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<document id="9B80E237A6837E7E07386072B1B0FBA1" ID-DOI="10.5281/zenodo.13155283" ID-ISSN="0068-547X" ID-Zenodo-Dep="13155283" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538343316" checkinUser="felipe" docAuthor="Deuve, Thierry, Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2016" docId="03D6879DFFA6FFBF3C2C60AAFE497663" docLanguage="en" docName="ProcCalifAcadSci.63.12.341-455.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (12)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Trechus (Trechus) qiqiensis Deuve &amp; Kavanaugh &amp; Liang 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="395" masterDocId="FFEFFFE5FF95FF893C576071FFCE7750" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species." masterLastPageNumber="455" masterPageNumber="341" pageNumber="392" updateTime="1722545240047" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="7E9E396E37C4D7AB30A1EECD5E4CEA51" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="76D63983DCD4C8A3440F3B0BE9E6557D">
<mods:title id="170748B133141F86C7EE8D7DD74EA8B1">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
</mods:titleInfo>
<mods:name id="D14B77F2B2D1AE7929448552A3AE2541" type="personal">
<mods:role id="AD7585A94C2F26CFE93D608E46FEF3D5">
<mods:roleTerm id="E54FF644E1F47076436A2A604ECD5CA7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3AC7DFC4F5EE708CED1D910DC6548BE8">Deuve, Thierry</mods:namePart>
<mods:affiliation id="D992CF592FC7336306A1D33300BDA022">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
</mods:name>
<mods:name id="76132B78E17CFB72FB3DA37D38F2E1C1" type="personal">
<mods:role id="03A098162EBFABE6F931F905606E0C0F">
<mods:roleTerm id="CC7A2DFB070162D1DF1F81F264CB813D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EDF6C93FB3F0DC2AF89AD525AA1F1C64">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="3BC64E7A589D6423A7C48EE776621578">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="555E893B5FA0B488262B3C07905314AE" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
</mods:name>
<mods:name id="48C931B680DB1620DB6855C662BA23D1" type="personal">
<mods:role id="7693BAD8B9CA8451641A98878AEBA567">
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<mods:namePart id="452275F6A37567AA8C237C5D656CCB95">Liang, Hongbin</mods:namePart>
<mods:affiliation id="82E57E5EA78F7304BB3566B6AD2891C5">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:typeOfResource id="64964F311DFA1F73C8A97AC91D5D27CF">text</mods:typeOfResource>
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<mods:title id="7514368728906A04FB3CA64B8A1E2518">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="C311DC98B967E5BD4C5E2B74451EAB9A">
<mods:date id="1F0A081C73E13AC0D0A0708A8F5AA916">2016</mods:date>
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<mods:number id="C244631DF39C57DC8460FD39DDAAA180">2016-10-14</mods:number>
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<mods:number id="E565A957FF623E7E7AB6BE4786ED482A">63</mods:number>
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<mods:number id="689D27922C940EB8B565582317E05744">12</mods:number>
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<treatment id="03D6879DFFA6FFBF3C2C60AAFE497663" LSID="urn:lsid:plazi:treatment:03D6879DFFA6FFBF3C2C60AAFE497663" httpUri="http://treatment.plazi.org/id/03D6879DFFA6FFBF3C2C60AAFE497663" lastPageId="54" lastPageNumber="395" pageId="51" pageNumber="392">
<subSubSection id="C3656500FFA6FFBA3C2C60AAFCD077A3" box="[123,798,219,243]" pageId="51" pageNumber="392" type="nomenclature">
<paragraph id="8BC0368BFFA6FFBA3C2C60AAFCD077A3" blockId="51.[123,798,219,273]" box="[123,798,219,243]" pageId="51" pageNumber="392">
<heading id="D08881E7FFA6FFBA3C2C60AAFCD077A3" box="[123,798,219,243]" fontSize="10" level="2" pageId="51" pageNumber="392" reason="4">
<emphasis id="B90BEA99FFA6FFBA3C2C60AAFCD077A3" bold="true" box="[123,798,219,243]" pageId="51" pageNumber="392">
<taxonomicName id="4C7F4D08FFA6FFBA3C2C60AAFE3377A3" ID-CoL="8M8J2" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,509,219,243]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="392" phylum="Arthropoda" rank="species" species="qiqiensis" subGenus="Trechus">
<emphasis id="B90BEA99FFA6FFBA3C2C60AAFF1A77A3" bold="true" box="[123,212,219,243]" italics="true" pageId="51" pageNumber="392">Trechus</emphasis>
(
<emphasis id="B90BEA99FFA6FFBA3CB360AAFEF377A3" bold="true" box="[228,317,219,243]" italics="true" pageId="51" pageNumber="392">Trechus</emphasis>
)
<emphasis id="B90BEA99FFA6FFBA3D1960AAFE6077A3" bold="true" box="[334,430,219,243]" italics="true" pageId="51" pageNumber="392">qiqiensis</emphasis>
Deuve
</taxonomicName>
and Kavanaugh,
<taxonomicNameLabel id="A23857E2FFA6FFBA3E9F60AAFCD077A3" box="[712,798,219,243]" pageId="51" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFA6FFBA3C2C608AFE4F7641" box="[123,385,251,273]" pageId="51" pageNumber="392" type="description">
<paragraph id="8BC0368BFFA6FFBA3C2C608AFE4F7641" blockId="51.[123,798,219,273]" box="[123,385,251,273]" pageId="51" pageNumber="392">
(
<figureCitation id="13442A0EFFA6FFBA3CD5608AFF1D7640" box="[130,211,251,273]" captionStart="FIGURE" captionStartId="52.[171,183,1583,1602]" captionTargetBox="[135,1176,216,1570]" captionTargetId="figure-12@52.[120,1176,216,1570]" captionTargetPageId="52" captionText="FIGURE 20. Trechus qiqiensis sp. nov.; a. Dorsal habitus (CASENT1007384). b. Median lobe of aedeagus of male (CASENT1007384), left lateral aspect. c. Map of locality records (red circles) for T. qiqiensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155322" httpUri="https://zenodo.org/record/13155322/files/figure.png" pageId="51" pageNumber="392">Figs. 20</figureCitation>
,
