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<document id="2AF208DCE43EE6CAD27467865E87CF99" ID-CLB-Dataset="26455" ID-DOI="10.11646/zootaxa.4619.2.2" ID-GBIF-Dataset="eee17be5-c4f7-4e12-b9c7-cdf61fd1289b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3249579" ID-ZooBank="E9CFFBFF-E437-4919-9E59-730E87875B62" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1560916788776" checkinUser="plazi" docAuthor="Dumas, Leandro Lourenço &amp; Nessimian, Jorge Luiz" docDate="2019" docId="F556879FE75DFF9C0785FDCE2E0B4CA4" docLanguage="en" docName="zootaxa.4619.2.2.pdf" docOrigin="Zootaxa 4619 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helicopsyche (Feropsyche) shaamunensu Dumas &amp; Nessimian 2019, new species" docType="treatment" docUuid="D30B23D9-C044-4B3C-A913-649AB8FF2CA8" docUuidSource="ZooBank" docVersion="15" lastPageNumber="243" masterDocId="096FFFE7E751FF920712FFCF2B494C5F" masterDocTitle="New species of Helicopsyche von Siebold 1856 (Trichoptera: Helicopsychidae) from Brazil, including the redescription of Helicopsyche (Feropsyche) planorboides Machado 1957" masterLastPageNumber="250" masterPageNumber="231" pageNumber="243" updateTime="1698680714656" updateUser="plazi">
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<mods:title id="7853D53AD5822C1845012408B5C2F642">New species of Helicopsyche von Siebold 1856 (Trichoptera: Helicopsychidae) from Brazil, including the redescription of Helicopsyche (Feropsyche) planorboides Machado 1957</mods:title>
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<mods:namePart id="E6CF2EB48C2422E8297A005B68A634F6">Dumas, Leandro Lourenço</mods:namePart>
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<mods:namePart id="DF684D2F93442550C7366745E0D5A0A7">Nessimian, Jorge Luiz</mods:namePart>
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<treatment id="F556879FE75DFF9C0785FDCE2E0B4CA4" ID-DOI="http://doi.org/10.5281/zenodo.5945279" ID-GBIF-Taxon="158297389" ID-Zenodo-Dep="5945279" LSID="urn:lsid:zoobank.org:act:D30B23D9-C044-4B3C-A913-649AB8FF2CA8" httpUri="http://treatment.plazi.org/id/F556879FE75DFF9C0785FDCE2E0B4CA4" lastPageId="14" lastPageNumber="243" pageId="12" pageNumber="243">
<subSubSection id="35E56502E75DFF9E0785FDCE28664E44" box="[151,815,513,540]" pageId="12" pageNumber="243" type="nomenclature">
<paragraph id="7D403689E75DFF9E0785FDCE28664E44" blockId="12.[151,979,513,604]" box="[151,815,513,540]" pageId="12" pageNumber="243">
<heading id="260881E5E75DFF9E0785FDCE28664E44" bold="true" box="[151,815,513,540]" fontSize="11" level="1" pageId="12" pageNumber="243" reason="1">
<emphasis id="4F8BEA9BE75DFF9E0785FDCE28664E44" bold="true" box="[151,815,513,540]" pageId="12" pageNumber="243">
<taxonomicName id="BAFF4D0AE75DFF9E0785FDCE29D84E43" authority="Dumas &amp; Nessimian, 2019" authorityName="Dumas &amp; Nessimian" authorityYear="2019" box="[151,657,513,540]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="shaamunensu" status="sp. nov." subGenus="Feropsyche">
<emphasis id="4F8BEA9BE75DFF9E0785FDCE2A714E43" bold="true" box="[151,312,513,540]" italics="true" pageId="12" pageNumber="243">Helicopsyche</emphasis>
(
<emphasis id="4F8BEA9BE75DFF9E0658FDCE2A9F4E43" bold="true" box="[330,470,513,540]" italics="true" pageId="12" pageNumber="243">Feropsyche</emphasis>
)
<emphasis id="4F8BEA9BE75DFF9E06F5FDCE29D84E43" bold="true" box="[487,657,513,540]" italics="true" pageId="12" pageNumber="243">shaamunensu</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="54B857E0E75DFF9E058DFDCE28664E44" box="[671,815,513,539]" pageId="12" pageNumber="243" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="35E56502E75DFF9E0785FDEA289A4E03" pageId="12" pageNumber="243" type="description">
<paragraph id="7D403689E75DFF9E0785FDEA2A0C4E1F" blockId="12.[151,979,513,604]" box="[151,325,549,576]" pageId="12" pageNumber="243">
(
