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<document ID-DOI="10.5252/geodiversitas2019v41a15" ID-GBIF-Dataset="1e89d723-7f54-4d6b-85dd-6bcd573eca32" ID-ISSN="1638-9395" ID-Zenodo-Dep="3694209" ID-ZooBank="urn:lsid:zoobank.org:pub:9DD3CC29-3AEA-44B8-8E8F-6AD882DF5B1C" checkinTime="1583172268754" checkinUser="valdenar" docAuthor="Bonis, Louis de, Gardin, Axelle &amp; Blondel, Cécile" docDate="2019" docId="03A487991A63FF9AFC19F9396D66FC5F" docLanguage="en" docName="Geodiversitas.41.15.601-621.pdf.imf" docOrigin="Geodiversitas 41 (15)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Amphicynodon Filhol 1881" docType="treatment" docVersion="8" lastPageNumber="603" masterDocId="FF9DFFE11A60FF9EFFADFF8D6E62FF8E" masterDocTitle="Carnivora from the early Oligocene of the Phosphorites du Quercy in southwestern France" masterLastPageNumber="621" masterPageNumber="601" pageNumber="602" updateTime="1675702587573" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Carnivora from the early Oligocene of the Phosphorites du Quercy in southwestern France</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bonis, Louis de</mods:namePart>
<mods:affiliation>Palevoprim: laboratoire de Paléontologie, Évolution, Paléoécosystèmes, Paléoprimatologie, Bâtiment B 35 TSA 51106, 6 rue Michel Brunet F- 86073 Poitiers cedex 9 (France)</mods:affiliation>
<mods:nameIdentifier type="email">louis.debonis@univ-poitiers.fr</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gardin, Axelle</mods:namePart>
<mods:affiliation>Palevoprim: laboratoire de Paléontologie, Évolution, Paléoécosystèmes, Paléoprimatologie, Bâtiment B 35 TSA 51106, 6 rue Michel Brunet F- 86073 Poitiers cedex 9 (France)</mods:affiliation>
<mods:nameIdentifier type="email">axelle.gardin@univ-poitiers.fr</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Blondel, Cécile</mods:namePart>
<mods:affiliation>Palevoprim: laboratoire de Paléontologie, Évolution, Paléoécosystèmes, Paléoprimatologie, Bâtiment B 35 TSA 51106, 6 rue Michel Brunet F- 86073 Poitiers cedex 9 (France)</mods:affiliation>
<mods:nameIdentifier type="email">cecile.blondel@univ-poitiers.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Geodiversitas</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-09-10</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>41</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>15</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>601</mods:start>
<mods:end>621</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5252/geodiversitas2019v41a15</mods:identifier>
<mods:identifier type="GBIF-Dataset">1e89d723-7f54-4d6b-85dd-6bcd573eca32</mods:identifier>
<mods:identifier type="ISSN">1638-9395</mods:identifier>
<mods:identifier type="Zenodo-Dep">3694209</mods:identifier>
<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:9DD3CC29-3AEA-44B8-8E8F-6AD882DF5B1C</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3703495" ID-GBIF-Taxon="162367543" ID-Zenodo-Dep="3703495" LSID="urn:lsid:plazi:treatment:03A487991A63FF9AFC19F9396D66FC5F" httpUri="http://treatment.plazi.org/id/03A487991A63FF9AFC19F9396D66FC5F" lastPageId="4" lastPageNumber="603" pageId="3" pageNumber="602">
<subSubSection box="[948,1320,1716,1742]" pageId="3" pageNumber="602" type="nomenclature">
<paragraph blockId="3.[948,1320,1716,1742]" box="[948,1320,1716,1742]" pageId="3" pageNumber="602">
<heading box="[948,1320,1716,1742]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="602" reason="2">
Genus
<taxonomicName authority="Filhol, 1881" authorityName="Filhol" authorityYear="1881" box="[1025,1320,1716,1742]" class="Mammalia" family="Ursidae" genus="Amphicynodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="3" pageNumber="602" phylum="Chordata" rank="genus">
<emphasis box="[1025,1180,1716,1742]" italics="true" pageId="3" pageNumber="602">Amphicynodon</emphasis>
Filhol, 1881
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="602" type="type_taxon">
<paragraph blockId="3.[813,1456,1785,1836]" pageId="3" pageNumber="602">
<typeStatus box="[813,863,1785,1809]" pageId="3" pageNumber="602">
<collectionCode box="[813,863,1785,1809]" pageId="3" pageNumber="602">TYPE</collectionCode>
</typeStatus>
<collectionCode box="[871,944,1790,1807]" pageId="3" pageNumber="602">SPECIES</collectionCode>
. —
<emphasis box="[994,1172,1785,1809]" italics="true" pageId="3" pageNumber="602">Cynodon velaunus</emphasis>
Aymard, 1846 (by original designation).
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="603" pageId="3" pageNumber="602" type="diagnosis">
<paragraph blockId="3.[813,1456,1872,2029]" lastBlockId="4.[132,775,217,426]" lastPageId="4" lastPageNumber="603" pageId="3" pageNumber="602">
DIAGNOSIS. —
<taxonomicName authorityName="Simpson" authorityYear="1945" box="[973,1182,1872,1896]" class="Mammalia" family="Amphicynodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="3" pageNumber="602" phylum="Chordata" rank="family">Amphicynodontidae</taxonomicName>
with a relatively low skull with a faint sagittal crest slightly more developed distally; elongate muzzle; ossified bullae; shallow suprameatal fossa; alisphenoid canal present. Dental formula 3/3 I, 1/1 C, 4/4 P, 2-3/
<specimenCount box="[1332,1380,1952,1976]" pageId="3" pageNumber="602" type="generic">3 M.</specimenCount>
Simple premolars P1-P3 and p1-p3; short P4 with robust protocone and short metastyle; triangular M1 with tiny parastyle without paraconule but with metaconule; M2 more or less reduced; p4 with pacd or not; grouped and not very tall trigonid of m1, horizontal wear of the cuspids, metaconid as high as the paraconid; not very elongate m2 with paraconid present or not, metaconid as high or higher than the protoconid; small rounded m3.