<figureCitation id="13442A0EFFA6FFBA3C8A608AFECD7641" box="[221,259,251,273]" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="51" pageNumber="392">37b</figureCitation>
,
<figureCitation id="13442A0EFFA6FFBA3D59608AFEFD7640" box="[270,307,251,272]" captionStart="FIGURE" captionStartId="104.[171,183,1626,1645]" captionTargetBox="[134,1163,217,1607]" captionTargetId="figure-12@104.[134,1163,217,1607]" captionTargetPageId="104" captionText="FIGURE 42. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Cikai Township, Qiqi Trail at No. 12 Bridge Camp area, elevation 2775 m; habitat in which specimens of Trechus qiqiensis sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. b. Fugong County, Shiyueliang Township, 11.5 km above Shibali on Shibali Road, elevation 3290 m; habitat in which the holotype of Trechus pseudoqiqiensis sp. nov. was collect- ed. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155370" httpUri="https://zenodo.org/record/13155370/files/figure.png" pageId="51" pageNumber="392">42a</figureCitation>
,
<figureCitation id="13442A0EFFA6FFBA3D6A608AFEB57641" box="[317,379,251,273]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="51" pageNumber="392">4648</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFA6FFBA3CFC6155FF74734D" pageId="51" pageNumber="392" type="materials_examined">
<paragraph id="8BC0368BFFA6FFBA3CFC6155FC2B742C" blockId="51.[123,1174,292,1725]" pageId="51" pageNumber="392">
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</emphasis>
.—
<materialsCitation id="3B173CD6FFA6FFBA3DD56155FE5F76EC" collectingDate="2000-07-15" collectingDateMax="2000-07-19" collectingDateMin="2000-07-15" collectorName="C. E. Griswold" country="China" county="Gaoligong Shan" elevation="2775" latitude="27.71503" location="Nujiang State Nature Reserve" longLatPrecision="1" longitude="98.50244" municipality="Nujiang Prefecture" pageId="51" pageNumber="392" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FFA6FFBA3DD56155FE26766C" box="[386,488,292,316]" pageId="51" pageNumber="392" type="holotype">Holotype</typeStatus>
, a male, in IOZ, labeled: “CASENT 1007384”/ “
<collectingCountry id="F368761BFFA6FFBA3FAB6155FB9B766C" box="[1020,1109,292,316]" name="China" pageId="51" pageNumber="392">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFA6FFBA38366155FEC5760C" country="China" name="Yunnan" pageId="51" pageNumber="392">Yunnan Province</collectingRegion>
,
<collectingCounty id="62A14E07FFA6FFBA3D416135FE09760C" box="[278,455,324,348]" pageId="51" pageNumber="392">Gaoligong Shan</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFA6FFBA3D856135FD6E760C" box="[466,672,324,348]" pageId="51" pageNumber="392">Nujiang Prefecture</collectingMunicipality>
,
<location id="8EA06050FFA6FFBA3EFB6135FC3E760B" LSID="urn:lsid:plazi:treatment:03D6879DFFA6FFBF3C2C60AAFE497663:8EA06050FFA6FFBA3EFB6135FC3E760B" box="[684,1008,324,348]" country="China" county="Gaoligong Shan" latitude="27.71503" longLatPrecision="1" longitude="98.50244" municipality="Nujiang Prefecture" name="Nujiang State Nature Reserve" pageId="51" pageNumber="392" stateProvince="Yunnan">Nujiang State Nature Reserve</location>
, No. 12
<location id="8EA06050FFA6FFBA381A6135FF3D762B" LSID="urn:lsid:plazi:treatment:03D6879DFFA6FFBF3C2C60AAFE497663:8EA06050FFA6FFBA381A6135FF3D762B" country="China" county="Gaoligong Shan" latitude="27.71503" longLatPrecision="1" longitude="98.50244" municipality="Nujiang Prefecture" name="Bridge Camp area" pageId="51" pageNumber="392" stateProvince="Yunnan">Bridge Camp area</location>
, 16.3 airkm
<locationDeviation id="9A722FA9FFA6FFBA3DD66115FDFD762C" box="[385,563,356,380]" location="Gongshan" pageId="51" pageNumber="392">W of Gongshan</locationDeviation>
,”/ “
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/
<geoCoordinate id="EE4B504CFFA6FFBA3EBF6115FCA7762C" box="[744,873,356,380]" degrees="98.50244" direction="east" orientation="longitude" pageId="51" pageNumber="392" precision="1" value="98.50244">E98.50244°</geoCoordinate>
,
<quantity id="4C879B6EFFA6FFBA3F206115FC03762B" box="[887,973,356,380]" metricMagnitude="3" metricUnit="m" metricValue="2.775" pageId="51" pageNumber="392" unit="m" value="2775.0">
<elevation id="0052D1B8FFA6FFBA3F206115FC03762B" box="[887,973,356,380]" metricMagnitude="3" metricUnit="m" metricValue="2.775" pageId="51" pageNumber="392" unit="m" value="2775.0">2775 m</elevation>
</quantity>
,
<date id="FFC1104BFFA6FFBA3F8C6115FB5F762C" box="[987,1169,356,380]" pageId="51" pageNumber="392" value="2000-07-15" valueMax="2000-07-19" valueMin="2000-07-15">
<collectingDate id="EF85E9A3FFA6FFBA3F8C6115FB5F762C" box="[987,1169,356,380]" pageId="51" pageNumber="392" value="2000-07-15" valueMax="2000-07-19" valueMin="2000-07-15">15-19 July 2000</collectingDate>
</date>
, Stop#00-23A, D.H.
<taxonomicName id="4C7F4D08FFA6FFBA3D0561F5FE0076CC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[338,462,388,412]" pageId="51" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFA6FFBA3D8E61F5FDBB76CC" box="[473,629,388,412]" pageId="51" pageNumber="392">C.E. Griswold</collectorName>
, Liang H.-B., D. Ubick, &amp; Dong D.-Z. collectors”/ “IMAGE” [green label]/ “
</materialsCitation>
<typeStatus id="54C48829FFA6FFBA3DC661D5FDEE76EC" box="[401,544,420,444]" pageId="51" pageNumber="392" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FFA6FFBA3E7F61D5FCE176EB" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[552,815,420,444]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="392" phylum="Arthropoda" rank="species" species="qiqiensis">
<emphasis id="B90BEA99FFA6FFBA3E7F61D5FD2C76EC" box="[552,738,420,444]" italics="true" pageId="51" pageNumber="392">Trechus qiqiensis</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FFA6FFBA3F8D61D5FBE376EB" box="[986,1069,420,443]" pageId="51" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
designat- ed 2016” [red label].