<figureCitation id="E5C42A0CE75DFF9E078DFDEA2A404E1F" box="[159,265,549,576]" captionStart="FIGURE 6" captionStartId="13.[151,250,1932,1957]" captionTargetBox="[188,1399,167,1913]" captionTargetId="figure@13.[186,1400,166,1913]" captionTargetPageId="13" captionText="FIGURE 6. Helicopsyche (Feropsyche) shaamunensu, new species. 6A6E, Male genitalia (holotype): 6A, left lateral; 6B, dorsal; 6C, ventral; 6D, phallus, left lateral; 6E, phallus, ventral. 6F, Abdominal sterna VI and VII, left lateral. 6G, Abdominal sternum VI ventral process, ventral. Scales = 0.5 mm." httpUri="https://zenodo.org/record/3249591/files/figure.png" pageId="12" pageNumber="243">Figures 6</figureCitation>
A-G)
</paragraph>
<paragraph id="7D403689E75DFF9E0785FD8D289A4E03" blockId="12.[151,979,513,604]" box="[151,979,578,604]" pageId="12" pageNumber="243">
<uri id="096E3A8BE75DFF9E0785FD8D289A4E03" box="[151,979,578,604]" pageId="12" pageNumber="243">
urn:lsid:zoobank.org:act:
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</paragraph>
</subSubSection>
<subSubSection id="35E56502E75DFF9E0785FD4F2F5D4F58" pageId="12" pageNumber="243" type="diagnosis">
<paragraph id="7D403689E75DFF9E0785FD4F2F5D4F58" blockId="12.[151,1437,640,2035]" pageId="12" pageNumber="243">
<emphasis id="4F8BEA9BE75DFF9E0785FD4F2A434EC4" bold="true" box="[151,266,640,667]" pageId="12" pageNumber="243">Diagnosis</emphasis>
.
<taxonomicName id="BAFF4D0AE75DFF9E0608FD4E29C84EC4" authority="Dumas &amp; Nessimian, 2019" authorityName="Dumas &amp; Nessimian" authorityYear="2019" box="[282,641,641,667]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="shaamunensu" status="sp. nov." subGenus="Feropsyche">
<emphasis id="4F8BEA9BE75DFF9E0608FD4E2AF84EC4" box="[282,433,641,667]" italics="true" pageId="12" pageNumber="243">Helicopsyche</emphasis>
(
<emphasis id="4F8BEA9BE75DFF9E06D0FD4D2A914EC4" box="[450,472,642,667]" italics="true" pageId="12" pageNumber="243">F.</emphasis>
)
<emphasis id="4F8BEA9BE75DFF9E06FBFD4E29C84EC4" box="[489,641,641,667]" italics="true" pageId="12" pageNumber="243">shaamunensu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54B857E0E75DFF9E059BFD4E29AD4EC4" box="[649,740,641,667]" pageId="12" pageNumber="243" rank="species">
<emphasis id="4F8BEA9BE75DFF9E059BFD4E29AD4EC4" bold="true" box="[649,740,641,667]" pageId="12" pageNumber="243">sp. nov.</emphasis>
</taxonomicNameLabel>
resembles
<taxonomicName id="BAFF4D0AE75DFF9E0477FD4E2A5A4EE0" authority="Holzenthal, Bahnik &amp; Calor 2016" authorityName="Holzenthal, Bahnik &amp; Calor" authorityYear="2016" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="lazzariae" subGenus="Feropsyche">
<emphasis id="4F8BEA9BE75DFF9E0477FD4E28B54EC4" box="[869,1020,641,667]" italics="true" pageId="12" pageNumber="243">Helicopsyche</emphasis>
(
<emphasis id="4F8BEA9BE75DFF9E031CFD4D2F6A4EC4" box="[1038,1059,642,667]" italics="true" pageId="12" pageNumber="243">F.</emphasis>
)
<emphasis id="4F8BEA9BE75DFF9E0326FD4E2FD24EC4" box="[1076,1179,641,667]" italics="true" pageId="12" pageNumber="243">lazzariae</emphasis>
Holzenthal, Bahnik &amp; Calor 2016
</taxonomicName>
by the overall deltoid shape of the primary branches of inferior appendages. The new species can be easily distinguished from
<taxonomicName id="BAFF4D0AE75DFF9E0681FD0529004EBC" authorityName="Holzenthal, Bahnik &amp; Calor" authorityYear="2016" box="[403,585,713,739]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="lazzariae" subGenus="Feropsyche">
<emphasis id="4F8BEA9BE75DFF9E0681FD052AE64EBC" box="[403,431,714,739]" italics="true" pageId="12" pageNumber="243">H.</emphasis>
(
<emphasis id="4F8BEA9BE75DFF9E06ACFD052A9A4EBC" box="[446,467,714,739]" italics="true" pageId="12" pageNumber="243">F.</emphasis>
)
<emphasis id="4F8BEA9BE75DFF9E06F0FD0629004EBC" box="[482,585,713,739]" italics="true" pageId="12" pageNumber="243">lazzariae</emphasis>
</taxonomicName>
by the presence of basomesal lobes of inferior appendages, which are absent in the second species, and by the shape and setation of tergum X in dorsal view.
