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<document id="8380285F7024B648C76337EE18097D25" ID-ISSN="0068-547X" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E26573041FFA8FF8EFF49B9120B47" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Lebia (Metalebia) ranomafanae Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="1" lastPageNumber="257" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="257" updateTime="1726771413727" updateUser="felipe" zenodo-license-document="CC-BY-4.0">
<mods:mods id="340D2F24CB519F8D188B04A22E689072" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3BB36029E1E7C43CF91D93D46D0CA7A2">
<mods:title id="E163B59EC6DF30A3FE30A9513894EF0D">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="AAACFF7CAE9E2B58BC75BCBF61B86D2C" type="personal">
<mods:role id="90EC63C1CFD4760E38919DE3DB376572">
<mods:roleTerm id="2633214A3D8EE9726509A0A04A8E676D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C659B80AD30D419BC8D49D873BC866E0">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="380F6FA1034250E4E5130E1FCDCF5DC2">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
<mods:name id="1104938DFF0BF74993898E4A22056562" type="personal">
<mods:role id="C74DA2842E802C8262C4165DB712E80C">
<mods:roleTerm id="99B81CB26AF8238AE82053E51C3388FE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5FBB92D8C0950E521B10A118176BA482">Rainio, Johanna</mods:namePart>
<mods:affiliation id="E11F169EBF7850B810513D796E6FD671">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
<mods:typeOfResource id="8C9732818DF40211D6F79434FF26C98C">text</mods:typeOfResource>
<mods:relatedItem id="43ED1E3DFE5774DBCC7B8F26BA14AE60" type="host">
<mods:titleInfo id="7B6FFA9AE42A1C6C78362FAA5E171C9D">
<mods:title id="19BD7DF0C5FE90A8E1F1E88E8A36CE1D">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="5C39A0DAD220A01262767ED7D77F970F">
<mods:date id="83A088DFADCC58D020478932322C3AE4">2016</mods:date>
<mods:detail id="339E25124FBDB172C5ED00329682420D" type="pubDate">
<mods:number id="3F61AEEC983502868284B3C1AB36C0A2">2016-04-29</mods:number>
</mods:detail>
<mods:detail id="147687DAC1CED255106DE46570361277" type="volume">
<mods:number id="147EBA5217BD8C565153B13C5E31ED40">63</mods:number>
</mods:detail>
<mods:detail id="F57DBB5C974E9E2A09ED85CAE03DCDCB" type="issue">
<mods:number id="AFD5E7E143C61C346B2435080662B16C">7</mods:number>
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<mods:start id="0304CAE09A304BD9173E22E41A520FF1">201</mods:start>
<mods:end id="360918C9E43A5BF6AFE0D9AA407352FB">268</mods:end>
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<mods:classification id="7C7B2D91CAC97C5A08D0F7BA67C08488">journal article</mods:classification>
<mods:identifier id="E8909802DA6662AF3417AE26026EB97A" type="ISSN">0068-547X</mods:identifier>
</mods:mods>
<treatment id="0A3E26573041FFA8FF8EFF49B9120B47" LSID="urn:lsid:plazi:treatment:0A3E26573041FFA8FF8EFF49B9120B47" httpUri="http://treatment.plazi.org/id/0A3E26573041FFA8FF8EFF49B9120B47" lastPageNumber="257" pageId="56" pageNumber="257">
<subSubSection id="CA8DC4CA3041FFA8FF8EFF49BE330DE8" box="[123,848,219,243]" pageId="56" pageNumber="257" type="nomenclature">
<paragraph id="822897413041FFA8FF8EFF49BE330DE8" blockId="56.[123,848,219,273]" box="[123,848,219,243]" pageId="56" pageNumber="257">
<heading id="D960202D3041FFA8FF8EFF49BE330DE8" box="[123,848,219,243]" fontSize="10" level="2" pageId="56" pageNumber="257" reason="4">
<emphasis id="B0E34B533041FFA8FF8EFF49BE330DE8" bold="true" box="[123,848,219,243]" pageId="56" pageNumber="257">
<taxonomicName id="4597ECC23041FFA8FF8EFF49BF8F0DE8" ID-CoL="6P429" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,748,219,243]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov." subGenus="Metalebia">
<emphasis id="B0E34B533041FFA8FF8EFF49BDDA0DE8" bold="true" box="[123,185,219,243]" italics="true" pageId="56" pageNumber="257">Lebia</emphasis>
(
<emphasis id="B0E34B533041FFA8FF3CFF49BC540DE8" bold="true" box="[201,311,219,243]" italics="true" pageId="56" pageNumber="257">Metalebia</emphasis>
)
<emphasis id="B0E34B533041FFA8FEB2FF49BCB80DE8" bold="true" box="[327,475,219,243]" italics="true" pageId="56" pageNumber="257">ranomafanae</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F6283041FFA8FD0FFF49BE330DE8" box="[762,848,219,243]" pageId="56" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3041FFA8FF8EFF69BDBB0C0B" box="[123,216,251,273]" pageId="56" pageNumber="257" type="description">
<paragraph id="822897413041FFA8FF8EFF69BDBB0C0B" blockId="56.[123,848,219,273]" box="[123,216,251,273]" pageId="56" pageNumber="257">
<figureCitation id="1AAC8BC43041FFA8FF8EFF69BDBB0C0B" box="[123,216,251,273]" captionStart="FIGURE" captionStartId="57.[155,167,1476,1495]" captionTargetBox="[137,1174,222,1457]" captionTargetId="figure-68@57.[118,1174,222,1472]" captionTargetPageId="57" captionText="FIGURE 34. Digital images of holotype male of Lebia (Metalebia) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 0.5 mm." pageId="56" pageNumber="257">Figure 34</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3041FFA8FF5EFEA7B9120B47" pageId="56" pageNumber="257" type="materials_examined">
<paragraph id="822897413041FFA8FF5EFEA7BE410F14" blockId="56.[123,1174,309,1628]" pageId="56" pageNumber="257">
<emphasis id="B0E34B533041FFA8FF5EFEA7BC390C50" bold="true" box="[171,346,309,333]" pageId="56" pageNumber="257">
<smallCapsWord id="84CE019D3041FFA8FF5EFEA7BD870C57" baselines="327,327" box="[171,228,309,333]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="56" pageNumber="257">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D3041FFA8FF18FEAABC390C50" baselines="326" box="[237,346,312,331]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="56" pageNumber="257">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C3041FFA8FE77FEA7BC8B0CB5" collectingDate="1988-10-22" collectingDateMax="1988-10-31" collectingDateMin="1988-10-22" collectorName="W. E. Steiner" country="Madagascar" elevation="1100" location="Ranomafana" pageId="56" pageNumber="257" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E33041FFA8FE77FEA7BC860C56" box="[386,485,309,333]" pageId="56" pageNumber="257" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43041FFA8FE02FEA7BF1A0C57" box="[503,633,309,333]" captionStart="FIGURE" captionStartId="57.[155,167,1476,1495]" captionTargetBox="[137,1174,222,1457]" captionTargetId="figure-68@57.[118,1174,222,1472]" captionTargetPageId="57" captionText="FIGURE 34. Digital images of holotype male of Lebia (Metalebia) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 0.5 mm." pageId="56" pageNumber="257">Figs.34A–B</figureCitation>
), a male, in NMNH, labeled: “
<collectingCountry id="FA80D7D13041FFA8FC25FEA7B9ED0C56" box="[976,1166,309,333]" name="Madagascar" pageId="56" pageNumber="257">MADAGASCAR</collectingCountry>
: Prov.
<collectingRegion id="405359A33041FFA8FF4DFEC7BC230C76" box="[184,320,341,365]" country="Madagascar" name="Fianarantsoa" pageId="56" pageNumber="257">Fianarantsoa</collectingRegion>
,
<quantity id="456F3AA43041FFA8FEB9FEC4BCE20C76" box="[332,385,341,365]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="56" pageNumber="257" unit="km" value="7.0">7 km</quantity>
W
<location id="8748C19A3041FFA8FE50FEC7BF4F0C76" LSID="urn:lsid:plazi:treatment:0A3E26573041FFA8FF8EFF49B9120B47:8748C19A3041FFA8FE50FEC7BF4F0C76" box="[421,556,341,365]" country="Madagascar" name="Ranomafana" pageId="56" pageNumber="257" stateProvince="Fianarantsoa">Ranomafana</location>
,
<quantity id="456F3AA43041FFA8FDCDFEC7BFE10C76" box="[568,642,341,365]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="56" pageNumber="257" unit="m" value="1100.0">
<elevation id="09BA70723041FFA8FDCDFEC7BFE10C76" box="[568,642,341,365]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="56" pageNumber="257" unit="m" value="1100.0">1100m</elevation>
</quantity>
<date id="F629B1813041FFA8FD72FEC7BE3F0C76" box="[647,860,341,365]" pageId="56" pageNumber="257" value="1988-10-22" valueMax="1988-10-31" valueMin="1988-10-22">
<collectingDate id="E66D48693041FFA8FD72FEC7BE3F0C76" box="[647,860,341,365]" pageId="56" pageNumber="257" value="1988-10-22" valueMax="1988-10-31" valueMin="1988-10-22">22-31 October 1988</collectingDate>
</date>
<collectorName id="2F62F2973041FFA8FC94FEC4BE8E0C76" box="[865,1005,341,365]" pageId="56" pageNumber="257">W. E. Steiner</collectorName>
”/ “
<collectingMethod id="5BD6EF563041FFA8FBE7FEC7B9F60C76" box="[1042,1173,341,365]" pageId="56" pageNumber="257">Malaise trap</collectingMethod>
in small clearing, montane rain forest”/ “
<typeStatus id="5D2C29E33041FFA8FDD8FEE4BFD90C95" box="[557,698,374,398]" pageId="56" pageNumber="257" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23041FFA8FD35FEE4BDBC0CB5" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov." subGenus="Metalebia">
<emphasis id="B0E34B533041FFA8FD35FEE4BF9E0C95" box="[704,765,374,398]" italics="true" pageId="56" pageNumber="257">Lebia</emphasis>
(
<emphasis id="B0E34B533041FFA8FCF9FEE4BE1B0C95" box="[780,888,374,398]" italics="true" pageId="56" pageNumber="257">Metalebia</emphasis>
)
<emphasis id="B0E34B533041FFA8FC72FEE4B9760C95" box="[903,1045,374,398]" italics="true" pageId="56" pageNumber="257">ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283041FFA8FF1CFE05BC470CB5" box="[233,292,407,430]" pageId="56" pageNumber="257" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label]
</materialsCitation>
.
<materialsCitation id="32FF9D1C3041FFA8FE0DFE05BE780F14" collectingDate="1990-02-23" collectingDateMax="1990-02-28" collectingDateMin="1990-02-23" collectionCode="CAS" collectorName="W. E. Steiner" country="Madagascar" elevation="900" location="Ranomafana" pageId="56" pageNumber="257" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="5D2C29E33041FFA8FE0DFE05BF030CB4" box="[504,608,407,431]" pageId="56" pageNumber="257" type="paratype">Paratypes</typeStatus>
(only 1): a female (in
<collectionCode id="E4860F843041FFA8FCABFE04BEF10CB5" box="[862,914,406,430]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="56" pageNumber="257" type="Museum">CAS</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13041FFA8FBF3FE04BDD10CD4" name="Madagascar" pageId="56" pageNumber="257">MADAGASCAR</collectingCountry>
: Prov.
<collectingRegion id="405359A33041FFA8FEFCFE24BCF20CD5" box="[265,401,438,462]" country="Madagascar" name="Fianarantsoa" pageId="56" pageNumber="257">Fianarantsoa</collectingRegion>
,
<quantity id="456F3AA43041FFA8FE51FE25BCBC0CD5" box="[420,479,438,462]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="56" pageNumber="257" unit="km" value="7.0">7 km</quantity>
W
<location id="8748C19A3041FFA8FDE5FE24BFF40CD5" LSID="urn:lsid:plazi:treatment:0A3E26573041FFA8FF8EFF49B9120B47:8748C19A3041FFA8FDE5FE24BFF40CD5" box="[528,663,438,462]" country="Madagascar" name="Ranomafana" pageId="56" pageNumber="257" stateProvince="Fianarantsoa">Ranomafana</location>
,
<quantity id="456F3AA43041FFA8FD5CFE25BF850CD5" box="[681,742,439,463]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="56" pageNumber="257" unit="m" value="900.0">
<elevation id="09BA70723041FFA8FD5CFE25BF850CD5" box="[681,742,439,463]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="56" pageNumber="257" unit="m" value="900.0">900m</elevation>
</quantity>
<date id="F629B1813041FFA8FD07FE25BEBD0CD4" box="[754,990,438,463]" pageId="56" pageNumber="257" value="1990-02-23" valueMax="1990-02-28" valueMin="1990-02-23">
<collectingDate id="E66D48693041FFA8FD07FE25BEBD0CD4" box="[754,990,438,463]" pageId="56" pageNumber="257" value="1990-02-23" valueMax="1990-02-28" valueMin="1990-02-23">23-28 February 1990</collectingDate>
</date>
<collectorName id="2F62F2973041FFA8FC1FFE25B9E00CD5" box="[1002,1155,438,462]" pageId="56" pageNumber="257">W. E. Steiner</collectorName>
”/ “
<collectingMethod id="5BD6EF563041FFA8FF72FE45BC790CF5" box="[135,282,471,495]" pageId="56" pageNumber="257">Malaise trap</collectingMethod>
in small clearing, montane rain forest”/ “
<typeStatus id="5D2C29E33041FFA8FCDEFE45BED10CF4" box="[811,946,471,495]" pageId="56" pageNumber="257" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC23041FFA8FC36FE45BC970F14" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov." subGenus="Metalebia">
<emphasis id="B0E34B533041FFA8FC36FE45B9620CF4" box="[963,1025,471,495]" italics="true" pageId="56" pageNumber="257">Lebia</emphasis>
(
<emphasis id="B0E34B533041FFA8FBE9FE45B9EF0CF4" box="[1052,1164,471,495]" italics="true" pageId="56" pageNumber="257">Metalebia</emphasis>
)
<emphasis id="B0E34B533041FFA8FF8EFE65BC6A0F14" box="[123,265,503,527]" italics="true" pageId="56" pageNumber="257">ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283041FFA8FE0EFE6ABF550F14" box="[507,566,504,527]" pageId="56" pageNumber="257" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="822897413041FFA8FF5EFD8AB9500F34" blockId="56.[123,1174,309,1628]" box="[171,1075,536,560]" pageId="56" pageNumber="257">
<materialsCitation id="32FF9D1C3041FFA8FF5EFD8AB94F0F2B" box="[171,1068,536,560]" country="Madagascar" location="Ranomafana National Park" pageId="56" pageNumber="257" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533041FFA8FF5EFD8ABC340F35" bold="true" box="[171,343,536,560]" pageId="56" pageNumber="257">
<smallCapsWord id="84CE019D3041FFA8FF5EFD8ABD870F35" baselines="554,553" box="[171,228,536,560]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="56" pageNumber="257">
<typeStatus id="5D2C29E33041FFA8FF5EFD8ABD870F35" box="[171,228,536,560]" pageId="56" pageNumber="257">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3041FFA8FF19FD89BC340F35" baselines="553" box="[236,343,539,558]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="56" pageNumber="257">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13041FFA8FE8BFD8ABC9C0F2B" box="[382,511,536,560]" name="Madagascar" pageId="56" pageNumber="257">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33041FFA8FDF8FD8ABF9F0F2B" box="[525,764,536,560]" country="Madagascar" name="Fianarantsoa" pageId="56" pageNumber="257">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3041FFA8FCFFFD8AB94F0F2B" LSID="urn:lsid:plazi:treatment:0A3E26573041FFA8FF8EFF49B9120B47:8748C19A3041FFA8FCFFFD8AB94F0F2B" box="[778,1068,536,560]" country="Madagascar" name="Ranomafana National Park" pageId="56" pageNumber="257" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413041FFA8FF5EFDAABECA0F6B" blockId="56.[123,1174,309,1628]" pageId="56" pageNumber="257">
<emphasis id="B0E34B533041FFA8FF5EFDAABC910F54" bold="true" box="[171,498,568,592]" pageId="56" pageNumber="257">
<smallCapsWord id="84CE019D3041FFA8FF5EFDAABC4D0F55" baselines="586,585" box="[171,302,568,592]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="56" pageNumber="257">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3041FFA8FEC3FDA9BC310F55" baselines="585" box="[310,338,571,590]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="56" pageNumber="257">OF</smallCapsWord>
<smallCapsWord id="84CE019D3041FFA8FEACFDA9BCCF0F55" baselines="585" box="[345,428,571,590]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="56" pageNumber="257">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3041FFA8FE40FDAEBC910F54" baselines="586" box="[437,498,572,591]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="56" pageNumber="257">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23041FFA8FD0CFDAABEE40F4B" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[761,903,568,592]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533041FFA8FD0CFDAABEE40F4B" box="[761,903,568,592]" italics="true" pageId="56" pageNumber="257">ranomafanae</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the national park in which the
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was collected.
</paragraph>
<paragraph id="822897413041FFA8FF5EFDEBBFC508E0" blockId="56.[123,1174,309,1628]" pageId="56" pageNumber="257">
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<emphasis id="B0E34B533041FFA8FF5EFDEBBC210F94" bold="true" box="[171,322,633,657]" pageId="56" pageNumber="257">RECOGNITION</emphasis>
</smallCapsWord>
.— Size slightly small for genus, SBL of male =
<quantity id="456F3AA43041FFA8FCA3FDEBBECA0F8B" box="[854,937,633,657]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="56" pageNumber="257" unit="mm" value="4.3">4.3 mm</quantity>
, of female =
<quantity id="456F3AA43041FFA8FBC9FDEBB9ED0F8B" box="[1084,1166,633,657]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="56" pageNumber="257" unit="mm" value="4.2">4.2 mm</quantity>
. Members of this species (
<figureCitation id="1AAC8BC43041FFA8FE67FD0BBC910FAA" box="[402,498,665,689]" captionStart="FIGURE" captionStartId="57.[155,167,1476,1495]" captionTargetBox="[137,1174,222,1457]" captionTargetId="figure-68@57.[118,1174,222,1472]" captionTargetPageId="57" captionText="FIGURE 34. Digital images of holotype male of Lebia (Metalebia) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 0.5 mm." pageId="56" pageNumber="257">Fig. 34A</figureCitation>
) key to subgenus
<taxonomicName id="4597ECC23041FFA8FD4FFD0BBEAE0FAA" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" box="[698,973,665,689]" pageId="56" pageNumber="257" rank="subGenus" subGenus="Metalebia">
<emphasis id="B0E34B533041FFA8FD4FFD0BBE450FAA" box="[698,806,665,689]" italics="true" pageId="56" pageNumber="257">Metalebia</emphasis>
Jeannel (1949)
</taxonomicName>
and the
<taxonomicName id="4597ECC23041FFA8FBD9FD0BBDB80FCA" authorityName="Chaudoir" authorityYear="1850" class="Insecta" family="Carabidae" genus="Assadecma" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B0E34B533041FFA8FBD9FD0BBDB80FCA" italics="true" pageId="56" pageNumber="257">madagascariensis</emphasis>
</taxonomicName>
group in Jeannel’s (1949) key: the basal elytral border is absent medially between the base of stria 3 and the scutellum; the elytral striae are deep and the elytral intervals convex and impunctate; the forebody (head plus pronotum) has no metallic reflection and the elytra are uniform in color (but not necessarily in reflection); and the frons is slightly convex and without distinct frontal furrows between the eyes. Within the
<taxonomicName id="4597ECC23041FFA8FD66FCA9BE370E48" authorityName="Chaudoir" authorityYear="1850" box="[659,852,827,851]" class="Insecta" family="Carabidae" genus="Assadecma" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B0E34B533041FFA8FD66FCA9BE370E48" box="[659,852,827,851]" italics="true" pageId="56" pageNumber="257">madagascariensis</emphasis>
</taxonomicName>
group,
<taxonomicName id="4597ECC23041FFA8FC53FCAEB9330E48" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[934,1104,827,851]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533041FFA8FC53FCAEB9330E48" box="[934,1104,827,851]" italics="true" pageId="56" pageNumber="257">L. ranomafanae</emphasis>
</taxonomicName>
members can be distinguished from those of all species except
<taxonomicName id="4597ECC23041FFA8FD16FCCEBE1B0E68" authorityName="Jeannel" authorityYear="1949" box="[739,888,859,883]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="alluaudana">
<emphasis id="B0E34B533041FFA8FD16FCCEBE1B0E68" box="[739,888,859,883]" italics="true" pageId="56" pageNumber="257">L. alluaudana</emphasis>
</taxonomicName>
,
<taxonomicName id="4597ECC23041FFA8FC70FCCEB9540E68" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[901,1079,859,883]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="apicoviolacea" status="sp. nov.">
<emphasis id="B0E34B533041FFA8FC70FCCEB9540E68" box="[901,1079,859,883]" italics="true" pageId="56" pageNumber="257">L. apicoviolacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283041FFA8FBC8FCCEB9EC0E68" box="[1085,1167,860,883]" pageId="56" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4597ECC23041FFA8FF8EFCEFBD850E8F" authorityName="Alluaud" authorityYear="1936" box="[123,230,893,916]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="mirana">
<emphasis id="B0E34B533041FFA8FF8EFCEFBD850E8F" box="[123,230,893,916]" italics="true" pageId="56" pageNumber="257">L. mirana</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC23041FFA8FEEAFCEFBCE10E8F" authorityName="Jeannel" authorityYear="1949" box="[287,386,892,916]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="tanala">
<emphasis id="B0E34B533041FFA8FEEAFCEFBCE10E8F" box="[287,386,892,916]" italics="true" pageId="56" pageNumber="257">L. tanala</emphasis>
</taxonomicName>
by the presence of elytral metallic reflection (metallic reflection absent from the elytra of members of the remaining seven species). They are distinguished from members of
<taxonomicName id="4597ECC23041FFA8FF6CFC2FBC670ECF" authorityName="Alluaud" authorityYear="1936" box="[153,260,957,980]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="mirana">
<emphasis id="B0E34B533041FFA8FF6CFC2FBC670ECF" box="[153,260,957,980]" italics="true" pageId="56" pageNumber="257">L. mirana</emphasis>
</taxonomicName>
by their reddish labrum (black in
<taxonomicName id="4597ECC23041FFA8FD8EFC2FBF860ECF" authorityName="Alluaud" authorityYear="1936" box="[635,741,957,980]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="mirana">
<emphasis id="B0E34B533041FFA8FD8EFC2FBF860ECF" box="[635,741,957,980]" italics="true" pageId="56" pageNumber="257">L. mirana</emphasis>
</taxonomicName>
members) and from those of
<taxonomicName id="4597ECC23041FFA8FBDEFC2FBDD30EEE" authorityName="Jeannel" authorityYear="1949" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="alluaudana">
<emphasis id="B0E34B533041FFA8FBDEFC2FBDD30EEE" italics="true" pageId="56" pageNumber="257">L. alluaudana</emphasis>
</taxonomicName>
by their small size and relatively short and broad elytra, with ratio of elytra length to width = 1.3 (size larger, SBL of males = 4.6–5.0 mm, of females 4.6–
<quantity id="456F3AA43041FFA8FCF8FC6FBE3C090E" box="[781,863,1021,1045]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="56" pageNumber="257" unit="mm" value="5.3">5.3 mm</quantity>
, and elytra relatively longer, with ratio EL/EW = 1.4, in
<taxonomicName id="4597ECC23041FFA8FE59FB8DBF26092D" authorityName="Jeannel" authorityYear="1949" box="[428,581,1054,1078]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="alluaudana">
<emphasis id="B0E34B533041FFA8FE59FB8DBF26092D" box="[428,581,1054,1078]" italics="true" pageId="56" pageNumber="257">L. alluaudana</emphasis>
</taxonomicName>
members). They are distinguished from members of
<taxonomicName id="4597ECC23041FFA8FF8EFBADBDB8094D" authorityName="Jeannel" authorityYear="1949" box="[123,219,1086,1110]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="tanala">
<emphasis id="B0E34B533041FFA8FF8EFBADBDB8094D" box="[123,219,1086,1110]" italics="true" pageId="56" pageNumber="257">L. tanala</emphasis>
</taxonomicName>
in having very dark and relatively faint elytral metallic reflection, pronotum with anteri- or angles indistinct, smoothly rounded from apical to lateral margin and not at all projected anteriorly beyond apical margin, lateral margins nearly parallel in posterior two-thirds and not or only very slightly sinuate anterior to obtusely angulate hind angles (elytral metallic reflection more distinct, pronotum with anterior angles broadly round but slightly projected anteriorly beyond apical margin, lateral margins distinctly sinuate anterior to sharp, rectangular hind angles slightly projected laterally in members of
<taxonomicName id="4597ECC23041FFA8FE21FA93BF5A0803" authorityName="Jeannel" authorityYear="1949" box="[468,569,1280,1304]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="tanala">
<emphasis id="B0E34B533041FFA8FE21FA93BF5A0803" box="[468,569,1280,1304]" italics="true" pageId="56" pageNumber="257">L. tanala</emphasis>
</taxonomicName>
). Finally, they can be distinguished from members of
<taxonomicName id="4597ECC23041FFA8FF8EFAB0BC4D0822" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,302,1313,1337]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="apicoviolacea">
<emphasis id="B0E34B533041FFA8FF8EFAB0BC4D0822" box="[123,302,1313,1337]" italics="true" pageId="56" pageNumber="257">L. apicoviolacea</emphasis>
</taxonomicName>
by slightly larger size and the faint, concolorous dark blue-green elytral metallic reflection and (size slightly smaller, SBL of male =
<quantity id="456F3AA43041FFA8FD59FAD3BF9C0842" box="[684,767,1345,1369]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="56" pageNumber="257" unit="mm" value="3.8">3.8 mm</quantity>
, of female = 4.0 mm, and elytra with distinct tricolored metallic reflection, green basally and on medial two-thirds, brassy on lateral one-third and violet in apical one-fifth, in
<taxonomicName id="4597ECC23041FFA8FDE7FA11BFA50881" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[530,710,1410,1434]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="257" phylum="Arthropoda" rank="species" species="apicoviolacea">
<emphasis id="B0E34B533041FFA8FDE7FA11BFA50881" box="[530,710,1410,1434]" italics="true" pageId="56" pageNumber="257">L. apicoviolacea</emphasis>
</taxonomicName>
members. Shape of the median lobe of the male aedeagus (
<figureCitation id="1AAC8BC43041FFA8FEDDFA30BCC408A1" box="[296,423,1442,1466]" captionStart="FIGURE" captionStartId="57.[155,167,1476,1495]" captionTargetBox="[137,1174,222,1457]" captionTargetId="figure-68@57.[118,1174,222,1472]" captionTargetPageId="57" captionText="FIGURE 34. Digital images of holotype male of Lebia (Metalebia) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 0.5 mm." pageId="56" pageNumber="257">Fig. 34C–D</figureCitation>
) is also diagnostic, with the shaft broadest at basal one-third, slightly and evenly arcuate on ventral margin in lateral view (
<figureCitation id="1AAC8BC43041FFA8FD4EFA51BE7A08C0" box="[699,793,1475,1499]" captionStart="FIGURE" captionStartId="57.[155,167,1476,1495]" captionTargetBox="[137,1174,222,1457]" captionTargetId="figure-68@57.[118,1174,222,1472]" captionTargetPageId="57" captionText="FIGURE 34. Digital images of holotype male of Lebia (Metalebia) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 0.5 mm." pageId="56" pageNumber="257">Fig. 34C</figureCitation>
) and apex parallel-side basally and broadly rounded apically in dorsal view (
<figureCitation id="1AAC8BC43041FFA8FDC3FA71BFF508E0" box="[566,662,1507,1531]" captionStart="FIGURE" captionStartId="57.[155,167,1476,1495]" captionTargetBox="[137,1174,222,1457]" captionTargetId="figure-68@57.[118,1174,222,1472]" captionTargetPageId="57" captionText="FIGURE 34. Digital images of holotype male of Lebia (Metalebia) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 0.5 mm." pageId="56" pageNumber="257">Fig. 34D</figureCitation>
).
</paragraph>
<paragraph id="822897413041FFA8FF5EF991B9690B00" blockId="56.[123,1174,309,1628]" box="[171,1034,1539,1564]" pageId="56" pageNumber="257">
<emphasis id="B0E34B533041FFA8FF5EF991BC920B01" bold="true" box="[171,497,1539,1563]" pageId="56" pageNumber="257">
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<smallCapsWord id="84CE019D3041FFA8FEA8F995BC920B01" baselines="1557" box="[349,497,1543,1562]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="56" pageNumber="257">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E33041FFA8FC8AF996BECE0B07" box="[895,941,1540,1564]" pageId="56" pageNumber="257">type</typeStatus>
locality.
</paragraph>
<paragraph id="822897413041FFA8FF5EF9B6B9120B47" blockId="56.[123,1174,309,1628]" pageId="56" pageNumber="257">
<emphasis id="B0E34B533041FFA8FF5EF9B6BCC00B21" bold="true" box="[171,419,1572,1596]" pageId="56" pageNumber="257">
<smallCapsWord id="84CE019D3041FFA8FF5EF9B6BC6A0B21" baselines="1590,1589" box="[171,265,1572,1596]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="56" pageNumber="257">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D3041FFA8FEFAF9B5BCC00B21" baselines="1589" box="[271,419,1575,1594]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="56" pageNumber="257">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Both specimens of the
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series were collected in malaise traps in secondary montane rainforest in the Talatakely area at elevations ranging from
<quantity id="456F3AA43041FFA8FC3BF9D6B9090B47" box="[974,1130,1604,1628]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="56" pageNumber="257" unit="m" value="1000.0" valueMax="1100.0" valueMin="900.0">900 to 1100 m</quantity>
.
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<mods:title id="E30A72B93CA52EAA9A852B91466B1A75">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<taxonomicName id="4597ECC23042FFABFF8EFA33BF3808A2" ID-CoL="3CG64" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,603,1441,1465]" class="Insecta" family="Carabidae" genus="Eunostus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B0E34B533042FFABFF8EFA33BC2908A2" bold="true" box="[123,330,1441,1465]" italics="true" pageId="59" pageNumber="260">Eunostus minimus</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F6283042FFABFD9CFA33BFDC08A2" box="[617,703,1441,1465]" pageId="59" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3042FFABFF8EFA53BE350B1A" pageId="59" pageNumber="260" type="description">
<paragraph id="822897413042FFABFF8EFA53BC6608CD" blockId="59.[123,703,1441,1495]" box="[123,261,1473,1495]" pageId="59" pageNumber="260">
<figureCitation id="1AAC8BC43042FFABFF8EFA53BC6608CD" box="[123,261,1473,1495]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStartId-0="60.[155,167,1162,1181]" captionStartId-1="61.[155,167,825,844]" captionTargetBox-0="[130,1176,216,1147]" captionTargetBox-1="[120,1176,216,803]" captionTargetId-0="figure-1103@60.[120,1176,216,1159]" captionTargetId-1="figure-1453@61.[120,1176,216,803]" captionTargetPageId-0="60" captionTargetPageId-1="61" captionText-0="FIGURE 36. Digital images of holotype male of Eunostus minimus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line A = 1.0 mm." captionText-1="FIGURE 37. Digital images of Eunostus minimus Kavanaugh &amp; Rainio sp. nov. A–B. Apical part of left hind femur, oblique medial aspect; A. Holotype male; B. Female; C–D. Aedeagus of holotype male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 0.5 mm." pageId="59" pageNumber="260">Figures 36–37</figureCitation>
</paragraph>
<paragraph id="822897413042FFABFF8EFA7FBE350B1A" blockId="59.[123,854,1516,1538]" box="[123,854,1516,1538]" pageId="59" pageNumber="260">
<emphasis id="B0E34B533042FFABFF8EFA7FBC430B1A" box="[123,288,1516,1538]" italics="true" pageId="59" pageNumber="260">
<taxonomicName id="4597ECC23042FFABFF8EFA7FBDB00B19" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,211,1517,1538]" class="Insecta" family="Carabidae" genus="Eunostus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="genus" status="n. sp.">Eunostus</taxonomicName>
<taxonomicNameLabel id="ABD0F6283042FFABFF2CFA7FBC6E0B19" box="[217,269,1517,1538]" pageId="59" pageNumber="260" rank="species">n. sp.</taxonomicNameLabel>
1
</emphasis>
; Rainio 2009: 31, Rainio 2012: 72 (informal designation).
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3042FFADFF5EF9B7BF1E0823" lastPageId="61" lastPageNumber="262" pageId="59" pageNumber="260" type="materials_examined">
<paragraph id="822897413042FFACFF5EF9B7BC1808A3" blockId="59.[123,1174,1573,1726]" lastBlockId="60.[123,1174,1246,1725]" lastPageId="60" lastPageNumber="261" pageId="59" pageNumber="260">
<emphasis id="B0E34B533042FFABFF5EF9B7BC390B20" bold="true" box="[171,346,1573,1597]" pageId="59" pageNumber="260">
<smallCapsWord id="84CE019D3042FFABFF5EF9B7BD870B27" baselines="1591,1591" box="[171,228,1573,1597]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="59" pageNumber="260">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D3042FFABFF18F9BABC390B20" baselines="1590" box="[237,346,1576,1595]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="59" pageNumber="260">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C3042FFABFE74F9B7BEA70BA6" collectingDate="2003-03-27" collectingDateMax="2003-03-31" collectingDateMin="2003-03-27" collectorName="Fisher" country="Madagascar" county="Parc National de Ranomafana" elevation="1100" latitude="-21.289999" location="California Acad. of Sciences" longLatPrecision="21" longitude="47.433334" municipality="Vatoharanana River" pageId="59" pageNumber="260" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E33042FFABFE74F9B7BC870B26" box="[385,484,1573,1597]" pageId="59" pageNumber="260" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43042FFABFE00F9B7BF1C0B27" box="[501,639,1573,1597]" captionStart="FIGURE" captionStartId="60.[155,167,1162,1181]" captionTargetBox="[130,1176,216,1147]" captionTargetId="figure-1103@60.[120,1176,216,1159]" captionTargetPageId="60" captionText="FIGURE 36. Digital images of holotype male of Eunostus minimus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line A = 1.0 mm." pageId="59" pageNumber="260">Figs. 36A–B</figureCitation>
), a male, in CAS, labeled: “CASENT 8069195”/ “
<collectingCountry id="FA80D7D13042FFABFF73F9D7BC270B46" box="[134,324,1605,1629]" name="Madagascar" pageId="59" pageNumber="260">MADAGASCAR</collectingCountry>
:
<collectingRegion id="405359A33042FFABFEA6F9D7BCB80B46" box="[339,475,1605,1629]" country="Madagascar" name="Fianarantsoa" pageId="59" pageNumber="260">Fianarantsoa</collectingRegion>
<collectingCounty id="6B49EFCD3042FFABFE17F9D4BE400B46" box="[482,803,1605,1629]" pageId="59" pageNumber="260">Parc National de Ranomafana</collectingCounty>
,
<collectingMunicipality id="624C0D3B3042FFABFCC5F9D7B9600B46" box="[816,1027,1605,1629]" pageId="59" pageNumber="260">Vatoharanana River</collectingMunicipality>
,
<locationDeviation id="939A8E633042FFABFBE4F9D7BC500B66" location="Ranomafana" pageId="59" pageNumber="260">
<quantity id="456F3AA43042FFABFBE4F9D7B9380B46" box="[1041,1115,1605,1629]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="59" pageNumber="260" unit="km" value="4.1">4.1 km</quantity>
231° SW Ranomafana
</locationDeviation>
elev
<quantity id="456F3AA43042FFABFE86F9F7BCDF0B66" box="[371,444,1637,1661]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="59" pageNumber="260" unit="m" value="1100.0">
<elevation id="09BA70723042FFABFE86F9F7BCDF0B66" box="[371,444,1637,1661]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="59" pageNumber="260" unit="m" value="1100.0">1100m</elevation>
</quantity>
<date id="F629B1813042FFABFE33F9F7BF1A0B66" box="[454,633,1637,1661]" pageId="59" pageNumber="260" value="2003-03-27" valueMax="2003-03-31" valueMin="2003-03-27">
<collectingDate id="E66D48693042FFABFE33F9F7BF1A0B66" box="[454,633,1637,1661]" pageId="59" pageNumber="260" value="2003-03-27" valueMax="2003-03-31" valueMin="2003-03-27">27-31 Mar 2003</collectingDate>
</date>
”/ “
<geoCoordinate id="E7A3F1863042FFABFD56F9F7BE420B66" box="[675,801,1637,1661]" degrees="21" direction="south" minutes="17" orientation="latitude" pageId="59" pageNumber="260" precision="15" seconds="24" value="-21.289999">21°17’24”S</geoCoordinate>
<geoCoordinate id="E7A3F1863042FFABFCDEF9F7BEDB0B66" box="[811,952,1637,1661]" degrees="047" direction="east" minutes="26" orientation="longitude" pageId="59" pageNumber="260" precision="15" seconds="00" value="47.433334">047°26’00”E</geoCoordinate>
<location id="8748C19A3042FFABFC37F9F7BDBB0B86" LSID="urn:lsid:plazi:treatment:0A3E26573042FFADFF8EFA33BF1E0823:8748C19A3042FFABFC37F9F7BDBB0B86" country="Madagascar" county="Parc National de Ranomafana" latitude="-21.289999" longLatPrecision="21" longitude="47.433334" municipality="Vatoharanana River" name="California Acad. of Sciences" pageId="59" pageNumber="260" stateProvince="Fianarantsoa">California Acad. of Sciences</location>
coll.
<collectorName id="2F62F2973042FFABFEE2F917BC3A0B86" box="[279,345,1669,1693]" pageId="59" pageNumber="260">Fisher</collectorName>
, Griswold et al. sifted litter, montane rainforest collection code: BLF8400”/ “
<typeStatus id="5D2C29E33042FFABFF72F934BC770BA5" box="[135,276,1702,1726]" pageId="59" pageNumber="260" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23042FFABFEEFF934BFA40BA6" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[282,711,1701,1726]" class="Insecta" family="Carabidae" genus="Eunostus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B0E34B533042FFABFEEFF934BCBE0BA6" box="[282,477,1702,1725]" italics="true" pageId="59" pageNumber="260">Eunostus minimus</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283042FFABFD3BF934BE640BA6" box="[718,775,1702,1725]" pageId="59" pageNumber="260" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label]
</materialsCitation>
.
<materialsCitation id="32FF9D1C3042FFACFC27F934BC1708A3" collectingDate="2001-12-21" collectingDateMax="2004-08-24" collectingDateMin="2001-12-21" collectionCode="MNHN, CAS" collectorName="R. Harin'Hala California Acad &amp; F. Ratalata &amp; J. Rainio" country="Madagascar" county="Ranomafana" elevation="690" lastPageId="60" lastPageNumber="261" latitude="-21.2485" location="Ranomafana" longLatPrecision="12" longitude="47.452168" municipality="Harin'Hala California Acad of Sciences" pageId="59" pageNumber="260" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="5D2C29E33042FFABFC27F934B9590BA5" box="[978,1082,1702,1726]" pageId="59" pageNumber="260" type="paratype">Paratypes</typeStatus>
(total of 2):
<specimenCount id="94915CC83045FFACFF5FFB4CBD9D09ED" box="[170,254,1246,1270]" count="1" pageId="60" pageNumber="261" type="male">1 male</specimenCount>
(in
<collectionCode id="E4860F843045FFACFECAFB4CBCF109EE" box="[319,402,1246,1269]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="60" pageNumber="261">MNHN</collectionCode>
) labeled “CASENT 8069129”/ “
<collectingCountry id="FA80D7D13045FFACFCD3FB4CBE8B09ED" box="[806,1000,1246,1270]" name="Madagascar" pageId="60" pageNumber="261">MADAGASCAR</collectingCountry>
:
<collectingRegion id="405359A33045FFACFBF4FB4CB9ED09ED" box="[1025,1166,1246,1270]" country="Madagascar" name="Fianarantsoa" pageId="60" pageNumber="261">Fianarantsoa</collectingRegion>
,
<collectingCounty id="6B49EFCD3045FFACFF8EFB6CBC61080D" box="[123,258,1278,1302]" pageId="60" pageNumber="261">Ranomafana</collectingCounty>
JIRAMA water works,
<geoCoordinate id="E7A3F1863045FFACFDF1FB6CBF1E080D" box="[516,637,1278,1302]" degrees="21" direction="south" minutes="14. 91" orientation="latitude" pageId="60" pageNumber="261" precision="9" value="-21.2485">21°14.91’S</geoCoordinate>
,
<geoCoordinate id="E7A3F1863045FFACFD7EFB6CBE70080D" box="[651,787,1278,1302]" degrees="047" direction="east" minutes="27. 13" orientation="longitude" pageId="60" pageNumber="261" precision="9" value="47.452168">047°27.13’E</geoCoordinate>
”/ “
<date id="F629B1813045FFACFCCCFB6CB94A080D" box="[825,1065,1278,1302]" pageId="60" pageNumber="261" value="2001-12-21" valueMax="2001-12-24" valueMin="2001-12-21">
<collectingDate id="E66D48693045FFACFCCCFB6CB94A080D" box="[825,1065,1278,1302]" pageId="60" pageNumber="261" value="2001-12-21" valueMax="2001-12-24" valueMin="2001-12-21">21-24 December 2001</collectingDate>
</date>
collector:
<collectorName id="2F62F2973045FFACFF8EFA8DBCA5082C" box="[123,454,1311,1335]" pageId="60" pageNumber="261">
R.
<collectingMunicipality id="624C0D3B3045FFACFF6EFA8DBCA5082C" box="[155,454,1311,1335]" pageId="60" pageNumber="261">Harin’Hala California Acad</collectingMunicipality>
</collectorName>
of Sciences malaise trap near river elev.
<quantity id="456F3AA43045FFACFC73FA8DBEA8082D" box="[902,971,1311,1335]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="60" pageNumber="261" unit="m" value="690.0">
<elevation id="09BA70723045FFACFC73FA8DBEA8082D" box="[902,971,1311,1335]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="60" pageNumber="261" unit="m" value="690.0">690 m</elevation>
</quantity>
, MA-02-09D-08”;
<specimenCount id="94915CC83045FFACFF8EFAADBDBA084C" box="[123,217,1343,1367]" count="1" pageId="60" pageNumber="261" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843045FFACFEF0FAADBC5A084C" box="[261,313,1343,1367]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="60" pageNumber="261" type="Museum">CAS</collectionCode>
) labeled “CASENT 1051146”/ “
<collectingCountry id="FA80D7D13045FFACFD69FAADBE39084C" box="[668,858,1343,1367]" name="Madagascar" pageId="60" pageNumber="261">MADAGASCAR</collectingCountry>
<location id="8748C19A3045FFACFC97FAADBE8A084C" LSID="urn:lsid:plazi:treatment:0A3E26573042FFADFF8EFA33BF1E0823:8748C19A3045FFACFC97FAADBE8A084C" box="[866,1001,1343,1367]" country="Madagascar" county="Ranomafana" latitude="-21.2485" longLatPrecision="12" longitude="47.452168" municipality="Harin'Hala California Acad of Sciences" name="Ranomafana" pageId="60" pageNumber="261" stateProvince="Fianarantsoa">Ranomafana</location>
NP, Vatoharana trail S 600
<date id="F629B1813045FFACFF0DFAF2BC010863" box="[248,354,1376,1400]" pageId="60" pageNumber="261" value="2004-08-24">
<collectingDate id="E66D48693045FFACFF0DFAF2BC010863" box="[248,354,1376,1400]" pageId="60" pageNumber="261" value="2004-08-24">24.8.2004</collectingDate>
</date>
”/ “collected by hand in daytime,
<collectorName id="2F62F2973045FFACFD2CFAF2BE2F0863" box="[729,844,1376,1400]" pageId="60" pageNumber="261">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973045FFACFC81FAF2BEB60863" box="[884,981,1376,1400]" pageId="60" pageNumber="261">J. Rainio</collectorName>
collectors”. Both
<typeStatus id="5D2C29E33045FFACFF8EFA13BD820882" box="[123,225,1409,1433]" pageId="60" pageNumber="261" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="5D2C29E33045FFACFDC0FA13BFDA0882" box="[565,697,1409,1433]" pageId="60" pageNumber="261" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC23045FFACFD34FA13B9130883" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[705,1136,1408,1433]" class="Insecta" family="Carabidae" genus="Eunostus" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="261" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B0E34B533045FFACFD34FA13BEE70883" box="[705,900,1409,1432]" italics="true" pageId="60" pageNumber="261">Eunostus minimus</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283045FFACFB82FA13BDEC08A3" pageId="60" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<caption id="D6E8C7C93045FFACFF6EFB18BCBC09AC" pageId="60" pageNumber="261" startId="60.[155,167,1162,1181]" targetBox="[130,1176,216,1147]" targetPageId="60" targetType="figure">
<paragraph id="822897413045FFACFF6EFB18BCBC09AC" blockId="60.[123,1173,1162,1207]" pageId="60" pageNumber="261">
<smallCapsWord id="84CE019D3045FFACFF6EFB18BDBB0980" baselines="1176,1175" box="[155,216,1162,1181]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="60" pageNumber="261">FIGURE</smallCapsWord>
36. Digital images of holotype male of
<taxonomicName id="4597ECC23045FFACFDB4FB18BEC10986" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[577,930,1162,1181]" class="Insecta" family="Carabidae" genus="Eunostus" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="261" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B0E34B533045FFACFDB4FB18BFBC0987" box="[577,735,1162,1180]" italics="true" pageId="60" pageNumber="261">Eunostus minimus</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283045FFACFC5FFB18BE8D0986" box="[938,1006,1162,1181]" pageId="60" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels. Scale line A = 1.0 mm.
</paragraph>
</caption>
<paragraph id="822897413045FFACFF5EFA50B95008C2" blockId="60.[123,1174,1246,1725]" box="[171,1075,1473,1498]" pageId="60" pageNumber="261">
<materialsCitation id="32FF9D1C3045FFACFF5EFA50B94F08C2" box="[171,1068,1473,1498]" country="Madagascar" location="Ranomafana National Park" pageId="60" pageNumber="261" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533045FFACFF5EFA50BC3408C3" bold="true" box="[171,343,1474,1498]" pageId="60" pageNumber="261">
<smallCapsWord id="84CE019D3045FFACFF5EFA50BD8708C3" baselines="1492,1491" box="[171,228,1474,1498]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="60" pageNumber="261">
<typeStatus id="5D2C29E33045FFACFF5EFA50BD8708C3" box="[171,228,1474,1498]" pageId="60" pageNumber="261">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3045FFACFF19FA57BC3408C3" baselines="1491" box="[236,343,1477,1496]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="60" pageNumber="261">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13045FFACFE8BFA53BC9C08C2" box="[382,511,1473,1497]" name="Madagascar" pageId="60" pageNumber="261">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33045FFACFDF8FA53BF9F08C2" box="[525,764,1473,1497]" country="Madagascar" name="Fianarantsoa" pageId="60" pageNumber="261">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3045FFACFCFFFA53B94F08C2" LSID="urn:lsid:plazi:treatment:0A3E26573042FFADFF8EFA33BF1E0823:8748C19A3045FFACFCFFFA53B94F08C2" box="[778,1068,1473,1497]" country="Madagascar" name="Ranomafana National Park" pageId="60" pageNumber="261" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413045FFACFF5EFA70BFC20B20" blockId="60.[123,1174,1246,1725]" pageId="60" pageNumber="261">
<emphasis id="B0E34B533045FFACFF5EFA70BC8F08E2" bold="true" box="[171,492,1506,1530]" pageId="60" pageNumber="261">
<smallCapsWord id="84CE019D3045FFACFF5EFA70BC4D08E2" baselines="1524,1524" box="[171,302,1506,1530]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="60" pageNumber="261">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3045FFACFEC1FA74BC3308E2" baselines="1524" box="[308,336,1510,1529]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="60" pageNumber="261">OF</smallCapsWord>
<smallCapsWord id="84CE019D3045FFACFEA0FA74BCCB08E2" baselines="1524" box="[341,424,1510,1529]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="60" pageNumber="261">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3045FFACFE5AFA74BC8F08E2" baselines="1524" box="[431,492,1510,1529]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="60" pageNumber="261">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23045FFACFD18FA71BE2408E1" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[749,839,1507,1530]" class="Insecta" family="Carabidae" genus="Eunostus" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="261" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B0E34B533045FFACFD18FA71BE2408E1" box="[749,839,1507,1530]" italics="true" pageId="60" pageNumber="261">minimus</emphasis>
</taxonomicName>
, is the Latin adjective meaning smallest, a reference to fact that the known specimens of this species are smaller than those of any of the other known species that we have examined.
</paragraph>
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<emphasis id="B0E34B533045FFACFF5EF9D6BC210B41" bold="true" box="[171,322,1604,1628]" pageId="60" pageNumber="261">RECOGNITION</emphasis>
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.— Small for genus, SBL of male =
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, of female 5.0
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5.
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. Members of this species (
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) do not key out in Jeannel’s (1949) key for
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<emphasis id="B0E34B533045FFACFC9AF9F7BEB20B67" box="[879,977,1637,1660]" italics="true" pageId="60" pageNumber="261">Eunostus</emphasis>
species.
</taxonomicName>
They are much smaller and shinier than members of any other described species. Their heads are slightly narrower, with the tempora more evenly rounded to the back of the head, their antennal scapes shorter and thicker than in other species, thickest slightly distal to mid-length. Lateral margins of the pronotum are smoothly rounded in anterior half with a moderately short and deep subbasal sinuation anterior to slightly obtuse to rectangular hind angles. The elytral setae are generally sparser than in other species and arranged in 2 more or less distinct longitudinal rows on each interval. Male (
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) and female (
<figureCitation id="1AAC8BC43044FFADFE33FBB2BF450923" box="[454,550,1056,1080]" captionStart="FIGURE" captionStartId="61.[155,167,825,844]" captionTargetBox="[120,1176,216,803]" captionTargetId="figure-1453@61.[120,1176,216,803]" captionTargetPageId="61" captionText="FIGURE 37. Digital images of Eunostus minimus Kavanaugh &amp; Rainio sp. nov. A–B. Apical part of left hind femur, oblique medial aspect; A. Holotype male; B. Female; C–D. Aedeagus of holotype male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 0.5 mm." pageId="61" pageNumber="262">Fig. 37B</figureCitation>
) hind femora are sexual dimorphic, each unique among Malagasy species for each sex. Median lobe of male genitalia
<figureCitation id="1AAC8BC43044FFADFCEAFBD2BECB0943" box="[799,936,1088,1112]" captionStart="FIGURE" captionStartId="61.[155,167,825,844]" captionTargetBox="[120,1176,216,803]" captionTargetId="figure-1453@61.[120,1176,216,803]" captionTargetPageId="61" captionText="FIGURE 37. Digital images of Eunostus minimus Kavanaugh &amp; Rainio sp. nov. A–B. Apical part of left hind femur, oblique medial aspect; A. Holotype male; B. Female; C–D. Aedeagus of holotype male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 0.5 mm." pageId="61" pageNumber="262">Figs. 37C–D</figureCitation>
markedly different in shape from those of the other species for which males are known (see Jeannel 1949, Fig. 513d–e).
</paragraph>
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37. Digital images of
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<emphasis id="B0E34B533044FFADFE56FCABBF220E50" box="[419,577,825,843]" italics="true" pageId="61" pageNumber="262">Eunostus minimus</emphasis>
Kavanaugh &amp; Rainio
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A–B. Apical part of left hind femur, oblique medial aspect; A. Holotype male; B. Female; C–D. Aedeagus of holotype male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 0.5 mm.
</paragraph>
</caption>
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</emphasis>
.— At present, known only from the
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locality.
</paragraph>
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<emphasis id="B0E34B533044FFADFF5EFB32BCC609AC" bold="true" box="[171,421,1184,1208]" pageId="61" pageNumber="262">
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</emphasis>
.— Specimens of the
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series were collected in both primary montane rainforest in the Vatoharanana area (at elevations ranging from of
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) and in secondary montane rainforest
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north of Ranomafana village at an elevation of
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.
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The specimens were collected in a malaise trap and sifted from forest litter, respectively, at the primary forest site and by hand at the secondary forest site.
</paragraph>
</subSubSection>
</treatment>
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<emphasis id="B0E34B533043FFAAFF8EFF49BDDA0DE8" bold="true" box="[123,185,219,243]" italics="true" pageId="58" pageNumber="259">Lebia</emphasis>
(
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)
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Kavanaugh and Rainio
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,
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.—
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Fianaran. Pr.
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HQ. area
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, at night
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&amp;
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”/ “
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<taxonomicName id="4597ECC23043FFAAFE34FEE7B9680C96" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[449,1035,373,397]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="259" phylum="Arthropoda" rank="species" species="apicoviolacea" status="sp. nov." subGenus="Metalebia">
<emphasis id="B0E34B533043FFAAFE34FEE7BC9D0C96" box="[449,510,373,397]" italics="true" pageId="58" pageNumber="259">Lebia</emphasis>
(
<emphasis id="B0E34B533043FFAAFDFBFEE7BF190C96" box="[526,634,373,397]" italics="true" pageId="58" pageNumber="259">Metalebia</emphasis>
)
<emphasis id="B0E34B533043FFAAFD7FFEE7BE430C96" box="[650,800,373,397]" italics="true" pageId="58" pageNumber="259">apicoviolacea</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283043FFAAFBE6FEE4B92E0C96" box="[1043,1101,374,397]" pageId="58" pageNumber="259" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label]
</materialsCitation>
.
<materialsCitation id="32FF9D1C3043FFAAFEF1FE04BC730F16" collectingDate="1988-10-06" collectingDateMax="1988-10-21" collectingDateMin="1988-10-06" collectionCode="NMHN" collectorName="W. E. Steiner" country="Madagascar" elevation="1100" location="Ranomafana" pageId="58" pageNumber="259" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="5D2C29E33043FFAAFEF1FE04BC0F0CB5" box="[260,364,406,430]" pageId="58" pageNumber="259" type="paratype">Paratypes</typeStatus>
(only 1): a female (in
<collectionCode id="E4860F843043FFAAFD70FE04BFB50CB6" box="[645,726,406,429]" pageId="58" pageNumber="259">NMHN</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13043FFAAFCA6FE07B9720CB6" box="[851,1041,405,429]" name="Madagascar" pageId="58" pageNumber="259">MADAGASCAR</collectingCountry>
: Prov.
<collectingRegion id="405359A33043FFAAFB9FFE07BD820CD6" country="Madagascar" name="Fianarantsoa" pageId="58" pageNumber="259">Fianarantsoa</collectingRegion>
,
<quantity id="456F3AA43043FFAAFF18FE24BC410CD6" box="[237,290,437,461]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="58" pageNumber="259" unit="km" value="7.0">7 km</quantity>
W
<location id="8748C19A3043FFAAFEB3FE27BCAE0CD6" LSID="urn:lsid:plazi:treatment:0A3E26573043FFABFF8EFF49BE9D098D:8748C19A3043FFAAFEB3FE27BCAE0CD6" box="[326,461,437,461]" country="Madagascar" name="Ranomafana" pageId="58" pageNumber="259" stateProvince="Fianarantsoa">Ranomafana</location>
,
<quantity id="456F3AA43043FFAAFE2FFE24BF4A0CD6" box="[474,553,438,462]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="58" pageNumber="259" unit="m" value="1100.0">
<elevation id="09BA70723043FFAAFE2FFE24BF4A0CD6" box="[474,553,438,462]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="58" pageNumber="259" unit="m" value="1100.0">1100 m</elevation>
</quantity>
<date id="F629B1813043FFAAFDDBFE27BF950CD5" box="[558,758,437,462]" pageId="58" pageNumber="259" value="1988-10-06" valueMax="1988-10-21" valueMin="1988-10-06">
<collectingDate id="E66D48693043FFAAFDDBFE27BF950CD5" box="[558,758,437,462]" pageId="58" pageNumber="259" value="1988-10-06" valueMax="1988-10-21" valueMin="1988-10-06">6-21 October 1988</collectingDate>
</date>
<collectorName id="2F62F2973043FFAAFD0EFE24BEE40CD6" box="[763,903,437,461]" pageId="58" pageNumber="259">W. E. Steiner</collectorName>
”/ “At black light in montane rain forest”/ “
<typeStatus id="5D2C29E33043FFAAFECAFE44BCA00CF5" box="[319,451,470,494]" pageId="58" pageNumber="259" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC23043FFAAFE3FFE44B96C0CF5" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[458,1039,470,494]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="259" phylum="Arthropoda" rank="species" species="apicoviolacea" status="sp. nov." subGenus="Metalebia">
<emphasis id="B0E34B533043FFAAFE3FFE44BF640CF5" box="[458,519,470,494]" italics="true" pageId="58" pageNumber="259">Lebia</emphasis>
(
<emphasis id="B0E34B533043FFAAFDE3FE44BFE10CF5" box="[534,642,470,494]" italics="true" pageId="58" pageNumber="259">Metalebia</emphasis>
)
<emphasis id="B0E34B533043FFAAFD64FE44BE440CF5" box="[657,807,470,494]" italics="true" pageId="58" pageNumber="259">apicoviolacea</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283043FFAAFBE0FE44B92D0CF6" box="[1045,1102,470,493]" pageId="58" pageNumber="259" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="822897413043FFAAFF5EFD84B9500F35" blockId="58.[123,1174,308,623]" box="[171,1075,534,558]" pageId="58" pageNumber="259">
<materialsCitation id="32FF9D1C3043FFAAFF5EFD84B94F0F35" box="[171,1068,534,558]" country="Madagascar" location="Ranomafana National Park" pageId="58" pageNumber="259" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533043FFAAFF5EFD84BC340F36" bold="true" box="[171,343,534,558]" pageId="58" pageNumber="259">
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<typeStatus id="5D2C29E33043FFAAFF5EFD84BD870F36" box="[171,228,534,558]" pageId="58" pageNumber="259">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3043FFAAFF19FD88BC340F36" baselines="552" box="[236,343,538,557]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="58" pageNumber="259">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13043FFAAFE8BFD84BC9C0F35" box="[382,511,534,558]" name="Madagascar" pageId="58" pageNumber="259">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33043FFAAFDF8FD84BF9F0F35" box="[525,764,534,558]" country="Madagascar" name="Fianarantsoa" pageId="58" pageNumber="259">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3043FFAAFCFFFD84B94F0F35" LSID="urn:lsid:plazi:treatment:0A3E26573043FFABFF8EFF49BE9D098D:8748C19A3043FFAAFCFFFD84B94F0F35" box="[778,1068,534,558]" country="Madagascar" name="Ranomafana National Park" pageId="58" pageNumber="259" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413043FFABFF5EFDA5B9790C08" blockId="58.[123,1174,308,623]" lastBlockId="59.[123,1174,219,1174]" lastPageId="59" lastPageNumber="260" pageId="58" pageNumber="259">
<emphasis id="B0E34B533043FFAAFF5EFDA5BC900F56" bold="true" box="[171,499,567,591]" pageId="58" pageNumber="259">
<smallCapsWord id="84CE019D3043FFAAFF5EFDA5BC4D0F56" baselines="585,584" box="[171,302,567,591]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="58" pageNumber="259">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3043FFAAFEC3FDA8BC310F56" baselines="584" box="[310,338,570,589]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="58" pageNumber="259">OF</smallCapsWord>
<smallCapsWord id="84CE019D3043FFAAFEAFFDA8BCCE0F56" baselines="584" box="[346,429,570,589]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="58" pageNumber="259">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3043FFAAFE43FDA8BC900F56" baselines="584" box="[438,499,570,589]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="58" pageNumber="259">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<emphasis id="B0E34B533043FFAAFD0EFDA4BEFB0F55" box="[763,920,566,590]" italics="true" pageId="58" pageNumber="259">
<taxonomicName id="4597ECC23043FFAAFD0EFDA4BEF20F55" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[763,913,566,590]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="259" phylum="Arthropoda" rank="species" species="apicoviolacea">apicoviolacea</taxonomicName>
,
</emphasis>
is an adjective derived from the Latin words,
<emphasis id="B0E34B533043FFAAFE9BFDC5BCC30F75" box="[366,416,599,622]" italics="true" pageId="58" pageNumber="259">apex</emphasis>
, meaning tip, and
<emphasis id="B0E34B533043FFAAFD99FDC5BFB20F74" box="[620,721,599,623]" italics="true" pageId="58" pageNumber="259">violaceus</emphasis>
, meaning violet-colored. The name refers to the violet or purple metallic reflection of elytral apices, which contrasts distinctly with the green or bronze metallic reflection of the remainder of the elytra in members of this species.
</paragraph>
<caption id="D6E8C7C93043FFAAFF6EF9E4BC4E0BA7" pageId="58" pageNumber="259" startId="58.[155,167,1654,1673]" targetBox="[123,1173,636,1650]" targetPageId="58" targetType="figure">
<paragraph id="822897413043FFAAFF6EF9E4BC4E0BA7" blockId="58.[123,1174,1653,1724]" pageId="58" pageNumber="259">
<smallCapsWord id="84CE019D3043FFAAFF6EF9E4BDBB0B9C" baselines="1668,1667" box="[155,216,1654,1673]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="58" pageNumber="259">FIGURE</smallCapsWord>
35. Digital images of holotype male of
<taxonomicName id="4597ECC23043FFAAFDD8F9E7B9630B93" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[557,1024,1653,1672]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="259" phylum="Arthropoda" rank="species" species="apicoviolacea" status="sp. nov." subGenus="Metalebia">
<emphasis id="B0E34B533043FFAAFDD8F9E7BF3D0B93" box="[557,606,1653,1672]" italics="true" pageId="58" pageNumber="259">Lebia</emphasis>
(
<emphasis id="B0E34B533043FFAAFD9FF9E7BFA30B93" box="[618,704,1653,1672]" italics="true" pageId="58" pageNumber="259">Metalebia</emphasis>
)
<emphasis id="B0E34B533043FFAAFD38F9E7BE260B93" box="[717,837,1653,1672]" italics="true" pageId="58" pageNumber="259">apicoviolacea</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283043FFAAFBF0F9E4B9250B92" box="[1029,1094,1654,1673]" pageId="58" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm.
</paragraph>
</caption>
<paragraph id="822897413042FFABFF5EFE89B909090E" blockId="59.[123,1174,219,1174]" pageId="59" pageNumber="260">
<smallCapsWord id="84CE019D3042FFABFF5EFE89BC210C29" baselines="301,301" box="[171,322,283,307]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="59" pageNumber="260">
<emphasis id="B0E34B533042FFABFF5EFE89BC210C29" bold="true" box="[171,322,283,307]" pageId="59" pageNumber="260">RECOGNITION</emphasis>
</smallCapsWord>
.— Size moderately small for genus, SBL of male =
<quantity id="456F3AA43042FFABFC7FFE89BEBD0C28" box="[906,990,283,307]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="59" pageNumber="260" unit="mm" value="3.8">3.8 mm</quantity>
, of female = 4.0 mm. Members of this species (
<figureCitation id="1AAC8BC43042FFABFE31FEA9BF400C4F" box="[452,547,315,340]" captionStart="FIGURE" captionStartId="58.[155,167,1654,1673]" captionTargetBox="[123,1173,636,1650]" captionTargetId="figure-809@58.[123,1173,636,1650]" captionTargetPageId="58" captionText="FIGURE 35. Digital images of holotype male of Lebia (Metalebia) apicoviolacea Kavanaugh &amp; Rainio sp. nov. A. Habi- tus, dorsal aspect; B. Labels; C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." pageId="59" pageNumber="260">Fig. 35A</figureCitation>
) key to subgenus
<taxonomicName id="4597ECC23042FFABFD10FEA9BE950C4F" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" box="[741,1014,315,340]" pageId="59" pageNumber="260" rank="subGenus" subGenus="Metalebia">
<emphasis id="B0E34B533042FFABFD10FEA9BE320C48" box="[741,849,315,339]" italics="true" pageId="59" pageNumber="260">Metalebia</emphasis>
Jeannel (1949)
</taxonomicName>
and the
<taxonomicName id="4597ECC23042FFABFBA4FEA9BC630C68" authorityName="Chaudoir" authorityYear="1850" class="Insecta" family="Carabidae" genus="Assadecma" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B0E34B533042FFABFBA4FEA9BC630C68" italics="true" pageId="59" pageNumber="260">madagascariensis</emphasis>
</taxonomicName>
group in Jeannel’s (1949) key: the basal elytral border is absent medially between the base of stria 3 and the scutellum; the elytral striae are deep and the elytral intervals convex and impunctate; the forebody has no metallic reflection and the elytra are uniform in color (but not necessarily in reflection); and the frons is slightly convex and without distinct frontal furrows between the eyes. Within the
<taxonomicName id="4597ECC23042FFABFEA6FE4EBF770CEF" authorityName="Chaudoir" authorityYear="1850" box="[339,532,476,500]" class="Insecta" family="Carabidae" genus="Assadecma" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B0E34B533042FFABFEA6FE4EBF770CEF" box="[339,532,476,500]" italics="true" pageId="59" pageNumber="260">madagascariensis</emphasis>
</taxonomicName>
group,
<taxonomicName id="4597ECC23042FFABFD91FE4FBE750CEF" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[612,790,476,500]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="apicoviolacea">
<emphasis id="B0E34B533042FFABFD91FE4FBE750CEF" box="[612,790,476,500]" italics="true" pageId="59" pageNumber="260">L. apicoviolacea</emphasis>
</taxonomicName>
members can be distinguished from those of all species except
<taxonomicName id="4597ECC23042FFABFE6DFE6FBF4D0F0F" authorityName="Jeannel" authorityYear="1949" box="[408,558,508,532]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="alluaudana">
<emphasis id="B0E34B533042FFABFE6DFE6FBF4D0F0F" box="[408,558,508,532]" italics="true" pageId="59" pageNumber="260">L. alluaudana</emphasis>
</taxonomicName>
,
<taxonomicName id="4597ECC23042FFABFDC9FE6FBE630F0F" authority="Alluaud" authorityName="Alluaud" authorityYear="1936" box="[572,768,508,532]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="mirana">
<emphasis id="B0E34B533042FFABFDC9FE6FBFC60F0F" box="[572,677,509,532]" italics="true" pageId="59" pageNumber="260">L. mirana</emphasis>
Alluaud
</taxonomicName>
,
<taxonomicName id="4597ECC23042FFABFCFBFE6FBEDB0F0F" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[782,952,508,532]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533042FFABFCFBFE6FBEDB0F0F" box="[782,952,508,532]" italics="true" pageId="59" pageNumber="260">L. ranomafanae</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC23042FFABFC19FE6FB92F0F0F" authorityName="Jeannel" authorityYear="1949" box="[1004,1100,508,532]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="tanala">
<emphasis id="B0E34B533042FFABFC19FE6FB92F0F0F" box="[1004,1100,508,532]" italics="true" pageId="59" pageNumber="260">L. tanala</emphasis>
</taxonomicName>
by the presence of elytral metallic reflection (metallic reflection absent from the elytra of members of the remaining seven species). They are distinguished from members of
<taxonomicName id="4597ECC23042FFABFC9AFDAFBEBF0F4F" authorityName="Alluaud" authorityYear="1936" box="[879,988,573,596]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="mirana">
<emphasis id="B0E34B533042FFABFC9AFDAFBEBF0F4F" box="[879,988,573,596]" italics="true" pageId="59" pageNumber="260">L. mirana</emphasis>
</taxonomicName>
by their reddish labrum (black in
<taxonomicName id="4597ECC23042FFABFEC6FDCFBCFF0F6F" authorityName="Alluaud" authorityYear="1936" box="[307,412,605,628]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="mirana">
<emphasis id="B0E34B533042FFABFEC6FDCFBCFF0F6F" box="[307,412,605,628]" italics="true" pageId="59" pageNumber="260">L. mirana</emphasis>
</taxonomicName>
members) and from those of
<taxonomicName id="4597ECC23042FFABFD2FFDCFBE130F6F" authorityName="Jeannel" authorityYear="1949" box="[730,880,604,628]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="alluaudana">
<emphasis id="B0E34B533042FFABFD2FFDCFBE130F6F" box="[730,880,604,628]" italics="true" pageId="59" pageNumber="260">L. alluaudana</emphasis>
</taxonomicName>
by their small size and relatively short and broad elytra, with ratio of elytra length to width = 1.2–1.3 (size larger, SBL of males = 4.6–5.0 mm, of females 4.6–
<quantity id="456F3AA43042FFABFDEFFD0EBF130FAF" box="[538,624,668,693]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="59" pageNumber="260" unit="mm" value="5.3">5.3 mm</quantity>
, and elytra relatively longer, with ratio of elytra length to width = 1.4, in
<taxonomicName id="4597ECC23042FFABFE69FD2CBF5B0FCE" authorityName="Jeannel" authorityYear="1949" box="[412,568,701,725]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="alluaudana">
<emphasis id="B0E34B533042FFABFE69FD2CBF5B0FCE" box="[412,568,701,725]" italics="true" pageId="59" pageNumber="260">L. alluaudana</emphasis>
</taxonomicName>
members). They are distinguished from members of
<taxonomicName id="4597ECC23042FFABFF8EFD4CBDBE0FEE" authorityName="Jeannel" authorityYear="1949" box="[123,221,733,757]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="tanala">
<emphasis id="B0E34B533042FFABFF8EFD4CBDBE0FEE" box="[123,221,733,757]" italics="true" pageId="59" pageNumber="260">L. tanala</emphasis>
</taxonomicName>
in having the pronotum with lateral margins slightly sinuate anterior to slightly obtuse hind angles (lateral margins distinctly sinuate anterior to sharp, rectangular hind angles slightly projected laterally in members of
<taxonomicName id="4597ECC23042FFABFE08FC8CBF010E2E" authorityName="Jeannel" authorityYear="1949" box="[509,610,797,821]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="tanala">
<emphasis id="B0E34B533042FFABFE08FC8CBF010E2E" box="[509,610,797,821]" italics="true" pageId="59" pageNumber="260">L. tanala</emphasis>
</taxonomicName>
) and from members of both
<taxonomicName id="4597ECC23042FFABFC41FC8CB9000E2E" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[948,1123,797,821]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533042FFABFC41FC8CB9000E2E" box="[948,1123,797,821]" italics="true" pageId="59" pageNumber="260">L. ranomafanae</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC23042FFABFF8EFCACBDBF0E4E" authorityName="Jeannel" authorityYear="1949" box="[123,220,829,853]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="260" phylum="Arthropoda" rank="species" species="tanala">
<emphasis id="B0E34B533042FFABFF8EFCACBDBF0E4E" box="[123,220,829,853]" italics="true" pageId="59" pageNumber="260">L. tanala</emphasis>
</taxonomicName>
in having elytra with distinct tricolored metallic reflection, green basally and on medial two-thirds, brassy on lateral one-third and violet in apical one-fifth (elytral metallic reflection uniformly faint or distinct metallic green or blue in members of
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<emphasis id="B0E34B533042FFABFCF4FCECBEC80E8D" box="[769,939,894,918]" italics="true" pageId="59" pageNumber="260">L. ranomafanae</emphasis>
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and
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<emphasis id="B0E34B533042FFABFC2BFCECB95D0E8E" box="[990,1086,893,917]" italics="true" pageId="59" pageNumber="260">L. tanala</emphasis>
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). Shape of the median lobe of the male aedeagus (
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) is also diagnostic, with the shaft very slightly arcuate, dorsal and ventral margins more or less parallel in basal two-thirds in lateral view (
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) and apex moderate in length and relatively broader in both lateral and dorsal (
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) views than in males of other species of this species group (compare with Jeannel 1949, Fig. 440a-l).
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<smallCapsWord id="84CE019D3042FFABFEA8FBB3BC92092F" baselines="1071" box="[349,497,1057,1076]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="59" pageNumber="260">DISTRIBUTION</smallCapsWord>
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.— At present, known only from the
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locality.
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.— Both specimens of the
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series were collected in secondary montane rainforest in the Talatakely area at elevations ranging from
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. Both were collect- ed at night, one by hand from vegetation, and the other attracted to ultraviolet light.
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This genus is endemic to
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(
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<smallCapsWord id="84CE019D3044FFADFF1EF9ECBC3B0B8A" baselines="1676" box="[235,344,1662,1681]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="61" pageNumber="262">MATERIAL</smallCapsWord>
</emphasis>
.—
<typeStatus id="5D2C29E33044FFADFE75F9E8BC800B89" box="[384,483,1658,1682]" pageId="61" pageNumber="262" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43044FFADFE06F9E8BF170B89" box="[499,628,1658,1683]" captionStart="FIGURE" captionStartId="62.[155,167,1459,1478]" captionTargetBox="[152,1176,222,1433]" captionTargetId="figure-507@62.[120,1176,216,1456]" captionTargetPageId="62" captionText="FIGURE 38. Digital images of holotype female of Erephognathus ranomafanae Kavanaugh and Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." pageId="61" pageNumber="262">Figs.38A–B</figureCitation>
), a female, in CAS, labeled: “CASENT 8068531”/
</paragraph>
<paragraph id="822897413044FFAFFF8EF909BE130C28" blockId="61.[123,1173,1658,1715]" lastBlockId="63.[123,1174,219,883]" lastPageId="63" lastPageNumber="264" pageId="61" pageNumber="262">
“
<materialsCitation id="32FF9D1C3044FFAFFF72F909BE0A0C28" collectingDate="2000-12-05" collectingDateMax="2001-11-15" collectingDateMin="2000-12-05" collectionCode="CAS" collectorName="R. Harin'Hala California Acad &amp; Fisher &amp; Both" country="Madagascar" county="Province" elevation="1110" lastPageId="63" lastPageNumber="264" latitude="-21.226166" location="Vohiparara" longLatPrecision="12" longitude="47.369835" municipality="Parc National Ranomafana" pageId="61" pageNumber="262" specimenCount="2" specimenCount-female="2" stateProvince="Fianarantsoa" typeStatus="paratype">
<collectingCountry id="FA80D7D13044FFADFF72F909BC260BA8" box="[135,325,1691,1715]" name="Madagascar" pageId="61" pageNumber="262">MADAGASCAR</collectingCountry>
:
<collectingCounty id="6B49EFCD3044FFADFEACF908BCDB0BA9" box="[345,440,1690,1714]" pageId="61" pageNumber="262">Province</collectingCounty>
<collectingRegion id="405359A33044FFADFE30F908BF2E0BA9" box="[453,589,1690,1714]" country="Madagascar" name="Fianarantsoa" pageId="61" pageNumber="262">Fianarantsoa</collectingRegion>
,
<collectingMunicipality id="624C0D3B3044FFADFD97F909BEEC0BA9" box="[610,911,1690,1714]" pageId="61" pageNumber="262">Parc National Ranomafana</collectingMunicipality>
,
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, at broken bridge, el
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</quantity>
<date id="F629B1813047FFAEFEC8F996BF420B07" box="[317,545,1540,1564]" pageId="62" pageNumber="263" value="2001-11-08" valueMax="2001-11-15" valueMin="2001-11-08">
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</date>
”/ “
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,
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collector:
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R.
<location id="8748C19A3047FFAEFC1DF996BC630B27" LSID="urn:lsid:plazi:treatment:0A3E26573044FFAFFF8EF9B3BE7F0E68:8748C19A3047FFAEFC1DF996BC630B27" country="Madagascar" county="Province" latitude="-21.226166" longLatPrecision="12" longitude="47.369835" municipality="Parc National Ranomafana" name="Harin'Hala California Acad of Sciences" pageId="62" pageNumber="263" stateProvince="Fianarantsoa">Harin’Hala California Acad</location>
</collectorName>
of Sciences malaise trap in high altitude rainforest, MA-02-09A-02”/ “
<typeStatus id="5D2C29E33047FFAEFBFCF9B7B9F50B26" box="[1033,1174,1573,1597]" pageId="62" pageNumber="263" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23047FFAEFF8EF9D7BFFE0B46" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,669,1605,1629]" class="Insecta" family="Carabidae" genus="Erephognathus" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="263" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533047FFAEFF8EF9D7BCD10B46" box="[123,434,1605,1629]" italics="true" pageId="62" pageNumber="263">Erephognathus ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283047FFAEFD50F9D7BFBC0B47" box="[677,735,1605,1628]" pageId="62" pageNumber="263" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
<typeStatus id="5D2C29E33047FFAEFC5EF9D7B9700B46" box="[939,1043,1605,1629]" pageId="62" pageNumber="263" type="paratype">Paratypes</typeStatus>
(total of 2):
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(in
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) labeled same as
<typeStatus id="5D2C29E33047FFAEFDFBF9F7BF080B66" box="[526,619,1637,1661]" pageId="62" pageNumber="263" type="holotype">holotype</typeStatus>
, except “CASENT 8068532”;
<specimenCount id="94915CC83047FFAEFC33F9F7B9450B66" box="[966,1062,1637,1661]" count="1" pageId="62" pageNumber="263" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843047FFAEFBACF9F7B9EE0B66" box="[1113,1165,1637,1661]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="62" pageNumber="263" type="Museum">CAS</collectionCode>
) labeled “CASENT 8068672”/; “
<collectingCountry id="FA80D7D13047FFAEFE20F917BFF00B86" box="[469,659,1669,1693]" name="Madagascar" pageId="62" pageNumber="263">MADAGASCAR</collectingCountry>
:
<collectingRegion id="405359A33047FFAEFD54F917BECD0B86" box="[673,942,1669,1693]" country="Madagascar" name="Antananarivo" pageId="62" pageNumber="263">Province d’Antananarivo</collectingRegion>
<locationDeviation id="939A8E633047FFAEFC43F917BDA10BA5" location="Andranomay" pageId="62" pageNumber="263">
<quantity id="456F3AA43047FFAEFC43F917BE8F0B86" box="[950,1004,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="62" pageNumber="263" unit="km" value="3.0">3 km</quantity>
41° NE Andranomay
</locationDeviation>
,
<locationDeviation id="939A8E633047FFAEFF2DF934BF210BA6" box="[216,578,1701,1726]" location="Anjozorobe" pageId="62" pageNumber="263">
<quantity id="456F3AA43047FFAEFF2DF934BC540BA6" box="[216,311,1701,1726]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="62" pageNumber="263" unit="km" value="11.5">11.5 km</quantity>
147° SSE Anjozorobe
</locationDeviation>
elev
<quantity id="456F3AA43047FFAEFD78F934BF840BA6" box="[653,743,1702,1726]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="62" pageNumber="263" unit="m" value="1300.0">
<elevation id="09BA70723047FFAEFD78F934BF840BA6" box="[653,743,1702,1726]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="62" pageNumber="263" unit="m" value="1300.0">1300 m</elevation>
</quantity>
<date id="F629B1813047FFAEFD03F937BE8B0BA5" box="[758,1000,1701,1726]" pageId="62" pageNumber="263" value="2000-12-05" valueMax="2000-12-13" valueMin="2000-12-05">
<collectingDate id="E66D48693047FFAEFD03F937BE8B0BA5" box="[758,1000,1701,1726]" pageId="62" pageNumber="263" value="2000-12-05" valueMax="2000-12-13" valueMin="2000-12-05">5-13 December 2000</collectingDate>
</date>
”/ “
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<geoCoordinate id="E7A3F1863046FFAFFF8EFF49BD990DE9" box="[123,250,219,243]" degrees="47" direction="east" minutes="57" orientation="longitude" pageId="63" pageNumber="264" precision="15" seconds="38" value="47.960556">47°57’38”E</geoCoordinate>
coll.
<collectorName id="2F62F2973046FFAFFECDFF49BC190DE8" box="[312,378,219,243]" pageId="63" pageNumber="264">Fisher</collectorName>
, Griswold et al.
<location id="8748C19A3046FFAFFDC5FF49BE070DE8" LSID="urn:lsid:plazi:treatment:0A3E26573044FFAFFF8EF9B3BE7F0E68:8748C19A3046FFAFFDC5FF49BE070DE8" box="[560,868,219,243]" country="Madagascar" county="Province" latitude="-21.226166" longLatPrecision="12" longitude="47.369835" municipality="Parc National Ranomafana" name="California Acad. of Sciences" pageId="63" pageNumber="264" stateProvince="Fianarantsoa">California Acad. of Sciences</location>
montane rainforest,
<collectingMethod id="5BD6EF563046FFAFFBB1FF49BDC70C08" pageId="63" pageNumber="264">malaise trap</collectingMethod>
, coll. code: BLF2375”.
<collectorName id="2F62F2973046FFAFFE59FF69BC830C08" box="[428,480,251,275]" pageId="63" pageNumber="264">Both</collectorName>
<typeStatus id="5D2C29E33046FFAFFE12FF6EBF2E0C0F" box="[487,589,252,276]" pageId="63" pageNumber="264" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="5D2C29E33046FFAFFC50FF69B94A0C08" box="[933,1065,251,275]" pageId="63" pageNumber="264" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC23046FFAFFBC7FF69BF200C28" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Erephognathus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="264" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533046FFAFFBC7FF69BC340C28" italics="true" pageId="63" pageNumber="264">Erephognathus ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283046FFAFFDBFFE8EBFE70C28" box="[586,644,284,307]" pageId="63" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<caption id="D6E8C7C93047FFAEFF6EFA21BF0E08FB" pageId="62" pageNumber="263" startId="62.[155,167,1459,1478]" targetBox="[152,1176,222,1433]" targetPageId="62" targetType="figure">
<paragraph id="822897413047FFAEFF6EFA21B9F608DD" blockId="62.[123,1173,1459,1504]" box="[155,1173,1459,1478]" pageId="62" pageNumber="263">
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38. Digital images of holotype female of
<taxonomicName id="4597ECC23047FFAEFD9EFA21B92008DD" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[619,1091,1459,1478]" class="Insecta" family="Carabidae" genus="Erephognathus" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="263" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533047FFAEFD9EFA21BE0A08DD" box="[619,873,1459,1478]" italics="true" pageId="62" pageNumber="263">Erephognathus ranomafanae</emphasis>
Kavanaugh and Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283047FFAEFBBBFA21B9F608DD" box="[1102,1173,1459,1478]" pageId="62" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="822897413047FFAEFF8EFA5FBF0E08FB" blockId="62.[123,1173,1459,1504]" box="[123,621,1484,1504]" pageId="62" pageNumber="263">A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm.</paragraph>
</caption>
<paragraph id="822897413046FFAFFF5EFEA9B9500C48" blockId="63.[123,1174,219,883]" box="[171,1075,315,339]" pageId="63" pageNumber="264">
<materialsCitation id="32FF9D1C3046FFAFFF5EFEA9B94F0C48" box="[171,1068,315,339]" country="Madagascar" location="Ranomafana National Park" pageId="63" pageNumber="264" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533046FFAFFF5EFEA9BC340C4A" bold="true" box="[171,343,315,339]" pageId="63" pageNumber="264">
<smallCapsWord id="84CE019D3046FFAFFF5EFEA9BD870C49" baselines="333,333" box="[171,228,315,339]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="63" pageNumber="264">
<typeStatus id="5D2C29E33046FFAFFF5EFEA9BD870C49" box="[171,228,315,339]" pageId="63" pageNumber="264">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3046FFAFFF19FEACBC340C4A" baselines="332" box="[236,343,318,337]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="63" pageNumber="264">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13046FFAFFE8BFEA9BC9C0C48" box="[382,511,315,339]" name="Madagascar" pageId="63" pageNumber="264">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33046FFAFFDF8FEA9BF9F0C48" box="[525,764,315,339]" country="Madagascar" name="Fianarantsoa" pageId="63" pageNumber="264">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3046FFAFFCFFFEA9B94F0C48" LSID="urn:lsid:plazi:treatment:0A3E26573044FFAFFF8EF9B3BE7F0E68:8748C19A3046FFAFFCFFFEA9B94F0C48" box="[778,1068,315,339]" country="Madagascar" name="Ranomafana National Park" pageId="63" pageNumber="264" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413046FFAFFF5EFEC9BECA0C88" blockId="63.[123,1174,219,883]" pageId="63" pageNumber="264">
<emphasis id="B0E34B533046FFAFFF5EFEC9BC910C69" bold="true" box="[171,498,347,371]" pageId="63" pageNumber="264">
<smallCapsWord id="84CE019D3046FFAFFF5EFEC9BC4D0C6A" baselines="365,364" box="[171,302,347,371]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="63" pageNumber="264">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3046FFAFFEC3FECCBC310C6A" baselines="364" box="[310,338,350,369]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="63" pageNumber="264">OF</smallCapsWord>
<smallCapsWord id="84CE019D3046FFAFFEACFECCBCCF0C6A" baselines="364" box="[345,428,350,369]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="63" pageNumber="264">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3046FFAFFE40FECDBC910C69" baselines="365" box="[437,498,351,370]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="63" pageNumber="264">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23046FFAFFD0CFEC9BEE40C68" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[761,903,347,371]" class="Insecta" family="Carabidae" genus="Erephognathus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="264" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533046FFAFFD0CFEC9BEE40C68" box="[761,903,347,371]" italics="true" pageId="63" pageNumber="264">ranomafanae</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the national park in which the
<typeStatus id="5D2C29E33046FFAFFD29FEEEBE690C8F" box="[732,778,380,404]" pageId="63" pageNumber="264">type</typeStatus>
was collected.