<materialsCitation id="3B173CD6FFA6FFBA3D3C61B5FF31756C" collectionCode="CAS, IOZ, MNHN" country="China" pageId="51" pageNumber="392" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54C48829FFA6FFBA3D3C61B5FE1D768C" box="[363,467,452,476]" pageId="51" pageNumber="392" type="paratype">Paratypes</typeStatus>
(at total of 11):
<specimenCount id="9D79FD02FFA6FFBA3ED861B5FD2A768C" box="[655,740,452,476]" count="2" pageId="51" pageNumber="392" type="male">2 males</specimenCount>
and
<specimenCount id="9D79FD02FFA6FFBA3F4861B5FC47768C" box="[799,905,452,476]" count="4" pageId="51" pageNumber="392" type="female">4 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFA6FFBA3FED61B5FC3E768C" box="[954,1008,452,476]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="51" pageNumber="392" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFA6FFBA3FA961B5FBE3768C" box="[1022,1069,452,476]" pageId="51" pageNumber="392">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFFA6FFBA386C61B5FB40768C" box="[1083,1166,452,476]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="51" pageNumber="392">MNHN</collectionCode>
) labeled: same as
<typeStatus id="54C48829FFA6FFBA3D6C6195FE5576AC" box="[315,411,484,508]" pageId="51" pageNumber="392" type="holotype">holotype</typeStatus>
, except first label “CASENT 1007380” to “CASENT 1007381” and “CASENT 1007382” to “CASENT 1007383” and “CASENT 1007385” to “CASENT 1007386”, respectively
</materialsCitation>
;
<materialsCitation id="3B173CD6FFA6FFBA3D5B6255FC4575CC" collectingDate="2002-10-01" collectingDateMax="2002-10-02" collectingDateMin="2002-10-01" collectionCode="CAS, IOZ" collectorName="P. A. Marek" country="China" county="Gongshan County" elevation="3370" latitude="27.77032" location="Cikai Township" longLatPrecision="1" longitude="98.44661" pageId="51" pageNumber="392" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFA6FFBA3D5B6255FE98756C" box="[268,342,548,572]" count="1" pageId="51" pageNumber="392" type="male">1 male</specimenCount>
and
<specimenCount id="9D79FD02FFA6FFBA3DD96255FE39756C" box="[398,503,548,572]" count="2" pageId="51" pageNumber="392" type="female">2 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFA6FFBA3E716255FD92756C" box="[550,604,548,572]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="51" pageNumber="392" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFA6FFBA3E3D6255FD56756C" box="[618,664,548,572]" pageId="51" pageNumber="392">IOZ</collectionCode>
) labeled: “CASENT 1010345” and “CASENT 1010346” to “CASENT 1010347”/ “
<collectingCountry id="F368761BFFA6FFBA3DA96235FD99750C" box="[510,599,580,604]" name="China" pageId="51" pageNumber="392">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFA6FFBA3E376234FD7A750C" box="[608,692,581,604]" country="China" name="Yunnan" pageId="51" pageNumber="392">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFA6FFBA3EE96235FC4E750C" box="[702,896,580,604]" pageId="51" pageNumber="392">Gongshan County</collectingCounty>
,
<location id="8EA06050FFA6FFBA3FDD6235FBFB750C" LSID="urn:lsid:plazi:treatment:03D6879DFFA6FFBF3C2C60AAFE497663:8EA06050FFA6FFBA3FDD6235FBFB750C" box="[906,1077,580,604]" country="China" county="Gongshan County" latitude="27.77032" longLatPrecision="1" longitude="98.44661" name="Cikai Township" pageId="51" pageNumber="392" stateProvince="Yunnan">Cikai Township</location>
,
<locationDeviation id="9A722FA9FFA6FFBA38686235FDFF752C" location="Gongshan on Dulong Valley Road" pageId="51" pageNumber="392">
<quantity id="4C879B6EFFA6FFBA38686235FB5B750C" box="[1087,1173,580,604]" metricMagnitude="4" metricUnit="m" metricValue="5.26" pageId="51" pageNumber="392" unit="km" value="52.6">52.6 km</quantity>
W of Gongshan on Dulong Valley Road
</locationDeviation>
,
<quantity id="4C879B6EFFA6FFBA3E6A6215FD1E752C" box="[573,720,612,636]" metricMagnitude="3" metricUnit="m" metricValue="3.37" metricValueMax="3.38" metricValueMin="3.36" pageId="51" pageNumber="392" unit="m" value="3370.0" valueMax="3380.0" valueMin="3360.0">
<elevation id="0052D1B8FFA6FFBA3E6A6215FD1E752C" box="[573,720,612,636]" metricMagnitude="3" metricUnit="m" metricValue="3.37" metricValueMax="3.38" metricValueMin="3.36" pageId="51" pageNumber="392" unit="m" value="3370.0" valueMax="3380.0" valueMin="3360.0">3360-3380 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFFA6FFBA3EAD6215FCB1752C" box="[762,895,612,636]" degrees="27.77032" direction="north" orientation="latitude" pageId="51" pageNumber="392" precision="1" value="27.77032">N27.77032°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFA6FFBA3FD06215FBC9752C" box="[903,1031,612,636]" degrees="98.44661" direction="east" orientation="longitude" pageId="51" pageNumber="392" precision="1" value="98.44661">E98.44661°</geoCoordinate>
,
<date id="FFC1104BFFA6FFBA38446215FF7D75CC" pageId="51" pageNumber="392" value="2002-10-01" valueMax="2002-10-02" valueMin="2002-10-01">
<collectingDate id="EF85E9A3FFA6FFBA38446215FF7D75CC" pageId="51" pageNumber="392" value="2002-10-01" valueMax="2002-10-02" valueMin="2002-10-01">1-2 October 2002</collectingDate>
</date>
, Stop #DHK-2002-034, D.H.