</paragraph>
</subSubSection>
<subSubSection id="35E56502E75DFF9E07D5FCDF2A764B1F" pageId="12" pageNumber="243" type="description">
<paragraph id="7D403689E75DFF9E07D5FCDF29494FBF" blockId="12.[151,1437,640,2035]" pageId="12" pageNumber="243">
<emphasis id="4F8BEA9BE75DFF9E07D5FCDF2A184F74" bold="true" box="[199,337,784,811]" pageId="12" pageNumber="243">Description</emphasis>
.
<emphasis id="4F8BEA9BE75DFF9E064DFCDE2AD54F74" box="[351,412,785,811]" italics="true" pageId="12" pageNumber="243">Adult</emphasis>
. Similar to
<taxonomicName id="BAFF4D0AE75DFF9E050DFCDD299B4F74" authority="Dumas &amp; Nessimian, 2019" authorityName="Dumas &amp; Nessimian" authorityYear="2019" box="[543,722,785,811]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="234" phylum="Arthropoda" rank="species" species="bendego" status="sp. nov." subGenus="Feropsyche">
<emphasis id="4F8BEA9BE75DFF9E050DFCDD297A4F74" box="[543,563,786,811]" italics="true" pageId="12" pageNumber="243">H</emphasis>
. (
<emphasis id="4F8BEA9BE75DFF9E0559FCDD29154F74" box="[587,604,786,811]" italics="true" pageId="12" pageNumber="243">F</emphasis>
.)
<emphasis id="4F8BEA9BE75DFF9E0561FCDE299B4F74" box="[627,722,785,811]" italics="true" pageId="12" pageNumber="243">bendego</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54B857E0E75DFF9E05CBFCDE287B4F74" box="[729,818,785,811]" pageId="12" pageNumber="243" rank="species">
<emphasis id="4F8BEA9BE75DFF9E05CBFCDE287B4F74" bold="true" box="[729,818,785,811]" pageId="12" pageNumber="243">sp. nov.</emphasis>
</taxonomicNameLabel>
except as follows: Overall color (in alcohol) uniformly golden-brown; antennae, palps, and legs pale yellow. Maxillary palps with proximal segment longer than distal one. Each foreleg anterior apical tibial spur about
<date id="09411049E75DFF9E05D8FC9629AC4F2C" box="[714,741,857,883]" pageId="12" pageNumber="243">2x</date>
longer than posterior spur. Male forewings each
<quantity id="BA079B6CE75DFF9E021FFC962E2D4F2C" box="[1293,1380,857,883]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="12" pageNumber="243" unit="mm" value="3.2">3.2 mm</quantity>
long (
<typeStatus id="A244882BE75DFF9E078DFCB22A4A4FC8" box="[159,259,893,919]" pageId="12" pageNumber="243" type="holotype">holotype</typeStatus>
male). Abdominal sternum VI ventral process long, about as long as segment VI, slender, tubular along its length, oriented posteroventrad, slightly wider at apex, nearly straight in lateral view with apex subacute, with lamellae apically (
<figureCitation id="E5C42A0CE75DFF9E0677FC0A2ABA4FBF" box="[357,499,965,992]" captionStart="FIGURE 6" captionStartId="13.[151,250,1932,1957]" captionTargetBox="[188,1399,167,1913]" captionTargetId="figure@13.[186,1400,166,1913]" captionTargetPageId="13" captionText="FIGURE 6. Helicopsyche (Feropsyche) shaamunensu, new species. 6A6E, Male genitalia (holotype): 6A, left lateral; 6B, dorsal; 6C, ventral; 6D, phallus, left lateral; 6E, phallus, ventral. 6F, Abdominal sterna VI and VII, left lateral. 6G, Abdominal sternum VI ventral process, ventral. Scales = 0.5 mm." httpUri="https://zenodo.org/record/3249591/files/figure.png" pageId="12" pageNumber="243">Figs. 6F, 6G</figureCitation>
).
