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<document id="72AB8F32FF41BD53C1CDB3B8B1515A35" ID-DOI="10.5281/zenodo.5391440" ID-ISSN="1638-9387" ID-Zenodo-Dep="5391440" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630315127817" checkinUser="marcus" docAuthor="Monniot, Françoise &amp; Monniot, Claude" docDate="2001" docId="F57D87A3FFF9311CEA51FDD4FE5C13A3" docLanguage="en" docName="Zoosystema.2001.23.2.201-383.pdf" docOrigin="Zoosystema 23 (2)" docStyle="DocumentStyle:64A3174DE33E974D4EB6573A0A793B12.4:Zoosystema.2000-2008.journal_article.type2" docStyleId="64A3174DE33E974D4EB6573A0A793B12" docStyleName="Zoosystema.2000-2008.journal_article.type2" docStyleVersion="4" docTitle="Eudistoma fucatum Monniot &amp; Monniot 2001, n. sp." docType="treatment" docVersion="3" lastPageNumber="240" masterDocId="0944FFDBFFDF313BEB47FF93FFBE1602" masterDocTitle="Ascidians from the tropical western Pacific" masterLastPageNumber="383" masterPageNumber="201" pageNumber="239" updateTime="1699223418854" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="1202D3E8EEAB9E38906E331102CF7796" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A978BC67D98ACF2506B8742C75B83E20">
<mods:title id="5D046F6C6BF6775CC586CD3D0CC7486C">Ascidians from the tropical western Pacific</mods:title>
</mods:titleInfo>
<mods:name id="C9B8E635EDFAECF6D385FA4D3CA323A3" type="personal">
<mods:role id="9725A6614ECE56411BBEF1D2A1E128D9">
<mods:roleTerm id="E85CAE14170FBB4545538DFF4E25D0EE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="840BED5C672448BC7988E2E7D828D65E">Monniot, Françoise</mods:namePart>
</mods:name>
<mods:name id="406F53FDFDEFABFE767749AD656EC298" type="personal">
<mods:role id="5D0C6F1AC3801D9EFE00217817F3D6C2">
<mods:roleTerm id="F96DCE727A70E6BE49D906D3A5259A14">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C569ED1F0A03EB64AECFCAB15EDDBA18">Monniot, Claude</mods:namePart>
<mods:affiliation id="5C9E26014A570A3A029EC24CA240538C">UPESA 8044, Laboratoire de Biologie des Invertébrés marins et Malacologie, Muséum national d’Histoire naturelle, 55 rue Buffon, F- 75005 Paris (France) monniot @ mnhn. fr.</mods:affiliation>
<mods:nameIdentifier id="7C1D505E5B20F099A8E7217F73B32CAC" type="email">monniot@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="668EFD4C54CD57CA6531F1680A48E41C">text</mods:typeOfResource>
<mods:relatedItem id="5835F2EA93097ECD310ADD65305699CE" type="host">
<mods:titleInfo id="E1996E9AEEFF1A1C5E9580BEAA172176">
<mods:title id="5CCCEB9EFECDE13B41321791468F608C">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="7686A7E4D70158066F1175C9DA0513E2">
<mods:date id="F18FE4D5DA2E700D6FC2EF3F24D9DB42">2001</mods:date>
<mods:detail id="6039EE71D9D4247D55306C48948B13AC" type="volume">
<mods:number id="8CD747202B2C41494D8C3E54E021BB1A">23</mods:number>
</mods:detail>
<mods:detail id="AF79F2E959AC1CB5B613685DBC5F52EF" type="issue">
<mods:number id="5C5CC103D03B285C97B6F7464772786A">2</mods:number>
</mods:detail>
<mods:extent id="4BCD9EF62968E55CC017B67756F9B4C7" unit="page">
<mods:start id="B97874B3D638F54BDA610543869DD8F2">201</mods:start>
<mods:end id="1E64C565723BB80404D4D1F8C09612E3">383</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="75FF54DB6B0E248F6135C79BFCD4CD0D">journal article</mods:classification>
<mods:identifier id="141B5DCD988651A507638BAA1EAB5283" type="DOI">10.5281/zenodo.5391440</mods:identifier>
<mods:identifier id="4EAA7962020A708992B881782C3424B3" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="E39434B554F2D34C3C0CE8ED95CBE9D1" type="Zenodo-Dep">5391440</mods:identifier>
</mods:mods>