<taxonomicName id="4C7F4D08FFA6FFBA3DA362F5FDA375CC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[500,621,644,668]" pageId="51" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
&amp;
<collectorName id="268A535DFFA6FFBA3ED862F5FCC775CC" box="[655,777,644,668]" pageId="51" pageNumber="392">P.A. Marek</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFA6FFBA3FD862F5FDA575AC" collectingDate="2004-11-13" collectionCode="CAS" country="China" county="Gongshan County" elevation="3330" latitude="27.78075" location="Cikai Township" longLatPrecision="1" longitude="98.47" pageId="51" pageNumber="392" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFA6FFBA3FD862F5FC1C75CC" box="[911,978,644,668]" count="1" pageId="51" pageNumber="392" type="male">1 male</specimenCount>
(in
<collectionCode id="ED6EAE4EFFA6FFBA3FAA62F5FBFC75CC" box="[1021,1074,644,668]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="51" pageNumber="392" type="Museum">CAS</collectionCode>
) labeled: “CASENT 1015626”/ “
<collectingCountry id="F368761BFFA6FFBA3D2F62D5FE1F75EC" box="[376,465,676,700]" name="China" pageId="51" pageNumber="392">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFA6FFBA3D8A62D4FDFF75EC" box="[477,561,677,700]" country="China" name="Yunnan" pageId="51" pageNumber="392">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFA6FFBA3E6A62D5FCCF75EC" box="[573,769,676,700]" pageId="51" pageNumber="392">Gongshan County</collectingCounty>
,
<location id="8EA06050FFA6FFBA3F5A62D5FC7475EC" LSID="urn:lsid:plazi:treatment:03D6879DFFA6FFBF3C2C60AAFE497663:8EA06050FFA6FFBA3F5A62D5FC7475EC" box="[781,954,676,700]" country="China" county="Gongshan County" latitude="27.78075" longLatPrecision="1" longitude="98.47" name="Cikai Township" pageId="51" pageNumber="392" stateProvince="Yunnan">Cikai Township</location>
,
<locationDeviation id="9A722FA9FFA6FFBA3F9162D5FE73758C" location="Gongshan on Dulong Valley Road" pageId="51" pageNumber="392">
<quantity id="4C879B6EFFA6FFBA3F9162D5FBC475EC" box="[966,1034,676,700]" metricMagnitude="4" metricUnit="m" metricValue="4.8" pageId="51" pageNumber="392" unit="km" value="48.0">48 km</quantity>
W of Gongshan on Dulong Valley Road
</locationDeviation>
,
<quantity id="4C879B6EFFA6FFBA3D9C62B5FDEF758C" box="[459,545,708,732]" metricMagnitude="3" metricUnit="m" metricValue="3.3299999999999996" pageId="51" pageNumber="392" unit="m" value="3330.0">
<elevation id="0052D1B8FFA6FFBA3D9C62B5FDEF758C" box="[459,545,708,732]" metricMagnitude="3" metricUnit="m" metricValue="3.3299999999999996" pageId="51" pageNumber="392" unit="m" value="3330.0">3330 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFFA6FFBA3E1B62B5FD1E758C" box="[588,720,708,732]" degrees="27.78075" direction="north" orientation="latitude" pageId="51" pageNumber="392" precision="1" value="27.78075">N27.78075°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFA6FFBA3E8F62B5FC94758C" box="[728,858,708,732]" degrees="98.47000" direction="east" orientation="longitude" pageId="51" pageNumber="392" precision="1" value="98.47">E98.47000°</geoCoordinate>
,
<date id="FFC1104BFFA6FFBA3F3F62B5FBF1758C" box="[872,1087,708,732]" pageId="51" pageNumber="392" value="2004-11-13">
<collectingDate id="EF85E9A3FFA6FFBA3F3F62B5FBF1758C" box="[872,1087,708,732]" pageId="51" pageNumber="392" value="2004-11-13">13 November 2004</collectingDate>
</date>
, Stop # DHK-2004-086, D.H.
<taxonomicName id="4C7F4D08FFA6FFBA3D2F6295FE3F75AC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[376,497,740,764]" pageId="51" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
collector”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFA6FFBA3E2B6295FC2E742C" collectingDate="2006-08-19" collectionCode="IOZ" collectorName="Y. Liu &amp; All" country="China" county="Gongshan County" elevation="3750" latitude="27.97686" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" pageId="51" pageNumber="392" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFA6FFBA3E2B6295FD0475AC" box="[636,714,740,764]" count="1" pageId="51" pageNumber="392" type="male">1 male</specimenCount>
(in
<collectionCode id="ED6EAE4EFFA6FFBA3F566295FCE175AC" box="[769,815,740,764]" pageId="51" pageNumber="392">IOZ</collectionCode>
) labeled: “CASENT 1025832”/ “
<collectingCountry id="F368761BFFA6FFBA3CD36375FF13744C" box="[132,221,772,796]" name="China" pageId="51" pageNumber="392">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFA6FFBA3CB06374FEF5744C" box="[231,315,773,796]" country="China" name="Yunnan" pageId="51" pageNumber="392">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFA6FFBA3D116375FDC7744C" box="[326,521,772,796]" pageId="51" pageNumber="392">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFA6FFBA3E446375FCD2744C" box="[531,796,772,796]" pageId="51" pageNumber="392">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FFA6FFBA3FE76375FBA7744C" LSID="urn:lsid:plazi:treatment:03D6879DFFA6FFBF3C2C60AAFE497663:8EA06050FFA6FFBA3FE76375FBA7744C" box="[944,1129,772,796]" country="China" county="Gongshan County" latitude="27.97686" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="51" pageNumber="392" stateProvince="Yunnan">Kawakarpu Shan</location>
,
<locationDeviation id="9A722FA9FFA6FFBA38236375FE46746C" location="Chukuai Lake" pageId="51" pageNumber="392">
<quantity id="4C879B6EFFA6FFBA38236375FF53746C" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="51" pageNumber="392" unit="km" value="0.3">0.3 km</quantity>
SW of Chukuai Lake
</locationDeviation>
at campsite,”/ “
<geoCoordinate id="EE4B504CFFA6FFBA3E636355FD76746C" box="[564,696,804,828]" degrees="27.97686" direction="north" orientation="latitude" pageId="51" pageNumber="392" precision="1" value="27.97686">N27.97686°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFA6FFBA3E976355FC8F746C" box="[704,833,804,828]" degrees="98.47799" direction="east" orientation="longitude" pageId="51" pageNumber="392" precision="1" value="98.47799">E98.47799°</geoCoordinate>
,
<quantity id="4C879B6EFFA6FFBA3F1B6355FC6E746C" box="[844,928,804,828]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="51" pageNumber="392" unit="m" value="3750.0">
<elevation id="0052D1B8FFA6FFBA3F1B6355FC6E746C" box="[844,928,804,828]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="51" pageNumber="392" unit="m" value="3750.0">3750 m</elevation>
</quantity>
,
<date id="FFC1104BFFA6FFBA3FFB6355FB97746C" box="[940,1113,804,828]" pageId="51" pageNumber="392" value="2006-08-19">
<collectingDate id="EF85E9A3FFA6FFBA3FFB6355FB97746C" box="[940,1113,804,828]" pageId="51" pageNumber="392" value="2006-08-19">19 August 2006</collectingDate>
</date>
, Stop #DHK-2006-082,
<collectorName id="268A535DFFA6FFBA3D156334FE47740C" box="[322,393,836,860]" pageId="51" pageNumber="392">Y. Liu</collectorName>
collector”.
