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<document id="3D71C168D4924C79C28D5882FCBB3C73" ID-DOI="10.5281/zenodo.13799439" ID-ISSN="0068-547X" ID-Zenodo-Dep="13799439" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E2657307FFF98FF8EFA4EBF6C0FAE" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Caelostomus latus Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="209" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="207" updateTime="1726771873529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="EDE99968B05675747610843E424745EF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9B09A72D61E587C6AEA88F1833B77B67">
<mods:title id="E426337213571C5CB8AA0A47D23779B5">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="263056243198961CAE173E19BA0C6DDB" type="personal">
<mods:role id="300B5743B16CC462381518D85E93537A">
<mods:roleTerm id="AA8BC65DB199747561FA5969C527B391">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D0AFEB6212E3479D1901C27D0706C03B">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="5BCF09DA57A9839E6F1E1D76EFC7CE4B">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
<mods:name id="76E34DBBBE3EBBAB5A99D67083129A12" type="personal">
<mods:role id="23081F5E1258202D775604F6118F78A3">
<mods:roleTerm id="B6F70A602336FBE541F7C768CA509366">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0A125961254B653E6048F10D632D8FD6">Rainio, Johanna</mods:namePart>
<mods:affiliation id="6B2C06F0879272E3CD8D2D5487C291A2">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
<mods:typeOfResource id="BA49AA25EDDE52EE2CB5E4D33C1B59A5">text</mods:typeOfResource>
<mods:relatedItem id="DBDD93A012CA7C36158462F04537606B" type="host">
<mods:titleInfo id="436965FA4D14D1A0ED2EF547F6E92F28">
<mods:title id="825B12E9558AF771BE0A004A7BC6E662">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="9FA74C94B3E04B048D3F4E9CC924D5B3">
<mods:date id="7B7C28DCEA01E56DE09D3324F65C4461">2016</mods:date>
<mods:detail id="A9FAEA990EC1D1DE820E6FED9BCD0EB6" type="pubDate">
<mods:number id="FEE2A3160A0328DC113E3636335C8846">2016-04-29</mods:number>
</mods:detail>
<mods:detail id="7B98A4410DAC119A9F70FE59316427A8" type="volume">
<mods:number id="6FF28A5B3804C31264A26862B259DC38">63</mods:number>
</mods:detail>
<mods:detail id="65ABFA2785557E9A986FD081268B92F0" type="issue">
<mods:number id="DD7F0ED457E726209C54C4B85ADA5B76">7</mods:number>
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<mods:start id="052E3FF0D49B7529DDC5BC75693DF5A8">201</mods:start>
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</mods:extent>
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<mods:classification id="2F85C0EBAF90CBFF995EB3D08EA609B8">journal article</mods:classification>
<mods:identifier id="6ECAF9AECA5E77BAFB67263E32C570B7" type="DOI">10.5281/zenodo.13799439</mods:identifier>
<mods:identifier id="79B9EAFA9704E7058674A2BA3105A126" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="F805B1F02D20531E8948037FF91F5547" type="Zenodo-Dep">13799439</mods:identifier>
</mods:mods>
<treatment id="0A3E2657307FFF98FF8EFA4EBF6C0FAE" LSID="urn:lsid:plazi:treatment:0A3E2657307FFF98FF8EFA4EBF6C0FAE" httpUri="http://treatment.plazi.org/id/0A3E2657307FFF98FF8EFA4EBF6C0FAE" lastPageId="8" lastPageNumber="209" pageId="6" pageNumber="207">
<subSubSection id="CA8DC4CA307FFF96FF8EFA4EBFDA08EE" box="[123,697,1500,1525]" pageId="6" pageNumber="207" type="nomenclature">
<paragraph id="82289741307FFF96FF8EFA4EBFDA08EE" blockId="6.[123,697,1500,1555]" box="[123,697,1500,1525]" pageId="6" pageNumber="207">
<heading id="D960202D307FFF96FF8EFA4EBFDA08EE" box="[123,697,1500,1525]" fontSize="10" level="2" pageId="6" pageNumber="207" reason="4">
<emphasis id="B0E34B53307FFF96FF8EFA4EBFDA08EE" bold="true" box="[123,697,1500,1525]" pageId="6" pageNumber="207">
