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<document id="A07D42EE45F666AFF34F1F06681887A3" ID-CLB-Dataset="45915" ID-DOI="10.11646/zootaxa.3793.3.2" ID-GBIF-Dataset="f7d7c961-c2e2-48c4-a369-0de93c56e1ad" ID-ISSN="1175-5326" ID-Zenodo-Dep="250385" ID-ZooBank="F6183C35-516B-4FEC-A6A3-E8F5DEAF1B2C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460413376537" checkinUser="plazi" docAuthor="Yao, Zhiyuan, Li, Shuqiang &amp; Jäger, Peter" docDate="2014" docId="805F565F7D1AFFF36D90FD275AA61C0D" docLanguage="en" docName="zt03793p349.pdf" docOrigin="Zootaxa 3793 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Belisana Thorell 1898" docType="treatment" docVersion="9" lastPageNumber="332" masterDocId="7C662E277D1BFFF26D07FF935B5C1806" masterDocTitle="Four new species of pholcine spiders (Araneae: Pholcidae) from Southeast Asia" masterLastPageNumber="349" masterPageNumber="331" pageNumber="332" updateTime="1698328215378" updateUser="ExternalLinkService">
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<mods:title id="7BD27015CC003EB25B7C3BDB744D48C6">Four new species of pholcine spiders (Araneae: Pholcidae) from Southeast Asia</mods:title>
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<mods:namePart id="2F948216877A7650D6C478A07BFA51F3">Yao, Zhiyuan</mods:namePart>
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<mods:namePart id="9E65E0F16D9654CC1178B37EED5FB649">Li, Shuqiang</mods:namePart>
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<treatment id="805F565F7D1AFFF36D90FD275AA61C0D" ID-DOI="http://doi.org/10.5281/zenodo.6135883" ID-GBIF-Taxon="119424159" ID-Zenodo-Dep="6135883" LSID="urn:lsid:plazi:treatment:805F565F7D1AFFF36D90FD275AA61C0D" httpUri="http://treatment.plazi.org/id/805F565F7D1AFFF36D90FD275AA61C0D" lastPageNumber="332" pageId="1" pageNumber="332">
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<paragraph id="0849E7497D1AFFF36D90FD275AA81ACB" blockId="1.[151,500,692,717]" box="[151,500,692,717]" pageId="1" pageNumber="332">
<heading id="530150257D1AFFF36D90FD275AA81ACB" bold="true" box="[151,500,692,717]" fontSize="10" level="2" pageId="1" pageNumber="332" reason="2">
<emphasis id="3A823B5B7D1AFFF36D90FD275AA81ACB" bold="true" box="[151,500,692,717]" pageId="1" pageNumber="332">
Genus
<taxonomicName id="CFF69CCA7D1AFFF36DEDFD275AA81ACB" ID-CoL="398Y" ID-ENA="1184437" authority="Thorell, 1898" authorityName="Thorell" authorityYear="1898" box="[234,500,692,717]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="3A823B5B7D1AFFF36DEDFD275A131ACB" bold="true" box="[234,335,692,717]" italics="true" pageId="1" pageNumber="332">Belisana</emphasis>
<bibRefCitation id="6C679AB87D1AFFF36C51FD275AA81ACB" author="Thorell" box="[342,500,692,717]" pageId="1" pageNumber="332" refString="Thorell, T. (1898) Viaggio di Leonardo Fea in Birmania e regioni vicine. LXXX. Secondo saggio sui Ragni birmani. II. Retitelariae et Orbitelariae. Annali del Museo Civico di Storia Naturale di Genova, 39, 271 - 378." type="journal article" year="1898">Thorell, 1898</bibRefCitation>
</taxonomicName>
</emphasis>
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<paragraph id="0849E7497D1AFFF36D90FD6F5FC21B14" blockId="1.[151,1182,764,786]" box="[151,1182,764,786]" pageId="1" pageNumber="332">
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<taxonomicName id="CFF69CCA7D1AFFF36D90FD6F5AF31B14" ID-CoL="398Y" authority="Thorell 1898: 278" authorityName="Thorell" authorityPageNumber="278" authorityYear="1898" box="[151,431,764,786]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="3A823B5B7D1AFFF36D90FD6F5BAC1B14" box="[151,240,764,786]" italics="true" pageId="1" pageNumber="332">Belisana</emphasis>
<treatmentCitation id="8957C1587D1AFFF36DF1FD6F5AF31B14" author="Thorell" box="[246,431,764,786]" page="278" pageId="1" pageNumber="332" year="1898">
<bibRefCitation id="6C679AB87D1AFFF36DF1FD6F5A211B14" author="Thorell" box="[246,381,764,786]" pageId="1" pageNumber="332" refString="Thorell, T. (1898) Viaggio di Leonardo Fea in Birmania e regioni vicine. LXXX. Secondo saggio sui Ragni birmani. II. Retitelariae et Orbitelariae. Annali del Museo Civico di Storia Naturale di Genova, 39, 271 - 378." type="journal article" year="1898">Thorell 1898</bibRefCitation>
: 278
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</taxonomicName>
.