<collectorName id="268A535DFFA6FFBA3E586335FDFF740C" box="[527,561,836,860]" pageId="51" pageNumber="392">All</collectorName>
<typeStatus id="54C48829FFA6FFBA3E6A6334FD6D740D" box="[573,675,837,861]" pageId="51" pageNumber="392" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="54C48829FFA6FFBA38596335FB5B740C" box="[1038,1173,836,860]" pageId="51" pageNumber="392" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C7F4D08FFA6FFBA3C2C6315FE4C742C" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,386,868,892]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="392" phylum="Arthropoda" rank="species" species="qiqiensis">
<emphasis id="B90BEA99FFA6FFBA3C2C6315FEFA742C" box="[123,308,868,892]" italics="true" pageId="51" pageNumber="392">Trechus qiqiensis</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FFA6FFBA3E7B6314FDB1742C" box="[556,639,869,892]" pageId="51" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFA6FFBA3CFC63F5FE2474EC" blockId="51.[123,1174,292,1725]" pageId="51" pageNumber="392">
<materialsCitation id="3B173CD6FFA6FFBA3CFC63F5FE2874EC" country="China" county="Gongshan County" elevation="2775" latitude="27.71503" location="Camp" longLatPrecision="1" longitude="98.50244" municipality="Qiqi Trail" pageId="51" pageNumber="392" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B90BEA99FFA6FFBA3CFC63F5FEAD74CB" bold="true" box="[171,355,900,924]" pageId="51" pageNumber="392">
<smallCapsWord id="8D26A057FFA6FFBA3CFC63F5FF2874CB" baselines="918,918" box="[171,230,900,924]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="51" pageNumber="392">
<typeStatus id="54C48829FFA6FFBA3CFC63F5FF2874CB" box="[171,230,900,924]" pageId="51" pageNumber="392">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FFA6FFBA3CA163F9FEAD74CB" baselines="918" box="[246,355,904,923]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="51" pageNumber="392">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="F368761BFFA6FFBA3DC163F5FE1474CC" box="[406,474,900,924]" name="China" pageId="51" pageNumber="392">China</collectingCountry>
,
<collectingRegion id="49BBF869FFA6FFBA3DB863F4FD8B74CC" box="[495,581,901,924]" country="China" name="Yunnan" pageId="51" pageNumber="392">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFA6FFBA3E0D63F5FCE574CD" box="[602,811,900,925]" pageId="51" pageNumber="392">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFA6FFBA3F6863F5FC7A74CC" box="[831,948,900,924]" pageId="51" pageNumber="392">Qiqi Trail</collectingMunicipality>
at No. 12
<location id="8EA06050FFA6FFBA381963F4FB5C74CD" LSID="urn:lsid:plazi:treatment:03D6879DFFA6FFBF3C2C60AAFE497663:8EA06050FFA6FFBA381963F4FB5C74CD" box="[1102,1170,901,925]" country="China" county="Gongshan County" latitude="27.71503" longLatPrecision="1" longitude="98.50244" municipality="Qiqi Trail" name="Camp" pageId="51" pageNumber="392" stateProvince="Yunnan">Camp</location>
,
<geoCoordinate id="EE4B504CFFA6FFBA3C2C63D5FF3374EC" box="[123,253,932,956]" degrees="27.71503" direction="north" orientation="latitude" pageId="51" pageNumber="392" precision="1" value="27.71503">N27.71503°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFA6FFBA3D5263D5FE4874EC" box="[261,390,932,956]" degrees="98.50244" direction="east" orientation="longitude" pageId="51" pageNumber="392" precision="1" value="98.50244">E98.50244°</geoCoordinate>
,
<quantity id="4C879B6EFFA6FFBA3DC663D5FE2874EC" box="[401,486,932,956]" metricMagnitude="3" metricUnit="m" metricValue="2.775" pageId="51" pageNumber="392" unit="m" value="2775.0">
<elevation id="0052D1B8FFA6FFBA3DC663D5FE2874EC" box="[401,486,932,956]" metricMagnitude="3" metricUnit="m" metricValue="2.775" pageId="51" pageNumber="392" unit="m" value="2775.0">2775 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFA6FFBA3CFC63B4FF74734D" blockId="51.[123,1174,292,1725]" pageId="51" pageNumber="392">
<emphasis id="B90BEA99FFA6FFBA3CFC63B4FE3F748B" bold="true" box="[171,497,965,989]" pageId="51" pageNumber="392">
<smallCapsWord id="8D26A057FFA6FFBA3CFC63B4FEE0748B" baselines="983,982" box="[171,302,965,989]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="51" pageNumber="392">DERIVATION</smallCapsWord>
<smallCapsWord id="8D26A057FFA6FFBA3D6163B9FE9C748B" baselines="982" box="[310,338,968,987]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="51" pageNumber="392">OF</smallCapsWord>
<smallCapsWord id="8D26A057FFA6FFBA3D0E63B9FE62748B" baselines="982" box="[345,428,968,987]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="51" pageNumber="392">SPECIES</smallCapsWord>
<smallCapsWord id="8D26A057FFA6FFBA3DE363B9FE3F748B" baselines="982" box="[436,497,968,987]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="51" pageNumber="392">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4C7F4D08FFA6FFBA3EAE63B4FC96748C" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[761,856,965,988]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="392" phylum="Arthropoda" rank="species" species="qiqiensis">
<emphasis id="B90BEA99FFA6FFBA3EAE63B4FC96748C" box="[761,856,965,988]" italics="true" pageId="51" pageNumber="392">qiqiensis</emphasis>
</taxonomicName>
, is derived from the name of the ancient trail, Qiqi Trail, passing through the
<typeStatus id="54C48829FFA6FFBA3E2A6394FD6574AD" box="[637,683,997,1021]" pageId="51" pageNumber="392">type</typeStatus>
locality, and the Latin suffix,
<emphasis id="B90BEA99FFA6FFBA3FBB6394FBE474AC" box="[1004,1066,997,1020]" italics="true" pageId="51" pageNumber="392">-ensis</emphasis>
, denoting place.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFA6FFBA3CFC6454FD1C722D" pageId="51" pageNumber="392" type="diagnosis">
<paragraph id="8BC0368BFFA6FFBA3CFC6454FD1C722D" blockId="51.[123,1174,292,1725]" pageId="51" pageNumber="392">
<smallCapsWord id="8D26A057FFA6FFBA3CFC6454FEEE736B" baselines="1079,1078" box="[171,288,1061,1085]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="51" pageNumber="392">
<emphasis id="B90BEA99FFA6FFBA3CFC6454FEEE736B" bold="true" box="[171,288,1061,1085]" pageId="51" pageNumber="392">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFFA6FFBA3E106454FD6D736D" box="[583,675,1061,1085]" captionStart="FIGURE" captionStartId="52.[171,183,1583,1602]" captionTargetBox="[135,1176,216,1570]" captionTargetId="figure-12@52.[120,1176,216,1570]" captionTargetPageId="52" captionText="FIGURE 20. Trechus qiqiensis sp. nov.; a. Dorsal habitus (CASENT1007384). b. Median lobe of aedeagus of male (CASENT1007384), left lateral aspect. c. Map of locality records (red circles) for T. qiqiensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155322" httpUri="https://zenodo.org/record/13155322/files/figure.png" pageId="51" pageNumber="392">Fig. 20a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size relatively large (BL = 4.0 to
<quantity id="4C879B6EFFA6FFBA3CE96414FEDD732D" box="[190,275,1125,1149]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="51" pageNumber="392" unit="mm" value="4.2">4.2 mm</quantity>