<treatment id="F57D87A3FFF9311CEA51FDD4FE5C13A3" LSID="urn:lsid:plazi:treatment:F57D87A3FFF9311CEA51FDD4FE5C13A3" httpUri="http://treatment.plazi.org/id/F57D87A3FFF9311CEA51FDD4FE5C13A3" lastPageId="39" lastPageNumber="240" pageId="38" pageNumber="239">
<subSubSection id="35CE653EFFF9311DEA51FDD4FD931462" box="[278,557,582,609]" pageId="38" pageNumber="239" type="nomenclature">
<paragraph id="7D6B36B5FFF9311DEA51FDD4FD931462" blockId="38.[278,557,582,640]" box="[278,557,582,609]" pageId="38" pageNumber="239">
<heading id="262381D9FFF9311DEA51FDD4FD931462" box="[278,557,582,609]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="239" reason="2">
<taxonomicName id="BAD44D36FFF9311DEA51FDD4FE551463" ID-CoL="3C297" authority="Monniot &amp; Monniot, 2001" authorityName="Monniot &amp; Monniot" authorityYear="2001" box="[278,491,583,609]" class="Ascidiacea" family="Polycitoridae" genus="Eudistoma" kingdom="Animalia" order="Aplousobranchia" pageId="38" pageNumber="239" phylum="Chordata" rank="species" species="fucatum" status="sp. nov.">
<emphasis id="4FA0EAA7FFF9311DEA51FDD4FE551463" bold="true" box="[278,491,583,609]" italics="true" pageId="38" pageNumber="239">Eudistoma fucatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="549357DCFFF9311DEAB5FDD5FD931462" box="[498,557,582,608]" pageId="38" pageNumber="239" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="35CE653EFFF9311DEA0AFDF5FE4B1482" box="[333,501,614,640]" pageId="38" pageNumber="239" type="description">
<paragraph id="7D6B36B5FFF9311DEA0AFDF5FE4B1482" blockId="38.[278,557,582,640]" box="[333,501,614,640]" pageId="38" pageNumber="239">
(
<figureCitation id="E5EF2A30FFF9311DEA12FDF5FE191482" box="[341,423,614,640]" captionStart="FIG" captionStartId="39.[151,162,1021,1038]" captionTargetBox="[151,684,166,983]" captionTargetId="figure-403@39.[133,709,139,1014]" captionTargetPageId="39" captionText="FIG. 29. — Eudistoma fucatum n. sp.; A, B two parts of a zooid; C, base of the abdomen. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391497" httpUri="https://zenodo.org/record/5391497/files/figure.png" pageId="38" pageNumber="239">Figs 29</figureCitation>
;
<figureCitation id="E5EF2A30FFF9311DEAF4FDF5FE531482" box="[435,493,614,640]" captionStart="FIG" captionStartId="157.[151,162,1565,1582]" captionTargetBox="[162,1240,227,1527]" captionTargetId="figure-71@157.[162,1261,227,1550]" captionTargetPageId="157" captionText="FIG. 116. — A, Clavelina robusta Kott, 1990; B, Citorclinum laboutei Monniot F. &amp; Millar, 1988; C, Cystodytes hapu Monniot C. &amp; Monniot F., 1987; D, Eudistoma album Monniot F., 1988; E, Eudistoma fluorescens n. sp.; F, Eudistoma fucatum n. sp." figureDoi="http://doi.org/10.5281/zenodo.5391665" httpUri="https://zenodo.org/record/5391665/files/figure.png" pageId="38" pageNumber="239">116F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="35CE653EFFF9311DEBD0FD0DFE5514E8" pageId="38" pageNumber="239" type="materials_examined">
<paragraph id="7D6B36B5FFF9311DEBD0FD0DFE5514E8" blockId="38.[151,684,670,786]" pageId="38" pageNumber="239">
<collectionCode id="1BC5AE70FFF9311DEBD0FD0DFEF814B6" box="[151,326,670,693]" pageId="38" pageNumber="239">
<typeStatus id="A26F8817FFF9311DEBD0FD0DFF7014B6" box="[151,206,670,693]" pageId="38" pageNumber="239">TYPE</typeStatus>
MATERIAL
</collectionCode>
. —
<emphasis id="4FA0EAA7FFF9311DEAC6FD0DFE7414B4" bold="true" box="[385,458,670,694]" pageId="38" pageNumber="239">
<collectingCountry id="05C37625FFF9311DEAC6FD0DFE7A14B4" box="[385,452,670,694]" name="Palau" pageId="38" pageNumber="239">Palau</collectingCountry>
.