</paragraph>
<paragraph id="7D403689E75DFF9E07D5FC262A764B1F" blockId="12.[151,1437,640,2035]" pageId="12" pageNumber="243">
<emphasis id="4F8BEA9BE75DFF9E07D5FC262A22485C" box="[199,363,1001,1027]" italics="true" pageId="12" pageNumber="243">Male genitalia</emphasis>
. Segment IX short ventrally; in lateral view with anteromesal margin well-developed, anterodorsal margin strongly convex near dorsum, and anteroventral margin slightly convex; lateral apodeme well-developed, located midlaterally in the segment (
<figureCitation id="E5C42A0CE75DFF9E053FFBFE29CE4813" box="[557,647,1073,1100]" captionStart="FIGURE 6" captionStartId="13.[151,250,1932,1957]" captionTargetBox="[188,1399,167,1913]" captionTargetId="figure@13.[186,1400,166,1913]" captionTargetPageId="13" captionText="FIGURE 6. Helicopsyche (Feropsyche) shaamunensu, new species. 6A6E, Male genitalia (holotype): 6A, left lateral; 6B, dorsal; 6C, ventral; 6D, phallus, left lateral; 6E, phallus, ventral. 6F, Abdominal sterna VI and VII, left lateral. 6G, Abdominal sternum VI ventral process, ventral. Scales = 0.5 mm." httpUri="https://zenodo.org/record/3249591/files/figure.png" pageId="12" pageNumber="243">Fig. 6A</figureCitation>
); in dorsal view with anterior margin strongly concave, lateral margins concave (
<figureCitation id="E5C42A0CE75DFF9E0611FB9A2A15482F" box="[259,348,1109,1136]" captionStart="FIGURE 6" captionStartId="13.[151,250,1932,1957]" captionTargetBox="[188,1399,167,1913]" captionTargetId="figure@13.[186,1400,166,1913]" captionTargetPageId="13" captionText="FIGURE 6. Helicopsyche (Feropsyche) shaamunensu, new species. 6A6E, Male genitalia (holotype): 6A, left lateral; 6B, dorsal; 6C, ventral; 6D, phallus, left lateral; 6E, phallus, ventral. 6F, Abdominal sterna VI and VII, left lateral. 6G, Abdominal sternum VI ventral process, ventral. Scales = 0.5 mm." httpUri="https://zenodo.org/record/3249591/files/figure.png" pageId="12" pageNumber="243">Fig. 6B</figureCitation>
); in ventral view subrectangular, with anterior and posterior margins slightly concave (
<figureCitation id="E5C42A0CE75DFF9E0224FB9A2EC6482F" box="[1334,1423,1109,1136]" captionStart="FIGURE 6" captionStartId="13.[151,250,1932,1957]" captionTargetBox="[188,1399,167,1913]" captionTargetId="figure@13.[186,1400,166,1913]" captionTargetPageId="13" captionText="FIGURE 6. Helicopsyche (Feropsyche) shaamunensu, new species. 6A6E, Male genitalia (holotype): 6A, left lateral; 6B, dorsal; 6C, ventral; 6D, phallus, left lateral; 6E, phallus, ventral. 6F, Abdominal sterna VI and VII, left lateral. 6G, Abdominal sternum VI ventral process, ventral. Scales = 0.5 mm." httpUri="https://zenodo.org/record/3249591/files/figure.png" pageId="12" pageNumber="243">Fig. 6C</figureCitation>
). Segment X, in lateral view, subtriangular, strongly tapering to apex from apical 2/3 (
<figureCitation id="E5C42A0CE75DFF9E032BFBB62FDB48CB" box="[1081,1170,1145,1172]" captionStart="FIGURE 6" captionStartId="13.[151,250,1932,1957]" captionTargetBox="[188,1399,167,1913]" captionTargetId="figure@13.[186,1400,166,1913]" captionTargetPageId="13" captionText="FIGURE 6. Helicopsyche (Feropsyche) shaamunensu, new species. 6A6E, Male genitalia (holotype): 6A, left lateral; 6B, dorsal; 6C, ventral; 6D, phallus, left lateral; 6E, phallus, ventral. 6F, Abdominal sterna VI and VII, left lateral. 6G, Abdominal sternum VI ventral process, ventral. Scales = 0.5 mm." httpUri="https://zenodo.org/record/3249591/files/figure.png" pageId="12" pageNumber="243">Fig. 6A</figureCitation>
); in dorsal view slightly narrowed near base, about as long as inferior appendages, mesodorsal borders inverted V-shaped, bearing 13 short, stout setae subapically and pair of clusters of short, stout setae apically, apex rounded on either side of slight mesal notch (