</paragraph>
<paragraph id="822897413046FFAFFF5EFE09BF4C0F68" blockId="63.[123,1174,219,883]" pageId="63" pageNumber="264">
<smallCapsWord id="84CE019D3046FFAFFF5EFE09BC210CAA" baselines="429,428" box="[171,322,411,435]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="63" pageNumber="264">
<emphasis id="B0E34B533046FFAFFF5EFE09BC210CAA" bold="true" box="[171,322,411,435]" pageId="63" pageNumber="264">RECOGNITION</emphasis>
</smallCapsWord>
.— Size moderate for genus, SBL of females = 7.5
<emphasis id="B0E34B533046FFAFFCAAFE09BE0B0CA8" bold="true" box="[863,872,411,435]" pageId="63" pageNumber="264">-</emphasis>
8.
<quantity id="456F3AA43046FFAFFC74FE09BEDD0CA8" box="[897,958,411,435]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="63" pageNumber="264" unit="mm" value="9.0">9 mm</quantity>
. In Jeannel’s (1949) key to species of
<taxonomicName id="4597ECC23046FFAFFECFFE29BCBF0CC8" authorityName="Alluaud" authorityYear="1932" box="[314,476,443,467]" class="Insecta" family="Carabidae" genus="Erephognathus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533046FFAFFECFFE29BCBF0CC8" box="[314,476,443,467]" italics="true" pageId="63" pageNumber="264">Erephognathus</emphasis>
</taxonomicName>
, members of
<taxonomicName id="4597ECC23046FFAFFD86FE2EBE430CC8" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[627,800,443,467]" class="Insecta" family="Carabidae" genus="Erephognathus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="264" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533046FFAFFD86FE2EBE430CC8" box="[627,800,443,467]" italics="true" pageId="63" pageNumber="264">E. ranomafanae</emphasis>
</taxonomicName>
key to
<taxonomicName id="4597ECC23046FFAFFC81FE2EBDDA0CE8" authority="(Fairmaire 1903)" baseAuthorityName="Fairmaire" baseAuthorityYear="1903" class="Insecta" family="Carabidae" genus="Erephognathus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="264" phylum="Arthropoda" rank="species" species="coerulescens">
<emphasis id="B0E34B533046FFAFFC81FE2EB97E0CC8" box="[884,1053,443,467]" italics="true" pageId="63" pageNumber="264">E. coerulescens</emphasis>
(
<bibRefCitation id="E606EAB03046FFAFFBDBFE29BDD30CE8" author="FAIRMAIRE, L." pageId="63" pageNumber="264" pagination="181 - 259" refId="ref30378" refString="FAIRMAIRE, L. 1903. Materiaux pour la faune coleopterique de la region Malagache (16 e Note). Annales de la Societe Entomologique de France 72: 181 - 259." type="journal article" year="1903">Fairmaire 1903</bibRefCitation>
)
</taxonomicName>
based on their short antennae (extended posteriorly only to the basal one-fourth of the elytra or less) and the absence of the tuft of setae from the apex of antennomere 11 (terminal antennomere) found in
<taxonomicName id="4597ECC23046FFAFFEC9FD8EBFFB0F28" authority="Alluaud (1936 a)" authorityName="Alluaud" authorityYear="1936" box="[316,664,539,563]" class="Insecta" family="Carabidae" genus="Erephognathus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="264" phylum="Arthropoda" rank="species" species="margarithrix">
<emphasis id="B0E34B533046FFAFFEC9FD8EBC800F28" box="[316,483,539,563]" italics="true" pageId="63" pageNumber="264">E. margarithrix</emphasis>
<bibRefCitation id="E606EAB03046FFAFFE1DFD89BFFB0F28" author="ALLUAUD, C." box="[488,664,539,563]" pageId="63" pageNumber="264" pagination="1 - 13" refId="ref28980" refString="ALLUAUD, C. 1936 a. Carabidae recueillis a Madagascar par MM. Syrig, G. Olsoufieff, Vadon, R. Catala, etc. Afra, Cahiers d'Entomologie 11: 1 - 13." type="journal article" year="1936">Alluaud (1936a)</bibRefCitation>
</taxonomicName>
members. The pronotum (
<figureCitation id="1AAC8BC43046FFAFFC43FD89B9760F28" box="[950,1045,539,563]" captionStart="FIGURE" captionStartId="62.[155,167,1459,1478]" captionTargetBox="[152,1176,222,1433]" captionTargetId="figure-507@62.[120,1176,216,1456]" captionTargetPageId="62" captionText="FIGURE 38. Digital images of holotype female of Erephognathus ranomafanae Kavanaugh and Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." pageId="63" pageNumber="264">Fig. 38A</figureCitation>
) is distinctly longer, narrower and less convex and with anterior angles more projected, narrowed and angulate than in members of
<taxonomicName id="4597ECC23046FFAFFE75FDCEBF4A0F68" baseAuthorityName="Fairmaire" baseAuthorityYear="1903" box="[384,553,603,627]" class="Insecta" family="Carabidae" genus="Erephognathus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="264" phylum="Arthropoda" rank="species" species="coerulescens">
<emphasis id="B0E34B533046FFAFFE75FDCEBF4A0F68" box="[384,553,603,627]" italics="true" pageId="63" pageNumber="264">E. coerulescens</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="822897413046FFAFFF5EFDE9B92F0E28" blockId="63.[123,1174,219,883]" pageId="63" pageNumber="264">
<emphasis id="B0E34B533046FFAFFF5EFDE9BC970F89" bold="true" box="[171,500,635,659]" pageId="63" pageNumber="264">
<smallCapsWord id="84CE019D3046FFAFFF5EFDE9BC350F89" baselines="653,653" box="[171,342,635,659]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="63" pageNumber="264">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="84CE019D3046FFAFFE95FDEDBC970F89" baselines="653" box="[352,500,639,658]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="63" pageNumber="264">DISTRIBUTION</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C3046FFAFFDE9FDEEB9250E28" country="Madagascar" county="Ranomafana National Park" location="Ranomafana" municipality="Anjozorobe" pageId="63" pageNumber="264" specimenCount="2" specimenCount-female="2" stateProvince="Antananarivo">
We have examined specimens from only two areas.
<specimenCount id="94915CC83046FFAFFB93FDEEBDA70FA8" count="2" pageId="63" pageNumber="264" type="female">Two female</specimenCount>
specimens were collected in
<collectingCounty id="6B49EFCD3046FFAFFDFDFD09BE4D0FA8" box="[520,814,667,691]" pageId="63" pageNumber="264">Ranomafana National Park</collectingCounty>
, the
<typeStatus id="5D2C29E33046FFAFFC92FD0EBEF60FAF" box="[871,917,668,692]" pageId="63" pageNumber="264">type</typeStatus>
locality, on the eastern slope off the central plateau in the northeastern part of the southern half of the island of
<collectingCountry id="FA80D7D13046FFAFFBD9FD29BDFF0FE8" name="Madagascar" pageId="63" pageNumber="264">Madagascar</collectingCountry>
. A third specimen, also a female, was collected near
<collectingMunicipality id="624C0D3B3046FFAFFD1EFD49BE0A0FE8" box="[747,873,731,755]" pageId="63" pageNumber="264">Anjozorobe</collectingMunicipality>
, in
<collectingRegion id="405359A33046FFAFFC60FD49B9EC0FE8" box="[917,1167,731,755]" country="Madagascar" name="Antananarivo" pageId="63" pageNumber="264">Antananarivo Province</collectingRegion>
, slightly north of the center of the island and just over
<quantity id="456F3AA43046FFAFFD4DFD69BE6A0E08" box="[696,777,763,787]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="63" pageNumber="264" unit="km" value="300.0">300 km</quantity>
north of
<location id="8748C19A3046FFAFFC99FD69BE900E08" LSID="urn:lsid:plazi:treatment:0A3E26573044FFAFFF8EF9B3BE7F0E68:8748C19A3046FFAFFC99FD69BE900E08" box="[876,1011,763,787]" country="Madagascar" county="Ranomafana National Park" municipality="Anjozorobe" name="Ranomafana" pageId="63" pageNumber="264" stateProvince="Antananarivo">Ranomafana</location>
along the eastern slope off the central plateau.
<location id="8748C19A3046FFAFFE29FC8EBC9E0E28" LSID="urn:lsid:plazi:treatment:0A3E26573044FFAFFF8EF9B3BE7F0E68:8748C19A3046FFAFFE29FC8EBC9E0E28" box="[476,509,796,819]" country="Madagascar" county="Ranomafana National Park" municipality="Anjozorobe" name="No" pageId="63" pageNumber="264" stateProvince="Antananarivo">No</location>
specimens have been recorded from intervening areas
</materialsCitation>
.
</paragraph>
<paragraph id="822897413046FFAFFF5EFCA9BE7F0E68" blockId="63.[123,1174,219,883]" pageId="63" pageNumber="264">
<emphasis id="B0E34B533046FFAFFF5EFCA9BCC60E49" bold="true" box="[171,421,827,851]" pageId="63" pageNumber="264">
<smallCapsWord id="84CE019D3046FFAFFF5EFCA9BC6A0E49" baselines="845,845" box="[171,265,827,851]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="63" pageNumber="264">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D3046FFAFFEE4FCADBCC60E49" baselines="845" box="[273,421,831,850]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="63" pageNumber="264">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Specimens of the
<typeStatus id="5D2C29E33046FFAFFD7BFCAEBFDF0E4F" box="[654,700,828,852]" pageId="63" pageNumber="264">type</typeStatus>
series were all collected in malaise traps in montane rainforest at elevations ranging from
<quantity id="456F3AA43046FFAFFD98FCC9BE750E68" box="[621,790,859,883]" metricMagnitude="3" metricUnit="m" metricValue="1.205" metricValueMax="1.3" metricValueMin="1.1099999999999999" pageId="63" pageNumber="264" unit="m" value="1205.0" valueMax="1300.0" valueMin="1110.0">1110 to 1300 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

338
data/0A/3E/26/0A3E26573049FFA3FF8EFA93BE8D09FD.xml Normal file
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@ -0,0 +1,338 @@
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<mods:title id="F1FD87F2FA115F9E4E0759769460BCC7">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:namePart id="F399DBEEF43E0A2FBB519478DCE3AFD4">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="43B053A177432E89CAF36A72234B3079">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:namePart id="79CF9606ED7DD525AC015AB728B8B43A">Rainio, Johanna</mods:namePart>
<mods:affiliation id="1CC8C15FD70A845ADE96BB7A2790B416">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
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<subSubSection id="CA8DC4CA3049FFA0FF8EFA93BE270801" box="[123,836,1281,1306]" pageId="48" pageNumber="249" type="nomenclature">
<paragraph id="822897413049FFA0FF8EFA93BE270801" blockId="48.[123,836,1281,1336]" box="[123,836,1281,1306]" pageId="48" pageNumber="249">
<heading id="D960202D3049FFA0FF8EFA93BE270801" box="[123,836,1281,1306]" fontSize="10" level="2" pageId="48" pageNumber="249" reason="4">
<emphasis id="B0E34B533049FFA0FF8EFA93BE270801" bold="true" box="[123,836,1281,1306]" pageId="48" pageNumber="249">
<taxonomicName id="4597ECC23049FFA0FF8EFA93BF830802" ID-CoL="4P2M6" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,736,1281,1306]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533049FFA0FF8EFA93BCAC0802" bold="true" box="[123,463,1281,1305]" italics="true" pageId="48" pageNumber="249">Pseudomasoreus ranomafanae</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F6283049FFA0FD1BFA90BE270801" box="[750,836,1282,1306]" pageId="48" pageNumber="249" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3049FFA0FF8EFAB0BFDA089B" pageId="48" pageNumber="249" type="description">
<paragraph id="822897413049FFA0FF8EFAB0BC2F082C" blockId="48.[123,836,1281,1336]" box="[123,332,1314,1336]" pageId="48" pageNumber="249">
<figureCitation id="1AAC8BC43049FFA0FF8EFAB0BD82082C" box="[123,225,1314,1336]" captionStart="FIGURE" captionStartId="49.[155,167,1542,1561]" captionTargetBox="[139,1172,230,1525]" captionTargetId="figure-67@49.[120,1176,216,1539]" captionTargetPageId="49" captionText="FIGURE 29. Digital images of holotype male of Pseudomasoreus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line A = 1.0 mm." pageId="48" pageNumber="249">Figures 29</figureCitation>
,
<figureCitation id="1AAC8BC43049FFA0FF1BFAB0BC2F082C" box="[238,332,1314,1335]" captionStart="FIGURE" captionStartId="50.[155,167,1556,1575]" captionTargetBox="[121,1176,219,1729]" captionTargetId="figure-1468@50.[600,1176,326,1728]" captionTargetPageId="50" captionText="FIGURE 30. Digital images of Pseudomasoreus spp. A–C. Holotype male of P. ranomafanae Kavanaugh &amp; Rainio sp. nov.; D, E. P. catalai Jean- nel (from Ranomafana National Park). A. Right labi- al palpus, ventral aspect; B–E. Aedeagus of male genitalia; B, D. Left lateral aspect; C, E. Ventral aspect. Scale lines = 0.5 mm." pageId="48" pageNumber="249">30A, 30C</figureCitation>
</paragraph>
<paragraph id="822897413049FFA0FF8EFADFBFCF0879" blockId="48.[123,697,1357,1409]" box="[123,684,1357,1379]" pageId="48" pageNumber="249">
<emphasis id="B0E34B533049FFA0FF8EFADFBC460879" box="[123,293,1357,1378]" italics="true" pageId="48" pageNumber="249">
Deuveilla
<taxonomicNameLabel id="ABD0F6283049FFA0FF2AFADFBC710879" box="[223,274,1357,1378]" pageId="48" pageNumber="249" rank="species">n. sp.</taxonomicNameLabel>
1
</emphasis>
; Rainio 2009: 31(informal designation).
</paragraph>
<paragraph id="822897413049FFA0FF8EFAF9BFDA089B" blockId="48.[123,697,1357,1409]" box="[123,697,1387,1409]" pageId="48" pageNumber="249">
<taxonomicName id="4597ECC23049FFA0FF8EFAF9BDAC089A" authorityName="Bonelli" authorityYear="1810" box="[123,207,1387,1409]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="subFamily" subFamily="Lebiinae">Lebiinae</taxonomicName>
(n. sp. 1); Rainio 2012: 72 (informal designation).
</paragraph>
</subSubSection>
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</emphasis>
.—
<materialsCitation id="32FF9D1C3049FFA0FE77FA36BDDB0B86" collectingDate="2001-01-02" collectingDateMax="2001-01-22" collectingDateMin="2001-01-02" collectorName="D. H. &amp; K. M Kavanugh &amp; R. L. Brett &amp; E. Elsom &amp; F. Vargas &amp; R. Ranaivosolo &amp; E. F. Randrianirina &amp; N. Rasoamananana &amp; T. J. Ravelomanana &amp; H. C. Raveloson" country="Madagascar" elevation="900" latitude="-21.25041" location="Talatakely area" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" pageId="48" pageNumber="249" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E33049FFA0FE77FA36BC8608A7" box="[386,485,1444,1468]" pageId="48" pageNumber="249" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43049FFA0FE02FA36BF1A08A7" box="[503,633,1444,1468]" captionStart="FIGURE" captionStartId="49.[155,167,1542,1561]" captionTargetBox="[139,1172,230,1525]" captionTargetId="figure-67@49.[120,1176,216,1539]" captionTargetPageId="49" captionText="FIGURE 29. Digital images of holotype male of Pseudomasoreus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line A = 1.0 mm." pageId="48" pageNumber="249">Figs.29A–B</figureCitation>
), a male, in CAS, labeled: “CASENT 1004471”/ “
<collectingCountry id="FA80D7D13049FFA0FF73FA56BC2708C7" box="[134,324,1476,1500]" name="Madagascar" pageId="48" pageNumber="249">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A33049FFA0FEA0FA56BF2408C7" box="[341,583,1476,1500]" country="Madagascar" name="Fianarantsoa" pageId="48" pageNumber="249">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B3049FFA0FDADFA56BEE308C7" box="[600,896,1476,1500]" pageId="48" pageNumber="249">Ranomafana National Park</collectingMunicipality>
,
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,
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<elevation id="09BA70723049FFA0FBB2FA56B9EC08C7" box="[1095,1167,1476,1500]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="48" pageNumber="249" unit="m" value="900.0">900 m</elevation>
</quantity>
, mixed tropical forest,
<date id="F629B1813049FFA0FE9EFA77BF5708E6" box="[363,564,1509,1533]" pageId="48" pageNumber="249" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">
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</date>
,”/ “
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/
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, Stop# DHK-01-001,
<collectorName id="2F62F2973049FFA0FB94FA77BC280B06" pageId="48" pageNumber="249">D.H. &amp; K.M Kavanugh</collectorName>
,
<collectorName id="2F62F2973049FFA0FE95F997BCB10B07" box="[352,466,1541,1564]" pageId="48" pageNumber="249">R.L. Brett</collectorName>
,
<collectorName id="2F62F2973049FFA0FE1DF997BF320B06" box="[488,593,1541,1565]" pageId="48" pageNumber="249">E. Elsom</collectorName>
,
<collectorName id="2F62F2973049FFA0FD93F997BFAC0B06" box="[614,719,1541,1565]" pageId="48" pageNumber="249">F. Vargas</collectorName>
,
<collectorName id="2F62F2973049FFA0FD11F997BEEC0B06" box="[740,911,1541,1565]" pageId="48" pageNumber="249">R. Ranaivosolo</collectorName>
,”/ “
<collectorName id="2F62F2973049FFA0FC31F997B9EC0B06" box="[964,1167,1541,1565]" pageId="48" pageNumber="249">E.F. Randrianirina</collectorName>
,
<collectorName id="2F62F2973049FFA0FF8EF9B7BC2F0B27" box="[123,332,1573,1596]" pageId="48" pageNumber="249">N. Rasoamananana</collectorName>
,
<collectorName id="2F62F2973049FFA0FEA8F9B7BF480B26" box="[349,555,1573,1597]" pageId="48" pageNumber="249">T.J. Ravelomanana</collectorName>
, and
<collectorName id="2F62F2973049FFA0FD98F9B7BE780B26" box="[621,795,1573,1597]" pageId="48" pageNumber="249">H.C. Raveloson</collectorName>
collectors”/ “collected in daytime from large accumulation of dead leaf, twig, and branch debris on and extended up to
<quantity id="456F3AA43049FFA0FBD7F9D7B92E0B47" box="[1058,1101,1605,1629]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="48" pageNumber="249" unit="m" value="2.0">
<elevation id="09BA70723049FFA0FBD7F9D7B92E0B47" box="[1058,1101,1605,1629]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="48" pageNumber="249" unit="m" value="2.0">2 m</elevation>
</quantity>
above forest floor”/ “
<typeStatus id="5D2C29E33049FFA0FEEDF9F7BCC60B66" box="[280,421,1637,1661]" pageId="48" pageNumber="249" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23049FFA0FE5EF9F7BEBF0B66" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[427,988,1637,1661]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533049FFA0FE5EF9F7BF900B66" box="[427,755,1637,1661]" italics="true" pageId="48" pageNumber="249">Pseudomasoreus ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283049FFA0FC16F9F4B97F0B66" box="[995,1052,1638,1661]" pageId="48" pageNumber="249" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label]
</materialsCitation>
.
<materialsCitation id="32FF9D1C3049FFA0FF3EF914BD870BA5" collectionCode="CAS" country="Madagascar" pageId="48" pageNumber="249" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="5D2C29E33049FFA0FF3EF914BC500B85" box="[203,307,1670,1694]" pageId="48" pageNumber="249" type="paratype">Paratypes</typeStatus>
(total of 8):
<specimenCount id="94915CC83049FFA0FE21F917BF410B86" box="[468,546,1669,1693]" count="1" pageId="48" pageNumber="249" type="male">1 male</specimenCount>
(in
<collectionCode id="E4860F843049FFA0FDAFF917BFED0B86" box="[602,654,1669,1693]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="48" pageNumber="249" type="Museum">CAS</collectionCode>
) labeled same as
<typeStatus id="5D2C29E33049FFA0FC90F917BEA10B86" box="[869,962,1669,1693]" pageId="48" pageNumber="249" type="holotype">holotype</typeStatus>
except “CASENT 1004472”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3049FFA0FF07F934B9ED0BA6" box="[242,1166,1701,1726]" collectionCode="CAS" country="Madagascar" pageId="48" pageNumber="249" specimenCount="2" specimenCount-male="2" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="94915CC83049FFA0FF07F934BC270BA6" box="[242,324,1701,1726]" count="2" pageId="48" pageNumber="249" type="male">2 males</specimenCount>
(in
<collectionCode id="E4860F843049FFA0FE9AF934BCC00BA5" box="[367,419,1702,1726]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="48" pageNumber="249" type="Museum">CAS</collectionCode>
) labeled “CASENT 1004003” and “CASENT 1004004”, respectively
</materialsCitation>
/
</paragraph>
<caption id="D6E8C7C93048FFA1FF6EF994BF5E0B28" pageId="49" pageNumber="250" startId="49.[155,167,1542,1561]" targetBox="[139,1172,230,1525]" targetPageId="49" targetType="figure">
<paragraph id="822897413048FFA1FF6EF994BF5E0B28" blockId="49.[123,1174,1542,1587]" pageId="49" pageNumber="250">
<smallCapsWord id="84CE019D3048FFA1FF6EF994BDBB0B03" baselines="1556,1556" box="[155,216,1542,1561]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="49" pageNumber="250">FIGURE</smallCapsWord>
29. Digital images of holotype male of
<taxonomicName id="4597ECC23048FFA1FDC0F994BE980B02" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[565,1019,1542,1561]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="250" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533048FFA1FDC0F994BE5D0B02" box="[565,830,1542,1561]" italics="true" pageId="49" pageNumber="250">Pseudomasoreus ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283048FFA1FBF7F994B9270B02" box="[1026,1092,1542,1561]" pageId="49" pageNumber="250" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels. Scale line A = 1.0 mm.
</paragraph>
</caption>
<caption id="D6E8C7C9304BFFA2FF6EF986BC160BDA" pageId="50" pageNumber="251" startId="50.[155,167,1556,1575]" targetBox="[121,1176,219,1729]" targetPageId="50" targetType="figure">
<paragraph id="82289741304BFFA2FF6EF986BF580B3C" blockId="50.[123,571,1556,1729]" box="[155,571,1556,1575]" pageId="50" pageNumber="251">
<smallCapsWord id="84CE019D304BFFA2FF6EF986BDBB0B3D" baselines="1570,1570" box="[155,216,1556,1575]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="50" pageNumber="251">FIGURE</smallCapsWord>
30. Digital images of
<taxonomicName id="4597ECC2304BFFA2FE5EF986BF580B3C" box="[427,571,1556,1575]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="251" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B0E34B53304BFFA2FE5EF986BF580B3C" box="[427,571,1556,1575]" italics="true" pageId="50" pageNumber="251">Pseudomasoreus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="82289741304BFFA2FF8EF9BCBF580B5A" blockId="50.[123,571,1556,1729]" box="[123,571,1581,1601]" pageId="50" pageNumber="251">
spp. A–C. Holotype male of
<taxonomicName id="4597ECC2304BFFA2FE53F9BCBF580B5A" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[422,571,1582,1601]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="251" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53304BFFA2FE53F9BCBF580B5A" box="[422,571,1582,1601]" italics="true" pageId="50" pageNumber="251">P. ranomafanae</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="82289741304BFFA2FF8EF9D5BC160BDA" blockId="50.[123,571,1556,1729]" pageId="50" pageNumber="251">
Kavanaugh &amp; Rainio
<taxonomicNameLabel id="ABD0F628304BFFA2FEC2F9D5BC1B0B41" box="[311,376,1607,1626]" pageId="50" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
; D, E.
<taxonomicName id="4597ECC2304BFFA2FE42F9DABDF60B6F" authority="Jeannel" authorityName="Jeannel" authorityYear="1949" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="251" phylum="Arthropoda" rank="species" species="catalai">
<emphasis id="B0E34B53304BFFA2FE42F9DABF6B0B41" box="[439,520,1607,1626]" italics="true" pageId="50" pageNumber="251">P. catalai</emphasis>
Jeannel
</taxonomicName>
(from Ranomafana National Park). A. Right labial palpus, ventral aspect; B–E. Aedeagus of male genitalia; B, D. Left lateral aspect; C, E. Ventral aspect. Scale lines = 0.5 mm.
</paragraph>
</caption>
<paragraph id="82289741304AFFA3FF5EFF49BECA0C08" blockId="51.[123,1174,219,1254]" pageId="51" pageNumber="252">
<emphasis id="B0E34B53304AFFA3FF5EFF49BC910DEA" bold="true" box="[171,498,219,243]" pageId="51" pageNumber="252">
<smallCapsWord id="84CE019D304AFFA3FF5EFF49BC4D0DEA" baselines="237,236" box="[171,302,219,243]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="51" pageNumber="252">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D304AFFA3FEC3FF4CBC310DEA" baselines="236" box="[310,338,222,241]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="51" pageNumber="252">OF</smallCapsWord>
<smallCapsWord id="84CE019D304AFFA3FEACFF4CBCCF0DEA" baselines="236" box="[345,428,222,241]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="51" pageNumber="252">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D304AFFA3FE40FF4CBC910DEA" baselines="236" box="[437,498,222,241]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="51" pageNumber="252">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC2304AFFA3FD0CFF49BEE40DE8" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[761,903,219,243]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="252" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53304AFFA3FD0CFF49BEE40DE8" box="[761,903,219,243]" italics="true" pageId="51" pageNumber="252">ranomafanae</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the national park in which the
<typeStatus id="5D2C29E3304AFFA3FD29FF6EBE690C0F" box="[732,778,252,276]" pageId="51" pageNumber="252">type</typeStatus>
was collected.
</paragraph>
<paragraph id="82289741304AFFA3FF5EFE8EB9F60EBB" blockId="51.[123,1174,219,1254]" pageId="51" pageNumber="252">
<smallCapsWord id="84CE019D304AFFA3FF5EFE8EBC210C28" baselines="302,302" box="[171,322,284,308]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="51" pageNumber="252">
<emphasis id="B0E34B53304AFFA3FF5EFE8EBC210C28" bold="true" box="[171,322,284,308]" pageId="51" pageNumber="252">RECOGNITION</emphasis>
</smallCapsWord>
.— Size large for genus, SBL of males = 9.7–10.
<quantity id="456F3AA4304AFFA3FCA6FE8EBEF20C2F" box="[851,913,284,308]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="51" pageNumber="252" unit="mm" value="6.0">6 mm</quantity>
, of females =
<quantity id="456F3AA4304AFFA3FBDBFE8EB9ED0C2F" box="[1070,1166,284,308]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="51" pageNumber="252" unit="mm" value="10.3">10.3 mm</quantity>
. In
<bibRefCitation id="E606EAB0304AFFA3FF6FFEAFBC230C4E" author="MATEU, J." box="[154,320,317,341]" pageId="51" pageNumber="252" pagination="15 - 23" refId="ref31841" refString="MATEU, J. 1980. Notes sur les Lebiens malagaches (5 e note). Sur trois nouveaux Pseudomasoreus Desbrochers [Coleoptera, Carabidae]. Revue Francaise d'Entomologie (N. S.) 2: 15 - 23." type="journal article" year="1980">Mateu’s (1980)</bibRefCitation>
key to the Malagasy species of this genus, members of
<taxonomicName id="4597ECC2304AFFA3FC5BFEACB93A0C4E" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[942,1113,317,341]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="252" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53304AFFA3FC5BFEACB93A0C4E" box="[942,1113,317,341]" italics="true" pageId="51" pageNumber="252">P. ranomafanae</emphasis>
</taxonomicName>
(
<figureCitation id="1AAC8BC4304AFFA3FB9EFEAFBDCB0C6D" captionStart="FIGURE" captionStartId="49.[155,167,1542,1561]" captionTargetBox="[139,1172,230,1525]" captionTargetId="figure-67@49.[120,1176,216,1539]" captionTargetPageId="49" captionText="FIGURE 29. Digital images of holotype male of Pseudomasoreus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line A = 1.0 mm." pageId="51" pageNumber="252">Fig. 29A</figureCitation>
) key to
<taxonomicName id="4597ECC2304AFFA3FEF5FECFBFA60C6D" authority="Jeannel (1941 b)" authorityName="Jeannel" authorityYear="1941" box="[256,709,349,374]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="252" phylum="Arthropoda" rank="species" species="decorsei">
<emphasis id="B0E34B53304AFFA3FEF5FECFBF770C6E" box="[256,532,349,373]" italics="true" pageId="51" pageNumber="252">Pseudomasoreus decorsei</emphasis>
<bibRefCitation id="E606EAB0304AFFA3FDEFFECFBFA60C6D" author="JEANNEL, R." box="[538,709,349,374]" pageId="51" pageNumber="252" pagination="62 - 67" refId="ref30833" refString="JEANNEL, R. 1941 b. Sur le genre Pseudomasoreus Desbrochers. Bulletin de la Societe Entomologique de France 1941: 62 - 67." type="journal article" year="1941">Jeannel (1941b)</bibRefCitation>
</taxonomicName>
based on the following combination of features: size large, SBL =
<quantity id="456F3AA4304AFFA3FE88FEECBCAD0C8D" box="[381,462,382,406]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="51" pageNumber="252" unit="mm" value="8.5">8.5 mm</quantity>
or more; elytra concolorous brown or piceous, without pale areas. However, they differ from the unique
<typeStatus id="5D2C29E3304AFFA3FDFBFE0DBF080CAC" box="[526,619,415,439]" pageId="51" pageNumber="252" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4597ECC2304AFFA3FD7BFE0DBE610CAC" authorityName="Jeannel" authorityYear="1941" box="[654,770,415,439]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="252" phylum="Arthropoda" rank="species" species="decorsei">
<emphasis id="B0E34B53304AFFA3FD7BFE0DBE610CAC" box="[654,770,415,439]" italics="true" pageId="51" pageNumber="252">P. decorsei</emphasis>
</taxonomicName>
in having even larger body size (SBL =
<quantity id="456F3AA4304AFFA3FF65FE2DBD860CCC" box="[144,229,447,471]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="51" pageNumber="252" unit="mm" value="8.5">8.5 mm</quantity>
in
<taxonomicName id="4597ECC2304AFFA3FEFBFE52BCE50CCC" authorityName="Jeannel" authorityYear="1941" box="[270,390,447,471]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="252" phylum="Arthropoda" rank="species" species="decorsei">
<emphasis id="B0E34B53304AFFA3FEFBFE52BCE50CCC" box="[270,390,447,471]" italics="true" pageId="51" pageNumber="252">P. decorsei</emphasis>
</taxonomicName>
type), pronotum narrower overall and especially basally and with less rounded, more evident hind angles (pronotum broad, rounded, with hind angles effaced by smoothly rounded transition from lateral to basal margin in
<taxonomicName id="4597ECC2304AFFA3FD42FD93BE4F0F02" authorityName="Jeannel" authorityYear="1941" box="[695,812,513,537]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="252" phylum="Arthropoda" rank="species" species="decorsei">
<emphasis id="B0E34B53304AFFA3FD42FD93BE4F0F02" box="[695,812,513,537]" italics="true" pageId="51" pageNumber="252">P. decorsei</emphasis>
</taxonomicName>
type), and elytral lateral margins nearly parallel (more evenly rounded in
<taxonomicName id="4597ECC2304AFFA3FDDEFDB0BFC30F22" authorityName="Jeannel" authorityYear="1941" box="[555,672,545,569]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="252" phylum="Arthropoda" rank="species" species="decorsei">
<emphasis id="B0E34B53304AFFA3FDDEFDB0BFC30F22" box="[555,672,545,569]" italics="true" pageId="51" pageNumber="252">P. decorsei</emphasis>
</taxonomicName>
type). Members of
<taxonomicName id="4597ECC2304AFFA3FC80FDB0B97D0F22" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[885,1054,545,569]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="252" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53304AFFA3FC80FDB0B97D0F22" box="[885,1054,545,569]" italics="true" pageId="51" pageNumber="252">P. ranomafanae</emphasis>
</taxonomicName>
are actually more similar to those of
<taxonomicName id="4597ECC2304AFFA3FE6CFDD1BC9A0F41" box="[409,505,578,602]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="252" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B53304AFFA3FE6CFDD1BC9A0F41" box="[409,505,578,602]" italics="true" pageId="51" pageNumber="252">P. deuvei</emphasis>
</taxonomicName>
, which was not included in Mateu’s key. They share large size, general body form with long, parallel-sided elytra, and antennomere 3 glabrous except for the apical whorl of fixed setae (additional pubescence seen in apical part of antennomere
<quantity id="456F3AA4304AFFA3FC06FD11B97E0F80" box="[1011,1053,643,667]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="51" pageNumber="252" unit="in" value="3.0">3 in</quantity>
most other species of genus). However, their body size is slightly smaller on average (SBL = 10.2–11.
<quantity id="456F3AA4304AFFA3FB9CFD31B9F50FA7" box="[1129,1174,675,700]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="51" pageNumber="252" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="4597ECC2304AFFA3FF8EFD57BDBE0FC7" box="[123,221,708,732]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="252" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B53304AFFA3FF8EFD57BDBE0FC7" box="[123,221,708,732]" italics="true" pageId="51" pageNumber="252">P. deuvei</emphasis>
</taxonomicName>
members), the pronotum is narrower overall and more narrowed basally, with the hind angles more distinct than in
<taxonomicName id="4597ECC2304AFFA3FE52FD74BF640FE6" box="[423,519,741,765]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="252" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B53304AFFA3FE52FD74BF640FE6" box="[423,519,741,765]" italics="true" pageId="51" pageNumber="252">P. deuvei</emphasis>
</taxonomicName>
members and the lateral margins either straight or with slight sinuation anterior to the hind angles (lateral margins not at all sinuate in the latter). Also, the penultimate labial palpomeres in all specimens of the type series have three or four setae (
<figureCitation id="1AAC8BC4304AFFA3FBD3FCB4B9E50E25" box="[1062,1158,806,830]" captionStart="FIGURE" captionStartId="49.[155,167,1542,1561]" captionTargetBox="[139,1172,230,1525]" captionTargetId="figure-67@49.[120,1176,216,1539]" captionTargetPageId="49" captionText="FIGURE 29. Digital images of holotype male of Pseudomasoreus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line A = 1.0 mm." pageId="51" pageNumber="252">Fig. 29C</figureCitation>
), whereas members of
<taxonomicName id="4597ECC2304AFFA3FE94FCD5BCA10E44" box="[353,450,839,863]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="252" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B53304AFFA3FE94FCD5BCA10E44" box="[353,450,839,863]" italics="true" pageId="51" pageNumber="252">P. deuvei</emphasis>
</taxonomicName>
have only two such setae, a feature particularly noted by Casale in his original description. Finally, the median lobe of male genitalia (
<figureCitation id="1AAC8BC4304AFFA3FC9AFCF5BE960E64" box="[879,1013,871,895]" captionStart="FIGURE" captionStartId="50.[155,167,1556,1575]" captionTargetBox="[121,1176,219,1729]" captionTargetId="figure-1468@50.[600,1176,326,1728]" captionTargetPageId="50" captionText="FIGURE 30. Digital images of Pseudomasoreus spp. A–C. Holotype male of P. ranomafanae Kavanaugh &amp; Rainio sp. nov.; D, E. P. catalai Jean- nel (from Ranomafana National Park). A. Right labi- al palpus, ventral aspect; B–E. Aedeagus of male genitalia; B, D. Left lateral aspect; C, E. Ventral aspect. Scale lines = 0.5 mm." pageId="51" pageNumber="252">Figs. 30A,C</figureCitation>
) has the shaft slightly less arcuate and the apex slightly longer on more rounded apically than in
<taxonomicName id="4597ECC2304AFFA3FC05FC1BB9330EBB" box="[1008,1104,904,928]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="252" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B53304AFFA3FC05FC1BB9330EBB" box="[1008,1104,904,928]" italics="true" pageId="51" pageNumber="252">P. deuvei</emphasis>
</taxonomicName>
males
</paragraph>
<paragraph id="82289741304AFFA3FF8EFC3BBF930939" blockId="51.[123,1174,219,1254]" pageId="51" pageNumber="252">
(see
<bibRefCitation id="E606EAB0304AFFA3FF59FC3ABC4C0EDA" author="CASALE, A." box="[172,303,936,961]" pageId="51" pageNumber="252" pagination="117 - 124" refId="ref29659" refString="CASALE, A. 1998. Pseudomasoreus deuvei new species from Madagascar, with notes about some genera of afrotropical and Madagascan Cymindini. Bolletino della Societa Entomologica Italiana 130: 117 - 124." type="journal article" year="1998">Casale 1998</bibRefCitation>
,
<figureCitation id="1AAC8BC4304AFFA3FEC9FC3ABCFA0EDB" box="[316,409,936,961]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStartId-0="7.[155,167,1251,1270]" captionStartId-1="9.[155,167,1384,1403]" captionTargetBox-0="[120,1176,216,1233]" captionTargetBox-1="[140,1176,216,1362]" captionTargetId-0="figure-985@7.[120,1176,216,1235]" captionTargetId-1="figure-703@9.[120,1176,216,1381]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionText-0="FIGURE 4. Digital images of holotype female of Caelostomus latus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Head, left lateral aspect (arrow points to tranverse depression between eyes); D. Right anterior tarsus, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." captionText-1="FIGURE 5. Digital images of holotype female of Caelostomus rotundiformis Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Right anterior tarsus, dorsal aspect. Scale line A = 1.0 mm, C = 0.5 mm." pageId="51" pageNumber="252">Figs.4–5</figureCitation>
). For future comparative purposes, we also provide digital photographs of the median lobe of the genitalia of a male
<taxonomicName id="4597ECC2304AFFA3FDA0FC5BBE9F0EFA" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" box="[597,1020,969,993]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="252" phylum="Arthropoda" rank="species" species="catalai">
<emphasis id="B0E34B53304AFFA3FDA0FC5BBE3A0EFA" box="[597,857,969,993]" italics="true" pageId="51" pageNumber="252">Pseudomasoreus catalai</emphasis>
Jeannel (1949)
</taxonomicName>
(
<figureCitation id="1AAC8BC4304AFFA3FBFEFC5BB9EF0EFA" box="[1035,1164,969,993]" captionStart="FIGURE" captionStartId="50.[155,167,1556,1575]" captionTargetBox="[121,1176,219,1729]" captionTargetId="figure-1468@50.[600,1176,326,1728]" captionTargetPageId="50" captionText="FIGURE 30. Digital images of Pseudomasoreus spp. A–C. Holotype male of P. ranomafanae Kavanaugh &amp; Rainio sp. nov.; D, E. P. catalai Jean- nel (from Ranomafana National Park). A. Right labi- al palpus, ventral aspect; B–E. Aedeagus of male genitalia; B, D. Left lateral aspect; C, E. Ventral aspect. Scale lines = 0.5 mm." pageId="51" pageNumber="252">Figs. 30B,D</figureCitation>
) [male previously unknown] to complement the illustrations of other
<taxonomicName id="4597ECC2304AFFA3FCAFFC78B9010919" box="[858,1122,1002,1026]" pageId="51" pageNumber="252">
<emphasis id="B0E34B53304AFFA3FCAFFC78B96D0919" box="[858,1038,1002,1026]" italics="true" pageId="51" pageNumber="252">Pseudomasoreus</emphasis>
species
</taxonomicName>
provided by Jeannel (1949),
<bibRefCitation id="E606EAB0304AFFA3FE79FB99BF7D0938" author="MATEU, J." box="[396,542,1034,1059]" pageId="51" pageNumber="252" pagination="15 - 23" refId="ref31841" refString="MATEU, J. 1980. Notes sur les Lebiens malagaches (5 e note). Sur trois nouveaux Pseudomasoreus Desbrochers [Coleoptera, Carabidae]. Revue Francaise d'Entomologie (N. S.) 2: 15 - 23." type="journal article" year="1980">Mateu (1980)</bibRefCitation>
and
<bibRefCitation id="E606EAB0304AFFA3FDA6FB98BF8A0938" author="CASALE, A." box="[595,745,1034,1059]" pageId="51" pageNumber="252" pagination="117 - 124" refId="ref29659" refString="CASALE, A. 1998. Pseudomasoreus deuvei new species from Madagascar, with notes about some genera of afrotropical and Madagascan Cymindini. Bolletino della Societa Entomologica Italiana 130: 117 - 124." type="journal article" year="1998">Casale (1998)</bibRefCitation>
.
</paragraph>
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<emphasis id="B0E34B53304AFFA3FF5EFBB9BC92095A" bold="true" box="[171,497,1067,1091]" pageId="51" pageNumber="252">
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<smallCapsWord id="84CE019D304AFFA3FEA8FBBCBC92095A" baselines="1084" box="[349,497,1070,1089]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="51" pageNumber="252">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E3304AFFA3FC8AFBB9BECE0958" box="[895,941,1067,1091]" pageId="51" pageNumber="252">type</typeStatus>
locality.
</paragraph>
<paragraph id="82289741304AFFA3FF5EFBDEBE8D09FD" blockId="51.[123,1174,219,1254]" pageId="51" pageNumber="252">
<emphasis id="B0E34B53304AFFA3FF5EFBDEBCC00979" bold="true" box="[171,419,1100,1124]" pageId="51" pageNumber="252">
<smallCapsWord id="84CE019D304AFFA3FF5EFBDEBC6A0979" baselines="1118,1117" box="[171,265,1100,1124]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="51" pageNumber="252">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D304AFFA3FEFAFBDDBCC00979" baselines="1117" box="[271,419,1103,1122]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="51" pageNumber="252">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Specimens of this species were are collected in secondary montane rainforest in the Talatakely and Vohiparara areas at elevations ranging from
<quantity id="456F3AA4304AFFA3FC4CFBFEB922099F" box="[953,1089,1132,1156]" metricMagnitude="3" metricUnit="m" metricValue="1.035" metricValueMax="1.17" metricValueMin="0.9" pageId="51" pageNumber="252" unit="m" value="1035.0" valueMax="1170.0" valueMin="900.0">900–1170 m</quantity>
. Adults were collected by hand, by beating live vegetation and suspended dead leaves and twigs, particularly dead leaves suspended is leaf whirls of
<taxonomicName id="4597ECC2304AFFA3FDABFB3FBF8C09DE" box="[606,751,1197,1221]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="51" pageNumber="252" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B0E34B53304AFFA3FDABFB3FBFAA09DE" box="[606,713,1197,1221]" italics="true" pageId="51" pageNumber="252">Pandanus</emphasis>
sp.
</taxonomicName>
<specimenCount id="94915CC8304AFFA3FD00FB3CBEF809DE" box="[757,923,1197,1221]" count="2" pageId="51" pageNumber="252" type="generic">Two specimens</specimenCount>
were found in daytime in masses of dead leaves, twigs and branch debris accumulated on the forest floor.
</paragraph>
</subSubSection>
</treatment>
</document>

143
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<mods:namePart id="61BE9E10B6EA7251D192D06F4B96F1F2">Rainio, Johanna</mods:namePart>
<mods:affiliation id="87D4EB678AC3B34DE34DD371E4A95E81">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
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<heading id="D960202D304AFFA3FF8EFA35BF8D08DB" box="[123,750,1447,1472]" fontSize="10" level="2" pageId="51" pageNumber="252" reason="4">
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<taxonomicName id="4597ECC2304AFFA3FF8EFA35BFE908A4" ID-CoL="HMB8" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,650,1447,1472]" class="Insecta" family="Carabidae" genus="Assadecma" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="252" phylum="Arthropoda" rank="species" species="basilewskyi" status="sp. nov.">
<emphasis id="B0E34B53304AFFA3FF8EFA35BC1A08A4" bold="true" box="[123,377,1447,1471]" italics="true" pageId="51" pageNumber="252">Assadecma basilewskyi</emphasis>
Kavanaugh and Rainio
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,
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, 32
</paragraph>
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<smallCapsWord id="84CE019D304AFFA3FF19F994BC3A0B02" baselines="1556" box="[236,345,1542,1561]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="51" pageNumber="252">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C304AFFA3FE74F990BF040B67" collectingDate="1994-04-18" collectorName="E. Rajeriarison &amp; R. Malenky" country="Madagascar" elevation="940" latitude="-21.266666" location="Ranomafana" longLatPrecision="1264" longitude="47.416668" pageId="51" pageNumber="252" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E3304AFFA3FE74F990BC870B01" box="[385,484,1538,1562]" pageId="51" pageNumber="252" type="holotype">Holotype</typeStatus>
(
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), a male, in NMNH, labeled: “
<collectingCountry id="FA80D7D1304AFFA3FC25F990B9ED0B01" box="[976,1166,1538,1562]" name="Madagascar" pageId="51" pageNumber="252">MADAGASCAR</collectingCountry>
: Prov.
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,
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W
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,
<quantity id="456F3AA4304AFFA3FDCBF9B1BF180B21" box="[574,635,1571,1595]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="51" pageNumber="252" unit="m" value="940.0">
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,
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<date id="F629B181304AFFA3FCB3F9B1BEB90B20" box="[838,986,1571,1595]" pageId="51" pageNumber="252" value="1994-04-18">
<collectingDate id="E66D4869304AFFA3FCB3F9B1BEB90B20" box="[838,986,1571,1595]" pageId="51" pageNumber="252" value="1994-04-18">18 April 1994</collectingDate>
</date>
”/ “
<collectingMethod id="5BD6EF56304AFFA3FBF5F9B1B9190B21" box="[1024,1146,1571,1595]" pageId="51" pageNumber="252">pitfall traps</collectingMethod>
in montane rainforest,
<collectorName id="2F62F297304AFFA3FE96F9D6BF6F0B47" box="[355,524,1604,1628]" pageId="51" pageNumber="252">E. Rajeriarison</collectorName>
&amp;
<collectorName id="2F62F297304AFFA3FDB5F9D6BFA40B47" box="[576,711,1604,1628]" pageId="51" pageNumber="252">R. Malenky</collectorName>
”/ “
<typeStatus id="5D2C29E3304AFFA3FD03F9D6BEE00B47" box="[758,899,1604,1628]" pageId="51" pageNumber="252" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC2304AFFA3FC66F9D6BC020B67" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Assadecma" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="252" phylum="Arthropoda" rank="species" species="basilewskyi" status="sp. nov.">
<emphasis id="B0E34B53304AFFA3FC66F9D6B9F60B47" box="[915,1173,1604,1628]" italics="true" pageId="51" pageNumber="252">Assadecma basilewskyi</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
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2015” [red label]
</materialsCitation>
.
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<typeStatus id="5D2C29E3304AFFA3FD82F9F7BFBC0B66" box="[631,735,1637,1661]" pageId="51" pageNumber="252" type="paratype">Paratypes</typeStatus>
(total of 2):
<specimenCount id="94915CC8304AFFA3FC98F9F6BED40B67" box="[877,951,1636,1660]" count="1" pageId="51" pageNumber="252" type="male">1 male</specimenCount>
(in
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) labeled same as
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(in
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) labeled “CASENT 1051147”/ “
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,
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,
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,
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,
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, mixed tropical forest, 26
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</paragraph>
</subSubSection>
</treatment>
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320
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<mods:title id="135FB5673CCA527C452CA6CBD5AC82F1">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:affiliation id="54C164D6A6CA47E4B835F4ED3E689CE9">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:namePart id="1D74625445B413665EC305C16417A717">Rainio, Johanna</mods:namePart>
<mods:affiliation id="BF93A774DF156E56D7D4FD61D5967467">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
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<emphasis id="B0E34B53304FFFA6FF8EFD0BBDDA0FAA" bold="true" box="[123,185,665,689]" italics="true" pageId="54" pageNumber="255">Lebia</emphasis>
(
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)
<emphasis id="B0E34B53304FFFA6FEB2FD0BBCA80FAA" bold="true" box="[327,459,665,689]" italics="true" pageId="54" pageNumber="255">laterolucida</emphasis>
Kavanaugh and Rainio
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,
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.—
<materialsCitation id="32FF9D1C304FFFA6FE8AFD66B9620E76" collectingDate="1999-12-21" collectorName="M. E. Irwin &amp; E. I. Schlinger" country="Madagascar" elevation="1000" latitude="-21.258333" location="Nacional Ranomafana Bell Vue Trail" longLatPrecision="126" longitude="47.426666" municipality="Parc Nacional Ranomafana Bell Vue" pageId="54" pageNumber="255" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E3304FFFA6FE8AFD66BC810E17" box="[383,482,756,780]" pageId="54" pageNumber="255" type="holotype">Holotype</typeStatus>
(
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), a female, in CAS, labeled: “CASENT 1006164”/
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<collectingRegion id="405359A3304FFFA6FEB6FC86BF6E0E37" box="[323,525,788,812]" country="Madagascar" name="Fianarantsoa" pageId="54" pageNumber="255">Fianarantsoa Prov.</collectingRegion>
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Parc
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Trail, tropical forest
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<collectorName id="2F62F297304FFFA6FE42FCA4BF440E56" box="[439,551,821,845]" pageId="54" pageNumber="255">M.E.Irwin</collectorName>
and
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MEI 99-MA-7
<date id="F629B181304FFFA6FC63FCA7B97E0E56" box="[918,1053,821,845]" pageId="54" pageNumber="255" value="1999-12-21">
<collectingDate id="E66D4869304FFFA6FC63FCA7B97E0E56" box="[918,1053,821,845]" pageId="54" pageNumber="255" value="1999-12-21">21-XII-1999</collectingDate>
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”/ “
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<emphasis id="B0E34B53304FFFA6FF30FCC4BC610E75" box="[197,258,854,878]" italics="true" pageId="54" pageNumber="255">Lebia</emphasis>
(
<emphasis id="B0E34B53304FFFA6FEE7FCC4BC1D0E75" box="[274,382,854,878]" italics="true" pageId="54" pageNumber="255">Metalebia</emphasis>
)
<emphasis id="B0E34B53304FFFA6FE7BFCC4BF6C0E75" box="[398,527,854,878]" italics="true" pageId="54" pageNumber="255">laterolucida</emphasis>
Kavanaugh &amp; Rainio
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2015” [red label].
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<paragraph id="82289741304FFFA6FF5EFCE5BDDD090B" blockId="54.[123,1174,756,1725]" pageId="54" pageNumber="255">
<materialsCitation id="32FF9D1C304FFFA6FF5EFCE5BDD4090B" collectingDate="1998-04-13" collectionCode="CAS" collectorName="M. E. Irwin &amp; E. I. Schlinger" country="Madagascar" elevation="850" latitude="-21.233334" location="Talatakely area" longLatPrecision="1264" longitude="47.366665" municipality="Ranomafana National Park" pageId="54" pageNumber="255" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="5D2C29E3304FFFA6FF5EFCE5BC700E94" box="[171,275,887,911]" pageId="54" pageNumber="255" type="paratype">Paratypes</typeStatus>
(only 1): a female (in
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) labelled “CASENT 1049030”/ “
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,
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,
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,
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,
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,
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,”/ “
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<collectingMethod id="5BD6EF56304FFFA6FF28FC25BC000ED4" box="[221,355,951,975]" pageId="54" pageNumber="255">Malaise trap</collectingMethod>
in mixed tropical forest,
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and
<collectorName id="2F62F297304FFFA6FCD6FC2ABEDB0ED4" box="[803,952,951,975]" pageId="54" pageNumber="255">E.I. Schlinger</collectorName>
collectors, Stop 98- MAD-1”/ “
<typeStatus id="5D2C29E3304FFFA6FF00FC4ABC1A0EEB" box="[245,377,984,1008]" pageId="54" pageNumber="255" type="paratype">PARATYPE</typeStatus>
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<emphasis id="B0E34B53304FFFA6FE75FC4ABCDE0EEB" box="[384,445,984,1008]" italics="true" pageId="54" pageNumber="255">Lebia</emphasis>
(
<emphasis id="B0E34B53304FFFA6FE38FC4ABF5A0EEB" box="[461,569,984,1008]" italics="true" pageId="54" pageNumber="255">Metalebia</emphasis>
)
<emphasis id="B0E34B53304FFFA6FDBCFC4ABFA90EEB" box="[585,714,984,1008]" italics="true" pageId="54" pageNumber="255">laterolucida</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628304FFFA6FC4EFC4BBE960EEB" box="[955,1013,985,1008]" pageId="54" pageNumber="255" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="82289741304FFFA6FF5EFB8BB950092A" blockId="54.[123,1174,756,1725]" box="[171,1075,1049,1073]" pageId="54" pageNumber="255">
<emphasis id="B0E34B53304FFFA6FF5EFB8BBC34092B" bold="true" box="[171,343,1049,1073]" pageId="54" pageNumber="255">
<smallCapsWord id="84CE019D304FFFA6FF5EFB8BBD87092B" baselines="1067,1067" box="[171,228,1049,1073]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="54" pageNumber="255">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D304FFFA6FF19FB8FBC34092B" baselines="1067" box="[236,343,1053,1072]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="54" pageNumber="255">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D1304FFFA6FE8BFB8BBC9C092A" box="[382,511,1049,1073]" name="Madagascar" pageId="54" pageNumber="255">Madagascar</collectingCountry>
,
<collectingRegion id="405359A3304FFFA6FDF8FB8BBF9F092A" box="[525,764,1049,1073]" country="Madagascar" name="Fianarantsoa" pageId="54" pageNumber="255">Fianarantsoa Province</collectingRegion>
, Ranomafana National Park.
</paragraph>
<paragraph id="82289741304FFFA6FF5EFBA8BF5109A8" blockId="54.[123,1174,756,1725]" pageId="54" pageNumber="255">
<emphasis id="B0E34B53304FFFA6FF5EFBA8BC9A094B" bold="true" box="[171,505,1082,1106]" pageId="54" pageNumber="255">
<smallCapsWord id="84CE019D304FFFA6FF5EFBA8BC4D094B" baselines="1100,1099" box="[171,302,1082,1106]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="54" pageNumber="255">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D304FFFA6FECDFBAFBC37094B" baselines="1099" box="[312,340,1085,1104]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="54" pageNumber="255">OF</smallCapsWord>
<smallCapsWord id="84CE019D304FFFA6FEABFBAFBCD2094B" baselines="1099" box="[350,433,1085,1104]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="54" pageNumber="255">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D304FFFA6FE49FBAFBC9A094B" baselines="1099" box="[444,505,1085,1104]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="54" pageNumber="255">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC2304FFFA6FCF2FBA8BEEB0949" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[775,904,1082,1106]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="laterolucida">
<emphasis id="B0E34B53304FFFA6FCF2FBA8BEEB0949" box="[775,904,1082,1106]" italics="true" pageId="54" pageNumber="255">laterolucida</emphasis>
</taxonomicName>
, is an adjective derived from the Latin words,
<emphasis id="B0E34B53304FFFA6FE9FFBC9BCCD0968" box="[362,430,1115,1139]" italics="true" pageId="54" pageNumber="255">lateris</emphasis>
, meaning side, and
<emphasis id="B0E34B53304FFFA6FD71FBC9BFB20968" box="[644,721,1115,1139]" italics="true" pageId="54" pageNumber="255">lucidus</emphasis>
, meaning bright or full of light. The name refers to the pale, translucent lateral pronotal margins, which contrast markedly with the dark pronotal disc in members of this species.
</paragraph>
<paragraph id="82289741304FFFA7FF5EFB2EBC090B0E" blockId="54.[123,1174,756,1725]" lastBlockId="55.[123,1174,1366,1659]" lastPageId="55" lastPageNumber="256" pageId="54" pageNumber="255">
<smallCapsWord id="84CE019D304FFFA6FF5EFB2EBC2109C8" baselines="1230,1230" box="[171,322,1212,1236]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="54" pageNumber="255">
<emphasis id="B0E34B53304FFFA6FF5EFB2EBC2109C8" bold="true" box="[171,322,1212,1236]" pageId="54" pageNumber="255">RECOGNITION</emphasis>
</smallCapsWord>
.— Size moderate for genus, SBL of females = 4.5–5.0 mm. Members of this species (
<figureCitation id="1AAC8BC4304FFFA6FF2CFB4FBC5B09EE" box="[217,312,1245,1269]" captionStart="FIGURE" captionStartId="55.[155,167,1281,1300]" captionTargetBox="[120,1176,233,1259]" captionTargetId="figure-694@55.[120,1176,216,1278]" captionTargetPageId="55" captionText="FIGURE 33. Digital images of holotype female of Lebia (Metalebia) laterolucida Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." pageId="54" pageNumber="255">Fig. 33A</figureCitation>
) key to subgenus
<taxonomicName id="4597ECC2304FFFA6FDF5FB4FBE7009EE" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" box="[512,787,1245,1269]" pageId="54" pageNumber="255" rank="subGenus" subGenus="Metalebia">
<emphasis id="B0E34B53304FFFA6FDF5FB4FBF0F09EE" box="[512,620,1245,1269]" italics="true" pageId="54" pageNumber="255">Metalebia</emphasis>
Jeannel (1949)
</taxonomicName>
and the
<taxonomicName id="4597ECC2304FFFA6FC87FB4FB95009EE" authorityName="Basilewsky" authorityYear="1982" box="[882,1075,1245,1269]" class="Insecta" family="Carabidae" genus="Assadecma" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B0E34B53304FFFA6FC87FB4FB95009EE" box="[882,1075,1245,1269]" italics="true" pageId="54" pageNumber="255">madagascariensis</emphasis>
</taxonomicName>
group in Jeannel’s (1949) key: the basal elytral border is absent medially between the base of stria 3 and the scutellum; the elytral striae are deep and the elytral intervals convex and impunctate; the forebody (head plus pronotum) has no metallic reflection and the elytra are uniform in color; and the frons is slightly convex and without distinct frontal furrows between the eyes. Within the
<taxonomicName id="4597ECC2304FFFA6FBD9FACDBDB80883" authorityName="Basilewsky" authorityYear="1982" class="Insecta" family="Carabidae" genus="Assadecma" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B0E34B53304FFFA6FBD9FACDBDB80883" italics="true" pageId="54" pageNumber="255">madagascariensis</emphasis>
</taxonomicName>
group,
<taxonomicName id="4597ECC2304FFFA6FEC7FA13BCB10883" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[306,466,1408,1432]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="laterolucida">
<emphasis id="B0E34B53304FFFA6FEC7FA13BCB10883" box="[306,466,1408,1432]" italics="true" pageId="54" pageNumber="255">L. laterolucida</emphasis>
</taxonomicName>
members are distinguished from those of
<taxonomicName id="4597ECC2304FFFA6FC51FA13BDA108A2" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="alluaudana">
<emphasis id="B0E34B53304FFFA6FC51FA13B95F0883" box="[932,1084,1408,1432]" italics="true" pageId="54" pageNumber="255">L. alluaudana</emphasis>
Jeannel (1949)
</taxonomicName>
,
<taxonomicName id="4597ECC2304FFFA6FF20FA33BCEE08A2" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[213,397,1441,1465]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="apicoviolacea" status="sp. nov.">
<emphasis id="B0E34B53304FFFA6FF20FA33BCEE08A2" box="[213,397,1441,1465]" italics="true" pageId="54" pageNumber="255">L. apicoviolacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABD0F628304FFFA6FE6CFA33BC9208A3" box="[409,497,1441,1464]" pageId="54" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4597ECC2304FFFA6FDF1FA33BE5708A2" authority="Alluaud (1936 a)" authorityName="Alluaud" authorityYear="1936" box="[516,820,1441,1465]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="mirana">
<emphasis id="B0E34B53304FFFA6FDF1FA33BF1008A3" box="[516,627,1441,1464]" italics="true" pageId="54" pageNumber="255">L. mirana</emphasis>
<bibRefCitation id="E606EAB0304FFFA6FD8BFA33BE5708A2" author="ALLUAUD, C." box="[638,820,1441,1465]" pageId="54" pageNumber="255" pagination="1 - 13" refId="ref28980" refString="ALLUAUD, C. 1936 a. Carabidae recueillis a Madagascar par MM. Syrig, G. Olsoufieff, Vadon, R. Catala, etc. Afra, Cahiers d'Entomologie 11: 1 - 13." type="journal article" year="1936">Alluaud (1936a)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4597ECC2304FFFA6FCB2FA33BE9408A2" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[839,1015,1441,1465]" class="Insecta" family="Carabidae" genus="Pseudomasoreus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B53304FFFA6FCB2FA33BE9408A2" box="[839,1015,1441,1465]" italics="true" pageId="54" pageNumber="255">L. ranomafanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABD0F628304FFFA6FBF1FA33B93F08A3" box="[1028,1116,1441,1464]" pageId="54" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4597ECC2304FFFA6FF8EFA50BCE708C2" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" box="[123,388,1473,1497]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="tanala">
<emphasis id="B0E34B53304FFFA6FF8EFA50BDBF08C2" box="[123,220,1473,1497]" italics="true" pageId="54" pageNumber="255">L. tanala</emphasis>
Jeannel (1949)
</taxonomicName>
by the absence of metallic reflection on the elytra and punctation of the anterior dorsum of the head (both features present in members of the other five species). They differ from members of
<taxonomicName id="4597ECC2304FFFA6FE9FF991BFB60B00" authority="(Fairmaire) (1889)" authorityName="" authorityYear="1889" baseAuthorityName="Fairmaire" baseAuthorityYear="1889" box="[362,725,1538,1563]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="sulcipennis">
<emphasis id="B0E34B53304FFFA6FE9FF991BF620B01" box="[362,513,1538,1562]" italics="true" pageId="54" pageNumber="255">L. sulcipennis</emphasis>
(Fairmaire) (1889)
</taxonomicName>
,
<taxonomicName id="4597ECC2304FFFA6FD10F991BE800B00" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" box="[741,995,1538,1563]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="nana">
<emphasis id="B0E34B53304FFFA6FD10F991BE5A0B01" box="[741,825,1539,1562]" italics="true" pageId="54" pageNumber="255">L. nana</emphasis>
Jeannel (1949)
</taxonomicName>
, and
<taxonomicName id="4597ECC2304FFFA6FBD6F991BC7A0B20" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="B0E34B53304FFFA6FBD6F991B9F60B01" box="[1059,1173,1539,1562]" italics="true" pageId="54" pageNumber="255">L. perrieri</emphasis>
Jeannel (1949)
</taxonomicName>
in having the pronotum broad, ratio PW/PL = at least 1.5, and with lateral margins distinctly rounded at least in the anterior half pronotum (narrower and less rounded anteriorly in members of the three other species). From adults of
<taxonomicName id="4597ECC2304FFFA6FD43F9F7BECB0B67" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" box="[694,936,1636,1660]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="rufa">
<emphasis id="B0E34B53304FFFA6FD43F9F7BE630B67" box="[694,768,1636,1660]" italics="true" pageId="54" pageNumber="255">L. rufa</emphasis>
Jeannel (1949)
</taxonomicName>
, they differ in having the pronotal basal lobe distinctly margined (absent from the former).
<taxonomicName id="4597ECC2304FFFA6FC94F917B9450B86" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[865,1062,1669,1693]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="laterolucida">
<emphasis id="B0E34B53304FFFA6FC94F917B9450B86" box="[865,1062,1669,1693]" italics="true" pageId="54" pageNumber="255">Lebia laterolucida</emphasis>
</taxonomicName>
adults are most similar to those of the two remaining species in Jeannel’s key,
<taxonomicName id="4597ECC2304FFFA7FC77F934BDA10875" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" lastPageId="55" lastPageNumber="256" order="Coleoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="brunneipennis">
<emphasis id="B0E34B53304FFFA6FC77F934B9580BA6" box="[898,1083,1701,1725]" italics="true" pageId="54" pageNumber="255">L. brunneipennis</emphasis>
Jeannel (1949)
</taxonomicName>
and
<taxonomicName id="4597ECC2304EFFA7FF01FAC4BFE40875" authority="Chaudoir (1850)" authorityName="Chaudoir" authorityYear="1850" box="[244,647,1366,1390]" class="Insecta" family="Carabidae" genus="Assadecma" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="256" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B0E34B53304EFFA7FF01FAC4BCB20875" box="[244,465,1366,1390]" italics="true" pageId="55" pageNumber="256">L. madagascariensis</emphasis>
<bibRefCitation id="E606EAB0304EFFA7FE22FAC4BFE40875" author="CHAUDOIR, M." box="[471,647,1366,1390]" pageId="55" pageNumber="256" pagination="3 - 85" refId="ref29725" refString="CHAUDOIR, M. 1850. Memoire sur la famille des Carabiques. 2 e partie. Bulletin de la Societe Imperiale des Naturalistes de Moscou 23: 3 - 85." type="journal article" year="1850">Chaudoir (1850)</bibRefCitation>
</taxonomicName>
, but differ from them in having the dorsum of the head, pronotum and elytra dark piceous, with markedly contrasting pale yellow lateral pronotal margins (dorsum uniformly reddish brown and the lateral pronotal margins not or less markedly contrasting in color in the together two species). In addition, they are distinctly smaller than
<taxonomicName id="4597ECC2304EFFA7FB75FA29BC5F08EF" authorityName="Chaudoir" authorityYear="1850" class="Insecta" family="Carabidae" genus="Assadecma" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="256" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B0E34B53304EFFA7FB75FA29BC5F08EF" italics="true" pageId="55" pageNumber="256">L. madagascariensis</emphasis>
</taxonomicName>
adults and have distinct elytral microsculpture (absent or at least very faint in
<taxonomicName id="4597ECC2304EFFA7FB75FA4FBC770B0D" authorityName="Jeannel" authorityYear="1949" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="256" phylum="Arthropoda" rank="species" species="brunneipennis">
<emphasis id="B0E34B53304EFFA7FB75FA4FBC770B0D" italics="true" pageId="55" pageNumber="256">L. brunneipennis</emphasis>
</taxonomicName>
adults).
</paragraph>
<caption id="D6E8C7C9304EFFA7FF6EFA93BF4A0835" pageId="55" pageNumber="256" startId="55.[155,167,1281,1300]" targetBox="[120,1176,233,1259]" targetPageId="55" targetType="figure">
<paragraph id="82289741304EFFA7FF6EFA93BF4A0835" blockId="55.[123,1174,1281,1326]" pageId="55" pageNumber="256">
<smallCapsWord id="84CE019D304EFFA7FF6EFA93BDBB0809" baselines="1295,1294" box="[155,216,1281,1300]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="55" pageNumber="256">FIGURE</smallCapsWord>
33. Digital images of holotype female of
<taxonomicName id="4597ECC2304EFFA7FDC8FA93B963080F" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[573,1024,1281,1300]" class="Insecta" family="Carabidae" genus="Lebia" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="256" phylum="Arthropoda" rank="species" species="laterolucida" status="sp. nov." subGenus="Metalebia">
<emphasis id="B0E34B53304EFFA7FDC8FA93BF0D080F" box="[573,622,1281,1300]" italics="true" pageId="55" pageNumber="256">Lebia</emphasis>
(
<emphasis id="B0E34B53304EFFA7FD8FFA93BFB3080F" box="[634,720,1281,1300]" italics="true" pageId="55" pageNumber="256">Metalebia</emphasis>
)
<emphasis id="B0E34B53304EFFA7FD28FA93BE27080F" box="[733,836,1281,1300]" italics="true" pageId="55" pageNumber="256">laterolucida</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628304EFFA7FBF0FA93B925080F" box="[1029,1094,1281,1300]" pageId="55" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm.
</paragraph>
</caption>
<paragraph id="82289741304EFFA7FF5EF98DB9690B2C" blockId="55.[123,1174,1366,1659]" box="[171,1034,1567,1592]" pageId="55" pageNumber="256">
<emphasis id="B0E34B53304EFFA7FF5EF98DBC920B2D" bold="true" box="[171,497,1567,1591]" pageId="55" pageNumber="256">
<smallCapsWord id="84CE019D304EFFA7FF5EF98DBC350B2D" baselines="1585,1585" box="[171,342,1567,1591]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="55" pageNumber="256">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="84CE019D304EFFA7FEA8F9B1BC920B2D" baselines="1585" box="[349,497,1571,1590]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="55" pageNumber="256">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E3304EFFA7FC8AF9B2BECE0B23" box="[895,941,1568,1592]" pageId="55" pageNumber="256">type</typeStatus>
locality.
</paragraph>
<paragraph id="82289741304EFFA7FF5EF9D3BFBD0B61" blockId="55.[123,1174,1366,1659]" pageId="55" pageNumber="256">
<emphasis id="B0E34B53304EFFA7FF5EF9D3BCC70B4C" bold="true" box="[171,420,1601,1625]" pageId="55" pageNumber="256">
<smallCapsWord id="84CE019D304EFFA7FF5EF9D3BC6A0B4C" baselines="1619,1618" box="[171,265,1601,1625]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="55" pageNumber="256">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D304EFFA7FEE5F9D6BCC70B4C" baselines="1618" box="[272,420,1604,1623]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="55" pageNumber="256">DISTRIBUTION</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C304EFFA7FE39F9D3BFB40B61" location="Talatakely area" pageId="55" pageNumber="256" specimenCount="1">
Both specimens of the
<typeStatus id="5D2C29E3304EFFA7FD37F9D3BF930B42" box="[706,752,1601,1625]" pageId="55" pageNumber="256">type</typeStatus>
series were collected in mixed secondary tropical forest in the
<location id="8748C19A304EFFA7FE71F9F0BF450B61" LSID="urn:lsid:plazi:treatment:0A3E2657304FFFA7FF8EFD0BBFBD0B61:8748C19A304EFFA7FE71F9F0BF450B61" box="[388,550,1634,1658]" name="Talatakely area" pageId="55" pageNumber="256">Talatakely area</location>
in malaise traps
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="76A9D5CF0758D49945DC0D0A8B7C55FD" ID-ISSN="0068-547X" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E26573051FFBAFF8EF999BC920B24" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Eurydera ocellata Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="1" lastPageNumber="243" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="241" updateTime="1726771413727" updateUser="felipe" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3A1C43D703CC2FFD33DC6DB70395B7BD">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:namePart id="6038DD71C598F50BA3D2C17CF0BD610B">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="E30C30709CE1EB218B4BAF66957BA582">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:namePart id="21CD27957CCC3D4F18D4F2F9F926CD2E">Rainio, Johanna</mods:namePart>
<mods:affiliation id="47D992999A9F1AE6E3E232C8EA59E3B6">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
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<mods:title id="A7E82F5B76D1C5CE99C0321350FE53FE">Proceedings of the California Academy of Sciences</mods:title>
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Kavanaugh and Rainio
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R.
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of Sciences malaise, secondary tropical forest MA-02-09C-16”/ “
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Kavanaugh &amp; Rainio
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W
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23. Digital images of holotype male of
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A. Habitus, dorsal aspect; B. Label. Scale line = 1.0 mm.
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</emphasis>
.—
<collectingCountry id="FA80D7D13050FFB9FE8BFA46BC9C08F7" box="[382,511,1492,1516]" name="Madagascar" pageId="41" pageNumber="242">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33050FFB9FDF8FA46BF9F08F7" box="[525,764,1492,1516]" country="Madagascar" name="Fianarantsoa" pageId="41" pageNumber="242">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3050FFB9FCFFFA46B94F08F7" LSID="urn:lsid:plazi:treatment:0A3E26573051FFBAFF8EF999BC920B24:8748C19A3050FFB9FCFFFA46B94F08F7" box="[778,1068,1492,1516]" country="Madagascar" name="Ranomafana National Park" pageId="41" pageNumber="242" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413050FFB9FF5EFA65BDBE0B6C" blockId="41.[123,1174,1318,1726]" pageId="41" pageNumber="242">
<emphasis id="B0E34B533050FFB9FF5EFA65BC8E0B16" bold="true" box="[171,493,1527,1551]" pageId="41" pageNumber="242">
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<smallCapsWord id="84CE019D3050FFB9FEC1FA68BC330B16" baselines="1544" box="[308,336,1530,1549]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="41" pageNumber="242">OF</smallCapsWord>
<smallCapsWord id="84CE019D3050FFB9FEA3FA68BCCA0B16" baselines="1544" box="[342,425,1530,1549]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="41" pageNumber="242">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3050FFB9FE45FA68BC8E0B16" baselines="1544" box="[432,493,1530,1549]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="41" pageNumber="242">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23050FFB9FD05FA65BE250B14" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[752,838,1527,1551]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="242" phylum="Arthropoda" rank="species" species="ocellata">
<emphasis id="B0E34B533050FFB9FD05FA65BE250B14" box="[752,838,1527,1551]" italics="true" pageId="41" pageNumber="242">ocellata</emphasis>
</taxonomicName>
, is the Latin adjective meaning having little eyes or marked with spots. The name refers to the small, faint dark spot on interval 3 within the subapical pale area of each elytron at the insertion point of the subapical discal setiferous pore.
</paragraph>
<paragraph id="822897413050FFBAFF5EF911BF670884" blockId="41.[123,1174,1318,1726]" lastBlockId="42.[123,1174,807,1599]" lastPageId="42" lastPageNumber="243" pageId="41" pageNumber="242">
<smallCapsWord id="84CE019D3050FFB9FF5EF911BC210B82" baselines="1685,1684" box="[171,322,1667,1691]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="41" pageNumber="242">
<emphasis id="B0E34B533050FFB9FF5EF911BC210B82" bold="true" box="[171,322,1667,1691]" pageId="41" pageNumber="242">RECOGNITION</emphasis>
</smallCapsWord>
.— Size small for genus, SBL of male and female =
<quantity id="456F3AA43050FFB9FC75F911BEB00B81" box="[896,979,1667,1691]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="41" pageNumber="242" unit="mm" value="9.2">9.2 mm</quantity>
. Members of this species (
<figureCitation id="1AAC8BC43050FFB9FF22F937BC560BA5" box="[215,309,1701,1726]" captionStart="FIGURE" captionStartId="41.[155,167,1239,1258]" captionTargetBox="[128,1176,230,1218]" captionTargetId="figure-897@41.[120,1176,216,1236]" captionTargetPageId="41" captionText="FIGURE 23. Digital images of holotype male of Eurydera ocellata Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Label. Scale line = 1.0 mm." pageId="41" pageNumber="242">Fig. 23A</figureCitation>
) key to
<taxonomicName id="4597ECC23050FFB9FE78F937BE330BA5" authority="Coquerel (1851)" authorityName="Coquerel" authorityYear="1851" box="[397,848,1701,1726]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="242" phylum="Arthropoda" rank="species" species="mormolycoides">
<emphasis id="B0E34B533050FFB9FE78F937BFF80BA6" box="[397,667,1701,1725]" italics="true" pageId="41" pageNumber="242">Eurydera mormolycoides</emphasis>
<bibRefCitation id="E606EAB03050FFB9FD54F937BE330BA5" author="COQUEREL, C." box="[673,848,1701,1726]" pageId="41" pageNumber="242" pagination="86 - 90" refId="ref29865" refString="COQUEREL, C. 1851. Coleopteres nouveaux de Madagascar. Reuue et Magasin de Zoologie Pure et Appliquee. (2 e Serie) 3: 86 - 90." type="journal article" year="1851">Coquerel (1851)</bibRefCitation>
</taxonomicName>
in Jeannel’s (1949) key on the basis of the following combination of features: size small; head (
<figureCitation id="1AAC8BC43053FFBAFCB2FCB5BECB0E5B" box="[839,936,807,832]" captionStart="FIGURE" captionStartId="42.[155,167,729,748]" captionTargetBox="[120,1168,229,707]" captionTargetId="figure-2005@42.[120,1176,216,707]" captionTargetPageId="42" captionText="FIGURE 24. Digital images of holotype male of Eurydera ocellata Kavanaugh &amp; Rainio sp. nov. A. Head, dorsal aspect; B–C. Aedeagus of male genitalia; B. Left lateral aspect; C. Dorsal aspect. Scale lines = 0.5 mm." pageId="42" pageNumber="243">Fig. 24A</figureCitation>
) with frontal furrows deeply impressed, smooth, divergent posteriorly and without a transverse impression uniting them, posterior part of frons smooth, narrow and triangular, without lateral grooves prolonged posteriorly beyond the posterior border of the eyes; mandibles short and broad, lateral border not broadly explanate, lateral margin only slightly convex; pronotum cordate, distinctly narrowed basally, lateral margins angulate at insertions of midlateral setae; elytra short, round and black, each with a reddish subapical spot tangential to the medial suture as well as subhumeral and subapicolateral reddish areas and reddish apical spines, elytral intervals moderately convex; median lobe of male genitalia without a large, angulate projection on the ventral margin of the shaft near the base. They differ from
<taxonomicName id="4597ECC23053FFBAFF09FBDABCDC0944" authorityName="Coquerel" authorityYear="1851" box="[252,447,1095,1119]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="mormolycoides">
<emphasis id="B0E34B533053FFBAFF09FBDABCDC0944" box="[252,447,1095,1119]" italics="true" pageId="42" pageNumber="243">E. mormolycoides</emphasis>
</taxonomicName>
members in having: dorsal surface slightly shiny (dull in
<taxonomicName id="4597ECC23053FFBAFBB4FBDABD900964" authorityName="Coquerel" authorityYear="1851" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="mormolycoides">
<emphasis id="B0E34B533053FFBAFBB4FBDABD900964" italics="true" pageId="42" pageNumber="243">E. mormolycoides</emphasis>
</taxonomicName>
members); pronotum with lateral explanations pale testaceous (darker in
<taxonomicName id="4597ECC23053FFBAFBFAFBFABDDC0984" authorityName="Coquerel" authorityYear="1851" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="mormolycoides">
<emphasis id="B0E34B533053FFBAFBFAFBFABDDC0984" italics="true" pageId="42" pageNumber="243">E. mormolycoides</emphasis>
</taxonomicName>
members), anterior angles broadly rounded (more narrowly rounded in
<taxonomicName id="4597ECC23053FFBAFC26FB1AB9F60984" authorityName="Coquerel" authorityYear="1851" box="[979,1173,1159,1183]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="mormolycoides">
<emphasis id="B0E34B533053FFBAFC26FB1AB9F60984" box="[979,1173,1159,1183]" italics="true" pageId="42" pageNumber="243">E. mormolycoides</emphasis>
</taxonomicName>
members) and lateral margins with shallow but distinct sinuation anterior to hind angles (sinuation absent from or less distinct in
<taxonomicName id="4597ECC23053FFBAFE4EFB5ABF1F09C4" authorityName="Coquerel" authorityYear="1851" box="[443,636,1223,1247]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="mormolycoides">
<emphasis id="B0E34B533053FFBAFE4EFB5ABF1F09C4" box="[443,636,1223,1247]" italics="true" pageId="42" pageNumber="243">E. mormolycoides</emphasis>
</taxonomicName>
members); elytra with larger but less distinct (less contrasting with rest of elytra) pale subapical macula with black eye spot on interval three (macula extended further laterally and lateroanteriorly than in
<taxonomicName id="4597ECC23053FFBAFD13FA9ABECA0804" authorityName="Coquerel" authorityYear="1851" box="[742,937,1287,1311]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="mormolycoides">
<emphasis id="B0E34B533053FFBAFD13FA9ABECA0804" box="[742,937,1287,1311]" italics="true" pageId="42" pageNumber="243">E. mormolycoides</emphasis>
</taxonomicName>
members) and apical spine about half as long as spine in
<taxonomicName id="4597ECC23053FFBAFDFDFABABFA90824" authorityName="Coquerel" authorityYear="1851" box="[520,714,1319,1343]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="mormolycoides">
<emphasis id="B0E34B533053FFBAFDFDFABABFA90824" box="[520,714,1319,1343]" italics="true" pageId="42" pageNumber="243">E. mormolycoides</emphasis>
</taxonomicName>
members. Finally, the median lobe of the male genitalia (
<figureCitation id="1AAC8BC43053FFBAFED4FAD5BCCA087B" box="[289,425,1351,1376]" captionStart="FIGURE" captionStartId="42.[155,167,729,748]" captionTargetBox="[120,1168,229,707]" captionTargetId="figure-2005@42.[120,1176,216,707]" captionTargetPageId="42" captionText="FIGURE 24. Digital images of holotype male of Eurydera ocellata Kavanaugh &amp; Rainio sp. nov. A. Head, dorsal aspect; B–C. Aedeagus of male genitalia; B. Left lateral aspect; C. Dorsal aspect. Scale lines = 0.5 mm." pageId="42" pageNumber="243">Figs. 24B–C</figureCitation>
) has a thicker shaft throughout and an apex shorter and thicker in lateral view (
<figureCitation id="1AAC8BC43053FFBAFF19FAF5BC2A0864" box="[236,329,1383,1408]" captionStart="FIGURE" captionStartId="42.[155,167,729,748]" captionTargetBox="[120,1168,229,707]" captionTargetId="figure-2005@42.[120,1176,216,707]" captionTargetPageId="42" captionText="FIGURE 24. Digital images of holotype male of Eurydera ocellata Kavanaugh &amp; Rainio sp. nov. A. Head, dorsal aspect; B–C. Aedeagus of male genitalia; B. Left lateral aspect; C. Dorsal aspect. Scale lines = 0.5 mm." pageId="42" pageNumber="243">Fig. 24B</figureCitation>
) and broader and straighter in dorsal view (
<figureCitation id="1AAC8BC43053FFBAFCEFFAF5BE14089B" box="[794,887,1383,1408]" captionStart="FIGURE" captionStartId="42.[155,167,729,748]" captionTargetBox="[120,1168,229,707]" captionTargetId="figure-2005@42.[120,1176,216,707]" captionTargetPageId="42" captionText="FIGURE 24. Digital images of holotype male of Eurydera ocellata Kavanaugh &amp; Rainio sp. nov. A. Head, dorsal aspect; B–C. Aedeagus of male genitalia; B. Left lateral aspect; C. Dorsal aspect. Scale lines = 0.5 mm." pageId="42" pageNumber="243">Fig. 24C</figureCitation>
) than in
<taxonomicName id="4597ECC23053FFBAFC20FAFAB9F60864" authorityName="Coquerel" authorityYear="1851" box="[981,1173,1383,1407]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="mormolycoides">
<emphasis id="B0E34B533053FFBAFC20FAFAB9F60864" box="[981,1173,1383,1407]" italics="true" pageId="42" pageNumber="243">E. mormolycoides</emphasis>
</taxonomicName>
males (see Jeannel 1949, Fig. 492b).
</paragraph>
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24. Digital images of holotype male of
<taxonomicName id="4597ECC23053FFBAFDDBFD4ABEE00FF0" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[558,899,728,747]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="ocellata" status="sp. nov.">
<emphasis id="B0E34B533053FFBAFDDBFD4ABFAA0FF0" box="[558,713,728,747]" italics="true" pageId="42" pageNumber="243">Eurydera ocellata</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283053FFBAFC7CFD4BBEA90FF7" box="[905,970,729,748]" pageId="42" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
A. Head, dorsal aspect;
</paragraph>
<paragraph id="822897413053FFBAFF8EFD60BED70E1E" blockId="42.[123,1173,728,773]" box="[123,948,754,773]" pageId="42" pageNumber="243">B–C. Aedeagus of male genitalia; B. Left lateral aspect; C. Dorsal aspect. Scale lines = 0.5 mm.</paragraph>
</caption>
<paragraph id="822897413053FFBAFF5EFA35B96908A4" blockId="42.[123,1174,807,1599]" box="[171,1034,1447,1472]" pageId="42" pageNumber="243">
<emphasis id="B0E34B533053FFBAFF5EFA35BC9208A5" bold="true" box="[171,497,1447,1471]" pageId="42" pageNumber="243">
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<smallCapsWord id="84CE019D3053FFBAFEA8FA39BC9208A5" baselines="1465" box="[349,497,1451,1470]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="42" pageNumber="243">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E33053FFBAFC8AFA3ABECE08DB" box="[895,941,1448,1472]" pageId="42" pageNumber="243">type</typeStatus>
locality.
</paragraph>
<paragraph id="822897413053FFBAFF5EFA5ABC920B24" blockId="42.[123,1174,807,1599]" pageId="42" pageNumber="243">
<emphasis id="B0E34B533053FFBAFF5EFA5ABCC608C5" bold="true" box="[171,421,1480,1504]" pageId="42" pageNumber="243">
<smallCapsWord id="84CE019D3053FFBAFF5EFA5ABC6A08C5" baselines="1498,1497" box="[171,265,1480,1504]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="42" pageNumber="243">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D3053FFBAFEE4FA59BCC608C5" baselines="1497" box="[273,421,1483,1502]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="42" pageNumber="243">DISTRIBUTION</smallCapsWord>
</emphasis>
.— The
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specimen was collected in a malaise trap in secondary montane rainforest in the Talatakely area at an elevation of
<quantity id="456F3AA43053FFBAFCE1FA7ABE0708E4" box="[788,868,1512,1536]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="42" pageNumber="243" unit="m" value="1030.0">1030 m</quantity>
. The
<typeStatus id="5D2C29E33053FFBAFC55FA7ABE9F0B1B" box="[928,1020,1512,1536]" pageId="42" pageNumber="243" type="paratype">paratype</typeStatus>
specimen was collected at ultraviolet light in secondary montane rainforest near the junction of the Namorona River and a small tributary stream.