<taxonomicName id="4597ECC2307FFF96FF8EFA4EBF3608EE" ID-CoL="5X55H" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,597,1500,1525]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
<emphasis id="B0E34B53307FFF96FF8EFA4EBC2708EF" bold="true" box="[123,324,1500,1524]" italics="true" pageId="6" pageNumber="207">Caelostomus latus</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F628307FFF96FD96FA4FBFDA08EE" box="[611,697,1501,1525]" pageId="6" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA307FFF96FF8EFA6FBDAF0B09" box="[123,204,1533,1555]" pageId="6" pageNumber="207" type="description">
<paragraph id="82289741307FFF96FF8EFA6FBDAF0B09" blockId="6.[123,697,1500,1555]" box="[123,204,1533,1555]" pageId="6" pageNumber="207">
<figureCitation id="1AAC8BC4307FFF96FF8EFA6FBDAF0B09" box="[123,204,1533,1555]" captionStart="FIGURE" captionStartId="7.[155,167,1251,1270]" captionTargetBox="[120,1176,216,1233]" captionTargetId="figure-985@7.[120,1176,216,1235]" captionTargetPageId="7" captionText="FIGURE 4. Digital images of holotype female of Caelostomus latus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Head, left lateral aspect (arrow points to tranverse depression between eyes); D. Right anterior tarsus, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799449" httpUri="https://zenodo.org/record/13799449/files/figure.png" pageId="6" pageNumber="207">Figure 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA307FFF98FF5EF9A3BF6C0FAE" lastPageId="8" lastPageNumber="209" pageId="6" pageNumber="207" type="materials_examined">
<paragraph id="82289741307FFF96FF5EF9A3B9F50B52" blockId="6.[123,1174,1585,1705]" box="[171,1174,1585,1609]" pageId="6" pageNumber="207">
<emphasis id="B0E34B53307FFF96FF5EF9A3BC3E0B5C" bold="true" box="[171,349,1585,1609]" pageId="6" pageNumber="207">
<smallCapsWord id="84CE019D307FFF96FF5EF9A3BD870B5C" baselines="1603,1602" box="[171,228,1585,1609]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="6" pageNumber="207">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D307FFF96FF05F9A6BC3E0B5C" baselines="1602" box="[240,349,1588,1607]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="6" pageNumber="207">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C307FFF96FE71F9A3B9F50B52" box="[388,1174,1585,1609]" country="Madagascar" location="MADAGASCAR" pageId="6" pageNumber="207" specimenCount="1" typeStatus="holotype">
<typeStatus id="5D2C29E3307FFF96FE71F9A3BC840B52" box="[388,487,1585,1609]" pageId="6" pageNumber="207" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC4307FFF96FE0EF9A3BF180B53" box="[507,635,1585,1609]" captionStart="FIGURE" captionStartId="7.[155,167,1251,1270]" captionTargetBox="[120,1176,216,1233]" captionTargetId="figure-985@7.[120,1176,216,1235]" captionTargetPageId="7" captionText="FIGURE 4. Digital images of holotype female of Caelostomus latus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Head, left lateral aspect (arrow points to tranverse depression between eyes); D. Right anterior tarsus, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799449" httpUri="https://zenodo.org/record/13799449/files/figure.png" pageId="6" pageNumber="207">Figs. 4A–B</figureCitation>
), a female, in CAS, labeled: “
<collectingCountry id="FA80D7D1307FFF96FC2DF9A3B9F50B52" box="[984,1174,1585,1609]" name="Madagascar" pageId="6" pageNumber="207">MADAGASCAR</collectingCountry>
</materialsCitation>
</paragraph>
<paragraph id="82289741307FFF96FF8EF9C3B9F60B72" blockId="6.[123,1174,1585,1705]" box="[123,1173,1617,1641]" pageId="6" pageNumber="207">
Ranomafana NP. Talatakely, Trail C,
<specimenCount id="94915CC8307FFF96FDC4F9C3BFE70B73" box="[561,644,1617,1641]" count="350" pageId="6" pageNumber="207" type="generic">
<quantity id="456F3AA4307FFF96FDC4F9C3BF1D0B73" box="[561,638,1617,1641]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="207" unit="m" value="350.0">350 m</quantity>
.