</emphasis>
<collectingRegion id="BF10F857FFF9311DEA91FD0DFDA214B7" box="[470,540,670,693]" country="Palau" name="Koror" pageId="38" pageNumber="239">Koror</collectingRegion>
, SW end of Uchelbeluu Reef, wall,
<geoCoordinate id="18E05072FFF9311DEA39FD2AFE5614D3" box="[382,488,697,721]" degrees="7" direction="north" minutes="16.54" orientation="latitude" pageId="38" pageNumber="239" precision="9" value="7.2756667">7°16.54’N</geoCoordinate>
,
<geoCoordinate id="18E05072FFF9311DEAB3FD2AFDCC14D3" box="[500,626,697,721]" degrees="134" direction="east" minutes="31.58" orientation="longitude" pageId="38" pageNumber="239" precision="9" value="134.52634">134°31.58’E</geoCoordinate>
,
<quantity id="BA2C9B50FFF9311DE939FD2BFD1614CD" box="[638,680,696,719]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="38" pageNumber="239" unit="m" value="7.0">7 m</quantity>
, 25.IX.97 (
<collectionCode id="1BC5AE70FFF9311DEBBAFD41FEEF14E8" box="[253,337,722,746]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="38" pageNumber="239">MNHN</collectionCode>
A3
<collectionCode id="1BC5AE70FFF9311DEA3DFD42FE1114EA" box="[378,431,721,744]" pageId="38" pageNumber="239">EUD</collectionCode>
230).
</paragraph>
</subSubSection>
<subSubSection id="35CE653EFFF9311DEBD0FD69FD3C1513" box="[151,642,762,786]" pageId="38" pageNumber="239" type="etymology">
<paragraph id="7D6B36B5FFF9311DEBD0FD69FD3C1513" blockId="38.[151,684,670,786]" box="[151,642,762,786]" pageId="38" pageNumber="239">
ETYMOLOGY. — From the Latin
<emphasis id="4FA0EAA7FFF9311DEA9EFD69FDA41510" box="[473,538,762,786]" italics="true" pageId="38" pageNumber="239">fucatus</emphasis>
: make up.
</paragraph>
</subSubSection>
<subSubSection id="35CE653EFFF9311CEBD0FCD4FE5C13A3" lastPageId="39" lastPageNumber="240" pageId="38" pageNumber="239" type="description">
<paragraph id="7D6B36B5FFF9311DEBD0FCD4FE96155D" blockId="38.[151,685,839,1666]" box="[151,296,839,865]" pageId="38" pageNumber="239">DESCRIPTION</paragraph>
<paragraph id="7D6B36B5FFF9311DEBD0FCFBFDC113E0" blockId="38.[151,685,839,1666]" pageId="38" pageNumber="239">
The colonies are firm oval cushions with a sand-free surface. They are fixed by a part of their base, which is encrusted with some sand; there is no sand in the internal tunic. The colour in life was green (
<figureCitation id="E5EF2A30FFF9311DEBD8FC7BFEAE1200" box="[159,272,1000,1026]" captionStart="FIG" captionStartId="157.[151,162,1565,1582]" captionTargetBox="[162,1240,227,1527]" captionTargetId="figure-71@157.[162,1261,227,1550]" captionTargetPageId="157" captionText="FIG. 116. — A, Clavelina robusta Kott, 1990; B, Citorclinum laboutei Monniot F. &amp; Millar, 1988; C, Cystodytes hapu Monniot C. &amp; Monniot F., 1987; D, Eudistoma album Monniot F., 1988; E, Eudistoma fluorescens n. sp.; F, Eudistoma fucatum n. sp." figureDoi="http://doi.org/10.5281/zenodo.5391665" httpUri="https://zenodo.org/record/5391665/files/figure.png" pageId="38" pageNumber="239">Fig. 116F</figureCitation>
) and turned brown in formalin. The zooids are arranged in circular systems that are very close to each other, with the cloacal apertures tightly grouped in the centre of each system. The tunic is very resistant in the upper layer of the colonies, more gelatinous inside when zooids are strongly retracted. The zooids (