), apterous; body color reddish-brown; eyes slightly convex; tempora glabrous, short and convex; pronotum transverse (ratio PW/PL = 1.44), basal angles rectangular and sharp; elytra with striae 1 to 4 deeply impressed on disc but attenuated apically, remaining striae successively less distinct, the outermost effaced or nearly so; recurrent stria terminated anteriorly by a slight convexity in line with stria 7; two discal setae present, inserted on interval 3 next to stria 3; preapical seta present and inserted in a discal position on interval 3, next to either stria 2 or
<quantity id="4C879B6EFFA6FFBA38466574FBF4724D" box="[1041,1082,1285,1309]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="51" pageNumber="392" unit="in" value="3.0">3 in</quantity>
forward position near apical one-fourth of elytra; median lobe of male aedeagus large and robust, subapically bent ventrally, apical lamella sinuate and reflexed dorsally (
<figureCitation id="13442A0EFFA6FFBA3F186534FC61720D" box="[847,943,1349,1373]" captionStart="FIGURE" captionStartId="52.[171,183,1583,1602]" captionTargetBox="[135,1176,216,1570]" captionTargetId="figure-12@52.[120,1176,216,1570]" captionTargetPageId="52" captionText="FIGURE 20. Trechus qiqiensis sp. nov.; a. Dorsal habitus (CASENT1007384). b. Median lobe of aedeagus of male (CASENT1007384), left lateral aspect. c. Map of locality records (red circles) for T. qiqiensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155322" httpUri="https://zenodo.org/record/13155322/files/figure.png" pageId="51" pageNumber="392">Fig. 20b</figureCitation>
), endophallus with a more heavily sclerotized voluminous lobe or scaly fold.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFA6FFBF3CFC65F4FE497663" lastPageId="54" lastPageNumber="395" pageId="51" pageNumber="392" type="description">
<paragraph id="8BC0368BFFA6FFBA3CFC65F4FE8E728D" blockId="51.[123,1174,292,1725]" pageId="51" pageNumber="392">
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<emphasis id="B90BEA99FFA6FFBA3CFC65F4FEF472CC" bold="true" box="[171,314,1413,1437]" pageId="51" pageNumber="392">DESCRIPTION</emphasis>
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.— Size relatively large, BL = 4.0 to
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. Body color reddish-brown, shiny, appendages concolorous but paler, palpi pale yellow. Body surface smooth, only head capsule faintly alutaceous.
</paragraph>
<paragraph id="8BC0368BFFA6FFBC3CFC6594FDD77624" blockId="51.[123,1174,292,1725]" lastBlockId="53.[123,1174,219,1726]" lastPageId="53" lastPageNumber="394" pageId="51" pageNumber="392">
Head. Moderate in size; eyes only slightly projected, slightly more convex, their diameter less than twice as long as tempora, the latter short, convex and glabrous. Frons slightly flattened; frontal furrows deep, curved, continuous but attenuated posterior to the eyes. Two pairs of supraorbital setae present, the anterior pair inserted in foveae. Clypeus with four setae. Labrum with six setae, anterior margin distinctly concave. Mandibles short; right mandible distinctly tridentate with middle tooth closer to basal tooth (premolar) than to distal tooth, left mandible with a small slightly bifid process.
<collectingRegion id="49BBF869FFA6FFBA3D4566D7FEA271ED" box="[274,364,1702,1725]" country="Myanmar" name="Chin" pageId="51" pageNumber="392">Mentum</collectingRegion>
and submentum not fused but suture between them only faintly impressed.
<collectingRegion id="49BBF869FFA0FFBC3C2C60AAFF1B77A2" box="[123,213,219,242]" country="Myanmar" name="Chin" pageId="53" pageNumber="394">Mentum</collectingRegion>
with medial tooth broad and apically truncate, about one-half the length of the lateral lobes. Submentum with six setae anteriorly. Gula wide. Genae with a single seta ventrally on each side. Antennae short, extended posteriorly to basal one-sixth of elytra, with about 2 or 3 antennomeres extended posteriorly beyond basal pronotal margin; antennomeres 2 and 3 about equal in length, antennomere 4 slightly shorter.
</paragraph>
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20.
<taxonomicName id="4C7F4D08FFA1FFBD3D46665EFE667111" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[273,424,1583,1602]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="393" phylum="Arthropoda" rank="species" species="qiqiensis" status="sp. nov.">
<emphasis id="B90BEA99FFA1FFBD3D46665EFE667111" box="[273,424,1583,1602]" italics="true" pageId="52" pageNumber="393">Trechus qiqiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFA1FFBD3DF8665EFE397112" box="[431,503,1583,1602]" pageId="52" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1007384). b. Median lobe of aedeagus of male (CASENT1007384), left lateral aspect. c. Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FFA1FFBD3F656638FC5C710B" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[818,914,1609,1627]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="393" phylum="Arthropoda" rank="species" species="qiqiensis">
<emphasis id="B90BEA99FFA1FFBD3F656638FC5C710B" box="[818,914,1609,1627]" italics="true" pageId="52" pageNumber="393">T. qiqiensis</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFFA0FFBC3CFC610CFDAA75C6" blockId="53.[123,1174,219,1726]" pageId="53" pageNumber="394">Pronotum. Transverse (ratio PW/PL = 1.44), narrowed posteriorly, greatest width near anteri- or one-fourth; lateral margins markedly rounded, with a short sinuation just anterior to basal angles, the latter small, rectangular and sharp. Disc convex, smooth and glabrous, median longitudinal impression finely impressed, but continuous between anterior and posterior margins; basal foveae distinct, circular; median basal area faintly rugulose, delimited laterally by short, oblique furrows. Basal margin nearly rectilinear, slightly projected medially. Lateral borders of pronotum moderately slender, regular, narrowly and regularly reflexed, lateral grooves narrow but distinctly impressed. Single midlateral setae on each side inserted slightly anterior to middle; single basolateral seta on each side, inserted at basal angle.</paragraph>
<paragraph id="8BC0368BFFA0FFBC3CFC62EFFDA674A8" blockId="53.[123,1174,219,1726]" pageId="53" pageNumber="394">
Elytra. Ovoid, humeri distinct but rounded. Disc convex and smooth. Striae finely punctate, intervals faintly convex; striae 1 to 4 deeply impressed but attenuated apically, stria 5 more faintly impressed, striae 6 to 8 effaced or nearly so. Parascutellar striole present. Recurrent stria terminated anteriorly by a slight convexity at the presumed location of stria 7 apically. Basal setiferous pore present at common origin of striae 1 and 2. Two discal setae present and inserted next to stria 3, one at anterior one-fourth and one near middle of elytra. Preapical seta present and inserted in a discal position on interval 3, next to either stria 2 or
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forward position at apical one-fourth of elytra in most specimens; in a few specimens two preapical setae present, one inserted farther forward next to stria 3, as a third discal seta, the second inserted less far forward and next to stria 2. Umbilicate setal series with setae of humeral group equidistance from each other and setae of medi- an group inserted slightly posterior to middle.