</specimenCount>
<date id="F629B181307FFF96FD66F9C3BF9D0B72" box="[659,766,1617,1641]" pageId="6" pageNumber="207" value="2005-05-10">10.5.2005</date>
”/ “
<typeStatus id="5D2C29E3307FFF96FCD9F9C3BEDA0B72" box="[812,953,1617,1641]" pageId="6" pageNumber="207" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC2307FFF96FC3DF9C3B9F60B72" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[968,1173,1617,1641]" pageId="6" pageNumber="207" status="sp. n.">
<emphasis id="B0E34B53307FFF96FC3DF9C3B9F60B72" box="[968,1173,1617,1641]" italics="true" pageId="6" pageNumber="207">Caelostomus latus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="82289741307FFF96FF8EF9E3B9500BB3" blockId="6.[123,1174,1585,1705]" pageId="6" pageNumber="207">
Kavanaugh &amp; Rainio
<taxonomicNameLabel id="ABD0F628307FFF96FE93F9E3BCC30B93" box="[358,416,1649,1672]" pageId="6" pageNumber="207" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
<materialsCitation id="32FF9D1C307FFF96FF5EF903B94F0BB2" box="[171,1068,1681,1705]" country="Madagascar" location="Ranomafana National Park" pageId="6" pageNumber="207" specimenCount="1" stateProvince="Fianarantsoa">
<emphasis id="B0E34B53307FFF96FF5EF903BC340BBC" bold="true" box="[171,343,1681,1705]" pageId="6" pageNumber="207">
<smallCapsWord id="84CE019D307FFF96FF5EF903BD870BBC" baselines="1699,1698" box="[171,228,1681,1705]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="6" pageNumber="207">
<typeStatus id="5D2C29E3307FFF96FF5EF903BD870BBC" box="[171,228,1681,1705]" pageId="6" pageNumber="207">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D307FFF96FF19F906BC340BBC" baselines="1698" box="[236,343,1684,1703]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="6" pageNumber="207">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D1307FFF96FE8BF903BC9C0BB2" box="[382,511,1681,1705]" name="Madagascar" pageId="6" pageNumber="207">Madagascar</collectingCountry>
,
<collectingRegion id="405359A3307FFF96FDF8F903BF9F0BB2" box="[525,764,1681,1705]" country="Madagascar" name="Fianarantsoa" pageId="6" pageNumber="207">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A307FFF96FCFFF903B94F0BB2" LSID="urn:lsid:plazi:treatment:0A3E2657307FFF98FF8EFA4EBF6C0FAE:8748C19A307FFF96FCFFF903B94F0BB2" box="[778,1068,1681,1705]" country="Madagascar" name="Ranomafana National Park" pageId="6" pageNumber="207" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<caption id="D6E8C7C9307EFF97FF6EFB71BF2C0831" ID-DOI="http://doi.org/10.5281/zenodo.13799449" ID-Zenodo-Dep="13799449" httpUri="https://zenodo.org/record/13799449/files/figure.png" pageId="7" pageNumber="208" startId="7.[155,167,1251,1270]" targetBox="[120,1176,216,1233]" targetPageId="7" targetType="figure">
<paragraph id="82289741307EFF97FF6EFB71BF2C0831" blockId="7.[123,1173,1251,1322]" pageId="7" pageNumber="208">
<smallCapsWord id="84CE019D307EFF97FF6EFB71BDBB09EE" baselines="1265,1265" box="[155,216,1251,1270]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="7" pageNumber="208">FIGURE</smallCapsWord>
4. Digital images of holotype female of
<taxonomicName id="4597ECC2307EFF97FDB1FB71BEC709ED" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[580,932,1251,1270]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
<emphasis id="B0E34B53307EFF97FDB1FB71BF8009ED" box="[580,739,1251,1270]" italics="true" pageId="7" pageNumber="208">Caelostomus latus</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628307EFF97FC59FB71BE8C09ED" box="[940,1007,1251,1270]" pageId="7" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels; C. Head, left lateral aspect (arrow points to tranverse depression between eyes); D. Right anterior tarsus, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm.