<typeStatus id="D74D59EB7D1AFFF36CBEFD6E5AB71B14" box="[441,491,765,786]" pageId="1" pageNumber="332">Type</typeStatus>
species
<taxonomicName id="CFF69CCA7D1AFFF36F46FD6F58FF1B14" ID-CoL="68GYK" authority="Thorell, 1898" authorityName="Thorell" authorityYear="1898" box="[577,931,764,786]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="332" phylum="Arthropoda" rank="species" species="tauricornis">
<emphasis id="3A823B5B7D1AFFF36F46FD6F584D1B14" box="[577,785,764,786]" italics="true" pageId="1" pageNumber="332">Belisana tauricornis</emphasis>
<bibRefCitation id="6C679AB87D1AFFF36E10FD6F58FF1B14" author="Thorell" box="[791,931,764,786]" pageId="1" pageNumber="332" refString="Thorell, T. (1898) Viaggio di Leonardo Fea in Birmania e regioni vicine. LXXX. Secondo saggio sui Ragni birmani. II. Retitelariae et Orbitelariae. Annali del Museo Civico di Storia Naturale di Genova, 39, 271 - 378." type="journal article" year="1898">Thorell, 1898</bibRefCitation>
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, by original designation.
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<emphasis id="3A823B5B7D1AFFF36D90FCAD5A4D1B51" bold="true" box="[151,273,830,855]" pageId="1" pageNumber="332">Diagnosis.</emphasis>
This genus resembles
<taxonomicName id="CFF69CCA7D1AFFF36F17FCAD59F41B50" box="[528,680,830,854]" class="Arachnida" family="Pholcidae" genus="Spermophora" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="3A823B5B7D1AFFF36F17FCAD59F41B50" box="[528,680,830,854]" italics="true" pageId="1" pageNumber="332">Spermophora</emphasis>
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, but can be distinguished by the dorsal apophysis on the pedipalpal femur (usually present;
<figureCitation id="90CDFBCC7D1AFFF36CA5FCF15AB51B7D" box="[418,489,866,891]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1911,1934]" captionTargetBox="[216,1370,198,1890]" captionTargetId="figure@2.[216,1371,193,1891]" captionTargetPageId="2" captionText="FIGURES 1 4. Belisana protumida spec. nov., holotype male. 1 2 Pedipalp (1 prolateral view, 2 retrolateral view; arrow points at a slender apophysis on pedipalpal trochanter); 3 4 Distal part of procursus (3 prolateral view, 4 retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = membranous flap, pr = procursus." httpUri="https://zenodo.org/record/250386/files/figure.png" pageId="1" pageNumber="332">Figs 2</figureCitation>
,
<figureCitation id="90CDFBCC7D1AFFF36CF2FCF0594F1B7D" box="[501,531,867,891]" captionStart="FIGURES 12 16" captionStartId="4.[151,264,1886,1909]" captionTargetBox="[224,1363,180,1860]" captionTargetId="figure@4.[220,1366,179,1865]" captionTargetPageId="4" captionText="FIGURES 12 16. Belisana protumida spec. nov., holotype male (12 13), paratype male (14), and paratype female (15 16). 12 13 Pedipalp (12 prolateral view, 13 retrolateral view; arrow points at a slender apophysis on pedipalpal trochanter); 14 Chelicerae, frontal view; 15 Epigyne, ventral view; 16 Vulva, dorsal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, f = membranous flap, pa = proximo-lateral apophysis, pp = pore plate, pr = procursus." httpUri="https://zenodo.org/record/250388/files/figure.png" pageId="1" pageNumber="332">13</figureCitation>
), by the absence of posterior pockets on the female opisthosoma, by the absence of sclerotized ventral flaps on the procursus, and by the absence of serrated bulbal apophyses (