</paragraph>
<paragraph id="8BC0368BFFA0FFBC3CFC6470FD6A7368" blockId="53.[123,1174,219,1726]" pageId="53" pageNumber="394">Legs. Medium proportions, slightly short, protibiae with longitudinal furrow. Male protarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFFA0FFBC3CFC6430FB857329" blockId="53.[123,1174,219,1726]" pageId="53" pageNumber="394">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of males apically with one pair of paramedial setae, of females with two pairs.</paragraph>
<paragraph id="8BC0368BFFA0FFBC3CFC64F0FBE6738A" blockId="53.[123,1174,219,1726]" pageId="53" pageNumber="394">
Male aedeagus. Median lobe (
<figureCitation id="13442A0EFFA0FFBC3E5364F0FDA873C9" box="[516,614,1153,1177]" captionStart="FIGURE" captionStartId="52.[171,183,1583,1602]" captionTargetBox="[135,1176,216,1570]" captionTargetId="figure-12@52.[120,1176,216,1570]" captionTargetPageId="52" captionText="FIGURE 20. Trechus qiqiensis sp. nov.; a. Dorsal habitus (CASENT1007384). b. Median lobe of aedeagus of male (CASENT1007384), left lateral aspect. c. Map of locality records (red circles) for T. qiqiensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155322" httpUri="https://zenodo.org/record/13155322/files/figure.png" pageId="53" pageNumber="394">Fig. 20b</figureCitation>
) large and robust, with a moderate-sized sagittal aileron; shaft subapically bent ventrally; apical lamella sinuate and reflexed dorsally, bluntly point- ed apically; endophallus with a more heavily sclerotized voluminous lobe or scaly fold.
</paragraph>
<paragraph id="8BC0368BFFA0FFBC3CFC6493FD73716C" blockId="53.[123,1174,219,1726]" pageId="53" pageNumber="394">
<emphasis id="B90BEA99FFA0FFBC3CFC6493FE6973A9" bold="true" box="[171,423,1250,1274]" pageId="53" pageNumber="394">
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</emphasis>
.— Members of this species have been found at elevations ranging from
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in a variety of habitats. At the lowest elevation (
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), specimens were collected by sifting meager leaf litter and mosses on the forest floor and on rotting logs (
<figureCitation id="13442A0EFFA0FFBC387A6552FB44726B" box="[1069,1162,1315,1339]" captionStart="FIGURE" captionStartId="104.[171,183,1626,1645]" captionTargetBox="[134,1163,217,1607]" captionTargetId="figure-12@104.[134,1163,217,1607]" captionTargetPageId="104" captionText="FIGURE 42. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Cikai Township, Qiqi Trail at No. 12 Bridge Camp area, elevation 2775 m; habitat in which specimens of Trechus qiqiensis sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. b. Fugong County, Shiyueliang Township, 11.5 km above Shibali on Shibali Road, elevation 3290 m; habitat in which the holotype of Trechus pseudoqiqiensis sp. nov. was collect- ed. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155370" httpUri="https://zenodo.org/record/13155370/files/figure.png" pageId="53" pageNumber="394">Fig. 42a</figureCitation>
). This area had abundant conifers (
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<emphasis id="B90BEA99FFA0FFBC3DAF6532FDFD720B" box="[504,563,1347,1371]" italics="true" pageId="53" pageNumber="394">Abies</emphasis>
</taxonomicName>
,
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<emphasis id="B90BEA99FFA0FFBC3E116532FD4A720B" box="[582,644,1347,1371]" italics="true" pageId="53" pageNumber="394">Thuja</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C7F4D08FFA0FFBC3E9E6535FCAF720B" box="[713,865,1347,1371]" pageId="53" pageNumber="394">
<emphasis id="B90BEA99FFA0FFBC3E9E6535FCC8720B" box="[713,774,1348,1371]" italics="true" pageId="53" pageNumber="394">Picea</emphasis>
species
</taxonomicName>
) that formed a partly open canopy. Litter and mosses were moist and substrate underneath was composed of crumbling granitic sand. Specimens of
<taxonomicName id="4C7F4D08FFA0FFBC3DF065F2FD2572CB" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[423,747,1411,1435]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="394" phylum="Arthropoda" rank="species" species="monochaeta" status="sp. nov.">
<emphasis id="B90BEA99FFA0FFBC3DF065F2FD2572CB" box="[423,747,1411,1435]" italics="true" pageId="53" pageNumber="394">Trechepaphiopsis monochaeta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFA0FFBC3EA565F5FC8A72CB" box="[754,836,1412,1435]" pageId="53" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
were also collected in the same litter samples at this site. At higher elevations (above
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), specimens were found under stones in meadows, talus slopes, in low
<taxonomicName id="4C7F4D08FFA0FFBC3D8B65B5FDB4728C" box="[476,634,1476,1500]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="53" pageNumber="394" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FFA0FFBC3D8B65B5FDB4728C" box="[476,634,1476,1500]" italics="true" pageId="53" pageNumber="394">Rhododendron</emphasis>
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thickets, and on heath-covered tundra slopes (
<figureCitation id="13442A0EFFA0FFBC383D65B5FF6972AC" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="53" pageNumber="394">Fig. 37b</figureCitation>
). In such areas,
<taxonomicName id="4C7F4D08FFA0FFBC3D096594FE1472AC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[350,474,1509,1532]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="394" phylum="Arthropoda" rank="species" species="qiqiensis">
<emphasis id="B90BEA99FFA0FFBC3D096594FE1472AC" box="[350,474,1509,1532]" italics="true" pageId="53" pageNumber="394">T. qiqiensis</emphasis>
</taxonomicName>
specimens were found syntopic with those of
<taxonomicName id="4C7F4D08FFA0FFBC3FB56595FE4D714C" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="394" phylum="Arthropoda" rank="species" species="balli" subGenus="Gaoligongtrechus">
<emphasis id="B90BEA99FFA0FFBC3FB56595FB5872AC" box="[994,1174,1508,1532]" italics="true" pageId="53" pageNumber="394">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FFA0FFBC3CD36675FE8A714C" box="[132,324,1540,1564]" italics="true" pageId="53" pageNumber="394">Gaoligongtrechus</emphasis>
)
<emphasis id="B90BEA99FFA0FFBC3D056675FE4D714C" box="[338,387,1540,1564]" italics="true" pageId="53" pageNumber="394">balli</emphasis>
</taxonomicName>
and
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<emphasis id="B90BEA99FFA0FFBC3DE16675FDA4714C" box="[438,618,1540,1564]" italics="true" pageId="53" pageNumber="394">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FFA0FFBC3E2E6674FD7F714C" box="[633,689,1541,1564]" italics="true" pageId="53" pageNumber="394">s. str.</emphasis>