<figureCitation id="E5C42A0CE75DFF9E063DFB2A2AC0495F" box="[303,393,1253,1280]" captionStart="FIGURE 6" captionStartId="13.[151,250,1932,1957]" captionTargetBox="[188,1399,167,1913]" captionTargetId="figure@13.[186,1400,166,1913]" captionTargetPageId="13" captionText="FIGURE 6. Helicopsyche (Feropsyche) shaamunensu, new species. 6A6E, Male genitalia (holotype): 6A, left lateral; 6B, dorsal; 6C, ventral; 6D, phallus, left lateral; 6E, phallus, ventral. 6F, Abdominal sterna VI and VII, left lateral. 6G, Abdominal sternum VI ventral process, ventral. Scales = 0.5 mm." httpUri="https://zenodo.org/record/3249591/files/figure.png" pageId="12" pageNumber="243">Fig. 6B</figureCitation>
). Superior appendages originate dorsolaterally, setose, rounded in lateral view, club-like and laterally directed in dorsal view (
<figureCitation id="E5C42A0CE75DFF9E051EFAC629EB497B" box="[524,674,1289,1316]" captionStart="FIGURE 6" captionStartId="13.[151,250,1932,1957]" captionTargetBox="[188,1399,167,1913]" captionTargetId="figure@13.[186,1400,166,1913]" captionTargetPageId="13" captionText="FIGURE 6. Helicopsyche (Feropsyche) shaamunensu, new species. 6A6E, Male genitalia (holotype): 6A, left lateral; 6B, dorsal; 6C, ventral; 6D, phallus, left lateral; 6E, phallus, ventral. 6F, Abdominal sterna VI and VII, left lateral. 6G, Abdominal sternum VI ventral process, ventral. Scales = 0.5 mm." httpUri="https://zenodo.org/record/3249591/files/figure.png" pageId="12" pageNumber="243">Figs. 6A, 6B</figureCitation>
). Basal plate of inferior appendages triangular in lateral and ventral views, tapering to apex, slightly surpassing anteroventral margin of segment IX (
<figureCitation id="E5C42A0CE75DFF9E0305FAE22FE04917" box="[1047,1193,1325,1352]" captionStart="FIGURE 6" captionStartId="13.[151,250,1932,1957]" captionTargetBox="[188,1399,167,1913]" captionTargetId="figure@13.[186,1400,166,1913]" captionTargetPageId="13" captionText="FIGURE 6. Helicopsyche (Feropsyche) shaamunensu, new species. 6A6E, Male genitalia (holotype): 6A, left lateral; 6B, dorsal; 6C, ventral; 6D, phallus, left lateral; 6E, phallus, ventral. 6F, Abdominal sterna VI and VII, left lateral. 6G, Abdominal sternum VI ventral process, ventral. Scales = 0.5 mm." httpUri="https://zenodo.org/record/3249591/files/figure.png" pageId="12" pageNumber="243">Figs. 6A, 6C</figureCitation>
); primary branches of inferior appendages large, covered by long, stout setae; in lateral view each deltoid, with distal portion about 2.5x broader than its proximal portion, dorsal margin slightly convex, ventral margin slightly convex near base, with 3 stout setae, and strongly convex near apex, with 56 stout setae, posterodorsal margin concave, slightly undulated, with sparse long, stout setae along apical margin, apex pointed (
<figureCitation id="E5C42A0CE75DFF9E0465FA72289B4987" box="[887,978,1469,1496]" captionStart="FIGURE 6" captionStartId="13.[151,250,1932,1957]" captionTargetBox="[188,1399,167,1913]" captionTargetId="figure@13.[186,1400,166,1913]" captionTargetPageId="13" captionText="FIGURE 6. Helicopsyche (Feropsyche) shaamunensu, new species. 6A6E, Male genitalia (holotype): 6A, left lateral; 6B, dorsal; 6C, ventral; 6D, phallus, left lateral; 6E, phallus, ventral. 6F, Abdominal sterna VI and VII, left lateral. 6G, Abdominal sternum VI ventral process, ventral. Scales = 0.5 mm." httpUri="https://zenodo.org/record/3249591/files/figure.png" pageId="12" pageNumber="243">Fig. 6A</figureCitation>
); in dorsal view curved mesad, tapering apically, dorsal margin projecting at midlength, bearing several stout setae along apical margin, apex subacute, with single short, stout seta (