</paragraph>
</caption>
<paragraph id="82289741307EFF97FF5EFAD7BCA20886" blockId="7.[123,1174,1349,1726]" pageId="7" pageNumber="208">
<emphasis id="B0E34B53307EFF97FF5EFAD7BC940847" bold="true" box="[171,503,1349,1373]" pageId="7" pageNumber="208">
<smallCapsWord id="84CE019D307EFF97FF5EFAD7BC4D0840" baselines="1367,1366" box="[171,302,1349,1373]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="7" pageNumber="208">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D307EFF97FEC2FADABC300840" baselines="1366" box="[311,339,1352,1371]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="7" pageNumber="208">OF</smallCapsWord>
<smallCapsWord id="84CE019D307EFF97FEA9FADABCCC0840" baselines="1366" box="[348,431,1352,1371]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="7" pageNumber="208">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D307EFF97FE4FFADBBC940847" baselines="1367" box="[442,503,1353,1372]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="7" pageNumber="208">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC2307EFF97FCF7FAD7BE550846" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[770,822,1349,1373]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B0E34B53307EFF97FCF7FAD7BE550846" box="[770,822,1349,1373]" italics="true" pageId="7" pageNumber="208">latus</emphasis>
</taxonomicName>
, is the Latin adjective meaning broad or wide, a reference to relatively broad pronotum of the
<typeStatus id="5D2C29E3307EFF97FCE7FAF7BE0C0866" box="[786,879,1381,1405]" pageId="7" pageNumber="208" type="holotype">holotype</typeStatus>
specimen and, presumably, other members of this species.
</paragraph>
<paragraph id="82289741307EFF98FF5EFA37BF3B0F4F" blockId="7.[123,1174,1349,1726]" lastBlockId="8.[123,1174,219,693]" lastPageId="8" lastPageNumber="209" pageId="7" pageNumber="208">
<smallCapsWord id="84CE019D307EFF97FF5EFA37BC2108A0" baselines="1463,1462" box="[171,322,1445,1469]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="7" pageNumber="208">
<emphasis id="B0E34B53307EFF97FF5EFA37BC2108A0" bold="true" box="[171,322,1445,1469]" pageId="7" pageNumber="208">RECOGNITION</emphasis>
</smallCapsWord>
.— Size moderate for genus, SBL =
<quantity id="456F3AA4307EFF97FD31FA37BE7708A6" box="[708,788,1445,1469]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="7" pageNumber="208" unit="mm" value="6.3">6.3 mm</quantity>
. Based on Jeannel’s (1948) key, this species is a member of subgenus
<taxonomicName id="4597ECC2307EFF97FE1EFA57BF1608C6" authorityName="MacLeay" authorityYear="1825" box="[491,629,1477,1501]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B0E34B53307EFF97FE1EFA57BF1608C6" box="[491,629,1477,1501]" italics="true" pageId="7" pageNumber="208">Caelostomus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABD0F628307EFF97FD8BFA54BFDB08C6" box="[638,696,1478,1501]" pageId="7" pageNumber="208" sensu="stricto">s. str.</taxonomicNameLabel>
It’s members (
<figureCitation id="1AAC8BC4307EFF97FCAAFA57BED008C6" box="[863,947,1477,1501]" captionStart="FIGURE" captionStartId="7.[155,167,1251,1270]" captionTargetBox="[120,1176,216,1233]" captionTargetId="figure-985@7.[120,1176,216,1235]" captionTargetPageId="7" captionText="FIGURE 4. Digital images of holotype female of Caelostomus latus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Head, left lateral aspect (arrow points to tranverse depression between eyes); D. Right anterior tarsus, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799449" httpUri="https://zenodo.org/record/13799449/files/figure.png" pageId="7" pageNumber="208">Fig. 4A</figureCitation>
) share with those of other members of the subgenus the following features: head with frons and frontal furrows smooth, impunctate; elytral base distinctly margined from humerus medially to base of stria 3 or 2; elytral striae 6 and 7 deeply impressed throughout, not effaced apically; lateral elytral intervals narrow, moderately convex; apical part of umbilicate series of setiferous pores not inserted in a deep groove; and female apical abdominal ventrite with (two or) three pairs of setae subapically. Within the subgenus, the following features are shared only with members of two species of Jeannel’s
<taxonomicName id="4597ECC2307EFF97FF8EF937BC7F0BA6" authorityName="Tschitscherine" authorityYear="1899" box="[123,284,1701,1725]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="convexiusculus">