<bibRefCitation id="6C679AB87D1AFFF369B8FC155E0B1B99" author="Huber" box="[1215,1367,902,927]" pageId="1" pageNumber="332" refString="Huber, B. A. (2005 a) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica, 155, 1 - 126." type="journal article" year="2005" yearSuffix="a">Huber 2005a</bibRefCitation>
). This genus can be distinguished from
<taxonomicName id="CFF69CCA7D1AFFF36F1FFC3858F91BC5" authority="Wunderlich, 1992" authorityName="Wunderlich" authorityYear="1992" box="[536,933,938,963]" class="Arachnida" family="Pholcidae" genus="Spermophorides" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="3A823B5B7D1AFFF36F1FFC38598C1BC5" box="[536,720,939,963]" italics="true" pageId="1" pageNumber="332">Spermophorides</emphasis>
<bibRefCitation id="6C679AB87D1AFFF36FDEFC3958F91BC5" author="Wunderlich" box="[729,933,938,963]" pageId="1" pageNumber="332" refString="Wunderlich, J. (1992) Die Spinnen-Fauna der Makaronesischen Inseln: Taxonomie, Okologie, Biogeographie und Evolution. Beitrage zur Araneologie, 1, 1 - 619." type="journal article" year="1992">Wunderlich, 1992</bibRefCitation>
</taxonomicName>
by the absence of pockets on the posterior epigynal plate and by the regular attachment site (i.e. rather proximal than distal) of the procursus on the cymbium (
<figureCitation id="90CDFBCC7D1AFFF36D98FC615A5F1C0D" box="[159,259,1010,1035]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1911,1934]" captionTargetBox="[216,1370,198,1890]" captionTargetId="figure@2.[216,1371,193,1891]" captionTargetPageId="2" captionText="FIGURES 1 4. Belisana protumida spec. nov., holotype male. 1 2 Pedipalp (1 prolateral view, 2 retrolateral view; arrow points at a slender apophysis on pedipalpal trochanter); 3 4 Distal part of procursus (3 prolateral view, 4 retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = membranous flap, pr = procursus." httpUri="https://zenodo.org/record/250386/files/figure.png" pageId="1" pageNumber="332">Figs 12</figureCitation>
,
<figureCitation id="90CDFBCC7D1AFFF36C08FC605A041C0D" box="[271,344,1011,1035]" captionStart="FIGURES 12 16" captionStartId="4.[151,264,1886,1909]" captionTargetBox="[224,1363,180,1860]" captionTargetId="figure@4.[220,1366,179,1865]" captionTargetPageId="4" captionText="FIGURES 12 16. Belisana protumida spec. nov., holotype male (12 13), paratype male (14), and paratype female (15 16). 12 13 Pedipalp (12 prolateral view, 13 retrolateral view; arrow points at a slender apophysis on pedipalpal trochanter); 14 Chelicerae, frontal view; 15 Epigyne, ventral view; 16 Vulva, dorsal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, f = membranous flap, pa = proximo-lateral apophysis, pp = pore plate, pr = procursus." httpUri="https://zenodo.org/record/250388/files/figure.png" pageId="1" pageNumber="332">1213</figureCitation>
;
<bibRefCitation id="6C679AB87D1AFFF36C63FC615AB21C0D" author="Huber" box="[356,494,1010,1035]" pageId="1" pageNumber="332" refString="Huber, B. A. (2007) Two new genera of small, six-eyed pholcid spiders from West Africa, and first record of Spermophorides for mainland Africa (Araneae: Pholcidae). Zootaxa, 1635, 23 - 43." type="journal article" year="2007">Huber 2007</bibRefCitation>
).
</paragraph>
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