<figureCitation id="E5EF2A30FFF9311DEA5AFB3BFE2212C3" box="[285,412,1191,1218]" captionStart="FIG" captionStartId="39.[151,162,1021,1038]" captionTargetBox="[151,684,166,983]" captionTargetId="figure-403@39.[133,709,139,1014]" captionTargetPageId="39" captionText="FIG. 29. — Eudistoma fucatum n. sp.; A, B two parts of a zooid; C, base of the abdomen. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391497" httpUri="https://zenodo.org/record/5391497/files/figure.png" pageId="38" pageNumber="239">Fig. 29A, B</figureCitation>
) are perpendicular to the colony surface, but the abdomens often cross each other. When relaxed the zooids are
<quantity id="BA2C9B50FFF9311DE93AFB74FF641323" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="38" pageNumber="239" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7 to 8 mm</quantity>
long, but only
<quantity id="BA2C9B50FFF9311DEACCFA9BFE5B1323" box="[395,485,1287,1314]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="38" pageNumber="239" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3-4 mm</quantity>
when contracted. The cloacal siphon is longer than the oral siphon. Both siphons are black in formalin (
<figureCitation id="E5EF2A30FFF9311DE970FADBFD1E1360" box="[567,672,1352,1378]" captionStart="FIG" captionStartId="39.[151,162,1021,1038]" captionTargetBox="[151,684,166,983]" captionTargetId="figure-403@39.[133,709,139,1014]" captionTargetPageId="39" captionText="FIG. 29. — Eudistoma fucatum n. sp.; A, B two parts of a zooid; C, base of the abdomen. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391497" httpUri="https://zenodo.org/record/5391497/files/figure.png" pageId="38" pageNumber="239">Fig. 29A</figureCitation>
), and in some zooids the pigment extends onto the upper part of the thorax (hence the species name:
<emphasis id="4FA0EAA7FFF9311DEBD0FA34FF5C13C3" box="[151,226,1447,1473]" italics="true" pageId="38" pageNumber="239">fucatus</emphasis>
means make-up). The remainder of the body is colourless and lies in a vitreous tunic.
</paragraph>
<caption id="29AB663DFFF9311DE98DFAEDFC6413A6" ID-DOI="http://doi.org/10.5281/zenodo.5391495" ID-Zenodo-Dep="5391495" httpUri="https://zenodo.org/record/5391495/files/figure.png" pageId="38" pageNumber="239" startId="38.[714,725,1406,1423]" targetBox="[714,1245,166,1368]" targetPageId="38">
<paragraph id="7D6B36B5FFF9311DE98DFAEDFC6413A6" blockId="38.[714,1249,1406,1444]" pageId="38" pageNumber="239">
FIG. 28. —
<taxonomicName id="BAD44D36FFF9311DE869FAEDFC49138D" authority="Monniot &amp; Monniot, 2001" authorityName="Monniot &amp; Monniot" authorityYear="2001" box="[814,1015,1406,1423]" class="Ascidiacea" family="Polycitoridae" genus="Eudistoma" kingdom="Animalia" order="Aplousobranchia" pageId="38" pageNumber="239" phylum="Chordata" rank="species" species="fluorescens" status="sp. nov.">
<emphasis id="4FA0EAA7FFF9311DE869FAEDFC49138D" box="[814,1015,1406,1423]" italics="true" pageId="38" pageNumber="239">Eudistoma fluorescens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="549357DCFFF9311DE8B8FAEDFB8B138D" box="[1023,1077,1406,1423]" pageId="38" pageNumber="239" rank="species">n. sp.</taxonomicNameLabel>
;
<emphasis id="4FA0EAA7FFF9311DEF79FAEDFBF5138D" bold="true" box="[1086,1099,1406,1423]" pageId="38" pageNumber="239">A</emphasis>
, zooid;
<emphasis id="4FA0EAA7FFF9311DEFD2FAEDFB1C138D" bold="true" box="[1173,1186,1406,1423]" pageId="38" pageNumber="239">B</emphasis>
, larva. Scale bars: A, 1 mm; B, 0.5 mm.