<figureCitation id="E5C42A0CE75DFF9E06B0F9CA2AB24A7F" box="[418,507,1541,1568]" captionStart="FIGURE 6" captionStartId="13.[151,250,1932,1957]" captionTargetBox="[188,1399,167,1913]" captionTargetId="figure@13.[186,1400,166,1913]" captionTargetPageId="13" captionText="FIGURE 6. Helicopsyche (Feropsyche) shaamunensu, new species. 6A6E, Male genitalia (holotype): 6A, left lateral; 6B, dorsal; 6C, ventral; 6D, phallus, left lateral; 6E, phallus, ventral. 6F, Abdominal sterna VI and VII, left lateral. 6G, Abdominal sternum VI ventral process, ventral. Scales = 0.5 mm." httpUri="https://zenodo.org/record/3249591/files/figure.png" pageId="12" pageNumber="243">Fig. 6B</figureCitation>
); in ventral view, with inner margin sinuous, with 3 stout setae basally and row of 45 long, stout setae subapically (
<figureCitation id="E5C42A0CE75DFF9E0502F9E6292E4A1B" box="[528,615,1577,1604]" captionStart="FIGURE 6" captionStartId="13.[151,250,1932,1957]" captionTargetBox="[188,1399,167,1913]" captionTargetId="figure@13.[186,1400,166,1913]" captionTargetPageId="13" captionText="FIGURE 6. Helicopsyche (Feropsyche) shaamunensu, new species. 6A6E, Male genitalia (holotype): 6A, left lateral; 6B, dorsal; 6C, ventral; 6D, phallus, left lateral; 6E, phallus, ventral. 6F, Abdominal sterna VI and VII, left lateral. 6G, Abdominal sternum VI ventral process, ventral. Scales = 0.5 mm." httpUri="https://zenodo.org/record/3249591/files/figure.png" pageId="12" pageNumber="243">Fig. 6C</figureCitation>
); basomesal lobes of inferior appendages well-developed, narrow basally, widening apically, bifid at apex and protruding beyond primary branches in lateral view (
<figureCitation id="E5C42A0CE75DFF9E0396F9822F974A37" box="[1156,1246,1613,1640]" captionStart="FIGURE 6" captionStartId="13.[151,250,1932,1957]" captionTargetBox="[188,1399,167,1913]" captionTargetId="figure@13.[186,1400,166,1913]" captionTargetPageId="13" captionText="FIGURE 6. Helicopsyche (Feropsyche) shaamunensu, new species. 6A6E, Male genitalia (holotype): 6A, left lateral; 6B, dorsal; 6C, ventral; 6D, phallus, left lateral; 6E, phallus, ventral. 6F, Abdominal sterna VI and VII, left lateral. 6G, Abdominal sternum VI ventral process, ventral. Scales = 0.5 mm." httpUri="https://zenodo.org/record/3249591/files/figure.png" pageId="12" pageNumber="243">Fig. 6A</figureCitation>
); in ventral view each of these lobes bearing 34 stout setae on upper apical surface and with small lateral protuberance bearing 23 short, stout setae at inner margin (
<figureCitation id="E5C42A0CE75DFF9E0506F95A29244AEF" box="[532,621,1685,1712]" captionStart="FIGURE 6" captionStartId="13.[151,250,1932,1957]" captionTargetBox="[188,1399,167,1913]" captionTargetId="figure@13.[186,1400,166,1913]" captionTargetPageId="13" captionText="FIGURE 6. Helicopsyche (Feropsyche) shaamunensu, new species. 6A6E, Male genitalia (holotype): 6A, left lateral; 6B, dorsal; 6C, ventral; 6D, phallus, left lateral; 6E, phallus, ventral. 6F, Abdominal sterna VI and VII, left lateral. 6G, Abdominal sternum VI ventral process, ventral. Scales = 0.5 mm." httpUri="https://zenodo.org/record/3249591/files/figure.png" pageId="12" pageNumber="243">Fig. 6C</figureCitation>
). Phallus tubular, strongly down-curved along its length; phallobase narrow, narrowest in middle, inflated basodorsally, with apicoventral margin elongate, strongly sclerotized, rounded apically, semi-membranous apically in lateral view; endotheca large, long, membranous lobe extending dorsally over endophallus; endophallic membranes not inflated; phallotremal sclerite of moderate size, slightly C-shaped (