</paragraph>
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346
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<heading id="D960202D3057FFBEFF8EFF49BFD90DE8" box="[123,698,219,243]" fontSize="10" level="2" pageId="46" pageNumber="247" reason="4">
<emphasis id="B0E34B533057FFBEFF8EFF49BFD90DE8" bold="true" box="[123,698,219,243]" pageId="46" pageNumber="247">
<taxonomicName id="4597ECC23057FFBEFF8EFF49BF350DE8" ID-CoL="3D44Z" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,598,219,243]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="247" phylum="Arthropoda" rank="species" species="simplica" status="sp. nov.">
<emphasis id="B0E34B533057FFBEFF8EFF49BC250DE8" bold="true" box="[123,326,219,243]" italics="true" pageId="46" pageNumber="247">Eurydera simplica</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F6283057FFBEFD91FF49BFD90DE8" box="[612,698,219,243]" pageId="46" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3057FFBEFF8EFF6FBE3A0C7E" pageId="46" pageNumber="247" type="description">
<paragraph id="822897413057FFBEFF8EFF6FBC660C08" blockId="46.[123,698,219,275]" box="[123,261,253,275]" pageId="46" pageNumber="247">
<figureCitation id="1AAC8BC43057FFBEFF8EFF6FBC660C08" box="[123,261,253,275]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStartId-0="46.[155,167,1667,1686]" captionStartId-1="47.[155,167,783,802]" captionTargetBox-0="[149,1176,526,1636]" captionTargetBox-1="[120,1164,230,765]" captionTargetId-0="figure-537@46.[120,1176,528,1664]" captionTargetId-1="figure-2124@47.[120,1176,216,765]" captionTargetPageId-0="46" captionTargetPageId-1="47" captionText-0="FIGURE 27. Digital images of holotype male of Eurydera simplica Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." captionText-1="FIGURE 28. Digital images of holotype male of Eurydera simplica Kavanaugh &amp; Rainio sp. nov. A. Head, dorsal aspect; B–C. Median lobe of aedeagus of genitalia; B. Left lateral aspect; C. Dorsal aspect. Scale lines = 0.5 mm." pageId="46" pageNumber="247">Figures 27–28</figureCitation>
</paragraph>
<paragraph id="822897413057FFBEFF8EFEBFBE710C58" blockId="46.[123,857,301,358]" box="[123,786,301,323]" pageId="46" pageNumber="247">
<taxonomicName id="4597ECC23057FFBEFF8EFEBFBD940C58" box="[123,247,301,323]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="247" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B0E34B533057FFBEFF8EFEBFBD940C58" box="[123,247,301,323]" italics="true" pageId="46" pageNumber="247">Eurydera sp.</emphasis>
</taxonomicName>
; Rainio and Niemelä 2006: 227 (informal designation).
</paragraph>
<paragraph id="822897413057FFBEFF8EFEC2BE3A0C7E" blockId="46.[123,857,301,358]" box="[123,857,336,358]" pageId="46" pageNumber="247">
<emphasis id="B0E34B533057FFBEFF8EFEC2BC410C7E" box="[123,290,336,357]" italics="true" pageId="46" pageNumber="247">
<taxonomicName id="4597ECC23057FFBEFF8EFEC2BDB50C7E" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,214,336,357]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="247" phylum="Arthropoda" rank="genus" status="n. sp.">Eurydera</taxonomicName>
<taxonomicNameLabel id="ABD0F6283057FFBEFF29FEC2BC730C7E" box="[220,272,336,357]" pageId="46" pageNumber="247" rank="species">n. sp.</taxonomicNameLabel>
3
</emphasis>
; Rainio 2009: 31, Rainio 2012: 72 (informal designation).
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3057FFA0FF5EFE1CBC7D0FEE" lastPageId="48" lastPageNumber="249" pageId="46" pageNumber="247" type="materials_examined">
<paragraph id="822897413057FFBFFF5EFE1CBFAA0ECC" blockId="46.[123,1174,398,526]" lastBlockId="47.[123,1174,864,1719]" lastPageId="47" lastPageNumber="248" pageId="46" pageNumber="247">
<emphasis id="B0E34B533057FFBEFF5EFE1CBC3D0CBF" bold="true" box="[171,350,398,422]" pageId="46" pageNumber="247">
<smallCapsWord id="84CE019D3057FFBEFF5EFE1CBD870CBF" baselines="416,415" box="[171,228,398,422]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="46" pageNumber="247">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D3057FFBEFF04FE03BC3D0CBF" baselines="415" box="[241,350,401,420]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="46" pageNumber="247">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C3057FFBEFE73FE1CBDB00F15" collectingDate="2004-03-08" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="900" location="Talatakely" pageId="46" pageNumber="247" specimenCount="1" typeStatus="holotype">
<typeStatus id="5D2C29E33057FFBEFE73FE1CBC8A0CBD" box="[390,489,398,422]" pageId="46" pageNumber="247" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43057FFBEFE0BFE1CBFE30CBE" box="[510,640,398,422]" captionStart="FIGURE" captionStartId="46.[155,167,1667,1686]" captionTargetBox="[149,1176,526,1636]" captionTargetId="figure-537@46.[120,1176,528,1664]" captionTargetPageId="46" captionText="FIGURE 27. Digital images of holotype male of Eurydera simplica Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." pageId="46" pageNumber="247">Figs.27A–B</figureCitation>
), a male, in CAS, labeled: “
<collectingCountry id="FA80D7D13057FFBEFC24FE1CB9EC0CBD" box="[977,1167,398,422]" name="Madagascar" pageId="46" pageNumber="247">MADAGASCAR</collectingCountry>
, Ranomafana NP.,
<location id="8748C19A3057FFBEFEC1FE22BCC20CD3" LSID="urn:lsid:plazi:treatment:0A3E26573057FFA0FF8EFF49BC7D0FEE:8748C19A3057FFBEFEC1FE22BCC20CD3" box="[308,417,432,456]" country="Madagascar" name="Talatakely" pageId="46" pageNumber="247">Talatakely</location>
,
<quantity id="456F3AA43057FFBEFE58FE22BC930CD3" box="[429,496,432,456]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="46" pageNumber="247" unit="m" value="900.0">
<elevation id="09BA70723057FFBEFE58FE22BC930CD3" box="[429,496,432,456]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="46" pageNumber="247" unit="m" value="900.0">900 m</elevation>
</quantity>
, Trail SP at
<quantity id="456F3AA43057FFBEFD85FE22BFC60CD3" box="[624,677,432,456]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="247" unit="m" value="50.0">
<elevation id="09BA70723057FFBEFD85FE22BFC60CD3" box="[624,677,432,456]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="247" unit="m" value="50.0">50 m</elevation>
</quantity>
mark,
<date id="F629B1813057FFBEFD1BFE22BE050CD3" box="[750,870,432,456]" pageId="46" pageNumber="247" value="2004-03-08">
<collectingDate id="E66D48693057FFBEFD1BFE22BE050CD3" box="[750,870,432,456]" pageId="46" pageNumber="247" value="2004-03-08">08.03.2004</collectingDate>
</date>
”/ “collected by hand in daytime in secondary montane rainforest,
<collectorName id="2F62F2973057FFBEFDD9FE41BFC10CF0" box="[556,674,467,491]" pageId="46" pageNumber="247">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973057FFBEFD38FE41BE520CF0" box="[717,817,467,491]" pageId="46" pageNumber="247">J. Rainio</collectorName>
collectors”/ “106” [handwritten label]/ “
</materialsCitation>
<materialsCitation id="32FF9D1C3057FFBFFF26FE64BE120EAC" collectingDate="2000-06-15" collectionCode="MNHN" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="900" lastPageId="47" lastPageNumber="248" location="Talatakely" pageId="46" pageNumber="247" specimenCount="2" typeStatus="holotype">
<typeStatus id="5D2C29E33057FFBEFF26FE64BC030F15" box="[211,352,502,526]" pageId="46" pageNumber="247" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23057FFBEFE92FE67BE7F0F16" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[359,796,501,526]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="247" phylum="Arthropoda" rank="species" species="simplica" status="sp. nov.">
<emphasis id="B0E34B533057FFBEFE92FE67BF4D0F16" box="[359,558,501,525]" italics="true" pageId="46" pageNumber="247">Eurydera simplica</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283057FFBEFCD1FE64BE3D0F16" box="[804,862,502,525]" pageId="46" pageNumber="247" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
<typeStatus id="5D2C29E33057FFBEFBDBFE64B9F50F15" box="[1070,1174,502,526]" pageId="46" pageNumber="247" type="paratype">Paratypes</typeStatus>
(only 1): a female (in
<collectionCode id="E4860F843056FFBFFE98FCF3BCDD0E63" box="[365,446,865,888]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="47" pageNumber="248">MNHN</collectionCode>
) labeled ““
<collectingCountry id="FA80D7D13056FFBFFDC8FCF3BF980E62" box="[573,763,865,889]" name="Madagascar" pageId="47" pageNumber="248">MADAGASCAR</collectingCountry>
, Ranomafana NP.,
<location id="8748C19A3056FFBFFC38FCF2B9590E63" LSID="urn:lsid:plazi:treatment:0A3E26573057FFA0FF8EFF49BC7D0FEE:8748C19A3056FFBFFC38FCF2B9590E63" box="[973,1082,864,888]" country="Madagascar" name="Talatakely" pageId="47" pageNumber="248">Talatakely</location>
,
<quantity id="456F3AA43056FFBFFBBCFCF3B9EC0E63" box="[1097,1167,865,889]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="47" pageNumber="248" unit="m" value="900.0">
<elevation id="09BA70723056FFBFFBBCFCF3B9EC0E63" box="[1097,1167,865,889]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="47" pageNumber="248" unit="m" value="900.0">900 m</elevation>
</quantity>
, Trail B at
<quantity id="456F3AA43056FFBFFF1EFCEDBC5D0E8D" box="[235,318,895,919]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="248" unit="m" value="1000.0">
<elevation id="09BA70723056FFBFFF1EFCEDBC5D0E8D" box="[235,318,895,919]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="248" unit="m" value="1000.0">1000 m</elevation>
</quantity>
mark,
<date id="F629B1813056FFBFFE79FCEDBF600E8C" box="[396,515,895,919]" pageId="47" pageNumber="248" value="2000-06-15">
<collectingDate id="E66D48693056FFBFFE79FCEDBF600E8C" box="[396,515,895,919]" pageId="47" pageNumber="248" value="2000-06-15">15.06.2000</collectingDate>
</date>
”/ “collected by hand in daytime in secondary montane rainforest,
<collectorName id="2F62F2973056FFBFFF30FC32BC560EAC" box="[197,309,927,951]" pageId="47" pageNumber="248">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973056FFBFFEADFC32BCD50EAC" box="[344,438,927,951]" pageId="47" pageNumber="248">J. Rainio</collectorName>
collectors”/ “7/13” [handwritten label]/ “
</materialsCitation>
<typeStatus id="5D2C29E33056FFBFFC84FC0DBE960EAC" box="[881,1013,927,951]" pageId="47" pageNumber="248" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC23056FFBFFC09FC0DBCF80ECC" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="species" species="simplica" status="sp. nov.">
<emphasis id="B0E34B533056FFBFFC09FC0DBDD30ECC" italics="true" pageId="47" pageNumber="248">Eurydera simplica</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283056FFBFFE56FC52BCBE0ECC" box="[419,477,960,983]" pageId="47" pageNumber="248" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label].
</paragraph>
<caption id="D6E8C7C93057FFBEFF6EF911BCA80BAB" pageId="46" pageNumber="247" startId="46.[155,167,1667,1686]" targetBox="[149,1176,526,1636]" targetPageId="46" targetType="figure">
<paragraph id="822897413057FFBEFF6EF911BCA80BAB" blockId="46.[123,1173,1666,1712]" pageId="46" pageNumber="247">
<smallCapsWord id="84CE019D3057FFBEFF6EF911BDBB0B8F" baselines="1681,1680" box="[155,216,1667,1686]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="46" pageNumber="247">FIGURE</smallCapsWord>
27. Digital images of holotype male of
<taxonomicName id="4597ECC23057FFBEFDB5F910BEC00B8E" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[576,931,1666,1686]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="247" phylum="Arthropoda" rank="species" species="simplica" status="sp. nov.">
<emphasis id="B0E34B533057FFBEFDB5F910BF820B8E" box="[576,737,1666,1685]" italics="true" pageId="46" pageNumber="247">Eurydera simplica</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283057FFBEFC5EF911BE8D0B8D" box="[939,1006,1667,1686]" pageId="46" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm.
</paragraph>
</caption>
<caption id="D6E8C7C93056FFBFFF6EFC9DB96A0E27" pageId="47" pageNumber="248" startId="47.[155,167,783,802]" targetBox="[120,1164,230,765]" targetPageId="47" targetType="figure">
<paragraph id="822897413056FFBFFF6EFC9DB9F60E39" blockId="47.[123,1173,783,828]" box="[155,1173,783,802]" pageId="47" pageNumber="248">
<smallCapsWord id="84CE019D3056FFBFFF6EFC9DBDBB0E3A" baselines="797,797" box="[155,216,783,802]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="47" pageNumber="248">FIGURE</smallCapsWord>
28. Digital images of holotype male of
<taxonomicName id="4597ECC23056FFBFFDD9FC9DBEE60E39" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[556,901,783,802]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="species" species="simplica" status="sp. nov.">
<emphasis id="B0E34B533056FFBFFDD9FC9DBFA90E39" box="[556,714,783,802]" italics="true" pageId="47" pageNumber="248">Eurydera simplica</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283056FFBFFC7FFC9DBEA80E39" box="[906,971,783,802]" pageId="47" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
A. Head, dorsal aspect;
</paragraph>
<paragraph id="822897413056FFBFFF8EFCBBB96A0E27" blockId="47.[123,1173,783,828]" box="[123,1033,808,828]" pageId="47" pageNumber="248">B–C. Median lobe of aedeagus of genitalia; B. Left lateral aspect; C. Dorsal aspect. Scale lines = 0.5 mm.</paragraph>
</caption>
<paragraph id="822897413056FFBFFF5EFC4DB9500EED" blockId="47.[123,1174,864,1719]" box="[171,1075,991,1015]" pageId="47" pageNumber="248">
<materialsCitation id="32FF9D1C3056FFBFFF5EFC4DB94F0EEC" box="[171,1068,991,1015]" country="Madagascar" location="Ranomafana National Park" pageId="47" pageNumber="248" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533056FFBFFF5EFC4DBC340EEE" bold="true" box="[171,343,991,1015]" pageId="47" pageNumber="248">
<smallCapsWord id="84CE019D3056FFBFFF5EFC4DBD870EED" baselines="1009,1009" box="[171,228,991,1015]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="47" pageNumber="248">
<typeStatus id="5D2C29E33056FFBFFF5EFC4DBD870EED" box="[171,228,991,1015]" pageId="47" pageNumber="248">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3056FFBFFF19FC70BC340EEE" baselines="1008" box="[236,343,994,1013]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="47" pageNumber="248">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13056FFBFFE8BFC4DBC9C0EEC" box="[382,511,991,1015]" name="Madagascar" pageId="47" pageNumber="248">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33056FFBFFDF8FC4DBF9F0EEC" box="[525,764,991,1015]" country="Madagascar" name="Fianarantsoa" pageId="47" pageNumber="248">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3056FFBFFCFFFC4DB94F0EEC" LSID="urn:lsid:plazi:treatment:0A3E26573057FFA0FF8EFF49BC7D0FEE:8748C19A3056FFBFFCFFFC4DB94F0EEC" box="[778,1068,991,1015]" country="Madagascar" name="Ranomafana National Park" pageId="47" pageNumber="248" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413056FFBFFF5EFC6DBDD0096C" blockId="47.[123,1174,864,1719]" pageId="47" pageNumber="248">
<emphasis id="B0E34B533056FFBFFF5EFC6DBC90090E" bold="true" box="[171,499,1023,1047]" pageId="47" pageNumber="248">
<smallCapsWord id="84CE019D3056FFBFFF5EFC6DBC4D090E" baselines="1041,1040" box="[171,302,1023,1047]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="47" pageNumber="248">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3056FFBFFEC3FB90BC31090E" baselines="1040" box="[310,338,1026,1045]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="47" pageNumber="248">OF</smallCapsWord>
<smallCapsWord id="84CE019D3056FFBFFEAFFB90BCCE090E" baselines="1040" box="[346,429,1026,1045]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="47" pageNumber="248">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3056FFBFFE43FB90BC90090E" baselines="1040" box="[438,499,1026,1045]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="47" pageNumber="248">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23056FFBFFD0FFC6DBE37090C" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[762,852,1023,1047]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="species" species="simplica">
<emphasis id="B0E34B533056FFBFFD0FFC6DBE37090C" box="[762,852,1023,1047]" italics="true" pageId="47" pageNumber="248">simplica</emphasis>
</taxonomicName>
, is an adjective derived from the Latin word,
<emphasis id="B0E34B533056FFBFFEDEFB8DBC1F092C" box="[299,380,1055,1079]" italics="true" pageId="47" pageNumber="248">simplex</emphasis>
, meaning simple. The name refers to the relative simplicity of form and surface macrosculpture of member of this species compared with those of other species of
<taxonomicName id="4597ECC23056FFBFFBACFBD2BDCF096C" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533056FFBFFBACFBD2BDCF096C" italics="true" pageId="47" pageNumber="248">Eurydera</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="822897413056FFA0FF5EFBEDB9F50F6F" blockId="47.[123,1174,864,1719]" lastBlockId="48.[123,1174,219,758]" lastPageId="48" lastPageNumber="249" pageId="47" pageNumber="248">
<smallCapsWord id="84CE019D3056FFBFFF5EFBEDBC21098E" baselines="1169,1168" box="[171,322,1151,1175]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="47" pageNumber="248">
<emphasis id="B0E34B533056FFBFFF5EFBEDBC21098E" bold="true" box="[171,322,1151,1175]" pageId="47" pageNumber="248">RECOGNITION</emphasis>
</smallCapsWord>
.— Size average for genus, SBL of male =
<quantity id="456F3AA43056FFBFFCFCFBEDBE04098D" box="[777,871,1151,1175]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="47" pageNumber="248" unit="mm" value="13.1">13.1 mm</quantity>
, of female = 12.4. Members of this species (
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) are unlike those of any other species in the genus. In Jeannel’s (1949) key species of
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<emphasis id="B0E34B533056FFBFFEE9FB2DBCE209CC" box="[284,385,1215,1239]" italics="true" pageId="47" pageNumber="248">Eurydera</emphasis>
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, they key to his
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<emphasis id="B0E34B533056FFBFFDC2FB2DBFF209CC" box="[567,657,1215,1239]" italics="true" pageId="47" pageNumber="248">unicolor</emphasis>
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species group, based on the following features: size medium; head (
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) with posterior part of frons formed as a broad, flat, smooth, elevat- ed plateau, laterally delimited by deep furrows extended posteriorly beyond the level of the posterior margin of the eye then slightly and arcuately convergent posteriorly and terminated abruptly at the level of the posterior margin of the ocular swelling and just less than halfway from the midline to the lateral margin of the head, area between the furrow smooth and only slightly elevated; mandibles moderately short and broad; median lobe of male genitalia without a large, angulate projection on the ventral margin of the shaft near the base. Jeannel included only
<taxonomicName id="4597ECC23056FFBFFC3BFA0DBC6208CC" authority="(Klug 1833)" baseAuthorityName="Klug" baseAuthorityYear="1833" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B0E34B533056FFBFFC3BFA0DB9F608AC" box="[974,1173,1439,1463]" italics="true" pageId="47" pageNumber="248">Eurydera unicolor</emphasis>
(
<bibRefCitation id="E606EAB03056FFBFFF71FA2DBD9B08CC" author="KLUG, J. C. F." box="[132,248,1471,1495]" pageId="47" pageNumber="248" pagination="91 - 223" refId="ref30967" refString="KLUG, J. C. F. 1833. Bericht uber eine auf Madagascar veranstaltete. Sammlung von Insecten aus der Ordnung Coleoptera. Abhandlungen der Koniglichen Preussischen Akademie der Wissenschaften 1832 - 33: 91 - 223 + 5 pls." type="book chapter" year="1833">Klug 1833</bibRefCitation>
)
</taxonomicName>
in this species group, but
<bibRefCitation id="E606EAB03056FFBFFDDBFA52BFA708CC" author="MATEU, J." box="[558,708,1471,1495]" pageId="47" pageNumber="248" pagination="845 - 855" refId="ref31792" refString="MATEU, J. 1973. Notes sur les Lebiens malagaches (Col. Caraboidea) (3 e note). Le genre Eurydera Castelnau. Bulletin du Museum National d'Histoire Naturelle (Ser. 3) No. 139, Zoologie 103: 845 - 855." type="journal article" year="1973">Mateu (1973)</bibRefCitation>
added another species,
<taxonomicName id="4597ECC23056FFBFFC33FA2DBDDC08ED" authority="Mateu" authorityName="Mateu" authorityYear="1973" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="species" species="fossulata">
<emphasis id="B0E34B533056FFBFFC33FA2DB9F608CC" box="[966,1173,1471,1495]" italics="true" pageId="47" pageNumber="248">Eurydera fossulata</emphasis>
Mateu
</taxonomicName>
to the group, in part based on the shared lack of sexual dimorphism in the front tarsi in members of these two species. Unlike males of most
<taxonomicName id="4597ECC23056FFBFFD75FA6DBE5B0B0C" box="[640,824,1535,1559]" pageId="47" pageNumber="248">
<emphasis id="B0E34B533056FFBFFD75FA6DBF860B0C" box="[640,741,1535,1559]" italics="true" pageId="47" pageNumber="248">Eurydera</emphasis>
species
</taxonomicName>
, those of
<taxonomicName id="4597ECC23056FFBFFC56F992B9790B0C" baseAuthorityName="Klug" baseAuthorityYear="1833" box="[931,1050,1535,1559]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B0E34B533056FFBFFC56F992B9790B0C" box="[931,1050,1535,1559]" italics="true" pageId="47" pageNumber="248">E. unicolor</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC23056FFBFFBBAF992BDDF0B2C" authorityName="Mateu" authorityYear="1973" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="species" species="fossulata">
<emphasis id="B0E34B533056FFBFFBBAF992BDDF0B2C" italics="true" pageId="47" pageNumber="248">E. fossulata</emphasis>
</taxonomicName>
have tarsomeres 1 to 3 no wider than in females and without ventral adhesive setae. Members of
<taxonomicName id="4597ECC23056FFBFFF25F9D2BC290B4C" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[208,330,1599,1623]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="248" phylum="Arthropoda" rank="species" species="simplica">
<emphasis id="B0E34B533056FFBFFF25F9D2BC290B4C" box="[208,330,1599,1623]" italics="true" pageId="47" pageNumber="248">E. simplica</emphasis>
</taxonomicName>
are easily distinguished from those of these two species, as well as from all other described
<taxonomicName id="4597ECC23056FFBFFED3F9CDBCBC0B6C" box="[294,479,1631,1655]" pageId="47" pageNumber="248">
<emphasis id="B0E34B533056FFBFFED3F9CDBCE80B6C" box="[294,395,1631,1655]" italics="true" pageId="47" pageNumber="248">Eurydera</emphasis>
species
</taxonomicName>
, by the uniquely shaped pronotum (
<figureCitation id="1AAC8BC43056FFBFFC90F9CDBEA70B6C" box="[869,964,1631,1655]" captionStart="FIGURE" captionStartId="46.[155,167,1667,1686]" captionTargetBox="[149,1176,526,1636]" captionTargetId="figure-537@46.[120,1176,528,1664]" captionTargetPageId="46" captionText="FIGURE 27. Digital images of holotype male of Eurydera simplica Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." pageId="47" pageNumber="248">Fig. 27A</figureCitation>
), which is relatively small and short for the genus, with the disc smooth, lateral explanation very narrow, lateral margins with very short and shallow sinuation anterior to markedly obtuse hind angles, and basal margin obliquely angulate laterally. Unlike males of
<taxonomicName id="4597ECC23049FFA0FD72FF4EBF9C0DE8" baseAuthorityName="Klug" baseAuthorityYear="1833" box="[647,767,219,243]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B0E34B533049FFA0FD72FF4EBF9C0DE8" box="[647,767,219,243]" italics="true" pageId="48" pageNumber="249">E. unicolor</emphasis>
</taxonomicName>
and
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<emphasis id="B0E34B533049FFA0FCC1FF4EBED10DE8" box="[820,946,219,243]" italics="true" pageId="48" pageNumber="249">E. fossulata</emphasis>
</taxonomicName>
, those of
<taxonomicName id="4597ECC23049FFA0FBE8FF4EB9F60DE8" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[1053,1173,219,243]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="simplica">
<emphasis id="B0E34B533049FFA0FBE8FF4EB9F60DE8" box="[1053,1173,219,243]" italics="true" pageId="48" pageNumber="249">E. simplica</emphasis>
</taxonomicName>
have front tarsi with tarsomeres 1 to 3 wider than in females and with at least a few adhesive setae ventrally, which may call into question a close relationship with these two these species. Members of
<taxonomicName id="4597ECC23049FFA0FF6FFEAEBC770C48" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[154,276,315,339]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="simplica">
<emphasis id="B0E34B533049FFA0FF6FFEAEBC770C48" box="[154,276,315,339]" italics="true" pageId="48" pageNumber="249">E. simplica</emphasis>
</taxonomicName>
also differ from those of
<taxonomicName id="4597ECC23049FFA0FDC4FEAEBFC80C48" baseAuthorityName="Klug" baseAuthorityYear="1833" box="[561,683,315,339]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B0E34B533049FFA0FDC4FEAEBFC80C48" box="[561,683,315,339]" italics="true" pageId="48" pageNumber="249">E. unicolor</emphasis>
</taxonomicName>
in having the pronotum with anterior angles much less projected anteriorly. They differ from members of
<taxonomicName id="4597ECC23049FFA0FCE2FECEBEF50C6F" authorityName="Mateu" authorityYear="1973" box="[791,918,348,372]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="fossulata">
<emphasis id="B0E34B533049FFA0FCE2FECEBEF50C6F" box="[791,918,348,372]" italics="true" pageId="48" pageNumber="249">E. fossulata</emphasis>
</taxonomicName>
in having the posterior part of the head, between the lateral furrows, smooth and slightly convex, whereas
<taxonomicName id="4597ECC23049FFA0FBE0FEEFB9F60C8F" authorityName="Mateu" authorityYear="1973" box="[1045,1173,380,404]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="fossulata">
<emphasis id="B0E34B533049FFA0FBE0FEEFB9F60C8F" box="[1045,1173,380,404]" italics="true" pageId="48" pageNumber="249">E. fossulata</emphasis>
</taxonomicName>
members have a deep, transverse medial depression, the bottom of which is flat and covered with deep transverse creases, on that part of the head. They also have the elytral intervals without the roughened surfaces typical of members of several species in genus, including
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<emphasis id="B0E34B533049FFA0FC49FE4FB9590CEF" box="[956,1082,476,500]" italics="true" pageId="48" pageNumber="249">E. fossulata</emphasis>
</taxonomicName>
(at least subapically), and the sutural spines are very long and straight (similar to those in
<taxonomicName id="4597ECC23049FFA0FC1BFE6FB9040F0F" baseAuthorityName="Klug" baseAuthorityYear="1833" box="[1006,1127,508,532]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B0E34B533049FFA0FC1BFE6FB9040F0F" box="[1006,1127,508,532]" italics="true" pageId="48" pageNumber="249">E. unicolor</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC23049FFA0FF8EFD8FBD990F2E" authorityName="Mateu" authorityYear="1973" box="[123,250,541,565]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="fossulata">
<emphasis id="B0E34B533049FFA0FF8EFD8FBD990F2E" box="[123,250,541,565]" italics="true" pageId="48" pageNumber="249">E. fossulata</emphasis>
</taxonomicName>
members). Finally, the median lobe of male genitalia of the
<typeStatus id="5D2C29E33049FFA0FC63FD8FBE900F2E" box="[918,1011,541,565]" pageId="48" pageNumber="249" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4597ECC23049FFA0FBEEFD8FB9F60F2E" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[1051,1173,541,565]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="simplica">
<emphasis id="B0E34B533049FFA0FBEEFD8FB9F60F2E" box="[1051,1173,541,565]" italics="true" pageId="48" pageNumber="249">E. simplica</emphasis>
</taxonomicName>
(
<figureCitation id="1AAC8BC43049FFA0FF71FDAFBC620F4E" box="[132,257,573,597]" captionStart="FIGURE" captionStartId="47.[155,167,783,802]" captionTargetBox="[120,1164,230,765]" captionTargetId="figure-2124@47.[120,1176,216,765]" captionTargetPageId="47" captionText="FIGURE 28. Digital images of holotype male of Eurydera simplica Kavanaugh &amp; Rainio sp. nov. A. Head, dorsal aspect; B–C. Median lobe of aedeagus of genitalia; B. Left lateral aspect; C. Dorsal aspect. Scale lines = 0.5 mm." pageId="48" pageNumber="249">Fig. 28B–C</figureCitation>
) has the shaft much narrower basally and the apical orifice shorter than in
<emphasis id="B0E34B533049FFA0FBDEFDAFBDF00F6E" italics="true" pageId="48" pageNumber="249">E. unicol- or</emphasis>
males (see Jeannel 1949, Fig. 495). No male of
<taxonomicName id="4597ECC23049FFA0FD6EFDCCBE7A0F6E" authorityName="Mateu" authorityYear="1973" box="[667,793,605,629]" class="Insecta" family="Carabidae" genus="Eurydera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="fossulata">
<emphasis id="B0E34B533049FFA0FD6EFDCCBE7A0F6E" box="[667,793,605,629]" italics="true" pageId="48" pageNumber="249">E. fossulata</emphasis>
</taxonomicName>
has been available for comparison.
</paragraph>
<paragraph id="822897413049FFA0FF5EFDEFB9690F8E" blockId="48.[123,1174,219,758]" box="[171,1034,637,662]" pageId="48" pageNumber="249">
<emphasis id="B0E34B533049FFA0FF5EFDEFBC920F88" bold="true" box="[171,497,637,661]" pageId="48" pageNumber="249">
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<smallCapsWord id="84CE019D3049FFA0FEA8FD12BC920F88" baselines="654" box="[349,497,640,659]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="48" pageNumber="249">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
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locality.
</paragraph>
<paragraph id="822897413049FFA0FF5EFD0FBC7D0FEE" blockId="48.[123,1174,219,758]" pageId="48" pageNumber="249">
<emphasis id="B0E34B533049FFA0FF5EFD0FBCC60FAF" bold="true" box="[171,421,669,693]" pageId="48" pageNumber="249">
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<smallCapsWord id="84CE019D3049FFA0FEE4FD33BCC60FAF" baselines="687" box="[273,421,673,692]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="48" pageNumber="249">DISTRIBUTION</smallCapsWord>
</emphasis>
.— The two known specimens of this species both were collected by hand in daytime in secondary montane rainforest in the Talatakely area at elevations ranging from
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.
</paragraph>
</subSubSection>
</treatment>
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<mods:classification id="51F8CBFEA6B4B2C7448BB8DB23D43B5E">journal article</mods:classification>
<mods:identifier id="1D492D76A7D2B05272A96A644844637D" type="ISSN">0068-547X</mods:identifier>
</mods:mods>
<treatment id="0A3E26573058FFB3FF8EFCB7B95E09C5" LSID="urn:lsid:plazi:treatment:0A3E26573058FFB3FF8EFCB7B95E09C5" httpUri="http://treatment.plazi.org/id/0A3E26573058FFB3FF8EFCB7B95E09C5" lastPageId="35" lastPageNumber="236" pageId="33" pageNumber="234">
<subSubSection id="CA8DC4CA3058FFB1FF8EFCB7BE620E26" box="[123,769,805,829]" pageId="33" pageNumber="234" type="nomenclature">
<paragraph id="822897413058FFB1FF8EFCB7BE620E26" blockId="33.[123,769,805,859]" box="[123,769,805,829]" pageId="33" pageNumber="234">
<heading id="D960202D3058FFB1FF8EFCB7BE620E26" box="[123,769,805,829]" fontSize="10" level="2" pageId="33" pageNumber="234" reason="4">
<emphasis id="B0E34B533058FFB1FF8EFCB7BE620E26" bold="true" box="[123,769,805,829]" pageId="33" pageNumber="234">
<taxonomicName id="4597ECC23058FFB1FF8EFCB7BFFD0E26" ID-CoL="6QRM4" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,670,805,829]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="bimaculata" status="sp. nov.">
<emphasis id="B0E34B533058FFB1FF8EFCB7BCEE0E26" bold="true" box="[123,397,805,829]" italics="true" pageId="33" pageNumber="234">Madecassina bimaculata</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F6283058FFB1FD5EFCB7BE620E26" box="[683,769,805,829]" pageId="33" pageNumber="234" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3058FFB1FF8EFCD7BDBB0E40" box="[123,216,837,859]" pageId="33" pageNumber="234" type="description">
<paragraph id="822897413058FFB1FF8EFCD7BDBB0E40" blockId="33.[123,769,805,859]" box="[123,216,837,859]" pageId="33" pageNumber="234">
<figureCitation id="1AAC8BC43058FFB1FF8EFCD7BDBB0E40" box="[123,216,837,859]" captionStart="FIGURE" captionStartId="34.[155,167,1377,1396]" captionTargetBox="[120,1176,216,1355]" captionTargetId="figure-856@34.[120,1176,216,1374]" captionTargetPageId="34" captionText="FIGURE 19. Digital images of holotype male of Madecassina bimaculata Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels C–D. Median lobe of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." pageId="33" pageNumber="234">Figure 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3058FFB3FF5EFCEDB95E09C5" lastPageId="35" lastPageNumber="236" pageId="33" pageNumber="234" type="materials_examined">
<paragraph id="822897413058FFB3FF5EFCEDBC8C0CCE" blockId="33.[123,1174,895,1726]" lastBlockId="35.[123,1174,219,1247]" lastPageId="35" lastPageNumber="236" pageId="33" pageNumber="234">
<emphasis id="B0E34B533058FFB1FF5EFCEDBC390E8E" bold="true" box="[171,346,895,919]" pageId="33" pageNumber="234">
<smallCapsWord id="84CE019D3058FFB1FF5EFCEDBD870E8D" baselines="913,913" box="[171,228,895,919]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="33" pageNumber="234">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D3058FFB1FF18FC10BC390E8E" baselines="912" box="[237,346,898,917]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="33" pageNumber="234">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C3058FFB1FE77FCEDBE53090C" collectingDate="1998-04-13" collectorName="D. H. Kavanaugh" country="Madagascar" elevation="900" latitude="-21.248333" location="Talatakely area" longLatPrecision="126" longitude="47.426666" municipality="Ranomafana National Park" pageId="33" pageNumber="234" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E33058FFB1FE77FCEDBC860E8C" box="[386,485,895,919]" pageId="33" pageNumber="234" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43058FFB1FE02FCEDBF1A0E8D" box="[503,633,895,919]" captionStart="FIGURE" captionStartId="34.[155,167,1377,1396]" captionTargetBox="[120,1176,216,1355]" captionTargetId="figure-856@34.[120,1176,216,1374]" captionTargetPageId="34" captionText="FIGURE 19. Digital images of holotype male of Madecassina bimaculata Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels C–D. Median lobe of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." pageId="33" pageNumber="234">Figs.19A–B</figureCitation>
), a male, in CAS, labeled: “CASENT 1049022”/ “
<collectingCountry id="FA80D7D13058FFB1FF72FC0DBC260EAC" box="[135,325,927,951]" name="Madagascar" pageId="33" pageNumber="234">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A33058FFB1FEA0FC0DBF240EAC" box="[341,583,927,951]" country="Madagascar" name="Fianarantsoa" pageId="33" pageNumber="234">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B3058FFB1FDADFC0DBEE30EAC" box="[600,896,927,951]" pageId="33" pageNumber="234">Ranomafana National Park</collectingMunicipality>
,
<location id="8748C19A3058FFB1FC64FC0DB9550EAC" LSID="urn:lsid:plazi:treatment:0A3E26573058FFB3FF8EFCB7B95E09C5:8748C19A3058FFB1FC64FC0DB9550EAC" box="[913,1078,927,951]" country="Madagascar" latitude="-21.248333" longLatPrecision="126" longitude="47.426666" municipality="Ranomafana National Park" name="Talatakely area" pageId="33" pageNumber="234" stateProvince="Fianarantsoa">Talatakely area</location>
,
<quantity id="456F3AA43058FFB1FBB2FC0DB9EC0EAC" box="[1095,1167,927,951]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="234" unit="m" value="900.0">
<elevation id="09BA70723058FFB1FBB2FC0DB9EC0EAC" box="[1095,1167,927,951]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="234" unit="m" value="900.0">900 m</elevation>
</quantity>
, mixed tropical forest,
<date id="F629B1813058FFB1FE93FC52BC990EC3" box="[358,506,959,984]" pageId="33" pageNumber="234" value="1998-04-13">
<collectingDate id="E66D48693058FFB1FE93FC52BC990EC3" box="[358,506,959,984]" pageId="33" pageNumber="234" value="1998-04-13">13 April 1998</collectingDate>
</date>
,
<geoCoordinate id="E7A3F1863058FFB1FDFDFC52BF100EC3" box="[520,627,960,984]" degrees="21" direction="south" minutes="14. 9" orientation="latitude" pageId="33" pageNumber="234" precision="92" value="-21.248333">21°14.9’S</geoCoordinate>
<geoCoordinate id="E7A3F1863058FFB1FD8FFC52BF840ECC" box="[634,743,959,984]" degrees="47" direction="east" minutes="25. 6" orientation="longitude" pageId="33" pageNumber="234" precision="92" value="47.426666">47°25.6’E</geoCoordinate>
,”/ “Stop # 98-29,
<collectorName id="2F62F2973058FFB1FC5EFC52B9060ECC" box="[939,1125,959,983]" pageId="33" pageNumber="234">D. H. Kavanaugh</collectorName>
collector collected by beating suspended clusters of dead leaves and twigs”/ “
<typeStatus id="5D2C29E33058FFB1FC4CFC72B9250EE3" box="[953,1094,992,1016]" pageId="33" pageNumber="234" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23058FFB1FBA5FC72BF500903" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="bimaculata" status="sp. nov.">
<emphasis id="B0E34B533058FFB1FBA5FC72BC2A0903" italics="true" pageId="33" pageNumber="234">Madecassina bimaculata</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283058FFB1FDCCFB92BF10090C" box="[569,627,1024,1047]" pageId="33" pageNumber="234" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label]
</materialsCitation>
.
<materialsCitation id="32FF9D1C3058FFB1FCC8FB92BC1909C1" collectingDate="2001-01-02" collectingDateMax="2001-01-22" collectingDateMin="2001-01-02" collectionCode="CAS" collectorName="D. H. &amp; K. M Kavanaugh &amp; R. L. Brett &amp; E. Elsom &amp; F. Vargas &amp; R. Ranaivosolo &amp; E. F. Randrianirina &amp; N. Rasoamananana &amp; T. J. Ravelomanana &amp; H. C. Raveloson" country="Madagascar" elevation="900" latitude="-21.25041" location="Talatakely area" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" pageId="33" pageNumber="234" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="5D2C29E33058FFB1FCC8FB92BEC60903" box="[829,933,1024,1048]" pageId="33" pageNumber="234" type="paratype">Paratypes</typeStatus>
(total of 16):
<specimenCount id="94915CC83058FFB1FBCCFB92B9F50903" box="[1081,1174,1024,1048]" count="1" pageId="33" pageNumber="234" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843058FFB1FF57FBB2BDB50923" box="[162,214,1056,1080]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="33" pageNumber="234" type="Museum">CAS</collectionCode>
) labeled “CASENT 1004001”/ “
<collectingCountry id="FA80D7D13058FFB1FDB5FBB2BF9D0923" box="[576,766,1056,1080]" name="Madagascar" pageId="33" pageNumber="234">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A33058FFB1FCFBFBB2BE9C0923" box="[782,1023,1056,1080]" country="Madagascar" name="Fianarantsoa" pageId="33" pageNumber="234">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B3058FFB1FBFAFBB2BC730943" pageId="33" pageNumber="234">Ranomafana National Park</collectingMunicipality>
,
<location id="8748C19A3058FFB1FEEAFBD2BCA10943" LSID="urn:lsid:plazi:treatment:0A3E26573058FFB3FF8EFCB7B95E09C5:8748C19A3058FFB1FEEAFBD2BCA10943" box="[287,450,1088,1112]" country="Madagascar" latitude="-21.25041" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" name="Talatakely area" pageId="33" pageNumber="234" stateProvince="Fianarantsoa">Talatakely area</location>
,
<quantity id="456F3AA43058FFB1FE27FBD3BF740943" box="[466,535,1089,1113]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="234" unit="m" value="900.0">
<elevation id="09BA70723058FFB1FE27FBD3BF740943" box="[466,535,1089,1113]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="234" unit="m" value="900.0">900 m</elevation>
</quantity>
, mixed tropical forest,
<date id="F629B1813058FFB1FCEDFBD3BE810942" box="[792,994,1089,1113]" pageId="33" pageNumber="234" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">
<collectingDate id="E66D48693058FFB1FCEDFBD3BE810942" box="[792,994,1089,1113]" pageId="33" pageNumber="234" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">2-22 January 2001</collectingDate>
</date>
,”/ “
<geoCoordinate id="E7A3F1863058FFB1FBE5FBD3B9ED0942" box="[1040,1166,1088,1113]" degrees="21. 25041" direction="south" orientation="latitude" pageId="33" pageNumber="234" precision="1" value="-21.25041">21.25041°S</geoCoordinate>
/
<geoCoordinate id="E7A3F1863058FFB1FF8EFBF3BD9A0963" box="[123,249,1121,1145]" degrees="47. 41945" direction="east" orientation="longitude" pageId="33" pageNumber="234" precision="1" value="47.41945">47.41945°E</geoCoordinate>
, Stop# DHK-01-001,
<collectorName id="2F62F2973058FFB1FE0FFBF3BE700962" box="[506,787,1121,1145]" pageId="33" pageNumber="234">D.H. &amp; K.M Kavanaugh</collectorName>
,
<collectorName id="2F62F2973058FFB1FCD2FBF3BEFB0963" box="[807,920,1121,1144]" pageId="33" pageNumber="234">R.L. Brett</collectorName>
,
<collectorName id="2F62F2973058FFB1FC59FBF3B9700962" box="[940,1043,1121,1145]" pageId="33" pageNumber="234">E. Elsom</collectorName>
,
<collectorName id="2F62F2973058FFB1FBD2FBF3B9EC0962" box="[1063,1167,1121,1145]" pageId="33" pageNumber="234">F. Vargas</collectorName>
,
<collectorName id="2F62F2973058FFB1FF8EFB13BC430982" box="[123,288,1153,1177]" pageId="33" pageNumber="234">R. Ranaivosolo</collectorName>
,”/ “
<collectorName id="2F62F2973058FFB1FEBBFB13BF700982" box="[334,531,1153,1177]" pageId="33" pageNumber="234">E.F. Randrianirina</collectorName>
,
<collectorName id="2F62F2973058FFB1FDD4FB13BF930983" box="[545,752,1153,1176]" pageId="33" pageNumber="234">N. Rasoamananana</collectorName>
,
<collectorName id="2F62F2973058FFB1FD0BFB13BEA90982" box="[766,970,1153,1177]" pageId="33" pageNumber="234">T.J. Ravelomanana</collectorName>
, and
<collectorName id="2F62F2973058FFB1FBF2FB13BDC309A2" pageId="33" pageNumber="234">H.C. Raveloson</collectorName>
collectors”/ “collected in daytime by beating live understory vegetation and/or dead leaves and other suspended debris”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3058FFB1FE7DFB50B95C0801" collectingDate="1998-11-10" collectingDateMax="1998-11-16" collectingDateMin="1998-11-10" collectionCode="CAS" collectorName="V. F. Lee &amp; K. J. Ribardo" country="Madagascar" elevation="900" latitude="-21.248333" location="Trail" longLatPrecision="126" longitude="47.426666" municipality="Ranomafana National Park" pageId="33" pageNumber="234" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="94915CC83058FFB1FE7DFB50BC8709C2" box="[392,484,1217,1242]" count="1" pageId="33" pageNumber="234" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843058FFB1FDFBFB50BF2109C1" box="[526,578,1218,1242]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="33" pageNumber="234" type="Museum">CAS</collectionCode>
) labeled “CASENT 1049023”/ “
<collectingCountry id="FA80D7D13058FFB1FC55FB50B93D09C1" box="[928,1118,1218,1242]" name="Madagascar" pageId="33" pageNumber="234">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A33058FFB1FB9FFB53BC2A09E1" country="Madagascar" name="Fianarantsoa" pageId="33" pageNumber="234">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B3058FFB1FEADFB70BF1E09E1" box="[344,637,1250,1274]" pageId="33" pageNumber="234">Ranomafana National Park</collectingMunicipality>
, Talatakely area,
<quantity id="456F3AA43058FFB1FCB5FB70BEE509E2" box="[832,902,1250,1274]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="234" unit="m" value="900.0">
<elevation id="09BA70723058FFB1FCB5FB70BEE509E2" box="[832,902,1250,1274]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="234" unit="m" value="900.0">900 m</elevation>
</quantity>
,
<location id="8748C19A3058FFB1FC60FB70BEAB09E1" LSID="urn:lsid:plazi:treatment:0A3E26573058FFB3FF8EFCB7B95E09C5:8748C19A3058FFB1FC60FB70BEAB09E1" box="[917,968,1250,1274]" country="Madagascar" latitude="-21.248333" longLatPrecision="126" longitude="47.426666" municipality="Ranomafana National Park" name="Trail" pageId="33" pageNumber="234" stateProvince="Fianarantsoa">Trail</location>
FF,
<geoCoordinate id="E7A3F1863058FFB1FC0EFB70B90409E1" box="[1019,1127,1250,1274]" degrees="21" direction="south" minutes="14. 9" orientation="latitude" pageId="33" pageNumber="234" precision="92" value="-21.248333">21°14.9’S</geoCoordinate>
<geoCoordinate id="E7A3F1863058FFB1FB85FB70BDA10801" degrees="47" direction="east" minutes="25. 6" orientation="longitude" pageId="33" pageNumber="234" precision="92" value="47.426666">47° 25.6’E</geoCoordinate>
,”/ “
<date id="F629B1813058FFB1FF1AFA90BC820801" box="[239,481,1282,1306]" pageId="33" pageNumber="234" value="1998-11-10" valueMax="1998-11-16" valueMin="1998-11-10">
<collectingDate id="E66D48693058FFB1FF1AFA90BC820801" box="[239,481,1282,1306]" pageId="33" pageNumber="234" value="1998-11-10" valueMax="1998-11-16" valueMin="1998-11-10">10-16 November 1998</collectingDate>
</date>
, beaten from vines,
<collectorName id="2F62F2973058FFB1FD49FA91BE7E0801" box="[700,797,1283,1306]" pageId="33" pageNumber="234">V. F. Lee</collectorName>
&amp;
<collectorName id="2F62F2973058FFB1FCCAFA91BEAC0801" box="[831,975,1282,1306]" pageId="33" pageNumber="234">K. J. Ribardo</collectorName>
collector”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3058FFB1FBB8FA90B9ED0860" collectingDate="1998-04-24" collectionCode="CAS" collectorName="D. H. Kavanaugh" country="Madagascar" elevation="900" latitude="-21.255" location="Talatakely area" longLatPrecision="126" longitude="47.431667" municipality="Ranomafana" pageId="33" pageNumber="234" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="94915CC83058FFB1FBB8FA90B9F50801" box="[1101,1174,1282,1306]" count="1" pageId="33" pageNumber="234" type="male">1 male</specimenCount>
(in
<collectionCode id="E4860F843058FFB1FF55FAB0BDB70821" box="[160,212,1314,1338]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="33" pageNumber="234" type="Museum">CAS</collectionCode>
) labeled “CASENT 1049024”/ “
<collectingCountry id="FA80D7D13058FFB1FDCCFAB0BF940821" box="[569,759,1314,1338]" name="Madagascar" pageId="33" pageNumber="234">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A33058FFB1FCF0FAB0BE970821" box="[773,1012,1314,1338]" country="Madagascar" name="Fianarantsoa" pageId="33" pageNumber="234">Fianarantsoa Province</collectingRegion>
, Parc National
<collectingMunicipality id="624C0D3B3058FFB1FF8EFAD0BC610841" box="[123,258,1346,1370]" pageId="33" pageNumber="234">Ranomafana</collectingMunicipality>
,
<location id="8748C19A3058FFB1FEECFAD0BCA70841" LSID="urn:lsid:plazi:treatment:0A3E26573058FFB3FF8EFCB7B95E09C5:8748C19A3058FFB1FEECFAD0BCA70841" box="[281,452,1346,1370]" country="Madagascar" latitude="-21.255" longLatPrecision="126" longitude="47.431667" municipality="Ranomafana" name="Talatakely area" pageId="33" pageNumber="234" stateProvince="Fianarantsoa">Talatakely area</location>
,
<quantity id="456F3AA43058FFB1FE29FAD1BF490841" box="[476,554,1347,1371]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="234" unit="m" value="900.0">
<elevation id="09BA70723058FFB1FE29FAD1BF490841" box="[476,554,1347,1371]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="234" unit="m" value="900.0">900 m</elevation>
</quantity>
, mixed tropical forest,
<date id="F629B1813058FFB1FCBEFAD1BE900840" box="[843,1011,1346,1371]" pageId="33" pageNumber="234" value="1998-04-24">
<collectingDate id="E66D48693058FFB1FCBEFAD1BE900840" box="[843,1011,1346,1371]" pageId="33" pageNumber="234" value="1998-04-24">24 April 1998</collectingDate>
</date>
,”/ “
<geoCoordinate id="E7A3F1863058FFB1FBDFFAD1B9F50840" box="[1066,1174,1346,1371]" degrees="21" direction="south" minutes="15. 3" orientation="latitude" pageId="33" pageNumber="234" precision="92" value="-21.255">21°15.3’S</geoCoordinate>
<geoCoordinate id="E7A3F1863058FFB1FF8EFAF1BD8B0861" box="[123,232,1379,1403]" degrees="47" direction="east" minutes="25. 9" orientation="longitude" pageId="33" pageNumber="234" precision="92" value="47.431667">47°25.9’E</geoCoordinate>
,”/ “Stop # 98-76,
<collectorName id="2F62F2973058FFB1FE5BFAF1BF090860" box="[430,618,1379,1403]" pageId="33" pageNumber="234">D. H. Kavanaugh</collectorName>
collector collected at night on standing tree trunk”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3058FFB1FF8EFA11BCB108E7" collectingDate="2003-03-09" collectingDateMax="2003-03-20" collectingDateMin="2003-03-09" collectionCode="CAS" collectorName="R. Harin'Hala California Academy" country="Madagascar" elevation="1130" latitude="-21.250834" location="Ranomafana National Park" longLatPrecision="12" longitude="47.407166" municipality="Province" pageId="33" pageNumber="234" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="94915CC83058FFB1FF8EFA11BDAF0880" box="[123,204,1411,1435]" count="1" pageId="33" pageNumber="234" type="male">1 male</specimenCount>
(in
<collectionCode id="E4860F843058FFB1FEFCFA11BC5E0880" box="[265,317,1411,1435]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="33" pageNumber="234" type="Museum">CAS</collectionCode>
) labeled “CASENT 8068817”/ “
<collectingCountry id="FA80D7D13058FFB1FD36FA11BEE20880" box="[707,897,1411,1435]" name="Madagascar" pageId="33" pageNumber="234">MADAGASCAR</collectingCountry>
:
<collectingMunicipality id="624C0D3B3058FFB1FC6DFA11BE940880" box="[920,1015,1411,1435]" pageId="33" pageNumber="234">Province</collectingMunicipality>
<collectingRegion id="405359A33058FFB1FBF3FA11B9ED0880" box="[1030,1166,1411,1435]" country="Madagascar" name="Fianarantsoa" pageId="33" pageNumber="234">Fianarantsoa</collectingRegion>
,
<location id="8748C19A3058FFB1FF8EFA31BCCD08A0" LSID="urn:lsid:plazi:treatment:0A3E26573058FFB3FF8EFCB7B95E09C5:8748C19A3058FFB1FF8EFA31BCCD08A0" box="[123,430,1443,1467]" country="Madagascar" latitude="-21.250834" longLatPrecision="12" longitude="47.407166" municipality="Province" name="Ranomafana National Park" pageId="33" pageNumber="234" stateProvince="Fianarantsoa">Ranomafana National Park</location>
, radio tower at forest edge, elev
<quantity id="456F3AA43058FFB1FCB8FA31BEC408A0" box="[845,935,1443,1467]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="33" pageNumber="234" unit="m" value="1130.0">
<elevation id="09BA70723058FFB1FCB8FA31BEC408A0" box="[845,935,1443,1467]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="33" pageNumber="234" unit="m" value="1130.0">1130 m</elevation>
</quantity>
<date id="F629B1813058FFB1FC43FA31B9E108A0" box="[950,1154,1443,1467]" pageId="33" pageNumber="234" value="2003-03-09" valueMax="2003-03-20" valueMin="2003-03-09">
<collectingDate id="E66D48693058FFB1FC43FA31B9E108A0" box="[950,1154,1443,1467]" pageId="33" pageNumber="234" value="2003-03-09" valueMax="2003-03-20" valueMin="2003-03-09">9-20 March 2003</collectingDate>
</date>
”/ “
<geoCoordinate id="E7A3F1863058FFB1FF72FA56BC6308C7" box="[135,256,1475,1500]" degrees="21" direction="south" minutes="15. 05" orientation="latitude" pageId="33" pageNumber="234" precision="9" value="-21.250834">21°15.05’S</geoCoordinate>
,
<geoCoordinate id="E7A3F1863058FFB1FEF8FA56BCE408C0" box="[269,391,1476,1500]" degrees="47" direction="east" minutes="24. 43" orientation="longitude" pageId="33" pageNumber="234" precision="9" value="47.407166">47°24.43’E</geoCoordinate>
collector:
<collectorName id="2F62F2973058FFB1FE0CFA56BE0E08C7" box="[505,877,1476,1500]" pageId="33" pageNumber="234">R. Harin’Hala California Academy</collectorName>
of Sciences malaise, mixed tropical forest MA-02-09B-55”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3058FFB1FE16FA76BE3B0B46" collectingDate="1998-04-18" collectionCode="CAS" collectorName="D. H. Kavanaugh" country="Madagascar" elevation="1150" latitude="-21.213333" location="Vohiparara area" longLatPrecision="126" longitude="47.383335" municipality="Ranomafana National Park" pageId="33" pageNumber="234" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="94915CC83058FFB1FE16FA76BF4C08E7" box="[483,559,1508,1532]" count="1" pageId="33" pageNumber="234" type="male">1 male</specimenCount>
(in
<collectionCode id="E4860F843058FFB1FD97FA76BFF508E7" box="[610,662,1508,1532]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="33" pageNumber="234" type="Museum">CAS</collectionCode>
) labeled “CASENT 1049026”/ “
<collectingCountry id="FA80D7D13058FFB1FBF3FA76BDD10B07" name="Madagascar" pageId="33" pageNumber="234">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A33058FFB1FF4AF996BCCE0B07" box="[191,429,1540,1564]" country="Madagascar" name="Fianarantsoa" pageId="33" pageNumber="234">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B3058FFB1FE4FF996BFB80B07" box="[442,731,1540,1564]" pageId="33" pageNumber="234">Ranomafana National Park</collectingMunicipality>
,
<location id="8748C19A3058FFB1FD1DF996BEEC0B07" LSID="urn:lsid:plazi:treatment:0A3E26573058FFB3FF8EFCB7B95E09C5:8748C19A3058FFB1FD1DF996BEEC0B07" box="[744,911,1540,1564]" country="Madagascar" latitude="-21.213333" longLatPrecision="126" longitude="47.383335" municipality="Ranomafana National Park" name="Vohiparara area" pageId="33" pageNumber="234" stateProvince="Fianarantsoa">Vohiparara area</location>
,
<quantity id="456F3AA43058FFB1FC68F996BE8E0B07" box="[925,1005,1540,1564]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="33" pageNumber="234" unit="m" value="1150.0">
<elevation id="09BA70723058FFB1FC68F996BE8E0B07" box="[925,1005,1540,1564]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="33" pageNumber="234" unit="m" value="1150.0">1150 m</elevation>
</quantity>
, mixed tropical forest,
<date id="F629B1813058FFB1FF33F9B6BC3F0B27" box="[198,348,1572,1596]" pageId="33" pageNumber="234" value="1998-04-18">
<collectingDate id="E66D48693058FFB1FF33F9B6BC3F0B27" box="[198,348,1572,1596]" pageId="33" pageNumber="234" value="1998-04-18">18 April 1998</collectingDate>
</date>
,
<geoCoordinate id="E7A3F1863058FFB1FE9EF9B6BCB40B27" box="[363,471,1572,1596]" degrees="21" direction="south" minutes="12. 8" orientation="latitude" pageId="33" pageNumber="234" precision="92" value="-21.213333">21°12.8’S</geoCoordinate>
<geoCoordinate id="E7A3F1863058FFB1FE2BF9B6BF280B27" box="[478,587,1572,1596]" degrees="47" direction="east" minutes="23. 0" orientation="longitude" pageId="33" pageNumber="234" precision="92" value="47.383335">47°23.0’E</geoCoordinate>
,”/ “Stop# 98-52,
<collectorName id="2F62F2973058FFB1FCF9F9B7BEAB0B27" box="[780,968,1572,1596]" pageId="33" pageNumber="234">D. H. Kavanaugh</collectorName>
collector collected by beating live vegetation and suspended dead leaves and twigs”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3058FFB2FC9EF9D7BDA308C0" collectingDate="1998-04-29" collectionCode="CAS" collectorName="D. H. Kavanaugh" country="Madagascar" elevation="1050" lastPageId="34" lastPageNumber="235" latitude="-21.278334" location="Ambatovory" longLatPrecision="126" longitude="47.435" municipality="Ranomafana National Park" pageId="33" pageNumber="234" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="94915CC83058FFB1FC9EF9D7BEAF0B46" box="[875,972,1605,1629]" count="1" pageId="33" pageNumber="234" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843058FFB1FBF5F9D7B9570B46" box="[1024,1076,1605,1629]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="33" pageNumber="234" type="Museum">CAS</collectionCode>
) labeled “CASENT 1049025”/ “
<collectingCountry id="FA80D7D13058FFB1FE70F9F7BF200B66" box="[389,579,1637,1661]" name="Madagascar" pageId="33" pageNumber="234">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A33058FFB1FDA2F9F7BE2F0B66" box="[599,844,1637,1661]" country="Madagascar" name="Fianarantsoa" pageId="33" pageNumber="234">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B3058FFB1FC95F9F7B9EC0B66" box="[864,1167,1637,1661]" pageId="33" pageNumber="234">Ranomafana National Park</collectingMunicipality>
, Vatoharanana area,
<quantity id="456F3AA43058FFB1FEA5F917BCC00B86" box="[336,419,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="33" pageNumber="234" unit="m" value="1050.0">
<elevation id="09BA70723058FFB1FEA5F917BCC00B86" box="[336,419,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="33" pageNumber="234" unit="m" value="1050.0">1050 m</elevation>
</quantity>
,
<location id="8748C19A3058FFB1FE44F917BF5B0B86" LSID="urn:lsid:plazi:treatment:0A3E26573058FFB3FF8EFCB7B95E09C5:8748C19A3058FFB1FE44F917BF5B0B86" box="[433,568,1669,1693]" country="Madagascar" latitude="-21.278334" longLatPrecision="126" longitude="47.435" municipality="Ranomafana National Park" name="Ambatovory" pageId="33" pageNumber="234" stateProvince="Fianarantsoa">Ambatovory</location>
stream,
<date id="F629B1813058FFB1FD62F917BE4C0B86" box="[663,815,1669,1693]" pageId="33" pageNumber="234" value="1998-04-29">
<collectingDate id="E66D48693058FFB1FD62F917BE4C0B86" box="[663,815,1669,1693]" pageId="33" pageNumber="234" value="1998-04-29">29 April 1998</collectingDate>
</date>
,
<geoCoordinate id="E7A3F1863058FFB1FCCBF917BEC90B86" box="[830,938,1669,1693]" degrees="21" direction="south" minutes="16. 7" orientation="latitude" pageId="33" pageNumber="234" precision="92" value="-21.278334">21°16.7’S</geoCoordinate>
<geoCoordinate id="E7A3F1863058FFB1FC46F917B94B0B86" box="[947,1064,1669,1693]" degrees="47" direction="east" minutes="26. 1" orientation="longitude" pageId="33" pageNumber="234" precision="92" value="47.435">47° 26.1’E</geoCoordinate>
,”/ “Stop# 98-101,
<collectorName id="2F62F2973058FFB1FF22F934BCFB0BA6" box="[215,408,1701,1725]" pageId="33" pageNumber="234">D. H. Kavanaugh</collectorName>
collector collected by beating suspended clusters of dead leaves and twigs”
</materialsCitation>
;
<materialsCitation id="32FF9D1C305BFFB2FF21FA51BFA30B46" collectingDate="2003-03-27" collectingDateMax="2003-03-31" collectingDateMin="2003-03-27" collectionCode="CAS" collectorName="Fisher" country="Madagascar" county="Parc National de Ranomafana" elevation="1100" latitude="-21.256666" location="California Academy of Sciences" longLatPrecision="21" longitude="47.433334" municipality="Vatoharanana River" pageId="34" pageNumber="235" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="94915CC8305BFFB2FF21FA51BC4808C0" box="[212,299,1475,1499]" count="2" pageId="34" pageNumber="235" type="male">2 males</specimenCount>
and
<specimenCount id="94915CC8305BFFB2FE9FFA51BCAE08C0" box="[362,461,1475,1499]" count="1" pageId="34" pageNumber="235" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F84305BFFB2FDF7FA51BF5508C0" box="[514,566,1475,1499]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="34" pageNumber="235" type="Museum">CAS</collectionCode>
and EM) labeled “CASENT 8072308” or “CASENT 8072309” or “CASENT 8072307”, respectively/ “
<collectingCountry id="FA80D7D1305BFFB2FD55FA71BE3D08E0" box="[672,862,1507,1531]" name="Madagascar" pageId="34" pageNumber="235">MADAGASCAR</collectingCountry>
:
<collectingRegion id="405359A3305BFFB2FC9AFA71BE9408E0" box="[879,1015,1507,1531]" country="Madagascar" name="Fianarantsoa" pageId="34" pageNumber="235">Fianarantsoa</collectingRegion>
<collectingCounty id="6B49EFCD305BFFB2FBF4FA76BC460B07" pageId="34" pageNumber="235">Parc National de Ranomafana</collectingCounty>
,
<collectingMunicipality id="624C0D3B305BFFB2FEC0F996BF680B07" box="[309,523,1540,1564]" pageId="34" pageNumber="235">Vatoharanana River</collectingMunicipality>
,
<locationDeviation id="939A8E63305BFFB2FDE9F996BE0A0B07" box="[540,873,1540,1564]" location="Ranomafana" pageId="34" pageNumber="235">
<quantity id="456F3AA4305BFFB2FDE9F996BF0A0B07" box="[540,617,1540,1564]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="34" pageNumber="235" unit="km" value="4.1">4.1 km</quantity>
231° SW Ranomafana
</locationDeviation>
elev
<quantity id="456F3AA4305BFFB2FC5FF996BE9E0B00" box="[938,1021,1540,1564]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="34" pageNumber="235" unit="m" value="1100.0">
<elevation id="09BA7072305BFFB2FC5FF996BE9E0B00" box="[938,1021,1540,1564]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="34" pageNumber="235" unit="m" value="1100.0">1100 m</elevation>
</quantity>
<date id="F629B181305BFFB2FBF2F996BDD30B27" pageId="34" pageNumber="235" value="2003-03-27" valueMax="2003-03-31" valueMin="2003-03-27">
<collectingDate id="E66D4869305BFFB2FBF2F996BDD30B27" pageId="34" pageNumber="235" value="2003-03-27" valueMax="2003-03-31" valueMin="2003-03-27">27-31 March 2003</collectingDate>
</date>
”/ “
<geoCoordinate id="E7A3F186305BFFB2FF2FF9B6BC3B0B27" box="[218,344,1572,1596]" degrees="21" direction="south" minutes="15" orientation="latitude" pageId="34" pageNumber="235" precision="15" seconds="24" value="-21.256666">21°15’24”S</geoCoordinate>
,
<geoCoordinate id="E7A3F186305BFFB2FE9CF9B6BC8A0B27" box="[361,489,1572,1596]" degrees="47" direction="east" minutes="26" orientation="longitude" pageId="34" pageNumber="235" precision="15" seconds="00" value="47.433334">47°26’00”E</geoCoordinate>
<location id="8748C19A305BFFB2FE01F9B6BE3B0B27" LSID="urn:lsid:plazi:treatment:0A3E26573058FFB3FF8EFCB7B95E09C5:8748C19A305BFFB2FE01F9B6BE3B0B27" box="[500,856,1572,1596]" country="Madagascar" county="Parc National de Ranomafana" latitude="-21.256666" longLatPrecision="21" longitude="47.433334" municipality="Vatoharanana River" name="California Academy of Sciences" pageId="34" pageNumber="235" stateProvince="Fianarantsoa">California Academy of Sciences</location>
coll.
<collectorName id="2F62F297305BFFB2FC69F9B6BEBD0B27" box="[924,990,1572,1596]" pageId="34" pageNumber="235">Fisher</collectorName>
, Griswold et al. beating low veg., montane rainforest code: BLF8401”
</materialsCitation>
;
<materialsCitation id="32FF9D1C305BFFB2FD3BF9D7BE950B86" collectingDate="2005-07-25" collectionCode="MNHN" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="150" location="Vatoharanana" municipality="Ranomafana" pageId="34" pageNumber="235" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="94915CC8305BFFB2FD3BF9D7BE740B47" box="[718,791,1604,1629]" count="1" pageId="34" pageNumber="235" type="male">1 male</specimenCount>
(in
<collectionCode id="E4860F84305BFFB2FCB1F9D7BEF60B47" box="[836,917,1605,1628]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="34" pageNumber="235">MNHN</collectionCode>
) labeled “
<collectingCountry id="FA80D7D1305BFFB2FBF3F9D7BDD10B66" name="Madagascar" pageId="34" pageNumber="235">MADAGASCAR</collectingCountry>
,
<collectingMunicipality id="624C0D3B305BFFB2FF35F9F7BC240B66" box="[192,327,1637,1661]" pageId="34" pageNumber="235">Ranomafana</collectingMunicipality>
NP.,
<location id="8748C19A305BFFB2FE76F9F7BF770B66" LSID="urn:lsid:plazi:treatment:0A3E26573058FFB3FF8EFCB7B95E09C5:8748C19A305BFFB2FE76F9F7BF770B66" box="[387,532,1637,1661]" country="Madagascar" municipality="Ranomafana" name="Vatoharanana" pageId="34" pageNumber="235">Vatoharanana</location>
, Trail K at
<quantity id="456F3AA4305BFFB2FD60F9F7BFB90B67" box="[661,730,1637,1661]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="235" unit="m" value="150.0">
<elevation id="09BA7072305BFFB2FD60F9F7BFB90B67" box="[661,730,1637,1661]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="235" unit="m" value="150.0">150 m</elevation>
</quantity>
mark,
<date id="F629B181305BFFB2FCD2F9F7BEF10B66" box="[807,914,1637,1661]" pageId="34" pageNumber="235" value="2005-07-25">
<collectingDate id="E66D4869305BFFB2FCD2F9F7BEF10B66" box="[807,914,1637,1661]" pageId="34" pageNumber="235" value="2005-07-25">25.7.2005</collectingDate>
</date>
”/ “collected by hand in daytime from dead leaves of bushes and trees,
<collectorName id="2F62F297305BFFB2FD8BF914BF920B86" box="[638,753,1669,1693]" pageId="34" pageNumber="235">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F297305BFFB2FCEDF914BE1A0B86" box="[792,889,1669,1693]" pageId="34" pageNumber="235">J. Rainio</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="32FF9D1C305BFFB3FBF3F917BC5E0C08" collectingDate="2005-07-28" collectionCode="MNHN, MZF" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="50" lastPageId="35" lastPageNumber="236" location="Talatakely" municipality="Ranomafana" pageId="34" pageNumber="235" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="94915CC8305BFFB2FBF3F917B90C0B86" box="[1030,1135,1669,1693]" count="2" pageId="34" pageNumber="235" type="female">2 females</specimenCount>
(in
<collectionCode id="E4860F84305BFFB2FF8EF934BDAF0BA6" box="[123,204,1702,1725]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="34" pageNumber="235">MNHN</collectionCode>
and
<collectionCode id="E4860F84305BFFB2FF0AF934BC550BA6" box="[255,310,1702,1725]" pageId="34" pageNumber="235">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D1305BFFB2FE51F934BF010BA5" box="[420,610,1702,1726]" name="Madagascar" pageId="34" pageNumber="235">MADAGASCAR</collectingCountry>
,
<collectingMunicipality id="624C0D3B305BFFB2FD9BF937BF960BA6" box="[622,757,1701,1725]" pageId="34" pageNumber="235">Ranomafana</collectingMunicipality>
NP.,
<location id="8748C19A305BFFB2FCD8F937BEF90BA6" LSID="urn:lsid:plazi:treatment:0A3E26573058FFB3FF8EFCB7B95E09C5:8748C19A305BFFB2FCD8F937BEF90BA6" box="[813,922,1701,1725]" country="Madagascar" municipality="Ranomafana" name="Talatakely" pageId="34" pageNumber="235">Talatakely</location>
, Trail SP at
<quantity id="456F3AA4305BFFB2FBE9F937B9310BA6" box="[1052,1106,1701,1725]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="235" unit="m" value="50.0">
<elevation id="09BA7072305BFFB2FBE9F937B9310BA6" box="[1052,1106,1701,1725]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="235" unit="m" value="50.0">50 m</elevation>
</quantity>
mark,
<date id="F629B181305AFFB3FF8EFF49BD910DE8" box="[123,242,219,243]" pageId="35" pageNumber="236" value="2005-07-28">
<collectingDate id="E66D4869305AFFB3FF8EFF49BD910DE8" box="[123,242,219,243]" pageId="35" pageNumber="236" value="2005-07-28">28.07.2005</collectingDate>
</date>
”/ “collected by hand in daytime from dead leaves of bushes and trees,
<collectorName id="2F62F297305AFFB3FC1AFF49B9020DE8" box="[1007,1121,219,243]" pageId="35" pageNumber="236">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F297305AFFB3FB71FF49BDA10C08" pageId="35" pageNumber="236">J. Rainio</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="32FF9D1C305AFFB3FEB9FF69BE6D0C4F" collectingDate="2005-01-28" collectionCode="MNHN, MZF" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="555" location="Valohoaka" municipality="Ranomafana" pageId="35" pageNumber="236" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="94915CC8305AFFB3FEB9FF69BCD70C08" box="[332,436,251,275]" count="2" pageId="35" pageNumber="236" type="female">2 females</specimenCount>
(in
<collectionCode id="E4860F84305AFFB3FE15FF6EBF520C08" box="[480,561,252,275]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="35" pageNumber="236">MNHN</collectionCode>
and
<collectionCode id="E4860F84305AFFB3FD92FF6EBFFD0C08" box="[615,670,252,275]" pageId="35" pageNumber="236">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D1305AFFB3FCFAFF69BEAE0C08" box="[783,973,251,275]" name="Madagascar" pageId="35" pageNumber="236">MADAGASCAR</collectingCountry>
,
<collectingMunicipality id="624C0D3B305AFFB3FC2EFF69B9010C08" box="[987,1122,251,275]" pageId="35" pageNumber="236">Ranomafana</collectingMunicipality>
NP.,
<location id="8748C19A305AFFB3FF8EFE8EBD8F0C2F" LSID="urn:lsid:plazi:treatment:0A3E26573058FFB3FF8EFCB7B95E09C5:8748C19A305AFFB3FF8EFE8EBD8F0C2F" box="[123,236,284,308]" country="Madagascar" municipality="Ranomafana" name="Valohoaka" pageId="35" pageNumber="236">Valohoaka</location>
, Trail B at
<quantity id="456F3AA4305AFFB3FE9CFE8EBCCC0C28" box="[361,431,284,308]" metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="35" pageNumber="236" unit="m" value="555.0">
<elevation id="09BA7072305AFFB3FE9CFE8EBCCC0C28" box="[361,431,284,308]" metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="35" pageNumber="236" unit="m" value="555.0">555 m</elevation>
</quantity>
mark,
<date id="F629B181305AFFB3FE09FE8EBF100C2F" box="[508,627,284,308]" pageId="35" pageNumber="236" value="2005-01-28">
<collectingDate id="E66D4869305AFFB3FE09FE8EBF100C2F" box="[508,627,284,308]" pageId="35" pageNumber="236" value="2005-01-28">28.01.2005</collectingDate>
</date>
”/ “collected by hand in daytime from dead leaves of bushes and trees,
<collectorName id="2F62F297305AFFB3FEABFEAFBCB20C4F" box="[350,465,316,340]" pageId="35" pageNumber="236">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F297305AFFB3FE0CFEAFBF390C4F" box="[505,602,316,340]" pageId="35" pageNumber="236">J. Rainio</collectorName>
collectors”; and
</materialsCitation>
<materialsCitation id="32FF9D1C305AFFB3FCE2FEAEBC8B0CCE" collectingDate="2005-11-16" collectionCode="MNHN" collectorName="F. Ratalata &amp; J. Rainio &amp; All" country="Madagascar" elevation="555" location="Valohoaka" municipality="Ranomafana" pageId="35" pageNumber="236" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="94915CC8305AFFB3FCE2FEAEBEE20C4F" box="[791,897,316,340]" count="2" pageId="35" pageNumber="236" type="female">2 females</specimenCount>
(in
<collectionCode id="E4860F84305AFFB3FC44FEAFB9610C4F" box="[945,1026,317,340]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="35" pageNumber="236">MNHN</collectionCode>
and
<collectionCode id="E4860F84305AFFB3FBCEFEAFB9EF0C4F" box="[1083,1164,317,340]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="35" pageNumber="236">MNHN</collectionCode>
) labeled “
<collectingCountry id="FA80D7D1305AFFB3FF2CFECFBCF40C6E" box="[217,407,349,373]" name="Madagascar" pageId="35" pageNumber="236">MADAGASCAR</collectingCountry>
,
<collectingMunicipality id="624C0D3B305AFFB3FE51FECEBF480C6F" box="[420,555,348,372]" pageId="35" pageNumber="236">Ranomafana</collectingMunicipality>
NP.,
<location id="8748C19A305AFFB3FDA8FECEBFAD0C6F" LSID="urn:lsid:plazi:treatment:0A3E26573058FFB3FF8EFCB7B95E09C5:8748C19A305AFFB3FDA8FECEBFAD0C6F" box="[605,718,348,372]" country="Madagascar" municipality="Ranomafana" name="Valohoaka" pageId="35" pageNumber="236">Valohoaka</location>
, Trail B at
<quantity id="456F3AA4305AFFB3FCB1FECEBEE40C6F" box="[836,903,348,372]" metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="35" pageNumber="236" unit="m" value="555.0">
<elevation id="09BA7072305AFFB3FCB1FECEBEE40C6F" box="[836,903,348,372]" metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="35" pageNumber="236" unit="m" value="555.0">555 m</elevation>
</quantity>
mark,
<date id="F629B181305AFFB3FC25FECEB9250C6F" box="[976,1094,348,373]" pageId="35" pageNumber="236" value="2005-11-16">
<collectingDate id="E66D4869305AFFB3FC25FECEB9250C6F" box="[976,1094,348,373]" pageId="35" pageNumber="236" value="2005-11-16">16.11.2005</collectingDate>
</date>
”/ “collected by hand in daytime from dead leaves of bushes and trees,
<collectorName id="2F62F297305AFFB3FCD9FEEFBEFE0C8E" box="[812,925,381,405]" pageId="35" pageNumber="236">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F297305AFFB3FC35FEEFB97C0C8E" box="[960,1055,381,405]" pageId="35" pageNumber="236">J. Rainio</collectorName>
collector”.
<collectorName id="2F62F297305AFFB3FF8EFE0FBDFE0CAE" box="[123,157,413,437]" pageId="35" pageNumber="236">All</collectorName>
<typeStatus id="5D2C29E3305AFFB3FF51FE0CBC690CAD" box="[164,266,414,438]" pageId="35" pageNumber="236" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="5D2C29E3305AFFB3FD95FE0CBF870CAD" box="[608,740,414,438]" pageId="35" pageNumber="236" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC2305AFFB3FD19FE0FBDA10CCD" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="bimaculata" status="sp. nov.">
<emphasis id="B0E34B53305AFFB3FD19FE0FBE9B0CAE" box="[748,1016,413,437]" italics="true" pageId="35" pageNumber="236">Madecassina bimaculata</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628305AFFB3FF3CFE2CBC600CCE" box="[201,259,446,469]" pageId="35" pageNumber="236" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<caption id="D6E8C7C9305BFFB2FF6EFAF3BC6308BC" pageId="34" pageNumber="235" startId="34.[155,167,1377,1396]" targetBox="[120,1176,216,1355]" targetPageId="34" targetType="figure">
<paragraph id="82289741305BFFB2FF6EFAF3BC6308BC" blockId="34.[123,1173,1377,1447]" pageId="34" pageNumber="235">
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19. Digital images of holotype male of
<taxonomicName id="4597ECC2305BFFB2FDDBFAF3BEDC086F" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[558,959,1377,1396]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="bimaculata" status="sp. nov.">
<emphasis id="B0E34B53305BFFB2FDDBFAF3BE67086F" box="[558,772,1377,1396]" italics="true" pageId="34" pageNumber="235">Madecassina bimaculata</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628305BFFB2FC31FAF3B965086F" box="[964,1030,1377,1396]" pageId="34" pageNumber="235" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels C–D. Median lobe of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm.
</paragraph>
</caption>
<paragraph id="82289741305AFFB3FF5EFE4CB9500CED" blockId="35.[123,1174,219,1247]" box="[171,1075,478,502]" pageId="35" pageNumber="236">
<materialsCitation id="32FF9D1C305AFFB3FF5EFE4CB94F0CED" box="[171,1068,478,502]" country="Madagascar" location="Ranomafana National Park" pageId="35" pageNumber="236" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B53305AFFB3FF5EFE4CBC340CEF" bold="true" box="[171,343,478,502]" pageId="35" pageNumber="236">
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<typeStatus id="5D2C29E3305AFFB3FF5EFE4CBD870CEE" box="[171,228,478,502]" pageId="35" pageNumber="236">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D305AFFB3FF19FE73BC340CEF" baselines="495" box="[236,343,481,500]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="35" pageNumber="236">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D1305AFFB3FE8BFE4CBC9C0CED" box="[382,511,478,502]" name="Madagascar" pageId="35" pageNumber="236">Madagascar</collectingCountry>
,
<collectingRegion id="405359A3305AFFB3FDF8FE4CBF9F0CED" box="[525,764,478,502]" country="Madagascar" name="Fianarantsoa" pageId="35" pageNumber="236">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A305AFFB3FCFFFE4CB94F0CED" LSID="urn:lsid:plazi:treatment:0A3E26573058FFB3FF8EFCB7B95E09C5:8748C19A305AFFB3FCFFFE4CB94F0CED" box="[778,1068,478,502]" country="Madagascar" name="Ranomafana National Park" pageId="35" pageNumber="236" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="82289741305AFFB3FF5EFE6DBE790F4C" blockId="35.[123,1174,219,1247]" pageId="35" pageNumber="236">
<emphasis id="B0E34B53305AFFB3FF5EFE6DBC880F0E" bold="true" box="[171,491,511,535]" pageId="35" pageNumber="236">
<smallCapsWord id="84CE019D305AFFB3FF5EFE6DBC4D0F0E" baselines="529,528" box="[171,302,511,535]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="35" pageNumber="236">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D305AFFB3FEC6FD90BC2C0F0E" baselines="528" box="[307,335,514,533]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="35" pageNumber="236">OF</smallCapsWord>
<smallCapsWord id="84CE019D305AFFB3FEA1FD90BCC40F0E" baselines="528" box="[340,423,514,533]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="35" pageNumber="236">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D305AFFB3FE5BFD90BC880F0E" baselines="528" box="[430,491,514,533]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="35" pageNumber="236">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC2305AFFB3FD1FFE6CBE010F0D" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[746,866,510,534]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="bimaculata">
<emphasis id="B0E34B53305AFFB3FD1FFE6CBE010F0D" box="[746,866,510,534]" italics="true" pageId="35" pageNumber="236">bimaculata</emphasis>
</taxonomicName>
, is an adjective derived from the Latin words,
<emphasis id="B0E34B53305AFFB3FEC0FD8DBC370F2C" box="[309,340,543,567]" italics="true" pageId="35" pageNumber="236">bis</emphasis>
, meaning two, and
<emphasis id="B0E34B53305AFFB3FDD8FD8DBF1F0F2C" box="[557,636,543,567]" italics="true" pageId="35" pageNumber="236">macula</emphasis>
, meaning spot or spotted. The name refers to the pair of subapical elytral spots seen in members of this species.
</paragraph>
<paragraph id="82289741305AFFB3FF5EFDF2BC030927" blockId="35.[123,1174,219,1247]" pageId="35" pageNumber="236">
<smallCapsWord id="84CE019D305AFFB3FF5EFDF2BC210F6D" baselines="626,625" box="[171,322,608,632]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="35" pageNumber="236">
<emphasis id="B0E34B53305AFFB3FF5EFDF2BC210F6D" bold="true" box="[171,322,608,632]" pageId="35" pageNumber="236">RECOGNITION</emphasis>
</smallCapsWord>
.— Size average for genus, SBL of males = 4.7–
<quantity id="456F3AA4305AFFB3FCA1FDCDBEC40F6C" box="[852,935,607,631]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="35" pageNumber="236" unit="mm" value="5.5">5.5 mm</quantity>
, of females = 5.3–5.