<emphasis id="B0E34B53307EFF97FF8EF937BC7F0BA6" box="[123,284,1701,1725]" italics="true" pageId="7" pageNumber="208">convexiusculus</emphasis>
</taxonomicName>
group, namely
<taxonomicName id="4597ECC2307EFF97FE39F937BE970BA6" authority="Tschitscherine (1899)" authorityName="Tschitscherine" authorityYear="1899" box="[460,1012,1701,1725]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="convexiusculus">
<emphasis id="B0E34B53307EFF97FE39F937BE630BA6" box="[460,768,1701,1725]" italics="true" pageId="7" pageNumber="208">Caelostomus convexiusculus</emphasis>
<bibRefCitation id="E606EAB0307EFF97FCFFF937BE970BA6" author="TSCHITSCHERINE, T. S." box="[778,1012,1701,1725]" pageId="7" pageNumber="208" pagination="50 - 82" refId="ref32397" refString="TSCHITSCHERINE, T. S. 1899. Memoire sur qulques Platysmatini nouveaux ou peu connus d'Afrique et de Madagascar. Annales de la Societe Entomologique de France 68: 50 - 82." type="journal article" year="1899">Tschitschérine (1899)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4597ECC2307EFF98FBDBF937BCD70DE8" authority="Jeannel (1948)" authorityName="Jeannel" authorityYear="1948" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" lastPageId="8" lastPageNumber="209" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B0E34B53307EFF98FBDBF937BC6A0DE8" italics="true" lastPageId="8" lastPageNumber="209" pageId="7" pageNumber="208">Caelostomus alluaudi</emphasis>
Jeannel (1948)
</taxonomicName>
: SBL greater than 6.0 mm, elytra short and broad, ratio EL/EW = 1.19, elytral disc convex; and elytral intervals distinctly punctate throughout, including in apical part. The
<typeStatus id="5D2C29E33071FF98FF15FE89BC5E0C28" box="[224,317,283,307]" pageId="8" pageNumber="209" type="holotype">holotype</typeStatus>
female of
<taxonomicName id="4597ECC23071FF98FE5BFE8EBF620C28" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[430,513,283,307]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B0E34B533071FF98FE5BFE8EBF620C28" box="[430,513,283,307]" italics="true" pageId="8" pageNumber="209">C. latus</emphasis>
</taxonomicName>
is slightly larger than members of these other two species, has a distinct and broad transverse depression between the eyes seen best in lateral view (
<figureCitation id="1AAC8BC43071FF98FBCCFEA9B9EE0C4F" box="[1081,1165,315,340]" captionStart="FIGURE" captionStartId="7.[155,167,1251,1270]" captionTargetBox="[120,1176,216,1233]" captionTargetId="figure-985@7.[120,1176,216,1235]" captionTargetPageId="7" captionText="FIGURE 4. Digital images of holotype female of Caelostomus latus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Head, left lateral aspect (arrow points to tranverse depression between eyes); D. Right anterior tarsus, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799449" httpUri="https://zenodo.org/record/13799449/files/figure.png" pageId="8" pageNumber="209">Fig. 4C</figureCitation>
) (depression absent from members of the other two species). The pronotum in the
<typeStatus id="5D2C29E33071FF98FBE2FECEB9170C6F" box="[1047,1140,348,372]" pageId="8" pageNumber="209" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4597ECC23071FF98FF8EFEEEBDAE0C8F" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,205,380,404]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B0E34B533071FF98FF8EFEEEBDAE0C8F" box="[123,205,380,404]" italics="true" pageId="8" pageNumber="209">C. latus</emphasis>
</taxonomicName>
is much wider than in
<taxonomicName id="4597ECC23071FF98FE4BFEEEBF1E0C8F" authorityName="Tschitscherine" authorityYear="1899" box="[446,637,380,404]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="convexiusculus">
<emphasis id="B0E34B533071FF98FE4BFEEEBF1E0C8F" box="[446,637,380,404]" italics="true" pageId="8" pageNumber="209">C. convexiusculus</emphasis>
</taxonomicName>
members, slightly wider than in
<taxonomicName id="4597ECC23071FF98FC2FFEEEB9320C8F" authorityName="Jeannel" authorityYear="1948" box="[986,1105,380,404]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B0E34B533071FF98FC2FFEEEB9320C8F" box="[986,1105,380,404]" italics="true" pageId="8" pageNumber="209">C. alluaudi</emphasis>
</taxonomicName>