</paragraph>
</caption>
<paragraph id="7D6B36B5FFF9311DEBD0FA7BFDB01000" blockId="38.[151,685,839,1666]" box="[151,526,1512,1538]" pageId="38" pageNumber="239">The thoracic musculature is dense.</paragraph>
<paragraph id="7D6B36B5FFF9311DEBD0F99BFDDE1083" blockId="38.[151,685,839,1666]" pageId="38" pageNumber="239">The dorsal part of the first row of stigmata is curved. The second row has 25 stigmata on each side. There is an unperforated area anterior to the branchial sac but not in the posterior part.</paragraph>
<paragraph id="7D6B36B5FFF9311CE98DFA7BFF45127D" blockId="38.[714,1247,1511,1666]" lastBlockId="39.[151,684,1127,1442]" lastPageId="39" lastPageNumber="240" pageId="38" pageNumber="239">
The abdomen is at least six times longer than the thorax (
<figureCitation id="E5EF2A30FFF9311DE85BF99BFC241023" box="[796,922,1543,1570]" captionStart="FIG" captionStartId="39.[151,162,1021,1038]" captionTargetBox="[151,684,166,983]" captionTargetId="figure-403@39.[133,709,139,1014]" captionTargetPageId="39" captionText="FIG. 29. — Eudistoma fucatum n. sp.; A, B two parts of a zooid; C, base of the abdomen. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391497" httpUri="https://zenodo.org/record/5391497/files/figure.png" pageId="38" pageNumber="239">Fig. 29A, B</figureCitation>
). The stomach is posterior and short (
<figureCitation id="E5EF2A30FFF9311DE84AF9BBFCD01040" box="[781,878,1576,1602]" captionStart="FIG" captionStartId="39.[151,162,1021,1038]" captionTargetBox="[151,684,166,983]" captionTargetId="figure-403@39.[133,709,139,1014]" captionTargetPageId="39" captionText="FIG. 29. — Eudistoma fucatum n. sp.; A, B two parts of a zooid; C, base of the abdomen. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5391497" httpUri="https://zenodo.org/record/5391497/files/figure.png" pageId="38" pageNumber="239">Fig. 29C</figureCitation>
). The gut loop is straight. There are 30 to 40 round testis follicles in the gut loop. The ovary was not developed and no larvae were found. REMARKS
</paragraph>
<caption id="29AB663DFFF8311CEBD0FC6EFE441221" ID-DOI="http://doi.org/10.5281/zenodo.5391497" ID-Zenodo-Dep="5391497" httpUri="https://zenodo.org/record/5391497/files/figure.png" pageId="39" pageNumber="240" startId="39.[151,162,1021,1038]" targetBox="[151,684,166,983]" targetPageId="39">
<paragraph id="7D6B36B5FFF8311CEBD0FC6EFE441221" blockId="39.[151,685,1021,1059]" pageId="39" pageNumber="240">
FIG. 29. —
<taxonomicName id="BAD44D36FFF8311CEBB4FC6EFE2B120C" authority="Monniot &amp; Monniot, 2001" authorityName="Monniot &amp; Monniot" authorityYear="2001" box="[243,405,1021,1038]" class="Ascidiacea" family="Polycitoridae" genus="Eudistoma" kingdom="Animalia" order="Aplousobranchia" pageId="39" pageNumber="240" phylum="Chordata" rank="species" species="fucatum" status="sp. nov.">
<emphasis id="4FA0EAA7FFF8311CEBB4FC6EFE2B120C" box="[243,405,1021,1038]" italics="true" pageId="39" pageNumber="240">Eudistoma fucatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="549357DCFFF8311CEADDFC6EFE73120C" box="[410,461,1021,1038]" pageId="39" pageNumber="240" rank="species">n. sp.</taxonomicNameLabel>
;
<emphasis id="4FA0EAA7FFF8311CEA93FC6EFE5F120C" bold="true" box="[468,481,1021,1038]" pageId="39" pageNumber="240">A</emphasis>
,
<emphasis id="4FA0EAA7FFF8311CEAABFC6EFE47120C" bold="true" box="[492,505,1021,1038]" pageId="39" pageNumber="240">B</emphasis>
two parts of a zooid;
<emphasis id="4FA0EAA7FFF8311CEBD0FB81FF1B1221" bold="true" box="[151,165,1042,1059]" pageId="39" pageNumber="240">C</emphasis>
, base of the abdomen. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="7D6B36B5FFF8311CEBD0FB1BFE5C13A3" blockId="39.[151,684,1127,1442]" pageId="39" pageNumber="240">This species is mostly characterised by the very constant pigmentation of the tip of the siphons. The green colour of the colony in life and the arrangement of the zooids are also distinctive. The systems are so tightly packed together that the distance between the oral and cloacal openings of a zooid is greater than the distance between the oral openings of adjacent zooids belonging to different systems.</paragraph>
</subSubSection>
</treatment>
</document>
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