<quantity id="456F3AA4305AFFB3FB7DFDCDBDC70F8C" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="35" pageNumber="236" unit="mm" value="6.0">6 mm</quantity>
. Members of this species (
<figureCitation id="1AAC8BC4305AFFB3FE3EFD12BF4F0F83" box="[459,556,640,664]" captionStart="FIGURE" captionStartId="34.[155,167,1377,1396]" captionTargetBox="[120,1176,216,1355]" captionTargetId="figure-856@34.[120,1176,216,1374]" captionTargetPageId="34" captionText="FIGURE 19. Digital images of holotype male of Madecassina bimaculata Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels C–D. Median lobe of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." pageId="35" pageNumber="236">Fig. 19A</figureCitation>
) key most closely, to
<taxonomicName id="4597ECC2305AFFB3FCEBFD12B9F60F83" authority="Alluaud (1897)" authorityName="Alluaud" authorityYear="1897" box="[798,1173,640,664]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="B0E34B53305AFFB3FCEBFD12BE8B0F83" box="[798,1000,640,664]" italics="true" pageId="35" pageNumber="236">Madecassina picta</emphasis>
<bibRefCitation id="E606EAB0305AFFB3FC05FD12B9F60F83" author="ALLUAUD, C." box="[1008,1173,640,664]" pageId="35" pageNumber="236" pagination="165 - 181" refId="ref28722" refString="ALLUAUD, C. 1897. Descriptions de Carabiques nouveaux de Madagascar. Annales de la Societe Entomologique de France 66: 165 - 181." type="journal article" year="1897">Alluaud (1897)</bibRefCitation>
</taxonomicName>
in Jeannel’s (1949) key based on the following combination of features: pronotum impunctate, elytra with two subapical pale spots and striae deeply impressed and legs testaceous in color. The only feature used in the key with which
<taxonomicName id="4597ECC2305AFFB3FDF2FD70BFFF0FE2" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[519,668,737,761]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="bimaculata">
<emphasis id="B0E34B53305AFFB3FDF2FD70BFFF0FE2" box="[519,668,737,761]" italics="true" pageId="35" pageNumber="236">T. bimaculata</emphasis>
</taxonomicName>
members differ is the presence of is metallic reflection on the forebody—members of
<taxonomicName id="4597ECC2305AFFB3FDC7FC90BFE20E02" authorityName="Alluaud" authorityYear="1897" box="[562,641,770,793]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="B0E34B53305AFFB3FDC7FC90BFE20E02" box="[562,641,770,793]" italics="true" pageId="35" pageNumber="236">T. picta</emphasis>
</taxonomicName>
have evident aeneous or greenish metallic reflection on the head and pronotum, whereas
<taxonomicName id="4597ECC2305AFFB3FDC8FCB1BFB10E21" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[573,722,802,826]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="bimaculata">
<emphasis id="B0E34B53305AFFB3FDC8FCB1BFB10E21" box="[573,722,802,826]" italics="true" pageId="35" pageNumber="236">T. bimaculata</emphasis>
</taxonomicName>
members are without metallic reflection. These two species are otherwise very similar externally; however the pronotum is proportionately slightly longer and more distinctly narrowed basally and the hind angles more sharply defined and rectangular in
<taxonomicName id="4597ECC2305AFFB3FEE1FC16BCC50E80" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[276,422,899,923]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="bimaculata">
<emphasis id="B0E34B53305AFFB3FEE1FC16BCC50E80" box="[276,422,899,923]" italics="true" pageId="35" pageNumber="236">T. bimaculata</emphasis>
</taxonomicName>
members than in
<taxonomicName id="4597ECC2305AFFB3FD91FC16BFD10E80" authorityName="Alluaud" authorityYear="1897" box="[612,690,900,923]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="B0E34B53305AFFB3FD91FC16BFD10E80" box="[612,690,900,923]" italics="true" pageId="35" pageNumber="236">T. picta</emphasis>
</taxonomicName>
members (which have the hind angles slightly obtuse). Also, the apex of the median lobe of the male genitalia is shorter, thicker and straighter in lateral aspect (
<figureCitation id="1AAC8BC4305AFFB3FEC6FC56BCF20EC7" box="[307,401,964,988]" captionStart="FIGURE" captionStartId="34.[155,167,1377,1396]" captionTargetBox="[120,1176,216,1355]" captionTargetId="figure-856@34.[120,1176,216,1374]" captionTargetPageId="34" captionText="FIGURE 19. Digital images of holotype male of Madecassina bimaculata Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels C–D. Median lobe of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." pageId="35" pageNumber="236">Fig. 19C</figureCitation>
) and more bluntly triangular in dorsal aspect (
<figureCitation id="1AAC8BC4305AFFB3FC71FC56BE800EC0" box="[900,995,964,988]" captionStart="FIGURE" captionStartId="34.[155,167,1377,1396]" captionTargetBox="[120,1176,216,1355]" captionTargetId="figure-856@34.[120,1176,216,1374]" captionTargetPageId="34" captionText="FIGURE 19. Digital images of holotype male of Madecassina bimaculata Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels C–D. Median lobe of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." pageId="35" pageNumber="236">Fig. 19D</figureCitation>
) in
<taxonomicName id="4597ECC2305AFFB3FBFAFC57BDF30EE7" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="bimaculata">
<emphasis id="B0E34B53305AFFB3FBFAFC57BDF30EE7" italics="true" pageId="35" pageNumber="236">T. bimaculata</emphasis>
</taxonomicName>
males than in those of
<taxonomicName id="4597ECC2305AFFB3FE55FC77BC970EE7" authorityName="Alluaud" authorityYear="1897" box="[416,500,997,1020]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="B0E34B53305AFFB3FE55FC77BC970EE7" box="[416,500,997,1020]" italics="true" pageId="35" pageNumber="236">T. picta</emphasis>
</taxonomicName>
males, which have the apex longer, narrower and distinctly hooked dorsally in lateral aspect and more narrowly rounded in dorsal aspect (see Jeannel 1949, Figs.
<quantity id="456F3AA4305AFFB3FF42FBB7BD8F0926" box="[183,236,1061,1085]" metricMagnitude="-1" metricUnit="kg" metricValue="4.74" pageId="35" pageNumber="236" unit="g" value="474.0">474g</quantity>
and 474i).
</paragraph>
<paragraph id="82289741305AFFB3FF5EFBD7B9690946" blockId="35.[123,1174,219,1247]" box="[171,1034,1093,1118]" pageId="35" pageNumber="236">
<emphasis id="B0E34B53305AFFB3FF5EFBD7BC920947" bold="true" box="[171,497,1093,1117]" pageId="35" pageNumber="236">
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</emphasis>
.— At present, known only from the
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locality.
</paragraph>
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.— Specimens of this species have been collected in primary montane rainforest in the Vatoharanana and Valohoaka areas and in secondary montane rainforest in the Talatakely and Vohiparara areas, at elevations ranging from
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. They have been collected mainly by beating live vegetation and suspended clusters of dead leaves and twigs.
</paragraph>
</subSubSection>
</treatment>
</document>

222
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<mods:title id="D491AEF0CBB6D98BB0FD4C1A8B1A6F42">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:affiliation id="65BAFCFAA4EF52262416DCC78D0F5E41">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:namePart id="AED4E960EAD0DE7A3AF7DFC5CFC6951B">Rainio, Johanna</mods:namePart>
<mods:affiliation id="229E3A67CD28C64BDADA526F6E7A5586">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
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<emphasis id="B0E34B53305AFFB3FF8EFA95BCA20804" bold="true" box="[123,449,1287,1311]" italics="true" pageId="35" pageNumber="236">Madecassina quadrimaculata</emphasis>
Kavanaugh and Rainio
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,
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.—
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(
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), a male, in CAS, labeled: “
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, Ranomafana NP., montane rainforest near
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,
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in from forest edge,”/ “
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,
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, collected by hand in daytime,
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collector”/ “
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<emphasis id="B0E34B53305AFFB3FE85FA51BFCF08C0" box="[368,684,1475,1499]" italics="true" pageId="35" pageNumber="236">Madecassina quadrimaculata</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
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2015” [red label].
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.—
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,
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,
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.
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</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC2305AFFB3FD1AF991BEFA0B00" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[751,921,1539,1563]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="quadrimaculata">
<emphasis id="B0E34B53305AFFB3FD1AF991BEFA0B00" box="[751,921,1539,1563]" italics="true" pageId="35" pageNumber="236">quadrimaculata</emphasis>
</taxonomicName>
, is an adjective derived from the Latin words,
<emphasis id="B0E34B53305AFFB3FE80F9B7BCB20B27" box="[373,465,1573,1596]" italics="true" pageId="35" pageNumber="236">quattuor</emphasis>
, meaning four, and
<emphasis id="B0E34B53305AFFB3FD47F9B6BE620B27" box="[690,769,1572,1596]" italics="true" pageId="35" pageNumber="236">macula</emphasis>
, meaning spot or spotted. The name refers to the two (subbasal and subapical) pairs of elytral spots seen in members of this species.
</paragraph>
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<emphasis id="B0E34B53305AFFB3FF5EF9F7BC210B60" bold="true" box="[171,322,1637,1661]" pageId="35" pageNumber="236">RECOGNITION</emphasis>
</smallCapsWord>
.— Size average for genus, SBL =
<quantity id="456F3AA4305AFFB3FD34F9F6BE770B67" box="[705,788,1636,1661]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="35" pageNumber="236" unit="mm" value="5.2">5.2 mm</quantity>
. The
<typeStatus id="5D2C29E3305AFFB3FCA3F9F7BED00B66" box="[854,947,1637,1661]" pageId="35" pageNumber="236" type="holotype">holotype</typeStatus>
male of this species (
<figureCitation id="1AAC8BC4305AFFB3FF71F917BD870B86" box="[132,228,1669,1693]" captionStart="FIGURE" captionStartId="36.[155,167,1570,1589]" captionTargetBox="[144,1176,216,1552]" captionTargetId="figure-53@36.[120,1176,216,1567]" captionTargetPageId="36" captionText="FIGURE 20. Digital images of holotype male of Madecassina quadrimaculata Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." pageId="35" pageNumber="236">Fig. 20A</figureCitation>
) keys to
<taxonomicName id="4597ECC2305AFFB3FEB9F917BF9C0B86" authority="Alluaud (1899 b)" authorityName="Alluaud" authorityYear="1899" box="[332,767,1669,1693]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B0E34B53305AFFB3FEB9F917BF270B86" box="[332,580,1669,1693]" italics="true" pageId="35" pageNumber="236">Madecassina maculata</emphasis>
<bibRefCitation id="E606EAB0305AFFB3FDB9F917BF9C0B86" author="ALLUAUD, C." box="[588,767,1669,1693]" pageId="35" pageNumber="236" pagination="341 - 344" refId="ref28787" refString="ALLUAUD, C. 1899 b. Contributions a la faune entomologique de la Region malagache. 6 e Note. Bulletin de la Societe Entomologique de France 1899: 341 - 344." type="journal article" year="1899">Alluaud (1899b)</bibRefCitation>
</taxonomicName>
in Jeannel’s (1949) key based on the following combination of features: body black, forebody without metallic reflection, pronotum punctate laterally, and elytra shiny with four pale spots. It differs from members of
<taxonomicName id="4597ECC2305CFFB5FC06FF4EB9190DE8" authorityName="Alluaud" authorityYear="1899" box="[1011,1146,219,243]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B0E34B53305CFFB5FC06FF4EB9190DE8" box="[1011,1146,219,243]" italics="true" pageId="37" pageNumber="238">M. maculata</emphasis>
</taxonomicName>
in having a smaller body size (SBL = 5.5–6.0 mm in
<taxonomicName id="4597ECC2305CFFB5FD56FF6EBE480C0F" authorityName="Alluaud" authorityYear="1899" box="[675,811,252,276]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B0E34B53305CFFB5FD56FF6EBE480C0F" box="[675,811,252,276]" italics="true" pageId="37" pageNumber="238">M. maculata</emphasis>
</taxonomicName>
), the pronotum punctate laterally and laterobasally only, with conspicuous transverse grooves and ridges on disc (in
<taxonomicName id="4597ECC2305CFFB5FC1DFE8FB90D0C2F" authorityName="Alluaud" authorityYear="1899" box="[1000,1134,284,308]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B0E34B53305CFFB5FC1DFE8FB90D0C2F" box="[1000,1134,284,308]" italics="true" pageId="37" pageNumber="238">M. maculata</emphasis>
</taxonomicName>
the pronotum is punctate on the disc also, between the ridges, in contrast to what Jeannel said in his key), elytral intervals shinier due to more shallowly impressed isodiametric microsculpture (elytral intervals duller due to more deeply impressed isodiametric microsculpture in
<taxonomicName id="4597ECC2305CFFB5FC27FEEDB9380C8D" authorityName="Alluaud" authorityYear="1899" box="[978,1115,382,406]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B0E34B53305CFFB5FC27FEEDB9380C8D" box="[978,1115,382,406]" italics="true" pageId="37" pageNumber="238">M. maculata</emphasis>
</taxonomicName>
), the elytral spots much more distinct (much less so in
<taxonomicName id="4597ECC2305CFFB5FD65FE0DBE7B0CAC" authorityName="Alluaud" authorityYear="1899" box="[656,792,415,439]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B0E34B53305CFFB5FD65FE0DBE7B0CAC" box="[656,792,415,439]" italics="true" pageId="37" pageNumber="238">M. maculata</emphasis>
</taxonomicName>
) and median lobe of male genitalia with apex hooked dorsally and thicker in lateral aspect (
<figureCitation id="1AAC8BC4305CFFB5FD16FE2DBE200CCC" box="[739,835,447,471]" captionStart="FIGURE" captionStartId="36.[155,167,1570,1589]" captionTargetBox="[144,1176,216,1552]" captionTargetId="figure-53@36.[120,1176,216,1567]" captionTargetPageId="36" captionText="FIGURE 20. Digital images of holotype male of Madecassina quadrimaculata Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." pageId="37" pageNumber="238">Fig. 20C</figureCitation>
) and slightly swollen and bent right in dorsal aspect (
<figureCitation id="1AAC8BC4305CFFB5FE9FFE72BCAB0CEC" box="[362,456,480,504]" captionStart="FIGURE" captionStartId="36.[155,167,1570,1589]" captionTargetBox="[144,1176,216,1552]" captionTargetId="figure-53@36.[120,1176,216,1567]" captionTargetPageId="36" captionText="FIGURE 20. Digital images of holotype male of Madecassina quadrimaculata Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." pageId="37" pageNumber="238">Fig. 20D</figureCitation>
) (apex hooked but thinner in lateral aspect and more parallel-sided and straight in dorsal aspect in
<taxonomicName id="4597ECC2305CFFB5FE3FFD93BF310F03" authorityName="Alluaud" authorityYear="1899" box="[458,594,512,536]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B0E34B53305CFFB5FE3FFD93BF310F03" box="[458,594,512,536]" italics="true" pageId="37" pageNumber="238">M. maculata</emphasis>
</taxonomicName>
males) (see Jeannel 1949, Fig. 474f).
</paragraph>
<caption id="D6E8C7C9305DFFB4FF6EF9B0BC630B73" pageId="36" pageNumber="237" startId="36.[155,167,1570,1589]" targetBox="[144,1176,216,1552]" targetPageId="36" targetType="figure">
<paragraph id="82289741305DFFB4FF6EF9B0BC630B73" blockId="36.[123,1173,1570,1640]" pageId="36" pageNumber="237">
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20. Digital images of holotype male of
<taxonomicName id="4597ECC2305DFFB4FDDBF9B0BE8B0B2E" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[558,1000,1570,1589]" class="Insecta" family="Carabidae" genus="Madecassina" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="quadrimaculata" status="sp. nov.">
<emphasis id="B0E34B53305DFFB4FDDBF9B0BE4F0B2E" box="[558,812,1570,1589]" italics="true" pageId="36" pageNumber="237">Madecassina quadrimaculata</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628305DFFB4FC18F9B0B94C0B2E" box="[1005,1071,1570,1589]" pageId="36" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels C–D. Aedeagus of male genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm.
</paragraph>
</caption>
<paragraph id="82289741305CFFB5FF5EFDB3B9690F22" blockId="37.[123,1174,219,634]" box="[171,1034,545,570]" pageId="37" pageNumber="238">
<emphasis id="B0E34B53305CFFB5FF5EFDB3BC920F2C" bold="true" box="[171,497,545,569]" pageId="37" pageNumber="238">
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<smallCapsWord id="84CE019D305CFFB5FEA8FDB6BC920F2C" baselines="562" box="[349,497,548,567]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="37" pageNumber="238">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E3305CFFB5FC8AFDB0BECE0F21" box="[895,941,546,570]" pageId="37" pageNumber="238">type</typeStatus>
locality.
</paragraph>
<paragraph id="82289741305CFFB5FF5EFDD0B9710F61" blockId="37.[123,1174,219,634]" pageId="37" pageNumber="238">
<emphasis id="B0E34B53305CFFB5FF5EFDD0BCC00F43" bold="true" box="[171,419,578,602]" pageId="37" pageNumber="238">
<smallCapsWord id="84CE019D305CFFB5FF5EFDD0BC6A0F43" baselines="596,595" box="[171,265,578,602]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="37" pageNumber="238">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D305CFFB5FEFAFDD7BCC00F43" baselines="595" box="[271,419,581,600]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="37" pageNumber="238">DISTRIBUTION</smallCapsWord>
</emphasis>
.— The unique
<typeStatus id="5D2C29E3305CFFB5FDBCFDD0BFC50F41" box="[585,678,578,602]" pageId="37" pageNumber="238" type="holotype">holotype</typeStatus>
was collected in montane rainforest near Tsinjorano village at a distance of
<quantity id="456F3AA4305CFFB5FE35FDF0BF670F61" box="[448,516,610,634]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="238" unit="m" value="100.0">100 m</quantity>
inside the forest edge at an elevation of
<quantity id="456F3AA4305CFFB5FC4FFDF0B9680F61" box="[954,1035,610,634]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="37" pageNumber="238" unit="m" value="1050.0">1050 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="E282F59D7D479621CA026AA46FBCAA3C">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:name id="93CC98E11C513A96C9A4C6C0DA63879A" type="personal">
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<mods:namePart id="10176EEA54F057484C5F57FAECC14748">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="687E3532114272F232A1789D879033B2">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:namePart id="3B62D1635A4BCFE51900EFB409DD4CD9">Rainio, Johanna</mods:namePart>
<mods:affiliation id="00D50AE6A7D92EB1AE697C3B6F560399">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
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<heading id="D960202D305CFFB5FF8EFB4BBF9809EA" box="[123,763,1240,1265]" fontSize="10" level="2" pageId="37" pageNumber="238" reason="4">
<emphasis id="B0E34B53305CFFB5FF8EFB4BBF9809EA" bold="true" box="[123,763,1240,1265]" pageId="37" pageNumber="238">
<taxonomicName id="4597ECC2305CFFB5FF8EFB4BBFFB09EA" ID-CoL="77WH4" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,664,1240,1265]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B53305CFFB5FF8EFB4BBCE409EB" bold="true" box="[123,391,1240,1265]" italics="true" pageId="37" pageNumber="238">Pristacrus ranomafanae</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
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<subSubSection id="CA8DC4CA305CFFB5FF8EFB6BBE010820" pageId="37" pageNumber="238" type="description">
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–22
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<emphasis id="B0E34B53305CFFB5FF8EFAB4BC4F0821" box="[123,300,1317,1339]" italics="true" pageId="37" pageNumber="238">
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<taxonomicNameLabel id="ABD0F628305CFFB5FF10FAB4BC7A0820" box="[229,281,1318,1339]" pageId="37" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
1
</emphasis>
; Rainio 2009: 33, Rainio 2012: 73 (informal designation).
</paragraph>
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.—
<materialsCitation id="32FF9D1C305CFFB5FE77FACDBE670B20" collectingDate="2001-01-02" collectingDateMax="2001-01-22" collectingDateMin="2001-01-02" collectorName="D. H. &amp; K. M Kavanugh &amp; R. L. Brett &amp; E. Elsom &amp; F. Vargas &amp; R. Ranaivosolo &amp; E. F. Randrianirina &amp; N. Rasoamananana &amp; T. J. Ravelomanana &amp; H. C. Raveloson" country="Madagascar" elevation="900" latitude="-21.25041" location="Talatakely area" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" pageId="37" pageNumber="238" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E3305CFFB5FE77FACDBC86086C" box="[386,485,1375,1399]" pageId="37" pageNumber="238" type="holotype">Holotype</typeStatus>
(
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), a male, in CAS, labeled: “CASENT 1003833”/ “
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,
<collectingRegion id="405359A3305CFFB5FEA0FA12BF240883" box="[341,583,1408,1432]" country="Madagascar" name="Fianarantsoa" pageId="37" pageNumber="238">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B305CFFB5FDADFA12BEE30883" box="[600,896,1408,1432]" pageId="37" pageNumber="238">Ranomafana National Park</collectingMunicipality>
,
<location id="8748C19A305CFFB5FC64FA12B955088C" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305CFFB5FC64FA12B955088C" box="[913,1078,1408,1432]" country="Madagascar" latitude="-21.25041" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" name="Talatakely area" pageId="37" pageNumber="238" stateProvince="Fianarantsoa">Talatakely area</location>
,
<quantity id="456F3AA4305CFFB5FBB2FA12B9EC088C" box="[1095,1167,1408,1432]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="37" pageNumber="238" unit="m" value="900.0">
<elevation id="09BA7072305CFFB5FBB2FA12B9EC088C" box="[1095,1167,1408,1432]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="37" pageNumber="238" unit="m" value="900.0">900 m</elevation>
</quantity>
, mixed tropical forest,
<date id="F629B181305CFFB5FE9EFA33BF5708A2" box="[363,564,1441,1465]" pageId="37" pageNumber="238" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">
<collectingDate id="E66D4869305CFFB5FE9EFA33BF5708A2" box="[363,564,1441,1465]" pageId="37" pageNumber="238" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">2-22 January 2001</collectingDate>
</date>
,”/ “
<geoCoordinate id="E7A3F186305CFFB5FD96FA33BF8308A2" box="[611,736,1440,1465]" degrees="21. 25041" direction="south" orientation="latitude" pageId="37" pageNumber="238" precision="1" value="-21.25041">21.25041°S</geoCoordinate>
/
<geoCoordinate id="E7A3F186305CFFB5FD05FA33BE0C08A3" box="[752,879,1440,1465]" degrees="47. 41945" direction="east" orientation="longitude" pageId="37" pageNumber="238" precision="1" value="47.41945">47.41945°E</geoCoordinate>
, Stop# DHK-01-001,
<collectorName id="2F62F297305CFFB5FB97FA33BC2808C2" pageId="37" pageNumber="238">D.H. &amp; K.M Kavanugh</collectorName>
,
<collectorName id="2F62F297305CFFB5FE95FA50BCB108C2" box="[352,466,1474,1497]" pageId="37" pageNumber="238">R.L. Brett</collectorName>
,
<collectorName id="2F62F297305CFFB5FE1DFA50BF3208C2" box="[488,593,1473,1497]" pageId="37" pageNumber="238">E. Elsom</collectorName>
,
<collectorName id="2F62F297305CFFB5FD93FA50BFAC08C1" box="[614,719,1474,1498]" pageId="37" pageNumber="238">F. Vargas</collectorName>
,
<collectorName id="2F62F297305CFFB5FD11FA50BEEC08C2" box="[740,911,1473,1497]" pageId="37" pageNumber="238">R. Ranaivosolo</collectorName>
,”/ “
<collectorName id="2F62F297305CFFB5FC31FA50B9EC08C2" box="[964,1167,1473,1497]" pageId="37" pageNumber="238">E.F. Randrianirina</collectorName>
,
<collectorName id="2F62F297305CFFB5FF8EFA70BC2408E2" box="[123,327,1506,1529]" pageId="37" pageNumber="238">N. Rasoamananana</collectorName>
,
<collectorName id="2F62F297305CFFB5FEA6FA70BF7E08E1" box="[339,541,1506,1530]" pageId="37" pageNumber="238">T.J. Ravelomanana</collectorName>
, and
<collectorName id="2F62F297305CFFB5FDA3FA70BF9C08E1" box="[598,767,1506,1530]" pageId="37" pageNumber="238">H.C. Raveloson</collectorName>
collectors”/ “collected at night on surface of rotting logs on forest floor or suspended just above it”/ “
<typeStatus id="5D2C29E3305CFFB5FC7FF991B9740B00" box="[906,1047,1539,1563]" pageId="37" pageNumber="238" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC2305CFFB5FBD3F991BC990B20" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B53305CFFB5FBD3F991BC6A0B20" italics="true" pageId="37" pageNumber="238">Pristacrus ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628305CFFB5FDF1F9B6BF230B20" box="[516,576,1572,1595]" pageId="37" pageNumber="238" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label]
</materialsCitation>
.
<materialsCitation id="32FF9D1C305CFFB5FCE1F9B6BC210B47" collectionCode="CAS" country="Madagascar" pageId="37" pageNumber="238" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="5D2C29E3305CFFB5FCE1F9B6BE1F0B27" box="[788,892,1572,1596]" pageId="37" pageNumber="238" type="paratype">Paratypes</typeStatus>
(total of 19):
<specimenCount id="94915CC8305CFFB5FBEEF9B1B9050B20" box="[1051,1126,1571,1595]" count="1" pageId="37" pageNumber="238" type="male">1 male</specimenCount>
and
<specimenCount id="94915CC8305CFFB5FF8EF9D6BDBA0B47" box="[123,217,1604,1628]" count="1" pageId="37" pageNumber="238" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F84305CFFB5FEF0F9D6BC5A0B47" box="[261,313,1604,1628]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="37" pageNumber="238" type="Museum">CAS</collectionCode>
)
</materialsCitation>
<materialsCitation id="32FF9D1C305CFFB5FEBCF9D6BF0D0B67" collectionCode="CAS" country="Madagascar" pageId="37" pageNumber="238" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="paratype">
labelled “CASENT 1003832” or “CASENT 1003831”, respectively, otherwise labeled same as
<typeStatus id="5D2C29E3305CFFB5FEC7F9F6BCEC0B67" box="[306,399,1636,1660]" pageId="37" pageNumber="238" type="holotype">holotype</typeStatus>
;
<specimenCount id="94915CC8305CFFB5FE55F9F6BF630B67" box="[416,512,1636,1660]" count="1" pageId="37" pageNumber="238" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F84305CFFB5FDC4F9F6BF060B67" box="[561,613,1636,1660]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="37" pageNumber="238" type="Museum">CAS</collectionCode>
)
</materialsCitation>
<materialsCitation id="32FF9D1C305CFFB6FD82F9F6B9F60B0C" collectingDate="2001-01-02" collectingDateMax="2001-01-22" collectingDateMin="2001-01-02" collectionCode="CAS" collectorName="D. H. &amp; K. M Kavanugh &amp; R. L. Brett &amp; E. Elsom &amp; F. Vargas &amp; R. Ranaivosolo &amp; E. F. Randrianirina &amp; N. Rasoamananana &amp; T. J. Ravelomanana &amp; H. C. Raveloson" country="Madagascar" elevation="900" lastPageId="38" lastPageNumber="239" latitude="-21.25041" location="Talatakely area" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" pageId="37" pageNumber="238" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
labeled “CASENT 1004470”/ “
<collectingCountry id="FA80D7D1305CFFB5FC24F9F6B9EC0B67" box="[977,1167,1636,1660]" name="Madagascar" pageId="37" pageNumber="238">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A3305CFFB5FF8EF917BC090B86" box="[123,362,1669,1693]" country="Madagascar" name="Fianarantsoa" pageId="37" pageNumber="238">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B305CFFB5FE8DF917BFFF0B86" box="[376,668,1669,1693]" pageId="37" pageNumber="238">Ranomafana National Park</collectingMunicipality>
,
<location id="8748C19A305CFFB5FD5CF917BE2F0B87" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305CFFB5FD5CF917BE2F0B87" box="[681,844,1669,1693]" country="Madagascar" latitude="-21.25041" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" name="Talatakely area" pageId="37" pageNumber="238" stateProvince="Fianarantsoa">Talatakely area</location>
,
<quantity id="456F3AA4305CFFB5FCAEF917BEC30B87" box="[859,928,1669,1693]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="37" pageNumber="238" unit="m" value="900.0">
<elevation id="09BA7072305CFFB5FCAEF917BEC30B87" box="[859,928,1669,1693]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="37" pageNumber="238" unit="m" value="900.0">900 m</elevation>
</quantity>
, mixed tropical forest,
<date id="F629B181305CFFB5FF8EF934BC5C0BA5" box="[123,319,1702,1726]" pageId="37" pageNumber="238" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">
<collectingDate id="E66D4869305CFFB5FF8EF934BC5C0BA5" box="[123,319,1702,1726]" pageId="37" pageNumber="238" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">2-22 January 2001</collectingDate>
</date>
,”/ “
<geoCoordinate id="E7A3F186305CFFB5FE9EF934BC8A0BA5" box="[363,489,1701,1726]" degrees="21. 25041" direction="south" orientation="latitude" pageId="37" pageNumber="238" precision="1" value="-21.25041">21.25041°S</geoCoordinate>
/
<geoCoordinate id="E7A3F186305CFFB5FE03F934BF160BA6" box="[502,629,1701,1726]" degrees="47. 41945" direction="east" orientation="longitude" pageId="37" pageNumber="238" precision="1" value="47.41945">47.41945°E</geoCoordinate>
, Stop# DHK-01-001,
<collectorName id="2F62F297305CFFB5FC95F934B9390BA6" box="[864,1114,1701,1726]" pageId="37" pageNumber="238">D.H. &amp; K.M Kavanugh</collectorName>
,
<collectorName id="2F62F297305CFFB6FB93F934BDD308CD" lastPageId="38" lastPageNumber="239" pageId="37" pageNumber="238">R.L. Brett</collectorName>
,
<collectorName id="2F62F297305FFFB6FF35FA2DBC4708CC" box="[192,292,1471,1495]" pageId="38" pageNumber="239">E. Elsom</collectorName>
,
<collectorName id="2F62F297305FFFB6FEC1FA2DBCFB08CC" box="[308,408,1471,1495]" pageId="38" pageNumber="239">F. Vargas</collectorName>
,
<collectorName id="2F62F297305FFFB6FE52FA2DBF2D08CC" box="[423,590,1471,1495]" pageId="38" pageNumber="239">R. Ranaivosolo</collectorName>
,”/ “
<collectorName id="2F62F297305FFFB6FD88FA2DBE2708CC" box="[637,836,1471,1495]" pageId="38" pageNumber="239">E.F. Randrianirina</collectorName>
,
<collectorName id="2F62F297305FFFB6FCA6FA2DB94008CD" box="[851,1059,1471,1494]" pageId="38" pageNumber="239">N. Rasoamananana</collectorName>
,
<collectorName id="2F62F297305FFFB6FBC7FA2DBD8D08EC" pageId="38" pageNumber="239">T.J. Ravelomanana</collectorName>
, and
<collectorName id="2F62F297305FFFB6FED2FA4DBCB308EC" box="[295,464,1503,1527]" pageId="38" pageNumber="239">H.C. Raveloson</collectorName>
collectors”/ “collected in daytime from large accumulation of dead leaf, twig, and branch debris on and extended up to
<quantity id="456F3AA4305FFFB6FD31FA6DBF920B0D" box="[708,753,1535,1559]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="38" pageNumber="239" unit="m" value="2.0">
<elevation id="09BA7072305FFFB6FD31FA6DBF920B0D" box="[708,753,1535,1559]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="38" pageNumber="239" unit="m" value="2.0">2 m</elevation>
</quantity>
above forest floor”;
<specimenCount id="94915CC8305FFFB6FC29FA6DB9440B0C" box="[988,1063,1535,1559]" count="1" pageId="38" pageNumber="239" type="male">1 male</specimenCount>
(in
<collectionCode id="E4860F84305FFFB6FBACFA6DB9EE0B0C" box="[1113,1165,1535,1559]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="38" pageNumber="239" type="Museum">CAS</collectionCode>
)
</materialsCitation>
<materialsCitation id="32FF9D1C305FFFB7FF8EF98DBF0E0DE8" collectingDate="2001-01-02" collectingDateMax="2001-01-22" collectingDateMin="2001-01-02" collectorName="D. H. &amp; K. M Kavanugh &amp; R. L. Brett &amp; E. Elsom &amp; F. Vargas &amp; R. Ranaivosolo &amp; E. F. Randrianirina &amp; N. Rasoamananana &amp; T. J. Ravelomanana &amp; H. C. Raveloson" country="Madagascar" elevation="900" lastPageId="39" lastPageNumber="240" latitude="-21.25041" location="Talatakely area" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" pageId="38" pageNumber="239" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
labeled “CASENT 1002838”/ “
<collectingCountry id="FA80D7D1305FFFB6FE22F98DBFF60B2C" box="[471,661,1567,1591]" name="Madagascar" pageId="38" pageNumber="239">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A3305FFFB6FD53F98DBEF40B2C" box="[678,919,1567,1591]" country="Madagascar" name="Fianarantsoa" pageId="38" pageNumber="239">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B305FFFB6FC5DF98DBDCF0B4C" pageId="38" pageNumber="239">Ranomafana National Park</collectingMunicipality>
,
<location id="8748C19A305FFFB6FF36F9ADBC120B4D" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305FFFB6FF36F9ADBC120B4D" box="[195,369,1599,1623]" country="Madagascar" latitude="-21.25041" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" name="Talatakely area" pageId="38" pageNumber="239" stateProvince="Fianarantsoa">Talatakely area</location>
,
<quantity id="456F3AA4305FFFB6FE7CF9ADBCB40B4D" box="[393,471,1599,1623]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="38" pageNumber="239" unit="m" value="900.0">
<elevation id="09BA7072305FFFB6FE7CF9ADBCB40B4D" box="[393,471,1599,1623]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="38" pageNumber="239" unit="m" value="900.0">900 m</elevation>
</quantity>
, mixed tropical forest,
<date id="F629B181305FFFB6FD0EF9ADBEBB0B4C" box="[763,984,1599,1623]" pageId="38" pageNumber="239" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">
<collectingDate id="E66D4869305FFFB6FD0EF9ADBEBB0B4C" box="[763,984,1599,1623]" pageId="38" pageNumber="239" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">2-22 January 2001</collectingDate>
</date>
,”/ “
<geoCoordinate id="E7A3F186305FFFB6FBFAF9ADB9ED0B4C" box="[1039,1166,1599,1623]" degrees="21. 25041" direction="south" orientation="latitude" pageId="38" pageNumber="239" precision="1" value="-21.25041">21.25041°S</geoCoordinate>
/
<geoCoordinate id="E7A3F186305FFFB6FF8EF9CDBD9A0B6D" box="[123,249,1631,1655]" degrees="47. 41945" direction="east" orientation="longitude" pageId="38" pageNumber="239" precision="1" value="47.41945">47.41945°E</geoCoordinate>
, Stop# DHK-01-001,
<collectorName id="2F62F297305FFFB6FE08F9CDBE6D0B6C" box="[509,782,1631,1655]" pageId="38" pageNumber="239">D.H. &amp; K.M Kavanugh</collectorName>
,
<collectorName id="2F62F297305FFFB6FCD7F9CDBEF70B6D" box="[802,916,1631,1654]" pageId="38" pageNumber="239">R.L. Brett</collectorName>
,
<collectorName id="2F62F297305FFFB6FC5CF9CDB9720B6C" box="[937,1041,1631,1655]" pageId="38" pageNumber="239">E. Elsom</collectorName>
,
<collectorName id="2F62F297305FFFB6FBD3F9CDB9EC0B6C" box="[1062,1167,1631,1655]" pageId="38" pageNumber="239">F. Vargas</collectorName>
,
<collectorName id="2F62F297305FFFB6FF8EF9EDBC430B8C" box="[123,288,1663,1687]" pageId="38" pageNumber="239">R. Ranaivosolo</collectorName>
,”/ “
<collectorName id="2F62F297305FFFB6FEBBF9EDBF700B8C" box="[334,531,1663,1687]" pageId="38" pageNumber="239">E.F. Randrianirina</collectorName>
,
<collectorName id="2F62F297305FFFB6FDD4F9EDBF930B8D" box="[545,752,1663,1686]" pageId="38" pageNumber="239">N. Rasoamananana</collectorName>
,
<collectorName id="2F62F297305FFFB6FD0BF9EDBEA90B8C" box="[766,970,1663,1687]" pageId="38" pageNumber="239">T.J. Ravelomanana</collectorName>
, and
<collectorName id="2F62F297305FFFB6FBF2F9EDBDC30BAD" pageId="38" pageNumber="239">H.C. Raveloson</collectorName>
collectors”/ “collected in daytime under loss bark of rotting log logs on forest floor or suspended just above it”;
<specimenCount id="94915CC8305EFFB7FE98FF49BCD50DE8" box="[365,438,219,243]" count="1" pageId="39" pageNumber="240" type="male">1 male</specimenCount>
and
<specimenCount id="94915CC8305EFFB7FE1EFF49BF0E0DE8" box="[491,621,219,243]" count="2" pageId="39" pageNumber="240" type="female">two females</specimenCount>
</materialsCitation>
</paragraph>
<caption id="D6E8C7C9305FFFB6FF6EFAFBBC80088D" pageId="38" pageNumber="239" startId="38.[155,167,1385,1404]" targetBox="[120,1176,216,1353]" targetPageId="38" targetType="figure">
<paragraph id="82289741305FFFB6FF6EFAFBBC80088D" blockId="38.[123,1173,1385,1430]" pageId="38" pageNumber="239">
<smallCapsWord id="84CE019D305FFFB6FF6EFAFBBDBB0861" baselines="1399,1398" box="[155,216,1385,1404]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="38" pageNumber="239">FIGURE</smallCapsWord>
21. Digital images of holotype male of
<taxonomicName id="4597ECC2305FFFB6FDC4FAFBBEDE0867" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[561,957,1385,1404]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="239" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B53305FFFB6FDC4FAFBBE620867" box="[561,769,1385,1404]" italics="true" pageId="38" pageNumber="239">Pristacrus ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628305FFFB6FC36FAFBB9660867" box="[963,1029,1385,1404]" pageId="38" pageNumber="239" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B Labels. Scale line = 1.0 mm.
</paragraph>
</caption>
<paragraph id="82289741305EFFB7FF8EFF69BF0E0C08" blockId="39.[123,1174,219,1715]" box="[123,621,251,275]" pageId="39" pageNumber="240">
(in
<collectionCode id="E4860F84305EFFB7FF45FF69BD860C08" box="[176,229,251,275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="39" pageNumber="240" type="Museum">CAS</collectionCode>
and
<collectionCode id="E4860F84305EFFB7FECCFF69BC120C09" box="[313,369,251,274]" pageId="39" pageNumber="240">MZF</collectionCode>
) labeled “CASENT
</paragraph>
<paragraph id="82289741305EFFB7FF8EFE89BF0D0C28" blockId="39.[123,1174,219,1715]" box="[123,622,283,307]" pageId="39" pageNumber="240">1003752”, “CASENT 1003751” or “CASENT</paragraph>
<paragraph id="82289741305EFFB7FF8EFEA9BF0D0C49" blockId="39.[123,1174,219,1715]" box="[123,622,315,339]" pageId="39" pageNumber="240">
1003753”, respectively/ “
<collectingCountry id="FA80D7D1305EFFB7FE53FEA9BF050C48" box="[422,614,315,339]" name="Madagascar" pageId="39" pageNumber="240">MADAGASCAR</collectingCountry>
,
</paragraph>
<paragraph id="82289741305EFFB7FF8EFEC9BF0D0C68" blockId="39.[123,1174,219,1715]" box="[123,622,347,371]" pageId="39" pageNumber="240">
<materialsCitation id="32FF9D1C305EFFB7FF8EFEC9BF0D0C68" box="[123,622,347,371]" location="Ranomafana" pageId="39" pageNumber="240" specimenCount="1" stateProvince="Fianarantsoa">
<collectingRegion id="405359A3305EFFB7FF8EFEC9BC0E0C68" box="[123,365,347,371]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A305EFFB7FE8AFEC9BF650C68" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FE8AFEC9BF650C68" box="[383,518,347,371]" name="Ranomafana" pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Ranomafana</location>
National
</materialsCitation>
</paragraph>
<paragraph id="82289741305EFFB7FF8EFEE9BF0E0CA8" blockId="39.[123,1174,219,1715]" pageId="39" pageNumber="240">
<materialsCitation id="32FF9D1C305EFFB7FF8EFEE9BF050CA8" collectingDate="2001-01-02" collectingDateMax="2001-01-22" collectingDateMin="2001-01-02" collectingRegion="Fianarantsoa Province" elevation="900" latitude="-21.25041" location="Park" municipality="Talatakely area" pageId="39" pageNumber="240" specimenCount="1" stateProvince="Fianarantsoa Province">
<location id="8748C19A305EFFB7FF8EFEE9BDCF0C88" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FF8EFEE9BDCF0C88" box="[123,172,379,403]" latitude="-21.25041" municipality="Talatakely area" name="Park" pageId="39" pageNumber="240" stateProvince="Fianarantsoa Province">Park</location>
,
<collectingMunicipality id="624C0D3B305EFFB7FF48FEE9BC000C89" box="[189,355,379,403]" pageId="39" pageNumber="240">Talatakely area</collectingMunicipality>
,
<quantity id="456F3AA4305EFFB7FE80FEE9BCDE0C89" box="[373,445,379,403]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">
<elevation id="09BA7072305EFFB7FE80FEE9BCDE0C89" box="[373,445,379,403]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">900 m</elevation>
</quantity>
, mixed tropical forest,
<date id="F629B181305EFFB7FF25FE09BCCD0CA8" box="[208,430,411,435]" pageId="39" pageNumber="240" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">
<collectingDate id="E66D4869305EFFB7FF25FE09BCCD0CA8" box="[208,430,411,435]" pageId="39" pageNumber="240" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">2-22 January 2001</collectingDate>
</date>
,”/ “
<geoCoordinate id="E7A3F186305EFFB7FE13FE09BF050CA8" box="[486,614,411,435]" degrees="21. 25041" direction="south" orientation="latitude" pageId="39" pageNumber="240" precision="1" value="-21.25041">21.25041°S</geoCoordinate>
</materialsCitation>
/
</paragraph>
<paragraph id="82289741305EFFB7FF8EFE29BF0D0CC9" blockId="39.[123,1174,219,1715]" box="[123,622,443,467]" pageId="39" pageNumber="240">
<geoCoordinate id="E7A3F186305EFFB7FF8EFE29BD9A0CC9" box="[123,249,443,467]" degrees="47. 41945" direction="east" orientation="longitude" pageId="39" pageNumber="240" precision="1" value="47.41945">47.41945°E</geoCoordinate>
, Stop# DHK-01-001, D.H. &amp; K.M
</paragraph>
<paragraph id="82289741305EFFB7FF8EFE49BF0D0CE9" blockId="39.[123,1174,219,1715]" box="[123,622,475,499]" pageId="39" pageNumber="240">Kavanugh, R.L. Brett, E. Elsom, F. Vargas,</paragraph>
<paragraph id="82289741305EFFB7FF8EFE69BF0D0F09" blockId="39.[123,1174,219,1715]" box="[123,622,507,531]" pageId="39" pageNumber="240">R. Ranaivosolo,”/ “E.F. Randrianirina,</paragraph>
<paragraph id="82289741305EFFB7FF8EFD89BF0D0F28" blockId="39.[123,1174,219,1715]" box="[123,622,539,563]" pageId="39" pageNumber="240">N. Rasoamananana, T.J. Ravelomanana, and</paragraph>
<paragraph id="82289741305EFFB7FF8EFDA9BF0D0F88" blockId="39.[123,1174,219,1715]" pageId="39" pageNumber="240">
H.C. Raveloson collectors”/ “collected in daytime on underside of rotting log logs on forest floor or suspended just above it”;
<specimenCount id="94915CC8305EFFB7FE0FFDE9BF250F88" box="[506,582,635,659]" count="1" pageId="39" pageNumber="240" type="male">1 male</specimenCount>
(in
</paragraph>
<paragraph id="82289741305EFFB7FF8EFD09BE6B0E18" blockId="39.[123,1174,219,1715]" pageId="39" pageNumber="240">
<materialsCitation id="32FF9D1C305EFFB7FF8EFD09BE600E18" collectionCode="CAS" collectorName="Rainio &amp; A. Left &amp; B. Dorsal &amp; Scale" county="Digital" elevation="900" location="Talatakely area" municipality="National Park" pageId="39" pageNumber="240" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<collectionCode id="E4860F84305EFFB7FF8EFD09BDCC0FA8" box="[123,175,667,691]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="39" pageNumber="240" type="Museum">CAS</collectionCode>
) labeled “CASENT 1004313”/ “MADA-
<smallCapsWord id="84CE019D305EFFB7FD5EFD31BF8B0FAE" baselines="689,689" box="[683,744,675,694]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="39" pageNumber="240">FIGURE</smallCapsWord>
22.
<collectingCounty id="6B49EFCD305EFFB7FCE3FD31BE320FAD" box="[790,849,675,694]" pageId="39" pageNumber="240">Digital</collectingCounty>
images of aedeagus of genitalia of GASCAR,
<collectingRegion id="405359A3305EFFB7FF0BFD29BC960FC8" box="[254,501,699,723]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa Province</collectingRegion>
, Ranoma- male
<typeStatus id="5D2C29E3305EFFB7FD35FD2EBE680FD4" box="[704,779,700,719]" pageId="39" pageNumber="240" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4597ECC2305EFFB7FCC0FD2FB91A0FD4" authority="Kavanaugh" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[821,1145,700,720]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53305EFFB7FCC0FD2FB96F0FCB" box="[821,1036,701,720]" italics="true" pageId="39" pageNumber="240">Pristacrus ranomafanae</emphasis>
Kavanaugh
</taxonomicName>
&amp; fana
<collectingMunicipality id="624C0D3B305EFFB7FF40FD49BC2C0FE8" box="[181,335,731,755]" pageId="39" pageNumber="240">National Park</collectingMunicipality>
,
<location id="8748C19A305EFFB7FE97FD49BF6A0FE9" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FE97FD49BF6A0FE9" box="[354,521,731,755]" county="Digital" municipality="National Park" name="Talatakely area" pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Talatakely area</location>
,
<quantity id="456F3AA4305EFFB7FDE8FD49BF040FE9" box="[541,615,731,755]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">
<elevation id="09BA7072305EFFB7FDE8FD49BF040FE9" box="[541,615,731,755]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">900 m</elevation>
</quantity>
,
<collectorName id="2F62F297305EFFB7FD7EFD44BFA70FF2" box="[651,708,726,745]" pageId="39" pageNumber="240">Rainio</collectorName>
<taxonomicNameLabel id="ABD0F628305EFFB7FD3CFD45BE680FF1" box="[713,779,727,746]" pageId="39" pageNumber="240" rank="species">sp. nov.</taxonomicNameLabel>
<collectorName id="2F62F297305EFFB7FCE5FD44BE2D0FF2" box="[784,846,726,746]" pageId="39" pageNumber="240">A. Left</collectorName>
lateral aspect;
<collectorName id="2F62F297305EFFB7FC3AFD45B9420FF2" box="[975,1057,726,746]" pageId="39" pageNumber="240">B. Dorsal</collectorName>
aspect.
<collectorName id="2F62F297305EFFB7FB92FD44B9F60FF2" box="[1127,1173,726,745]" pageId="39" pageNumber="240">Scale</collectorName>
line =
<quantity id="456F3AA4305EFFB7FD34FD62BE600E18" box="[705,771,752,771]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="39" pageNumber="240" unit="mm" value="0.5">0.5 mm</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="82289741305EFFB7FF8EFD69BF0E0E08" blockId="39.[123,1174,219,1715]" box="[123,621,763,787]" pageId="39" pageNumber="240">
mixed tropical forest,
<date id="F629B181305EFFB7FE88FD69BF300E08" box="[381,595,763,787]" pageId="39" pageNumber="240" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">2-22 January 2001</date>
,”/
</paragraph>
<paragraph id="82289741305EFFB7FF8EFC89BE4108A8" blockId="39.[123,1174,219,1715]" pageId="39" pageNumber="240">
“
<geoCoordinate id="E7A3F186305EFFB7FF72FC89BC6A0E28" box="[135,265,795,819]" degrees="21. 25041" direction="south" orientation="latitude" pageId="39" pageNumber="240" precision="1" value="-21.25041">21.25041°S</geoCoordinate>
/
<geoCoordinate id="E7A3F186305EFFB7FED3FC89BCCA0E29" box="[294,425,795,819]" degrees="47. 41945" direction="east" orientation="longitude" pageId="39" pageNumber="240" precision="1" value="47.41945">47.41945°E</geoCoordinate>
, Stop# DHK-01-001, D.H. &amp; K.M Kavanugh, R.L. Brett, E. Elsom, F. Vargas, R. Ranaivosolo,”/ “E.F. Randrianirina, N. Rasoamananana, T.J. Ravelomanana, and H.C. Raveloson collectors”/ “collected in daytime from leaf litter on forest floor”;
<materialsCitation id="32FF9D1C305EFFB7FF8EFCE9BC460EE8" collectingDate="1998-04-27" collectionCode="CAS" collectorName="A D. H. Kavanaugh" country="Madagascar" elevation="900" latitude="-21.255" location="Talatakely area" longLatPrecision="126" longitude="47.431667" municipality="Ranomafana National Park" pageId="39" pageNumber="240" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa">
<specimenCount id="94915CC8305EFFB7FF8EFCE9BDBC0E88" box="[123,223,891,915]" count="1" pageId="39" pageNumber="240" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F84305EFFB7FEEDFCE9BC2F0E88" box="[280,332,891,915]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="39" pageNumber="240" type="Museum">CAS</collectionCode>
) labeled “CASENT 1051144”/ “
<collectingCountry id="FA80D7D1305EFFB7FD3DFCE9BEE50E88" box="[712,902,891,915]" name="Madagascar" pageId="39" pageNumber="240">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A3305EFFB7FC6FFCE9B9EC0E88" box="[922,1167,891,915]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B305EFFB7FF8EFC09BCC80EA8" box="[123,427,923,947]" pageId="39" pageNumber="240">Ranomafana National Park</collectingMunicipality>
,
<location id="8748C19A305EFFB7FE4AFC09BF0B0EA9" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FE4AFC09BF0B0EA9" box="[447,616,923,947]" country="Madagascar" latitude="-21.255" longLatPrecision="126" longitude="47.431667" municipality="Ranomafana National Park" name="Talatakely area" pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Talatakely area</location>
,
<quantity id="456F3AA4305EFFB7FD88FC09BFAB0EA9" box="[637,712,923,947]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">
<elevation id="09BA7072305EFFB7FD88FC09BFAB0EA9" box="[637,712,923,947]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">900 m</elevation>
</quantity>
, mixed tropical forest,
<date id="F629B181305EFFB7FC2AFC09B9E10EA8" box="[991,1154,923,947]" pageId="39" pageNumber="240" value="1998-04-27">
<collectingDate id="E66D4869305EFFB7FC2AFC09B9E10EA8" box="[991,1154,923,947]" pageId="39" pageNumber="240" value="1998-04-27">27 April 1998</collectingDate>
</date>
”/ “
<geoCoordinate id="E7A3F186305EFFB7FF72FC29BD910EC8" box="[135,242,955,979]" degrees="21" direction="south" minutes="15. 3" orientation="latitude" pageId="39" pageNumber="240" precision="92" value="-21.255">21°15.3’S</geoCoordinate>
<geoCoordinate id="E7A3F186305EFFB7FF0CFC29BC050EC9" box="[249,358,955,979]" degrees="47" direction="east" minutes="25. 9" orientation="longitude" pageId="39" pageNumber="240" precision="92" value="47.431667">47°25.9’E</geoCoordinate>
, Stop # 98-85
<collectorName id="2F62F297305EFFB7FE09FC29BFAB0EC8" box="[508,712,955,979]" pageId="39" pageNumber="240">A D.H. Kavanaugh</collectorName>
collector collected in and under rotten logs on forest floor”
</materialsCitation>
;
<materialsCitation id="32FF9D1C305EFFB7FEC3FC49BF7B0948" collectingDate="2002-04-28" collectingDateMax="2002-05-05" collectingDateMin="2002-04-28" collectionCode="CAS" collectorName="R. Harin'Hala California Acad" country="Madagascar" county="Province" elevation="1020" latitude="-21.2665" location="Belle Vue" longLatPrecision="12" longitude="47.420166" municipality="Ranomafana" pageId="39" pageNumber="240" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa">
<specimenCount id="94915CC8305EFFB7FEC3FC49BCF40EE8" box="[310,407,987,1011]" count="1" pageId="39" pageNumber="240" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F84305EFFB7FE32FC49BC980EE8" box="[455,507,987,1011]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="39" pageNumber="240" type="Museum">CAS</collectionCode>
) labeled “CASENT 8068264”/ “
<collectingCountry id="FA80D7D1305EFFB7FC9DFC49B9450EE8" box="[872,1062,987,1011]" name="Madagascar" pageId="39" pageNumber="240">MADAGASCAR</collectingCountry>
:
<collectingCounty id="6B49EFCD305EFFB7FBC2FC49B9F50EE8" box="[1079,1174,987,1011]" pageId="39" pageNumber="240">Province</collectingCounty>
<collectingRegion id="405359A3305EFFB7FF8EFC69BC600908" box="[123,259,1019,1043]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa</collectingRegion>
, Parc National
<collectingMunicipality id="624C0D3B305EFFB7FE58FC69BF570908" box="[429,564,1019,1043]" pageId="39" pageNumber="240">Ranomafana</collectingMunicipality>
,
<location id="8748C19A305EFFB7FDB7FC69BFCE0909" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FDB7FC69BFCE0909" box="[578,685,1019,1043]" country="Madagascar" county="Province" latitude="-21.2665" longLatPrecision="12" longitude="47.420166" municipality="Ranomafana" name="Belle Vue" pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Belle Vue</location>
at
<location id="8748C19A305EFFB7FD3AFC69BE5F0908" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FD3AFC69BE5F0908" box="[719,828,1019,1043]" country="Madagascar" county="Province" latitude="-21.2665" longLatPrecision="12" longitude="47.420166" municipality="Ranomafana" name="Talatakely" pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Talatakely</location>
, elev
<quantity id="456F3AA4305EFFB7FC8AFC69BEB20909" box="[895,977,1019,1043]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="39" pageNumber="240" unit="m" value="1020.0">
<elevation id="09BA7072305EFFB7FC8AFC69BEB20909" box="[895,977,1019,1043]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="39" pageNumber="240" unit="m" value="1020.0">1020 m</elevation>
</quantity>
<date id="F629B181305EFFB7FC2CFC69BDD30928" pageId="39" pageNumber="240" value="2002-04-28" valueMax="2002-05-05" valueMin="2002-04-28">
<collectingDate id="E66D4869305EFFB7FC2CFC69BDD30928" pageId="39" pageNumber="240" value="2002-04-28" valueMax="2002-05-05" valueMin="2002-04-28">28 April – 5 May 2002</collectingDate>
</date>
”/ “
<geoCoordinate id="E7A3F186305EFFB7FF23FB89BC2C0928" box="[214,335,1051,1075]" degrees="21" direction="south" minutes="15. 99" orientation="latitude" pageId="39" pageNumber="240" precision="9" value="-21.2665">21°15.99’S</geoCoordinate>
<geoCoordinate id="E7A3F186305EFFB7FEA3FB89BCB30929" box="[342,464,1051,1075]" degrees="47" direction="east" minutes="25. 21" orientation="longitude" pageId="39" pageNumber="240" precision="9" value="47.420166">47°25.21’E</geoCoordinate>
collector:
<collectorName id="2F62F297305EFFB7FDB6FB89BEE90928" box="[579,906,1051,1075]" pageId="39" pageNumber="240">R. Harin’Hala California Acad</collectorName>
of Sciences malaise secondary tropical forest MA-02-09C-27”
</materialsCitation>
;
<materialsCitation id="32FF9D1C305EFFB7FDD0FBA9BF9A09A8" collectingDate="2001-11-15" collectingDateMax="2001-11-22" collectingDateMin="2001-11-15" collectionCode="CAS" collectorName="R. Harin'Hala California Acad" country="Madagascar" county="Province" elevation="1020" latitude="-21.2665" location="Belle Vue" longLatPrecision="12" longitude="47.420166" municipality="Ranomafana" pageId="39" pageNumber="240" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa">
<specimenCount id="94915CC8305EFFB7FDD0FBA9BFE20948" box="[549,641,1083,1107]" count="1" pageId="39" pageNumber="240" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F84305EFFB7FD5FFBA9BFBD0948" box="[682,734,1083,1107]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="39" pageNumber="240" type="Museum">CAS</collectionCode>
) labeled “CASENT 8068264”/ “
<collectingCountry id="FA80D7D1305EFFB7FBCEFBA9BD840968" name="Madagascar" pageId="39" pageNumber="240">MADAGASCAR</collectingCountry>
:
<collectingCounty id="6B49EFCD305EFFB7FF03FBC9BC360968" box="[246,341,1115,1139]" pageId="39" pageNumber="240">Province</collectingCounty>
<collectingRegion id="405359A3305EFFB7FEA8FBC9BC860968" box="[349,485,1115,1139]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa</collectingRegion>
, Parc National
<collectingMunicipality id="624C0D3B305EFFB7FD7BFBC9BE760968" box="[654,789,1115,1139]" pageId="39" pageNumber="240">Ranomafana</collectingMunicipality>
,
<location id="8748C19A305EFFB7FCD1FBC9BEED0969" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FCD1FBC9BEED0969" box="[804,910,1115,1139]" country="Madagascar" county="Province" latitude="-21.2665" longLatPrecision="12" longitude="47.420166" municipality="Ranomafana" name="Belle Vue" pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Belle Vue</location>
at
<location id="8748C19A305EFFB7FC45FBC9B97E0968" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FC45FBC9B97E0968" box="[944,1053,1115,1139]" country="Madagascar" county="Province" latitude="-21.2665" longLatPrecision="12" longitude="47.420166" municipality="Ranomafana" name="Talatakely" pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Talatakely</location>
, elev
<quantity id="456F3AA4305EFFB7FB95FBC9BDF30989" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="39" pageNumber="240" unit="m" value="1020.0">
<elevation id="09BA7072305EFFB7FB95FBC9BDF30989" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="39" pageNumber="240" unit="m" value="1020.0">1020 m</elevation>
</quantity>
<date id="F629B181305EFFB7FF6DFBE9BCED0988" box="[152,398,1147,1171]" pageId="39" pageNumber="240" value="2001-11-15" valueMax="2001-11-22" valueMin="2001-11-15">
<collectingDate id="E66D4869305EFFB7FF6DFBE9BCED0988" box="[152,398,1147,1171]" pageId="39" pageNumber="240" value="2001-11-15" valueMax="2001-11-22" valueMin="2001-11-15">15-22 November 2001</collectingDate>
</date>
”/ “
<geoCoordinate id="E7A3F186305EFFB7FE43FBE9BF4C0988" box="[438,559,1147,1171]" degrees="21" direction="south" minutes="15. 99" orientation="latitude" pageId="39" pageNumber="240" precision="9" value="-21.2665">21°15.99’S</geoCoordinate>
<geoCoordinate id="E7A3F186305EFFB7FDCDFBE9BFD10989" box="[568,690,1147,1171]" degrees="47" direction="east" minutes="25. 21" orientation="longitude" pageId="39" pageNumber="240" precision="9" value="47.420166">47°25.21’E</geoCoordinate>
collector:
<collectorName id="2F62F297305EFFB7FCDCFBE9B9150988" box="[809,1142,1147,1171]" pageId="39" pageNumber="240">R. Harin’Hala California Acad</collectorName>
of Sciences malaise secondary tropical forest MA-02-09C-03”
</materialsCitation>
;
<materialsCitation id="32FF9D1C305EFFB7FCF2FB09BF8C09E8" collectingDate="1990-02-01" collectingDateMax="1990-02-09" collectingDateMin="1990-02-01" collectionCode="NMNH" collectorName="W. E. Steiner" country="Madagascar" elevation="900" location="Prov." pageId="39" pageNumber="240" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa">
<specimenCount id="94915CC8305EFFB7FCF2FB09BE2C09A8" box="[775,847,1179,1203]" count="1" pageId="39" pageNumber="240" type="male">1 male</specimenCount>
(in
<collectionCode id="E4860F84305EFFB7FC8FFB09BEA809A9" box="[890,971,1179,1202]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="39" pageNumber="240" type="Museum">NMNH</collectionCode>
) labeled “
<collectingCountry id="FA80D7D1305EFFB7FBCEFB09BD8409C8" name="Madagascar" pageId="39" pageNumber="240">MADAGASCAR</collectingCountry>
:
<location id="8748C19A305EFFB7FF00FB29BC4E09C9" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FF00FB29BC4E09C9" box="[245,301,1211,1234]" country="Madagascar" name="Prov." pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Prov.</location>
<collectingRegion id="405359A3305EFFB7FEC6FB29BCD809C8" box="[307,443,1211,1235]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa</collectingRegion>
,
<locationDeviation id="939A8E63305EFFB7FE3CFB29BFC809C8" box="[457,683,1211,1235]" location="Ranomafana" pageId="39" pageNumber="240">
<quantity id="456F3AA4305EFFB7FE3CFB29BC9C09C8" box="[457,511,1211,1235]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="39" pageNumber="240" unit="km" value="7.0">7 km</quantity>
W Ranomafana
</locationDeviation>
,
<quantity id="456F3AA4305EFFB7FD4DFB29BF9F09C9" box="[696,764,1211,1235]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">
<elevation id="09BA7072305EFFB7FD4DFB29BF9F09C9" box="[696,764,1211,1235]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">900 m</elevation>
</quantity>
<date id="F629B181305EFFB7FCF7FB29BEAA09C8" box="[770,969,1211,1235]" pageId="39" pageNumber="240" value="1990-02-01" valueMax="1990-02-09" valueMin="1990-02-01">
<collectingDate id="E66D4869305EFFB7FCF7FB29BEAA09C8" box="[770,969,1211,1235]" pageId="39" pageNumber="240" value="1990-02-01" valueMax="1990-02-09" valueMin="1990-02-01">1-9 February 1990</collectingDate>
</date>
<collectorName id="2F62F297305EFFB7FC3AFB29B93E09C8" box="[975,1117,1211,1235]" pageId="39" pageNumber="240">W. E. Steiner</collectorName>
”/ “at black light in montane rainforest near river and stream”
</materialsCitation>
;
<materialsCitation id="32FF9D1C305EFFB7FCF7FB49BEC00828" collectionCode="CAS" collectorName="Col. K. Will" country="Madagascar" elevation="900" latitude="-21.25041" location="Fianarantsoa" longLatPrecision="1" longitude="47.41945" pageId="39" pageNumber="240" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa">
<specimenCount id="94915CC8305EFFB7FCF7FB49BE2D09E8" box="[770,846,1243,1267]" count="1" pageId="39" pageNumber="240" type="male">1 male</specimenCount>
(in
<collectionCode id="E4860F84305EFFB7FC77FB49BED509E8" box="[898,950,1243,1267]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="39" pageNumber="240" type="Museum">CAS</collectionCode>
) labeled “CASENT 8005762”/ “
<geoCoordinate id="E7A3F186305EFFB7FF0AFB69BC1E0808" box="[255,381,1275,1299]" degrees="21. 25041" direction="south" orientation="latitude" pageId="39" pageNumber="240" precision="1" value="-21.25041">21.25041°S</geoCoordinate>
/
<geoCoordinate id="E7A3F186305EFFB7FE79FB69BF680809" box="[396,523,1275,1299]" degrees="47. 41945" direction="east" orientation="longitude" pageId="39" pageNumber="240" precision="1" value="47.41945">47.41945°E</geoCoordinate>
<collectingCountry id="FA80D7D1305EFFB7FDE6FB69BFB20808" box="[531,721,1275,1299]" name="Madagascar" pageId="39" pageNumber="240">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A3305EFFB7FD15FB69BE0B0808" box="[736,872,1275,1299]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa</collectingRegion>
Prov Ranomafana National Park, Talatakely area,
<quantity id="456F3AA4305EFFB7FE9CFA89BCC50829" box="[361,422,1307,1331]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">
<elevation id="09BA7072305EFFB7FE9CFA89BCC50829" box="[361,422,1307,1331]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">900m</elevation>
</quantity>
, 10:1:2001
<collectorName id="2F62F297305EFFB7FDD3FA89BFCA0828" box="[550,681,1307,1331]" pageId="39" pageNumber="240">Col. K. Will</collectorName>
headlamp search logs”
</materialsCitation>
;
<materialsCitation id="32FF9D1C305EFFB7FC44FA89BE7808A8" collectionCode="EMEC, MNHN" collectorName="Col. K. Will &amp; All" country="Madagascar" elevation="900" latitude="-21.25041" location="Fianarantsoa" longLatPrecision="1" longitude="47.41945" pageId="39" pageNumber="240" specimenCount="6" specimenCount-female="1" specimenCount-male="5" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="94915CC8305EFFB7FC44FA89B9670828" box="[945,1028,1307,1331]" count="5" pageId="39" pageNumber="240" type="male">5 males</specimenCount>
and
<specimenCount id="94915CC8305EFFB7FBCDFA89B9F50828" box="[1080,1174,1307,1331]" count="1" pageId="39" pageNumber="240" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F84305EFFB7FF6BFAA9BD8B0848" box="[158,232,1339,1363]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="39" pageNumber="240" type="University or college">EMEC</collectionCode>
and
<collectionCode id="E4860F84305EFFB7FEEFFAA9BC080849" box="[282,363,1339,1362]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="39" pageNumber="240">MNHN</collectionCode>
) labeled “CASENT 8005762”/ “
<geoCoordinate id="E7A3F186305EFFB7FD32FAA9BE260848" box="[711,837,1339,1363]" degrees="21. 25041" direction="south" orientation="latitude" pageId="39" pageNumber="240" precision="1" value="-21.25041">21.25041°S</geoCoordinate>
/
<geoCoordinate id="E7A3F186305EFFB7FCB9FAA9BEA80849" box="[844,971,1339,1363]" degrees="47. 41945" direction="east" orientation="longitude" pageId="39" pageNumber="240" precision="1" value="47.41945">47.41945°E</geoCoordinate>
<collectingCountry id="FA80D7D1305EFFB7FC24FAA9B9EC0848" box="[977,1167,1339,1363]" name="Madagascar" pageId="39" pageNumber="240">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A3305EFFB7FF8EFAC9BC600868" box="[123,259,1371,1395]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa</collectingRegion>
Prov Ranomafana National Park, Talatakely area,
<quantity id="456F3AA4305EFFB7FCECFAC9BE350869" box="[793,854,1371,1395]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">
<elevation id="09BA7072305EFFB7FCECFAC9BE350869" box="[793,854,1371,1395]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">900m</elevation>
</quantity>
, 10:1:2001
<collectorName id="2F62F297305EFFB7FC21FAC9B9370868" box="[980,1108,1371,1395]" pageId="39" pageNumber="240">Col. K. Will</collectorName>
headlamp search logs”.
<collectorName id="2F62F297305EFFB7FE90FAE9BCE40888" box="[357,391,1403,1427]" pageId="39" pageNumber="240">All</collectorName>
<typeStatus id="5D2C29E3305EFFB7FE6CFAEEBF63088F" box="[409,512,1404,1428]" pageId="39" pageNumber="240" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="5D2C29E3305EFFB7FC7AFAE9B9770888" box="[911,1044,1403,1427]" pageId="39" pageNumber="240" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC2305EFFB7FBD0FAEEBC9708A8" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B53305EFFB7FBD0FAEEBC6A08A8" italics="true" pageId="39" pageNumber="240">Pristacrus ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628305EFFB7FE0EFA0EBF5508A8" box="[507,566,1436,1459]" pageId="39" pageNumber="240" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="82289741305EFFB7FF5EFA29BECA0B09" blockId="39.[123,1174,219,1715]" pageId="39" pageNumber="240">
<materialsCitation id="32FF9D1C305EFFB7FF5EFA29BEC10B08" country="Madagascar" location="Ranomafana National Park" pageId="39" pageNumber="240" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B53305EFFB7FF5EFA29BC3408CA" bold="true" box="[171,343,1467,1491]" pageId="39" pageNumber="240">
<smallCapsWord id="84CE019D305EFFB7FF5EFA29BD8708C9" baselines="1485,1485" box="[171,228,1467,1491]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="39" pageNumber="240">
<typeStatus id="5D2C29E3305EFFB7FF5EFA29BD8708C9" box="[171,228,1467,1491]" pageId="39" pageNumber="240">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D305EFFB7FF19FA2CBC3408CA" baselines="1484" box="[236,343,1470,1489]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="39" pageNumber="240">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D1305EFFB7FE8BFA29BC9C08C8" box="[382,511,1467,1491]" name="Madagascar" pageId="39" pageNumber="240">Madagascar</collectingCountry>
,
<collectingRegion id="405359A3305EFFB7FDF8FA29BF9F08C8" box="[525,764,1467,1491]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A305EFFB7FCFFFA29B94F08C8" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FCFFFA29B94F08C8" box="[778,1068,1467,1491]" country="Madagascar" name="Ranomafana National Park" pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Ranomafana National Park</location>
.
<emphasis id="B0E34B53305EFFB7FF5EFA49BC9108EA" bold="true" box="[171,498,1499,1523]" pageId="39" pageNumber="240">
<smallCapsWord id="84CE019D305EFFB7FF5EFA49BC4D08EA" baselines="1517,1516" box="[171,302,1499,1523]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="39" pageNumber="240">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D305EFFB7FEC3FA4CBC3108EA" baselines="1516" box="[310,338,1502,1521]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="39" pageNumber="240">OF</smallCapsWord>
<smallCapsWord id="84CE019D305EFFB7FEACFA4CBCCF08EA" baselines="1516" box="[345,428,1502,1521]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="39" pageNumber="240">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D305EFFB7FE40FA4CBC9108EA" baselines="1516" box="[437,498,1502,1521]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="39" pageNumber="240">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC2305EFFB7FD0CFA49BEE408E8" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[761,903,1499,1523]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53305EFFB7FD0CFA49BEE408E8" box="[761,903,1499,1523]" italics="true" pageId="39" pageNumber="240">ranomafanae</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the national park in which the
<typeStatus id="5D2C29E3305EFFB7FD29FA69BE690B08" box="[732,778,1531,1555]" pageId="39" pageNumber="240">type</typeStatus>
was collected
</materialsCitation>
.
</paragraph>
<paragraph id="82289741305EFFB8FF5EF989BDAF0EE2" blockId="39.[123,1174,219,1715]" lastBlockId="40.[123,1174,219,1275]" lastPageId="40" lastPageNumber="241" pageId="39" pageNumber="240">
<smallCapsWord id="84CE019D305EFFB7FF5EF989BC210B2A" baselines="1581,1580" box="[171,322,1563,1587]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="39" pageNumber="240">
<emphasis id="B0E34B53305EFFB7FF5EF989BC210B2A" bold="true" box="[171,322,1563,1587]" pageId="39" pageNumber="240">RECOGNITION</emphasis>
</smallCapsWord>
.— Size small for genus, SBL of males = 8.3–8.
<quantity id="456F3AA4305EFFB7FCABF989BEFC0B29" box="[862,927,1563,1587]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="39" pageNumber="240" unit="mm" value="5.0">5 mm</quantity>
, of female =
<quantity id="456F3AA4305EFFB7FBCDF989B9ED0B29" box="[1080,1166,1563,1587]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="39" pageNumber="240" unit="mm" value="8.6">8.6 mm</quantity>
. Members of this species (
<figureCitation id="1AAC8BC4305EFFB7FE67F9A9BC910B48" box="[402,498,1595,1619]" captionStart="FIGURE" captionStartId="38.[155,167,1385,1404]" captionTargetBox="[120,1176,216,1353]" captionTargetId="figure-677@38.[120,1176,216,1372]" captionTargetPageId="38" captionText="FIGURE 21. Digital images of holotype male of Pristacrus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels. Scale line = 1.0 mm." pageId="39" pageNumber="240">Fig. 21A</figureCitation>
) are distinguished from those of all other species in the genus except
<taxonomicName id="4597ECC2305EFFB7FF3CF9CEBF500B68" authority="Klug (1833)" authorityName="Klug" authorityYear="1833" box="[201,563,1627,1651]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="binotatus">
<emphasis id="B0E34B53305EFFB7FF3CF9CEBCC70B68" box="[201,420,1627,1651]" italics="true" pageId="39" pageNumber="240">Pristacrus binotatus</emphasis>
<bibRefCitation id="E606EAB0305EFFB7FE58F9C9BF500B68" author="KLUG, J. C. F." box="[429,563,1627,1651]" pageId="39" pageNumber="240" pagination="91 - 223" refId="ref30967" refString="KLUG, J. C. F. 1833. Bericht uber eine auf Madagascar veranstaltete. Sammlung von Insecten aus der Ordnung Coleoptera. Abhandlungen der Koniglichen Preussischen Akademie der Wissenschaften 1832 - 33: 91 - 223 + 5 pls." type="book chapter" year="1833">Klug (1833)</bibRefCitation>
</taxonomicName>
by their small body size (SBL greater than 10.0 mm in members of the three other species). They differ from members of
<taxonomicName id="4597ECC2305EFFB7FC94F9EEBD9C0BA8" authority="(Fairmaire 1887)" baseAuthorityName="Fairmaire" baseAuthorityYear="1887" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="semipiceus">
<emphasis id="B0E34B53305EFFB7FC94F9EEB92C0B88" box="[865,1103,1660,1683]" italics="true" pageId="39" pageNumber="240">Pristacrus semipiceus</emphasis>
(
<bibRefCitation id="E606EAB0305EFFB7FB97F9E9BD960BA8" author="FAIRMAIRE, L." pageId="39" pageNumber="240" pagination="56 - 57" refId="ref30177" refString="FAIRMAIRE, L. 1887. Diagnoses du Coleopteres nouveaux de Madagascar. Le Naturaliste (2 e Serie) 1: 56 - 57" type="journal article" year="1887">Fairmaire 1887</bibRefCitation>
)
</taxonomicName>
, in having much smaller body size (SBL = 13.6 to 14.
<quantity id="456F3AA4305EFFB7FCA3F909BEF50BA8" box="[854,918,1691,1715]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="39" pageNumber="240" unit="mm" value="5.0">5 mm</quantity>
in
<taxonomicName id="4597ECC2305EFFB7FC48F90EB92D0BA8" baseAuthorityName="Fairmaire" baseAuthorityYear="1887" box="[957,1102,1692,1715]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="semipiceus">
<emphasis id="B0E34B53305EFFB7FC48F90EB92D0BA8" box="[957,1102,1692,1715]" italics="true" pageId="39" pageNumber="240">P. semipiceus</emphasis>
</taxonomicName>
members), a shinier dorsal surface, shorter and broadly rounded mandibles and an impunctate (but longitudinally rugulose) frons (dorsal surface duller, mandibles distinctly longer and more tapered, and frons punctate as well as rugulose in
<taxonomicName id="4597ECC23051FFB8FDB7FE8EBFB00C28" baseAuthorityName="Fairmaire" baseAuthorityYear="1887" box="[578,723,284,307]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="semipiceus">
<emphasis id="B0E34B533051FFB8FDB7FE8EBFB00C28" box="[578,723,284,307]" italics="true" pageId="40" pageNumber="241">P. semipiceus</emphasis>
</taxonomicName>
members). They differ from members of
<taxonomicName id="4597ECC23051FFB8FF8EFEAFBD910C4F" baseAuthorityName="Gory and Laporte" baseAuthorityYear="1837" box="[123,242,316,340]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="laticollis">
<emphasis id="B0E34B533051FFB8FF8EFEAFBD910C4F" box="[123,242,316,340]" italics="true" pageId="40" pageNumber="241">P. laticollis</emphasis>
</taxonomicName>
in having slightly smaller body size (SBL = 10.2–11.
<quantity id="456F3AA43051FFB8FCD9FEAEBE0A0C48" box="[812,873,316,340]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="40" pageNumber="241" unit="mm" value="1.0">1 mm</quantity>
), slightly shinier dorsal surface, relatively longer and narrower pronotum and elytra that are broadly ovoid with lateral margin arcuate and lateral explanation broad throughout, especially at middle, and elytral apices distinctly angulate and toothed (duller dorsal surface, relatively broader and shorter pronotum and elytra that are nearly parallel-sided at middle with lateral explanation narrower throughout and elytral apices broadly rounded in
<taxonomicName id="4597ECC23051FFB8FE68FE4CBF750CEE" baseAuthorityName="Gory and Laporte" baseAuthorityYear="1837" box="[413,534,477,501]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="laticollis">
<emphasis id="B0E34B533051FFB8FE68FE4CBF750CEE" box="[413,534,477,501]" italics="true" pageId="40" pageNumber="241">P. laticollis</emphasis>
</taxonomicName>
members). They differ from the unique
<typeStatus id="5D2C29E33051FFB8FC24FE4CBE9C0CED" box="[977,1023,478,502]" pageId="40" pageNumber="241">type</typeStatus>
of
<taxonomicName id="4597ECC23051FFB8FBD3FE4CBCCF0F0D" authority="(Fairmaire 1892)" baseAuthorityName="Fairmaire" baseAuthorityYear="1892" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="B0E34B533051FFB8FBD3FE4CBD8D0F0E" italics="true" pageId="40" pageNumber="241">Pristacrus rotundatus</emphasis>
(
<bibRefCitation id="E606EAB03051FFB8FF0BFE6FBCC00F0D" author="FAIRMAIRE, L." box="[254,419,509,534]" pageId="40" pageNumber="241" pagination="clxviii - clxxii" refId="ref30226" refString="FAIRMAIRE, L. 1892. Descriptions de quelques Coleopteres de Madagascar, provenant de la region de Diego- Suarez, encore peu exploree, et qui lui ont ete communiques par notre collegue M. Rene Oberthur. Annales de la Societe Entomologique de France 61 (Bulletin Entomologique Seance du 22 Juin 1892): clxviii - clxxii." type="journal article" year="1892">Fairmaire 1892</bibRefCitation>
)
</taxonomicName>
in having smaller body size (SBL =
<quantity id="456F3AA43051FFB8FCC8FE6CBEFF0F0E" box="[829,924,510,534]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="40" pageNumber="241" unit="mm" value="11.9">11.9 mm</quantity>
in
<taxonomicName id="4597ECC23051FFB8FC37FE6CB9330F0E" baseAuthorityName="Fairmaire" baseAuthorityYear="1892" box="[962,1104,509,533]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="B0E34B533051FFB8FC37FE6CB9330F0E" box="[962,1104,509,533]" italics="true" pageId="40" pageNumber="241">P. rotundatus</emphasis>
</taxonomicName>
<typeStatus id="5D2C29E33051FFB8FBADFE6CB9E50F0D" box="[1112,1158,510,534]" pageId="40" pageNumber="241">type</typeStatus>
), shinier dorsal surface, elytra less broadly rounded and humeri less anteriorly projected (dorsal surface dull and elytra markedly broad and rounded and humeri more projected anteriorly in
<taxonomicName id="4597ECC23051FFB8FBCCFDADBDD60F6D" baseAuthorityName="Fairmaire" baseAuthorityYear="1892" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="B0E34B533051FFB8FBCCFDADBDD60F6D" italics="true" pageId="40" pageNumber="241">P. rotundatus</emphasis>
</taxonomicName>
<typeStatus id="5D2C29E33051FFB8FF4EFDCDBD8A0F6C" box="[187,233,607,631]" pageId="40" pageNumber="241">type</typeStatus>
). Members of
<taxonomicName id="4597ECC23051FFB8FE7CFDCDBF520F6D" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[393,561,606,630]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533051FFB8FE7CFDCDBF520F6D" box="[393,561,606,630]" italics="true" pageId="40" pageNumber="241">P. ranomafanae</emphasis>
</taxonomicName>
are very similar to those of
<taxonomicName id="4597ECC23051FFB8FCAAFDCDBEBF0F6D" authorityName="Klug" authorityYear="1833" box="[863,988,606,630]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="binotatus">
<emphasis id="B0E34B533051FFB8FCAAFDCDBEBF0F6D" box="[863,988,606,630]" italics="true" pageId="40" pageNumber="241">P. binotatus</emphasis>
</taxonomicName>
in size and overall form. Specimens of the former have the dorsal surface slightly shinier, the pronotum relatively broader basally, with subbasal sinuation of the lateral margins absent or only very faint and hind angles less sharply defined, and elytra with striae slightly more deeply impressed and intervals more convex and more roughened at the raised center than in most members of
<taxonomicName id="4597ECC23051FFB8FC3BFD72B9280FEC" authorityName="Klug" authorityYear="1833" box="[974,1099,735,759]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="binotatus">
<emphasis id="B0E34B533051FFB8FC3BFD72B9280FEC" box="[974,1099,735,759]" italics="true" pageId="40" pageNumber="241">P. binotatus</emphasis>
</taxonomicName>
. However, at least a few specimens of
<taxonomicName id="4597ECC23051FFB8FE15FC92BF3D0E0C" authorityName="Klug" authorityYear="1833" box="[480,606,767,791]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="binotatus">
<emphasis id="B0E34B533051FFB8FE15FC92BF3D0E0C" box="[480,606,767,791]" italics="true" pageId="40" pageNumber="241">P. binotatus</emphasis>
</taxonomicName>
share one or more of these features with
<taxonomicName id="4597ECC23051FFB8FBEBFC92BDD50E23" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="ranomofanae">
<emphasis id="B0E34B533051FFB8FBEBFC92BDD50E23" italics="true" pageId="40" pageNumber="241">P. ranomofanae</emphasis>
</taxonomicName>
members. They only features that most reliably distinguish members of
<taxonomicName id="4597ECC23051FFB8FC41FCB2B9380E23" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[948,1115,800,824]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533051FFB8FC41FCB2B9380E23" box="[948,1115,800,824]" italics="true" pageId="40" pageNumber="241">P. ranomafanae</emphasis>
</taxonomicName>
from those of
<taxonomicName id="4597ECC23051FFB8FF2BFCD3BC3C0E43" authorityName="Klug" authorityYear="1833" box="[222,351,832,856]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="binotatus">
<emphasis id="B0E34B533051FFB8FF2BFCD3BC3C0E43" box="[222,351,832,856]" italics="true" pageId="40" pageNumber="241">P. binotatus</emphasis>
</taxonomicName>
, as well as from members of the other species, are the male genitalia. In
<taxonomicName id="4597ECC23051FFB8FF8EFCF3BC470E63" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,292,864,888]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533051FFB8FF8EFCF3BC470E63" box="[123,292,864,888]" italics="true" pageId="40" pageNumber="241">P. ranomafanae</emphasis>
</taxonomicName>
males (Fig. 22A–B), the shaft of the median lobe is inflated subbasally but lacks the large, well-defined dorsal bulge seen in males of
<taxonomicName id="4597ECC23051FFB8FD45FC13BE4E0E83" authorityName="Klug" authorityYear="1833" box="[688,813,896,920]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="binotatus">
<emphasis id="B0E34B533051FFB8FD45FC13BE4E0E83" box="[688,813,896,920]" italics="true" pageId="40" pageNumber="241">P. binotatus</emphasis>
</taxonomicName>
(see Jeannel 1949, Fig. 486) and, in lateral aspect (
<figureCitation id="1AAC8BC43051FFB8FEC7FC33BCF20EA2" box="[306,401,929,953]" captionStart="FIGURE" captionStartId="38.[155,167,1385,1404]" captionTargetBox="[120,1176,216,1353]" captionTargetId="figure-677@38.[120,1176,216,1372]" captionTargetPageId="38" captionText="FIGURE 21. Digital images of holotype male of Pristacrus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels. Scale line = 1.0 mm." pageId="40" pageNumber="241">Fig. 21A</figureCitation>
), the apex is distinctly hooked dorsally (apex not hooked in
<taxonomicName id="4597ECC23051FFB8FBEDFC30B9F60EA2" authorityName="Klug" authorityYear="1833" box="[1048,1173,929,953]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="binotatus">
<emphasis id="B0E34B533051FFB8FBEDFC30B9F60EA2" box="[1048,1173,929,953]" italics="true" pageId="40" pageNumber="241">P. binotatus</emphasis>
</taxonomicName>
males. Males of the each of the other species also have the shaft of the median lobe uniquely shaped.
</paragraph>
<paragraph id="822897413051FFB8FF5EFB93B9690902" blockId="40.[123,1174,219,1275]" box="[171,1034,1025,1050]" pageId="40" pageNumber="241">
<emphasis id="B0E34B533051FFB8FF5EFB93BC920903" bold="true" box="[171,497,1025,1049]" pageId="40" pageNumber="241">
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<smallCapsWord id="84CE019D3051FFB8FEA8FB97BC920903" baselines="1043" box="[349,497,1029,1048]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="40" pageNumber="241">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E33051FFB8FC8AFB90BECE0901" box="[895,941,1026,1050]" pageId="40" pageNumber="241">type</typeStatus>
locality.
</paragraph>
<paragraph id="822897413051FFB8FF5EFBB0BCA909E0" blockId="40.[123,1174,219,1275]" pageId="40" pageNumber="241">
<emphasis id="B0E34B533051FFB8FF5EFBB0BCC70923" bold="true" box="[171,420,1058,1082]" pageId="40" pageNumber="241">
<smallCapsWord id="84CE019D3051FFB8FF5EFBB0BC6A0923" baselines="1076,1075" box="[171,265,1058,1082]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="40" pageNumber="241">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D3051FFB8FEE5FBB7BCC70923" baselines="1075" box="[272,420,1061,1080]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="40" pageNumber="241">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Specimens of the
<typeStatus id="5D2C29E33051FFB8FD78FBB0BFD80921" box="[653,699,1058,1082]" pageId="40" pageNumber="241">type</typeStatus>
series were all collected in secondary montane rainforest in the Talatakely area at elevations of
<quantity id="456F3AA43051FFB8FD47FBD0BE2C0941" box="[690,847,1090,1114]" metricMagnitude="2" metricUnit="m" metricValue="9.6" metricValueMax="10.2" metricValueMin="9.0" pageId="40" pageNumber="241" unit="m" value="960.0" valueMax="1020.0" valueMin="900.0">900 to 1020 m</quantity>
. The area is replete with invasive guava (
<taxonomicName id="4597ECC23051FFB8FEF7FBF0BC820961" box="[258,481,1122,1146]" class="Magnoliopsida" family="Myrtaceae" genus="Psidium" kingdom="Plantae" order="Myrtales" pageId="40" pageNumber="241" phylum="Tracheophyta" rank="species" species="cattleianum">
<emphasis id="B0E34B533051FFB8FEF7FBF0BC820961" box="[258,481,1122,1146]" italics="true" pageId="40" pageNumber="241">Psidium cattleianum</emphasis>
</taxonomicName>
). Specimens were found in daytime in masses of dead leaves, twigs and branch debris accumulated on the forest floor, in leaf litter, and under loose bark or on the underside of rotting logs on the forest floor. Several were found at night running on the surfaces of rotting logs on the forest floor; two were collected in malaise traps and one was attracted to and collected at an ultraviolet light.