members, and less depressed anterolaterally than in members of either of the other species; and the pronotal basal foveae are straighter and more sharply defined laterally than in members of the other two species. The elytra are less abruptly rounded laterally and apically than in
<taxonomicName id="4597ECC23071FF98FC20FE4FB92C0CEF" authorityName="Jeannel" authorityYear="1948" box="[981,1103,476,500]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B0E34B533071FF98FC20FE4FB92C0CEF" box="[981,1103,476,500]" italics="true" pageId="8" pageNumber="209">C. alluaudi</emphasis>
</taxonomicName>
members but similar to shape in to those in
<taxonomicName id="4597ECC23071FF98FDDBFE6FBF920F0F" authorityName="Tschitscherine" authorityYear="1899" box="[558,753,508,532]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="convexiusculus">
<emphasis id="B0E34B533071FF98FDDBFE6FBF920F0F" box="[558,753,508,532]" italics="true" pageId="8" pageNumber="209">C. convexiusculus</emphasis>
</taxonomicName>
members. Finally, anterior tarsomeres 1 to 3 of the female
<typeStatus id="5D2C29E33071FF98FEACFD8EBCD50F2F" box="[345,438,540,564]" pageId="8" pageNumber="209" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4597ECC23071FF98FE28FD8FBF520F2F" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[477,561,540,564]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B0E34B533071FF98FE28FD8FBF520F2F" box="[477,561,540,564]" italics="true" pageId="8" pageNumber="209">C. latus</emphasis>
</taxonomicName>
are distinctly toothed medioapically (
<figureCitation id="1AAC8BC43071FF98FC38FD8EB9430F2F" box="[973,1056,540,565]" captionStart="FIGURE" captionStartId="7.[155,167,1251,1270]" captionTargetBox="[120,1176,216,1233]" captionTargetId="figure-985@7.[120,1176,216,1235]" captionTargetPageId="7" captionText="FIGURE 4. Digital images of holotype female of Caelostomus latus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Head, left lateral aspect (arrow points to tranverse depression between eyes); D. Right anterior tarsus, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799449" httpUri="https://zenodo.org/record/13799449/files/figure.png" pageId="8" pageNumber="209">Fig. 4D</figureCitation>
), a feature not seen in females of the other two species.
</paragraph>
<paragraph id="822897413071FF98FF5EFDCFB9690F6F" blockId="8.[123,1174,219,693]" box="[171,1034,605,629]" pageId="8" pageNumber="209">
<emphasis id="B0E34B533071FF98FF5EFDCFBC920F68" bold="true" box="[171,497,605,629]" pageId="8" pageNumber="209">
<smallCapsWord id="84CE019D3071FF98FF5EFDCFBC350F68" baselines="623,622" box="[171,342,605,629]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="8" pageNumber="209">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="84CE019D3071FF98FEA8FDF2BC920F68" baselines="622" box="[349,497,608,627]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="8" pageNumber="209">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E33071FF98FC8AFDCFBECE0F6E" box="[895,941,605,629]" pageId="8" pageNumber="209">type</typeStatus>
locality.
</paragraph>
<paragraph id="822897413071FF98FF5EFDEFBF6C0FAE" blockId="8.[123,1174,219,693]" pageId="8" pageNumber="209">
<emphasis id="B0E34B533071FF98FF5EFDEFBCC50F88" bold="true" box="[171,422,637,661]" pageId="8" pageNumber="209">
<smallCapsWord id="84CE019D3071FF98FF5EFDEFBC6A0F88" baselines="655,654" box="[171,265,637,661]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="8" pageNumber="209">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D3071FF98FEE7FD12BCC50F88" baselines="654" box="[274,422,640,659]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="8" pageNumber="209">DISTRIBUTION</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C3071FF98FE3BFDEFBF640FAE" location="Talatakely area" pageId="8" pageNumber="209" specimenCount="1" typeStatus="holotype">
The unique
<typeStatus id="5D2C29E33071FF98FDA6FDEFBFD30F8E" box="[595,688,637,661]" pageId="8" pageNumber="209" type="holotype">holotype</typeStatus>
female was collected by hand in secondary montane forest in the
<location id="8748C19A3071FF98FE90FD0FBF640FAE" LSID="urn:lsid:plazi:treatment:0A3E2657307FFF98FF8EFA4EBF6C0FAE:8748C19A3071FF98FE90FD0FBF640FAE" box="[357,519,669,693]" name="Talatakely area" pageId="8" pageNumber="209">Talatakely area</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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