<figureCitation id="E5C42A0CE75DFF9E078DF8EA2A7B4B1F" box="[159,306,1829,1856]" captionStart="FIGURE 6" captionStartId="13.[151,250,1932,1957]" captionTargetBox="[188,1399,167,1913]" captionTargetId="figure@13.[186,1400,166,1913]" captionTargetPageId="13" captionText="FIGURE 6. Helicopsyche (Feropsyche) shaamunensu, new species. 6A6E, Male genitalia (holotype): 6A, left lateral; 6B, dorsal; 6C, ventral; 6D, phallus, left lateral; 6E, phallus, ventral. 6F, Abdominal sterna VI and VII, left lateral. 6G, Abdominal sternum VI ventral process, ventral. Scales = 0.5 mm." httpUri="https://zenodo.org/record/3249591/files/figure.png" pageId="12" pageNumber="243">Figs. 6D, 6E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="35E56502E75DFF9E07D5F8862ECF4BD8" pageId="12" pageNumber="243" type="materials_examined">
<paragraph id="7D403689E75DFF9E07D5F8862ECF4BD8" blockId="12.[151,1437,640,2035]" pageId="12" pageNumber="243">
<materialsCitation id="CD973CD4E75DFF9E07D5F8862ECF4BD8" ID-GBIF-Occurrence="2268317456" collectingDate="2016-09-13" collectionCode="DZRJ" collectorName="LL Dumas &amp; JL Nessimian &amp; JF Barbosa" country="Brazil" county="Nova Iguacu" latitude="-22.583918" location="Reserva Biologica do Tingua" longLatPrecision="1" longitude="-43.44542" municipality="Tingua" pageId="12" pageNumber="243" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
<emphasis id="4F8BEA9BE75DFF9E07D5F8862A3A4B3C" bold="true" box="[199,371,1865,1891]" pageId="12" pageNumber="243">
<typeStatus id="A244882BE75DFF9E07D5F8862A7D4B3C" box="[199,308,1865,1891]" pageId="12" pageNumber="243" type="holotype">Holotype</typeStatus>
male
</emphasis>
:
<emphasis id="4F8BEA9BE75DFF9E0692F88729E44B3C" bold="true" box="[384,685,1864,1891]" pageId="12" pageNumber="243">
<collectingCountry id="05E87619E75DFF9E0692F8872AB94B3C" box="[384,496,1864,1891]" name="Brazil" pageId="12" pageNumber="243">BRAZIL</collectingCountry>
:
<collectingRegion id="BF3BF86BE75DFF9E06E8F88729E14B3C" box="[506,680,1864,1891]" country="Brazil" name="Rio de Janeiro" pageId="12" pageNumber="243">Rio de Janeiro</collectingRegion>
:
</emphasis>
<collectingCounty id="94214E05E75DFF9E05A0F886280A4B3C" box="[690,835,1865,1891]" pageId="12" pageNumber="243">Nova Iguaçu</collectingCounty>
,
<collectingMunicipality id="9D24ACF3E75DFF9E045EF88628D44B3C" box="[844,925,1865,1891]" pageId="12" pageNumber="243">Tinguá</collectingMunicipality>
,
<location id="78206052E75DFF9E04B5F8862FA34B3C" LSID="urn:lsid:plazi:treatment:F556879FE75DFF9C0785FDCE2E0B4CA4:78206052E75DFF9E04B5F8862FA34B3C" box="[935,1258,1865,1891]" country="Brazil" county="Nova Iguacu" latitude="-22.583918" longLatPrecision="1" longitude="-43.44542" municipality="Tingua" name="Reserva Biologica do Tingua" pageId="12" pageNumber="243" stateProvince="Rio de Janeiro">Reserva Biológica do Tinguá</location>
, afluente do
<location id="78206052E75DFF9E0266F8862BA14BD8" LSID="urn:lsid:plazi:treatment:F556879FE75DFF9C0785FDCE2E0B4CA4:78206052E75DFF9E0266F8862BA14BD8" country="Brazil" county="Nova Iguacu" latitude="-22.583918" longLatPrecision="1" longitude="-43.44542" municipality="Tingua" name="Rio Tingua" pageId="12" pageNumber="243" stateProvince="Rio de Janeiro">Rio Tinguá</location>
,
<geoCoordinate id="18CB504EE75DFF9E07E6F8A32ADC4BD8" box="[244,405,1900,1927]" degrees="22" direction="south" minutes="35" orientation="latitude" pageId="12" pageNumber="243" precision="1" seconds="02.1" value="-22.583918">22°3502.1” S</geoCoordinate>
,
<geoCoordinate id="18CB504EE75DFF9E06B0F8A329044BD8" box="[418,589,1900,1927]" degrees="43" direction="west" minutes="26" orientation="longitude" pageId="12" pageNumber="243" precision="1" seconds="43.5" value="-43.44542">43°2643.5” W</geoCoordinate>
,
<collectingDate id="1905E9A1E75DFF9E054AF8A2299A4BD8" box="[600,723,1901,1927]" pageId="12" pageNumber="243" value="2016-09-13">
<date id="09411049E75DFF9E054AF8A2299A4BD8" box="[600,723,1901,1927]" pageId="12" pageNumber="243" value="2016-09-13">
13.ix.