</paragraph>
</subSubSection>
</treatment>
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409
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<mods:title id="C68F7203AE1BA04F114FFF3141AD437D">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:namePart id="484A1B3B9C9742ADA782A0BB6E2B94C7">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="A92C63799D7D0C6B9923599BA0E36F95">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:namePart id="EE37B3D66DE9F9F5785A789F6D95B346">Rainio, Johanna</mods:namePart>
<mods:affiliation id="CF9103EC1707D48388CEA6A4B96CFB9F">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
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<heading id="D960202D3060FF89FF8EFF49BE460DE8" box="[123,805,219,243]" fontSize="10" level="2" pageId="25" pageNumber="226" reason="4">
<taxonomicName id="4597ECC23060FF89FF8EFF49BFA20DE8" ID-CoL="775R9" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,705,219,243]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="deuvei" status="sp. nov." subGenus="Ripogena">
<emphasis id="B0E34B533060FF89FF8EFF49BC3A0DE8" bold="true" box="[123,345,219,243]" pageId="25" pageNumber="226">
<emphasis id="B0E34B533060FF89FF8EFF49BD830DE8" bold="true" box="[123,224,219,243]" italics="true" pageId="25" pageNumber="226">Perigona</emphasis>
(
<emphasis id="B0E34B533060FF89FF05FF49BC3A0DE8" bold="true" box="[240,345,219,243]" italics="true" pageId="25" pageNumber="226">Ripogena</emphasis>
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)
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<emphasis id="B0E34B533060FF89FE9CFF49BCD30DE8" bold="true" box="[361,432,219,243]" italics="true" pageId="25" pageNumber="226">deuvei</emphasis>
Kavanaugh and Rainio
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,
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<figureCitation id="1AAC8BC43060FF89FF8EFF69BDBB0C0B" box="[123,216,251,273]" captionStart="FIGURE" captionStartId="26.[155,167,1478,1497]" captionTargetBox="[139,1176,216,1455]" captionTargetId="figure-53@26.[120,1176,216,1455]" captionTargetPageId="26" captionText="FIGURE 14. Digital images of holotype male of Perigona (Ripogena) deuvei Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm." pageId="25" pageNumber="226">Figure 14</figureCitation>
</paragraph>
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<emphasis id="B0E34B533060FF89FF8EFEB5BC470C20" box="[123,292,294,316]" italics="true" pageId="25" pageNumber="226">
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<taxonomicNameLabel id="ABD0F6283060FF89FF28FEB5BC720C27" box="[221,273,295,316]" pageId="25" pageNumber="226" rank="species">n. sp.</taxonomicNameLabel>
1
</emphasis>
; Rainio 2009: 33 (informal designation).
</paragraph>
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<taxonomicName id="4597ECC23060FF89FF8EFED5BC240C40" authorityName="Jeannel" authorityYear="1941" box="[123,327,326,348]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="subGenus" subGenus="Ripogena">
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(
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)
</taxonomicName>
<emphasis id="B0E34B533060FF89FEB8FED5BCF70C40" box="[333,404,326,348]" italics="true" pageId="25" pageNumber="226">n. sp. 1</emphasis>
; Rainio 2012: 73, Rainio 2013: 96 (informal designation).
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</emphasis>
.—
<materialsCitation id="32FF9D1C3060FF89FE73FEEDBCA30CEC" collectingDate="2005-07-24" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="350" location="Vatoharanana" pageId="25" pageNumber="226" specimenCount="2" typeStatus="holotype">
<typeStatus id="5D2C29E33060FF89FE73FEEDBC8A0C8C" box="[390,489,383,407]" pageId="25" pageNumber="226" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43060FF89FE0BFEEDBFE30C8C" box="[510,640,383,408]" captionStart="FIGURE" captionStartId="26.[155,167,1478,1497]" captionTargetBox="[139,1176,216,1455]" captionTargetId="figure-53@26.[120,1176,216,1455]" captionTargetPageId="26" captionText="FIGURE 14. Digital images of holotype male of Perigona (Ripogena) deuvei Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm." pageId="25" pageNumber="226">Figs.14A–B</figureCitation>
), a male, in CAS, labeled: “
<collectingCountry id="FA80D7D13060FF89FC24FEEDB9EC0C8C" box="[977,1167,383,407]" name="Madagascar" pageId="25" pageNumber="226">MADAGASCAR</collectingCountry>
, Ranomafana NP.,
<location id="8748C19A3060FF89FECBFE0DBCAC0CAC" LSID="urn:lsid:plazi:treatment:0A3E26573060FF8BFF8EFF49BEB20CA8:8748C19A3060FF89FECBFE0DBCAC0CAC" box="[318,463,415,439]" country="Madagascar" name="Vatoharanana" pageId="25" pageNumber="226">Vatoharanana</location>
, Trail K at
<quantity id="456F3AA43060FF89FDBAFE0DBFF60CAC" box="[591,661,415,439]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="25" pageNumber="226" unit="m" value="350.0">
<elevation id="09BA70723060FF89FDBAFE0DBFF60CAC" box="[591,661,415,439]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="25" pageNumber="226" unit="m" value="350.0">350 m</elevation>
</quantity>
mark,
<date id="F629B1813060FF89FD17FE32BE390CAC" box="[738,858,415,440]" pageId="25" pageNumber="226" value="2005-07-24">
<collectingDate id="E66D48693060FF89FD17FE32BE390CAC" box="[738,858,415,440]" pageId="25" pageNumber="226" value="2005-07-24">24.07.2005</collectingDate>
</date>
”/ “collected by hand in daytime,
<collectorName id="2F62F2973060FF89FF42FE52BC4B0CC3" box="[183,296,448,472]" pageId="25" pageNumber="226">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973060FF89FEBCFE52BCC40CC3" box="[329,423,448,472]" pageId="25" pageNumber="226">J. Rainio</collectorName>
collectors”/ “
<typeStatus id="5D2C29E33060FF89FDCEFE52BFAB0CC3" box="[571,712,448,472]" pageId="25" pageNumber="226" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23060FF89FD3BFE52BDA10CE3" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="deuvei" status="sp. nov." subGenus="Ripogena">
<emphasis id="B0E34B533060FF89FD3BFE52BE520CCC" box="[718,817,448,471]" italics="true" pageId="25" pageNumber="226">Perigona</emphasis>
(
<emphasis id="B0E34B533060FF89FCB5FE52BEC50CCC" box="[832,934,448,471]" italics="true" pageId="25" pageNumber="226">Ripogena</emphasis>
)
<emphasis id="B0E34B533060FF89FC40FE52BE980CC3" box="[949,1019,448,472]" italics="true" pageId="25" pageNumber="226">deuvei</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283060FF89FF3CFE72BC610CEC" box="[201,258,480,503]" pageId="25" pageNumber="226" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label]
</materialsCitation>
.
<materialsCitation id="32FF9D1C3060FF89FE38FE72BE010F23" collectingDate="2005-10-06" collectionCode="MNHN" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="50" location="Vatoharanana" municipality="Trail" pageId="25" pageNumber="226" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="5D2C29E33060FF89FE38FE72BF560CE3" box="[461,565,480,504]" pageId="25" pageNumber="226" type="paratype">Paratypes</typeStatus>
(total of 4):
<specimenCount id="94915CC83060FF89FD49FE72BE790CE3" box="[700,794,480,504]" count="1" pageId="25" pageNumber="226" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843060FF89FCB0FE72BEF50CEC" box="[837,918,480,503]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="226">MNHN</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13060FF89FBF3FE72BDD10F03" name="Madagascar" pageId="25" pageNumber="226">MADAGASCAR</collectingCountry>
, Ranomafana NP.,
<location id="8748C19A3060FF89FE88FD92BF6D0F03" LSID="urn:lsid:plazi:treatment:0A3E26573060FF8BFF8EFF49BEB20CA8:8748C19A3060FF89FE88FD92BF6D0F03" box="[381,526,512,536]" country="Madagascar" municipality="Trail" name="Vatoharanana" pageId="25" pageNumber="226">Vatoharanana</location>
,
<collectingMunicipality id="624C0D3B3060FF89FDEFFD92BF2E0F03" box="[538,589,512,536]" pageId="25" pageNumber="226">Trail</collectingMunicipality>
J at
<quantity id="456F3AA43060FF89FD89FD92BFD10F0C" box="[636,690,512,536]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="226" unit="m" value="50.0">
<elevation id="09BA70723060FF89FD89FD92BFD10F0C" box="[636,690,512,536]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="226" unit="m" value="50.0">50 m</elevation>
</quantity>
mark,
<date id="F629B1813060FF89FD0EFD92BE060F03" box="[763,869,512,536]" pageId="25" pageNumber="226" value="2005-10-06">
<collectingDate id="E66D48693060FF89FD0EFD92BE060F03" box="[763,869,512,536]" pageId="25" pageNumber="226" value="2005-10-06">6.10.2005</collectingDate>
</date>
”/ “collected by hand in daytime,
<collectorName id="2F62F2973060FF89FF4FFDB3BC4D0F23" box="[186,302,544,568]" pageId="25" pageNumber="226">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973060FF89FEA0FDB3BCD40F23" box="[341,439,544,568]" pageId="25" pageNumber="226">J. Rainio</collectorName>
collectors”/ “239.” [handwritten label]
</materialsCitation>
;
<materialsCitation id="32FF9D1C3060FF89FC86FDB2B9720F63" collectingDate="2005-09-09" collectionCode="CAS" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="600" location="Vatoharanana" municipality="Trail" pageId="25" pageNumber="226" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="94915CC83060FF89FC86FDB2BEB00F23" box="[883,979,544,568]" count="1" pageId="25" pageNumber="226" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843060FF89FBF7FDB2B9550F23" box="[1026,1078,544,568]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="25" pageNumber="226" type="Museum">CAS</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13060FF89FF72FDD2BC260F43" box="[135,325,576,600]" name="Madagascar" pageId="25" pageNumber="226">MADAGASCAR</collectingCountry>
, Ranomafana NP.,
<location id="8748C19A3060FF89FDE0FDD2BFC50F43" LSID="urn:lsid:plazi:treatment:0A3E26573060FF8BFF8EFF49BEB20CA8:8748C19A3060FF89FDE0FDD2BFC50F43" box="[533,678,576,600]" country="Madagascar" municipality="Trail" name="Vatoharanana" pageId="25" pageNumber="226">Vatoharanana</location>
,
<collectingMunicipality id="624C0D3B3060FF89FD41FDD2BF840F43" box="[692,743,576,600]" pageId="25" pageNumber="226">Trail</collectingMunicipality>
S at
<quantity id="456F3AA43060FF89FCD5FDD2BE070F43" box="[800,868,576,600]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="226" unit="m" value="600.0">
<elevation id="09BA70723060FF89FCD5FDD2BE070F43" box="[800,868,576,600]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="226" unit="m" value="600.0">600 m</elevation>
</quantity>
mark,
<date id="F629B1813060FF89FC44FDD2B96D0F43" box="[945,1038,576,600]" pageId="25" pageNumber="226" value="2005-09-09">
<collectingDate id="E66D48693060FF89FC44FDD2B96D0F43" box="[945,1038,576,600]" pageId="25" pageNumber="226" value="2005-09-09">9.9.2005</collectingDate>
</date>
”/ “collected by hand in daytime,
<collectorName id="2F62F2973060FF89FE96FDF3BCB40F63" box="[355,471,608,632]" pageId="25" pageNumber="226">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973060FF89FDF5FDF3BF010F63" box="[512,610,608,632]" pageId="25" pageNumber="226">J. Rainio</collectorName>
collectors”/ “239.” [handwritten label]
</materialsCitation>
;
<materialsCitation id="32FF9D1C3060FF89FBD7FDF2BD830FC3" collectingDate="2005-07-24" collectionCode="MZF" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="600" location="Vatoharanana" municipality="Trail" pageId="25" pageNumber="226" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="94915CC83060FF89FBD7FDF2B90D0F63" box="[1058,1134,608,632]" count="1" pageId="25" pageNumber="226" type="male">1 male</specimenCount>
(in
<collectionCode id="E4860F843060FF89FF8EFD13BDD10F83" box="[123,178,641,664]" pageId="25" pageNumber="226">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13060FF89FEC4FD13BC8C0F82" box="[305,495,641,665]" name="Madagascar" pageId="25" pageNumber="226">MADAGASCAR</collectingCountry>
, Ranomafana NP.,
<location id="8748C19A3060FF89FD20FD12BE050F83" LSID="urn:lsid:plazi:treatment:0A3E26573060FF8BFF8EFF49BEB20CA8:8748C19A3060FF89FD20FD12BE050F83" box="[725,870,640,664]" country="Madagascar" municipality="Trail" name="Vatoharanana" pageId="25" pageNumber="226">Vatoharanana</location>
,
<collectingMunicipality id="624C0D3B3060FF89FC8EFD12BECD0F83" box="[891,942,640,664]" pageId="25" pageNumber="226">Trail</collectingMunicipality>
S at
<quantity id="456F3AA43060FF89FC08FD12B92A0F83" box="[1021,1097,640,664]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="226" unit="m" value="600.0">
<elevation id="09BA70723060FF89FC08FD12B92A0F83" box="[1021,1097,640,664]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="226" unit="m" value="600.0">600 m</elevation>
</quantity>
mark,
<date id="F629B1813060FF89FF8EFD33BD860FA3" box="[123,229,672,697]" pageId="25" pageNumber="226" value="2005-07-24">
<collectingDate id="E66D48693060FF89FF8EFD33BD860FA3" box="[123,229,672,697]" pageId="25" pageNumber="226" value="2005-07-24">24.7.2005</collectingDate>
</date>
”/ “collected by hand in daytime,
<collectorName id="2F62F2973060FF89FDA1FD33BFA40FA2" box="[596,711,673,697]" pageId="25" pageNumber="226">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973060FF89FD19FD33BE2F0FA2" box="[748,844,673,697]" pageId="25" pageNumber="226">J. Rainio</collectorName>
collectors”/ “253.” [handwritten label]
</materialsCitation>
;
<materialsCitation id="32FF9D1C3060FF89FF1BFD53BE320E22" collectingDate="2005-10-06" collectionCode="MZF" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="600" location="Vatoharanana" municipality="Trail" pageId="25" pageNumber="226" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="94915CC83060FF89FF1BFD53BC2F0FC2" box="[238,332,705,729]" count="1" pageId="25" pageNumber="226" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843060FF89FE8DFD53BCCC0FC3" box="[376,431,705,728]" pageId="25" pageNumber="226">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13060FF89FDD5FD53BFBD0FC2" box="[544,734,705,729]" name="Madagascar" pageId="25" pageNumber="226">MADAGASCAR</collectingCountry>
, Ranomafana NP.,
<location id="8748C19A3060FF89FC58FD53B95D0FC2" LSID="urn:lsid:plazi:treatment:0A3E26573060FF8BFF8EFF49BEB20CA8:8748C19A3060FF89FC58FD53B95D0FC2" box="[941,1086,705,729]" country="Madagascar" municipality="Trail" name="Vatoharanana" pageId="25" pageNumber="226">Vatoharanana</location>
,
<collectingMunicipality id="624C0D3B3060FF89FBB9FD53B91C0FC2" box="[1100,1151,705,729]" pageId="25" pageNumber="226">Trail</collectingMunicipality>
S at
<quantity id="456F3AA43060FF89FF6DFD73BDBD0FE3" box="[152,222,737,761]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="226" unit="m" value="600.0">
<elevation id="09BA70723060FF89FF6DFD73BDBD0FE3" box="[152,222,737,761]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="226" unit="m" value="600.0">600 m</elevation>
</quantity>
mark,
<date id="F629B1813060FF89FEDAFD73BCFA0FE2" box="[303,409,737,761]" pageId="25" pageNumber="226" value="2005-10-06">
<collectingDate id="E66D48693060FF89FEDAFD73BCFA0FE2" box="[303,409,737,761]" pageId="25" pageNumber="226" value="2005-10-06">6.10.2005</collectingDate>
</date>
”/ “collected by hand in daytime,
<collectorName id="2F62F2973060FF89FCE6FD73BEE40FE2" box="[787,903,737,761]" pageId="25" pageNumber="226">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973060FF89FC5AFD73B9720FE2" box="[943,1041,737,761]" pageId="25" pageNumber="226">J. Rainio</collectorName>
collectors”/ “245.” [handwritten label]. All
<typeStatus id="5D2C29E33060FF89FE22FC90BF5E0E01" box="[471,573,770,794]" pageId="25" pageNumber="226" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="5D2C29E33060FF89FC56FC93B9440E02" box="[931,1063,769,793]" pageId="25" pageNumber="226" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC23060FF89FBC7FC90BF480E22" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="deuvei" status="sp. nov." subGenus="Ripogena">
<emphasis id="B0E34B533060FF89FBC7FC90B9F60E02" box="[1074,1173,770,793]" italics="true" pageId="25" pageNumber="226">Perigona</emphasis>
(
<emphasis id="B0E34B533060FF89FF71FCB0BD890E22" box="[132,234,802,825]" italics="true" pageId="25" pageNumber="226">Ripogena</emphasis>
)
<emphasis id="B0E34B533060FF89FF0FFCB3BC230E22" box="[250,320,801,825]" italics="true" pageId="25" pageNumber="226">deuvei</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283060FF89FDC7FCB0BF0F0E22" box="[562,620,802,825]" pageId="25" pageNumber="226" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="822897413060FF89FF5EFCD0B9500E42" blockId="25.[123,1174,383,1726]" box="[171,1075,833,858]" pageId="25" pageNumber="226">
<materialsCitation id="32FF9D1C3060FF89FF5EFCD0B94F0E42" box="[171,1068,833,858]" country="Madagascar" location="Ranomafana National Park" pageId="25" pageNumber="226" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533060FF89FF5EFCD0BC340E43" bold="true" box="[171,343,834,858]" pageId="25" pageNumber="226">
<smallCapsWord id="84CE019D3060FF89FF5EFCD0BD870E43" baselines="852,851" box="[171,228,834,858]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="25" pageNumber="226">
<typeStatus id="5D2C29E33060FF89FF5EFCD0BD870E43" box="[171,228,834,858]" pageId="25" pageNumber="226">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3060FF89FF19FCD7BC340E43" baselines="851" box="[236,343,837,856]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="25" pageNumber="226">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13060FF89FE8BFCD3BC9C0E42" box="[382,511,833,857]" name="Madagascar" pageId="25" pageNumber="226">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33060FF89FDF8FCD3BF9F0E42" box="[525,764,833,857]" country="Madagascar" name="Fianarantsoa" pageId="25" pageNumber="226">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3060FF89FCFFFCD3B94F0E42" LSID="urn:lsid:plazi:treatment:0A3E26573060FF8BFF8EFF49BEB20CA8:8748C19A3060FF89FCFFFCD3B94F0E42" box="[778,1068,833,857]" country="Madagascar" name="Ranomafana National Park" pageId="25" pageNumber="226" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413060FF89FF5EFCF0BD8B0EE1" blockId="25.[123,1174,383,1726]" pageId="25" pageNumber="226">
<emphasis id="B0E34B533060FF89FF5EFCF0BC8D0E63" bold="true" box="[171,494,866,890]" pageId="25" pageNumber="226">
<smallCapsWord id="84CE019D3060FF89FF5EFCF0BC4D0E63" baselines="884,883" box="[171,302,866,890]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="25" pageNumber="226">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3060FF89FEC0FCF7BC320E63" baselines="883" box="[309,337,869,888]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="25" pageNumber="226">OF</smallCapsWord>
<smallCapsWord id="84CE019D3060FF89FEA2FCF7BCC90E63" baselines="883" box="[343,426,869,888]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="25" pageNumber="226">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3060FF89FE44FCF7BC8D0E63" baselines="883" box="[433,494,869,888]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="25" pageNumber="226">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23060FF89FD04FCF0BE540E61" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[753,823,866,890]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533060FF89FD04FCF0BE540E61" box="[753,823,866,890]" italics="true" pageId="25" pageNumber="226">deuvei</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the surname of Dr. Thierry Deuve of the Laboratoire d’Entomologie at the Muséum National d’Histoire Naturelle in Paris. We are pleased to name this new species in honor of Dr. Deuve in thanks for his friendship and kind hospitality during our several visits to MNHN in the course of this study.
</paragraph>
<paragraph id="822897413060FF8BFF5EFB90BE880C48" blockId="25.[123,1174,383,1726]" lastBlockId="27.[123,1174,219,436]" lastPageId="27" lastPageNumber="228" pageId="25" pageNumber="226">
<smallCapsWord id="84CE019D3060FF89FF5EFB90BC210902" baselines="1044,1044" box="[171,322,1026,1050]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="25" pageNumber="226">
<emphasis id="B0E34B533060FF89FF5EFB90BC210902" bold="true" box="[171,322,1026,1050]" pageId="25" pageNumber="226">RECOGNITION</emphasis>
</smallCapsWord>
.— Size average for subgenus, SBL of males = 3.5–
<quantity id="456F3AA43060FF89FC9BFB90BEA30901" box="[878,960,1026,1050]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="25" pageNumber="226" unit="mm" value="3.8">3.8 mm</quantity>
, of females 3.8–
<quantity id="456F3AA43060FF89FB81FB90BDC70921" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="25" pageNumber="226" unit="mm" value="3.9">3.9 mm</quantity>
. Members of this species (
<figureCitation id="1AAC8BC43060FF89FE32FBB0BF440920" box="[455,551,1058,1083]" captionStart="FIGURE" captionStartId="26.[155,167,1478,1497]" captionTargetBox="[139,1176,216,1455]" captionTargetId="figure-53@26.[120,1176,216,1455]" captionTargetPageId="26" captionText="FIGURE 14. Digital images of holotype male of Perigona (Ripogena) deuvei Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm." pageId="25" pageNumber="226">Fig. 14A</figureCitation>
) share with those of
<taxonomicName id="4597ECC23060FF89FCFEFBB1B9F60920" authority="Alluaud (1936)" authorityName="Alluaud" authorityYear="1936" box="[779,1173,1058,1083]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533060FF89FCFEFBB1BE890921" box="[779,1002,1058,1082]" italics="true" pageId="25" pageNumber="226">Perigona heterodera</emphasis>
Alluaud (1936)
</taxonomicName>
and
<taxonomicName id="4597ECC23060FF89FF5DFBD1BF750940" authority="Alluaud (1936)" authorityName="Alluaud" authorityYear="1936" box="[168,534,1091,1115]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="B0E34B533060FF89FF5DFBD1BC0E0941" box="[168,365,1091,1114]" italics="true" pageId="25" pageNumber="226">Perigona prasinus</emphasis>
Alluaud (1936)
</taxonomicName>
the following features: pronotum with lateral margins without sinuation anterior to hind angles, posterior margin slightly and smoothly convex, and hind angles either obtuse or broadly rounded. These features distinguish members of these three species from those of
<taxonomicName id="4597ECC23060FF89FEE2FB36BFDD09A0" authority="Alluaud (1936)" authorityName="Alluaud" authorityYear="1936" box="[279,702,1187,1211]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="bembidioides">
<emphasis id="B0E34B533060FF89FEE2FB36BF7209A0" box="[279,529,1187,1211]" italics="true" pageId="25" pageNumber="226">Perigona bembidioides</emphasis>
Alluaud (1936)
</taxonomicName>
and
<taxonomicName id="4597ECC23060FF89FD00FB36B9EC09A0" authority="Jeannel (1948)" authorityName="Jeannel" authorityYear="1948" box="[757,1167,1187,1211]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="viridimicans">
<emphasis id="B0E34B533060FF89FD00FB36BE8509A0" box="[757,998,1187,1211]" italics="true" pageId="25" pageNumber="226">Perigona viridimicans</emphasis>
Jeannel (1948)
</taxonomicName>
, which have distinct sinuation of the lateral margins anterior to rectangular hind angles and a straight basal margin. Members of
<taxonomicName id="4597ECC23060FF89FE19FB76BF2F09E0" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[492,588,1251,1275]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533060FF89FE19FB76BF2F09E0" box="[492,588,1251,1275]" italics="true" pageId="25" pageNumber="226">P. deuvei</emphasis>
</taxonomicName>
differ from those of
<taxonomicName id="4597ECC23060FF89FCD2FB76BEFF09E0" authorityName="Alluaud" authorityYear="1936" box="[807,924,1252,1275]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="B0E34B533060FF89FCD2FB76BEFF09E0" box="[807,924,1252,1275]" italics="true" pageId="25" pageNumber="226">P. prasinus</emphasis>
</taxonomicName>
in having smaller body size (SBL =
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in
<taxonomicName id="4597ECC23060FF89FE8FFA96BC910800" authorityName="Alluaud" authorityYear="1936" box="[378,498,1284,1307]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="B0E34B533060FF89FE8FFA96BC910800" box="[378,498,1284,1307]" italics="true" pageId="25" pageNumber="226">P. prasinus</emphasis>
</taxonomicName>
), narrower body form, and dorsum without metallic reflection (dorsum with metallic green reflection in
<taxonomicName id="4597ECC23060FF89FDC9FAB6BFD10820" authorityName="Alluaud" authorityYear="1936" box="[572,690,1316,1339]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="B0E34B533060FF89FDC9FAB6BFD10820" box="[572,690,1316,1339]" italics="true" pageId="25" pageNumber="226">P. prasinus</emphasis>
</taxonomicName>
) but elytra with moderate but distinct iridescence generated by microsculpture comprised of transverse microlines and marjkedly transverse sculpticells. They are most similar to members of
<taxonomicName id="4597ECC23060FF89FD6AFAF7BE4C0867" authorityName="Alluaud" authorityYear="1936" box="[671,815,1380,1404]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533060FF89FD6AFAF7BE4C0867" box="[671,815,1380,1404]" italics="true" pageId="25" pageNumber="226">P. heterodera</emphasis>
</taxonomicName>
in body form and size but differ from them in having the pronotum proportionately longer and male genitalia (
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) with shaft markedly narrowed basally, markedly inflated in apical two-thirds, and apex longer, slender and ventrally bent in lateral view (
<figureCitation id="1AAC8BC43060FF89FE0DFA56BF3B08C7" box="[504,600,1476,1500]" captionStart="FIGURE" captionStartId="26.[155,167,1478,1497]" captionTargetBox="[139,1176,216,1455]" captionTargetId="figure-53@26.[120,1176,216,1455]" captionTargetPageId="26" captionText="FIGURE 14. Digital images of holotype male of Perigona (Ripogena) deuvei Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm." pageId="25" pageNumber="226">Fig. 14C</figureCitation>
), slightly deflected left and slightly longer in dorsal view (
<figureCitation id="1AAC8BC43060FF89FF4AFA76BC7C08E7" box="[191,287,1508,1533]" captionStart="FIGURE" captionStartId="26.[155,167,1478,1497]" captionTargetBox="[139,1176,216,1455]" captionTargetId="figure-53@26.[120,1176,216,1455]" captionTargetPageId="26" captionText="FIGURE 14. Digital images of holotype male of Perigona (Ripogena) deuvei Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm." pageId="25" pageNumber="226">Fig. 14D</figureCitation>
) than in
<taxonomicName id="4597ECC23060FF89FE77FA77BF7208E7" authorityName="Alluaud" authorityYear="1936" box="[386,529,1508,1532]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533060FF89FE77FA77BF7208E7" box="[386,529,1508,1532]" italics="true" pageId="25" pageNumber="226">P. heterodera</emphasis>
</taxonomicName>
(see Figs. 351b-c in Jeannel 1948). They differ from members of
<taxonomicName id="4597ECC23060FF89FF3AF997BF1C0B06" authority="(Deuve 1998)" baseAuthorityName="Deuve" baseAuthorityYear="1998" box="[207,639,1541,1565]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B0E34B533060FF89FF3AF997BC830B06" box="[207,480,1541,1565]" italics="true" pageId="25" pageNumber="226">Perigona descarpentriesi</emphasis>
(
<bibRefCitation id="E606EAB03060FF89FE07F997BF150B06" author="DEUVE, T." box="[498,630,1541,1565]" pageId="25" pageNumber="226" pagination="2 - 4" refId="ref30013" refString="DEUVE, T. 1998. Description d'un Perigoninae microphtalme de Madagascar [Coleoptera, Harpalidea]. Revue Francaise d'Entomologie (N. S.) 20: 2 - 4." type="journal article" year="1998">Deuve 1998</bibRefCitation>
)
</taxonomicName>
in having moderately large eyes (eyes markedly reduced eyes in size in
<taxonomicName id="4597ECC23060FF89FE84F9B4BF4C0B26" baseAuthorityName="Deuve" baseAuthorityYear="1998" box="[369,559,1573,1597]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B0E34B533060FF89FE84F9B4BF4C0B26" box="[369,559,1573,1597]" italics="true" pageId="25" pageNumber="226">P. descarpentriesi</emphasis>
</taxonomicName>
), pronotum distinctly narrower and with less rounded lateral margins, and elytra with only stria 1 deeply impressed, stria 2 and 3 shallowly impressed but evident on disk, striae 4 to 7 effaced, and stria 8 impressed only in apical one-third (striae 1–7 shallowly impressed but evident at least on disc and stria 8 deeply impressed throughout in
<taxonomicName id="4597ECC23060FF89FBDFF914BDBB0BA6" baseAuthorityName="Deuve" baseAuthorityYear="1998" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B0E34B533060FF89FBDFF914BDBB0BA6" italics="true" pageId="25" pageNumber="226">P. descarpentriesi</emphasis>
</taxonomicName>
members). Also, the apex of the median lobe of the male genitalia is narrower in lateral aspect in
<taxonomicName id="4597ECC23062FF8BFF15FF4EBC230DE8" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[224,320,219,243]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533062FF8BFF15FF4EBC230DE8" box="[224,320,219,243]" italics="true" pageId="27" pageNumber="228">P. deuvei</emphasis>
</taxonomicName>
males (Fig. 143C) than in those of
<taxonomicName id="4597ECC23062FF8BFD4EFF4EBE1A0DE8" baseAuthorityName="Deuve" baseAuthorityYear="1998" box="[699,889,219,243]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B0E34B533062FF8BFD4EFF4EBE1A0DE8" box="[699,889,219,243]" italics="true" pageId="27" pageNumber="228">P. descarpentriesi</emphasis>
</taxonomicName>
(see
<bibRefCitation id="E606EAB03062FF8BFC47FF4EB9500DE8" author="DEUVE, T." box="[946,1075,219,243]" pageId="27" pageNumber="228" pagination="2 - 4" refId="ref30013" refString="DEUVE, T. 1998. Description d'un Perigoninae microphtalme de Madagascar [Coleoptera, Harpalidea]. Revue Francaise d'Entomologie (N. S.) 20: 2 - 4." type="journal article" year="1998">Deuve 1998</bibRefCitation>
,
<figureCitation id="1AAC8BC43062FF8BFBB4FF49B91C0DE8" box="[1089,1151,219,243]" captionStart="FIGURE" captionStartId="5.[155,167,1152,1171]" captionTargetBox="[133,1174,245,1135]" captionTargetId="figure-997@5.[120,1174,216,1150]" captionTargetPageId="5" captionText="FIGURE 2. Digital images of holotype female of Brachypelus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." pageId="27" pageNumber="228">Fig. 2</figureCitation>
.). Finally,
<taxonomicName id="4597ECC23062FF8BFF26FF6EBC570C08" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[211,308,251,275]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533062FF8BFF26FF6EBC570C08" box="[211,308,251,275]" italics="true" pageId="27" pageNumber="228">P. deuvei</emphasis>
</taxonomicName>
members differ from those of the new species described below in having small- er body size, pronotum with lateral margination very narrow throughout, fewer elytral striae evident, and median lobe of the male genitalia with a longer and ventrally bent apex.
</paragraph>
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14. Digital images of holotype male of
<taxonomicName id="4597ECC23063FF8AFDC1FA54BE8608C2" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[564,997,1478,1497]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="deuvei" status="sp. nov." subGenus="Ripogena">
<emphasis id="B0E34B533063FF8AFDC1FA54BFE008C3" box="[564,643,1478,1496]" italics="true" pageId="26" pageNumber="227">Perigona</emphasis>
(
<emphasis id="B0E34B533063FF8AFD64FA54BF8008C3" box="[657,739,1478,1496]" italics="true" pageId="26" pageNumber="227">Ripogena</emphasis>
)
<emphasis id="B0E34B533063FF8AFD05FA54BE4B08C2" box="[752,808,1478,1497]" italics="true" pageId="26" pageNumber="227">deuvei</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283063FF8AFC1EFA54B94E08C2" box="[1003,1069,1478,1497]" pageId="26" pageNumber="227" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm.
</paragraph>
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<paragraph id="822897413062FF8BFF5EFEC9B9690C68" blockId="27.[123,1174,219,436]" box="[171,1034,347,372]" pageId="27" pageNumber="228">
<emphasis id="B0E34B533062FF8BFF5EFEC9BC920C69" bold="true" box="[171,497,347,371]" pageId="27" pageNumber="228">
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<smallCapsWord id="84CE019D3062FF8BFEA8FECDBC920C69" baselines="365" box="[349,497,351,370]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="27" pageNumber="228">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
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locality.
</paragraph>
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<emphasis id="B0E34B533062FF8BFF5EFEEEBCC70C89" bold="true" box="[171,420,380,404]" pageId="27" pageNumber="228">
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<smallCapsWord id="84CE019D3062FF8BFEE5FEEDBCC70C89" baselines="397" box="[272,420,383,402]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="27" pageNumber="228">DISTRIBUTION</smallCapsWord>
</emphasis>
.— All specimens of this species were collected by hand in daytime in primary montane rainforest in the Vatoharanana area at an elevation of
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.
</paragraph>
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<treatment id="0A3E26573062FF8CFF8EFE4EBC080B0A" LSID="urn:lsid:plazi:treatment:0A3E26573062FF8CFF8EFE4EBC080B0A" httpUri="http://treatment.plazi.org/id/0A3E26573062FF8CFF8EFE4EBC080B0A" lastPageId="28" lastPageNumber="228" pageId="27" pageNumber="228">
<subSubSection id="CA8DC4CA3062FF8BFF8EFE4EBE110CEF" box="[123,882,476,500]" pageId="27" pageNumber="228" type="nomenclature">
<paragraph id="822897413062FF8BFF8EFE4EBE110CEF" blockId="27.[123,882,476,530]" box="[123,882,476,500]" pageId="27" pageNumber="228">
<heading id="D960202D3062FF8BFF8EFE4EBE110CEF" box="[123,882,476,500]" fontSize="10" level="2" pageId="27" pageNumber="228" reason="4">
<taxonomicName id="4597ECC23062FF8BFF8EFE4EBE6D0CEF" ID-CoL="775QY" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,782,476,500]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov." subGenus="Ripogena">
<emphasis id="B0E34B533062FF8BFF8EFE4EBC3A0CEF" bold="true" box="[123,345,476,500]" pageId="27" pageNumber="228">
<emphasis id="B0E34B533062FF8BFF8EFE4EBD830CEF" bold="true" box="[123,224,476,500]" italics="true" pageId="27" pageNumber="228">Perigona</emphasis>
(
<emphasis id="B0E34B533062FF8BFF05FE4EBC3A0CEF" bold="true" box="[240,345,476,500]" italics="true" pageId="27" pageNumber="228">Ripogena</emphasis>
</emphasis>
)
<emphasis id="B0E34B533062FF8BFE9CFE4EBE6D0CEF" bold="true" box="[361,782,476,500]" pageId="27" pageNumber="228">
<emphasis id="B0E34B533062FF8BFE9CFE4EBC9E0CEF" bold="true" box="[361,509,476,500]" italics="true" pageId="27" pageNumber="228">ranomafanae</emphasis>
Kavanaugh and Rainio
</emphasis>
</taxonomicName>
<emphasis id="B0E34B533062FF8BFCFBFE4EBE110CEF" bold="true" box="[782,882,476,500]" pageId="27" pageNumber="228">
,
<taxonomicNameLabel id="ABD0F6283062FF8BFCE9FE4EBE110CEF" box="[796,882,476,500]" pageId="27" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3062FF8BFF8EFE6EBEA90F46" pageId="27" pageNumber="228" type="description">
<paragraph id="822897413062FF8BFF8EFE6EBDBB0F09" blockId="27.[123,882,476,530]" box="[123,216,508,530]" pageId="27" pageNumber="228">
<figureCitation id="1AAC8BC43062FF8BFF8EFE6EBDBB0F09" box="[123,216,508,530]" captionStart="FIGURE" captionStartId="28.[155,167,1395,1414]" captionTargetBox="[148,1176,233,1374]" captionTargetId="figure-419@28.[120,1176,216,1392]" captionTargetPageId="28" captionText="FIGURE 15. Digital images of holotype male of Perigona (Ripogena) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C. Aedeagus of genitalia, left lateral aspect [dissection not fully cleaned due to teneral nature of holotype]; D. Dorsal aspect. Scale lines A = 1.0 mm, C = 0.25 mm." pageId="27" pageNumber="228">Figure 15</figureCitation>
</paragraph>
<paragraph id="822897413062FF8BFF8EFDBABFD20F26" blockId="27.[123,970,551,605]" box="[123,689,551,573]" pageId="27" pageNumber="228">
<emphasis id="B0E34B533062FF8BFF8EFDBABC470F27" box="[123,292,551,573]" italics="true" pageId="27" pageNumber="228">
<taxonomicName id="4597ECC23062FF8BFF8EFDBABDB40F26" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,215,552,573]" pageId="27" pageNumber="224" rank="subGenus" status="n. sp." subGenus="Ripogena">Ripogena</taxonomicName>
<taxonomicNameLabel id="ABD0F6283062FF8BFF28FDBABC720F26" box="[221,273,552,573]" pageId="27" pageNumber="228" rank="species">n. sp.</taxonomicNameLabel>
2
</emphasis>
; Rainio 2009: 33 (informal designation).
</paragraph>
<paragraph id="822897413062FF8BFF8EFDDABEA90F46" blockId="27.[123,970,551,605]" box="[123,970,583,605]" pageId="27" pageNumber="228">
<taxonomicName id="4597ECC23062FF8BFF8EFDDABC240F47" authorityName="Jeannel" authorityYear="1941" box="[123,327,583,605]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="subGenus" subGenus="Ripogena">
<emphasis id="B0E34B533062FF8BFF8EFDDABDB70F46" box="[123,212,584,605]" italics="true" pageId="27" pageNumber="228">Perigona</emphasis>
(
<emphasis id="B0E34B533062FF8BFF16FDDABC5C0F46" box="[227,319,584,605]" italics="true" pageId="27" pageNumber="228">Ripogena</emphasis>
)
</taxonomicName>
<emphasis id="B0E34B533062FF8BFEB8FDDABCF70F47" box="[333,404,583,605]" italics="true" pageId="27" pageNumber="228">n. sp. 2</emphasis>
; Rainio 2012: 73, Rainio 2013: 96 (informal designation).
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3062FF8CFF5EFD13BC080B0A" lastPageId="28" lastPageNumber="229" pageId="27" pageNumber="228" type="materials_examined">
<paragraph id="822897413062FF8BFF5EFD13BE190E02" blockId="27.[123,1174,640,1726]" pageId="27" pageNumber="228">
<emphasis id="B0E34B533062FF8BFF5EFD13BC3A0F8C" bold="true" box="[171,345,641,665]" pageId="27" pageNumber="228">
<smallCapsWord id="84CE019D3062FF8BFF5EFD13BD870F8C" baselines="659,658" box="[171,228,641,665]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="27" pageNumber="228">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D3062FF8BFF19FD16BC3A0F8C" baselines="658" box="[236,345,644,663]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="27" pageNumber="228">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C3062FF8BFE74FD12BE190E02" collectingDate="1998-04-29" collectorName="D. H. Kavanaugh" country="Madagascar" county="Ranomafana National Park" elevation="1050" latitude="-21.278334" location="Abotovory" longLatPrecision="126" longitude="47.435" municipality="Vatoharanana area" pageId="27" pageNumber="228" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E33062FF8BFE74FD12BC870F83" box="[385,484,640,664]" pageId="27" pageNumber="228" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43062FF8BFE00FD12BF150F83" box="[501,630,640,665]" captionStart="FIGURE" captionStartId="28.[155,167,1395,1414]" captionTargetBox="[148,1176,233,1374]" captionTargetId="figure-419@28.[120,1176,216,1392]" captionTargetPageId="28" captionText="FIGURE 15. Digital images of holotype male of Perigona (Ripogena) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C. Aedeagus of genitalia, left lateral aspect [dissection not fully cleaned due to teneral nature of holotype]; D. Dorsal aspect. Scale lines A = 1.0 mm, C = 0.25 mm." pageId="27" pageNumber="228">Figs.15A–B</figureCitation>
), a male, in CAS, labeled: ““CASENT 1049017”/ “
<collectingCountry id="FA80D7D13062FF8BFF72FD33BC260FA2" box="[135,325,673,697]" name="Madagascar" pageId="27" pageNumber="228">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A33062FF8BFEA1FD33BF270FA2" box="[340,580,673,697]" country="Madagascar" name="Fianarantsoa" pageId="27" pageNumber="228">Fianarantsoa Province</collectingRegion>
,
<collectingCounty id="6B49EFCD3062FF8BFDA1FD33BE190FA2" box="[596,890,673,697]" pageId="27" pageNumber="228">Ranomafana National Park</collectingCounty>
,
<collectingMunicipality id="624C0D3B3062FF8BFC7CFD33B9330FA3" box="[905,1104,673,697]" pageId="27" pageNumber="228">Vatoharanana area</collectingMunicipality>
,
<quantity id="456F3AA43062FF8BFB95FD32BDF30FC3" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="27" pageNumber="228" unit="m" value="1050.0">
<elevation id="09BA70723062FF8BFB95FD32BDF30FC3" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="27" pageNumber="228" unit="m" value="1050.0">1050 m</elevation>
</quantity>
,
<location id="8748C19A3062FF8BFF6EFD53BC6C0FC2" LSID="urn:lsid:plazi:treatment:0A3E26573062FF8CFF8EFE4EBC080B0A:8748C19A3062FF8BFF6EFD53BC6C0FC2" box="[155,271,705,729]" country="Madagascar" county="Ranomafana National Park" latitude="-21.278334" longLatPrecision="126" longitude="47.435" municipality="Vatoharanana area" name="Abotovory" pageId="27" pageNumber="228" stateProvince="Fianarantsoa">Abotovory</location>
stream,
<date id="F629B1813062FF8BFE9DFD53BC990FC2" box="[360,506,705,729]" pageId="27" pageNumber="228" value="1998-04-29">
<collectingDate id="E66D48693062FF8BFE9DFD53BC990FC2" box="[360,506,705,729]" pageId="27" pageNumber="228" value="1998-04-29">29 April 1998</collectingDate>
</date>
”/ “
<geoCoordinate id="E7A3F1863062FF8BFDEAFD53BFE80FC2" box="[543,651,705,729]" degrees="21" direction="south" minutes="16. 7" orientation="latitude" pageId="27" pageNumber="228" precision="92" value="-21.278334">21°16.7’S</geoCoordinate>
<geoCoordinate id="E7A3F1863062FF8BFD64FD53BE670FC3" box="[657,772,705,729]" degrees="47" direction="east" minutes="26. 1" orientation="longitude" pageId="27" pageNumber="228" precision="92" value="47.435">47° 26.1’E</geoCoordinate>
, Stop # 98-100,
<collectorName id="2F62F2973062FF8BFC46FD53B9050FC2" box="[947,1126,705,729]" pageId="27" pageNumber="228">D.H. Kavanaugh</collectorName>
collector collected in large rotting log just under loose bark with cavities”/ “
<typeStatus id="5D2C29E33062FF8BFC68FD73B9490FE2" box="[925,1066,737,761]" pageId="27" pageNumber="228" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23062FF8BFBC7FD70BF100E02" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov." subGenus="Ripogena">
<emphasis id="B0E34B533062FF8BFBC7FD70B9F60FE2" box="[1074,1173,738,761]" italics="true" pageId="27" pageNumber="228">Perigona</emphasis>
(
<emphasis id="B0E34B533062FF8BFF71FC90BD890E02" box="[132,234,770,793]" italics="true" pageId="27" pageNumber="228">Ripogena</emphasis>
)
<emphasis id="B0E34B533062FF8BFF0FFC93BCEB0E02" box="[250,392,769,793]" italics="true" pageId="27" pageNumber="228">ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283062FF8BFD8EFC90BFD60E02" box="[635,693,770,793]" pageId="27" pageNumber="228" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
</materialsCitation>
</paragraph>
<paragraph id="822897413062FF8BFF5EFCB3B9500E22" blockId="27.[123,1174,640,1726]" box="[171,1075,801,825]" pageId="27" pageNumber="228">
<materialsCitation id="32FF9D1C3062FF8BFF5EFCB3B94F0E22" box="[171,1068,801,825]" country="Madagascar" location="Ranomafana National Park" pageId="27" pageNumber="228" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533062FF8BFF5EFCB3BC340E23" bold="true" box="[171,343,801,825]" pageId="27" pageNumber="228">
<smallCapsWord id="84CE019D3062FF8BFF5EFCB3BD870E23" baselines="819,819" box="[171,228,801,825]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="27" pageNumber="228">
<typeStatus id="5D2C29E33062FF8BFF5EFCB3BD870E23" box="[171,228,801,825]" pageId="27" pageNumber="228">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3062FF8BFF19FCB7BC340E23" baselines="819" box="[236,343,805,824]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="27" pageNumber="228">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13062FF8BFE8BFCB3BC9C0E22" box="[382,511,801,825]" name="Madagascar" pageId="27" pageNumber="228">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33062FF8BFDF8FCB3BF9F0E22" box="[525,764,801,825]" country="Madagascar" name="Fianarantsoa" pageId="27" pageNumber="228">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3062FF8BFCFFFCB3B94F0E22" LSID="urn:lsid:plazi:treatment:0A3E26573062FF8CFF8EFE4EBC080B0A:8748C19A3062FF8BFCFFFCB3B94F0E22" box="[778,1068,801,825]" country="Madagascar" name="Ranomafana National Park" pageId="27" pageNumber="228" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413062FF8BFF5EFCD0BECA0E62" blockId="27.[123,1174,640,1726]" pageId="27" pageNumber="228">
<emphasis id="B0E34B533062FF8BFF5EFCD0BC910E43" bold="true" box="[171,498,834,858]" pageId="27" pageNumber="228">
<smallCapsWord id="84CE019D3062FF8BFF5EFCD0BC4D0E43" baselines="852,851" box="[171,302,834,858]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="27" pageNumber="228">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3062FF8BFEC3FCD7BC310E43" baselines="851" box="[310,338,837,856]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="27" pageNumber="228">OF</smallCapsWord>
<smallCapsWord id="84CE019D3062FF8BFEACFCD7BCCF0E43" baselines="851" box="[345,428,837,856]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="27" pageNumber="228">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3062FF8BFE40FCD7BC910E43" baselines="851" box="[437,498,837,856]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="27" pageNumber="228">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23062FF8BFD0CFCD3BEE40E42" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[761,903,833,857]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533062FF8BFD0CFCD3BEE40E42" box="[761,903,833,857]" italics="true" pageId="27" pageNumber="228">ranomafanae</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the national park in which the
<typeStatus id="5D2C29E33062FF8BFD29FCF0BE690E61" box="[732,778,866,890]" pageId="27" pageNumber="228">type</typeStatus>
was collected.
</paragraph>
<paragraph id="822897413062FF8BFF5EFC10BFD50B66" blockId="27.[123,1174,640,1726]" pageId="27" pageNumber="228">
<smallCapsWord id="84CE019D3062FF8BFF5EFC10BC210E83" baselines="916,915" box="[171,322,898,922]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="27" pageNumber="228">
<emphasis id="B0E34B533062FF8BFF5EFC10BC210E83" bold="true" box="[171,322,898,922]" pageId="27" pageNumber="228">RECOGNITION</emphasis>
</smallCapsWord>
.— Size larger than average for subgenus, SBL =
<quantity id="456F3AA43062FF8BFCA6FC10BEC70E82" box="[851,932,898,922]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="27" pageNumber="228" unit="mm" value="4.4">4.4 mm</quantity>
. The
<typeStatus id="5D2C29E33062FF8BFC14FC10B95D0E81" box="[993,1086,898,922]" pageId="27" pageNumber="228" type="holotype">holotype</typeStatus>
male of this species (
<figureCitation id="1AAC8BC43062FF8BFEFEFC30BC0F0EA1" box="[267,364,930,954]" captionStart="FIGURE" captionStartId="28.[155,167,1395,1414]" captionTargetBox="[148,1176,233,1374]" captionTargetId="figure-419@28.[120,1176,216,1392]" captionTargetPageId="28" captionText="FIGURE 15. Digital images of holotype male of Perigona (Ripogena) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C. Aedeagus of genitalia, left lateral aspect [dissection not fully cleaned due to teneral nature of holotype]; D. Dorsal aspect. Scale lines A = 1.0 mm, C = 0.25 mm." pageId="27" pageNumber="228">Fig. 15A</figureCitation>
) shares with members of
<taxonomicName id="4597ECC23062FF8BFD65FC31B9410EA1" authority="Alluaud (1936)" authorityName="Alluaud" authorityYear="1936" box="[656,1058,930,954]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533062FF8BFD65FC31BE120EA1" box="[656,881,930,954]" italics="true" pageId="27" pageNumber="228">Perigona heterodera</emphasis>
Alluaud (1936)
</taxonomicName>
,
<taxonomicName id="4597ECC23062FF8BFBC7FC31BCE20EC1" authority="Alluaud (1936)" authorityName="Alluaud" authorityYear="1936" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="B0E34B533062FF8BFBC7FC31BDB40EC1" italics="true" pageId="27" pageNumber="228">Perigona prasinus</emphasis>
Alluaud (1936)
</taxonomicName>
and
<taxonomicName id="4597ECC23062FF8BFE43FC51BF740EC1" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[438,535,962,986]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="deuvei" status="sp. nov.">
<emphasis id="B0E34B533062FF8BFE43FC51BF740EC1" box="[438,535,962,986]" italics="true" pageId="27" pageNumber="228">P. deuvei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283062FF8BFDEBFC51BF120EC1" box="[542,625,963,986]" pageId="27" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
the following features: pronotum with lateral margins without sinuation anterior to hind angles, posterior margin slightly and smoothly convex, and hind angles either obtuse or broadly rounded. These features distinguish members of these four species from those of
<taxonomicName id="4597ECC23062FF8BFE98FBB1BE770920" authority="Alluaud (1936)" authorityName="Alluaud" authorityYear="1936" box="[365,788,1058,1083]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="bembidioides">
<emphasis id="B0E34B533062FF8BFE98FBB1BF040921" box="[365,615,1058,1082]" italics="true" pageId="27" pageNumber="228">Perigona bembidioides</emphasis>
Alluaud (1936)
</taxonomicName>
and
<taxonomicName id="4597ECC23062FF8BFCBEFBB1BDA10940" authority="Jeannel (1948)" authorityName="Jeannel" authorityYear="1948" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="viridimicans">
<emphasis id="B0E34B533062FF8BFCBEFBB1B95F0921" box="[843,1084,1058,1082]" italics="true" pageId="27" pageNumber="228">Perigona viridimicans</emphasis>
Jeannel (1948)
</taxonomicName>
, which have distinct sinuation of the lateral margins anterior to rectangular hind angles and a straight basal margin. Members of
<taxonomicName id="4597ECC23062FF8BFDE6FBF6BFDC0960" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[531,703,1123,1147]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533062FF8BFDE6FBF6BFDC0960" box="[531,703,1123,1147]" italics="true" pageId="27" pageNumber="228">P. ranomafanae</emphasis>
</taxonomicName>
differ from those of
<taxonomicName id="4597ECC23062FF8BFC59FBF6B9460961" authorityName="Alluaud" authorityYear="1936" box="[940,1061,1123,1147]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="B0E34B533062FF8BFC59FBF6B9460961" box="[940,1061,1123,1147]" italics="true" pageId="27" pageNumber="228">P. prasinus</emphasis>
</taxonomicName>
in having slightly smaller body size and dorsum without metallic reflection (dorsum with metallic green reflection in
<taxonomicName id="4597ECC23062FF8BFEF6FB36BC1909A0" authorityName="Alluaud" authorityYear="1936" box="[259,378,1188,1211]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="B0E34B533062FF8BFEF6FB36BC1909A0" box="[259,378,1188,1211]" italics="true" pageId="27" pageNumber="228">P. prasinus</emphasis>
</taxonomicName>
). They differ from members of
<taxonomicName id="4597ECC23062FF8BFD3AFB36BE3E09A0" authorityName="Alluaud" authorityYear="1936" box="[719,861,1187,1211]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533062FF8BFD3AFB36BE3E09A0" box="[719,861,1187,1211]" italics="true" pageId="27" pageNumber="228">P. heterodera</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC23062FF8BFC67FB36BE9109A0" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[914,1010,1187,1211]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533062FF8BFC67FB36BE9109A0" box="[914,1010,1187,1211]" italics="true" pageId="27" pageNumber="228">P. deuvei</emphasis>
</taxonomicName>
in having larg- er body size (SBL =
<quantity id="456F3AA43062FF8BFEAAFB51BCD009C0" box="[351,435,1219,1243]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="27" pageNumber="228" unit="mm" value="3.4">3.4 mm</quantity>
in
<taxonomicName id="4597ECC23062FF8BFE2FFB56BF0909C0" authorityName="Alluaud" authorityYear="1936" box="[474,618,1219,1243]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533062FF8BFE2FFB56BF0909C0" box="[474,618,1219,1243]" italics="true" pageId="27" pageNumber="228">P. heterodera</emphasis>
</taxonomicName>
, 3.5 to
<quantity id="456F3AA43062FF8BFD37FB51BE7609C0" box="[706,789,1219,1243]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="27" pageNumber="228" unit="mm" value="3.9">3.9 mm</quantity>
in
<taxonomicName id="4597ECC23062FF8BFCC9FB56BEFC09C0" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[828,927,1219,1243]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533062FF8BFCC9FB56BEFC09C0" box="[828,927,1219,1243]" italics="true" pageId="27" pageNumber="228">P. deuvei</emphasis>
</taxonomicName>
), pronotal hind angles more sharply angulate, lateral explanation of pronotum markedly widened and flattened in region of hind angles (pronotal hind angles rounded or at least less sharply angulate and lateral explanation only slightly wider near hind angles than anteriorly in
<taxonomicName id="4597ECC23062FF8BFD1CFAB6BE150827" authorityName="Alluaud" authorityYear="1936" box="[745,886,1316,1340]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533062FF8BFD1CFAB6BE150827" box="[745,886,1316,1340]" italics="true" pageId="27" pageNumber="228">P. heterodera</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC23062FF8BFC5CFAB6B96B0827" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[937,1032,1316,1340]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533062FF8BFC5CFAB6B96B0827" box="[937,1032,1316,1340]" italics="true" pageId="27" pageNumber="228">P. deuvei</emphasis>
</taxonomicName>
). The unique male
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of
<taxonomicName id="4597ECC23062FF8BFEC2FAD7BC830847" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[311,480,1348,1372]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533062FF8BFEC2FAD7BC830847" box="[311,480,1348,1372]" italics="true" pageId="27" pageNumber="228">P. ranomafanae</emphasis>
</taxonomicName>
is slightly teneral and the genitalia are very soft and only lightly sclerotized. However, the apex is sclerotized sufficiently to allow comparisons with those of
<taxonomicName id="4597ECC23062FF8BFBA7FAF7BDAD0887" authorityName="Alluaud" authorityYear="1936" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533062FF8BFBA7FAF7BDAD0887" italics="true" pageId="27" pageNumber="228">P. heterodera</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC23062FF8BFEF1FA17BC050887" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[260,358,1412,1436]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533062FF8BFEF1FA17BC050887" box="[260,358,1412,1436]" italics="true" pageId="27" pageNumber="228">P. deuvei</emphasis>
</taxonomicName>
males. In lateral view (
<figureCitation id="1AAC8BC43062FF8BFD92FA16BFA50887" box="[615,710,1412,1436]" captionStart="FIGURE" captionStartId="28.[155,167,1395,1414]" captionTargetBox="[148,1176,233,1374]" captionTargetId="figure-419@28.[120,1176,216,1392]" captionTargetPageId="28" captionText="FIGURE 15. Digital images of holotype male of Perigona (Ripogena) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C. Aedeagus of genitalia, left lateral aspect [dissection not fully cleaned due to teneral nature of holotype]; D. Dorsal aspect. Scale lines A = 1.0 mm, C = 0.25 mm." pageId="27" pageNumber="228">Fig. 15C</figureCitation>
), the apex is slightly longer than in
<taxonomicName id="4597ECC23062FF8BFBA5FA17BDAD08A7" authorityName="Alluaud" authorityYear="1936" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533062FF8BFBA5FA17BDAD08A7" italics="true" pageId="27" pageNumber="228">P. heterodera</emphasis>
</taxonomicName>
males (see Jeannel 1948, Fig. 351b) and distinctly shorter and straighter than in
<taxonomicName id="4597ECC23062FF8BFBC0FA37B9F508A7" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[1077,1174,1444,1468]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533062FF8BFBC0FA37B9F508A7" box="[1077,1174,1444,1468]" italics="true" pageId="27" pageNumber="228">P. deuvei</emphasis>
</taxonomicName>
males. Finally, they differ from members of
<taxonomicName id="4597ECC23062FF8BFD9CFA57BE4908C7" baseAuthorityName="Deuve" baseAuthorityYear="1998" box="[617,810,1476,1500]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B0E34B533062FF8BFD9CFA57BE4908C7" box="[617,810,1476,1500]" italics="true" pageId="27" pageNumber="228">P. descarpentriesi</emphasis>
</taxonomicName>
in having moderately large eyes (eyes markedly reduced eyes in size in
<taxonomicName id="4597ECC23062FF8BFDEEFA77BFBA08E6" baseAuthorityName="Deuve" baseAuthorityYear="1998" box="[539,729,1509,1533]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B0E34B533062FF8BFDEEFA77BFBA08E6" box="[539,729,1509,1533]" italics="true" pageId="27" pageNumber="228">P. descarpentriesi</emphasis>
</taxonomicName>
), pronotum widest anterior to middle and lateral margins nearly straight in posterior half (pronotum widest at middle and lateral margins evenly arcuate throughout
<taxonomicName id="4597ECC23062FF8BFE6CF9B4BF3B0B26" baseAuthorityName="Deuve" baseAuthorityYear="1998" box="[409,600,1573,1597]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B0E34B533062FF8BFE6CF9B4BF3B0B26" box="[409,600,1573,1597]" italics="true" pageId="27" pageNumber="228">P. descarpentriesi</emphasis>
</taxonomicName>
). Also, the apex of the median lobe of the male genitalia is narrower and more pointed in lateral aspect in
<taxonomicName id="4597ECC23062FF8BFD26F9D4BE1D0B46" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[723,894,1605,1629]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533062FF8BFD26F9D4BE1D0B46" box="[723,894,1605,1629]" italics="true" pageId="27" pageNumber="228">P. ranomafanae</emphasis>
</taxonomicName>
males (
<figureCitation id="1AAC8BC43062FF8BFC22F9D7B95B0B46" box="[983,1080,1605,1629]" captionStart="FIGURE" captionStartId="28.[155,167,1395,1414]" captionTargetBox="[148,1176,233,1374]" captionTargetId="figure-419@28.[120,1176,216,1392]" captionTargetPageId="28" captionText="FIGURE 15. Digital images of holotype male of Perigona (Ripogena) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C. Aedeagus of genitalia, left lateral aspect [dissection not fully cleaned due to teneral nature of holotype]; D. Dorsal aspect. Scale lines A = 1.0 mm, C = 0.25 mm." pageId="27" pageNumber="228">Fig. 15C</figureCitation>
) than in those of
<taxonomicName id="4597ECC23062FF8BFF2DF9F4BCF40B66" baseAuthorityName="Deuve" baseAuthorityYear="1998" box="[216,407,1637,1661]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B0E34B533062FF8BFF2DF9F4BCF40B66" box="[216,407,1637,1661]" italics="true" pageId="27" pageNumber="228">P. descarpentriesi</emphasis>
</taxonomicName>
(see
<bibRefCitation id="E606EAB03062FF8BFE24F9F4BF300B66" author="DEUVE, T." box="[465,595,1637,1661]" pageId="27" pageNumber="228" pagination="2 - 4" refId="ref30013" refString="DEUVE, T. 1998. Description d'un Perigoninae microphtalme de Madagascar [Coleoptera, Harpalidea]. Revue Francaise d'Entomologie (N. S.) 20: 2 - 4." type="journal article" year="1998">Deuve 1998</bibRefCitation>
,
<figureCitation id="1AAC8BC43062FF8BFD94F9F7BFC30B66" box="[609,672,1637,1661]" captionStart="FIGURE" captionStartId="5.[155,167,1152,1171]" captionTargetBox="[133,1174,245,1135]" captionTargetId="figure-997@5.[120,1174,216,1150]" captionTargetPageId="5" captionText="FIGURE 2. Digital images of holotype female of Brachypelus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." pageId="27" pageNumber="228">Fig. 2</figureCitation>
.).
</paragraph>
<paragraph id="822897413062FF8BFF5EF917B9690B86" blockId="27.[123,1174,640,1726]" box="[171,1034,1669,1694]" pageId="27" pageNumber="228">
<emphasis id="B0E34B533062FF8BFF5EF917BC920B87" bold="true" box="[171,497,1669,1693]" pageId="27" pageNumber="228">
<smallCapsWord id="84CE019D3062FF8BFF5EF917BC350B87" baselines="1687,1687" box="[171,342,1669,1693]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="27" pageNumber="228">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="84CE019D3062FF8BFEA8F91BBC920B87" baselines="1687" box="[349,497,1673,1692]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="27" pageNumber="228">DISTRIBUTION</smallCapsWord>
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.— At present, known only from the
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locality.
</paragraph>
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<emphasis id="B0E34B533062FF8BFF5EF934BCC40BA7" bold="true" box="[171,423,1702,1726]" pageId="27" pageNumber="228">
<smallCapsWord id="84CE019D3062FF8BFF5EF934BC6A0BA7" baselines="1720,1719" box="[171,265,1702,1726]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="27" pageNumber="228">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D3062FF8BFEE6F93BBCC40BA7" baselines="1719" box="[275,423,1705,1724]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="27" pageNumber="228">DISTRIBUTION</smallCapsWord>
</emphasis>
.— The unique
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was collected by hand in primary montane rainforest in the Vatoharanana area at an elevation of
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from under the loose bark of a large log on the forest floor.
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285
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<mods:title id="3A20BE4D682CCFBF44D8117A47235DD8">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:affiliation id="E89F4CA5E0306073F786711E821044C0">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:affiliation id="F674A1653C6EB868235E6EF86A86289A">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
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Kavanaugh and Rainio
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,
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</emphasis>
.—
<materialsCitation id="32FF9D1C3064FF8DFE8AFDF6BF980FC7" collectingDate="1988-10-22" collectingDateMax="1988-10-31" collectingDateMin="1988-10-22" collectorName="W. E. Steiner" country="Madagascar" elevation="1100" location="Ranomafana" pageId="29" pageNumber="230" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E33064FF8DFE8AFDF6BC810F67" box="[383,482,612,636]" pageId="29" pageNumber="230" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43064FF8DFE07FDF6BF100F67" box="[498,627,612,636]" captionStart="FIGURE" captionStartId="30.[155,167,1246,1265]" captionTargetBox="[120,1176,216,1220]" captionTargetId="figure-760@30.[120,1176,216,1239]" captionTargetPageId="30" captionText="FIGURE 16. Digital images of holotype female of Archicolliuris ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." pageId="29" pageNumber="230">Figs.16A–B</figureCitation>
), a female, in NMNH, labeled: “
<collectingCountry id="FA80D7D13064FF8DFC25FDF6B9ED0F67" box="[976,1166,612,636]" name="Madagascar" pageId="29" pageNumber="230">MADAGASCAR</collectingCountry>
: Prov.
<collectingRegion id="405359A33064FF8DFF4DFD16BC230F87" box="[184,320,644,668]" country="Madagascar" name="Fianarantsoa" pageId="29" pageNumber="230">Fianarantsoa</collectingRegion>
,
<quantity id="456F3AA43064FF8DFEB8FD17BCE10F87" box="[333,386,644,668]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="29" pageNumber="230" unit="km" value="7.0">7 km</quantity>
W
<location id="8748C19A3064FF8DFE52FD16BF4D0F87" LSID="urn:lsid:plazi:treatment:0A3E26573064FF8EFF8EFD99BEDB0B4C:8748C19A3064FF8DFE52FD16BF4D0F87" box="[423,558,644,668]" country="Madagascar" name="Ranomafana" pageId="29" pageNumber="230" stateProvince="Fianarantsoa">Ranomafana</location>
,
<quantity id="456F3AA43064FF8DFDCFFD16BFE70F87" box="[570,644,644,668]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="29" pageNumber="230" unit="m" value="1100.0">
<elevation id="09BA70723064FF8DFDCFFD16BFE70F87" box="[570,644,644,668]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="29" pageNumber="230" unit="m" value="1100.0">1100m</elevation>
</quantity>
<date id="F629B1813064FF8DFD7CFD16BE3C0F87" box="[649,863,644,668]" pageId="29" pageNumber="230" value="1988-10-22" valueMax="1988-10-31" valueMin="1988-10-22">
<collectingDate id="E66D48693064FF8DFD7CFD16BE3C0F87" box="[649,863,644,668]" pageId="29" pageNumber="230" value="1988-10-22" valueMax="1988-10-31" valueMin="1988-10-22">22-31 October 1988</collectingDate>
</date>
<collectorName id="2F62F2973064FF8DFC90FD17BE920F87" box="[869,1009,644,668]" pageId="29" pageNumber="230">W. E. Steiner</collectorName>
”/ “
<collectingMethod id="5BD6EF563064FF8DFBE3FD16BC0E0FA7" pageId="29" pageNumber="230">Flight intercept-yellow pan trap</collectingMethod>
, island in stream, montane rainforest”/ “
<typeStatus id="5D2C29E33064FF8DFC94FD36BE8C0FA7" box="[865,1007,676,700]" pageId="29" pageNumber="230" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23064FF8DFBF5FD36BC970FC7" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Archicolliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533064FF8DFBF5FD36BC6A0FC7" italics="true" pageId="29" pageNumber="230">Archicolliuris ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283064FF8DFE0EFD57BF550FC7" box="[507,566,709,732]" pageId="29" pageNumber="230" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
</materialsCitation>
</paragraph>
<paragraph id="822897413064FF8DFF5EFD77B9500FE7" blockId="29.[123,1174,612,1726]" box="[171,1075,740,765]" pageId="29" pageNumber="230">
<materialsCitation id="32FF9D1C3064FF8DFF5EFD77B94F0FE7" box="[171,1068,740,765]" country="Madagascar" location="Ranomafana National Park" pageId="29" pageNumber="230" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533064FF8DFF5EFD77BC340FE0" bold="true" box="[171,343,741,765]" pageId="29" pageNumber="230">
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<typeStatus id="5D2C29E33064FF8DFF5EFD77BD870FE0" box="[171,228,741,765]" pageId="29" pageNumber="230">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3064FF8DFF19FD7ABC340FE0" baselines="758" box="[236,343,744,763]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="29" pageNumber="230">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13064FF8DFE8BFD76BC9C0FE7" box="[382,511,740,764]" name="Madagascar" pageId="29" pageNumber="230">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33064FF8DFDF8FD76BF9F0FE7" box="[525,764,740,764]" country="Madagascar" name="Fianarantsoa" pageId="29" pageNumber="230">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3064FF8DFCFFFD76B94F0FE7" LSID="urn:lsid:plazi:treatment:0A3E26573064FF8EFF8EFD99BEDB0B4C:8748C19A3064FF8DFCFFFD76B94F0FE7" box="[778,1068,740,764]" country="Madagascar" name="Ranomafana National Park" pageId="29" pageNumber="230" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413064FF8DFF5EFC97BECA0E27" blockId="29.[123,1174,612,1726]" pageId="29" pageNumber="230">
<emphasis id="B0E34B533064FF8DFF5EFC97BC910E00" bold="true" box="[171,498,773,797]" pageId="29" pageNumber="230">
<smallCapsWord id="84CE019D3064FF8DFF5EFC97BC4D0E00" baselines="791,790" box="[171,302,773,797]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="29" pageNumber="230">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3064FF8DFEC3FC9ABC310E00" baselines="790" box="[310,338,776,795]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="29" pageNumber="230">OF</smallCapsWord>
<smallCapsWord id="84CE019D3064FF8DFEACFC9ABCCF0E00" baselines="790" box="[345,428,776,795]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="29" pageNumber="230">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3064FF8DFE40FC9ABC910E00" baselines="790" box="[437,498,776,795]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="29" pageNumber="230">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23064FF8DFD0CFC97BEE40E06" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[761,903,773,797]" class="Insecta" family="Carabidae" genus="Archicolliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533064FF8DFD0CFC97BEE40E06" box="[761,903,773,797]" italics="true" pageId="29" pageNumber="230">ranomafanae</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the national park in which the
<typeStatus id="5D2C29E33064FF8DFD29FCB7BE690E26" box="[732,778,805,829]" pageId="29" pageNumber="230">type</typeStatus>
was collected.
</paragraph>
<paragraph id="822897413064FF8DFF5EFCD7BFD10B86" blockId="29.[123,1174,612,1726]" pageId="29" pageNumber="230">
<smallCapsWord id="84CE019D3064FF8DFF5EFCD7BC210E40" baselines="855,854" box="[171,322,837,861]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="29" pageNumber="230">
<emphasis id="B0E34B533064FF8DFF5EFCD7BC210E40" bold="true" box="[171,322,837,861]" pageId="29" pageNumber="230">RECOGNITION</emphasis>
</smallCapsWord>
.— Size large for genus, SBL =
<quantity id="456F3AA43064FF8DFD53FCD7BF990E47" box="[678,762,837,861]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="29" pageNumber="230" unit="mm" value="8.2">8.2 mm</quantity>
. The
<typeStatus id="5D2C29E33064FF8DFCCBFCD6BEF80E47" box="[830,923,836,860]" pageId="29" pageNumber="230" type="holotype">holotype</typeStatus>
female of this species (
<figureCitation id="1AAC8BC43064FF8DFF71FCF6BD800E66" box="[132,227,868,893]" captionStart="FIGURE" captionStartId="30.[155,167,1246,1265]" captionTargetBox="[120,1176,216,1220]" captionTargetId="figure-760@30.[120,1176,216,1239]" captionTargetPageId="30" captionText="FIGURE 16. Digital images of holotype female of Archicolliuris ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." pageId="29" pageNumber="230">Fig. 16A</figureCitation>
) is easily distinguished from members of all other Malagasy odacanthine species based on numerous features. The following features used by Jeannel (1948) in his key to genera are shared with the three species that he included in “
<taxonomicName id="4597ECC23064FF8DFD79FC37BF910EA7" authorityName="Latreille &amp; Dejean" authorityYear="1822" box="[652,754,933,956]" class="Insecta" family="Carabidae" genus="Casnonia" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533064FF8DFD79FC37BF910EA7" box="[652,754,933,956]" italics="true" pageId="29" pageNumber="230">Casnonia</emphasis>
</taxonomicName>
”: integument glabrous (without pubescence), pronotum long, narrow and tubular anteriorly, impunctate, with one or more setae present along each lateral margin. The
<typeStatus id="5D2C29E33064FF8DFE34FC77BF7D0EE6" box="[449,542,997,1021]" pageId="29" pageNumber="230" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4597ECC23064FF8DFDB4FC77BF880EE6" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[577,747,997,1021]" class="Insecta" family="Carabidae" genus="Archicolliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533064FF8DFDB4FC77BF880EE6" box="[577,747,997,1021]" italics="true" pageId="29" pageNumber="230">A. ranomafanae</emphasis>
</taxonomicName>
differs from members of
<taxonomicName id="4597ECC23064FF8DFC0FFC77BC060906" authority="Gestro (1895)" authorityName="Gestro" authorityYear="1895" class="Insecta" family="Carabidae" genus="Casnonia" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="fairmairei">
<emphasis id="B0E34B533064FF8DFC0FFC77BDA30907" italics="true" pageId="29" pageNumber="230">Casnonia fairmairei</emphasis>
<bibRefCitation id="E606EAB03064FF8DFF3EFB97BC060906" author="GESTRO, R." box="[203,357,1029,1053]" pageId="29" pageNumber="230" pagination="247 - 478" refId="ref30448" refString="GESTRO, R. 1895. Esplorazione del Giuba e dei suoi affluienti compiuta dal Cap. V. Bottego durante gli anni 1892 - 93 sotto gli auspicii della Societa Geografica Italiana. Risultati zoologici. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova 35: 247 - 478." type="journal article" year="1895">Gestro (1895)</bibRefCitation>
</taxonomicName>
[currently included in genus
<taxonomicName id="4597ECC23064FF8DFD4DFB97B9630906" authority="Liebke (1931)" authorityName="Liebke" authorityYear="1931" box="[696,1024,1029,1053]" class="Insecta" family="Carabidae" genus="Erectocolliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533064FF8DFD4DFB97BE340906" box="[696,855,1029,1053]" italics="true" pageId="29" pageNumber="230">Erectocolliuris</emphasis>
<bibRefCitation id="E606EAB03064FF8DFC96FB97B9630906" author="LIEBKE, M." box="[867,1024,1029,1053]" pageId="29" pageNumber="230" pagination="280 - 301" refId="ref31583" refString="LIEBKE, M. 1931. Die Africanischen Arten der Gattung Colliuris Degeer (Col. Car.). Revue de Zoologie et de Botanique Africaines 20 (1930 - 31): 280 - 301." type="journal article" year="1931">Liebke (1931)</bibRefCitation>
</taxonomicName>
] in having a pronotum black in color and with only a single pair of midlateral setae (pronotum rufotestaceous in color and with 5 or 6 pairs of lateral setae present in
<taxonomicName id="4597ECC23064FF8DFD1FFBD4BE160946" authorityName="Gestro" authorityYear="1895" box="[746,885,1093,1117]" class="Insecta" family="Carabidae" genus="Casnonia" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="fairmairei">
<emphasis id="B0E34B533064FF8DFD1FFBD4BE160946" box="[746,885,1093,1117]" italics="true" pageId="29" pageNumber="230">E. fairmairei</emphasis>
</taxonomicName>
members). It differs from members
<taxonomicName id="4597ECC23064FF8DFF11FBF7BF920966" authority="Kunckel d'Herculais (1887)" authorityName="Kunckel d'Herculais" authorityYear="1887" box="[228,753,1125,1149]" class="Insecta" family="Carabidae" genus="Casnonia" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="coerulans">
<emphasis id="B0E34B533064FF8DFF11FBF7BCD80966" box="[228,443,1125,1149]" italics="true" pageId="29" pageNumber="230">Casnonia coerulans</emphasis>
Künckel d’Herculais (1887)
</taxonomicName>
[currently included in genus
<taxonomicName id="4597ECC23064FF8DFBC5FBF7BC300986" authority="Liebke (1931)" authorityName="Liebke" authorityYear="1931" class="Insecta" family="Carabidae" genus="Protocolliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533064FF8DFBC5FBF7BDD00986" italics="true" pageId="29" pageNumber="230">Protocolliuris</emphasis>
<bibRefCitation id="E606EAB03064FF8DFF4FFB17BC300986" author="LIEBKE, M." box="[186,339,1157,1181]" pageId="29" pageNumber="230" pagination="280 - 301" refId="ref31583" refString="LIEBKE, M. 1931. Die Africanischen Arten der Gattung Colliuris Degeer (Col. Car.). Revue de Zoologie et de Botanique Africaines 20 (1930 - 31): 280 - 301." type="journal article" year="1931">Liebke (1931)</bibRefCitation>
</taxonomicName>
in having the pronotum only about twice as long as wide (three times as long as wide in
<taxonomicName id="4597ECC23064FF8DFF03FB34BC1E09A6" authorityName="Kunckel d'Herculais" authorityYear="1887" box="[246,381,1189,1213]" class="Insecta" family="Carabidae" genus="Casnonia" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="coerulans">
<emphasis id="B0E34B533064FF8DFF03FB34BC1E09A6" box="[246,381,1189,1213]" italics="true" pageId="29" pageNumber="230">P. coerulans</emphasis>
</taxonomicName>
), the elytra without metallic reflection (elytra with a dark metallic blue reflection), and legs dark black to piceous, except trochanthers rufous and basal parts of all femora pale (legs pale in
<taxonomicName id="4597ECC23064FF8DFEADFB74BCBE09E6" authorityName="Kunckel d'Herculais" authorityYear="1887" box="[344,477,1253,1277]" class="Insecta" family="Carabidae" genus="Casnonia" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="coerulans">
<emphasis id="B0E34B533064FF8DFEADFB74BCBE09E6" box="[344,477,1253,1277]" italics="true" pageId="29" pageNumber="230">P. coerulans</emphasis>
</taxonomicName>
, except apical parts of femora dark). It is similar to members of the last of Jeannel’s “
<taxonomicName id="4597ECC23064FF8DFEABFA97BCA70807" authorityName="Latreille &amp; Dejean" authorityYear="1822" box="[350,452,1285,1308]" class="Insecta" family="Carabidae" genus="Casnonia" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533064FF8DFEABFA97BCA70807" box="[350,452,1285,1308]" italics="true" pageId="29" pageNumber="230">Casnonia</emphasis>
</taxonomicName>
” species,
<taxonomicName id="4597ECC23064FF8DFDC5FA97BECA0806" authority="Alluaud (1935)" authorityName="Alluaud" authorityYear="1935" box="[560,937,1285,1309]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFDC5FA97BE630806" box="[560,768,1285,1309]" italics="true" pageId="29" pageNumber="230">Colliuris olsoufieffi</emphasis>
<bibRefCitation id="E606EAB03064FF8DFCF0FA97BECA0806" author="ALLUAUD, C." box="[773,937,1285,1309]" pageId="29" pageNumber="230" pagination="1 - 27" refId="ref28945" refString="ALLUAUD, C. 1935. Carabidae recueillis a Madagascar par MM. A. Syrig et G. Olsoufieff de 1933 a 1935. Afra, Cahiers d'Entomologie 10: 1 - 27." type="journal article" year="1935">Alluaud (1935)</bibRefCitation>
</taxonomicName>
[currently included in
<taxonomicName id="4597ECC23064FF8DFF8EFAB7BC6D0826" authorityName="Liebke" authorityYear="1931" box="[123,270,1317,1341]" class="Insecta" family="Carabidae" genus="Archicolliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533064FF8DFF8EFAB7BC6D0826" box="[123,270,1317,1341]" italics="true" pageId="29" pageNumber="230">Archicolliuris</emphasis>
</taxonomicName>
] in having a shiny black dorsum (with the only pale areas present as elytral pale spots), the pronotum with transverse grooves and ridges in the basal half and anterior angles projected laterally, and the elytra with a deep transverse depression at the basal one-fourth and with three or four discal setiferous pores on elytral interval 3. However, it differs with members of
<taxonomicName id="4597ECC23064FF8DFF8EFA37BC6B08A6" authorityName="Alluaud" authorityYear="1935" box="[123,264,1445,1469]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFF8EFA37BC6B08A6" box="[123,264,1445,1469]" italics="true" pageId="29" pageNumber="230">A. olsoufieffi</emphasis>
</taxonomicName>
in several features: body size larger (SBL =
<quantity id="456F3AA43064FF8DFCF1FA37BE3908A6" box="[772,858,1445,1469]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="29" pageNumber="230" unit="mm" value="6.8">6.8 mm</quantity>
in
<taxonomicName id="4597ECC23064FF8DFC70FA37B97108A6" authorityName="Alluaud" authorityYear="1935" box="[901,1042,1445,1469]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFC70FA37B97108A6" box="[901,1042,1445,1469]" italics="true" pageId="29" pageNumber="230">A. olsoufieffi</emphasis>
</taxonomicName>
); pronotum impunctate (coarsely punctate laterally and basally in
<taxonomicName id="4597ECC23064FF8DFD40FA54BE5E08C6" authorityName="Alluaud" authorityYear="1935" box="[693,829,1477,1501]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFD40FA54BE5E08C6" box="[693,829,1477,1501]" italics="true" pageId="29" pageNumber="230">A. olsoufieffi</emphasis>
</taxonomicName>
), slightly less than twice as long as wide (2.5 times as long as wide in
<taxonomicName id="4597ECC23064FF8DFDE7FA74BFFF08E6" authorityName="Alluaud" authorityYear="1935" box="[530,668,1509,1533]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFDE7FA74BFFF08E6" box="[530,668,1509,1533]" italics="true" pageId="29" pageNumber="230">A. olsoufieffi</emphasis>
</taxonomicName>
), with lateral borders vaguely present but only as very faintly impressed lines (lateral borders absent from
<taxonomicName id="4597ECC23064FF8DFCFAF994BEF90B06" authorityName="Alluaud" authorityYear="1935" box="[783,922,1541,1565]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFCFAF994BEF90B06" box="[783,922,1541,1565]" italics="true" pageId="29" pageNumber="230">A. olsoufieffi</emphasis>
</taxonomicName>
members); elytra with only one pair of small pale spots, located at apical one-third on interval 4 (both subapical and subbasal pairs of spots present in
<taxonomicName id="4597ECC23064FF8DFE36F9D4BF2F0B46" authorityName="Alluaud" authorityYear="1935" box="[451,588,1605,1629]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFE36F9D4BF2F0B46" box="[451,588,1605,1629]" italics="true" pageId="29" pageNumber="230">A. olsoufieffi</emphasis>
</taxonomicName>
) and with three or four discal setiferous pores also on interval 5 (absent from
<taxonomicName id="4597ECC23064FF8DFE8EF9F4BF660B66" authorityName="Alluaud" authorityYear="1935" box="[379,517,1637,1661]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFE8EF9F4BF660B66" box="[379,517,1637,1661]" italics="true" pageId="29" pageNumber="230">A. olsoufieffi</emphasis>
</taxonomicName>
members); and legs dark, except pale at base of femora and on trochanters (legs pale thoughout in
<taxonomicName id="4597ECC23064FF8DFDECF914BFC10B86" authorityName="Alluaud" authorityYear="1935" box="[537,674,1669,1693]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B0E34B533064FF8DFDECF914BFC10B86" box="[537,674,1669,1693]" italics="true" pageId="29" pageNumber="230">A. olsoufieffi</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="822897413064FF8EFF5EF934B9EA08EA" blockId="29.[123,1174,612,1726]" lastBlockId="30.[123,1174,1328,1624]" lastPageId="30" lastPageNumber="231" pageId="29" pageNumber="230">
In addition, the
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of
<taxonomicName id="4597ECC23064FF8DFE04F934BFC10BA6" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[497,674,1701,1725]" class="Insecta" family="Carabidae" genus="Archicolliuris" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533064FF8DFE04F934BFC10BA6" box="[497,674,1701,1725]" italics="true" pageId="29" pageNumber="230">A. ranomafanae</emphasis>
</taxonomicName>
is unique among members of all Malagasy odcanthine species in having the elytra distinctly convex throughout, except in the area of the transverse depression, elytral intervals flat, and elytral striae 3 to 7 effaced except in the transverse depression and apically, where they are evident. Striae 1 and 2 are very shallowly impressed but evident (stria 2 less so) in and posterior to the transverse depression. The form and location of the transverse depression of the elytra and the nearly effaced striae are similar to these features of members and genus
<taxonomicName id="4597ECC23067FF8EFEA2FA48BFED08E9" authority="Liebke (1933)" authorityName="Liebke" authorityYear="1933" box="[343,654,1498,1522]" class="Insecta" family="Carabidae" genus="Mimocolliuris" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533067FF8EFEA2FA48BC8D08E9" box="[343,494,1498,1522]" italics="true" pageId="30" pageNumber="231">Mimocolliuris</emphasis>
<bibRefCitation id="E606EAB03067FF8EFE03FA48BFED08E9" author="LIEBKE, M." box="[502,654,1498,1522]" pageId="30" pageNumber="231" pagination="202 - 210" refId="ref31621" refString="LIEBKE, M. 1933. Neue Colliurinen. Stylops 2: 202 - 210." type="journal article" year="1933">Liebke (1933)</bibRefCitation>
</taxonomicName>
of the eastern Palearctic and Oriental Regions.
</paragraph>
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16. Digital images of holotype female of
<taxonomicName id="4597ECC23067FF8EFDCAFB4CBE8B09EA" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[575,1000,1246,1265]" class="Insecta" family="Carabidae" genus="Archicolliuris" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533067FF8EFDCAFB4CBE4F09EA" box="[575,812,1246,1265]" italics="true" pageId="30" pageNumber="231">Archicolliuris ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
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A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm.
</paragraph>
</caption>
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<emphasis id="B0E34B533067FF8EFF5EFA69BC920B09" bold="true" box="[171,497,1531,1555]" pageId="30" pageNumber="231">
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</emphasis>
.— At present, known only from the
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locality.
</paragraph>
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<emphasis id="B0E34B533067FF8EFF5EF98CBCC70B2F" bold="true" box="[171,420,1566,1590]" pageId="30" pageNumber="231">
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</emphasis>
.— The unique
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was collected in a flight intercept-yellow pan trap on an island in a stream in montane rainforest at an elevation of
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.