)
<emphasis id="B90BEA99FFA0FFBC3E976675FCE9714C" box="[704,807,1540,1564]" italics="true" pageId="53" pageNumber="394">griswoldi</emphasis>
</taxonomicName>
<emphasis id="B90BEA99FFA0FFBC3F706675FCE5714C" box="[807,811,1540,1564]" italics="true" pageId="53" pageNumber="394">,</emphasis>
and at highest elevation (
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) also with members of
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<emphasis id="B90BEA99FFA0FFBC3D3D6655FDAC716C" box="[362,610,1572,1596]" italics="true" pageId="53" pageNumber="394">Trechus gongshanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFA0FFBC3E3D6654FD73716C" box="[618,701,1573,1596]" pageId="53" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BC0368BFFA0FFBC3CFC6635FBDD71ED" blockId="53.[123,1174,219,1726]" pageId="53" pageNumber="394">
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<smallCapsWord id="8D26A057FFA0FFBC3D086639FE3D710B" baselines="1622" box="[351,499,1608,1627]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="53" pageNumber="394">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="8D26A057FFA0FFBC3DAA6639FD82710B" baselines="1622" box="[509,588,1608,1627]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="within" pageId="53" pageNumber="394">WITHIN</smallCapsWord>
<smallCapsWord id="8D26A057FFA0FFBC3E016639FD4E710B" baselines="1622" box="[598,640,1608,1627]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="the" pageId="53" pageNumber="394">THE</smallCapsWord>
</emphasis>
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<emphasis id="B90BEA99FFA0FFBC3EDD6635FCC1710B" bold="true" box="[650,783,1604,1628]" pageId="53" pageNumber="394">GAOLIGONG</emphasis>
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<materialsCitation id="3B173CD6FFA0FFBC3F4E6635FBEB71CD" county="Cikai Townships in Gongshan County" location="Bingzhiongluo" municipality="We" pageId="53" pageNumber="394" specimenCount="24" specimenCount-female="7" specimenCount-male="5" stateProvince="Shan">
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<emphasis id="B90BEA99FFA0FFBC3F4E6635FC9A710B" bold="true" box="[793,852,1604,1628]" pageId="53" pageNumber="394">
<collectingRegion id="49BBF869FFA0FFBC3F4E6635FC9A710B" box="[793,852,1604,1628]" country="Myanmar" name="Shan" pageId="53" pageNumber="394">SHAN</collectingRegion>
</emphasis>
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.—
<figureCitation id="13442A0EFFA0FFBC3FD76634FC10710D" box="[896,990,1605,1629]" captionStart="FIGURE" captionStartId="52.[171,183,1583,1602]" captionTargetBox="[135,1176,216,1570]" captionTargetId="figure-12@52.[120,1176,216,1570]" captionTargetPageId="52" captionText="FIGURE 20. Trechus qiqiensis sp. nov.; a. Dorsal habitus (CASENT1007384). b. Median lobe of aedeagus of male (CASENT1007384), left lateral aspect. c. Map of locality records (red circles) for T. qiqiensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155322" httpUri="https://zenodo.org/record/13155322/files/figure.png" pageId="53" pageNumber="394">Fig. 20c</figureCitation>
.
<collectingMunicipality id="6BA4ACF1FFA0FFBC3FBC6634FBC0710C" box="[1003,1038,1605,1628]" pageId="53" pageNumber="394">We</collectingMunicipality>
examined a total of
<specimenCount id="9D79FD02FFA0FFBC3C846614FEA8712D" box="[211,358,1637,1661]" count="12" pageId="53" pageNumber="394" type="generic">12 specimens</specimenCount>
(
<specimenCount id="9D79FD02FFA0FFBC3D2F6614FE03712D" box="[376,461,1637,1661]" count="5" pageId="53" pageNumber="394" type="male">5 males</specimenCount>
and
<specimenCount id="9D79FD02FFA0FFBC3E516614FDBA712D" box="[518,628,1637,1661]" count="7" pageId="53" pageNumber="394" type="female">7 females</specimenCount>
), all from
<location id="8EA06050FFA0FFBC3EBD6614FC47712D" LSID="urn:lsid:plazi:treatment:03D6879DFFA6FFBF3C2C60AAFE497663:8EA06050FFA0FFBC3EBD6614FC47712D" box="[746,905,1637,1661]" county="Cikai Townships in Gongshan County" municipality="We" name="Bingzhiongluo" pageId="53" pageNumber="394" stateProvince="Shan">Bingzhiongluo</location>
and
<collectingCounty id="62A14E07FFA0FFBC3F956614FEF071CD" pageId="53" pageNumber="394">Cikai Townships in Gongshan County</collectingCounty>
on the crest and eastern slope of the northern part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FFA0FFBC387A66F4FBC171ED" location="Core Area" pageId="53" pageNumber="394" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FFA0FFBC387A66F4FBAC71CD" box="[1069,1122,1669,1693]" country="Myanmar" name="Shan" pageId="53" pageNumber="394">Shan</collectingRegion>
(see
<typeStatus id="54C48829FFA0FFBC3C2C66D7FF7E71EE" box="[123,176,1702,1726]" pageId="53" pageNumber="394">Type</typeStatus>
material above for exact collection data). These localities are all in
<location id="8EA06050FFA0FFBC3FDC66D4FC3671EE" LSID="urn:lsid:plazi:treatment:03D6879DFFA6FFBF3C2C60AAFE497663:8EA06050FFA0FFBC3FDC66D4FC3671EE" box="[907,1016,1701,1726]" name="Core Area" pageId="53" pageNumber="394" stateProvince="Shan">Core Area</location>
2
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFA3FFBF3CFC60AAFE497663" blockId="54.[123,1173,219,307]" pageId="54" pageNumber="395">
<emphasis id="B90BEA99FFA3FFBF3CFC60AAFD9477A1" bold="true" box="[171,602,219,243]" pageId="54" pageNumber="395">
<smallCapsWord id="8D26A057FFA3FFBF3CFC60AAFEDC77A1" baselines="237,236" box="[171,274,219,243]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Overall" pageId="54" pageNumber="395">OVERALL</smallCapsWord>
<smallCapsWord id="8D26A057FFA3FFBF3D4E60AFFE0E77A1" baselines="236" box="[281,448,222,241]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="geographical" pageId="54" pageNumber="395">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FFA3FFBF3D9160AFFD9477A1" baselines="236" box="[454,602,222,241]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="54" pageNumber="395">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This species currently is known only from Gongshan County on the crest and eastern slope of the northern part of the Gaoligong Shan, in western
<collectingRegion id="49BBF869FFA3FFBF3C2C616DFEFB7663" box="[123,309,283,307]" country="China" name="Yunnan" pageId="54" pageNumber="395">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFA3FFBF3D17616AFE4D7663" box="[320,387,283,307]" name="China" pageId="54" pageNumber="395">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>