<quantity id="BA079B6CE75DFF9E0581F8A2299A4BD8" box="[659,723,1901,1927]" metricMagnitude="6" metricUnit="m" metricValue="2.0162329999999997" pageId="12" pageNumber="243" unit="m" value="2016233.0">2016</quantity>
</date>
</collectingDate>
, 233 m,
<collectorName id="D00A535FE75DFF9E0421F8A228E44BD8" box="[819,941,1901,1927]" pageId="12" pageNumber="243">LL Dumas</collectorName>
,
<collectorName id="D00A535FE75DFF9E04AAF8A22F194BD8" box="[952,1104,1901,1927]" pageId="12" pageNumber="243">JL Nessimian</collectorName>
&amp;
<collectorName id="D00A535FE75DFF9E0366F8A22FBB4BD8" box="[1140,1266,1901,1927]" pageId="12" pageNumber="243">JF Barbosa</collectorName>
leg. (
<collectionCode id="1BEEAE4CE75DFF9E0223F8A22E334BD8" box="[1329,1402,1901,1927]" pageId="12" pageNumber="243">DZRJ</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="35E56502E75DFF9E07D5F85F29264BF3" box="[199,623,1936,1964]" pageId="12" pageNumber="243" type="distribution">
<paragraph id="7D403689E75DFF9E07D5F85F29264BF3" blockId="12.[151,1437,640,2035]" box="[199,623,1936,1964]" pageId="12" pageNumber="243">
<emphasis id="4F8BEA9BE75DFF9E07D5F85F2A104BF4" bold="true" box="[199,345,1936,1963]" pageId="12" pageNumber="243">Distribution</emphasis>
.
<collectingCountry id="05E87619E75DFF9E0675F85E2AE24BF4" box="[359,427,1937,1963]" name="Brazil" pageId="12" pageNumber="243">Brazil</collectingCountry>
(
<collectingRegion id="BF3BF86BE75DFF9E06A9F85E292A4BF3" box="[443,611,1937,1964]" country="Brazil" name="Rio de Janeiro" pageId="12" pageNumber="243">Rio de Janeiro</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="35E56502E75DFF9C07D5F87A2E0B4CA4" lastPageId="14" lastPageNumber="245" pageId="12" pageNumber="243" type="etymology">
<paragraph id="7D403689E75DFF9C07D5F87A2E0B4CA4" blockId="12.[151,1437,640,2035]" lastBlockId="14.[151,1437,153,251]" lastPageId="14" lastPageNumber="245" pageId="12" pageNumber="243">
<emphasis id="4F8BEA9BE75DFF9E07D5F87A2A014B90" bold="true" box="[199,328,1973,1999]" pageId="12" pageNumber="243">Etymology</emphasis>
. This species is named for Sha-Amun-En-Su, an Egyptian priestess and singer who lived in Thebes during the first half of the 8th century B.C., responsible for ceremonial duties at the Temple of Karnak, dedicated to the god Amun. The wonderful wooden sarcophagus and its mummy were given as presents to the Brazilian emperor Dom Pedro II during his second travel to
<collectingCountry id="05E87619E75FFF9C05BFFF7229BA4C88" box="[685,755,189,215]" name="Egypt" pageId="14" pageNumber="245">Egypt</collectingCountry>
, in 1876, by the Khedive Ismail Pasha. In 1889 they became part of the great Egyptian artifacts collection of the Museu Nacional, one of the specialties of the museum.
</paragraph>
</subSubSection>
</treatment>
</document>