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="C96D684B6F692AC696012085ABA1BBAA">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:namePart id="C183EC55BF1FDBB95A9ED0B5D53F0AA5">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="5F8554CFADEB216FD7B04AE352A4B502">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:namePart id="2BF6A90E99BFCC648AA2ACC0A9D51E95">Rainio, Johanna</mods:namePart>
<mods:affiliation id="692FF6065A960E47AFAF33DCCC150EC8">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
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<taxonomicName id="4597ECC23066FF8FFF8EFE48BFDB0CE9" ID-CoL="56TVV" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,696,474,498]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="bimaculatoides" status="sp. nov.">
<emphasis id="B0E34B533066FF8FFF8EFE48BCC40CE9" bold="true" box="[123,423,474,498]" italics="true" pageId="31" pageNumber="232">Thysanotus bimaculatoides</emphasis>
Kavanaugh and Rainio
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,
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<smallCapsWord id="84CE019D3066FF8FFF07FDA3BC3C0F5F" baselines="575" box="[242,351,561,580]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="31" pageNumber="232">MATERIAL</smallCapsWord>
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.—
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(
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), a male, in CAS, labeled: “
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Ranomafana NP. Mangevo plot
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<date id="F629B1813066FF8FFDB5FDDCBFC90F7E" box="[576,682,589,614]" pageId="31" pageNumber="232" value="2005-04-20">20.4.2005</date>
”/ “240” [handwritten label]/ “
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17. A-D. Digital images of holotype male of
<taxonomicName id="4597ECC23066FF8FFD78F916B9260B8C" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[653,1093,1668,1688]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="bimaculatoides" status="sp. nov.">
<emphasis id="B0E34B533066FF8FFD78F916BE190B8C" box="[653,890,1668,1687]" italics="true" pageId="31" pageNumber="232">Thysanotus bimaculatoides</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283066FF8FFBA5F917B9F50B83" box="[1104,1174,1669,1688]" pageId="31" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="822897413066FF8FFF89F90CBFC30BA9" blockId="31.[124,1174,1668,1714]" box="[124,672,1694,1714]" pageId="31" pageNumber="232">A–D. A Habitus, Dorsal aspect; B. Labels. Scale line = 1.0 mm.</paragraph>
</caption>
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<paragraph id="822897413059FFB0FF6EFD31BCDE0FF2" blockId="32.[123,1174,675,745]" pageId="32" pageNumber="233">
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18. Digital images of aedeagus of male genitalia. A, C. Left lateral aspect; B, D. Dorsal aspect. A–B. Holotype male of
<taxonomicName id="4597ECC23059FFB0FF37FD2EBF0A0FD4" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[194,617,700,719]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatoides" status="sp. nov.">
<emphasis id="B0E34B533059FFB0FF37FD2EBCCF0FD4" box="[194,428,700,719]" italics="true" pageId="32" pageNumber="233">Thysanotus bimaculatoides</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283059FFB0FD9AFD2FBFD20FCB" box="[623,689,701,720]" pageId="32" pageNumber="233" rank="species">sp. nov.</taxonomicNameLabel>
; B, D.
<taxonomicName id="4597ECC23059FFB0FD07FD2EBE8E0FD4" authority="Alluaud" authorityName="Alluaud" authorityYear="1936" box="[754,1005,700,719]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B0E34B533059FFB0FD07FD2EBEC10FD4" box="[754,930,700,719]" italics="true" pageId="32" pageNumber="233">Thysanotus spinosus</emphasis>
Alluaud
</taxonomicName>
(from Ranomafana National Park). Scale lines = 0.5 mm.
</paragraph>
</caption>
<paragraph id="822897413059FFB0FF8EFC90BE550EC1" blockId="32.[123,1174,770,1725]" pageId="32" pageNumber="233">
<taxonomicName id="4597ECC23059FFB0FF8EFC90BFF30E01" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,656,770,794]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatoides" status="sp. nov.">
<emphasis id="B0E34B533059FFB0FF8EFC90BCC30E01" box="[123,416,770,794]" italics="true" pageId="32" pageNumber="233">Thysanotus bimaculatoides</emphasis>
<materialsCitation id="32FF9D1C3059FFB0FE5CFC90BFF30E01" box="[425,656,770,794]" collectingDate="2000-11" collectingDateMax="2005-04-22" collectingDateMin="2000-11" collectionCode="MNHN, CAS" collectorName="R. Harin'Hala &amp; M. E. Irwin" country="Madagascar" county="Mangevo" elevation="1600" latitude="-21.827501" location="Foret d'Ambalagoavy Nord" longLatPrecision="21" longitude="47.338886" municipality="Province" pageId="32" pageNumber="233" specimenCount="2" specimenCount-female="2" stateProvince="Fianarantsoa" typeStatus="paratype">Kavanaugh &amp; Rainio</materialsCitation>
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283059FFB0FD6CFC90BFB70E02" box="[665,724,770,793]" pageId="32" pageNumber="233" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
<typeStatus id="5D2C29E33059FFB0FC52FC90B96C0E01" box="[935,1039,770,794]" pageId="32" pageNumber="233" type="paratype">Paratypes</typeStatus>
(total of 2):
<specimenCount id="94915CC83059FFB0FF8EFCB0BD810E21" box="[123,226,802,826]" count="1" pageId="32" pageNumber="233" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843059FFB0FED7FCB0BC170E22" box="[290,372,802,825]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="32" pageNumber="233">MNHN</collectionCode>
) labeled “
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Ranomafana NP.
<collectingCounty id="6B49EFCD3059FFB0FC6CFCB0BE9C0E21" box="[921,1023,802,826]" pageId="32" pageNumber="233">Mangevo</collectingCounty>
plot
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<elevation id="09BA70723059FFB0FBBEFCB0B9F50E22" box="[1099,1174,802,826]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="32" pageNumber="233" unit="m" value="1600.0">1600m</elevation>
</quantity>
<date id="F629B1813059FFB0FF8EFCD0BD860E41" box="[123,229,834,858]" pageId="32" pageNumber="233" value="2005-04-22">
<collectingDate id="E66D48693059FFB0FF8EFCD0BD860E41" box="[123,229,834,858]" pageId="32" pageNumber="233" value="2005-04-22">22.4.2005</collectingDate>
</date>
”/ “240” [handwritten label];
<specimenCount id="94915CC83059FFB0FDEAFCD0BF1F0E41" box="[543,636,834,858]" count="1" pageId="32" pageNumber="233" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843059FFB0FD52FCD0BFB80E41" box="[679,731,834,858]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="32" pageNumber="233" type="Museum">CAS</collectionCode>
) labeled “CASENT 8069538”/ “
<collectingCountry id="FA80D7D13059FFB0FBCEFCD0BD840E61" name="Madagascar" pageId="32" pageNumber="233">MADAGASCAR</collectingCountry>
:
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<collectingRegion id="405359A33059FFB0FE91FCF0BC8F0E61" box="[356,492,866,890]" country="Madagascar" name="Fianarantsoa" pageId="32" pageNumber="233">Fianarantsoa</collectingRegion>
,
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,
<location id="8748C19A3059FFB0FCB4FCF0BEE80E61" LSID="urn:lsid:plazi:treatment:0A3E26573066FFB1FF8EFE48BDA50C48:8748C19A3059FFB0FCB4FCF0BEE80E61" box="[833,907,866,890]" country="Madagascar" county="Mangevo" latitude="-21.827501" longLatPrecision="21" longitude="47.338886" municipality="Province" name="Ikonga" pageId="32" pageNumber="233" stateProvince="Fianarantsoa">Ikonga</location>
,
<location id="8748C19A3059FFB0FC69FCF0B9680E61" LSID="urn:lsid:plazi:treatment:0A3E26573066FFB1FF8EFE48BDA50C48:8748C19A3059FFB0FC69FCF0B9680E61" box="[924,1035,866,890]" country="Madagascar" county="Mangevo" latitude="-21.827501" longLatPrecision="21" longitude="47.338886" municipality="Province" name="Abatombe" pageId="32" pageNumber="233" stateProvince="Fianarantsoa">Abatombe</location>
<date id="F629B1813059FFB0FBE3FCF1B9E10E61" box="[1046,1154,866,890]" pageId="32" pageNumber="233" value="2000-11">
<collectingDate id="E66D48693059FFB0FBE3FCF1B9E10E61" box="[1046,1154,866,890]" pageId="32" pageNumber="233" value="2000-11">Nov-2000</collectingDate>
</date>
”/ “
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,
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<location id="8748C19A3059FFB0FE62FC10BFF50E81" LSID="urn:lsid:plazi:treatment:0A3E26573066FFB1FF8EFE48BDA50C48:8748C19A3059FFB0FE62FC10BFF50E81" box="[407,662,898,922]" country="Madagascar" county="Mangevo" latitude="-21.827501" longLatPrecision="21" longitude="47.338886" municipality="Province" name="Calif. Acad. of Sciences" pageId="32" pageNumber="233" stateProvince="Fianarantsoa">Calif. Acad. of Sciences</location>
colls:
<collectorName id="2F62F2973059FFB0FD29FC11BE110E81" box="[732,882,898,922]" pageId="32" pageNumber="233">R. Harin’Hala</collectorName>
&amp;
<collectorName id="2F62F2973059FFB0FC66FC11B96B0E81" box="[915,1032,898,922]" pageId="32" pageNumber="233">M.E. Irwin</collectorName>
,
<collectingMethod id="5BD6EF563059FFB0FBE1FC10B9F60E81" box="[1044,1173,898,922]" pageId="32" pageNumber="233">malaise trap</collectingMethod>
<quantity id="456F3AA43059FFB0FF8EFC30BDDB0EA1" box="[123,184,930,954]" metricMagnitude="2" metricUnit="m" metricValue="6.25" pageId="32" pageNumber="233" unit="m" value="625.0">
<elevation id="09BA70723059FFB0FF8EFC30BDDB0EA1" box="[123,184,930,954]" metricMagnitude="2" metricUnit="m" metricValue="6.25" pageId="32" pageNumber="233" unit="m" value="625.0">625m</elevation>
</quantity>
MA-01-12-01”. Both
<typeStatus id="5D2C29E33059FFB0FE42FC31BF7E0EA0" box="[439,541,931,955]" pageId="32" pageNumber="233" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="5D2C29E33059FFB0FC79FC31B9730EA0" box="[908,1040,931,955]" pageId="32" pageNumber="233" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC23059FFB0FBE9FC30BF6A0EC0" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatoides" status="sp. nov.">
<emphasis id="B0E34B533059FFB0FBE9FC30BC7D0EC0" italics="true" pageId="32" pageNumber="233">Thysanotus bimaculatoides</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283059FFB0FDE5FC51BF290EC1" box="[528,586,963,986]" pageId="32" pageNumber="233" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label].
</paragraph>
<paragraph id="822897413059FFB0FF5EFC71B9500EE1" blockId="32.[123,1174,770,1725]" box="[171,1075,995,1019]" pageId="32" pageNumber="233">
<materialsCitation id="32FF9D1C3059FFB0FF5EFC71B94F0EE0" box="[171,1068,995,1019]" country="Madagascar" location="Ranomafana National Park" pageId="32" pageNumber="233" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533059FFB0FF5EFC71BC340EE2" bold="true" box="[171,343,995,1019]" pageId="32" pageNumber="233">
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<typeStatus id="5D2C29E33059FFB0FF5EFC71BD870EE2" box="[171,228,995,1019]" pageId="32" pageNumber="233">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3059FFB0FF19FC74BC340EE2" baselines="1012" box="[236,343,998,1017]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="32" pageNumber="233">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13059FFB0FE8BFC71BC9C0EE0" box="[382,511,995,1019]" name="Madagascar" pageId="32" pageNumber="233">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33059FFB0FDF8FC71BF9F0EE0" box="[525,764,995,1019]" country="Madagascar" name="Fianarantsoa" pageId="32" pageNumber="233">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3059FFB0FCFFFC71B94F0EE0" LSID="urn:lsid:plazi:treatment:0A3E26573066FFB1FF8EFE48BDA50C48:8748C19A3059FFB0FCFFFC71B94F0EE0" box="[778,1068,995,1019]" country="Madagascar" name="Ranomafana National Park" pageId="32" pageNumber="233" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413059FFB0FF5EFB91BF5D0960" blockId="32.[123,1174,770,1725]" pageId="32" pageNumber="233">
<emphasis id="B0E34B533059FFB0FF5EFB91BC8C0902" bold="true" box="[171,495,1027,1051]" pageId="32" pageNumber="233">
<smallCapsWord id="84CE019D3059FFB0FF5EFB91BC4D0902" baselines="1045,1044" box="[171,302,1027,1051]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="32" pageNumber="233">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3059FFB0FEC0FB94BC320902" baselines="1044" box="[309,337,1030,1049]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="32" pageNumber="233">OF</smallCapsWord>
<smallCapsWord id="84CE019D3059FFB0FEA2FB94BCC90902" baselines="1044" box="[343,426,1030,1049]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="32" pageNumber="233">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3059FFB0FE47FB94BC8C0902" baselines="1044" box="[434,495,1030,1049]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="32" pageNumber="233">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23059FFB0FD06FB91BEF50900" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[755,918,1027,1051]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatoides">
<emphasis id="B0E34B533059FFB0FD06FB91BEF50900" box="[755,918,1027,1051]" italics="true" pageId="32" pageNumber="233">bimaculatoides</emphasis>
</taxonomicName>
, is an adjective derived from the Latin words,
<emphasis id="B0E34B533059FFB0FE9CFBB1BCEB0920" box="[361,392,1059,1083]" italics="true" pageId="32" pageNumber="233">bis</emphasis>
, meaning two, and
<emphasis id="B0E34B533059FFB0FDAFFBB1BFCA0920" box="[602,681,1059,1083]" italics="true" pageId="32" pageNumber="233">macula</emphasis>
, meaning spot or spotted, and the Greek word
<emphasis id="B0E34B533059FFB0FF8EFBD1BDD00940" box="[123,179,1091,1115]" italics="true" pageId="32" pageNumber="233">eidos</emphasis>
, meaning like. The name refers to the similarity between members of this species and those of
<taxonomicName id="4597ECC23059FFB0FF6DFBF1BF540960" authority="Deuve (2010)" authorityName="Deuve" authorityYear="2010" box="[152,567,1123,1147]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B0E34B533059FFB0FF6DFBF1BCF80960" box="[152,411,1123,1147]" italics="true" pageId="32" pageNumber="233">Thysanotus bimaculatus</emphasis>
<bibRefCitation id="E606EAB03059FFB0FE56FBF6BF540960" author="DEUVE, T." box="[419,567,1123,1147]" pageId="32" pageNumber="233" pagination="165 - 176" refId="ref30046" refString="DEUVE, T. 2010. Note sur le genre Thysanotus Chaudoir et quelques autres Lebiinae Pericalini de Madagascar [Coleoptera, Caraboidea, Harpalidae]. Revue Francaise d'Entomologie (N. S.) 32: 165 - 176." type="journal article" year="2010">Deuve (2010)</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="822897413059FFB0FF5EFB11BEC10B66" blockId="32.[123,1174,770,1725]" pageId="32" pageNumber="233">
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<emphasis id="B0E34B533059FFB0FF5EFB11BC210981" bold="true" box="[171,322,1155,1179]" pageId="32" pageNumber="233">RECOGNITION</emphasis>
</smallCapsWord>
.— Size slightly less than average for genus, SBL of male =
<quantity id="456F3AA43059FFB0FC32FB11B97B0980" box="[967,1048,1155,1179]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="32" pageNumber="233" unit="mm" value="5.7">5.7 mm</quantity>
, of females =
<quantity id="456F3AA43059FFB0FF65FB31BD8709A0" box="[144,228,1187,1212]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="32" pageNumber="233" unit="mm" value="5.3">5.3 mm</quantity>
. Members of this species (
<figureCitation id="1AAC8BC43059FFB0FDE4FB31BF1109A7" box="[529,626,1187,1212]" captionStart="FIGURE" captionStartId="31.[157,169,1669,1688]" captionTargetBox="[121,1177,624,1665]" captionTargetId="figure-579@31.[121,1177,624,1665]" captionTargetPageId="31" captionText="FIGURE 17. A-D. Digital images of holotype male of Thysanotus bimaculatoides Kavanaugh &amp; Rainio sp. nov. A–D. A Habitus, Dorsal aspect; B. Labels. Scale line = 1.0 mm." pageId="32" pageNumber="233">Fig. 17A</figureCitation>
) key to
<taxonomicName id="4597ECC23059FFB0FD26FB31B91409A7" authority="Deuve (2010)" authorityName="Deuve" authorityYear="2010" box="[723,1143,1187,1212]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B0E34B533059FFB0FD26FB31BEBB09A0" box="[723,984,1187,1211]" italics="true" pageId="32" pageNumber="233">Thysanotus bimaculatus</emphasis>
<bibRefCitation id="E606EAB03059FFB0FC14FB36B91409A7" author="DEUVE, T." box="[993,1143,1188,1212]" pageId="32" pageNumber="233" pagination="165 - 176" refId="ref30046" refString="DEUVE, T. 2010. Note sur le genre Thysanotus Chaudoir et quelques autres Lebiinae Pericalini de Madagascar [Coleoptera, Caraboidea, Harpalidae]. Revue Francaise d'Entomologie (N. S.) 32: 165 - 176." type="journal article" year="2010">Deuve (2010)</bibRefCitation>
</taxonomicName>
in Deuve’s key based on the following combination of features: elytra shiny, intervals smooth, elytral microsculpture indistinct (comprised of faintly impressed microlines and markedly transverse sculpticells, each elytron with a large, pale macula near the base on intervals 2 to 4, also extended partially onto interval 1 and to scutellum at basomedial part of macula, elytral apex bluntly round- ed, not toothed. Members of
<taxonomicName id="4597ECC23059FFB0FE40FAD7BF100847" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[437,627,1348,1372]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatoides">
<emphasis id="B0E34B533059FFB0FE40FAD7BF100847" box="[437,627,1348,1372]" italics="true" pageId="32" pageNumber="233">T. bimaculatoides</emphasis>
</taxonomicName>
differ from the
<typeStatus id="5D2C29E33059FFB0FCD4FAD6BE1D0847" box="[801,894,1348,1372]" pageId="32" pageNumber="233" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4597ECC23059FFB0FC51FAD7B9220847" authorityName="Deuve" authorityYear="2010" box="[932,1089,1348,1372]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B0E34B533059FFB0FC51FAD7B9220847" box="[932,1089,1348,1372]" italics="true" pageId="32" pageNumber="233">T. bimaculatus</emphasis>
</taxonomicName>
in having slightly larger body size (SBL of
<typeStatus id="5D2C29E33059FFB0FDD1FAF6BFE20867" box="[548,641,1380,1404]" pageId="32" pageNumber="233" type="holotype">holotype</typeStatus>
female of
<taxonomicName id="4597ECC23059FFB0FCF5FAF7BEC20867" authorityName="Deuve" authorityYear="2010" box="[768,929,1380,1404]" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B0E34B533059FFB0FCF5FAF7BEC20867" box="[768,929,1380,1404]" italics="true" pageId="32" pageNumber="233">T. bimaculatus</emphasis>
</taxonomicName>
=
<quantity id="456F3AA43059FFB0FC4BFAF6B9700867" box="[958,1043,1380,1404]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="32" pageNumber="233" unit="mm" value="4.9">4.9 mm</quantity>
), pronotum slightly narrower, ratio of PW/PL = 1.1 (ratio of PW/PL =
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), narrower basally with subbasal sinuation of the lateral margin slightly longer and deeper and hind angles sharp- er, rectangular to distinctly acute (compare with
<bibRefCitation id="E606EAB03059FFB0FD76FA57BE6608C6" author="DEUVE, T." box="[643,773,1477,1501]" pageId="32" pageNumber="233" pagination="165 - 176" refId="ref30046" refString="DEUVE, T. 2010. Note sur le genre Thysanotus Chaudoir et quelques autres Lebiinae Pericalini de Madagascar [Coleoptera, Caraboidea, Harpalidae]. Revue Francaise d'Entomologie (N. S.) 32: 165 - 176." type="journal article" year="2010">Deuve 2010</bibRefCitation>
,
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). Deuve did not have a male specimen of
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<emphasis id="B0E34B533059FFB0FEF0FA77BCC108E6" box="[261,418,1509,1533]" italics="true" pageId="32" pageNumber="233">T. bimaculatus</emphasis>
</taxonomicName>
, so we cannot compare male genitalia of these two species. However, for future comparative purposes, we provide digital photographs of the genitalia (
<figureCitation id="1AAC8BC43059FFB0FC19F997B90C0B06" box="[1004,1135,1541,1565]" captionStart="FIGURE" captionStartId="32.[155,167,675,694]" captionTargetBox="[140,1163,222,658]" captionTargetId="figure-2339@32.[120,1176,216,672]" captionTargetPageId="32" captionText="FIGURE 18. Digital images of aedeagus of male genitalia. A, C. Left lateral aspect; B, D. Dorsal aspect. A–B. Holotype male of Thysanotus bimaculatoides Kavanaugh &amp; Rainio sp. nov.; B, D. Thysanotus spinosus Alluaud (from Ranomafana National Park). Scale lines = 0.5 mm." pageId="32" pageNumber="233">Figs. 18A,C</figureCitation>
) of the
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of
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<emphasis id="B0E34B533059FFB0FEDBF9B4BC8C0B26" box="[302,495,1573,1597]" italics="true" pageId="32" pageNumber="233">T. bimaculatoides</emphasis>
</taxonomicName>
as well as those of a male of
<taxonomicName id="4597ECC23059FFB0FCB9F9B7BDA60B46" authority="(Alluaud 1936 c)" baseAuthorityName="Alluaud" baseAuthorityYear="1936" class="Insecta" family="Carabidae" genus="Thysanotus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B0E34B533059FFB0FCB9F9B7B94F0B26" box="[844,1068,1573,1597]" italics="true" pageId="32" pageNumber="233">Thysanotus spinosus</emphasis>
(
<bibRefCitation id="E606EAB03059FFB0FBCAF9B7BDDF0B46" author="ALLUAUD, C." pageId="32" pageNumber="233" pagination="1 - 10" refId="ref29048" refString="ALLUAUD, C. 1936 c. Carabiques nouveaux de Madagascar. Afra, Cahiers d'Entomologie 12: 1 - 10." type="journal article" year="1936">Alluaud 1936c</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="1AAC8BC43059FFB0FF20F9D7BC340B46" box="[213,343,1605,1629]" captionStart="FIGURE" captionStartId="32.[155,167,675,694]" captionTargetBox="[140,1163,222,658]" captionTargetId="figure-2339@32.[120,1176,216,672]" captionTargetPageId="32" captionText="FIGURE 18. Digital images of aedeagus of male genitalia. A, C. Left lateral aspect; B, D. Dorsal aspect. A–B. Holotype male of Thysanotus bimaculatoides Kavanaugh &amp; Rainio sp. nov.; B, D. Thysanotus spinosus Alluaud (from Ranomafana National Park). Scale lines = 0.5 mm." pageId="32" pageNumber="233">Figs. 18B,D</figureCitation>
) [males of this species previously unknown], to complement the illustrations of other
<taxonomicName id="4597ECC23059FFB0FF23F9F7BCC70B66" box="[214,420,1637,1661]" pageId="32" pageNumber="233">
<emphasis id="B0E34B533059FFB0FF23F9F7BC2C0B66" box="[214,335,1637,1661]" italics="true" pageId="32" pageNumber="233">Thysanotus</emphasis>
species
</taxonomicName>
provided by Jeannel (1949) and
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.
</paragraph>
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</emphasis>
.— At present, known only from two localities,
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apart, in central
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. Both localities are on the eastern slope off the central plateau.
</paragraph>
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</emphasis>
.—
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and
<specimenCount id="94915CC83058FFB1FD51FF49BE500DE8" box="[676,819,219,243]" count="1" pageId="33" pageNumber="234" type="generic" typeStatus="paratype">one paratype</specimenCount>
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<materialsCitation id="32FF9D1C3058FFB1FCCAFF49BDA50C48" elevation="1600" location="Mangevo area" municipality="The" pageId="33" pageNumber="234" specimenCount="650" typeStatus="paratype">
were collected by hand in the
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, about
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distance in from the edge of montane rainforest, at an elevation of
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.
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other
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was collected in a
<collectingMethod id="5BD6EF563058FFB1FD99FE89BF8F0C28" box="[620,748,283,307]" pageId="33" pageNumber="234">malaise trap</collectingMethod>
in montane rainforest at an elevation of
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<elevation id="09BA70723058FFB1FF8EFEAEBDDC0C48" box="[123,191,316,340]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="33" pageNumber="234" unit="m" value="650.0">650 m</elevation>
</quantity>
.
</specimenCount>
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</paragraph>
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</treatment>
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Kavanaugh and Rainio
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1
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; Rainio 2009: 32, Rainio 2012: 74 (informal designation).
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.—
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<typeStatus id="5D2C29E33069FF80FE70FC89BC8B0E28" box="[389,488,795,819]" pageId="16" pageNumber="217" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43069FF80FE08FC89BFE80E28" box="[509,651,795,819]" captionStart="FIGURE" captionStartId="17.[155,167,1571,1590]" captionTargetBox="[139,1176,235,1551]" captionTargetId="figure-68@17.[120,1176,216,1569]" captionTargetPageId="17" captionText="FIGURE 10. Digital images of holotype male of Trichinillus (Mallopelmus) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." pageId="16" pageNumber="217">Figs. 10A–B</figureCitation>
), a male, in CAS, labeled: “
<collectingCountry id="FA80D7D13069FF80FC2DFC89B9F50E28" box="[984,1174,795,819]" name="Madagascar" pageId="16" pageNumber="217">MADAGASCAR</collectingCountry>
Ranomafana NP.
<location id="8748C19A3069FF80FEC6FCA9BC1B0E48" LSID="urn:lsid:plazi:treatment:0A3E26573069FF82FF8EFD07BCD40E48:8748C19A3069FF80FEC6FCA9BC1B0E48" box="[307,376,827,851]" country="Madagascar" name="Buffer" pageId="16" pageNumber="217">Buffer</location>
zone, degraded forest
<date id="F629B1813069FF80FD92FCAEBFBC0E4F" box="[615,735,828,852]" pageId="16" pageNumber="217" value="2002-10-14">
<collectingDate id="E66D48693069FF80FD92FCAEBFBC0E4F" box="[615,735,828,852]" pageId="16" pageNumber="217" value="2002-10-14">14.10.2002</collectingDate>
</date>
”/ “collected by hand in daytime,
<collectorName id="2F62F2973069FF80FBB7FCAEBDC20E6F" pageId="16" pageNumber="217">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973069FF80FF36FCCEBC420E6F" box="[195,289,860,884]" pageId="16" pageNumber="217">J. Rainio</collectorName>
collectors”/ “
<typeStatus id="5D2C29E33069FF80FE40FCCEBF210E6F" box="[437,578,860,884]" pageId="16" pageNumber="217" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23069FF80FDBDFCCEBDA10E8F" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov." subGenus="Mallopelmus">
<emphasis id="B0E34B533069FF80FDBDFCCEBFA00E6F" box="[584,707,860,884]" italics="true" pageId="16" pageNumber="217">Trichinillus</emphasis>
(
<emphasis id="B0E34B533069FF80FD27FCCEBE3E0E6F" box="[722,861,860,884]" italics="true" pageId="16" pageNumber="217">Mallopelmus</emphasis>
)
<emphasis id="B0E34B533069FF80FC99FCCEBE990E6F" box="[876,1018,860,884]" italics="true" pageId="16" pageNumber="217">ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283069FF80FF3FFCEFBC650E8F" box="[202,262,893,916]" pageId="16" pageNumber="217" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label]
</materialsCitation>
.
<materialsCitation id="32FF9D1C3069FF80FE22FCEFBE350EEE" collectingDate="2004-03-05" collectionCode="MNHN" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="600" location="Talatakely" pageId="16" pageNumber="217" specimenCount="600" typeStatus="paratype">
<typeStatus id="5D2C29E33069FF80FE22FCEFBF5C0E8E" box="[471,575,893,917]" pageId="16" pageNumber="217" type="paratype">Paratypes</typeStatus>
(only 1): a female (in
<collectionCode id="E4860F843069FF80FCCFFCEFBEE80E8F" box="[826,907,893,916]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="217">MNHN</collectionCode>
) labelled “
<collectingCountry id="FA80D7D13069FF80FBF3FCEEBDD10EAE" name="Madagascar" pageId="16" pageNumber="217">MADAGASCAR</collectingCountry>
Ranomafana NP.
<location id="8748C19A3069FF80FE84FC0EBCBD0EAF" LSID="urn:lsid:plazi:treatment:0A3E26573069FF82FF8EFD07BCD40E48:8748C19A3069FF80FE84FC0EBCBD0EAF" box="[369,478,924,948]" country="Madagascar" name="Talatakely" pageId="16" pageNumber="217">Talatakely</location>
, trail B,
<specimenCount id="94915CC83069FF80FDCEFC0EBFE60EAF" box="[571,645,924,948]" count="600" pageId="16" pageNumber="217" type="generic">
<quantity id="456F3AA43069FF80FDCEFC0EBF1D0EAF" box="[571,638,924,948]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="217" unit="m" value="600.0">
<elevation id="09BA70723069FF80FDCEFC0EBF1D0EAF" box="[571,638,924,948]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="217" unit="m" value="600.0">600 m</elevation>
</quantity>
.
</specimenCount>
<date id="F629B1813069FF80FD7EFC0EBE600EAE" box="[651,771,924,949]" pageId="16" pageNumber="217" value="2004-03-05">
<collectingDate id="E66D48693069FF80FD7EFC0EBE600EAE" box="[651,771,924,949]" pageId="16" pageNumber="217" value="2004-03-05">05.03.2004</collectingDate>
</date>
”/ “collected by hand in daytime from dead leaves in bushes,
<collectorName id="2F62F2973069FF80FE81FC2FBC850ECE" box="[372,486,957,981]" pageId="16" pageNumber="217">F. Ratalata</collectorName>
and
<collectorName id="2F62F2973069FF80FDE8FC2FBF1E0ECE" box="[541,637,957,981]" pageId="16" pageNumber="217">J. Rainio</collectorName>
collectors”/ “
<typeStatus id="5D2C29E33069FF80FCE1FC2FBEFB0ECE" box="[788,920,957,981]" pageId="16" pageNumber="217" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC23069FF80FC55FC2FBF4C0EEE" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov." subGenus="Mallopelmus">
<emphasis id="B0E34B533069FF80FC55FC2FB9780ECE" box="[928,1051,957,981]" italics="true" pageId="16" pageNumber="217">Trichinillus</emphasis>
(
<emphasis id="B0E34B533069FF80FBD9FC2FBDC50EEE" italics="true" pageId="16" pageNumber="217">Mallopelmus</emphasis>
)
<emphasis id="B0E34B533069FF80FF43FC4FBC270EEE" box="[182,324,989,1013]" italics="true" pageId="16" pageNumber="217">ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283069FF80FDC2FC4CBF120EEE" box="[567,625,990,1013]" pageId="16" pageNumber="217" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="822897413069FF80FF5EFC6CB950090E" blockId="16.[123,1174,795,1726]" box="[171,1075,1022,1046]" pageId="16" pageNumber="217">
<materialsCitation id="32FF9D1C3069FF80FF5EFC6CB94F090D" box="[171,1068,1022,1046]" country="Madagascar" location="Ranomafana National Park" pageId="16" pageNumber="217" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533069FF80FF5EFC6CBC34090F" bold="true" box="[171,343,1022,1046]" pageId="16" pageNumber="217">
<smallCapsWord id="84CE019D3069FF80FF5EFC6CBD87090F" baselines="1040,1039" box="[171,228,1022,1046]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="16" pageNumber="217">
<typeStatus id="5D2C29E33069FF80FF5EFC6CBD87090F" box="[171,228,1022,1046]" pageId="16" pageNumber="217">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3069FF80FF19FB93BC34090F" baselines="1039" box="[236,343,1025,1044]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="16" pageNumber="217">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13069FF80FE8BFC6CBC9C090D" box="[382,511,1022,1046]" name="Madagascar" pageId="16" pageNumber="217">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33069FF80FDF8FC6CBF9F090D" box="[525,764,1022,1046]" country="Madagascar" name="Fianarantsoa" pageId="16" pageNumber="217">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3069FF80FCFFFC6CB94F090D" LSID="urn:lsid:plazi:treatment:0A3E26573069FF82FF8EFD07BCD40E48:8748C19A3069FF80FCFFFC6CB94F090D" box="[778,1068,1022,1046]" country="Madagascar" name="Ranomafana National Park" pageId="16" pageNumber="217" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413069FF80FF5EFB8CBECA094D" blockId="16.[123,1174,795,1726]" pageId="16" pageNumber="217">
<emphasis id="B0E34B533069FF80FF5EFB8CBC91092F" bold="true" box="[171,498,1054,1078]" pageId="16" pageNumber="217">
<smallCapsWord id="84CE019D3069FF80FF5EFB8CBC4D092F" baselines="1072,1071" box="[171,302,1054,1078]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="16" pageNumber="217">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3069FF80FEC3FBB3BC31092F" baselines="1071" box="[310,338,1057,1076]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="16" pageNumber="217">OF</smallCapsWord>
<smallCapsWord id="84CE019D3069FF80FEACFBB3BCCF092F" baselines="1071" box="[345,428,1057,1076]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="16" pageNumber="217">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3069FF80FE40FBB3BC91092F" baselines="1071" box="[437,498,1057,1076]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="16" pageNumber="217">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23069FF80FD0CFB8CBEE4092D" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[761,903,1054,1078]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533069FF80FD0CFB8CBEE4092D" box="[761,903,1054,1078]" italics="true" pageId="16" pageNumber="217">ranomafanae</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the national park in which the
<typeStatus id="5D2C29E33069FF80FD29FBADBE69094C" box="[732,778,1087,1111]" pageId="16" pageNumber="217">type</typeStatus>
was collected.
</paragraph>
<paragraph id="822897413069FF82FF5EFBCDBCB30F68" blockId="16.[123,1174,795,1726]" lastBlockId="18.[123,1174,219,852]" lastPageId="18" lastPageNumber="219" pageId="16" pageNumber="217">
<smallCapsWord id="84CE019D3069FF80FF5EFBCDBC21096E" baselines="1137,1136" box="[171,322,1119,1143]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="16" pageNumber="217">
<emphasis id="B0E34B533069FF80FF5EFBCDBC21096E" bold="true" box="[171,322,1119,1143]" pageId="16" pageNumber="217">RECOGNITION</emphasis>
</smallCapsWord>
.— Slightly smaller than average for subgenus, SBL male =
<quantity id="456F3AA43069FF80FC21FBCDB944096D" box="[980,1063,1119,1143]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="16" pageNumber="217" unit="mm" value="7.2">7.2 mm</quantity>
, female =
<quantity id="456F3AA43069FF80FF8EFB12BDAE098C" box="[123,205,1151,1175]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="16" pageNumber="217" unit="mm" value="7.5">7.5 mm</quantity>
. The following combination of features identify members of this species as coelostomine pterostichines of the genus
<taxonomicName id="4597ECC23069FF80FE47FB0DBF4E09AC" authorityName="Straneo" authorityYear="1938" box="[434,557,1183,1207]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533069FF80FE47FB0DBF4E09AC" box="[434,557,1183,1207]" italics="true" pageId="16" pageNumber="217">Trichinillus</emphasis>
</taxonomicName>
, subgenus
<taxonomicName id="4597ECC23069FF80FD5AFB0DBE5909AC" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[687,826,1183,1207]" class="Insecta" family="Carabidae" genus="Mallopelmus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533069FF80FD5AFB0DBE5909AC" box="[687,826,1183,1207]" italics="true" pageId="16" pageNumber="217">Mallopelmus</emphasis>
</taxonomicName>
: antennae with antennomere 3 attached symmetrically or nearly symmetrically to antennomere 2; mentum with deep emargination, apex of median mental tooth distinctly posterior to apices of epilobes; elytron with epipleuron interrupted by internal plica subapically; parascutellar stria short but evident; striae smooth, impunctate; basal setiferious puncture situated at base of stria 2 + parascutellar stria; tarsomeres 1 to 4 of all legs with dense pads of setae ventrally; front and middle tarsomeres 1 to 3 with more or less faintly-defined longitudinal groove medially and depressed areas paralaterally on dorsal surface; male front tarsomeres 1 to 4 wide, dilated, not latero- or medioapically toothed; aedeagus of male with right face dorsal in repose, right paramere conchoid, left paramere reduced, short, apically digitiform. Of the three previously described species represented in Jeannel’s (1948) key to
<taxonomicName id="4597ECC23069FF80FF8EFA71BC3D08E0" box="[123,350,1507,1531]" pageId="16" pageNumber="217">
<emphasis id="B0E34B533069FF80FF8EFA71BC6508E0" box="[123,262,1507,1531]" italics="true" pageId="16" pageNumber="217">Mallopelmus</emphasis>
species
</taxonomicName>
, members of
<taxonomicName id="4597ECC23069FF80FE0AFA76BFCF08E0" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[511,684,1507,1531]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533069FF80FE0AFA76BFCF08E0" box="[511,684,1507,1531]" italics="true" pageId="16" pageNumber="217">T. ranomafanae</emphasis>
</taxonomicName>
(
<figureCitation id="1AAC8BC43069FF80FD34FA71BE4708E0" box="[705,804,1507,1531]" captionStart="FIGURE" captionStartId="17.[155,167,1571,1590]" captionTargetBox="[139,1176,235,1551]" captionTargetId="figure-68@17.[120,1176,216,1569]" captionTargetPageId="17" captionText="FIGURE 10. Digital images of holotype male of Trichinillus (Mallopelmus) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." pageId="16" pageNumber="217">Fig. 10A</figureCitation>
) are most similar to
<taxonomicName id="4597ECC23069FF80FBEFFA71BCC50B07" authority="(Alluaud 1936)" baseAuthorityName="Alluaud" baseAuthorityYear="1936" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="abacetoides">
<emphasis id="B0E34B533069FF80FBEFFA71BD990B07" italics="true" pageId="16" pageNumber="217">Trichinillus abacetoides</emphasis>
(Alluaud 1936)
</taxonomicName>
members based on the following shared features: head with temporal area long, nearly as long as the diameter of the eye, and joined to the neck region at a very obtuse (about 165°) angle; eyes moderately convex and projected; pronotum with basal area distinctly and abruptly depressed relative to disc; and elytral base without margination medial to base of stria 6. Members of the other two species,
<taxonomicName id="4597ECC23069FF80FDF6F917BE820B86" authority="(Alluaud 1936)" baseAuthorityName="Alluaud" baseAuthorityYear="1936" box="[515,993,1669,1693]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="dactyleuryoides">
<emphasis id="B0E34B533069FF80FDF6F917BE530B86" box="[515,816,1669,1693]" italics="true" pageId="16" pageNumber="217">Trichinillus dactyleuryoides</emphasis>
(Alluaud 1936)
</taxonomicName>
and
<taxonomicName id="4597ECC23069FF80FBEFF917BC110BA5" authority="(Jeannel 1948)" baseAuthorityName="Jeannel" baseAuthorityYear="1948" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="B0E34B533069FF80FBEFF917BDAD0BA6" italics="true" pageId="16" pageNumber="217">Trichinillus perrieri</emphasis>
(Jeannel 1948)
</taxonomicName>
, differ in having the head with the temporal area shorter, about one-third as long as the diameter of the eye, and joined to the neck region at a moderately obtuse (about 135°) angle, eyes more convex and projected, pronotum not or less depressed, especially medially, and margination of the elytra base more varied, extended medially to the base of stria
<quantity id="456F3AA4306BFF82FC00FE89B97C0C28" box="[1013,1055,283,307]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="18" pageNumber="219" unit="in" value="3.0">3 in</quantity>
most individuals). The apex of the median lobe of the male genitalia is also differently shaped in each of these species (compare with Figs. 191c-d and 191f-g, respectively, in Jeannel 1948). Members of
<taxonomicName id="4597ECC2306BFF82FF8EFEEEBC470C88" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,292,379,403]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53306BFF82FF8EFEEEBC470C88" box="[123,292,379,403]" italics="true" pageId="18" pageNumber="219">T. ranomafanae</emphasis>
</taxonomicName>
differ from those of
<taxonomicName id="4597ECC2306BFF82FDFDFEEEBFC10C88" baseAuthorityName="Alluaud" baseAuthorityYear="1936" box="[520,674,379,403]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="abacetoides">
<emphasis id="B0E34B53306BFF82FDFDFEEEBFC10C88" box="[520,674,379,403]" italics="true" pageId="18" pageNumber="219">T. abacetoides</emphasis>
</taxonomicName>
in having: slightly larger size (SBL =
<quantity id="456F3AA4306BFF82FBB6FEE9B9F60C88" box="[1091,1173,379,403]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="18" pageNumber="219" unit="mm" value="6.4">6.4 mm</quantity>
in
<taxonomicName id="4597ECC2306BFF82FF6FFE0EBC540CA8" baseAuthorityName="Alluaud" baseAuthorityYear="1936" box="[154,311,411,435]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="abacetoides">
<emphasis id="B0E34B53306BFF82FF6FFE0EBC540CA8" box="[154,311,411,435]" italics="true" pageId="18" pageNumber="219">T. abacetoides</emphasis>
</taxonomicName>
); pronotum with shallow but distinct subbasal sinuation of the lateral margin (absent from
<taxonomicName id="4597ECC2306BFF82FEFAFE2EBCC80CC8" baseAuthorityName="Alluaud" baseAuthorityYear="1936" box="[271,427,443,467]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="abacetoides">
<emphasis id="B0E34B53306BFF82FEFAFE2EBCC80CC8" box="[271,427,443,467]" italics="true" pageId="18" pageNumber="219">T. abacetoides</emphasis>
</taxonomicName>
members); elytra relatively shorter and wider, especially across the base, and humeri broadly rounded and not at all sloped (humeri more slope in
<taxonomicName id="4597ECC2306BFF82FC32FE4EB9030CE8" baseAuthorityName="Alluaud" baseAuthorityYear="1936" box="[967,1120,475,499]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="abacetoides">
<emphasis id="B0E34B53306BFF82FC32FE4EB9030CE8" box="[967,1120,475,499]" italics="true" pageId="18" pageNumber="219">T. abacetoides</emphasis>
</taxonomicName>
, and median lobe of male genitalia with apex straighter, slightly bent ventrally toward apex in lateral aspect (
<figureCitation id="1AAC8BC4306BFF82FF3AFD89BC4C0F28" box="[207,303,539,563]" captionStart="FIGURE" captionStartId="17.[155,167,1571,1590]" captionTargetBox="[139,1176,235,1551]" captionTargetId="figure-68@17.[120,1176,216,1569]" captionTargetPageId="17" captionText="FIGURE 10. Digital images of holotype male of Trichinillus (Mallopelmus) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." pageId="18" pageNumber="219">Fig. 10C</figureCitation>
) and slightly deflect right in dorsal view (
<figureCitation id="1AAC8BC4306BFF82FCF6FD89BE070F28" box="[771,868,539,563]" captionStart="FIGURE" captionStartId="17.[155,167,1571,1590]" captionTargetBox="[139,1176,235,1551]" captionTargetId="figure-68@17.[120,1176,216,1569]" captionTargetPageId="17" captionText="FIGURE 10. Digital images of holotype male of Trichinillus (Mallopelmus) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." pageId="18" pageNumber="219">Fig. 10D</figureCitation>
) (not recurved dorsally and then ventrally in lateral aspect and straight in dorsal aspect as in
<taxonomicName id="4597ECC2306BFF82FCB3FDAEBE810F48" baseAuthorityName="Alluaud" baseAuthorityYear="1936" box="[838,994,571,595]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="abacetoides">
<emphasis id="B0E34B53306BFF82FCB3FDAEBE810F48" box="[838,994,571,595]" italics="true" pageId="18" pageNumber="219">T. abacetoides</emphasis>
</taxonomicName>
males; compare with Fig. 191e in Jeannel 1948)
</paragraph>
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10. Digital images of holotype male of
<taxonomicName id="4597ECC23068FF81FDC7F9B1B92E0B2D" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[562,1101,1571,1590]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov." subGenus="Mallopelmus">
<emphasis id="B0E34B533068FF81FDC7F9B1BFF70B2D" box="[562,660,1571,1590]" italics="true" pageId="17" pageNumber="218">Trichinillus</emphasis>
(
<emphasis id="B0E34B533068FF81FD57F9B1BE720B2D" box="[674,785,1571,1590]" italics="true" pageId="17" pageNumber="218">Mallopelmus</emphasis>
)
<emphasis id="B0E34B533068FF81FCEBF9B1BEF30B2D" box="[798,912,1571,1590]" italics="true" pageId="17" pageNumber="218">ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283068FF81FBA6F9B1B9F60B2D" box="[1107,1173,1571,1590]" pageId="17" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm.
</paragraph>
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</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E3306BFF82FC8AFDEEBECE0F8F" box="[895,941,636,660]" pageId="18" pageNumber="219">type</typeStatus>
locality.
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<emphasis id="B0E34B53306BFF82FF5EFD09BCC50FA9" bold="true" box="[171,422,667,691]" pageId="18" pageNumber="219">
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<smallCapsWord id="84CE019D306BFF82FEE7FD0DBCC50FA9" baselines="685" box="[274,422,671,690]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="18" pageNumber="219">DISTRIBUTION</smallCapsWord>
</emphasis>
.— The male
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of
<taxonomicName id="4597ECC2306BFF82FD31FD0EBE0C0FA8" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[708,879,667,691]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53306BFF82FD31FD0EBE0C0FA8" box="[708,879,667,691]" italics="true" pageId="18" pageNumber="219">T. ranomafanae</emphasis>
</taxonomicName>
was collected in degraded secondary montane rainforest at an elevation of
<quantity id="456F3AA4306BFF82FD65FD29BFB50FC8" box="[656,726,699,723]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="219" unit="m" value="800.0">800 m</quantity>
, near a village in the peripheral zone of Ranomafana National Park about1000 meters from the main gate. The female
<typeStatus id="5D2C29E3306BFF82FC20FD4EB9520FEF" box="[981,1073,732,756]" pageId="18" pageNumber="219" type="paratype">paratype</typeStatus>
was collected in secondary montane rainforest in the Talatakely area, at an elevation of
<quantity id="456F3AA4306BFF82FC2DFD69B97E0E08" box="[984,1053,763,787]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="18" pageNumber="219" unit="m" value="900.0">900 m</quantity>
, at the
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mark along Trail B. Vegetation in this area was dominated by young trees (DBH &lt;
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) and guava (
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<emphasis id="B0E34B53306BFF82FF3EFCA9BCC40E48" box="[203,423,827,851]" italics="true" pageId="18" pageNumber="219">Psidium cattleianum</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

328
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<mods:title id="1E2574D71CD431AE4604294784E99067">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:affiliation id="371189D0FB323FF4C2CAE0C4EFAC8321">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:namePart id="2BAC9A240E4B495A1314BDE797B29E05">Rainio, Johanna</mods:namePart>
<mods:affiliation id="97DE00576D9257A451BD850894C6F9D4">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
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<emphasis id="B0E34B53306BFF82FF8EFBE3BC010992" bold="true" box="[123,354,1137,1161]" italics="true" pageId="18" pageNumber="219">Chlaenius kathrynae</emphasis>
Kavanaugh and Rainio
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,
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.—
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), a male, in CAS, labeled: “CASENT 1002782”/ “
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,
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,”/ “
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/
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, Stop# DHK-01-002,
<collectorName id="2F62F297306BFF82FB97FA94BC3D0826" pageId="18" pageNumber="219">D.H. &amp; K.M. Kavanaugh</collectorName>
,
<collectorName id="2F62F297306BFF82FE86FAB4BC850826" box="[371,486,1318,1341]" pageId="18" pageNumber="219">R.L. Brett</collectorName>
,
<collectorName id="2F62F297306BFF82FE09FAB4BF060826" box="[508,613,1317,1341]" pageId="18" pageNumber="219">E. Elsom</collectorName>
,
<collectorName id="2F62F297306BFF82FD8EFAB4BF870825" box="[635,740,1318,1342]" pageId="18" pageNumber="219">F. Vargas</collectorName>
, Ranaivosolo,”/ “
<collectorName id="2F62F297306BFF82FC46FAB4B9EC0826" box="[947,1167,1317,1341]" pageId="18" pageNumber="219">E.F. Randrianifirina</collectorName>
,
<collectorName id="2F62F297306BFF82FF8EFAD4BC2A0846" box="[123,329,1350,1373]" pageId="18" pageNumber="219">N. Rasoamananana</collectorName>
,
<collectorName id="2F62F297306BFF82FEA3FAD4BF420846" box="[342,545,1349,1373]" pageId="18" pageNumber="219">T.J. Ravelomanana</collectorName>
, and
<collectorName id="2F62F297306BFF82FDA9FAD4BE640846" box="[604,775,1349,1373]" pageId="18" pageNumber="219">H.C. Raveloson</collectorName>
collectors”/ “collected in daytime by treading marsh vegetation and shore”/ “
<typeStatus id="5D2C29E3306BFF82FDDAFAF4BFDF0865" box="[559,700,1382,1406]" pageId="18" pageNumber="219" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC2306BFF82FD30FAF7B9F60866" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[709,1173,1381,1406]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="kathrynae" status="sp. nov.">
<emphasis id="B0E34B53306BFF82FD30FAF7BEC60866" box="[709,933,1381,1405]" italics="true" pageId="18" pageNumber="219">Chlaenius kathrynae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628306BFF82FF8EFA14BDD60886" box="[123,181,1414,1437]" pageId="18" pageNumber="219" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label]
</materialsCitation>
.
<materialsCitation id="32FF9D1C306BFF82FE71FA14B9530886" box="[388,1072,1413,1438]" collectionCode="CAS, MNHN, MZF" country="Madagascar" pageId="18" pageNumber="219" specimenCount="4" specimenCount-male="4" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="5D2C29E3306BFF82FE71FA14BC8F0885" box="[388,492,1414,1438]" pageId="18" pageNumber="219" type="paratype">Paratypes</typeStatus>
(total of 8):
<specimenCount id="94915CC8306BFF82FD82FA17BFA90886" box="[631,714,1413,1437]" count="4" pageId="18" pageNumber="219" type="male">4 males</specimenCount>
(in
<collectionCode id="E4860F84306BFF82FD0DFA17BE4F0886" box="[760,812,1413,1437]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="219" type="Museum">CAS</collectionCode>
,
<collectionCode id="E4860F84306BFF82FCCFFA14BEE80886" box="[826,907,1414,1437]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="18" pageNumber="219">MNHN</collectionCode>
and
<collectionCode id="E4860F84306BFF82FC37FA14BE9A0886" box="[962,1017,1414,1437]" pageId="18" pageNumber="219">MZF</collectionCode>
) and
</materialsCitation>
<materialsCitation id="32FF9D1C306BFF82FBCDFA17BEF008C6" collectionCode="CAS" country="Madagascar" pageId="18" pageNumber="219" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="94915CC8306BFF82FBCDFA17B9F50886" box="[1080,1174,1413,1437]" count="1" pageId="18" pageNumber="219" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F84306BFF82FF6AFA37BDB008A6" box="[159,211,1445,1469]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="219" type="Museum">CAS</collectionCode>
) labeled same as
<typeStatus id="5D2C29E3306BFF82FE64FA37BC8D08A6" box="[401,494,1445,1469]" pageId="18" pageNumber="219" type="holotype">holotype</typeStatus>
except “CASENT 1002783”, “CASENT 1002784”, “CASENT 1002786”, “CASENT 1002787”, and “CASENT 1002785”, respectively
</materialsCitation>
;
<materialsCitation id="32FF9D1C306BFF82FC51FA57BCD40B45" collectingDate="1998-04-23" collectionCode="CAS" collectorName="D. H. Kavanaugh" country="Madagascar" elevation="1050" latitude="-21.226667" location="Vohiparara area" longLatPrecision="126" longitude="47.383335" municipality="Ranomafana National Park" pageId="18" pageNumber="219" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="94915CC8306BFF82FC51FA57BE8C08C6" box="[932,1007,1477,1501]" count="1" pageId="18" pageNumber="219" type="male">1 male</specimenCount>
(
<collectionCode id="E4860F84306BFF82FBF7FA57B95508C6" box="[1026,1078,1477,1501]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="219" type="Museum">CAS</collectionCode>
) labeled “CASENT 1049008”/ “
<collectingCountry id="FA80D7D1306BFF82FE70FA77BF2008E6" box="[389,579,1509,1533]" name="Madagascar" pageId="18" pageNumber="219">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A3306BFF82FDA2FA77BE2F08E6" box="[599,844,1509,1533]" country="Madagascar" name="Fianarantsoa" pageId="18" pageNumber="219">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B306BFF82FC95FA77B9EC08E6" box="[864,1167,1509,1533]" pageId="18" pageNumber="219">Ranomafana National Park</collectingMunicipality>
,
<location id="8748C19A306BFF82FF8EF997BC410B06" LSID="urn:lsid:plazi:treatment:0A3E2657306BFF84FF8EFBE3BF8A0985:8748C19A306BFF82FF8EF997BC410B06" box="[123,290,1541,1565]" country="Madagascar" latitude="-21.226667" longLatPrecision="126" longitude="47.383335" municipality="Ranomafana National Park" name="Vohiparara area" pageId="18" pageNumber="219" stateProvince="Fianarantsoa">Vohiparara area</location>
,
<quantity id="456F3AA4306BFF82FEC5F997BCE20B06" box="[304,385,1541,1565]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="18" pageNumber="219" unit="m" value="1050.0">
<elevation id="09BA7072306BFF82FEC5F997BCE20B06" box="[304,385,1541,1565]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="18" pageNumber="219" unit="m" value="1050.0">1050 m</elevation>
</quantity>
, mixed tropical forest,
<date id="F629B181306BFF82FD8EF994BE6C0B05" box="[635,783,1541,1566]" pageId="18" pageNumber="219" value="1998-04-23">
<collectingDate id="E66D4869306BFF82FD8EF994BE6C0B05" box="[635,783,1541,1566]" pageId="18" pageNumber="219" value="1998-04-23">23 April 1998</collectingDate>
</date>
,”/ “
<geoCoordinate id="E7A3F186306BFF82FCC9F994BECB0B05" box="[828,936,1541,1566]" degrees="21" direction="south" minutes="13. 6" orientation="latitude" pageId="18" pageNumber="219" precision="92" value="-21.226667">21°13.6’S</geoCoordinate>
/
<geoCoordinate id="E7A3F186306BFF82FC43F994B9400B06" box="[950,1059,1542,1566]" degrees="47" direction="east" minutes="23. 0" orientation="longitude" pageId="18" pageNumber="219" precision="92" value="47.383335">47°23.0’E</geoCoordinate>
,”/ “Stop # 98-71,
<collectorName id="2F62F297306BFF82FF32F9B4BC190B26" box="[199,378,1573,1597]" pageId="18" pageNumber="219">D.H. Kavanaugh</collectorName>
collector collected in soil cracks, under dirt clods, and under root clods in abandoned rice paddy area”
</materialsCitation>
;
<materialsCitation id="32FF9D1C306BFF82FE3EF9D4BF670BA6" collectingDate="1994-11-25" collectionCode="MTEC" collectorName="M. A. Ivie &amp; D. A. Pollack" country="Madagascar" latitude="-21.256666" location="Fianaran. Pr. Ranomafana" longLatPrecision="21" longitude="47.420834" pageId="18" pageNumber="219" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="94915CC8306BFF82FE3EF9D4BF5B0B46" box="[459,568,1605,1630]" count="2" pageId="18" pageNumber="219" type="female">2 females</specimenCount>
(in
<collectionCode id="E4860F84306BFF82FD85F9D4BFD90B45" box="[624,698,1606,1630]" country="USA" httpUri="http://grbio.org/cool/zm6z-4gpy" name="Montana State Entomology Collection" pageId="18" pageNumber="219" type="Museum">MTEC</collectionCode>
) labeled “
<collectingCountry id="FA80D7D1306BFF82FCC0F9D4BE900B45" box="[821,1011,1606,1630]" name="Madagascar" pageId="18" pageNumber="219">MADAGASCAR</collectingCountry>
:
<location id="8748C19A306BFF82FBFDF9D7BC610B66" LSID="urn:lsid:plazi:treatment:0A3E2657306BFF84FF8EFBE3BF8A0985:8748C19A306BFF82FBFDF9D7BC610B66" country="Madagascar" latitude="-21.256666" longLatPrecision="21" longitude="47.420834" name="Fianaran. Pr. Ranomafana" pageId="18" pageNumber="219">Fianaran. Pr. Ranomafana</location>
N.P. HQ. area
<geoCoordinate id="E7A3F186306BFF82FE50F9F4BF400B65" box="[421,547,1637,1662]" degrees="21" direction="south" minutes="15" orientation="latitude" pageId="18" pageNumber="219" precision="15" seconds="24" value="-21.256666">21°15’24”S</geoCoordinate>
<geoCoordinate id="E7A3F186306BFF82FDDFF9F4BFC90B66" box="[554,682,1637,1662]" degrees="47" direction="east" minutes="25" orientation="longitude" pageId="18" pageNumber="219" precision="15" seconds="15" value="47.420834">47°25’15”E</geoCoordinate>
<date id="F629B181306BFF82FD44F9F7BE580B65" box="[689,827,1637,1662]" pageId="18" pageNumber="219" value="1994-11-25">
<collectingDate id="E66D4869306BFF82FD44F9F7BE580B65" box="[689,827,1637,1662]" pageId="18" pageNumber="219" value="1994-11-25">25NOV1994</collectingDate>
</date>
, at night
<collectorName id="2F62F297306BFF82FC54F9F4B9690B66" box="[929,1034,1637,1662]" pageId="18" pageNumber="219">M.A. Ivie</collectorName>
&amp;
<collectorName id="2F62F297306BFF82FBDBF9F4BDC40B86" pageId="18" pageNumber="219">D.A. Pollack</collectorName>
”. All
<typeStatus id="5D2C29E3306BFF82FF1FF914BC330B85" box="[234,336,1670,1694]" pageId="18" pageNumber="219" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="5D2C29E3306BFF82FD5CF914BE4E0B85" box="[681,813,1670,1694]" pageId="18" pageNumber="219" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC2306BFF82FCC0F917BDBD0BA6" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="kathrynae" status="sp. nov.">
<emphasis id="B0E34B53306BFF82FCC0F917B9770B86" box="[821,1044,1669,1693]" italics="true" pageId="18" pageNumber="219">Chlaenius kathrynae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628306BFF82FF10F934BC7C0BA6" box="[229,287,1702,1725]" pageId="18" pageNumber="219" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<caption id="D6E8C7C9306AFF83FF6EF9C7B9450B99" pageId="19" pageNumber="220" startId="19.[155,167,1621,1640]" targetBox="[152,1175,216,1604]" targetPageId="19" targetType="figure">
<paragraph id="82289741306AFF83FF6EF9C7B9450B99" blockId="19.[123,1173,1620,1666]" pageId="19" pageNumber="220">
<smallCapsWord id="84CE019D306AFF83FF6EF9C7BDBB0B7D" baselines="1635,1634" box="[155,216,1621,1640]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="19" pageNumber="220">FIGURE</smallCapsWord>
11. Digital images of holotype male of
<taxonomicName id="4597ECC2306AFF83FDCFF9C6BECE0B7C" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[570,941,1620,1640]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="kathrynae" status="sp. nov.">
<emphasis id="B0E34B53306AFF83FDCFF9C6BF8E0B7C" box="[570,749,1620,1639]" italics="true" pageId="19" pageNumber="220">Chlaenius kathrynae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628306AFF83FC41F9C7BE940B73" box="[948,1015,1621,1640]" pageId="19" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
A Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm.
</paragraph>
</caption>
<paragraph id="82289741306DFF84FF5EFF49B9500DE9" blockId="20.[123,1174,219,1182]" box="[171,1075,219,243]" pageId="20" pageNumber="221">
<materialsCitation id="32FF9D1C306DFF84FF5EFF49B94F0DE8" box="[171,1068,219,243]" country="Madagascar" location="Ranomafana National Park" pageId="20" pageNumber="221" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B53306DFF84FF5EFF49BC340DEA" bold="true" box="[171,343,219,243]" pageId="20" pageNumber="221">
<smallCapsWord id="84CE019D306DFF84FF5EFF49BD870DE9" baselines="237,237" box="[171,228,219,243]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="20" pageNumber="221">
<typeStatus id="5D2C29E3306DFF84FF5EFF49BD870DE9" box="[171,228,219,243]" pageId="20" pageNumber="221">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D306DFF84FF19FF4CBC340DEA" baselines="236" box="[236,343,222,241]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="20" pageNumber="221">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D1306DFF84FE8BFF49BC9C0DE8" box="[382,511,219,243]" name="Madagascar" pageId="20" pageNumber="221">Madagascar</collectingCountry>
,
<collectingRegion id="405359A3306DFF84FDF8FF49BF9F0DE8" box="[525,764,219,243]" country="Madagascar" name="Fianarantsoa" pageId="20" pageNumber="221">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A306DFF84FCFFFF49B94F0DE8" LSID="urn:lsid:plazi:treatment:0A3E2657306BFF84FF8EFBE3BF8A0985:8748C19A306DFF84FCFFFF49B94F0DE8" box="[778,1068,219,243]" country="Madagascar" name="Ranomafana National Park" pageId="20" pageNumber="221" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="82289741306DFF84FF5EFF6EBCCB0C4F" blockId="20.[123,1174,219,1182]" pageId="20" pageNumber="221">
<emphasis id="B0E34B53306DFF84FF5EFF6EBC890C09" bold="true" box="[171,490,252,276]" pageId="20" pageNumber="221">
<smallCapsWord id="84CE019D306DFF84FF5EFF6EBC4D0C09" baselines="270,269" box="[171,302,252,276]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="20" pageNumber="221">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D306DFF84FEC6FF6DBC2C0C09" baselines="269" box="[307,335,255,274]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="20" pageNumber="221">OF</smallCapsWord>
<smallCapsWord id="84CE019D306DFF84FEA1FF6DBCC40C09" baselines="269" box="[340,423,255,274]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="20" pageNumber="221">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D306DFF84FE58FF6DBC890C09" baselines="269" box="[429,490,255,274]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="20" pageNumber="221">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC2306DFF84FD1CFF69BE370C08" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[745,852,251,275]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="kathrynae">
<emphasis id="B0E34B53306DFF84FD1CFF69BE370C08" box="[745,852,251,275]" italics="true" pageId="20" pageNumber="221">kathrynae</emphasis>
</taxonomicName>
, is a noun in the genitive case, derived from the first given name of Kathryn May Kavanaugh, daughter of DHK and one of the collectors of the
<typeStatus id="5D2C29E3306DFF84FEDAFEAFBC3E0C4E" box="[303,349,317,341]" pageId="20" pageNumber="221">type</typeStatus>
series.
</paragraph>
<paragraph id="82289741306DFF84FF5EFECFBF930EE7" blockId="20.[123,1174,219,1182]" pageId="20" pageNumber="221">
<smallCapsWord id="84CE019D306DFF84FF5EFECFBC210C68" baselines="367,366" box="[171,322,349,373]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="20" pageNumber="221">
<emphasis id="B0E34B53306DFF84FF5EFECFBC210C68" bold="true" box="[171,322,349,373]" pageId="20" pageNumber="221">RECOGNITION</emphasis>
</smallCapsWord>
.— Size slightly below average for genus, SBL of males = 10.2–10.
<quantity id="456F3AA4306DFF84FBDBFECFB90D0C6F" box="[1070,1134,349,373]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="20" pageNumber="221" unit="mm" value="8.0">8 mm</quantity>
, of females 9.8–11.
<quantity id="456F3AA4306DFF84FED2FEEFBC0B0C8F" box="[295,360,381,405]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="221" unit="mm" value="1.0">1 mm</quantity>
. Members of this species (
<figureCitation id="1AAC8BC4306DFF84FD6FFEEFBF980C8E" box="[666,763,381,405]" captionStart="FIGURE" captionStartId="19.[155,167,1621,1640]" captionTargetBox="[152,1175,216,1604]" captionTargetId="figure-67@19.[120,1176,216,1618]" captionTargetPageId="19" captionText="FIGURE 11. Digital images of holotype male of Chlaenius kathrynae Kavanaugh &amp; Rainio sp. nov. A Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm." pageId="20" pageNumber="221">Fig. 11A</figureCitation>
) share the following features, which place them in genus
<taxonomicName id="4597ECC2306DFF84FEAEFE0FBCAE0CAE" authorityName="Motschulsky" authorityYear="1860" box="[347,461,413,437]" class="Insecta" family="Carabidae" genus="Chlaenites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306DFF84FEAEFE0FBCAE0CAE" box="[347,461,413,437]" italics="true" pageId="20" pageNumber="221">Chlaenites</emphasis>
</taxonomicName>
sense Jeannel (1948): antennomere 3 longer than antennomere 4; maxillary palpi setose, penultimate labial palpomeres 4 or 5 setose; pronotum with basolateral setae inserted distinctly anterior to the hind angles; elytra with pubescence restricted to near the striae on disk (
<emphasis id="B0E34B53306DFF84FF43FE6DBDB40F0D" box="[182,215,511,534]" italics="true" pageId="20" pageNumber="221">i.e.</emphasis>
, not present on the centers of intervals), except more generally and densely present on lateral intervals and near apices; and tarsi asetose dorsally, tarsomeres 5 with two rows of stout setae ventrally. In Jeannel’s (1949) key to subgenera of
<taxonomicName id="4597ECC2306DFF84FD56FDADBE760F4C" authorityName="Motschulsky" authorityYear="1860" box="[675,789,575,599]" class="Insecta" family="Carabidae" genus="Chlaenites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306DFF84FD56FDADBE760F4C" box="[675,789,575,599]" italics="true" pageId="20" pageNumber="221">Chlaenites</emphasis>
</taxonomicName>
, members of this species key best, although not fully, to subgenus
<taxonomicName id="4597ECC2306DFF84FE20FDF2BE4F0F63" authority="Kuntzen (1919)" authorityName="Kuntzen" authorityYear="1919" box="[469,812,608,632]" class="Insecta" family="Carabidae" genus="Chlaeniostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306DFF84FE20FDF2BF1A0F63" box="[469,633,608,632]" italics="true" pageId="20" pageNumber="221">Chlaeniostenus</emphasis>
<bibRefCitation id="E606EAB0306DFF84FD77FDF2BE4F0F63" author="KUNTZEN, H." box="[642,812,608,632]" pageId="20" pageNumber="221" pagination="1918 - 19" refId="ref31152" refString="KUNTZEN, H. 1919. Die Carabinenfaune Deutsch-Sudwestafrikas. Mitteilungen aus dem Zoologischen Museum in Berlin 9 [1918 - 19]: 91 - 156." type="journal article" year="1919">Kuntzen (1919)</bibRefCitation>
</taxonomicName>
, which is now considered a subgenus of
<taxonomicName id="4597ECC2306DFF84FF2BFD12BC880F83" authority="(Lorenz 2005)" baseAuthorityName="Lorenz" baseAuthorityYear="2005" box="[222,491,640,664]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306DFF84FF2BFD12BC290F83" box="[222,330,640,664]" italics="true" pageId="20" pageNumber="221">Chlaenius</emphasis>
(
<bibRefCitation id="E606EAB0306DFF84FEAFFD12BC810F83" author="LORENZ, W." box="[346,482,640,664]" pageId="20" pageNumber="221" refId="ref31666" refString="LORENZ, W. 2005. Systematic List of Extant Ground Beetles of the World (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. W. Lorenz, Tutzing. 530 pp." type="book" year="2005">Lorenz 2005</bibRefCitation>
)
</taxonomicName>
. Shared with other Malagasy members of this subgenus are the following features: pronotum relatively narrow, more or less cordiform, narrowed basally, with the anterior angles not projected anterior of the basal margin and the anterolateral areas of the pronotum distinctly curved ventrally; and the setose punctures along the lateral margins of the elytral intervals not or only faintly evident, most clearly evident in the apical half only. Jeannel included three other species in this subgenus; namely,
<taxonomicName id="4597ECC2306DFF84FD92FCB0BEB40E21" authority="Klug (1833)" authorityName="Klug" authorityYear="1833" box="[615,983,802,826]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="attenuatus">
<emphasis id="B0E34B53306DFF84FD92FCB0BE290E21" box="[615,842,802,826]" italics="true" pageId="20" pageNumber="221">Chlaenius attenuatus</emphasis>
<bibRefCitation id="E606EAB0306DFF84FCA7FCB0BEB40E21" author="KLUG, J. C. F." box="[850,983,802,826]" pageId="20" pageNumber="221" pagination="91 - 223" refId="ref30967" refString="KLUG, J. C. F. 1833. Bericht uber eine auf Madagascar veranstaltete. Sammlung von Insecten aus der Ordnung Coleoptera. Abhandlungen der Koniglichen Preussischen Akademie der Wissenschaften 1832 - 33: 91 - 223 + 5 pls." type="book chapter" year="1833">Klug (1833)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4597ECC2306DFF84FC13FCB0BC110E41" authority="Chaudoir (1876)" authorityName="Chaudoir" authorityYear="1876" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="subovatus">
<emphasis id="B0E34B53306DFF84FC13FCB0BDD00E41" italics="true" pageId="20" pageNumber="221">Chlaenius subovatus</emphasis>
<bibRefCitation id="E606EAB0306DFF84FF48FCD0BC110E41" author="CHAUDOIR, M." box="[189,370,834,858]" pageId="20" pageNumber="221" pagination="3 - 315" refId="ref29840" refString="CHAUDOIR, M. 1876. Monographie des chleniens. Annali del Museo Civico di Storia Naturelle di Genova 8: 3 - 315." type="journal article" year="1876">Chaudoir (1876)</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4597ECC2306DFF84FE41FCD0BE400E41" authority="Dejean (1831)" authorityName="Dejean" authorityYear="1831" box="[436,803,834,858]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="sellatus">
<emphasis id="B0E34B53306DFF84FE41FCD0BF180E41" box="[436,635,834,858]" italics="true" pageId="20" pageNumber="221">Chlaenius sellatus</emphasis>
<bibRefCitation id="E606EAB0306DFF84FD73FCD0BE400E41" author="DEJEAN, P. F. M. A." box="[646,803,834,858]" pageId="20" pageNumber="221" refId="ref29934" refString="DEJEAN, P. F. M. A. 1831. Species general des coleopteres, de la collection de M. le Comte Dejean. Tome cinquieme. Mequignon-Marvis, Paris, France. viii + 883 pp." type="book" year="1831">Dejean (1831)</bibRefCitation>
</taxonomicName>
. Members of
<taxonomicName id="4597ECC2306DFF84FC36FCD1B9320E41" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[963,1105,834,858]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="kathrynae">
<emphasis id="B0E34B53306DFF84FC36FCD1B9320E41" box="[963,1105,834,858]" italics="true" pageId="20" pageNumber="221">C. kathrynae</emphasis>
</taxonomicName>
differ from those of the other species in lacking any trace of the pale lateral and apical elytral border distinct in members of the other three species. They are also smaller than members of the other species (SBL in the latter ranges from 11.9 to 15.
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), have slightly broader and less cordiform pronota and males have genitalia (
<figureCitation id="1AAC8BC4306DFF84FE6EFC56BF470EC0" box="[411,548,964,988]" captionStart="FIGURE" captionStartId="19.[155,167,1621,1640]" captionTargetBox="[152,1175,216,1604]" captionTargetId="figure-67@19.[120,1176,216,1618]" captionTargetPageId="19" captionText="FIGURE 11. Digital images of holotype male of Chlaenius kathrynae Kavanaugh &amp; Rainio sp. nov. A Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm." pageId="20" pageNumber="221">Figs. 11C–D</figureCitation>
) that are very different in form from those of any of the other species (compare with Jeannel’s (1949) Figs. 388c-i)
</paragraph>
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<emphasis id="B0E34B53306DFF84FF5EFB96BC920900" bold="true" box="[171,497,1028,1052]" pageId="20" pageNumber="221">
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</emphasis>
.— At present, known only from the
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locality.
</paragraph>
<paragraph id="82289741306DFF84FF5EFBB7BF8A0985" blockId="20.[123,1174,219,1182]" pageId="20" pageNumber="221">
<emphasis id="B0E34B53306DFF84FF5EFBB7BCC60920" bold="true" box="[171,421,1061,1085]" pageId="20" pageNumber="221">
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<smallCapsWord id="84CE019D306DFF84FEE4FBBABCC60920" baselines="1078" box="[273,421,1064,1083]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="20" pageNumber="221">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Most specimens of the
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series were collected in open habitats at the edges of mixed tropical forest in the Vohiparara area at an elevation of
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. Several were collected by treading down vegetation at the edge of a marsh, and
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was found under loose dirt and root clods in an abandoned rice paddy area.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="E9F81C47D574CC9DE04EC3A807E3B6F2">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:namePart id="656F2FDC3F9FEEAA400AE96F589BC55D">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="FBC2747BB1731E92EBF2B023B14106F4">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:namePart id="04EC2C1CB2C48AA7B672FC2B7FA22CF6">Rainio, Johanna</mods:namePart>
<mods:affiliation id="732501045AED898392B5EE42B190A61A">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
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<emphasis id="B0E34B53306DFF84FF8EFB54BC2C09C5" bold="true" box="[123,335,1222,1246]" italics="true" pageId="20" pageNumber="221">Chlaenius robertae</emphasis>
Kavanaugh and Rainio
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,
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.—
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), a male, in CAS, labeled: “CASENT 1002781”/ “
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,
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,
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,
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,”/ “
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/
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, Stop# DHK-01-002,
<collectorName id="2F62F297306DFF84FB94FAF0BC3D0881" pageId="20" pageNumber="221">D.H. &amp; K.M. Kavanaugh</collectorName>
,
<collectorName id="2F62F297306DFF84FE86FA10BC850882" box="[371,486,1410,1433]" pageId="20" pageNumber="221">R.L. Brett</collectorName>
,
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,
<collectorName id="2F62F297306DFF84FD8EFA10BF870881" box="[635,740,1410,1434]" pageId="20" pageNumber="221">F. Vargas</collectorName>
, Ranaivosolo,”/ “
<collectorName id="2F62F297306DFF84FC46FA10B9EC0881" box="[947,1167,1410,1434]" pageId="20" pageNumber="221">E.F. Randrianifirina</collectorName>
,
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,
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, and
<collectorName id="2F62F297306DFF84FDA9FA31BE6408A1" box="[604,775,1442,1466]" pageId="20" pageNumber="221">H.C. Raveloson</collectorName>
collectors”/ “collected in daytime by treading marsh vegetation and shore”/ “
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<taxonomicName id="4597ECC2306DFF84FD4DFA51B91308C0" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[696,1136,1475,1499]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="robertae" status="sp. nov.">
<emphasis id="B0E34B53306DFF84FD4DFA51BEE508C0" box="[696,902,1475,1499]" italics="true" pageId="20" pageNumber="221">Chlaenius robertae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628306DFF84FB82FA51BDEC08E0" pageId="20" pageNumber="221" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label]
</materialsCitation>
.
<materialsCitation id="32FF9D1C306DFF84FEADFA76BFE60B67" collectingDate="1998-04-27" collectionCode="CAS" collectorName="D. H. Kavanaugh" country="Madagascar" elevation="1050" latitude="-21.226667" location="Vohiparara area" longLatPrecision="126" longitude="47.383335" municipality="Ranomafana National Park" pageId="20" pageNumber="221" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="5D2C29E3306DFF84FEADFA76BCA308E7" box="[344,448,1508,1532]" pageId="20" pageNumber="221" type="paratype">Paratypes</typeStatus>
(only 1): a female (in
<collectionCode id="E4860F84306DFF84FD5BFA71BF8108E0" box="[686,738,1507,1531]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="221" type="Museum">CAS</collectionCode>
) labeled “CASENT1049009”/ “
<collectingCountry id="FA80D7D1306DFF84FBCEFA71BD840B07" name="Madagascar" pageId="20" pageNumber="221">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A3306DFF84FF0DF991BC8A0B00" box="[248,489,1539,1563]" country="Madagascar" name="Fianarantsoa" pageId="20" pageNumber="221">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B306DFF84FE0FF991BE410B00" box="[506,802,1539,1563]" pageId="20" pageNumber="221">Ranomafana National Park</collectingMunicipality>
,
<location id="8748C19A306DFF84FCC7F991BEBF0B00" LSID="urn:lsid:plazi:treatment:0A3E2657306DFF86FF8EFB54BE4608B9:8748C19A306DFF84FCC7F991BEBF0B00" box="[818,988,1539,1563]" country="Madagascar" latitude="-21.226667" longLatPrecision="126" longitude="47.383335" municipality="Ranomafana National Park" name="Vohiparara area" pageId="20" pageNumber="221" stateProvince="Fianarantsoa">Vohiparara area</location>
,
<quantity id="456F3AA4306DFF84FC1BF991B9210B00" box="[1006,1090,1539,1563]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="20" pageNumber="221" unit="m" value="1050.0">
<elevation id="09BA7072306DFF84FC1BF991B9210B00" box="[1006,1090,1539,1563]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="20" pageNumber="221" unit="m" value="1050.0">1050 m</elevation>
</quantity>
, mixed tropical forest,
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<collectingDate id="E66D4869306DFF84FEEAF9B6BCD70B27" box="[287,436,1572,1596]" pageId="20" pageNumber="221" value="1998-04-27">27 April 1998</collectingDate>
</date>
,”/ “
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/
<geoCoordinate id="E7A3F186306DFF84FDA9F9B6BFAA0B20" box="[604,713,1572,1596]" degrees="47" direction="east" minutes="23. 0" orientation="longitude" pageId="20" pageNumber="221" precision="92" value="47.383335">47°23.0’E</geoCoordinate>
,”/ “Stop # 98-91,
<collectorName id="2F62F297306DFF84FC65F9B6B9260B27" box="[912,1093,1572,1596]" pageId="20" pageNumber="221">D.H. Kavanaugh</collectorName>
collector collected on bare soil at night in abandoned rice paddy area”/ “
<typeStatus id="5D2C29E3306DFF84FCC8F9D6BEA20B47" box="[829,961,1604,1628]" pageId="20" pageNumber="221" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC2306DFF84FC3DF9D6BC3C0B66" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="robertae" status="sp. nov.">
<emphasis id="B0E34B53306DFF84FC3DF9D6B9F60B47" box="[968,1173,1604,1628]" italics="true" pageId="20" pageNumber="221">Chlaenius robertae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628306DFF84FE93F9F7BCC30B67" box="[358,416,1637,1660]" pageId="20" pageNumber="221" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="82289741306DFF84FF5EF917B9500B86" blockId="20.[123,1174,1313,1726]" box="[171,1075,1669,1693]" pageId="20" pageNumber="221">
<materialsCitation id="32FF9D1C306DFF84FF5EF917B94F0B86" box="[171,1068,1669,1693]" country="Madagascar" location="Ranomafana National Park" pageId="20" pageNumber="221" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B53306DFF84FF5EF917BC340B80" bold="true" box="[171,343,1669,1693]" pageId="20" pageNumber="221">
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<typeStatus id="5D2C29E3306DFF84FF5EF917BD870B87" box="[171,228,1669,1693]" pageId="20" pageNumber="221">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D306DFF84FF19F91ABC340B80" baselines="1686" box="[236,343,1672,1691]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="20" pageNumber="221">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D1306DFF84FE8BF917BC9C0B86" box="[382,511,1669,1693]" name="Madagascar" pageId="20" pageNumber="221">Madagascar</collectingCountry>
,
<collectingRegion id="405359A3306DFF84FDF8F917BF9F0B86" box="[525,764,1669,1693]" country="Madagascar" name="Fianarantsoa" pageId="20" pageNumber="221">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A306DFF84FCFFF917B94F0B86" LSID="urn:lsid:plazi:treatment:0A3E2657306DFF86FF8EFB54BE4608B9:8748C19A306DFF84FCFFF917B94F0B86" box="[778,1068,1669,1693]" country="Madagascar" name="Ranomafana National Park" pageId="20" pageNumber="221" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="82289741306DFF84FF5EF934B9F50BA6" blockId="20.[123,1174,1313,1726]" box="[171,1174,1701,1726]" pageId="20" pageNumber="221">
<emphasis id="B0E34B53306DFF84FF5EF934BC8D0BA7" bold="true" box="[171,494,1702,1726]" pageId="20" pageNumber="221">
<smallCapsWord id="84CE019D306DFF84FF5EF934BC4D0BA7" baselines="1720,1719" box="[171,302,1702,1726]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="20" pageNumber="221">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D306DFF84FEC1F93BBC330BA7" baselines="1719" box="[308,336,1705,1724]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="20" pageNumber="221">OF</smallCapsWord>
<smallCapsWord id="84CE019D306DFF84FEA3F93BBCCA0BA7" baselines="1719" box="[342,425,1705,1724]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="20" pageNumber="221">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D306DFF84FE44F93BBC8D0BA7" baselines="1719" box="[433,494,1705,1724]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="20" pageNumber="221">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC2306DFF84FD05F937BE280BA6" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[752,843,1701,1725]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="robertae">
<emphasis id="B0E34B53306DFF84FD05F937BE280BA6" box="[752,843,1701,1725]" italics="true" pageId="20" pageNumber="221">robertae</emphasis>
</taxonomicName>
, is a noun in the genitive case,
</paragraph>
<caption id="D6E8C7C9306CFF85FF6EF9FBB9450B8E" pageId="21" pageNumber="222" startId="21.[155,167,1641,1660]" targetBox="[130,1176,216,1618]" targetPageId="21" targetType="figure">
<paragraph id="82289741306CFF85FF6EF9FBB9450B8E" blockId="21.[123,1173,1640,1685]" pageId="21" pageNumber="222">
<smallCapsWord id="84CE019D306CFF85FF6EF9FBBDBB0B61" baselines="1655,1654" box="[155,216,1641,1660]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="21" pageNumber="222">FIGURE</smallCapsWord>
12. Digital images of holotype male of
<taxonomicName id="4597ECC2306CFF85FDC8F9FABEC60B60" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[573,933,1640,1659]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="222" phylum="Arthropoda" rank="species" species="robertae" status="sp. nov.">
<emphasis id="B0E34B53306CFF85FDC8F9FABF870B60" box="[573,740,1640,1659]" italics="true" pageId="21" pageNumber="222">Chlaenius robertae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628306CFF85FC59F9FBBE8C0B67" box="[940,1007,1641,1660]" pageId="21" pageNumber="222" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm.
</paragraph>
</caption>
<paragraph id="82289741306FFF86FF8EFF49BFCB0C08" blockId="22.[123,1174,219,1443]" pageId="22" pageNumber="223">
derived from the given name of Roberta L. Brett, former graduate student and technical assistant of DHK and one of the collectors of the
<typeStatus id="5D2C29E3306FFF86FDDAFF6EBF3E0C0F" box="[559,605,252,276]" pageId="22" pageNumber="223">type</typeStatus>
series.
