row of few granules. Chelicerae : (Fig. 4A ) Swollen in large males, similar to females in the small males. Segment I with three small prolateral tubercles and one large tubercle located medially or retrolaterally. Segment II with small setiferous granules, concentrated more distally; one tooth. Segment III with two teeth. Pedipalpus : Trochanter with a ventroapical setiferous tubercle. Femur with a ventral row of 6-7 setiferous tubercles of irregular size, the largest being the most apical; one proapical spine. Patella with a promedian spine. Tibia: retrolateral iiIIIi, prolateral IIII. Tarsus: retrolateral iIiIi, prolateral iIIi. Venter : Coxa I with a median row of 5-6 setiferous tubercles, the most median of which is larger; III-IV with small granules sparsely distributed. Rows of tubercles between the coxae II-III and III-IV. Genital area, anal operculum and free sternites with few granules. Legs : (Figs 4A , 11A, B ) Coxa I-II each one with a prolateral and a retrolateral apophysis. Coxa III unarmed. Coxa IV with setiferous granules distributed throughout its surface, with the largest clustered apically. Trochanters I-IV unarmed and with a few granules. Femora I-III unarmed and with few granules. Femur IV unarmed and with more abundant and larger granulation than those of the other femora. Patellae-tibiae I-IV unarmed with few granules. Tarsal segmentation: ( n =3) 7-8 (7), 16-17 (17), 15-17 (17), 16-17 (17). Penis : (18A-C) VP rectangular, with distal margin straight; VP wide and robust in lateral view. MS C1-C13(C14) subapical long and apically curved; MS A1 median short and straight; MS B1-B2 median-basal long and straight (longer than MS A, shorter than MS C); MS D1 very short, placed dorsally, near MS C. Lateral sacs very short (twice as long as wide), apically blunt, with short T3-like microsetae. Stylus with apex dorsoventrally expanded and with apical projections. Dorsal process absent. Promontory convex. - FEMALE: Measurements ( n =3) DSW: 4.1-4.3; DSL: 4.1-4.3; CL: 1.6-1.8 FIVL: 6.9-7.0. ChL: 1.6-1.7. (Fig. 24B ) Chelicerae small, not swollen. Femur IV with less dense granulation, with granules smaller than those of males. Tarsal segmentation: ( n =3) 7, 13-16, 15-17, 15-17. Diagnosis. As for the genus. Derivatio nominis. The specific epithet of masculine gender, in the genitive form, dedicated to the American director, producer and screenwriter Stanley Kubrick (1928-1999). Distribution. (Fig. 29A ) PERU. Cajamarca . Near Cutervo. Material examined. Type material : Holotype ♂ , ' PERU , Cajamarca , near Cutervo , 06°20′42″S 78°49′19″W , 20/V/2011 , R. Pinto-da-Rocha & D. Silva leg. (MUSM) - Paratypes 7 ♀ , 'ditto' (MUSM); Paratypes 2 ♂ , 7 ♀ , 'ditto' (MZSP 36989).