</paragraph>
<paragraph blockId="4.[132,775,217,426]" pageId="4" pageNumber="603">
The genus is present in the old Quercy collections represented by several species but also in the new early Oligocene localities (
<tableCitation box="[141,217,402,426]" captionStart="TABLE" captionStartId="5.[408,419,223,241]" captionTargetPageId="5" captionText="TABLE 1. — Measurements (in mm) of teeth of Amphicynodon Filhol, 1881, from the alveoli." httpUri="http://table.plazi.org/id/DF7266071A65FF9BFE35FF526AFFFF7C" pageId="4" pageNumber="603" tableUuid="DF7266071A65FF9BFE35FF526AFFFF7C">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="603" type="discussion">
<paragraph blockId="4.[132,775,472,977]" box="[132,226,472,498]" pageId="4" pageNumber="603">REMARK</paragraph>
<paragraph blockId="4.[132,775,472,977]" pageId="4" pageNumber="603">
<bibRefCitation box="[132,294,504,531]" pageId="4" pageNumber="603">Aymard (1846)</bibRefCitation>
created the species
<emphasis box="[492,610,504,531]" italics="true" pageId="4" pageNumber="603">C. velaunus</emphasis>
for a fossil from the early Oligocene locality Ronzon (
<collectingCountry box="[514,587,536,562]" name="France" pageId="4" pageNumber="603">France</collectingCountry>
) that he described a few years later (
<bibRefCitation author="Aymard" box="[320,469,568,595]" firstAuthor="Aymard" pageId="4" pageNumber="603" pagination="92 - 122" refId="ref16402" refString="AYMARD A. 1850. - Du Cynodon, mammifere carnassier fossile trouve dans les calcaires marneux de Ronzon, pres du Puy. Annales de la Societe d'Agriculture, Sciences, Arts et Commerce du Puy 15: 92 - 122. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 2293835 / f 95. item" type="journal article" year="1850">Aymard 1850</bibRefCitation>
). He added another species,
<emphasis box="[132,247,600,626]" italics="true" pageId="4" pageNumber="603">C. palustris</emphasis>
Aymard, 1853 for a hemi-mandible slightly larger than that of
<emphasis box="[260,379,632,658]" italics="true" pageId="4" pageNumber="603">C. velaunus</emphasis>
. Later (1881) Filhol, considering that
<emphasis box="[132,250,664,690]" italics="true" pageId="4" pageNumber="603">C. palustris</emphasis>
was very different from
<emphasis box="[519,641,664,690]" italics="true" pageId="4" pageNumber="603">C. velaunus</emphasis>
, created the new genus
<taxonomicName authorityName="Filhol" authorityYear="1881" box="[255,408,695,721]" class="Mammalia" family="Ursidae" genus="Amphicynodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="604" phylum="Chordata" rank="genus">
<emphasis box="[255,408,695,721]" italics="true" pageId="4" pageNumber="603">Amphicynodon</emphasis>
</taxonomicName>
for it. Finally, the name
<emphasis box="[684,775,695,721]" italics="true" pageId="4" pageNumber="603">Cynodon</emphasis>
being pre-occupied by a fish,
<bibRefCitation author="Trouessart" box="[466,664,728,754]" firstAuthor="Trouessart" pageId="4" pageNumber="603" refId="ref19259" refString="TROUESSART E. 1904. - Catalogus mammalium tam viventium quam fossium quinquinale supplementum. Vol. 1. R. Friedlander &amp; Sohn, Berlin: 1 - 546, 224. https: // doi. org / 10.5962 / bhl. title. 61437" type="journal volume" year="1904">Trouessart (1904)</bibRefCitation>
proposed the genus name
<emphasis box="[322,429,759,785]" italics="true" pageId="4" pageNumber="603">Aymardia</emphasis>
. However, following the rules of nomenclature,
<taxonomicName authority="Filhol, 1881" authorityName="Filhol" authorityYear="1881" box="[331,634,791,817]" class="Mammalia" family="Ursidae" genus="Amphicynodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="603" phylum="Chordata" rank="genus">
<emphasis box="[331,488,791,817]" italics="true" pageId="4" pageNumber="603">Amphicynodon</emphasis>
Filhol, 1881
</taxonomicName>
has priority, although
<bibRefCitation author="Teilhard" box="[232,400,823,850]" firstAuthor="Teilhard" pageId="4" pageNumber="603" pagination="103 - 190" refId="ref19194" refString="TEILHARD DE CHARDIN P. 1915. - Les carnassiers des phosphorites du Quercy. Annales de Paleontologie 9: 103 - 190." type="journal article" year="1915">Teilhard (1915)</bibRefCitation>
, without taking into account either Filhol or Trouessart, continued using
<emphasis box="[533,624,855,881]" italics="true" pageId="4" pageNumber="603">Cynodon</emphasis>
. Later, several species of
<taxonomicName authorityName="Filhol" authorityYear="1881" box="[237,389,887,913]" class="Mammalia" family="Ursidae" genus="Amphicynodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="604" phylum="Chordata" rank="genus">
<emphasis box="[237,389,887,913]" italics="true" pageId="4" pageNumber="603">Amphicynodon</emphasis>
</taxonomicName>
were described, principally from the old collections of the Quercy, thus lacking biostratigraphic support. We shall here focus on the specimens from MP 22.
</paragraph>
</subSubSection>
</treatment>
</document>