</paragraph>
<paragraph id="82289741306FFF86FF5EFE8EBE0F081B" blockId="22.[123,1174,219,1443]" pageId="22" pageNumber="223">
<smallCapsWord id="84CE019D306FFF86FF5EFE8EBC210C29" baselines="302,301" box="[171,322,284,308]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="22" pageNumber="223">
<emphasis id="B0E34B53306FFF86FF5EFE8EBC210C29" bold="true" box="[171,322,284,308]" pageId="22" pageNumber="223">RECOGNITION</emphasis>
</smallCapsWord>
.— Sized moderate for genus, SBL of male =
<quantity id="456F3AA4306FFF86FCCCFE8EBEFA0C28" box="[825,921,284,308]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="22" pageNumber="223" unit="mm" value="12.9">12.9 mm</quantity>
, of female =
<quantity id="456F3AA4306FFF86FBDBFE8EB9ED0C28" box="[1070,1166,284,308]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="22" pageNumber="223" unit="mm" value="13.3">13.3 mm</quantity>
. Members of this species (
<figureCitation id="1AAC8BC4306FFF86FE69FEAEBC9D0C4F" box="[412,510,316,340]" captionStart="FIGURE" captionStartId="21.[155,167,1641,1660]" captionTargetBox="[130,1176,216,1618]" captionTargetId="figure-67@21.[120,1176,216,1618]" captionTargetPageId="21" captionText="FIGURE 12. Digital images of holotype male of Chlaenius robertae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm." pageId="22" pageNumber="223">Fig. 12A</figureCitation>
) exhibit the following features, which place them in genus
<taxonomicName id="4597ECC2306FFF86FF8EFECFBC860C6E" authority="sensu Jeannel (1949)" authorityName="sensu Jeannel" authorityYear="1949" box="[123,485,349,373]" class="Insecta" family="Carabidae" genus="Chlaenites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306FFF86FF8EFECFBD8E0C6E" box="[123,237,349,373]" italics="true" pageId="22" pageNumber="223">Chlaenites</emphasis>
sensu Jeannel (1949)
</taxonomicName>
: antennomere 3 longer than antennomere 4; maxillary palpi setose; penultimate labial palpomeres 4 or 5 setose; pronotum with basolateral setae inserted distinctly anterior to the hind angles; elytra with pubescence restricted to near the striae on disk (i.e., not present on the centers of intervals), except more generally and densely present on lateral intervals and near apices; and tarsi asetose dorsally, tarsomeres 5 with two rows of stout setae ventrally. In Jeannel’s (1949) key to subgenera of
<taxonomicName id="4597ECC2306FFF86FDB1FE6DBFD50F0C" authorityName="sensu Jeannel" authorityYear="1949" box="[580,694,511,535]" class="Insecta" family="Carabidae" genus="Chlaenites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306FFF86FDB1FE6DBFD50F0C" box="[580,694,511,535]" italics="true" pageId="22" pageNumber="223">Chlaenites</emphasis>
</taxonomicName>
, members of this species key to his new subgenus
<taxonomicName id="4597ECC2306FFF86FF34FD8DBC320F2C" box="[193,337,543,567]" class="Insecta" family="Carabidae" genus="Chlaenitidius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306FFF86FF34FD8DBC320F2C" box="[193,337,543,567]" italics="true" pageId="22" pageNumber="223">Chlaenitidius</emphasis>
</taxonomicName>
, which is now considered a junior synonym of subgenus
<taxonomicName id="4597ECC2306FFF86FC34FD8DBC2D0F43" authority="Laferte-Senectere (1851)" authorityName="Laferte-Senectere" authorityYear="1851" class="Insecta" family="Carabidae" genus="Amblygenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306FFF86FC34FD8DB9280F2C" box="[961,1099,543,567]" italics="true" pageId="22" pageNumber="223">Amblygenius</emphasis>
<bibRefCitation id="E606EAB0306FFF86FBA6FD8DBC2D0F43" author="LAFERTE-SENECTERE, F. T." pageId="22" pageNumber="223" pagination="209 - 294" refId="ref31239" refString="LAFERTE-SENECTERE, F. T. 1851. Revision de la tribu des Pelmatellimanes de Dejean, coleopteres pentameres de la famille des Carabiques. Annales de la Societe Entomologique de France (2) 9: 209 - 294." type="journal article" year="1851">Laferté-Sénectère (1851)</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="E606EAB0306FFF86FEAAFDD2BC8B0F4C" author="LORENZ, W." box="[351,488,575,599]" pageId="22" pageNumber="223" refId="ref31666" refString="LORENZ, W. 2005. Systematic List of Extant Ground Beetles of the World (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. W. Lorenz, Tutzing. 530 pp." type="book" year="2005">Lorenz 2005</bibRefCitation>
), in having the pronotum relatively broad and the setose punctures along the lateral margins of the elytral intervals slightly foveate, especially in the apical half apically. Member s of
<taxonomicName id="4597ECC2306FFF86FE9EFD13BC850F83" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[363,486,640,664]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="robertae">
<emphasis id="B0E34B53306FFF86FE9EFD13BC850F83" box="[363,486,640,664]" italics="true" pageId="22" pageNumber="223">C. robertae</emphasis>
</taxonomicName>
differ distinctly from those of the each of the four species Jeannel included in this subgenus. They differ from members of
<taxonomicName id="4597ECC2306FFF86FCF0FD33B9F60FA2" authority="Faimaire (1901)" authorityName="Faimaire" authorityYear="1901" box="[773,1173,673,697]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="cupreolus">
<emphasis id="B0E34B53306FFF86FCF0FD33BE830FA2" box="[773,992,673,697]" italics="true" pageId="22" pageNumber="223">Chlaenius cupreolus</emphasis>
Faimaire (1901)
</taxonomicName>
in having striae deeply impressed and intervals moderately convex (striae superficial only and intervals flat in
<taxonomicName id="4597ECC2306FFF86FEDFFD70BCD70FE1" authorityName="Faimaire" authorityYear="1901" box="[298,436,738,762]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="cupreolus">
<emphasis id="B0E34B53306FFF86FEDFFD70BCD70FE1" box="[298,436,738,762]" italics="true" pageId="22" pageNumber="223">C. cupreolus</emphasis>
</taxonomicName>
members), head with metallic green reflection but pronotum black without a trace of metallic reflection (both head and pronotum with green metallic reflection in
<taxonomicName id="4597ECC2306FFF86FF8EFCB1BC620E20" authorityName="Faimaire" authorityYear="1901" box="[123,257,803,827]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="cupreolus">
<emphasis id="B0E34B53306FFF86FF8EFCB1BC620E20" box="[123,257,803,827]" italics="true" pageId="22" pageNumber="223">C. cupreolus</emphasis>
</taxonomicName>
members), and elytra without a pale lateral and apical margin (a thin pale margin present in
<taxonomicName id="4597ECC2306FFF86FF35FCD1BC2B0E40" authorityName="Faimaire" authorityYear="1901" box="[192,328,835,859]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="cupreolus">
<emphasis id="B0E34B53306FFF86FF35FCD1BC2B0E40" box="[192,328,835,859]" italics="true" pageId="22" pageNumber="223">C. cupreolus</emphasis>
</taxonomicName>
members). They differ from members of
<taxonomicName id="4597ECC2306FFF86FCFAFCD1B9EC0E40" authority="Alluaud (1919)" authorityName="Alluaud" authorityYear="1919" box="[783,1167,835,859]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="allacteus">
<emphasis id="B0E34B53306FFF86FCFAFCD1BE800E40" box="[783,995,835,859]" italics="true" pageId="22" pageNumber="223">Chlaenius allacteus</emphasis>
<bibRefCitation id="E606EAB0306FFF86FC1FFCD1B9EC0E40" author="ALLUAUD, C." box="[1002,1167,835,859]" pageId="22" pageNumber="223" pagination="50 - 53" refId="ref28857" refString="ALLUAUD, C. 1919. Contributions a l'etude des Carabiques d'Afrique et de Madagascar [Col.] XXII. Descriptions de quatre especes et d'un genre nouveaux de Chleniens. Bulletin de la Societe Entomologique de France 1919: 50 - 53." type="journal article" year="1919">Alluaud (1919)</bibRefCitation>
</taxonomicName>
, which is current classified in subgenus
<taxonomicName id="4597ECC2306FFF86FDDDFCF1BE380E67" authority="Alluaud (1933)" authorityName="Alluaud" authorityYear="1933" box="[552,859,867,892]" class="Insecta" family="Carabidae" genus="Oochlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306FFF86FDDDFCF1BFCC0E60" box="[552,687,867,891]" italics="true" pageId="22" pageNumber="223">Oochlaenius</emphasis>
<bibRefCitation id="E606EAB0306FFF86FD40FCF1BE380E67" author="ALLUAUD, C." box="[693,859,867,892]" pageId="22" pageNumber="223" pagination="10 - 20" refId="ref28924" refString="ALLUAUD, C. 1933 II. Chlaeniinae d'Afrique tropicale. Afra, Cahiers d'Entomologie 6: 10 - 20." type="journal article" year="1933">Alluaud (1933)</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="E606EAB0306FFF86FC99FCF6BE960E60" author="LORENZ, W." box="[876,1013,867,891]" pageId="22" pageNumber="223" refId="ref31666" refString="LORENZ, W. 2005. Systematic List of Extant Ground Beetles of the World (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. W. Lorenz, Tutzing. 530 pp." type="book" year="2005">Lorenz 2005</bibRefCitation>
), in having all elytral interval similarly and moderately convex (intervals 1, 3, 5 and 7 more convex and intervals 2, 4, 6 and 8 flat in
<taxonomicName id="4597ECC2306FFF86FEBBFC37BCAE0EA7" authorityName="Alluaud" authorityYear="1919" box="[334,461,932,956]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="allacteus">
<emphasis id="B0E34B53306FFF86FEBBFC37BCAE0EA7" box="[334,461,932,956]" italics="true" pageId="22" pageNumber="223">C. allacteus</emphasis>
</taxonomicName>
members) and, again, elytra without a pale lateral and apical margin (a wide pale margin present in
<taxonomicName id="4597ECC2306FFF86FE18FC57BF0F0EC6" authorityName="Alluaud" authorityYear="1919" box="[493,620,965,989]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="allacteus">
<emphasis id="B0E34B53306FFF86FE18FC57BF0F0EC6" box="[493,620,965,989]" italics="true" pageId="22" pageNumber="223">C. allacteus</emphasis>
</taxonomicName>
members). They differ from members of
<taxonomicName id="4597ECC2306FFF86FBDCFC57BCC50EE6" authority="Faimaire (1901)" authorityName="Faimaire" authorityYear="1901" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B0E34B53306FFF86FBDCFC57BD890EE6" italics="true" pageId="22" pageNumber="223">Chlaenius inaequalis</emphasis>
Faimaire (1901)
</taxonomicName>
in having the pronotum black without metallic reflection (metallic green reflection present on pronotum in
<taxonomicName id="4597ECC2306FFF86FDC9FB94BFAE0905" authorityName="Faimaire" authorityYear="1901" box="[572,717,1030,1054]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B0E34B53306FFF86FDC9FB94BFAE0905" box="[572,717,1030,1054]" italics="true" pageId="22" pageNumber="223">C. inaequalis</emphasis>
</taxonomicName>
members), a longer and narrower elytral form (shorter and broader elytra in
<taxonomicName id="4597ECC2306FFF86FE0AFBB4BFF30925" authorityName="Faimaire" authorityYear="1901" box="[511,656,1062,1086]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B0E34B53306FFF86FE0AFBB4BFF30925" box="[511,656,1062,1086]" italics="true" pageId="22" pageNumber="223">C. inaequalis</emphasis>
</taxonomicName>
members) and elytral epipleurae black (pale in
<taxonomicName id="4597ECC2306FFF86FF8EFBD5BC6E0945" authorityName="Faimaire" authorityYear="1901" box="[123,269,1094,1118]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B0E34B53306FFF86FF8EFBD5BC6E0945" box="[123,269,1094,1118]" italics="true" pageId="22" pageNumber="223">C. inaequalis</emphasis>
</taxonomicName>
members). Overall, members of
<taxonomicName id="4597ECC2306FFF86FD74FBD5BF9C0945" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[641,767,1094,1118]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="robertae">
<emphasis id="B0E34B53306FFF86FD74FBD5BF9C0945" box="[641,767,1094,1118]" italics="true" pageId="22" pageNumber="223">C. robertae</emphasis>
</taxonomicName>
are most similar to those the fourth species,
<taxonomicName id="4597ECC2306FFF86FF22FBF5BF270964" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" box="[215,580,1127,1151]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="lyperus">
<emphasis id="B0E34B53306FFF86FF22FBF5BCF90964" box="[215,410,1127,1151]" italics="true" pageId="22" pageNumber="223">Chlaenius lyperus</emphasis>
Jeannel (1949)
</taxonomicName>
, with which they share similar elytral shape, depth of striae and convexity of intervals. However, they differ from
<taxonomicName id="4597ECC2306FFF86FD0DFB1ABE070984" authorityName="Jeannel" authorityYear="1949" box="[760,868,1159,1183]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="lyperus">
<emphasis id="B0E34B53306FFF86FD0DFB1ABE070984" box="[760,868,1159,1183]" italics="true" pageId="22" pageNumber="223">C. lyperus</emphasis>
</taxonomicName>
members in having a pronotum broader basally than the latter and legs that are pale in color (dark in
<taxonomicName id="4597ECC2306FFF86FC5EFB3AB97809DB" authorityName="Jeannel" authorityYear="1949" box="[939,1051,1192,1216]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="lyperus">
<emphasis id="B0E34B53306FFF86FC5EFB3AB97809DB" box="[939,1051,1192,1216]" italics="true" pageId="22" pageNumber="223">C. lyperus</emphasis>
</taxonomicName>
members). Males of
<taxonomicName id="4597ECC2306FFF86FF17FB5BBC3D09FB" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[226,350,1224,1248]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="robertae">
<emphasis id="B0E34B53306FFF86FF17FB5BBC3D09FB" box="[226,350,1224,1248]" italics="true" pageId="22" pageNumber="223">C. robertae</emphasis>
</taxonomicName>
also have genitalia (
<figureCitation id="1AAC8BC4306FFF86FDB5FB5ABFA909FB" box="[576,714,1224,1248]" captionStart="FIGURE" captionStartId="21.[155,167,1641,1660]" captionTargetBox="[130,1176,216,1618]" captionTargetId="figure-67@21.[120,1176,216,1618]" captionTargetPageId="21" captionText="FIGURE 12. Digital images of holotype male of Chlaenius robertae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm." pageId="22" pageNumber="223">Figs. 12C–D</figureCitation>
) that are markedly different in form those of the other four species (compare with Jeannel’s (1949) Fig. 386a-d).
</paragraph>
<paragraph id="82289741306FFF86FF5EFA9BB969083B" blockId="22.[123,1174,219,1443]" box="[171,1034,1289,1313]" pageId="22" pageNumber="223">
<emphasis id="B0E34B53306FFF86FF5EFA9BBC920804" bold="true" box="[171,497,1289,1313]" pageId="22" pageNumber="223">
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</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E3306FFF86FC8AFA9BBECE083A" box="[895,941,1289,1313]" pageId="22" pageNumber="223">type</typeStatus>
locality.
</paragraph>
<paragraph id="82289741306FFF86FF5EFABBBE4608B9" blockId="22.[123,1174,219,1443]" pageId="22" pageNumber="223">
<emphasis id="B0E34B53306FFF86FF5EFABBBCC6085B" bold="true" box="[171,421,1321,1345]" pageId="22" pageNumber="223">
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</emphasis>
.— Both specimens of the
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series were collected in open habitats at the edges of mixed tropical forest in the Vohiparara area at elevations ranging from
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. One was collected in daytime by treading down vegetation at the edge of a marsh, the other was found at night running on bare soil in an abandoned rice paddy.
</paragraph>
</subSubSection>
</treatment>
</document>

239
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<mods:title id="57ED659307F8977B9542FF5537198867">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:affiliation id="679BA1150FC39BE4C47B5B82FCAC13C9">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:affiliation id="613B0BA9D630E9A6F12DD8A87CDA7841">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
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<emphasis id="B0E34B53306EFF87FF8EFF49BCCA0DE8" bold="true" box="[123,425,219,243]" italics="true" pageId="23" pageNumber="224">Omphreoides ranomafanae</emphasis>
Kavanaugh and Rainio
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,
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.—
<materialsCitation id="32FF9D1C306EFF87FE8AFEA7BF060CB6" collectingDate="1990-02-17" collectingDateMax="1990-02-22" collectingDateMin="1990-02-17" collectorName="W. E. Steiner" country="Madagascar" elevation="900" location="Ranomafana" pageId="23" pageNumber="224" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E3306EFF87FE8AFEA7BC810C56" box="[383,482,309,333]" pageId="23" pageNumber="224" type="holotype">Holotype</typeStatus>
(
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), a female, in NMNH, labeled: “
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: Prov.
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,
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W
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”/ “at black light in montane rainforest near river and stream”/ “
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<emphasis id="B0E34B53306EFF87FC99FEE4B9F60C95" box="[876,1173,374,398]" italics="true" pageId="23" pageNumber="224">Omphreoides ranomafanae</emphasis>
Kavanaugh &amp; Rainio
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2015” [red label].
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.—
<collectingCountry id="FA80D7D1306EFF87FE8BFE24BC9C0CD5" box="[382,511,438,462]" name="Madagascar" pageId="23" pageNumber="224">Madagascar</collectingCountry>
,
<collectingRegion id="405359A3306EFF87FDF8FE24BF9F0CD5" box="[525,764,438,462]" country="Madagascar" name="Fianarantsoa" pageId="23" pageNumber="224">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A306EFF87FCFFFE24B94F0CD5" LSID="urn:lsid:plazi:treatment:0A3E2657306EFF87FF8EFF49BEB4082C:8748C19A306EFF87FCFFFE24B94F0CD5" box="[778,1068,438,462]" country="Madagascar" name="Ranomafana National Park" pageId="23" pageNumber="224" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="82289741306EFF87FF5EFE45BECA0F15" blockId="23.[123,1174,309,1336]" pageId="23" pageNumber="224">
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<smallCapsWord id="84CE019D306EFF87FEACFE48BCCF0CF6" baselines="488" box="[345,428,474,493]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="23" pageNumber="224">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D306EFF87FE40FE48BC910CF6" baselines="488" box="[437,498,474,493]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="23" pageNumber="224">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC2306EFF87FD0CFE45BEE40CF4" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[761,903,471,495]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53306EFF87FD0CFE45BEE40CF4" box="[761,903,471,495]" italics="true" pageId="23" pageNumber="224">ranomafanae</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the national park in which the
<typeStatus id="5D2C29E3306EFF87FD29FE65BE690F14" box="[732,778,503,527]" pageId="23" pageNumber="224">type</typeStatus>
was collected.
</paragraph>
<paragraph id="82289741306EFF87FF5EFD85B94909CD" blockId="23.[123,1174,309,1336]" pageId="23" pageNumber="224">
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<emphasis id="B0E34B53306EFF87FF5EFD85BC210F35" bold="true" box="[171,322,535,559]" pageId="23" pageNumber="224">RECOGNITION</emphasis>
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.— Size larger than average for genus, SBL =
<quantity id="456F3AA4306EFF87FCB4FD85BEC00F34" box="[833,931,535,559]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="23" pageNumber="224" unit="mm" value="13.4">13.4 mm</quantity>
. The unique
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female of
<taxonomicName id="4597ECC2306EFF87FF04FDAABCC60F4B" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[241,421,568,592]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53306EFF87FF04FDAABCC60F4B" box="[241,421,568,592]" italics="true" pageId="23" pageNumber="224">O. ranomafanae</emphasis>
</taxonomicName>
(
<figureCitation id="1AAC8BC4306EFF87FE4FFDAABF7E0F4B" box="[442,541,568,592]" captionStart="FIGURE" captionStartId="24.[155,167,1561,1580]" captionTargetBox="[120,1176,216,1539]" captionTargetId="figure-53@24.[120,1176,216,1539]" captionTargetPageId="24" captionText="FIGURE 13. Digital images of holotype female of Omphreoides ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Forebody, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm" pageId="23" pageNumber="224">Fig. 13A</figureCitation>
) is clearly unlike members of any described species of
<taxonomicName id="4597ECC2306EFF87FF8EFDCABC690F6B" baseAuthorityName="Lorenz" baseAuthorityYear="2005" box="[123,266,600,624]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306EFF87FF8EFDCABC690F6B" box="[123,266,600,624]" italics="true" pageId="23" pageNumber="224">Omphreoides</emphasis>
</taxonomicName>
, with several features intermediate between the four typical species and
<taxonomicName id="4597ECC2306EFF87FBDAFDCAB9EC0F6B" authorityName="Alluaud" authorityYear="1910" box="[1071,1167,600,624]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="quodi">
<emphasis id="B0E34B53306EFF87FBDAFDCAB9EC0F6B" box="[1071,1167,600,624]" italics="true" pageId="23" pageNumber="224">O. quodi</emphasis>
</taxonomicName>
. The head of the
<typeStatus id="5D2C29E3306EFF87FED8FDEABCE90F8B" box="[301,394,632,656]" pageId="23" pageNumber="224" type="holotype">holotype</typeStatus>
(
<figureCitation id="1AAC8BC4306EFF87FE6EFDEABC990F8A" box="[411,506,632,657]" captionStart="FIGURE" captionStartId="24.[155,167,1561,1580]" captionTargetBox="[120,1176,216,1539]" captionTargetId="figure-53@24.[120,1176,216,1539]" captionTargetPageId="24" captionText="FIGURE 13. Digital images of holotype female of Omphreoides ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Forebody, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm" pageId="23" pageNumber="224">Fig. 13C</figureCitation>
) is longer and more slender than that of females of any other species except
<taxonomicName id="4597ECC2306EFF87FEEFFD0BBC140FAA" authorityName="Alluaud" authorityYear="1910" box="[282,375,665,689]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="quodi">
<emphasis id="B0E34B53306EFF87FEEFFD0BBC140FAA" box="[282,375,665,689]" italics="true" pageId="23" pageNumber="224">O. quodi</emphasis>
</taxonomicName>
, which has an even longer and narrower head (see Jeannel 1949, Fig. 364). In
<taxonomicName id="4597ECC2306EFF87FF62FD28BC260FCA" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[151,325,697,721]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53306EFF87FF62FD28BC260FCA" box="[151,325,697,721]" italics="true" pageId="23" pageNumber="224">O. ranomafanae</emphasis>
</taxonomicName>
, the tempora are straight, parallel in anterior their two-thirds, slightly more than twice as long as the diameter of eye, distinctly delimited posteriorly by narrowly round angles. The dorsal longitudinal grooves typical of all members of this genus, except those of
<taxonomicName id="4597ECC2306EFF87FBF2FD68BCF00E29" authority="Fairmaire (1896)" authorityName="Fairmaire" authorityYear="1896" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="bispinus">
<emphasis id="B0E34B53306EFF87FBF2FD68BDB70E29" italics="true" pageId="23" pageNumber="224">Omphreoides bispinus</emphasis>
<bibRefCitation id="E606EAB0306EFF87FF29FC88BCF00E29" author="FAIRMAIRE, L." box="[220,403,794,818]" pageId="23" pageNumber="224" pagination="48 - 50" refId="ref30287" refString="FAIRMAIRE, L. 1896. Description de six Carabiques de la region Malgache. Bulletin de la Societe Entomologique de France 65: 48 - 50." type="journal article" year="1896">Fairmaire (1896)</bibRefCitation>
</taxonomicName>
, are extended as sharply-defined grooves only about halfway from the point of insertion of the anterior supraorbital setae to that of posterior supraorbital setae and not continued on to base of the head as they are in
<taxonomicName id="4597ECC2306EFF87FD67FCC9B9060E68" authority="Alluaud (1899 a)" authorityName="Alluaud" authorityYear="1899" box="[658,1125,859,883]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="bucculentus">
<emphasis id="B0E34B53306EFF87FD67FCC9BECA0E68" box="[658,937,859,883]" italics="true" pageId="23" pageNumber="224">Omphreoides bucculentus</emphasis>
<bibRefCitation id="E606EAB0306EFF87FC47FCC9B9060E68" author="ALLUAUD, C." box="[946,1125,859,883]" pageId="23" pageNumber="224" pagination="410 - 413" refId="ref28748" refString="ALLUAUD, C. 1899 a. Sur quelques Coleoptera de Madagascar de famille Carabiques, recueillis par M. Mocquerys et acquis par le Museum. Bulletin du Museum National d'Histoire Naturelle 5: 410 - 413." type="journal article" year="1899">Alluaud (1899a)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4597ECC2306EFF87FF8EFCE9BF7D0E88" authority="Alluaud (1936)" authorityName="Alluaud" authorityYear="1936" box="[123,542,891,915]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B0E34B53306EFF87FF8EFCE9BC150E88" box="[123,374,891,915]" italics="true" pageId="23" pageNumber="224">Omphreoides distinctus</emphasis>
Alluaud (1936)
</taxonomicName>
. Posterior to the sharply-defined segments, broad and shallow depressions extend posteriorly, arcuately convergent and nearly joined in the midline near the back of the head. The pronotum (
<figureCitation id="1AAC8BC4306EFF87FE2BFC2EBF5F0ECF" box="[478,572,956,980]" captionStart="FIGURE" captionStartId="24.[155,167,1561,1580]" captionTargetBox="[120,1176,216,1539]" captionTargetId="figure-53@24.[120,1176,216,1539]" captionTargetPageId="24" captionText="FIGURE 13. Digital images of holotype female of Omphreoides ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Forebody, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm" pageId="23" pageNumber="224">Fig. 13C</figureCitation>
) is subquadrate, longer than wide (ratio PW/PL = 0.75), with a wavy diagonal row of sparse, coarse punctures (seen also in
<taxonomicName id="4597ECC2306EFF87FCC8FC4FBEA00EEF" authorityName="Alluaud" authorityYear="1936" box="[829,963,988,1012]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B0E34B53306EFF87FCC8FC4FBEA00EEF" box="[829,963,988,1012]" italics="true" pageId="23" pageNumber="224">O. distinctus</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC2306EFF87FC00FC4FB9310EEF" authorityName="Alluaud" authorityYear="1910" box="[1013,1106,988,1012]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="quodi">
<emphasis id="B0E34B53306EFF87FC00FC4FB9310EEF" box="[1013,1106,988,1012]" italics="true" pageId="23" pageNumber="224">O. quodi</emphasis>
</taxonomicName>
members but absent from those of
<taxonomicName id="4597ECC2306EFF87FE3EFC6FBF0D090E" authorityName="Alluaud" authorityYear="1899" box="[459,622,1021,1045]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="bucculentus">
<emphasis id="B0E34B53306EFF87FE3EFC6FBF0D090E" box="[459,622,1021,1045]" italics="true" pageId="23" pageNumber="224">O. bucculentus</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC2306EFF87FD5CFC6FBE45090E" authorityName="Fairmaire" authorityYear="1896" box="[681,806,1021,1045]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="bispinus">
<emphasis id="B0E34B53306EFF87FD5CFC6FBE45090E" box="[681,806,1021,1045]" italics="true" pageId="23" pageNumber="224">O. bispinus</emphasis>
</taxonomicName>
) extended from near the anterior transverse impression paramedially to the basal foveae posteriorly; lateral pronotal margins with short but distinct sinuations anterior to slightly obtuse hind angles; median longitudinal impression deep, wider than in all other
<taxonomicName id="4597ECC2306EFF87FE5EFBCCBF59096D" baseAuthorityName="Lorenz" baseAuthorityYear="2005" box="[427,570,1118,1142]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306EFF87FE5EFBCCBF59096D" box="[427,570,1118,1142]" italics="true" pageId="23" pageNumber="224">Omphreoides</emphasis>
</taxonomicName>
except
<taxonomicName id="4597ECC2306EFF87FD78FBCCBF89096D" authorityName="Alluaud" authorityYear="1910" box="[653,746,1118,1142]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="quodi">
<emphasis id="B0E34B53306EFF87FD78FBCCBF89096D" box="[653,746,1118,1142]" italics="true" pageId="23" pageNumber="224">O. quodi</emphasis>
</taxonomicName>
(in which the impression is even wider) and sparsely but coarsely punctate; lateral explanation narrow (narrower at middle than that in
<taxonomicName id="4597ECC2306EFF87FF8EFB0DBC7909AD" authorityName="Alluaud" authorityYear="1899" box="[123,282,1182,1206]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="bucculentus">
<emphasis id="B0E34B53306EFF87FF8EFB0DBC7909AD" box="[123,282,1182,1206]" italics="true" pageId="23" pageNumber="224">O. bucculentus</emphasis>
</taxonomicName>
members and similar to that in
<taxonomicName id="4597ECC2306EFF87FD92FB0DBF8E09AD" authorityName="Alluaud" authorityYear="1936" box="[615,749,1182,1206]" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B0E34B53306EFF87FD92FB0DBF8E09AD" box="[615,749,1182,1206]" italics="true" pageId="23" pageNumber="224">O. distinctus</emphasis>
</taxonomicName>
member. The elytra are relatively wider than in all other species; each elytron with a straight apical spine in line with interval 3.
</paragraph>
<paragraph id="82289741306EFF87FF5EFB4DB96909EC" blockId="23.[123,1174,309,1336]" box="[171,1034,1247,1272]" pageId="23" pageNumber="224">
<emphasis id="B0E34B53306EFF87FF5EFB4DBC9209EE" bold="true" box="[171,497,1247,1271]" pageId="23" pageNumber="224">
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<smallCapsWord id="84CE019D306EFF87FEA8FB70BC9209EE" baselines="1264" box="[349,497,1250,1269]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="23" pageNumber="224">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
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locality.
</paragraph>
<paragraph id="82289741306EFF87FF5EFA92BEB4082C" blockId="23.[123,1174,309,1336]" pageId="23" pageNumber="224">
<emphasis id="B0E34B53306EFF87FF5EFA92BCC6080D" bold="true" box="[171,421,1280,1304]" pageId="23" pageNumber="224">
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<smallCapsWord id="84CE019D306EFF87FEE4FA91BCC6080D" baselines="1297" box="[273,421,1283,1302]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="23" pageNumber="224">DISTRIBUTION</smallCapsWord>
</emphasis>
.— The unique
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was collected at ultraviolet light in montane rainforest near the junction of the Namorona River and a small tributary stream.
</paragraph>
</subSubSection>
</treatment>
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<mods:affiliation id="9CAE70607DD8F3DC4E2E4505520442F3">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
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<mods:classification id="5BA9B1EC3C462FFEDE76F7E9C307B6B0">journal article</mods:classification>
<mods:identifier id="6DF39DA5B811D1143879324856C4F9F4" type="ISSN">0068-547X</mods:identifier>
</mods:mods>
<treatment id="0A3E26573071FF9AFF8EFD4FB9350CCD" LSID="urn:lsid:plazi:treatment:0A3E26573071FF9AFF8EFD4FB9350CCD" httpUri="http://treatment.plazi.org/id/0A3E26573071FF9AFF8EFD4FB9350CCD" lastPageId="10" lastPageNumber="211" pageId="8" pageNumber="209">
<subSubSection id="CA8DC4CA3071FF98FF8EFD4FBE7F0FEE" box="[123,796,733,757]" pageId="8" pageNumber="209" type="nomenclature">
<paragraph id="822897413071FF98FF8EFD4FBE7F0FEE" blockId="8.[123,796,733,787]" box="[123,796,733,757]" pageId="8" pageNumber="209">
<heading id="D960202D3071FF98FF8EFD4FBE7F0FEE" box="[123,796,733,757]" fontSize="10" level="2" pageId="8" pageNumber="209" reason="4">
<emphasis id="B0E34B533071FF98FF8EFD4FBE7F0FEE" bold="true" box="[123,796,733,757]" pageId="8" pageNumber="209">
<taxonomicName id="4597ECC23071FF98FF8EFD4FBFDB0FEE" ID-CoL="68T5N" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,696,733,757]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="rotundiformis" status="sp. nov.">
<emphasis id="B0E34B533071FF98FF8EFD4FBCCB0FEE" bold="true" box="[123,424,733,757]" italics="true" pageId="8" pageNumber="209">Caelostomus rotundiformis</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F6283071FF98FD33FD4FBE7F0FEE" box="[710,796,733,757]" pageId="8" pageNumber="209" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3071FF98FF8EFD6FBEAC0E25" pageId="8" pageNumber="209" type="description">
<paragraph id="822897413071FF98FF8EFD6FBDAF0E08" blockId="8.[123,796,733,787]" box="[123,204,765,787]" pageId="8" pageNumber="209">
<figureCitation id="1AAC8BC43071FF98FF8EFD6FBDAF0E08" box="[123,204,765,787]" captionStart="FIGURE" captionStartId="9.[155,167,1384,1403]" captionTargetBox="[140,1176,216,1362]" captionTargetId="figure-703@9.[120,1176,216,1381]" captionTargetPageId="9" captionText="FIGURE 5. Digital images of holotype female of Caelostomus rotundiformis Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Right anterior tarsus, dorsal aspect. Scale line A = 1.0 mm, C = 0.5 mm." pageId="8" pageNumber="209">Figure 5</figureCitation>
</paragraph>
<paragraph id="822897413071FF98FF8EFCB5BEAC0E25" blockId="8.[123,975,807,831]" box="[123,975,807,831]" pageId="8" pageNumber="209">
<emphasis id="B0E34B533071FF98FF8EFCB5BC390E24" box="[123,346,807,831]" italics="true" pageId="8" pageNumber="209">
<taxonomicName id="4597ECC23071FF98FF8EFCB5BC660E24" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,261,807,831]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="genus" status="n. sp.">Caelostomus</taxonomicName>
<taxonomicNameLabel id="ABD0F6283071FF98FEF9FCB5BC250E25" box="[268,326,807,830]" pageId="8" pageNumber="209" rank="species">n. sp.</taxonomicNameLabel>
1
</emphasis>
; Rainio 2009: 31, Rainio 2012: 62 (informal designation).
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3071FF9AFF5EFCF0B9350CCD" lastPageId="10" lastPageNumber="211" pageId="8" pageNumber="209" type="materials_examined">
<paragraph id="822897413071FF98FF5EFCF0BFB40921" blockId="8.[123,1174,865,1726]" pageId="8" pageNumber="209">
<emphasis id="B0E34B533071FF98FF5EFCF0BC3E0E63" bold="true" box="[171,349,866,890]" pageId="8" pageNumber="209">
<smallCapsWord id="84CE019D3071FF98FF5EFCF0BD870E63" baselines="884,883" box="[171,228,866,890]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="8" pageNumber="209">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D3071FF98FF05FCF7BC3E0E63" baselines="883" box="[240,349,869,888]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="8" pageNumber="209">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C3071FF98FE71FCF0BDE40EC1" collectingDate="2000-06-07" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="50" location="Vatoharanana" pageId="8" pageNumber="209" specimenCount="1" typeStatus="holotype">
<typeStatus id="5D2C29E33071FF98FE71FCF0BC840E61" box="[388,487,866,890]" pageId="8" pageNumber="209" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43071FF98FE0EFCF0BF180E62" box="[507,635,865,890]" captionStart="FIGURE" captionStartId="9.[155,167,1384,1403]" captionTargetBox="[140,1176,216,1362]" captionTargetId="figure-703@9.[120,1176,216,1381]" captionTargetPageId="9" captionText="FIGURE 5. Digital images of holotype female of Caelostomus rotundiformis Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Right anterior tarsus, dorsal aspect. Scale line A = 1.0 mm, C = 0.5 mm." pageId="8" pageNumber="209">Figs. 5A–B</figureCitation>
), a female, in CAS, labeled: “
<collectingCountry id="FA80D7D13071FF98FC2DFCF0B9F50E61" box="[984,1174,866,890]" name="Madagascar" pageId="8" pageNumber="209">MADAGASCAR</collectingCountry>
Ranomafana NP.,
<location id="8748C19A3071FF98FECCFC10BCA90E81" LSID="urn:lsid:plazi:treatment:0A3E26573071FF9AFF8EFD4FB9350CCD:8748C19A3071FF98FECCFC10BCA90E81" box="[313,458,898,922]" country="Madagascar" name="Vatoharanana" pageId="8" pageNumber="209">Vatoharanana</location>
, Trail K at
<quantity id="456F3AA43071FF98FDB7FC10BF140E82" box="[578,631,898,922]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="209" unit="m" value="50.0">
<elevation id="09BA70723071FF98FDB7FC10BF140E82" box="[578,631,898,922]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="209" unit="m" value="50.0">50 m</elevation>
</quantity>
mark,
<date id="F629B1813071FF98FD35FC10BE5B0E81" box="[704,824,898,922]" pageId="8" pageNumber="209" value="2000-06-07">
<collectingDate id="E66D48693071FF98FD35FC10BE5B0E81" box="[704,824,898,922]" pageId="8" pageNumber="209" value="2000-06-07">07.06.2000</collectingDate>
</date>
”/ “collected by hand in daytime,
<collectorName id="2F62F2973071FF98FF8EFC30BD8D0EA1" box="[123,238,930,954]" pageId="8" pageNumber="209">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973071FF98FEE1FC30BC170EA1" box="[276,372,930,954]" pageId="8" pageNumber="209">J. Rainio</collectorName>
collectors”/ “13.” [handwritten label]/ “7.6.00 K 50” [handwritten label]/ “
</materialsCitation>
<typeStatus id="5D2C29E33071FF98FF72FC50BC770EC1" box="[135,276,962,986]" pageId="8" pageNumber="209" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23071FF98FEEFFC50BE460EC1" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[282,805,962,986]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="rotundiformis" status="sp. nov.">
<emphasis id="B0E34B533071FF98FEEFFC50BF5F0EC1" box="[282,572,962,986]" italics="true" pageId="8" pageNumber="209">Caelostomus rotundiformis</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283071FF98FCDEFC51BE070EC1" box="[811,868,963,986]" pageId="8" pageNumber="209" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
<materialsCitation id="32FF9D1C3071FF98FBDBFC51BE7D0901" collectingDate="2004-03-11" collectionCode="MNHN" collectorName="Johanna Rainio" country="Madagascar" location="MADAGASCAR" pageId="8" pageNumber="209" specimenCount="1" typeStatus="paratype">
<typeStatus id="5D2C29E33071FF98FBDBFC51B9F50EC0" box="[1070,1174,963,987]" pageId="8" pageNumber="209" type="paratype">Paratypes</typeStatus>
(only 1): a female (in
<collectionCode id="E4860F843071FF98FE8EFC71BCAF0EE1" box="[379,460,995,1018]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="8" pageNumber="209">MNHN</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13071FF98FDB1FC70BE610EE1" box="[580,770,994,1018]" name="Madagascar" pageId="8" pageNumber="209">MADAGASCAR</collectingCountry>
Ranomafana NP. Talatakely, C1450
<date id="F629B1813071FF98FF8EFB90BD870901" box="[123,228,1026,1050]" pageId="8" pageNumber="209" value="2004-03-11">
<collectingDate id="E66D48693071FF98FF8EFB90BD870901" box="[123,228,1026,1050]" pageId="8" pageNumber="209" value="2004-03-11">11.3.2004</collectingDate>
</date>
<collectorName id="2F62F2973071FF98FF1BFB90BCF50901" box="[238,406,1026,1050]" pageId="8" pageNumber="209">Johanna Rainio</collectorName>
leg.”/ “112.” [handwritten label]/ “
</materialsCitation>
<typeStatus id="5D2C29E33071FF98FCEBFB91BEC10900" box="[798,930,1027,1051]" pageId="8" pageNumber="209" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC23071FF98FC59FB90BCC90921" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="rotundiformis" status="sp. nov.">
<emphasis id="B0E34B533071FF98FC59FB90BDDC0920" italics="true" pageId="8" pageNumber="209">Caelostomus rotundiformis</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283071FF98FE44FBB1BC880921" box="[433,491,1059,1082]" pageId="8" pageNumber="209" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label].
</paragraph>
<paragraph id="822897413071FF98FF5EFBD1B9500941" blockId="8.[123,1174,865,1726]" box="[171,1075,1091,1115]" pageId="8" pageNumber="209">
<materialsCitation id="32FF9D1C3071FF98FF5EFBD1B94F0940" box="[171,1068,1091,1115]" country="Madagascar" location="Ranomafana National Park" pageId="8" pageNumber="209" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533071FF98FF5EFBD1BC340942" bold="true" box="[171,343,1091,1115]" pageId="8" pageNumber="209">
<smallCapsWord id="84CE019D3071FF98FF5EFBD1BD870942" baselines="1109,1108" box="[171,228,1091,1115]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="8" pageNumber="209">
<typeStatus id="5D2C29E33071FF98FF5EFBD1BD870942" box="[171,228,1091,1115]" pageId="8" pageNumber="209">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3071FF98FF19FBD4BC340942" baselines="1108" box="[236,343,1094,1113]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="8" pageNumber="209">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13071FF98FE8BFBD1BC9C0940" box="[382,511,1091,1115]" name="Madagascar" pageId="8" pageNumber="209">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33071FF98FDF8FBD1BF9F0940" box="[525,764,1091,1115]" country="Madagascar" name="Fianarantsoa" pageId="8" pageNumber="209">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3071FF98FCFFFBD1B94F0940" LSID="urn:lsid:plazi:treatment:0A3E26573071FF9AFF8EFD4FB9350CCD:8748C19A3071FF98FCFFFBD1B94F0940" box="[778,1068,1091,1115]" country="Madagascar" name="Ranomafana National Park" pageId="8" pageNumber="209" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413071FF98FF5EFBF1BEE409A0" blockId="8.[123,1174,865,1726]" pageId="8" pageNumber="209">
<emphasis id="B0E34B533071FF98FF5EFBF1BC970962" bold="true" box="[171,500,1123,1147]" pageId="8" pageNumber="209">
<smallCapsWord id="84CE019D3071FF98FF5EFBF1BC4D0962" baselines="1141,1140" box="[171,302,1123,1147]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="8" pageNumber="209">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3071FF98FEC2FBF4BC300962" baselines="1140" box="[311,339,1126,1145]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="8" pageNumber="209">OF</smallCapsWord>
<smallCapsWord id="84CE019D3071FF98FEAEFBF4BCCD0962" baselines="1140" box="[347,430,1126,1145]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="8" pageNumber="209">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3071FF98FE42FBF4BC970962" baselines="1140" box="[439,500,1126,1145]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="8" pageNumber="209">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23071FF98FD08FBF1BEEC0960" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[765,911,1123,1147]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="rotundiformis">
<emphasis id="B0E34B533071FF98FD08FBF1BEEC0960" box="[765,911,1123,1147]" italics="true" pageId="8" pageNumber="209">rotundiformis</emphasis>
</taxonomicName>
, is an adjective derived from the Latin words,
<emphasis id="B0E34B533071FF98FE83FB11BCB70980" box="[374,468,1155,1179]" italics="true" pageId="8" pageNumber="209">rotundus</emphasis>
, meaning round, and
<emphasis id="B0E34B533071FF98FD3CFB11BE6A0980" box="[713,777,1155,1179]" italics="true" pageId="8" pageNumber="209">forma</emphasis>
, meaning form or shape. The name refers to the very short, broad, rounded form of members of this species.
</paragraph>
<paragraph id="822897413071FF9AFF5EFB51BF050C2F" blockId="8.[123,1174,865,1726]" lastBlockId="10.[123,1174,219,471]" lastPageId="10" lastPageNumber="211" pageId="8" pageNumber="209">
<smallCapsWord id="84CE019D3071FF98FF5EFB51BC2109C1" baselines="1237,1237" box="[171,322,1219,1243]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="8" pageNumber="209">
<emphasis id="B0E34B533071FF98FF5EFB51BC2109C1" bold="true" box="[171,322,1219,1243]" pageId="8" pageNumber="209">RECOGNITION</emphasis>
</smallCapsWord>
.— Size moderately small for genus, SBL = 5.4–5.
<quantity id="456F3AA43071FF98FC9EFB51BEC909C0" box="[875,938,1219,1243]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="209" unit="mm" value="5.0">5 mm</quantity>
(females only), body form (
<figureCitation id="1AAC8BC43071FF98FF4BFB71BC6C09E7" box="[190,271,1251,1276]" captionStart="FIGURE" captionStartId="9.[155,167,1384,1403]" captionTargetBox="[140,1176,216,1362]" captionTargetId="figure-703@9.[120,1176,216,1381]" captionTargetPageId="9" captionText="FIGURE 5. Digital images of holotype female of Caelostomus rotundiformis Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Right anterior tarsus, dorsal aspect. Scale line A = 1.0 mm, C = 0.5 mm." pageId="8" pageNumber="209">Fig. 5A</figureCitation>
) short and broad, ratio SBL/EW = 1.97–2.01. This species is undoubtedly a member of genus
<taxonomicName id="4597ECC23071FF98FF2AFA91BC0A0800" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[223,361,1283,1307]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533071FF98FF2AFA91BC0A0800" box="[223,361,1283,1307]" italics="true" pageId="8" pageNumber="209">Caelostomus</emphasis>
</taxonomicName>
as conceived by Jeannel (1948), although we cannot confirm this based on his key because he used male features only in two couplets to distinguish
<taxonomicName id="4597ECC23071FF98FC54FAB6B9480827" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[929,1067,1316,1340]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533071FF98FC54FAB6B9480827" box="[929,1067,1316,1340]" italics="true" pageId="8" pageNumber="209">Caelostomus</emphasis>
</taxonomicName>
members from those of several other genera and we have only female specimens. However, we examined specimens of all species in all the other Malagasy caelostomine genera while at MNHN and confirmed that this species represents none of them. Observable female features, including body form short and broad, frontal furrows simple, not doubled, parascutellar striae of elytra indistinct, basal setiferous pore situated at base of elytral stria 3, tarsomeres not densely setose ventrally, and anterior tarsomere 4 not bilobed apically (
<figureCitation id="1AAC8BC43071FF98FDEFFA77BF0F08E6" box="[538,620,1508,1533]" captionStart="FIGURE" captionStartId="9.[155,167,1384,1403]" captionTargetBox="[140,1176,216,1362]" captionTargetId="figure-703@9.[120,1176,216,1381]" captionTargetPageId="9" captionText="FIGURE 5. Digital images of holotype female of Caelostomus rotundiformis Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Right anterior tarsus, dorsal aspect. Scale line A = 1.0 mm, C = 0.5 mm." pageId="8" pageNumber="209">Fig. 5C</figureCitation>
), are consistent with
<taxonomicName id="4597ECC23071FF98FCADFA77BE8108E6" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[856,994,1509,1533]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533071FF98FCADFA77BE8108E6" box="[856,994,1509,1533]" italics="true" pageId="8" pageNumber="209">Caelostomus</emphasis>
</taxonomicName>
females. Within this genus, members of
<taxonomicName id="4597ECC23071FF98FE8DF997BF4A0B06" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[376,553,1541,1565]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="rotundiformis">
<emphasis id="B0E34B533071FF98FE8DF997BF4A0B06" box="[376,553,1541,1565]" italics="true" pageId="8" pageNumber="209">C. rotundiformis</emphasis>
</taxonomicName>
share the following features with those of other species of the nominate subgenus: head with frons and frontal furrows smooth, impunctate; elytral base distinctly margined from humerus medially to base of stria 3 or 2; elytral striae 6 and 7 deeply impressed throughout, not effaced apically; lateral elytral intervals narrow, moderately convex; apical part of umbilicate series of setiferous pores not inserted in a deep groove; and female apical abdominal ventrite with (two or) three pairs of setae subapically. However, members of this species confound Jeannel’s (1948) key to species of
<taxonomicName id="4597ECC23070FF99FDACFA2ABF8008CB" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[601,739,1464,1488]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B0E34B533070FF99FDACFA2ABF8008CB" box="[601,739,1464,1488]" italics="true" pageId="9" pageNumber="210">Caelostomus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283070FF99FD1FFA2BBE4108CB" box="[746,802,1465,1488]" pageId="9" pageNumber="210" sensu="stricto">s. str.</taxonomicNameLabel>
Their short and broad elytra (ratio EL/EW = 1.13–1.15) are shared with members of the
<taxonomicName id="4597ECC23070FF99FD4AFA48BE0308E9" authorityName="Tschitscherine" authorityYear="1899" box="[703,864,1498,1522]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="convexiusculus">
<emphasis id="B0E34B533070FF99FD4AFA48BE0308E9" box="[703,864,1498,1522]" italics="true" pageId="9" pageNumber="210">convexiusculus</emphasis>
</taxonomicName>
species group, but their elytral striae are impuncate, whereas those of other species group members are markedly punctate throughout their length. Within the group, their small size (less than 6.0 mm) is shared with
<taxonomicName id="4597ECC23070FF99FF8EF9D2BF130B43" authority="(Tschitscherine 1903)" baseAuthorityName="Tschitscherine" baseAuthorityYear="1903" box="[123,624,1600,1624]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="humilis">
<emphasis id="B0E34B533070FF99FF8EF9D2BC0B0B43" box="[123,360,1600,1624]" italics="true" pageId="9" pageNumber="210">Caelostomus humilis</emphasis>
(
<bibRefCitation id="E606EAB03070FF99FE77F9D2BF050B43" author="TSCHITSCHERINE, T. S." box="[386,614,1600,1624]" pageId="9" pageNumber="210" pagination="38 - 64" refId="ref32431" refString="TSCHITSCHERINE, T. S. 1903. Nouvelles espouvelles especes de la tribu des Platysmatini. Horae Societatis Entomologicae Rossicae 36: 38 - 64." type="journal article" year="1903">Tschitschérine 1903</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4597ECC23070FF99FD72F9D2B9340B43" authority="Straneo (1940)" authorityName="Straneo" authorityYear="1940" box="[647,1111,1600,1624]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="minisculus">
<emphasis id="B0E34B533070FF99FD72F9D2BEFA0B43" box="[647,921,1600,1624]" italics="true" pageId="9" pageNumber="210">Caelostomus minisculus</emphasis>
<bibRefCitation id="E606EAB03070FF99FC5FF9D2B9340B43" author="STRANEO, S. L." box="[938,1111,1600,1624]" pageId="9" pageNumber="210" pagination="36 - 41" refId="ref32325" refString="STRANEO, S. L. 1940. Descrive i nuovi Caelostomus colasi, minusculus, oblongus e substriatus del Madagascar ed enuncia nuovi caraterri dei Caelostomini e del gen. All. Revue Francaise d'Entomologie 7: 36 - 41." type="journal article" year="1940">Straneo (1940)</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4597ECC23070FF99FF8EF9F0BF290B61" authority="Jeannel (1948)" authorityName="Jeannel" authorityYear="1948" box="[123,586,1634,1658]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="minutissimus">
<emphasis id="B0E34B533070FF99FF8EF9F0BCF80B62" box="[123,411,1634,1658]" italics="true" pageId="9" pageNumber="210">Caelostomus minutissimus</emphasis>
Jeannel (1948)
</taxonomicName>
; however, members of these three species are even smaller (SBL = 5.0 mm or less) and, again, have markedly punctate elytral striae. The striae in
<taxonomicName id="4597ECC23070FF99FF8EF934BC4E0BA6" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,301,1701,1725]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="rotundiformis">
<emphasis id="B0E34B533070FF99FF8EF934BC4E0BA6" box="[123,301,1701,1725]" italics="true" pageId="9" pageNumber="210">C. rotundiformis</emphasis>
</taxonomicName>
members are smooth, without evident punctures, and the shape of the pronotum (
<figureCitation id="1AAC8BC43073FF9AFF71FF49BDBB0DE8" box="[132,216,219,243]" captionStart="FIGURE" captionStartId="9.[155,167,1384,1403]" captionTargetBox="[140,1176,216,1362]" captionTargetId="figure-703@9.[120,1176,216,1381]" captionTargetPageId="9" captionText="FIGURE 5. Digital images of holotype female of Caelostomus rotundiformis Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Right anterior tarsus, dorsal aspect. Scale line A = 1.0 mm, C = 0.5 mm." pageId="10" pageNumber="211">Fig. 5A</figureCitation>
) is markedly different, with lateral margins evenly arcuate throughout their length and hind angles markedly obtuse. This species probably represents a distinct species group of its own within subgenus
<taxonomicName id="4597ECC23073FF9AFEC4FE8EBCD80C2F" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[305,443,284,308]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533073FF9AFEC4FE8EBCD80C2F" box="[305,443,284,308]" italics="true" pageId="10" pageNumber="211">Caelostomus</emphasis>
</taxonomicName>
in
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.
</paragraph>
<caption id="D6E8C7C93070FF99FF6EFAFABEB9088E" pageId="9" pageNumber="210" startId="9.[155,167,1384,1403]" targetBox="[140,1176,216,1362]" targetPageId="9" targetType="figure">
<paragraph id="822897413070FF99FF6EFAFABEB9088E" blockId="9.[123,1173,1384,1429]" pageId="9" pageNumber="210">
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5. Digital images of holotype female of
<taxonomicName id="4597ECC23070FF99FDC9FAFABE870860" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[572,996,1384,1403]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="rotundiformis" status="sp. nov.">
<emphasis id="B0E34B533070FF99FDC9FAFABE450860" box="[572,806,1384,1403]" italics="true" pageId="9" pageNumber="210">Caelostomus rotundiformis</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283070FF99FC1FFAFAB94E0860" box="[1002,1069,1384,1403]" pageId="9" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels; C. Right anterior tarsus, dorsal aspect. Scale line A = 1.0 mm, C = 0.5 mm.
</paragraph>
</caption>
<paragraph id="822897413073FF9AFF5EFEAEB9690C4F" blockId="10.[123,1174,219,471]" box="[171,1034,316,341]" pageId="10" pageNumber="211">
<emphasis id="B0E34B533073FF9AFF5EFEAEBC920C48" bold="true" box="[171,497,316,340]" pageId="10" pageNumber="211">
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</emphasis>
.— At present, known only from the
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locality.
</paragraph>
<paragraph id="822897413073FF9AFF5EFECFB9350CCD" blockId="10.[123,1174,219,471]" pageId="10" pageNumber="211">
<emphasis id="B0E34B533073FF9AFF5EFECFBCC60C68" bold="true" box="[171,421,349,373]" pageId="10" pageNumber="211">
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<smallCapsWord id="84CE019D3073FF9AFEE4FEF2BCC60C68" baselines="366" box="[273,421,352,371]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="10" pageNumber="211">DISTRIBUTION</smallCapsWord>
</emphasis>
.— The
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of this species was collected by hand in daytime in primary montane rain forest in the Vatoharanana area at an elevation of
<quantity id="456F3AA43073FF9AFC7DFEECBEB80C8E" box="[904,987,382,406]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="211" unit="m" value="1200.0">1200 m</quantity>
, along Trail K at the
<quantity id="456F3AA43073FF9AFF54FE0CBD9C0CAE" box="[161,255,414,438]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="211" unit="m" value="50.0">50 meter</quantity>
mark. The
<typeStatus id="5D2C29E33073FF9AFE8DFE0CBCB70CAD" box="[376,468,414,438]" pageId="10" pageNumber="211" type="paratype">paratype</typeStatus>
specimen was collected in a pitfall trap in secondary montane rain forest in the Talatakely area at an elevation of
<quantity id="456F3AA43073FF9AFD9BFE2DBFD10CCD" box="[622,690,447,471]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="211" unit="m" value="900.0">900 m</quantity>
, along Trail C at the
<quantity id="456F3AA43073FF9AFC6DFE2CB9720CCD" box="[920,1041,446,470]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="10" pageNumber="211" unit="m" value="1450.0">1450 meter</quantity>
mark.
</paragraph>
</subSubSection>
</treatment>
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266
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<mods:title id="DA45928BCCE6A64702837F46F8F2D900">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:affiliation id="2C6D7F4C156CD937FBFCC1BB1F54B03B">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
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<taxonomicName id="4597ECC23073FF9AFF8EFD4FBF150FEE" ID-CoL="6C9F6" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,630,733,758]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B0E34B533073FF9AFF8EFD4FBC070FED" bold="true" box="[123,356,733,758]" italics="true" pageId="10" pageNumber="211">Dactyleurys minimus</emphasis>
Kavanaugh and Rainio
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,
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,
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,
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.—
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(
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), a male, in CAS, labeled: “
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, Ranomafana NP, Talatakely, X at
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mark
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<collectingDate id="E66D48693073FF9AFD98FCCBBFB40E6B" box="[621,727,856,881]" pageId="10" pageNumber="211" value="2005-04-12">12.4.2005</collectingDate>
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”/ “collected by hand in daytime,
<collectorName id="2F62F2973073FF9AFBB4FCCBBDC20E8A" pageId="10" pageNumber="211">F. Ratalata</collectorName>
&amp; Johanna Rainio collectors”/ “
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<emphasis id="B0E34B533073FF9AFD78FCEBBE0D0E8A" box="[653,878,889,913]" italics="true" pageId="10" pageNumber="211">Dactyleurys minimus</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
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2015” [red label].
</materialsCitation>
</paragraph>
<paragraph id="822897413073FF9AFF5EFC28B9500ECA" blockId="10.[122,1174,824,1725]" box="[171,1075,954,978]" pageId="10" pageNumber="211">
<materialsCitation id="32FF9D1C3073FF9AFF5EFC28B94F0EC9" box="[171,1068,954,978]" country="Madagascar" location="Ranomafana National Park" pageId="10" pageNumber="211" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533073FF9AFF5EFC28BC340ECB" bold="true" box="[171,343,954,978]" pageId="10" pageNumber="211">
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<typeStatus id="5D2C29E33073FF9AFF5EFC28BD870ECA" box="[171,228,954,978]" pageId="10" pageNumber="211">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3073FF9AFF19FC2FBC340ECB" baselines="971" box="[236,343,957,976]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="10" pageNumber="211">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13073FF9AFE8BFC28BC9C0EC9" box="[382,511,954,978]" name="Madagascar" pageId="10" pageNumber="211">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33073FF9AFDF8FC28BF9F0EC9" box="[525,764,954,978]" country="Madagascar" name="Fianarantsoa" pageId="10" pageNumber="211">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3073FF9AFCFFFC28B94F0EC9" LSID="urn:lsid:plazi:treatment:0A3E26573073FF9BFF8EFD4FBCF10BA1:8748C19A3073FF9AFCFFFC28B94F0EC9" box="[778,1068,954,978]" country="Madagascar" name="Ranomafana National Park" pageId="10" pageNumber="211" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413073FF9AFF5EFC49BFC20928" blockId="10.[122,1174,824,1725]" pageId="10" pageNumber="211">
<emphasis id="B0E34B533073FF9AFF5EFC49BC8F0EEA" bold="true" box="[171,492,987,1011]" pageId="10" pageNumber="211">
<smallCapsWord id="84CE019D3073FF9AFF5EFC49BC4D0EEA" baselines="1005,1004" box="[171,302,987,1011]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="10" pageNumber="211">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3073FF9AFEC1FC4CBC330EEA" baselines="1004" box="[308,336,990,1009]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="10" pageNumber="211">OF</smallCapsWord>
<smallCapsWord id="84CE019D3073FF9AFEA0FC4CBCCB0EEA" baselines="1004" box="[341,424,990,1009]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="10" pageNumber="211">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3073FF9AFE5AFC4CBC8F0EEA" baselines="1004" box="[431,492,990,1009]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="10" pageNumber="211">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23073FF9AFD18FC49BE240EE9" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[749,839,987,1010]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B0E34B533073FF9AFD18FC49BE240EE9" box="[749,839,987,1010]" italics="true" pageId="10" pageNumber="211">minimus</emphasis>
</taxonomicName>
, is the Latin adjective meaning smallest, a reference to fact that the known adult male of this species is smaller than males of any of the other known species that we have examined.
</paragraph>
<paragraph id="822897413073FF9BFF5EFBAEBF0D08E1" blockId="10.[122,1174,824,1725]" lastBlockId="11.[123,1174,1347,1723]" lastPageId="11" lastPageNumber="212" pageId="10" pageNumber="211">
<smallCapsWord id="84CE019D3073FF9AFF5EFBAEBC210949" baselines="1102,1101" box="[171,322,1084,1108]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="10" pageNumber="211">
<emphasis id="B0E34B533073FF9AFF5EFBAEBC210949" bold="true" box="[171,322,1084,1108]" pageId="10" pageNumber="211">RECOGNITION</emphasis>
</smallCapsWord>
.— Smallest adult known for the genus, SBL =
<quantity id="456F3AA43073FF9AFCB2FBAEBEFA0948" box="[839,921,1084,1108]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="10" pageNumber="211" unit="mm" value="6.9">6.9 mm</quantity>
. The following combination of features identify members of this species as coelostomine pterostichines of the genus
<taxonomicName id="4597ECC23073FF9AFF8EFBEFBD9F098E" authorityName="Tschitscherine" authorityYear="1899" box="[123,252,1149,1173]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533073FF9AFF8EFBEFBD9F098E" box="[123,252,1149,1173]" italics="true" pageId="10" pageNumber="211">Dactyleurys</emphasis>
</taxonomicName>
: antennae with antennomere 3 attached symmetrically or nearly symmetrically to antennomere 2; mentum with deep emargination, apex of median mental tooth distinctly posterior to apices of epilobes; elytron with epipleuron interrupted by internal plica subapically; parascutellar stria not evident; basal setiferious puncture situated at base of stria 3; tarsomeres 1 to 4 of all legs with dense pads of setae ventrally; front and middle tarsomeres 1 to 3 smoothly convex dorsally; male front tarsomeres 1 to 4 wide, dilated, not latero- or medioapically toothed; aedeagus of male with right face dorsal in repose, right paramere conchoid, left paramere reduced, short, apically digitiform. Members of
<taxonomicName id="4597ECC23073FF9AFE35FAF3BF5F0863" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[448,572,1377,1400]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B0E34B533073FF9AFE35FAF3BF5F0863" box="[448,572,1377,1400]" italics="true" pageId="10" pageNumber="211">D. minimus</emphasis>
</taxonomicName>
(
<figureCitation id="1AAC8BC43073FF9AFDBBFAF2BFC10862" box="[590,674,1376,1401]" captionStart="FIGURE" captionStartId="11.[155,167,1273,1292]" captionTargetBox="[120,1176,216,1254]" captionTargetId="figure-920@11.[120,1176,216,1255]" captionTargetPageId="11" captionText="FIGURE 6. Digital images of holotype male of Dactyleurys minimus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." pageId="10" pageNumber="211">Fig. 6A</figureCitation>
) differ from those of
<taxonomicName id="4597ECC23073FF9AFC61FAF3B97C0863" authorityName="Tschitscherine. Scale" authorityYear="1899" box="[916,1055,1376,1400]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B0E34B533073FF9AFC61FAF3B97C0863" box="[916,1055,1376,1400]" italics="true" pageId="10" pageNumber="211">D. anomalus</emphasis>
</taxonomicName>
in the following features: smaller size (SBL =
<quantity id="456F3AA43073FF9AFDFEFA13BF3F0883" box="[523,604,1409,1433]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="10" pageNumber="211" unit="mm" value="6.9">6.9 mm</quantity>
compared with 7.7–9.
<quantity id="456F3AA43073FF9AFCB8FA13BEE80883" box="[845,907,1409,1433]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="211" unit="mm" value="4.0">4 mm</quantity>
in
<taxonomicName id="4597ECC23073FF9AFC5BFA10B9540882" authorityName="Tschitscherine. Scale" authorityYear="1899" box="[942,1079,1409,1433]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B0E34B533073FF9AFC5BFA10B9540882" box="[942,1079,1409,1433]" italics="true" pageId="10" pageNumber="211">D. anomalus</emphasis>
</taxonomicName>
); pronotum (
<figureCitation id="1AAC8BC43073FF9AFF40FA33BC6408A1" box="[181,263,1441,1466]" captionStart="FIGURE" captionStartId="14.[587,599,1619,1638]" captionTargetBox="[123,1176,216,1715]" captionTargetId="figure-1475@14.[552,1176,216,1601]" captionTargetPageId="14" captionText="FIGURE 8. Digital images of pronota of Dactyleurys spp., dorsal aspect. A. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C. D. anomalus Tschitschérine. Scale lines = 1.0 mm" pageId="10" pageNumber="211">Fig. 8A</figureCitation>
) relatively narrower, widest at middle (relatively wider (
<figureCitation id="1AAC8BC43073FF9AFC9AFA33BEA208A1" box="[879,961,1441,1466]" captionStart="FIGURE" captionStartId="14.[587,599,1619,1638]" captionTargetBox="[123,1176,216,1715]" captionTargetId="figure-1475@14.[552,1176,216,1601]" captionTargetPageId="14" captionText="FIGURE 8. Digital images of pronota of Dactyleurys spp., dorsal aspect. A. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C. D. anomalus Tschitschérine. Scale lines = 1.0 mm" pageId="10" pageNumber="211">Fig. 8C</figureCitation>
) and clearly widest anterior to middle in
<taxonomicName id="4597ECC23073FF9AFEA9FA51BC8608C1" authorityName="Tschitscherine. Scale" authorityYear="1899" box="[348,485,1474,1498]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B0E34B533073FF9AFEA9FA51BC8608C1" box="[348,485,1474,1498]" italics="true" pageId="10" pageNumber="211">D. anomalus</emphasis>
</taxonomicName>
); elytra faintly but evidently iridescence and elytral microsculpture markedly transverse with sculpticells great than 2.5 times as wide as long (elytra not iridescent and elytral microsculpture moderately transverse with sculpticells less than 2.5 times as wide as long in
<taxonomicName id="4597ECC23073FF9AFF1CF9B6BC120B20" authorityName="Tschitscherine. Scale" authorityYear="1899" box="[233,369,1571,1595]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B0E34B533073FF9AFF1CF9B6BC120B20" box="[233,369,1571,1595]" italics="true" pageId="10" pageNumber="211">D. anomalus</emphasis>
</taxonomicName>
); and median lobe of male genitalia (
<figureCitation id="1AAC8BC43073FF9AFCF4F9B1BE310B27" box="[769,850,1571,1596]" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." pageId="10" pageNumber="211">Fig. 9A</figureCitation>
) with shaft abruptly bent ventrally in apical fourth with apex nearly straight in lateral aspect (shaft nearly straight ventrally (
<figureCitation id="1AAC8BC43073FF9AFB9EF9D6BDF90B66" captionStart="FIGURE" captionStartId="14.[587,599,1619,1638]" captionTargetBox="[123,1176,216,1715]" captionTargetId="figure-1475@14.[552,1176,216,1601]" captionTargetPageId="14" captionText="FIGURE 8. Digital images of pronota of Dactyleurys spp., dorsal aspect. A. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C. D. anomalus Tschitschérine. Scale lines = 1.0 mm" pageId="10" pageNumber="211">Fig. 8C</figureCitation>
) in apical fourth with apex slightly recurved dorsally in
<taxonomicName id="4597ECC23073FF9AFD09F9F7BEE50B67" authorityName="Tschitscherine. Scale" authorityYear="1899" box="[764,902,1636,1660]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B0E34B533073FF9AFD09F9F7BEE50B67" box="[764,902,1636,1660]" italics="true" pageId="10" pageNumber="211">D. anomalus</emphasis>
</taxonomicName>
), apex narrowly rounded in dorsal aspect (
<figureCitation id="1AAC8BC43073FF9AFEC6F917BCE60B87" box="[307,389,1669,1693]" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." pageId="10" pageNumber="211">Fig. 9D</figureCitation>
) (more broadly rounded (
<figureCitation id="1AAC8BC43073FF9AFD6BF917BF8F0B87" box="[670,748,1669,1693]" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." pageId="10" pageNumber="211">Fig. 9F</figureCitation>
) in
<taxonomicName id="4597ECC23073FF9AFCEDF914BEC10B86" authorityName="Tschitscherine. Scale" authorityYear="1899" box="[792,930,1669,1693]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B0E34B533073FF9AFCEDF914BEC10B86" box="[792,930,1669,1693]" italics="true" pageId="10" pageNumber="211">D. anomalus</emphasis>
</taxonomicName>
). Members of
<taxonomicName id="4597ECC23073FF9AFBB6F914BDCF0BA6" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B0E34B533073FF9AFBB6F914BDCF0BA6" italics="true" pageId="10" pageNumber="211">D. minimus</emphasis>
</taxonomicName>
differ from those of the other new species,
<taxonomicName id="4597ECC23073FF9AFD7BF937BEC50BA6" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[654,934,1701,1725]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533073FF9AFD7BF937BEC50BA6" box="[654,934,1701,1725]" italics="true" pageId="10" pageNumber="211">Dactyleurys ranomafanae</emphasis>
</taxonomicName>
, in the following features: smaller size (SBL = 7.7–8.
<quantity id="456F3AA43072FF9BFE11FAD1BF460841" box="[484,549,1347,1371]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="212" unit="mm" value="3.0">3 mm</quantity>
in
<taxonomicName id="4597ECC23072FF9BFDBBFAD6BF9C0840" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[590,767,1347,1371]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533072FF9BFDBBFAD6BF9C0840" box="[590,767,1347,1371]" italics="true" pageId="11" pageNumber="212">D. ranomafanae</emphasis>
</taxonomicName>
); pronotum (
<figureCitation id="1AAC8BC43072FF9BFC61FAD1BE8A0840" box="[916,1001,1347,1371]" captionStart="FIGURE" captionStartId="14.[587,599,1619,1638]" captionTargetBox="[123,1176,216,1715]" captionTargetId="figure-1475@14.[552,1176,216,1601]" captionTargetPageId="14" captionText="FIGURE 8. Digital images of pronota of Dactyleurys spp., dorsal aspect. A. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C. D. anomalus Tschitschérine. Scale lines = 1.0 mm" pageId="11" pageNumber="212">Fig. 8A</figureCitation>
) relatively narrower and with evenly rounded lateral margin (slightly wider (
<figureCitation id="1AAC8BC43072FF9BFCC5FAF1BEE00861" box="[816,899,1379,1403]" captionStart="FIGURE" captionStartId="14.[587,599,1619,1638]" captionTargetBox="[123,1176,216,1715]" captionTargetId="figure-1475@14.[552,1176,216,1601]" captionTargetPageId="14" captionText="FIGURE 8. Digital images of pronota of Dactyleurys spp., dorsal aspect. A. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C. D. anomalus Tschitschérine. Scale lines = 1.0 mm" pageId="11" pageNumber="212">Fig. 8B</figureCitation>
) and with lateral margin slightly straighter in basal one-third in
<taxonomicName id="4597ECC23072FF9BFDD4FA16BFB20880" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[545,721,1411,1435]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533072FF9BFDD4FA16BFB20880" box="[545,721,1411,1435]" italics="true" pageId="11" pageNumber="212">D. ranomafanae</emphasis>
</taxonomicName>
); and median lobe of male genitalia (
<figureCitation id="1AAC8BC43072FF9BFB9EFA11BDF808A0" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." pageId="11" pageNumber="212">Fig. 9A</figureCitation>
) with apex nearly straight in lateral aspect (apex slightly recurved dorsally (
<figureCitation id="1AAC8BC43072FF9BFBE0FA31B90808A0" box="[1045,1131,1443,1467]" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." pageId="11" pageNumber="212">Fig. 9C</figureCitation>
) in
<taxonomicName id="4597ECC23072FF9BFF8EFA56BC4A08C0" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,297,1475,1499]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533072FF9BFF8EFA56BC4A08C0" box="[123,297,1475,1499]" italics="true" pageId="11" pageNumber="212">D. ranomafanae</emphasis>
</taxonomicName>
), apical part of shaft (
<figureCitation id="1AAC8BC43072FF9BFDE2FA51BF0B08C1" box="[535,616,1475,1499]" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." pageId="11" pageNumber="212">Fig. 9D</figureCitation>
) slightly deflected right in dorsal aspect (apical part of shaft (
<figureCitation id="1AAC8BC43072FF9BFF29FA71BC4808E1" box="[220,299,1507,1531]" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." pageId="11" pageNumber="212">Fig. 9E</figureCitation>
) straight in
<taxonomicName id="4597ECC23072FF9BFE5AFA76BF3D08E0" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[431,606,1507,1531]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533072FF9BFE5AFA76BF3D08E0" box="[431,606,1507,1531]" italics="true" pageId="11" pageNumber="212">D. ranomafanae</emphasis>
</taxonomicName>
).
</paragraph>
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6. Digital images of holotype male of
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<emphasis id="B0E34B533072FF9BFDC5FB6BBF850810" box="[560,742,1273,1292]" italics="true" pageId="11" pageNumber="212">Dactyleurys minimus</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
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A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm.
</paragraph>
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</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E33072FF9BFC8AF991BECE0B00" box="[895,941,1539,1563]" pageId="11" pageNumber="212">type</typeStatus>
locality.
</paragraph>
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<smallCapsWord id="84CE019D3072FF9BFEE4F9B4BCC60B22" baselines="1588" box="[273,421,1574,1593]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="11" pageNumber="212">DISTRIBUTION</smallCapsWord>
</emphasis>
.— The unique
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was collected by hand in daytime in secondary montane rain forest in the Talatakely area, at an elevation of
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, at the
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mark along Trail “X”. Vegetation in that area consisted of bamboo, small young trees and some larger trees with diameters at breast-height (DBH) of
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. Tree trunks and branches were heavily laden with epiphytes and lianas.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="6416862479F697748223A541CCEEE29F">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:affiliation id="E3EAF366F113BEFDE50C7ED7F4E28BC1">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:namePart id="98B930CB71A70574AD610465EE9C121B">Rainio, Johanna</mods:namePart>
<mods:affiliation id="15616D85358B3F02BEF9AFDD0C701E8A">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
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<taxonomicName id="4597ECC23075FF9CFF8EFF49BFCA0DE8" ID-CoL="6C9F7" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,681,219,243]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533075FF9CFF8EFF49BCFB0DE8" bold="true" box="[123,408,219,243]" italics="true" pageId="12" pageNumber="213">Dactyleurys ranomafanae</emphasis>
Kavanaugh and Rainio
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,
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,
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,
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</paragraph>
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<taxonomicNameLabel id="ABD0F6283075FF9CFF00FEB5BC4A0C27" box="[245,297,295,316]" pageId="12" pageNumber="213" rank="species">n. sp.</taxonomicNameLabel>
1
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; Rainio 2009: 31, Rainio 2012: 62, Rainio 2013: 95 (informal designation).
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.—
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<typeStatus id="5D2C29E33075FF9CFE77FECDBC860C6C" box="[386,485,351,375]" pageId="12" pageNumber="213" type="holotype">Holotype</typeStatus>
(
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), a male, in CAS, labeled: “CASENT 1009744”/ “
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,
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,
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,
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,
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<elevation id="09BA70723075FF9CFBB2FEEDB9EC0C8C" box="[1095,1167,383,407]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="213" unit="m" value="900.0">900 m</elevation>
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, mixed tropical forest,
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<collectingDate id="E66D48693075FF9CFE9EFE0DBF610CA3" box="[363,514,415,440]" pageId="12" pageNumber="213" value="1998-04-26">26 April 1998</collectingDate>
</date>
”/ “
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/
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, Stop #98-83
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collector collected by beating
<taxonomicName id="4597ECC23075FF9CFE49FE52BF440CC3" box="[444,551,448,472]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="genus">
<emphasis id="B0E34B533075FF9CFE49FE52BF440CC3" box="[444,551,448,472]" italics="true" pageId="12" pageNumber="213">Pandanus</emphasis>
</taxonomicName>
tree boluses with debris”/ “
<typeStatus id="5D2C29E33075FF9CFC8CFE52B9650CC3" box="[889,1030,448,472]" pageId="12" pageNumber="213" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23075FF9CFBE1FE52BC9B0CE3" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533075FF9CFBE1FE52BC6A0CE3" italics="true" pageId="12" pageNumber="213">Dactyleurys ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
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2015” [red label]
</materialsCitation>
.
<materialsCitation id="32FF9D1C3075FF9CFCF8FE72B9610F03" collectingDate="2005-04-21" collectionCode="MZF" country="Madagascar" location="Mangevo" municipality="Ranomafana" pageId="12" pageNumber="213" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="5D2C29E33075FF9CFCF8FE72BE160CE3" box="[781,885,480,504]" pageId="12" pageNumber="213" type="paratype">Paratypes</typeStatus>
(a total of 35):
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(in
<collectionCode id="E4860F843075FF9CFF8EFD92BDD10F0C" box="[123,178,512,535]" pageId="12" pageNumber="213">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13075FF9CFED1FD92BC810F03" box="[292,482,512,536]" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
<collectingMunicipality id="624C0D3B3075FF9CFE1CFD92BF130F03" box="[489,624,512,536]" pageId="12" pageNumber="213">Ranomafana</collectingMunicipality>
NP.
<location id="8748C19A3075FF9CFD53FD92BE680F03" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFD53FD92BE680F03" box="[678,779,512,536]" country="Madagascar" municipality="Ranomafana" name="Mangevo" pageId="12" pageNumber="213">Mangevo</location>
, plot 0 m.,
<date id="F629B1813075FF9CFC7EFD92BE950F03" box="[907,1014,512,536]" pageId="12" pageNumber="213" value="2005-04-21">
<collectingDate id="E66D48693075FF9CFC7EFD92BE950F03" box="[907,1014,512,536]" pageId="12" pageNumber="213" value="2005-04-21">21.4.2005</collectingDate>
</date>
”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFBE4FD92B97E0F23" collectingDate="2005-04-21" collectionCode="MZF" country="Madagascar" elevation="100" location="Mangevo" municipality="Ranomafana" pageId="12" pageNumber="213" specimenCount="101" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFBE4FD92B9130F03" box="[1041,1136,512,536]" count="1" pageId="12" pageNumber="213" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFF8EFDB3BDD10F23" box="[123,178,545,568]" pageId="12" pageNumber="213">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13075FF9CFED1FDB2BC810F23" box="[292,482,544,568]" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
<collectingMunicipality id="624C0D3B3075FF9CFE1FFDB2BF120F23" box="[490,625,544,568]" pageId="12" pageNumber="213">Ranomafana</collectingMunicipality>
NP.
<location id="8748C19A3075FF9CFD52FDB3BE6F0F22" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFD52FDB3BE6F0F22" box="[679,780,545,569]" country="Madagascar" municipality="Ranomafana" name="Mangevo" pageId="12" pageNumber="213">Mangevo</location>
, plot
<specimenCount id="94915CC83075FF9CFCB9FDB2BEFB0F23" box="[844,920,544,568]" count="100" pageId="12" pageNumber="213" type="generic">
<quantity id="456F3AA43075FF9CFCB9FDB2BEF20F23" box="[844,913,544,568]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="213" unit="m" value="100.0">
<elevation id="09BA70723075FF9CFCB9FDB2BEF20F23" box="[844,913,544,568]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="213" unit="m" value="100.0">100 m</elevation>
</quantity>
.
</specimenCount>
,
<date id="F629B1813075FF9CFC52FDB2B9720F23" box="[935,1041,544,568]" pageId="12" pageNumber="213" value="2005-04-21">
<collectingDate id="E66D48693075FF9CFC52FDB2B9720F23" box="[935,1041,544,568]" pageId="12" pageNumber="213" value="2005-04-21">21.4.2005</collectingDate>
</date>
”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFBD8FDB2BDDB0F63" collectionCode="CAS" country="Madagascar" pageId="12" pageNumber="213" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFBD8FDB2B9F60F23" box="[1069,1173,544,568]" count="2" pageId="12" pageNumber="213" type="female">2 females</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFF6AFDD2BDB00F43" box="[159,211,576,600]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="12" pageNumber="213" type="Museum">CAS</collectionCode>
) labeled same as
<typeStatus id="5D2C29E33075FF9CFE64FDD2BC8D0F43" box="[401,494,576,600]" pageId="12" pageNumber="213" type="holotype">holotype</typeStatus>
except “CASENT 1009742” and “CASENT 1009743”, respectively
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFF3EFDF2BF4E0FE2" collectingDate="1998-04-25" collectionCode="CAS" collectorName="D. H. Kavanaugh" country="Madagascar" elevation="900" latitude="-21.255" location="Talatakely area" longLatPrecision="126" longitude="47.431667" municipality="Ranomafana National Park" pageId="12" pageNumber="213" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFF3EFDF2BC400F63" box="[203,291,608,632]" count="3" pageId="12" pageNumber="213" type="male">3 males</specimenCount>
and
<specimenCount id="94915CC83075FF9CFE96FDF2BCB30F63" box="[355,464,608,632]" count="2" pageId="12" pageNumber="213" type="female">2 females</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFDF2FDF2BF580F63" box="[519,571,608,632]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="12" pageNumber="213" type="Museum">CAS</collectionCode>
) labeled “CASENT 1009737”, “CASENT 1009738”, “CASENT 1009739”, “CASENT 1009740” and “CASENT 1009741”, respectively/ “
<collectingCountry id="FA80D7D13075FF9CFBF3FD13BDD10FA2" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A33075FF9CFF34FD33BCD30FA2" box="[193,432,673,697]" country="Madagascar" name="Fianarantsoa" pageId="12" pageNumber="213">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B3075FF9CFE4AFD33BF870FA2" box="[447,740,673,697]" pageId="12" pageNumber="213">Ranomafana National Park</collectingMunicipality>
,
<location id="8748C19A3075FF9CFD07FD33BEF60FA3" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFD07FD33BEF60FA3" box="[754,917,673,697]" country="Madagascar" latitude="-21.255" longLatPrecision="126" longitude="47.431667" municipality="Ranomafana National Park" name="Talatakely area" pageId="12" pageNumber="213" stateProvince="Fianarantsoa">Talatakely area</location>
,
<quantity id="456F3AA43075FF9CFC50FD33BE890FA3" box="[933,1002,673,697]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="213" unit="m" value="900.0">
<elevation id="09BA70723075FF9CFC50FD33BE890FA3" box="[933,1002,673,697]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="213" unit="m" value="900.0">900 m</elevation>
</quantity>
, mixed tropical forest,
<date id="F629B1813075FF9CFF31FD53BC340FC2" box="[196,343,705,729]" pageId="12" pageNumber="213" value="1998-04-25">
<collectingDate id="E66D48693075FF9CFF31FD53BC340FC2" box="[196,343,705,729]" pageId="12" pageNumber="213" value="1998-04-25">25 April 1998</collectingDate>
</date>
”/ “
<geoCoordinate id="E7A3F1863075FF9CFE88FD53BC8B0FC2" box="[381,488,705,729]" degrees="21" direction="south" minutes="15. 3" orientation="latitude" pageId="12" pageNumber="213" precision="92" value="-21.255">21°15.3’S</geoCoordinate>
/
<geoCoordinate id="E7A3F1863075FF9CFE03FD53BF000FC3" box="[502,611,705,729]" degrees="47" direction="east" minutes="25. 9" orientation="longitude" pageId="12" pageNumber="213" precision="92" value="47.431667">47°25.9’E</geoCoordinate>
, Stop #98-78
<collectorName id="2F62F2973075FF9CFD0CFD53BECF0FC2" box="[761,940,705,729]" pageId="12" pageNumber="213">D.H. Kavanaugh</collectorName>
collector beaten from vegetation and suspended dead leaves”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFDCAFD73BC1C0EC1" collectingDate="2001-01-02" collectingDateMax="2001-01-22" collectingDateMin="2001-01-02" collectionCode="CAS, MNHN" collectorName="D. H. &amp; K. M Kavanaugh &amp; R. L. Brett &amp; E. Elsom &amp; F. Vargas &amp; R. Ranaivosolo &amp; E. F. Randrianirina &amp; N. Rasoamananana &amp; T. J. Ravelomanana &amp; H. C. Raveloson" country="Madagascar" elevation="900" latitude="-21.25041" location="Talatakely area" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" pageId="12" pageNumber="213" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFDCAFD73BFF50FE2" box="[575,662,737,761]" count="3" pageId="12" pageNumber="213" type="male">3 males</specimenCount>
and
<specimenCount id="94915CC83075FF9CFD26FD73BE5D0FE2" box="[723,830,737,761]" count="2" pageId="12" pageNumber="213" type="female">2 females</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFC87FD73BEC50FE2" box="[882,934,737,761]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="12" pageNumber="213" type="Museum">CAS</collectionCode>
and
<collectionCode id="E4860F843075FF9CFC16FD73B9570FE3" box="[995,1076,737,760]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="213">MNHN</collectionCode>
) labeled “CASENT 1009734”, “CASENT 1009735”, “CASENT 1009736”, “CASENT 1009732” and “CASENT 1009733”, respectively/ “
<collectingCountry id="FA80D7D13075FF9CFDD2FCB3BF850E22" box="[551,742,801,825]" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A33075FF9CFD08FCB3BE950E22" box="[765,1014,801,825]" country="Madagascar" name="Fianarantsoa" pageId="12" pageNumber="213">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B3075FF9CFBFBFCB3BC730E42" pageId="12" pageNumber="213">Ranomafana National Park</collectingMunicipality>
,
<location id="8748C19A3075FF9CFEEAFCD3BCA10E42" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFEEAFCD3BCA10E42" box="[287,450,833,857]" country="Madagascar" latitude="-21.25041" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" name="Talatakely area" pageId="12" pageNumber="213" stateProvince="Fianarantsoa">Talatakely area</location>
,
<quantity id="456F3AA43075FF9CFE27FCD0BF740E42" box="[466,535,834,858]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="213" unit="m" value="900.0">
<elevation id="09BA70723075FF9CFE27FCD0BF740E42" box="[466,535,834,858]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="213" unit="m" value="900.0">900 m</elevation>
</quantity>
, mixed tropical forest,
<date id="F629B1813075FF9CFCEDFCD0BE810E41" box="[792,994,834,858]" pageId="12" pageNumber="213" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">
<collectingDate id="E66D48693075FF9CFCEDFCD0BE810E41" box="[792,994,834,858]" pageId="12" pageNumber="213" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">2-22 January 2001</collectingDate>
</date>
,”/ “
<geoCoordinate id="E7A3F1863075FF9CFBE5FCD0B9ED0E41" box="[1040,1166,833,858]" degrees="21. 25041" direction="south" orientation="latitude" pageId="12" pageNumber="213" precision="1" value="-21.25041">21.25041°S</geoCoordinate>
/
<geoCoordinate id="E7A3F1863075FF9CFF8EFCF0BD9A0E62" box="[123,249,865,890]" degrees="47. 41945" direction="east" orientation="longitude" pageId="12" pageNumber="213" precision="1" value="47.41945">47.41945°E</geoCoordinate>
, Stop# DHK-01-001,
<collectorName id="2F62F2973075FF9CFE0FFCF0BE700E62" box="[506,787,865,890]" pageId="12" pageNumber="213">D.H. &amp; K.M Kavanaugh</collectorName>
,
<collectorName id="2F62F2973075FF9CFCD2FCF0BEFB0E62" box="[807,920,866,889]" pageId="12" pageNumber="213">R.L. Brett</collectorName>
,
<collectorName id="2F62F2973075FF9CFC59FCF0B9700E62" box="[940,1043,865,889]" pageId="12" pageNumber="213">E. Elsom</collectorName>
,
<collectorName id="2F62F2973075FF9CFBD2FCF0B9EC0E61" box="[1063,1167,866,890]" pageId="12" pageNumber="213">F. Vargas</collectorName>
,
<collectorName id="2F62F2973075FF9CFF8EFC10BC430E81" box="[123,288,898,922]" pageId="12" pageNumber="213">R. Ranaivosolo</collectorName>
,”/ “
<collectorName id="2F62F2973075FF9CFEBBFC10BF700E81" box="[334,531,898,922]" pageId="12" pageNumber="213">E.F. Randrianirina</collectorName>
,
<collectorName id="2F62F2973075FF9CFDD4FC10BF930E82" box="[545,752,898,921]" pageId="12" pageNumber="213">N. Rasoamananana</collectorName>
,
<collectorName id="2F62F2973075FF9CFD0BFC10BEA90E81" box="[766,970,898,922]" pageId="12" pageNumber="213">T.J. Ravelomanana</collectorName>
, and
<collectorName id="2F62F2973075FF9CFBF2FC10BDC30EA2" pageId="12" pageNumber="213">H.C. Raveloson</collectorName>
collectors”/ “collected in daytime by beating live understory vegetation and/or dead leaves and other suspended debris”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFE7AFC50BCC40EE1" collectingDate="2001-02-26" collectionCode="MZF" country="Madagascar" location="Talatakely" municipality="Ranomafana" pageId="12" pageNumber="213" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFE7AFC50BCBA0EC1" box="[399,473,962,986]" count="1" pageId="12" pageNumber="213" type="male">1 male</specimenCount>
and
<specimenCount id="94915CC83075FF9CFDE4FC50BF130EC1" box="[529,624,962,986]" count="1" pageId="12" pageNumber="213" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFD6BFC50BFB60EC2" box="[670,725,962,985]" pageId="12" pageNumber="213">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13075FF9CFCBDFC50B9650EC1" box="[840,1030,962,986]" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
<collectingMunicipality id="624C0D3B3075FF9CFBFAFC50B9F50EC1" box="[1039,1174,962,986]" pageId="12" pageNumber="213">Ranomafana</collectingMunicipality>
NP.
<location id="8748C19A3075FF9CFF5BFC70BC780EE1" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFF5BFC70BC780EE1" box="[174,283,994,1018]" country="Madagascar" municipality="Ranomafana" name="Talatakely" pageId="12" pageNumber="213">Talatakely</location>
,
<date id="F629B1813075FF9CFEC5FC70BCF90EE1" box="[304,410,994,1018]" pageId="12" pageNumber="213" value="2001-02-26">
<collectingDate id="E66D48693075FF9CFEC5FC70BCF90EE1" box="[304,410,994,1018]" pageId="12" pageNumber="213" value="2001-02-26">26.2.2001</collectingDate>
</date>
”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFE49FC70BFB40901" collectingDate="2000-06-08" collectionCode="MZF" country="Madagascar" elevation="1150" location="Talatakely" municipality="Ranomafana" pageId="12" pageNumber="213" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFE49FC70BF430EE1" box="[444,544,994,1018]" count="1" pageId="12" pageNumber="213" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFDAFFC71BFF20EE1" box="[602,657,995,1018]" pageId="12" pageNumber="213">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13075FF9CFCFAFC70BEAE0EE1" box="[783,973,994,1018]" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
<collectingMunicipality id="624C0D3B3075FF9CFC2EFC70B9010EE1" box="[987,1122,994,1018]" pageId="12" pageNumber="213">Ranomafana</collectingMunicipality>
NP.
<location id="8748C19A3075FF9CFF8EFB90BD8B0901" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFF8EFB90BD8B0901" box="[123,232,1026,1050]" country="Madagascar" municipality="Ranomafana" name="Talatakely" pageId="12" pageNumber="213">Talatakely</location>
, trail B
<quantity id="456F3AA43075FF9CFEBCFB90BCFF0901" box="[329,412,1026,1050]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="12" pageNumber="213" unit="m" value="1150.0">
<elevation id="09BA70723075FF9CFEBCFB90BCFF0901" box="[329,412,1026,1050]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="12" pageNumber="213" unit="m" value="1150.0">1150 m</elevation>
</quantity>
., in
<taxonomicName id="4597ECC23075FF9CFE24FB90BF180901" box="[465,635,1026,1050]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="species" species="undefined-8">
<emphasis id="B0E34B533075FF9CFE24FB90BF5F0901" box="[465,572,1026,1050]" italics="true" pageId="12" pageNumber="213">Pandanus</emphasis>
sp.
<collectingDate id="E66D48693075FF9CFD9BFB90BF180901" box="[622,635,1026,1050]" pageId="12" pageNumber="213" value="2000-06-08">8</collectingDate>
</taxonomicName>
.6.2000”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFD1DFB90B9ED0920" collectingDate="2000-06-16" collectionCode="MZF" country="Madagascar" elevation="380" location="Talatakely" municipality="Ranomafana" pageId="12" pageNumber="213" specimenCount="382" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFD1DFB90BE570901" box="[744,820,1026,1050]" count="1" pageId="12" pageNumber="213" type="male">1 male</specimenCount>
and
<specimenCount id="94915CC83075FF9CFC9BFB90BEAD0901" box="[878,974,1026,1050]" count="1" pageId="12" pageNumber="213" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFC0AFB91B9550901" box="[1023,1078,1027,1050]" pageId="12" pageNumber="213">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13075FF9CFF72FBB1BC260920" box="[135,325,1059,1083]" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
<collectingMunicipality id="624C0D3B3075FF9CFEA5FBB0BCB40921" box="[336,471,1058,1082]" pageId="12" pageNumber="213">Ranomafana</collectingMunicipality>
NP.
<location id="8748C19A3075FF9CFDE6FBB0BFE30921" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFDE6FBB0BFE30921" box="[531,640,1058,1082]" country="Madagascar" municipality="Ranomafana" name="Talatakely" pageId="12" pageNumber="213">Talatakely</location>
, trail BF
<specimenCount id="94915CC83075FF9CFD00FBB1BE260921" box="[757,837,1059,1083]" count="380" pageId="12" pageNumber="213" type="generic">
<quantity id="456F3AA43075FF9CFD00FBB1BE5D0921" box="[757,830,1059,1083]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="12" pageNumber="213" unit="m" value="380.0">
<elevation id="09BA70723075FF9CFD00FBB1BE5D0921" box="[757,830,1059,1083]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="12" pageNumber="213" unit="m" value="380.0">380 m</elevation>
</quantity>
.
</specimenCount>
, in
<taxonomicName id="4597ECC23075FF9CFC82FBB0B96F0921" box="[887,1036,1058,1082]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B0E34B533075FF9CFC82FBB0BE810921" box="[887,994,1058,1082]" italics="true" pageId="12" pageNumber="213">Pandanus</emphasis>
sp.
</taxonomicName>
<date id="F629B1813075FF9CFBE2FBB0B9E10920" box="[1047,1154,1058,1083]" pageId="12" pageNumber="213" value="2000-06-16">
<collectingDate id="E66D48693075FF9CFBE2FBB0B9E10920" box="[1047,1154,1058,1083]" pageId="12" pageNumber="213" value="2000-06-16">16.6.2000</collectingDate>
</date>
”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFF8EFBD1BF670960" collectingDate="2000-06-16" collectionCode="MZF" country="Madagascar" elevation="200" location="Talatakely" municipality="Ranomafana" pageId="12" pageNumber="213" specimenCount="202" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFF8EFBD1BDA60940" box="[123,197,1091,1115]" count="1" pageId="12" pageNumber="213" type="male">1 male</specimenCount>
and
<specimenCount id="94915CC83075FF9CFF08FBD1BC3E0940" box="[253,349,1091,1115]" count="1" pageId="12" pageNumber="213" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFE79FBD1BCA00941" box="[396,451,1091,1114]" pageId="12" pageNumber="213">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13075FF9CFDC3FBD1BF970940" box="[566,756,1091,1115]" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
<collectingMunicipality id="624C0D3B3075FF9CFD08FBD1BEE70940" box="[765,900,1091,1115]" pageId="12" pageNumber="213">Ranomafana</collectingMunicipality>
NP.
<location id="8748C19A3075FF9CFC4EFBD1B94B0940" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFC4EFBD1B94B0940" box="[955,1064,1091,1115]" country="Madagascar" municipality="Ranomafana" name="Talatakely" pageId="12" pageNumber="213">Talatakely</location>
, trail BM
<specimenCount id="94915CC83075FF9CFF8EFBF1BDA40961" box="[123,199,1123,1147]" count="200" pageId="12" pageNumber="213" type="generic">
<quantity id="456F3AA43075FF9CFF8EFBF1BDA30961" box="[123,192,1123,1147]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="213" unit="m" value="200.0">
<elevation id="09BA70723075FF9CFF8EFBF1BDA30961" box="[123,192,1123,1147]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="213" unit="m" value="200.0">200 m</elevation>
</quantity>
.
</specimenCount>
, in
<taxonomicName id="4597ECC23075FF9CFF06FBF1BCE60960" box="[243,389,1123,1147]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B0E34B533075FF9CFF06FBF1BC3D0960" box="[243,350,1123,1147]" italics="true" pageId="12" pageNumber="213">Pandanus</emphasis>
sp.
</taxonomicName>
<date id="F629B1813075FF9CFE7BFBF1BC9B0960" box="[398,504,1123,1147]" pageId="12" pageNumber="213" value="2000-06-16">
<collectingDate id="E66D48693075FF9CFE7BFBF1BC9B0960" box="[398,504,1123,1147]" pageId="12" pageNumber="213" value="2000-06-16">16.6.2000</collectingDate>
</date>
”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFDE1FBF1BE120980" collectingDate="2000-06-06" collectionCode="MZF" country="Madagascar" elevation="50" location="Talatakely" municipality="Ranomafana" pageId="12" pageNumber="213" specimenCount="53" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFDE1FBF1BF3D0960" box="[532,606,1123,1147]" count="1" pageId="12" pageNumber="213" type="male">1 male</specimenCount>
and
<specimenCount id="94915CC83075FF9CFD63FBF1BF9C0960" box="[662,767,1123,1147]" count="2" pageId="12" pageNumber="213" type="female">2 females</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFCDBFBF1BE060961" box="[814,869,1123,1146]" pageId="12" pageNumber="213">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13075FF9CFC2DFBF1B9F50960" box="[984,1174,1123,1147]" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
<collectingMunicipality id="624C0D3B3075FF9CFF8EFB11BC610980" box="[123,258,1155,1179]" pageId="12" pageNumber="213">Ranomafana</collectingMunicipality>
NP.
<location id="8748C19A3075FF9CFEC7FB11BCFC0980" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFEC7FB11BCFC0980" box="[306,415,1155,1179]" country="Madagascar" municipality="Ranomafana" name="Talatakely" pageId="12" pageNumber="213">Talatakely</location>
, trail FBF
<specimenCount id="94915CC83075FF9CFDE4FB11BF2E0981" box="[529,589,1155,1179]" count="50" pageId="12" pageNumber="213" type="generic">
<quantity id="456F3AA43075FF9CFDE4FB11BF240981" box="[529,583,1155,1179]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="213" unit="m" value="50.0">
<elevation id="09BA70723075FF9CFDE4FB11BF240981" box="[529,583,1155,1179]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="213" unit="m" value="50.0">50 m</elevation>
</quantity>
.
</specimenCount>
, in
<taxonomicName id="4597ECC23075FF9CFD81FB11BE760980" box="[628,789,1155,1179]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="species" species="undefined-6">
<emphasis id="B0E34B533075FF9CFD81FB11BFBC0980" box="[628,735,1155,1179]" italics="true" pageId="12" pageNumber="213">Pandanus</emphasis>
sp.
<collectingDate id="E66D48693075FF9CFCFDFB11BE760980" box="[776,789,1155,1179]" pageId="12" pageNumber="213" value="2000-06-06">6</collectingDate>
</taxonomicName>
.6.2000”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFC8BFB11B9ED09A0" collectingDate="2000-06-06" collectionCode="MZF" country="Madagascar" elevation="380" location="Talatakely" municipality="Ranomafana" pageId="12" pageNumber="213" specimenCount="381" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFC8BFB11BEB90980" box="[894,986,1155,1179]" count="1" pageId="12" pageNumber="213" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFBF6FB11B9590981" box="[1027,1082,1155,1178]" pageId="12" pageNumber="213">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13075FF9CFF72FB31BC2609A0" box="[135,325,1187,1211]" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
<collectingMunicipality id="624C0D3B3075FF9CFEA5FB31BCB409A0" box="[336,471,1187,1211]" pageId="12" pageNumber="213">Ranomafana</collectingMunicipality>
NP.
<location id="8748C19A3075FF9CFDE6FB31BFE309A0" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFDE6FB31BFE309A0" box="[531,640,1187,1211]" country="Madagascar" municipality="Ranomafana" name="Talatakely" pageId="12" pageNumber="213">Talatakely</location>
, trail FBF
<specimenCount id="94915CC83075FF9CFCF6FB31BE3009A0" box="[771,851,1187,1211]" count="380" pageId="12" pageNumber="213" type="generic">
<quantity id="456F3AA43075FF9CFCF6FB31BE2F09A0" box="[771,844,1187,1211]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="12" pageNumber="213" unit="m" value="380.0">
<elevation id="09BA70723075FF9CFCF6FB31BE2F09A0" box="[771,844,1187,1211]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="12" pageNumber="213" unit="m" value="380.0">380 m</elevation>
</quantity>
.
</specimenCount>
, in
<taxonomicName id="4597ECC23075FF9CFC70FB31B95109A0" box="[901,1074,1187,1211]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="species" species="undefined-6">
<emphasis id="B0E34B533075FF9CFC70FB31BE9309A0" box="[901,1008,1187,1211]" italics="true" pageId="12" pageNumber="213">Pandanus</emphasis>
sp.
<collectingDate id="E66D48693075FF9CFBD0FB31B95109A0" box="[1061,1074,1187,1211]" pageId="12" pageNumber="213" value="2000-06-06">6</collectingDate>
</taxonomicName>
.6.2000”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFF8EFB51BD9209E0" collectingDate="2001-03-22" collectionCode="MZF" country="Madagascar" elevation="50" location="Talatakely" municipality="Ranomafana" pageId="12" pageNumber="213" specimenCount="52" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFF8EFB51BD8A09C0" box="[123,233,1219,1243]" count="2" pageId="12" pageNumber="213" type="female">2 females</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFED4FB56BC3B09C0" box="[289,344,1220,1243]" pageId="12" pageNumber="213">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13075FF9CFE20FB51BFF009C0" box="[469,659,1219,1243]" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
<collectingMunicipality id="624C0D3B3075FF9CFD55FB51BE4409C0" box="[672,807,1219,1243]" pageId="12" pageNumber="213">Ranomafana</collectingMunicipality>
NP.
<location id="8748C19A3075FF9CFC92FB51BEB709C0" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFC92FB51BEB709C0" box="[871,980,1219,1243]" country="Madagascar" municipality="Ranomafana" name="Talatakely" pageId="12" pageNumber="213">Talatakely</location>
, trail SP
<specimenCount id="94915CC83075FF9CFBBEFB51B9EC09C0" box="[1099,1167,1219,1243]" count="50" pageId="12" pageNumber="213" type="generic">
<quantity id="456F3AA43075FF9CFBBEFB51B9EB09C0" box="[1099,1160,1219,1243]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="213" unit="m" value="50.0">
<elevation id="09BA70723075FF9CFBBEFB51B9EB09C0" box="[1099,1160,1219,1243]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="213" unit="m" value="50.0">50 m</elevation>
</quantity>
.
</specimenCount>
,
<date id="F629B1813075FF9CFF8EFB71BD8609E0" box="[123,229,1251,1275]" pageId="12" pageNumber="213" value="2001-03-22">
<collectingDate id="E66D48693075FF9CFF8EFB71BD8609E0" box="[123,229,1251,1275]" pageId="12" pageNumber="213" value="2001-03-22">22.3.2001</collectingDate>
</date>
”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFF0AFB71BC6F0807" collectingDate="2005-02-09" collectionCode="MZF" country="Madagascar" elevation="50" location="Talatakely" municipality="Ranomafana" pageId="12" pageNumber="213" specimenCount="51" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFF0AFB71BC3F09E0" box="[255,348,1251,1275]" count="1" pageId="12" pageNumber="213" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFE72FB76BCDD09E0" box="[391,446,1252,1275]" pageId="12" pageNumber="213">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13075FF9CFDD8FB71BF8809E0" box="[557,747,1251,1275]" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
<collectingMunicipality id="624C0D3B3075FF9CFD07FB71BE1A09E0" box="[754,889,1251,1275]" pageId="12" pageNumber="213">Ranomafana</collectingMunicipality>
NP.
<location id="8748C19A3075FF9CFC5EFB71B97B09E0" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFC5EFB71B97B09E0" box="[939,1048,1251,1275]" country="Madagascar" municipality="Ranomafana" name="Talatakely" pageId="12" pageNumber="213">Talatakely</location>
, trail SP
<specimenCount id="94915CC83075FF9CFB8EFB71BDF50800" count="50" pageId="12" pageNumber="213" type="generic">
<quantity id="456F3AA43075FF9CFB8EFB71BDF30800" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="213" unit="m" value="50.0">
<elevation id="09BA70723075FF9CFB8EFB71BDF30800" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="213" unit="m" value="50.0">50 m</elevation>
</quantity>
.
</specimenCount>
,
<date id="F629B1813075FF9CFF56FA96BC630800" box="[163,256,1283,1308]" pageId="12" pageNumber="213" value="2005-02-09">
<collectingDate id="E66D48693075FF9CFF56FA96BC630800" box="[163,256,1283,1308]" pageId="12" pageNumber="213" value="2005-02-09">9.2.2005</collectingDate>
</date>
”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFEECFA96BC790827" collectingDate="2005-04-12" collectionCode="MZF" country="Madagascar" elevation="130" location="Talatakely" municipality="Ranomafana" pageId="12" pageNumber="213" specimenCount="131" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFEECFA96BC030800" box="[281,352,1283,1308]" count="1" pageId="12" pageNumber="213" type="male">1 male</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFE7FFA96BCA20800" box="[394,449,1284,1307]" pageId="12" pageNumber="213">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13075FF9CFDDBFA96BF8F0807" box="[558,748,1284,1308]" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
<collectingMunicipality id="624C0D3B3075FF9CFD07FA91BE1A0800" box="[754,889,1283,1307]" pageId="12" pageNumber="213">Ranomafana</collectingMunicipality>
NP.
<location id="8748C19A3075FF9CFC5FFA91B9740800" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFC5FFA91B9740800" box="[938,1047,1283,1307]" country="Madagascar" municipality="Ranomafana" name="Talatakely" pageId="12" pageNumber="213">Talatakely</location>
, trail X
<specimenCount id="94915CC83075FF9CFB98FA96BDF50820" count="130" pageId="12" pageNumber="213" type="generic">
<quantity id="456F3AA43075FF9CFB98FA96BDF30820" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="213" unit="m" value="130.0">
<elevation id="09BA70723075FF9CFB98FA96BDF30820" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="213" unit="m" value="130.0">130 m</elevation>
</quantity>
.
</specimenCount>
,
<date id="F629B1813075FF9CFF56FAB6BC6D0820" box="[163,270,1315,1340]" pageId="12" pageNumber="213" value="2005-04-12">
<collectingDate id="E66D48693075FF9CFF56FAB6BC6D0820" box="[163,270,1315,1340]" pageId="12" pageNumber="213" value="2005-04-12">12.4.2005</collectingDate>
</date>
”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFEDDFAB6BC2A0847" collectingDate="2001-02-28" collectionCode="MNHN" country="Madagascar" elevation="900" location="Valohoaka" municipality="Ranomafana" pageId="12" pageNumber="213" specimenCount="901" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFEDDFAB6BC130827" box="[296,368,1316,1340]" count="1" pageId="12" pageNumber="213" type="male">1 male</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFE6FFAB6BC880820" box="[410,491,1316,1339]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="213">MNHN</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13075FF9CFDAFFAB6BE7B0827" box="[602,792,1316,1340]" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
<collectingMunicipality id="624C0D3B3075FF9CFCEBFAB6BEC60827" box="[798,933,1316,1340]" pageId="12" pageNumber="213">Ranomafana</collectingMunicipality>
NP.
<location id="8748C19A3075FF9CFC22FAB6B92B0827" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFC22FAB6B92B0827" box="[983,1096,1316,1340]" country="Madagascar" municipality="Ranomafana" name="Valohoaka" pageId="12" pageNumber="213">Valohoaka</location>
, trail F
<specimenCount id="94915CC83075FF9CFF8EFAD6BDA60840" box="[123,197,1348,1372]" count="900" pageId="12" pageNumber="213" type="generic">
<quantity id="456F3AA43075FF9CFF8EFAD6BDDD0840" box="[123,190,1348,1372]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="213" unit="m" value="900.0">
<elevation id="09BA70723075FF9CFF8EFAD6BDDD0840" box="[123,190,1348,1372]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="213" unit="m" value="900.0">900 m</elevation>
</quantity>
.
</specimenCount>
,
<date id="F629B1813075FF9CFF27FAD6BC5F0847" box="[210,316,1348,1372]" pageId="12" pageNumber="213" value="2001-02-28">
<collectingDate id="E66D48693075FF9CFF27FAD6BC5F0847" box="[210,316,1348,1372]" pageId="12" pageNumber="213" value="2001-02-28">28.2.2001</collectingDate>
</date>
”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFEA2FAD6BC0D0867" collectingDate="2002-10-16" collectionCode="MZF" country="Madagascar" elevation="850" location="Valohoaka" municipality="Ranomafana" pageId="12" pageNumber="213" specimenCount="851" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFEA2FAD6BCFD0847" box="[343,414,1348,1372]" count="1" pageId="12" pageNumber="213" type="male">1 male</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFE3CFAD6BF630840" box="[457,512,1348,1371]" pageId="12" pageNumber="213">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13075FF9CFD9AFAD6BE4E0847" box="[623,813,1348,1372]" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
<collectingMunicipality id="624C0D3B3075FF9CFCC6FAD6BED90847" box="[819,954,1348,1372]" pageId="12" pageNumber="213">Ranomafana</collectingMunicipality>
NP.
<location id="8748C19A3075FF9CFC18FAD6B93D0847" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFC18FAD6B93D0847" box="[1005,1118,1348,1372]" country="Madagascar" municipality="Ranomafana" name="Valohoaka" pageId="12" pageNumber="213">Valohoaka</location>
, trail F
<specimenCount id="94915CC83075FF9CFF64FAF6BDBF0860" box="[145,220,1380,1404]" count="850" pageId="12" pageNumber="213" type="generic">
<quantity id="456F3AA43075FF9CFF64FAF6BDB50860" box="[145,214,1380,1404]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="213" unit="m" value="850.0">
<elevation id="09BA70723075FF9CFF64FAF6BDB50860" box="[145,214,1380,1404]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="213" unit="m" value="850.0">850 m</elevation>
</quantity>
.
</specimenCount>
,
<date id="F629B1813075FF9CFF1FFAF6BC010867" box="[234,354,1380,1404]" pageId="12" pageNumber="213" value="2002-10-16">
<collectingDate id="E66D48693075FF9CFF1FFAF6BC010867" box="[234,354,1380,1404]" pageId="12" pageNumber="213" value="2002-10-16">16.10.2002</collectingDate>
</date>
”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFE88FAF6BF270887" collectingDate="2004-12-18" collectionCode="MNHN" country="Madagascar" elevation="850" location="Valohoaka" municipality="Ranomafana" pageId="12" pageNumber="213" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFE88FAF6BCA50867" box="[381,454,1380,1404]" count="1" pageId="12" pageNumber="213" type="male">1 male</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFE07FAF6BF200860" box="[498,579,1380,1403]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="213">MNHN</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13075FF9CFD40FAF6BE100867" box="[693,883,1380,1404]" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
<collectingMunicipality id="624C0D3B3075FF9CFC8FFAF6B9620867" box="[890,1025,1380,1404]" pageId="12" pageNumber="213">Ranomafana</collectingMunicipality>
NP.
<location id="8748C19A3075FF9CFBC0FAF6BDF70887" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFBC0FAF6BDF70887" country="Madagascar" municipality="Ranomafana" name="Valohoaka" pageId="12" pageNumber="213">Valohoaka</location>
, trail F
<quantity id="456F3AA43075FF9CFEF2FA16BC360887" box="[263,341,1412,1436]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="213" unit="m" value="850.0">
<elevation id="09BA70723075FF9CFEF2FA16BC360887" box="[263,341,1412,1436]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="213" unit="m" value="850.0">850 m</elevation>
</quantity>
down.,
<date id="F629B1813075FF9CFE4AFA16BF5B0887" box="[447,568,1412,1436]" pageId="12" pageNumber="213" value="2004-12-18">
<collectingDate id="E66D48693075FF9CFE4AFA16BF5B0887" box="[447,568,1412,1436]" pageId="12" pageNumber="213" value="2004-12-18">18.12.2004</collectingDate>
</date>
”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFDA9FA16BFBC08A7" collectingDate="2005-01-25" collectionCode="MNHN" country="Madagascar" elevation="850" location="Valohoaka" municipality="Ranomafana" pageId="12" pageNumber="213" specimenCount="851" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFDA9FA16BFA00887" box="[604,707,1412,1436]" count="1" pageId="12" pageNumber="213" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFCF7FA17BE370887" box="[770,852,1413,1436]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="213">MNHN</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13075FF9CFC22FA16B9F60887" box="[983,1173,1412,1436]" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
<collectingMunicipality id="624C0D3B3075FF9CFF8EFA36BC6108A7" box="[123,258,1444,1468]" pageId="12" pageNumber="213">Ranomafana</collectingMunicipality>
NP.
<location id="8748C19A3075FF9CFECEFA36BCCF08A7" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFECEFA36BCCF08A7" box="[315,428,1444,1468]" country="Madagascar" municipality="Ranomafana" name="Valohoaka" pageId="12" pageNumber="213">Valohoaka</location>
, trail F
<specimenCount id="94915CC83075FF9CFDFFFA36BF3B08A7" box="[522,600,1444,1468]" count="850" pageId="12" pageNumber="213" type="generic">
<quantity id="456F3AA43075FF9CFDFFFA36BF3108A7" box="[522,594,1444,1468]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="213" unit="m" value="850.0">
<elevation id="09BA70723075FF9CFDFFFA36BF3108A7" box="[522,594,1444,1468]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="213" unit="m" value="850.0">850 m</elevation>
</quantity>
.
</specimenCount>
,
<date id="F629B1813075FF9CFD9CFA36BFB008A7" box="[617,723,1444,1468]" pageId="12" pageNumber="213" value="2005-01-25">
<collectingDate id="E66D48693075FF9CFD9CFA36BFB008A7" box="[617,723,1444,1468]" pageId="12" pageNumber="213" value="2005-01-25">25.1.2005</collectingDate>
</date>
”
</materialsCitation>
;
<materialsCitation id="32FF9D1C3075FF9CFD04FA36BC8B0B07" collectingDate="2005-01-26" collectionCode="MNHN, MZF" collectorName="All" country="Madagascar" elevation="850" location="Valohoaka" pageId="12" pageNumber="213" specimenCount="853" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="94915CC83075FF9CFD04FA36BE2508A7" box="[753,838,1444,1468]" count="2" pageId="12" pageNumber="213" type="male">2 males</specimenCount>
and
<specimenCount id="94915CC83075FF9CFC74FA36BE8108A7" box="[897,994,1444,1468]" count="1" pageId="12" pageNumber="213" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843075FF9CFBE6FA37B90708A7" box="[1043,1124,1445,1468]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="213">MNHN</collectionCode>
and
<collectionCode id="E4860F843075FF9CFF8EFA57BDD108C7" box="[123,178,1477,1500]" pageId="12" pageNumber="213">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13075FF9CFEDFFA57BC8B08C6" box="[298,488,1477,1501]" name="Madagascar" pageId="12" pageNumber="213">MADAGASCAR</collectingCountry>
Ranomafana NP.
<location id="8748C19A3075FF9CFD42FA56BE4B08C7" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFD42FA56BE4B08C7" box="[695,808,1476,1500]" country="Madagascar" name="Valohoaka" pageId="12" pageNumber="213">Valohoaka</location>
, trail F
<specimenCount id="94915CC83075FF9CFC7EFA56BEB908C7" box="[907,986,1476,1500]" count="850" pageId="12" pageNumber="213" type="generic">
<quantity id="456F3AA43075FF9CFC7EFA56BEB008C7" box="[907,979,1476,1500]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="213" unit="m" value="850.0">
<elevation id="09BA70723075FF9CFC7EFA56BEB008C7" box="[907,979,1476,1500]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="213" unit="m" value="850.0">850 m</elevation>
</quantity>
.
</specimenCount>
,
<date id="F629B1813075FF9CFC19FA56B93508C7" box="[1004,1110,1476,1501]" pageId="12" pageNumber="213" value="2005-01-26">
<collectingDate id="E66D48693075FF9CFC19FA56B93508C7" box="[1004,1110,1476,1501]" pageId="12" pageNumber="213" value="2005-01-26">26.1.2005</collectingDate>
</date>
”.
<collectorName id="2F62F2973075FF9CFB86FA56B9F608C7" box="[1139,1173,1476,1500]" pageId="12" pageNumber="213">All</collectorName>
<typeStatus id="5D2C29E33075FF9CFF8EFA77BD8208E6" box="[123,225,1509,1533]" pageId="12" pageNumber="213" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="5D2C29E33075FF9CFDBCFA77BFAE08E6" box="[585,717,1509,1533]" pageId="12" pageNumber="213" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC23075FF9CFD2DFA77BDA10B06" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533075FF9CFD2DFA77BE9108E6" box="[728,1010,1509,1533]" italics="true" pageId="12" pageNumber="213">Dactyleurys ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283075FF9CFF3CF997BC600B07" box="[201,259,1541,1564]" pageId="12" pageNumber="213" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="822897413075FF9CFF5EF9B7B9500B27" blockId="12.[123,1174,351,1725]" box="[171,1075,1573,1597]" pageId="12" pageNumber="213">
<materialsCitation id="32FF9D1C3075FF9CFF5EF9B7B94F0B26" box="[171,1068,1573,1597]" country="Madagascar" location="Ranomafana National Park" pageId="12" pageNumber="213" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533075FF9CFF5EF9B7BC340B20" bold="true" box="[171,343,1573,1597]" pageId="12" pageNumber="213">
<smallCapsWord id="84CE019D3075FF9CFF5EF9B7BD870B27" baselines="1591,1591" box="[171,228,1573,1597]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="12" pageNumber="213">
<typeStatus id="5D2C29E33075FF9CFF5EF9B7BD870B27" box="[171,228,1573,1597]" pageId="12" pageNumber="213">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3075FF9CFF19F9BABC340B20" baselines="1590" box="[236,343,1576,1595]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="12" pageNumber="213">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13075FF9CFE8BF9B7BC9C0B26" box="[382,511,1573,1597]" name="Madagascar" pageId="12" pageNumber="213">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33075FF9CFDF8F9B7BF9F0B26" box="[525,764,1573,1597]" country="Madagascar" name="Fianarantsoa" pageId="12" pageNumber="213">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3075FF9CFCFFF9B7B94F0B26" LSID="urn:lsid:plazi:treatment:0A3E26573075FF9FFF8EFF49BCE50B42:8748C19A3075FF9CFCFFF9B7B94F0B26" box="[778,1068,1573,1597]" country="Madagascar" name="Ranomafana National Park" pageId="12" pageNumber="213" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413075FF9CFF5EF9D7BECA0B66" blockId="12.[123,1174,351,1725]" pageId="12" pageNumber="213">
<emphasis id="B0E34B533075FF9CFF5EF9D7BC910B40" bold="true" box="[171,498,1605,1629]" pageId="12" pageNumber="213">
<smallCapsWord id="84CE019D3075FF9CFF5EF9D7BC4D0B40" baselines="1623,1622" box="[171,302,1605,1629]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="12" pageNumber="213">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3075FF9CFEC3F9DABC310B40" baselines="1622" box="[310,338,1608,1627]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="12" pageNumber="213">OF</smallCapsWord>
<smallCapsWord id="84CE019D3075FF9CFEACF9DABCCF0B40" baselines="1622" box="[345,428,1608,1627]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="12" pageNumber="213">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3075FF9CFE40F9DABC910B40" baselines="1622" box="[437,498,1608,1627]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="12" pageNumber="213">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23075FF9CFD0CF9D7BEE40B46" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[761,903,1605,1629]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533075FF9CFD0CF9D7BEE40B46" box="[761,903,1605,1629]" italics="true" pageId="12" pageNumber="213">ranomafanae</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the national park in which the
<typeStatus id="5D2C29E33075FF9CFD29F9F4BE690B65" box="[732,778,1638,1662]" pageId="12" pageNumber="213">type</typeStatus>
was collected.
</paragraph>
<paragraph id="822897413075FF9FFF5EF917BF53086B" blockId="12.[123,1174,351,1725]" lastBlockId="15.[123,1174,1235,1625]" lastPageId="15" lastPageNumber="216" pageId="12" pageNumber="213">
<smallCapsWord id="84CE019D3075FF9CFF5EF917BC210B87" baselines="1687,1687" box="[171,322,1669,1693]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="12" pageNumber="213">
<emphasis id="B0E34B533075FF9CFF5EF917BC210B87" bold="true" box="[171,322,1669,1693]" pageId="12" pageNumber="213">RECOGNITION</emphasis>
</smallCapsWord>
.— Size average for genus, SBL of males = 7.7–8.
<quantity id="456F3AA43075FF9CFC9DF917BEC40B86" box="[872,935,1669,1693]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="213" unit="mm" value="2.0">2 mm</quantity>
, of females = 7.8–
<quantity id="456F3AA43075FF9CFB81F917BDC70BA6" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="12" pageNumber="213" unit="mm" value="8.4">8.4 mm</quantity>
. The following combination of features identify members of this species as coelostomine (
<figureCitation id="1AAC8BC43076FF9FFF71FB41BDB609F0" box="[132,213,1235,1259]" captionStart="FIGURE" captionStartId="14.[587,599,1619,1638]" captionTargetBox="[123,1176,216,1715]" captionTargetId="figure-1475@14.[552,1176,216,1601]" captionTargetPageId="14" captionText="FIGURE 8. Digital images of pronota of Dactyleurys spp., dorsal aspect. A. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C. D. anomalus Tschitschérine. Scale lines = 1.0 mm" pageId="15" pageNumber="216">Fig. 8B</figureCitation>
) relatively broader and with lateral margins slightly straighter in basal one-third (slightly narrower and with lateral margins more evenly rounded (
<figureCitation id="1AAC8BC43076FF9FFD2CFB66BE490816" box="[729,810,1268,1293]" captionStart="FIGURE" captionStartId="14.[587,599,1619,1638]" captionTargetBox="[123,1176,216,1715]" captionTargetId="figure-1475@14.[552,1176,216,1601]" captionTargetPageId="14" captionText="FIGURE 8. Digital images of pronota of Dactyleurys spp., dorsal aspect. A. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C. D. anomalus Tschitschérine. Scale lines = 1.0 mm" pageId="15" pageNumber="216">Fig. 8A</figureCitation>
) in
<taxonomicName id="4597ECC23076FF9FFCA0FB67BEAC0817" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[853,975,1269,1292]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B0E34B533076FF9FFCA0FB67BEAC0817" box="[853,975,1269,1292]" italics="true" pageId="15" pageNumber="216">D. minimus</emphasis>
</taxonomicName>
); and median lobe of male genitalia (
<figureCitation id="1AAC8BC43076FF9FFEB1FA84BCF50836" box="[324,406,1302,1326]" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." pageId="15" pageNumber="216">Fig. 9B</figureCitation>
) with apex slightly recurved dorsally in lateral aspect (apex (
<figureCitation id="1AAC8BC43076FF9FFBCCFA84B9EF0835" box="[1081,1164,1302,1326]" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." pageId="15" pageNumber="216">Fig. 9A</figureCitation>
) nearly straight in
<taxonomicName id="4597ECC23076FF9FFECEFAAABCDB0854" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[315,440,1336,1359]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B0E34B533076FF9FFECEFAAABCDB0854" box="[315,440,1336,1359]" italics="true" pageId="15" pageNumber="216">D. minimus</emphasis>
</taxonomicName>
), apical part of shaft (
<figureCitation id="1AAC8BC43076FF9FFD5BFAA5BF9D0854" box="[686,766,1335,1359]" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." pageId="15" pageNumber="216">Fig. 9E</figureCitation>
) straight (apical part of shaft slightly deflected right (
<figureCitation id="1AAC8BC43076FF9FFED3FACABC1B086B" box="[294,376,1368,1393]" captionStart="FIGURE" captionStartId="15.[155,167,1141,1160]" captionTargetBox="[147,1159,229,1126]" captionTargetId="figure-936@15.[120,1176,216,1138]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of aedeagus of male genitalia of Dactyleurys spp. A–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of D. minimus Kavanaugh &amp; Rainio sp. nov.; B, E. Holotype of D. ranomafanae Kavanaugh &amp; Rainio sp. nov.; C, F. D. anomalus Tschitschérine. Scale lines = 0.5 mm." pageId="15" pageNumber="216">Fig. 9D</figureCitation>
) in
<taxonomicName id="4597ECC23076FF9FFE50FACBBF43086B" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[421,544,1369,1392]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B0E34B533076FF9FFE50FACBBF43086B" box="[421,544,1369,1392]" italics="true" pageId="15" pageNumber="216">D. minimus</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="D6E8C7C93074FF9DFF6EFA63BC8B0B05" pageId="13" pageNumber="214" startId="13.[155,167,1521,1540]" targetBox="[139,1176,226,1499]" targetPageId="13" targetType="figure">
<paragraph id="822897413074FF9DFF6EFA63BC8B0B05" blockId="13.[123,1173,1521,1566]" pageId="13" pageNumber="214">
<smallCapsWord id="84CE019D3074FF9DFF6EFA63BDBB0B19" baselines="1535,1534" box="[155,216,1521,1540]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="13" pageNumber="214">FIGURE</smallCapsWord>
7. Digital images of holotype male of
<taxonomicName id="4597ECC23074FF9DFDD1FA63BEDD0B1F" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[548,958,1521,1540]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533074FF9DFDD1FA63BE600B1F" box="[548,771,1521,1540]" italics="true" pageId="13" pageNumber="214">Dactyleurys ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283074FF9DFC31FA63B9660B1F" box="[964,1029,1521,1540]" pageId="13" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm.
</paragraph>
</caption>
<caption id="D6E8C7C93077FF9EFDBEF9C1BE910BA9" pageId="14" pageNumber="215" startId="14.[587,599,1619,1638]" targetBox="[123,1176,216,1715]" targetPageId="14" targetType="figure">
<paragraph id="822897413077FF9EFDBEF9C1B9F60B64" blockId="14.[555,1173,1618,1714]" pageId="14" pageNumber="215">
<smallCapsWord id="84CE019D3077FF9EFDBEF9C1BFEB0B7F" baselines="1633,1632" box="[587,648,1619,1638]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="14" pageNumber="215">FIGURE</smallCapsWord>
8. Digital images of pronota of
<taxonomicName id="4597ECC23077FF9EFC48F9C0B9320B7D" box="[957,1105,1618,1638]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B0E34B533077FF9EFC48F9C0B9470B7E" box="[957,1060,1618,1637]" italics="true" pageId="14" pageNumber="215">Dactyleurys</emphasis>
spp.
</taxonomicName>
, dorsal aspect. A. Holotype of
<taxonomicName id="4597ECC23077FF9EFCF2F9FEB95E0B64" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[775,1085,1644,1663]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B0E34B533077FF9EFCF2F9FEBE130B65" box="[775,880,1644,1662]" italics="true" pageId="14" pageNumber="215">D. minimus</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283077FF9EFBBDF9FEB9EC0B64" box="[1096,1167,1644,1663]" pageId="14" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
;
</paragraph>
<paragraph id="822897413077FF9EFDDEF914B9F60B82" blockId="14.[555,1173,1618,1714]" box="[555,1173,1669,1689]" pageId="14" pageNumber="215">
B. Holotype of
<taxonomicName id="4597ECC23077FF9EFD3FF914B9540B83" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[714,1079,1669,1688]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533077FF9EFD3FF914BE020B83" box="[714,865,1669,1688]" italics="true" pageId="14" pageNumber="215">D. ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283077FF9EFBB1F914B9EC0B82" box="[1092,1167,1670,1689]" pageId="14" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
;
</paragraph>
<paragraph id="822897413077FF9EFDDEF90DBE910BA9" blockId="14.[555,1173,1618,1714]" box="[555,1010,1695,1714]" pageId="14" pageNumber="215">
C.
<taxonomicName id="4597ECC23077FF9EFDB1F932BE080BA9" authority="Tschitscherine. Scale" authorityName="Tschitscherine. Scale" authorityYear="1899" box="[580,875,1695,1714]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B0E34B533077FF9EFDB1F932BFD10BA9" box="[580,690,1695,1714]" italics="true" pageId="14" pageNumber="215">D. anomalus</emphasis>
Tschitschérine. Scale
</taxonomicName>
lines = 1.0 mm
</paragraph>
</caption>
<caption id="D6E8C7C93076FF9FFF6EFBE7BFEF09A0" pageId="15" pageNumber="216" startId="15.[155,167,1141,1160]" targetBox="[147,1159,229,1126]" targetPageId="15" targetType="figure">
<paragraph id="822897413076FF9FFF6EFBE7BFEF09A0" blockId="15.[123,1173,1140,1211]" pageId="15" pageNumber="216">
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9. Digital images of aedeagus of male genitalia of
<taxonomicName id="4597ECC23076FF9FFD78FBE6BE530993" box="[653,816,1140,1160]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B0E34B533076FF9FFD78FBE6BF97099C" box="[653,756,1140,1159]" italics="true" pageId="15" pageNumber="216">Dactyleurys</emphasis>
spp. A
</taxonomicName>
–C. left lateral aspect; D–F. dorsal aspect. A, D. Holotype of
<taxonomicName id="4597ECC23076FF9FFEE8FB1DBF5F09BA" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[285,572,1166,1185]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B0E34B533076FF9FFEE8FB1DBC1C09BA" box="[285,383,1167,1185]" italics="true" pageId="15" pageNumber="216">D. minimus</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283076FF9FFDB7FB1DBFE709B9" box="[578,644,1167,1186]" pageId="15" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
; B, E. Holotype of
<taxonomicName id="4597ECC23076FF9FFCDBFB1DB91409BA" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[814,1143,1166,1185]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533076FF9FFCDBFB1DBED809BA" box="[814,955,1166,1185]" italics="true" pageId="15" pageNumber="216">D. ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283076FF9FFB88FB1DBDFD09A0" pageId="15" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
; C, F.
<taxonomicName id="4597ECC23076FF9FFF2CFB3ABF6309A0" authority="Tschitscherine. Scale" authorityName="Tschitscherine. Scale" authorityYear="1899" box="[217,512,1192,1211]" class="Insecta" family="Carabidae" genus="Dactyleurys" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B0E34B533076FF9FFF2CFB3ABC2409A0" box="[217,327,1192,1211]" italics="true" pageId="15" pageNumber="216">D. anomalus</emphasis>
Tschitschérine. Scale
</taxonomicName>
lines = 0.5 mm.
</paragraph>
</caption>
<paragraph id="822897413076FF9FFF5EFAEBB969088A" blockId="15.[123,1174,1235,1625]" box="[171,1034,1401,1426]" pageId="15" pageNumber="216">
<emphasis id="B0E34B533076FF9FFF5EFAEBBC92088B" bold="true" box="[171,497,1401,1425]" pageId="15" pageNumber="216">
<smallCapsWord id="84CE019D3076FF9FFF5EFAEBBC35088B" baselines="1419,1419" box="[171,342,1401,1425]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="15" pageNumber="216">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="84CE019D3076FF9FFEA8FAEFBC92088B" baselines="1419" box="[349,497,1405,1424]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="15" pageNumber="216">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E33076FF9FFC8AFAE8BECE0889" box="[895,941,1402,1426]" pageId="15" pageNumber="216">type</typeStatus>
locality.
</paragraph>
<paragraph id="822897413076FF9FFF5EFA09BCE50B42" blockId="15.[123,1174,1235,1625]" pageId="15" pageNumber="216">
<emphasis id="B0E34B533076FF9FFF5EFA09BCC708AA" bold="true" box="[171,420,1435,1459]" pageId="15" pageNumber="216">
<smallCapsWord id="84CE019D3076FF9FFF5EFA09BC6A08AA" baselines="1453,1452" box="[171,265,1435,1459]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="15" pageNumber="216">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D3076FF9FFEE5FA0CBCC708AA" baselines="1452" box="[272,420,1438,1457]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="15" pageNumber="216">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Specimens of this species have been collected in both primary and secondary montane rainforest throughout RNP, at elevations ranging from of
<quantity id="456F3AA43076FF9FFC43FA2EB93208CF" box="[950,1105,1468,1492]" metricMagnitude="3" metricUnit="m" metricValue="1.04" metricValueMax="1.18" metricValueMin="0.9" pageId="15" pageNumber="216" unit="m" value="1040.0" valueMax="1180.0" valueMin="900.0">900 to 1180 m</quantity>
. Most of the specimens were collected by beating vegetation, especially plants of
<taxonomicName id="4597ECC23076FF9FFC55FA4CB95308EE" box="[928,1072,1502,1526]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="15" pageNumber="216" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B0E34B533076FF9FFC55FA4CB96808ED" box="[928,1035,1502,1526]" italics="true" pageId="15" pageNumber="216">Pandanus</emphasis>
sp.
</taxonomicName>
, and suspended accumulations of dead vegetative debris in shrubs and understory trees. One was collected in a malaise trap, another sifted from forest litter at the primary forest site, and another by hand at the secondary forest site.
</paragraph>
</subSubSection>
</treatment>
</document>

309
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<mods:title id="E008233179CE267B4082DE1FE2927DBF">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:namePart id="DFFB82C4AA2C8D07039CDFB6BB0634E4">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="5AFDB64A447608CFB08271ED969F7027">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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</mods:role>
<mods:namePart id="0AEFC71EDCA4AF73F109D2B9C6BAAF53">Rainio, Johanna</mods:namePart>
<mods:affiliation id="67077618F0DB9EBA1C6BE81B04A45AF4">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
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<treatment id="0A3E2657307DFF95FF8EFCD9BC8D0B62" LSID="urn:lsid:plazi:treatment:0A3E2657307DFF95FF8EFCD9BC8D0B62" httpUri="http://treatment.plazi.org/id/0A3E2657307DFF95FF8EFCD9BC8D0B62" lastPageId="5" lastPageNumber="206" pageId="4" pageNumber="205">
<subSubSection id="CA8DC4CA307DFF94FF8EFCD9BE760E7F" box="[123,789,843,868]" pageId="4" pageNumber="205" type="nomenclature">
<paragraph id="82289741307DFF94FF8EFCD9BE760E7F" blockId="4.[123,789,843,898]" box="[123,789,843,868]" pageId="4" pageNumber="205">
<heading id="D960202D307DFF94FF8EFCD9BE760E7F" box="[123,789,843,868]" fontSize="10" level="2" pageId="4" pageNumber="205" reason="4">
<emphasis id="B0E34B53307DFF94FF8EFCD9BE760E7F" bold="true" box="[123,789,843,868]" pageId="4" pageNumber="205">
<taxonomicName id="4597ECC2307DFF94FF8EFCD9BFD20E78" ID-CoL="MYCC" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,689,843,868]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B53307DFF94FF8EFCD9BCC30E78" bold="true" box="[123,416,843,867]" italics="true" pageId="4" pageNumber="205">Brachypelus ranomafanae</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F628307DFF94FD4BFCDEBE760E7F" box="[702,789,844,868]" pageId="4" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA307DFF94FF8EFCFEBFA40EB0" pageId="4" pageNumber="205" type="description">
<paragraph id="82289741307DFF94FF8EFCFEBD8E0E9A" blockId="4.[123,789,843,898]" box="[123,237,876,898]" pageId="4" pageNumber="205">
<figureCitation id="1AAC8BC4307DFF94FF8EFCFEBD8E0E9A" box="[123,237,876,898]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStartId-0="5.[155,167,1152,1171]" captionStartId-1="6.[155,167,1136,1155]" captionTargetBox-0="[133,1174,245,1135]" captionTargetBox-1="[306,980,229,1110]" captionTargetId-0="figure-997@5.[120,1174,216,1150]" captionTargetId-1="figure-647@6.[306,980,215,1110]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 2. Digital images of holotype female of Brachypelus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." captionText-1="FIGURE 3. Digital images of holotype female of Brachypelus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Head, dorsal aspect; B. Elytral base, dorsal aspect. Scale lines = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13799443" figureDoi-1="http://doi.org/10.5281/zenodo.13799447" httpUri-0="https://zenodo.org/record/13799443/files/figure.png" httpUri-1="https://zenodo.org/record/13799447/files/figure.png" pageId="4" pageNumber="205">Figures 2–3</figureCitation>
</paragraph>
<paragraph id="82289741307DFF94FF8EFC07BFA40EB0" blockId="4.[123,711,917,939]" box="[123,711,917,939]" pageId="4" pageNumber="205">
<emphasis id="B0E34B53307DFF94FF8EFC07BC5A0EB1" box="[123,313,917,939]" italics="true" pageId="4" pageNumber="205">
<taxonomicName id="4597ECC2307DFF94FF8EFC07BD900EB1" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,243,917,938]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="genus" status="n. sp.">Brachypelus</taxonomicName>
<taxonomicNameLabel id="ABD0F628307DFF94FF0CFC04BC4E0EB0" box="[249,301,918,939]" pageId="4" pageNumber="205" rank="species">n. sp.</taxonomicNameLabel>
1
</emphasis>
; Rainio 2009: 31 (informal designation).
</paragraph>
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<smallCapsWord id="84CE019D307DFF94FF05FC50BC3E0ECE" baselines="976" box="[240,349,962,981]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="4" pageNumber="205">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C307DFF94FE71FC2CBF80090C" collectingDate="2001-01-15" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="130" location="MADAGASCAR" pageId="4" pageNumber="205" specimenCount="130" typeStatus="holotype">
<typeStatus id="5D2C29E3307DFF94FE71FC2CBC840ECD" box="[388,487,958,982]" pageId="4" pageNumber="205" type="holotype">Holotype</typeStatus>
(
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), a female, in CAS, labeled: “
<collectingCountry id="FA80D7D1307DFF94FC2DFC2DB9F50ECC" box="[984,1174,959,983]" name="Madagascar" pageId="4" pageNumber="205">MADAGASCAR</collectingCountry>
Ranomafana NP. Talatakely, X,
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<quantity id="456F3AA4307DFF94FE39FC4DBF6C0EED" box="[460,527,991,1015]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="205" unit="m" value="130.0">
<elevation id="09BA7072307DFF94FE39FC4DBF6C0EED" box="[460,527,991,1015]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="205" unit="m" value="130.0">130 m</elevation>
</quantity>
.
</specimenCount>
<date id="F629B181307DFF94FDEEFC4DBFE60EEC" box="[539,645,991,1015]" pageId="4" pageNumber="205" value="2001-01-15">
<collectingDate id="E66D4869307DFF94FDEEFC4DBFE60EEC" box="[539,645,991,1015]" pageId="4" pageNumber="205" value="2001-01-15">15.1.2001</collectingDate>
</date>
”/ “
<collectingMethod id="5BD6EF56307DFF94FD5FFC4DBEF90EED" box="[682,922,991,1015]" pageId="4" pageNumber="205">collected in pitfall trap</collectingMethod>
on forest floor,
<collectorName id="2F62F297307DFF94FBB7FC4DBDC2090C" pageId="4" pageNumber="205">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F297307DFF94FF3DFB92BC4A090C" box="[200,297,1023,1047]" pageId="4" pageNumber="205">J. Rainio</collectorName>
collectors”/ “32.” [handwritten label]/ “
</materialsCitation>
<typeStatus id="5D2C29E3307DFF94FD16FC6DBE13090C" box="[739,880,1023,1047]" pageId="4" pageNumber="205" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC2307DFF94FC8CFC6DBC3C092C" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B53307DFF94FC8CFC6DB9F6090C" box="[889,1173,1023,1047]" italics="true" pageId="4" pageNumber="205">Brachypelus ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628307DFF94FE93FBB2BCC3092C" box="[358,416,1056,1079]" pageId="4" pageNumber="205" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
</paragraph>
<paragraph id="82289741307DFF94FF5EFBD2B950094C" blockId="4.[123,1174,958,1726]" box="[171,1075,1088,1112]" pageId="4" pageNumber="205">
<materialsCitation id="32FF9D1C307DFF94FF5EFBD2B94F0943" box="[171,1068,1088,1112]" country="Madagascar" location="Ranomafana National Park" pageId="4" pageNumber="205" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B53307DFF94FF5EFBD2BC34094D" bold="true" box="[171,343,1088,1112]" pageId="4" pageNumber="205">
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<typeStatus id="5D2C29E3307DFF94FF5EFBD2BD87094D" box="[171,228,1088,1112]" pageId="4" pageNumber="205">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D307DFF94FF19FBD1BC34094D" baselines="1105" box="[236,343,1091,1110]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="4" pageNumber="205">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D1307DFF94FE8BFBD2BC9C0943" box="[382,511,1088,1112]" name="Madagascar" pageId="4" pageNumber="205">Madagascar</collectingCountry>
,
<collectingRegion id="405359A3307DFF94FDF8FBD2BF9F0943" box="[525,764,1088,1112]" country="Madagascar" name="Fianarantsoa" pageId="4" pageNumber="205">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A307DFF94FCFFFBD2B94F0943" LSID="urn:lsid:plazi:treatment:0A3E2657307DFF95FF8EFCD9BC8D0B62:8748C19A307DFF94FCFFFBD2B94F0943" box="[778,1068,1088,1112]" country="Madagascar" name="Ranomafana National Park" pageId="4" pageNumber="205" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="82289741307DFF94FF5EFBF2BECA0983" blockId="4.[123,1174,958,1726]" pageId="4" pageNumber="205">
<emphasis id="B0E34B53307DFF94FF5EFBF2BC91096D" bold="true" box="[171,498,1120,1144]" pageId="4" pageNumber="205">
<smallCapsWord id="84CE019D307DFF94FF5EFBF2BC4D096D" baselines="1138,1137" box="[171,302,1120,1144]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="4" pageNumber="205">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D307DFF94FEC3FBF1BC31096D" baselines="1137" box="[310,338,1123,1142]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="4" pageNumber="205">OF</smallCapsWord>
<smallCapsWord id="84CE019D307DFF94FEACFBF1BCCF096D" baselines="1137" box="[345,428,1123,1142]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="4" pageNumber="205">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D307DFF94FE40FBF1BC91096D" baselines="1137" box="[437,498,1123,1142]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="4" pageNumber="205">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC2307DFF94FD0CFBF2BEE40963" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[761,903,1120,1144]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53307DFF94FD0CFBF2BEE40963" box="[761,903,1120,1144]" italics="true" pageId="4" pageNumber="205">ranomafanae</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the national park in which the
<typeStatus id="5D2C29E3307DFF94FD29FB13BE690982" box="[732,778,1153,1177]" pageId="4" pageNumber="205">type</typeStatus>
was collected.
</paragraph>
<paragraph id="82289741307DFF95FF5EFB33BC8008B4" blockId="4.[123,1174,958,1726]" lastBlockId="5.[123,1174,1229,1658]" lastPageId="5" lastPageNumber="206" pageId="4" pageNumber="205">
<smallCapsWord id="84CE019D307DFF94FF5EFB33BC2109AC" baselines="1203,1202" box="[171,322,1185,1209]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="4" pageNumber="205">
<emphasis id="B0E34B53307DFF94FF5EFB33BC2109AC" bold="true" box="[171,322,1185,1209]" pageId="4" pageNumber="205">RECOGNITION</emphasis>
</smallCapsWord>
.— Large for genus, SBL =
<quantity id="456F3AA4307DFF94FD99FB33BFDE09A3" box="[620,701,1185,1209]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="4" pageNumber="205" unit="mm" value="6.8">6.8 mm</quantity>
. Members of this species (
<figureCitation id="1AAC8BC4307DFF94FC2FFB33B94809A2" box="[986,1067,1185,1209]" captionStart="FIGURE" captionStartId="5.[155,167,1152,1171]" captionTargetBox="[133,1174,245,1135]" captionTargetId="figure-997@5.[120,1174,216,1150]" captionTargetPageId="5" captionText="FIGURE 2. Digital images of holotype female of Brachypelus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799443" httpUri="https://zenodo.org/record/13799443/files/figure.png" pageId="4" pageNumber="205">Fig. 2A</figureCitation>
) are most similar to those of
<taxonomicName id="4597ECC2307DFF94FEB2FB53BFA509C2" authority="Putzeys (1866)" authorityName="Putzeys" authorityYear="1866" box="[327,710,1217,1241]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B0E34B53307DFF94FEB2FB53BF7E09C2" box="[327,541,1217,1241]" italics="true" pageId="4" pageNumber="205">Brachypelus obesus</emphasis>
<bibRefCitation id="E606EAB0307DFF94FDD1FB53BFA509C2" author="PUTZEYS, J. A. A. H." box="[548,710,1217,1241]" pageId="4" pageNumber="205" pagination="1 - 242" refId="ref32024" refString="PUTZEYS, J. A. A. H. 1866. Revision generale des clivinides. Annales de la Societe Entomologique de Belgique 10: 1 - 242." type="journal article" year="1866">Putzeys (1866)</bibRefCitation>
</taxonomicName>
, with which they share large size, only six setae on the labrum, and a single discal setiferous pore on elytral interval 3 (at apical one-fifth). They differ from
<taxonomicName id="4597ECC2307DFF94FEC1FA90BCF80802" authorityName="Putzeys" authorityYear="1866" box="[308,411,1281,1305]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B0E34B53307DFF94FEC1FA90BCF80802" box="[308,411,1281,1305]" italics="true" pageId="4" pageNumber="205">B. obesus</emphasis>
</taxonomicName>
members in the following: transverse fronto-clypeal suture present and deeply impressed (
<figureCitation id="1AAC8BC4307DFF94FEB3FAB0BCFB0821" box="[326,408,1314,1338]" captionStart="FIGURE" captionStartId="6.[155,167,1136,1155]" captionTargetBox="[306,980,229,1110]" captionTargetId="figure-647@6.[306,980,215,1110]" captionTargetPageId="6" captionText="FIGURE 3. Digital images of holotype female of Brachypelus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Head, dorsal aspect; B. Elytral base, dorsal aspect. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799447" httpUri="https://zenodo.org/record/13799447/files/figure.png" pageId="4" pageNumber="205">Fig. 3A</figureCitation>
) (absent from
<taxonomicName id="4597ECC2307DFF94FDCCFAB1BFC30821" authorityName="Putzeys" authorityYear="1866" box="[569,672,1314,1338]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B0E34B53307DFF94FDCCFAB1BFC30821" box="[569,672,1314,1338]" italics="true" pageId="4" pageNumber="205">B. obesus</emphasis>
</taxonomicName>
members); frons with longitudinal frontal furrows narrower and shallower than in
<taxonomicName id="4597ECC2307DFF94FDF6FAD1BF0A0841" authorityName="Putzeys" authorityYear="1866" box="[515,617,1346,1370]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B0E34B53307DFF94FDF6FAD1BF0A0841" box="[515,617,1346,1370]" italics="true" pageId="4" pageNumber="205">B. obesus</emphasis>
</taxonomicName>
; posterior lateral margins of the eyes broadly round- ed (more narrowly and angularly rounded in
<taxonomicName id="4597ECC2307DFF94FD9DFAF1BFB20861" authorityName="Putzeys" authorityYear="1866" box="[616,721,1378,1402]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B0E34B53307DFF94FD9DFAF1BFB20861" box="[616,721,1378,1402]" italics="true" pageId="4" pageNumber="205">B. obesus</emphasis>
</taxonomicName>
members); elytral base (
<figureCitation id="1AAC8BC4307DFF94FC17FAF0B9570861" box="[994,1076,1378,1403]" captionStart="FIGURE" captionStartId="6.[155,167,1136,1155]" captionTargetBox="[306,980,229,1110]" captionTargetId="figure-647@6.[306,980,215,1110]" captionTargetPageId="6" captionText="FIGURE 3. Digital images of holotype female of Brachypelus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Head, dorsal aspect; B. Elytral base, dorsal aspect. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799447" httpUri="https://zenodo.org/record/13799447/files/figure.png" pageId="4" pageNumber="205">Fig. 3B</figureCitation>
) straight between intervals 3 and obliquely sloped laterally from interval 4 (elytral base straight between intervals 4 and gradually rounded laterally from interval
<quantity id="456F3AA4307DFF94FD2BFA31BE6B08A0" box="[734,776,1443,1467]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="205" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="4597ECC2307DFF94FCFAFA36BE1508A0" authorityName="Putzeys" authorityYear="1866" box="[783,886,1443,1467]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B0E34B53307DFF94FCFAFA36BE1508A0" box="[783,886,1443,1467]" italics="true" pageId="4" pageNumber="205">B. obesus</emphasis>
</taxonomicName>
members); elytra with lateral margin distinctly toothed at humerus (smoothly rounded at humerus, without a tooth in
<taxonomicName id="4597ECC2307DFF94FBBCFA56BDFE08E0" authorityName="Putzeys" authorityYear="1866" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B0E34B53307DFF94FBBCFA56BDFE08E0" italics="true" pageId="4" pageNumber="205">B. obesus</emphasis>
</taxonomicName>
members); and elytral intervals slightly convex apically (nearly flat in
<taxonomicName id="4597ECC2307DFF94FC47FA76B97808E7" authorityName="Putzeys" authorityYear="1866" box="[946,1051,1508,1532]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B0E34B53307DFF94FC47FA76B97808E7" box="[946,1051,1508,1532]" italics="true" pageId="4" pageNumber="205">B. obesus</emphasis>
</taxonomicName>
members). Others species with members of comparably large size include
<taxonomicName id="4597ECC2307DFF94FCEAF996BDA10B27" authority="Bulirsch et al. (2005)" authorityName="Bulirsch" authorityYear="2005" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="rolandi">
<emphasis id="B0E34B53307DFF94FCEAF996BE9A0B07" box="[799,1017,1540,1564]" italics="true" pageId="4" pageNumber="205">Brachypelus rolandi</emphasis>
<bibRefCitation id="E606EAB0307DFF94FBF5F996BDA10B27" author="BULIRSCH, P. &amp; J. JANAK &amp; P. MORAVEC" pageId="4" pageNumber="205" pagination="1 - 35" refId="ref29568" refString="BULIRSCH, P., J. JANAK, AND P. MORAVEC. 2005. New species and findings of Scaritinae (Coleoptera: Carabidae) from Madagascar. Studies and Reports of District Museum Prague-East, Taxonomical Series 1: 1 - 35." type="book chapter" year="2005">Bulirsch et al. (2005)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4597ECC2307DFF94FF3AF9B6BFF40B27" authority="Bulirsch et al. (2005)" authorityName="Bulirsch" authorityYear="2005" box="[207,663,1572,1596]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="betsileo">
<emphasis id="B0E34B53307DFF94FF3AF9B6BCCD0B27" box="[207,430,1572,1596]" italics="true" pageId="4" pageNumber="205">Brachypelus betsileo</emphasis>
<bibRefCitation id="E606EAB0307DFF94FE41F9B6BFF40B27" author="BULIRSCH, P. &amp; J. JANAK &amp; P. MORAVEC" box="[436,663,1572,1596]" pageId="4" pageNumber="205" pagination="1 - 35" refId="ref29568" refString="BULIRSCH, P., J. JANAK, AND P. MORAVEC. 2005. New species and findings of Scaritinae (Coleoptera: Carabidae) from Madagascar. Studies and Reports of District Museum Prague-East, Taxonomical Series 1: 1 - 35." type="book chapter" year="2005">Bulirsch et al. (2005)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4597ECC2307DFF94FD51F9B6B90B0B27" authority="Bulirsch et al. (2005)" authorityName="Bulirsch" authorityYear="2005" box="[676,1128,1572,1596]" class="Insecta" family="Carabidae" genus="Brachypelas" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="fischeri">
<emphasis id="B0E34B53307DFF94FD51F9B6BEE30B27" box="[676,896,1572,1596]" italics="true" pageId="4" pageNumber="205">Brachypelas fischeri</emphasis>
<bibRefCitation id="E606EAB0307DFF94FC73F9B6B90B0B27" author="BULIRSCH, P. &amp; J. JANAK &amp; P. MORAVEC" box="[902,1128,1572,1596]" pageId="4" pageNumber="205" pagination="1 - 35" refId="ref29568" refString="BULIRSCH, P., J. JANAK, AND P. MORAVEC. 2005. New species and findings of Scaritinae (Coleoptera: Carabidae) from Madagascar. Studies and Reports of District Museum Prague-East, Taxonomical Series 1: 1 - 35." type="book chapter" year="2005">Bulirsch et al. (2005)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4597ECC2307DFF94FF8EF9D6BFE90B46" authority="Bulirsch and Moravec (2009)" authorityName="Bulirsch and Moravec" authorityYear="2009" box="[123,650,1604,1629]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="janaki">
<emphasis id="B0E34B53307DFF94FF8EF9D6BC290B47" box="[123,330,1604,1628]" italics="true" pageId="4" pageNumber="205">Brachypelus janaki</emphasis>
<bibRefCitation id="E606EAB0307DFF94FEA5F9D6BFE90B46" author="BULIRSCH, P. &amp; P. MORAVEC" box="[336,650,1604,1629]" pageId="4" pageNumber="205" pagination="27 - 36" refId="ref29614" refString="BULIRSCH, P., AND P. MORAVEC. 2009. New species and findings of Scaritinae (Coleoptera: Carabidae) from Madagascar: Part 2. Studies and Reports of District Museum Prague-East, Taxonomical Series 5: 27 - 36." type="book chapter" year="2009">Bulirsch and Moravec (2009)</bibRefCitation>
</taxonomicName>
. However,
<taxonomicName id="4597ECC2307DFF94FCF0F9D7BED20B46" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[773,945,1605,1629]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53307DFF94FCF0F9D7BED20B46" box="[773,945,1605,1629]" italics="true" pageId="4" pageNumber="205">B. ranomafanae</emphasis>
</taxonomicName>
members differ from those of
<taxonomicName id="4597ECC2307DFF94FF2DF9F4BC270B66" box="[216,324,1637,1661]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="rolandi">
<emphasis id="B0E34B53307DFF94FF2DF9F4BC270B66" box="[216,324,1637,1661]" italics="true" pageId="4" pageNumber="205">B. rolandi</emphasis>
</taxonomicName>
in having the labrum with only six setae (seven in
<taxonomicName id="4597ECC2307DFF94FC98F9F4BEBA0B66" box="[877,985,1637,1661]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="rolandi">
<emphasis id="B0E34B53307DFF94FC98F9F4BEBA0B66" box="[877,985,1637,1661]" italics="true" pageId="4" pageNumber="205">B. rolandi</emphasis>
</taxonomicName>
members), elytra widest at middle (anterior to middle in
<taxonomicName id="4597ECC2307DFF94FDEAF914BFE80B86" box="[543,651,1669,1693]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="rolandi">
<emphasis id="B0E34B53307DFF94FDEAF914BFE80B86" box="[543,651,1669,1693]" italics="true" pageId="4" pageNumber="205">B. rolandi</emphasis>
</taxonomicName>
members), and elytral intervals moderately convex throughout (flattened apical in
<taxonomicName id="4597ECC2307DFF94FE0FF934BF040BA6" box="[506,615,1701,1725]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="rolandi">
<emphasis id="B0E34B53307DFF94FE0FF934BF040BA6" box="[506,615,1701,1725]" italics="true" pageId="4" pageNumber="205">B. rolandi</emphasis>
</taxonomicName>
members). Members of both
<taxonomicName id="4597ECC2307DFF94FC5EF934B97E0BA6" box="[939,1053,1701,1725]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="betsileo">
<emphasis id="B0E34B53307DFF94FC5EF934B97E0BA6" box="[939,1053,1701,1725]" italics="true" pageId="4" pageNumber="205">B. betsileo</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC2307DFF95FBA1F934BDD109FE" class="Insecta" family="Carabidae" genus="Brachypelas" kingdom="Animalia" lastPageId="5" lastPageNumber="206" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="fischeri">
<emphasis id="B0E34B53307DFF95FBA1F934BDD109FE" italics="true" lastPageId="5" lastPageNumber="206" pageId="4" pageNumber="205">B. fischeri</emphasis>
</taxonomicName>
have seven labral setae (whereas those of
<taxonomicName id="4597ECC2307CFF95FD85FB5CBE7809FE" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[624,795,1229,1253]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53307CFF95FD85FB5CBE7809FE" box="[624,795,1229,1253]" italics="true" pageId="5" pageNumber="206">B. ranomafanae</emphasis>
</taxonomicName>
have only six) and lack discal setiferous pores (those of
<taxonomicName id="4597ECC2307CFF95FE94FB62BF6E081C" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[353,525,1263,1287]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53307CFF95FE94FB62BF6E081C" box="[353,525,1263,1287]" italics="true" pageId="5" pageNumber="206">B. ranomafanae</emphasis>
</taxonomicName>
have a single setiferous pore in the apical one-fifth on interval 3. Finally,
<taxonomicName id="4597ECC2307CFF95FEE3FA83BCA10832" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[278,450,1297,1321]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53307CFF95FEE3FA83BCA10832" box="[278,450,1297,1321]" italics="true" pageId="5" pageNumber="206">B. ranomafanae</emphasis>
</taxonomicName>
members differ from those of
<taxonomicName id="4597ECC2307CFF95FCF8FA83BE0D0832" authorityName="Bulirsch and Moravec" authorityYear="2009" box="[781,878,1297,1321]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="janaki">
<emphasis id="B0E34B53307CFF95FCF8FA83BE0D0832" box="[781,878,1297,1321]" italics="true" pageId="5" pageNumber="206">B. janaki</emphasis>
</taxonomicName>
in having only six setae on the labrum (seven labral setae in
<taxonomicName id="4597ECC2307CFF95FE1CFAA1BF2F0851" authorityName="Bulirsch and Moravec" authorityYear="2009" box="[489,588,1330,1354]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="janaki">
<emphasis id="B0E34B53307CFF95FE1CFAA1BF2F0851" box="[489,588,1330,1354]" italics="true" pageId="5" pageNumber="206">B. janaki</emphasis>
</taxonomicName>
members), the elytral margin with a distinct humeral tooth (tooth absent from
<taxonomicName id="4597ECC2307CFF95FE7BFAC7BC910877" authorityName="Bulirsch and Moravec" authorityYear="2009" box="[398,498,1364,1388]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="janaki">
<emphasis id="B0E34B53307CFF95FE7BFAC7BC910877" box="[398,498,1364,1388]" italics="true" pageId="5" pageNumber="206">B. janaki</emphasis>
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members), and only a single discal setiferous pore on elytral interval 3, with intervals 5 and 7 without such punctures (five or more such punctures on intervals 3, 5 and
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members).
</paragraph>
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2. Digital images of holotype female of
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<emphasis id="B0E34B53307CFF95FDCBFB12BE410988" box="[574,802,1152,1171]" italics="true" pageId="5" pageNumber="206">Brachypelus ranomafanae</emphasis>
Kavanaugh &amp; Rainio
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A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm.
</paragraph>
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.— At present, known only from the
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locality.
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</emphasis>
.— The unique
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was collected in a pitfall trap placed in secondary montane rain forest in the Talatakely area, at an elevation of
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, at the
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mark along Trail “X”. Vegetation in that area consisted of bamboo, small young trees and some larger trees with diameters at breast-height (DBH) of
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. Tree trunks and branches were mostly covered with epiphytes and lianas.
</paragraph>
</subSubSection>
</treatment>
</document>

234
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<mods:title id="E426337213571C5CB8AA0A47D23779B5">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
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<mods:affiliation id="6B2C06F0879272E3CD8D2D5487C291A2">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
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<emphasis id="B0E34B53307FFF96FF8EFA4EBC2708EF" bold="true" box="[123,324,1500,1524]" italics="true" pageId="6" pageNumber="207">Caelostomus latus</emphasis>
Kavanaugh and Rainio
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,
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.—
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(
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), a female, in CAS, labeled: “
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Ranomafana NP. Talatakely, Trail C,
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.
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”/ “
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Kavanaugh &amp; Rainio
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2015” [red label].
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.—
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,
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,
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.
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4. Digital images of holotype female of
<taxonomicName id="4597ECC2307EFF97FDB1FB71BEC709ED" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[580,932,1251,1270]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
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Kavanaugh &amp; Rainio
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A. Habitus, dorsal aspect; B. Labels; C. Head, left lateral aspect (arrow points to tranverse depression between eyes); D. Right anterior tarsus, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm.
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</caption>
<paragraph id="82289741307EFF97FF5EFAD7BCA20886" blockId="7.[123,1174,1349,1726]" pageId="7" pageNumber="208">
<emphasis id="B0E34B53307EFF97FF5EFAD7BC940847" bold="true" box="[171,503,1349,1373]" pageId="7" pageNumber="208">
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<smallCapsWord id="84CE019D307EFF97FEC2FADABC300840" baselines="1366" box="[311,339,1352,1371]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="7" pageNumber="208">OF</smallCapsWord>
<smallCapsWord id="84CE019D307EFF97FEA9FADABCCC0840" baselines="1366" box="[348,431,1352,1371]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="7" pageNumber="208">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D307EFF97FE4FFADBBC940847" baselines="1367" box="[442,503,1353,1372]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="7" pageNumber="208">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC2307EFF97FCF7FAD7BE550846" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[770,822,1349,1373]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B0E34B53307EFF97FCF7FAD7BE550846" box="[770,822,1349,1373]" italics="true" pageId="7" pageNumber="208">latus</emphasis>
</taxonomicName>
, is the Latin adjective meaning broad or wide, a reference to relatively broad pronotum of the
<typeStatus id="5D2C29E3307EFF97FCE7FAF7BE0C0866" box="[786,879,1381,1405]" pageId="7" pageNumber="208" type="holotype">holotype</typeStatus>
specimen and, presumably, other members of this species.
</paragraph>
<paragraph id="82289741307EFF98FF5EFA37BF3B0F4F" blockId="7.[123,1174,1349,1726]" lastBlockId="8.[123,1174,219,693]" lastPageId="8" lastPageNumber="209" pageId="7" pageNumber="208">
<smallCapsWord id="84CE019D307EFF97FF5EFA37BC2108A0" baselines="1463,1462" box="[171,322,1445,1469]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="7" pageNumber="208">
<emphasis id="B0E34B53307EFF97FF5EFA37BC2108A0" bold="true" box="[171,322,1445,1469]" pageId="7" pageNumber="208">RECOGNITION</emphasis>
</smallCapsWord>
.— Size moderate for genus, SBL =
<quantity id="456F3AA4307EFF97FD31FA37BE7708A6" box="[708,788,1445,1469]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="7" pageNumber="208" unit="mm" value="6.3">6.3 mm</quantity>
. Based on Jeannel’s (1948) key, this species is a member of subgenus
<taxonomicName id="4597ECC2307EFF97FE1EFA57BF1608C6" authorityName="MacLeay" authorityYear="1825" box="[491,629,1477,1501]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B0E34B53307EFF97FE1EFA57BF1608C6" box="[491,629,1477,1501]" italics="true" pageId="7" pageNumber="208">Caelostomus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABD0F628307EFF97FD8BFA54BFDB08C6" box="[638,696,1478,1501]" pageId="7" pageNumber="208" sensu="stricto">s. str.</taxonomicNameLabel>
It’s members (
<figureCitation id="1AAC8BC4307EFF97FCAAFA57BED008C6" box="[863,947,1477,1501]" captionStart="FIGURE" captionStartId="7.[155,167,1251,1270]" captionTargetBox="[120,1176,216,1233]" captionTargetId="figure-985@7.[120,1176,216,1235]" captionTargetPageId="7" captionText="FIGURE 4. Digital images of holotype female of Caelostomus latus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Head, left lateral aspect (arrow points to tranverse depression between eyes); D. Right anterior tarsus, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799449" httpUri="https://zenodo.org/record/13799449/files/figure.png" pageId="7" pageNumber="208">Fig. 4A</figureCitation>
) share with those of other members of the subgenus the following features: head with frons and frontal furrows smooth, impunctate; elytral base distinctly margined from humerus medially to base of stria 3 or 2; elytral striae 6 and 7 deeply impressed throughout, not effaced apically; lateral elytral intervals narrow, moderately convex; apical part of umbilicate series of setiferous pores not inserted in a deep groove; and female apical abdominal ventrite with (two or) three pairs of setae subapically. Within the subgenus, the following features are shared only with members of two species of Jeannel’s
<taxonomicName id="4597ECC2307EFF97FF8EF937BC7F0BA6" authorityName="Tschitscherine" authorityYear="1899" box="[123,284,1701,1725]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="convexiusculus">
<emphasis id="B0E34B53307EFF97FF8EF937BC7F0BA6" box="[123,284,1701,1725]" italics="true" pageId="7" pageNumber="208">convexiusculus</emphasis>
</taxonomicName>
group, namely
<taxonomicName id="4597ECC2307EFF97FE39F937BE970BA6" authority="Tschitscherine (1899)" authorityName="Tschitscherine" authorityYear="1899" box="[460,1012,1701,1725]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="convexiusculus">
<emphasis id="B0E34B53307EFF97FE39F937BE630BA6" box="[460,768,1701,1725]" italics="true" pageId="7" pageNumber="208">Caelostomus convexiusculus</emphasis>
<bibRefCitation id="E606EAB0307EFF97FCFFF937BE970BA6" author="TSCHITSCHERINE, T. S." box="[778,1012,1701,1725]" pageId="7" pageNumber="208" pagination="50 - 82" refId="ref32397" refString="TSCHITSCHERINE, T. S. 1899. Memoire sur qulques Platysmatini nouveaux ou peu connus d'Afrique et de Madagascar. Annales de la Societe Entomologique de France 68: 50 - 82." type="journal article" year="1899">Tschitschérine (1899)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4597ECC2307EFF98FBDBF937BCD70DE8" authority="Jeannel (1948)" authorityName="Jeannel" authorityYear="1948" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" lastPageId="8" lastPageNumber="209" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B0E34B53307EFF98FBDBF937BC6A0DE8" italics="true" lastPageId="8" lastPageNumber="209" pageId="7" pageNumber="208">Caelostomus alluaudi</emphasis>
Jeannel (1948)
</taxonomicName>
: SBL greater than 6.0 mm, elytra short and broad, ratio EL/EW = 1.19, elytral disc convex; and elytral intervals distinctly punctate throughout, including in apical part. The
<typeStatus id="5D2C29E33071FF98FF15FE89BC5E0C28" box="[224,317,283,307]" pageId="8" pageNumber="209" type="holotype">holotype</typeStatus>
female of
<taxonomicName id="4597ECC23071FF98FE5BFE8EBF620C28" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[430,513,283,307]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B0E34B533071FF98FE5BFE8EBF620C28" box="[430,513,283,307]" italics="true" pageId="8" pageNumber="209">C. latus</emphasis>
</taxonomicName>
is slightly larger than members of these other two species, has a distinct and broad transverse depression between the eyes seen best in lateral view (
<figureCitation id="1AAC8BC43071FF98FBCCFEA9B9EE0C4F" box="[1081,1165,315,340]" captionStart="FIGURE" captionStartId="7.[155,167,1251,1270]" captionTargetBox="[120,1176,216,1233]" captionTargetId="figure-985@7.[120,1176,216,1235]" captionTargetPageId="7" captionText="FIGURE 4. Digital images of holotype female of Caelostomus latus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Head, left lateral aspect (arrow points to tranverse depression between eyes); D. Right anterior tarsus, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799449" httpUri="https://zenodo.org/record/13799449/files/figure.png" pageId="8" pageNumber="209">Fig. 4C</figureCitation>
) (depression absent from members of the other two species). The pronotum in the
<typeStatus id="5D2C29E33071FF98FBE2FECEB9170C6F" box="[1047,1140,348,372]" pageId="8" pageNumber="209" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4597ECC23071FF98FF8EFEEEBDAE0C8F" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,205,380,404]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B0E34B533071FF98FF8EFEEEBDAE0C8F" box="[123,205,380,404]" italics="true" pageId="8" pageNumber="209">C. latus</emphasis>
</taxonomicName>
is much wider than in
<taxonomicName id="4597ECC23071FF98FE4BFEEEBF1E0C8F" authorityName="Tschitscherine" authorityYear="1899" box="[446,637,380,404]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="convexiusculus">
<emphasis id="B0E34B533071FF98FE4BFEEEBF1E0C8F" box="[446,637,380,404]" italics="true" pageId="8" pageNumber="209">C. convexiusculus</emphasis>
</taxonomicName>
members, slightly wider than in
<taxonomicName id="4597ECC23071FF98FC2FFEEEB9320C8F" authorityName="Jeannel" authorityYear="1948" box="[986,1105,380,404]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B0E34B533071FF98FC2FFEEEB9320C8F" box="[986,1105,380,404]" italics="true" pageId="8" pageNumber="209">C. alluaudi</emphasis>
</taxonomicName>
members, and less depressed anterolaterally than in members of either of the other species; and the pronotal basal foveae are straighter and more sharply defined laterally than in members of the other two species. The elytra are less abruptly rounded laterally and apically than in
<taxonomicName id="4597ECC23071FF98FC20FE4FB92C0CEF" authorityName="Jeannel" authorityYear="1948" box="[981,1103,476,500]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B0E34B533071FF98FC20FE4FB92C0CEF" box="[981,1103,476,500]" italics="true" pageId="8" pageNumber="209">C. alluaudi</emphasis>
</taxonomicName>
members but similar to shape in to those in
<taxonomicName id="4597ECC23071FF98FDDBFE6FBF920F0F" authorityName="Tschitscherine" authorityYear="1899" box="[558,753,508,532]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="convexiusculus">
<emphasis id="B0E34B533071FF98FDDBFE6FBF920F0F" box="[558,753,508,532]" italics="true" pageId="8" pageNumber="209">C. convexiusculus</emphasis>
</taxonomicName>
members. Finally, anterior tarsomeres 1 to 3 of the female
<typeStatus id="5D2C29E33071FF98FEACFD8EBCD50F2F" box="[345,438,540,564]" pageId="8" pageNumber="209" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4597ECC23071FF98FE28FD8FBF520F2F" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[477,561,540,564]" class="Insecta" family="Carabidae" genus="Caelostomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B0E34B533071FF98FE28FD8FBF520F2F" box="[477,561,540,564]" italics="true" pageId="8" pageNumber="209">C. latus</emphasis>
</taxonomicName>
are distinctly toothed medioapically (
<figureCitation id="1AAC8BC43071FF98FC38FD8EB9430F2F" box="[973,1056,540,565]" captionStart="FIGURE" captionStartId="7.[155,167,1251,1270]" captionTargetBox="[120,1176,216,1233]" captionTargetId="figure-985@7.[120,1176,216,1235]" captionTargetPageId="7" captionText="FIGURE 4. Digital images of holotype female of Caelostomus latus Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Head, left lateral aspect (arrow points to tranverse depression between eyes); D. Right anterior tarsus, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799449" httpUri="https://zenodo.org/record/13799449/files/figure.png" pageId="8" pageNumber="209">Fig. 4D</figureCitation>
), a feature not seen in females of the other two species.
</paragraph>
<paragraph id="822897413071FF98FF5EFDCFB9690F6F" blockId="8.[123,1174,219,693]" box="[171,1034,605,629]" pageId="8" pageNumber="209">
<emphasis id="B0E34B533071FF98FF5EFDCFBC920F68" bold="true" box="[171,497,605,629]" pageId="8" pageNumber="209">
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<smallCapsWord id="84CE019D3071FF98FEA8FDF2BC920F68" baselines="622" box="[349,497,608,627]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="8" pageNumber="209">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E33071FF98FC8AFDCFBECE0F6E" box="[895,941,605,629]" pageId="8" pageNumber="209">type</typeStatus>
locality.
</paragraph>
<paragraph id="822897413071FF98FF5EFDEFBF6C0FAE" blockId="8.[123,1174,219,693]" pageId="8" pageNumber="209">
<emphasis id="B0E34B533071FF98FF5EFDEFBCC50F88" bold="true" box="[171,422,637,661]" pageId="8" pageNumber="209">
<smallCapsWord id="84CE019D3071FF98FF5EFDEFBC6A0F88" baselines="655,654" box="[171,265,637,661]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="8" pageNumber="209">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D3071FF98FEE7FD12BCC50F88" baselines="654" box="[274,422,640,659]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="8" pageNumber="209">DISTRIBUTION</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C3071FF98FE3BFDEFBF640FAE" location="Talatakely area" pageId="8" pageNumber="209" specimenCount="1" typeStatus="holotype">
The unique
<typeStatus id="5D2C29E33071FF98FDA6FDEFBFD30F8E" box="[595,688,637,661]" pageId="8" pageNumber="209" type="holotype">holotype</typeStatus>
female was collected by hand in secondary montane forest in the
<location id="8748C19A3071FF98FE90FD0FBF640FAE" LSID="urn:lsid:plazi:treatment:0A3E2657307FFF98FF8EFA4EBF6C0FAE:8748C19A3071FF98FE90FD0FBF640FAE" box="[357,519,669,693]" name="Talatakely area" pageId="8" pageNumber="209">Talatakely area</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>